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Modelling the response of sandy beaches to sea level rise is a major scientific challenge and several types of models can be applied. Process-based models are useful tools for understanding beach responses on short time scales, but they are computationally expensive and tend to accumulate errors when resolving the many short-term processes they contain. Alternatively, reduced-complexity models can be an interesting option for long-term modelling. Furthermore, to simulate the long-term response of beaches, it is necessary to combine different model forcing sources (wave and sea-level, e.g., from buoys or hindcast models). The aim of this contribution is to quantify the effect of using different forcing sources in morphodynamic modelling with these model types.Two numerical models are applied to the embayed microtidal beach of El Castell (Palamós, Catalunya, NW Mediterranean). The XBeach process-based model, which solves the full 2DH nearshore hydrodynamics and the corresponding bed evolution, and the Q2Dmorfo reduced-complexity model, which simulates the bed level variations by calculating the sediment fluxes parametrically directly from the wave field without resolving the currents. Both models are first calibrated using two topobathymetric surveys conducted in January and July 2020 and wave and sea-level data measured from an AWAC deployed at 14.5 m depth during those 6 months. Calibration is performed using the most sensitive parameters, i.e., those related to cross-shore transport. In XBeach, the surfbeat mode must be used to obtain realistic results. It generates an aleatory spectral wave time series at the boundary that includes groupiness. To handle the effect of this randomness, a total of 5 realizations are made for each set of parameter values and a mean bathymetry is computed out of these realizations. To assess the model performance, the Brier Skill Score (BSS) is calculated both for the modelled bathymetry and its coastline during the 6-month period. Moreover, the Standard deviation (STD) of the 5 realizations is also computed. The chosen optimum parameter setting is the one maximizing the BSS (0.36 and 0.79 for the bathymetry and shoreline respectively) and minimizing the STD so that the result is robust and reproducible. In Q2Dmorfo, the BSS of the simulated final coastline is calculated for each set of parameter values. The optimum parameter setting also produces a maximum BSS of 0.79.Once both models are calibrated, the other potential forcing sources are applied. They include wave and sea-level datasets from a sea-level and wind-waves 72-years hindcast generated with the hydrodynamic-wave coupled SCHISM model, a wave dataset from an offshore buoy propagated to the AWAC position using SWAN model and a sea-level dataset measured by the tidal gauge at Barcelona harbour. The results show a very significant sensitivity to the wave forcing source and much less sensitivity to the sea-level source. By using the dataset propagated from the buoy by SWAN, both models represent well the observed beach rotation, whereas using the dataset obtained with SCHISM, the beach rotation is systematically under-predicted by both models, giving negative BSS values. This is because SCHISM predicts wave angles biased to the west.

This paper investigates concentric traveling ionospheric disturbances (CTIDs) associated with the Tonga volcanic eruption. Results show that: (a) two types of CTIDs (CTID #1 and CTID #2) were identified that traveled radially from Tonga at the speed of 610–880 m/s (acoustic-mode) and 300–380 m/s (Lamb-mode), respectively. CTID #1 reached 3,800 and 5,000 km away from the eruption location toward the directions of New Zealand and Australia, respectively. CTID #2 propagated persistently for ∼9 hr over New Zealand and Australia. (b) The CTID #2 wavefront changed after 08:35 UT over New Zealand, possibly due to a combination of factors including the anisotropic propagation of CTID #2, the regional geomagnetic declination, and westward-moving Lamb waves. (c) Topside total electron content (TEC) enhancement with a magnitude over two TECu was observed from COSMIC-2 measurements. The enhancement agrees with CTID #1 peak from nearby ground-based TEC observations and could be related to the upward propagation of the F layer’s CTID #1 signatures.

11 pages, 4 figures, 1 table, supplementary data https://doi.org/10.1016/j.envres.2022.113549, Sex ratio depends on sex determination mechanisms and is a key demographic parameter determining population viability and resilience to natural and anthropogenic stressors. There is increasing evidence that the environment can alter sex ratio even in genetically sex-determined species (GSD), as elevated temperature can cause female-to-male sex reversal (neomales). Alarmingly, neomales are being discovered in natural populations of several fish, amphibian and reptile species worldwide. Understanding the basis of neomale development is important for conservation biology. Among GSD species, it is unknown whether those with chromosomal sex determination (CSD), the most common system, will better resist the influence of high temperature than those with polygenic sex determination (PSD). Here, we compared the effects of elevated temperature in two wild zebrafish strains, Nadia (NA) and Ekkwill (EKW), which have CSD with a ZZ/ZW system, against the AB laboratory strain, which has PSD. First, we uncovered novel sex genotypes and the results showed that, at control temperature, the masculinization rate roughly doubled with the addition of each Z chromosome, while some ZW and WW fish of the wild strains became neomales. Surprisingly, we found that at elevated temperatures WW fish were just as likely as ZW fish to become neomales and that all strains were equally susceptible to masculinization. These results demonstrate that the Z chromosome is not essential for male development and that the dose of W buffers masculinization at the control temperature but not at elevated temperature. Furthermore, at the elevated temperature the testes of neomales, but not of normal males, contained more spermatozoa than at the control temperature. Our results show in an unprecedented way that, in a global warming scenario, CSD species may not necessarily be better protected against the masculinizing effect of elevated temperature than PSD species, and reveal genotype-by-temperature interactions in male sex determination and spermatogenesis, This study was supported by the Spanish Ministry of Science and Innovation grant AGL2016-787107-R “Epimark” to FP and NIH grants R01 GM085318 and 5R35 GM139635 to JHP. With funding from the Spanish government through the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S). AV was supported by Spanish government scholarships (BES-2014-069051) with two short-term stay grants (EEBB-I-18-12947 and EEBB-I-17-12342) performed in the JHP lab. LR was supported by RYC2018-024017-I. RHN and MSR were supported by São Paulo Research Foundation (FAPESP 14/07620–7 and 17/15793–7)

A set of historical tide gauge sea‐level records from two locations in Santander (Northern Spain) and Alicante (Spanish Mediterranean coast) have been recovered from logbooks stored in national archives. Sea‐level measurements have been digitized, quality‐controlled and merged into three consistent sea‐level time series (two in Alicante and one in Santander) using high‐precision levelling information. The historical sea‐level record in Santander consists of a daily time series spanning the period 1876‐1924 and it is further connected to the record from the modern tide gauge station nearby, ensuring datum continuity up to the present. The sea‐level recording in Alicante started in 1870, with daily averaged values until the 1920s and hourly afterwards, and the tide gauges at the two Alicante sites are still operating, thereby providing one of the longest tide gauge sea‐level time series in the Mediterranean Sea. The long‐term consistency and reliability of the new records are discussed based on comparison with nearby tide gauge time series.

Every year the Caribbean Sea faces the passage of several tropical cyclones that generate coastal extreme sea levels with potential strong and hazardous impacts. In this work we simulate the storm surges and wind waves induced by a set of 1000 tropical cyclones over the Caribbean Sea that are representative of the present-day climate. These events have been extracted from a global database of synthetic tropical cyclones spanning a 10 000-year period. The atmospheric forcing fields, associated with the set of tropical cyclones, are used to feed a coupled hydrodynamic–wave model with high resolution (∼ 2 km) along the continental and island coasts. Given the large number of events modelled, our results allow detailed statistical analyses of the magnitude and mechanisms of coastal extreme sea levels, as well as the identification of one of the most exposed areas to both storm surges and extreme wind waves.

On 15 January 2022, around 4:30 UTC the eruption of the Hunga-Tonga volcano, in the South Pacific Ocean, generated a violent underwater explosion. In addition to tsunami waves that affected the Pacific coasts, the eruption created atmospheric pressure disturbances that spread out in the form of Lamb waves. The associated atmospheric pressure oscillations were detected in high-frequency in-situ observations all over the globe. Here we take advantage of the similarities in the propagation and characteristics between atmospheric Lamb waves and long ocean waves and we use a 2DH ocean numerical model to simulate the phenomenon. We compare the outputs of the numerical simulation with in-situ atmospheric pressure records and with remote satellite observations. The signal in the model matches the observed atmospheric pressure perturbations and reveals an excellent agreement in the wave arrival time between model and observations at hundreds of locations at different distances from the origin., This study was supported by the MOCCA project Grant RTI2018-093941-B-C31 funded by MCIN/AEI/ 10.13039/501100011033 and by "ERDF A way of making Europe". It was also supported by grants PGC2018-099285-B-C21 and PGC2018-099285-B-C22 funded by MCIN/AEI/10.13039/501100011033 and by “ERDF A way of making Europe” NextGenerationEU/PRTR. Angel Amores was funded by the Conselleria d’Educació, Universitat i Recerca del Govern Balear through the Direcció General de Política Universitària i Recerca and by the Fondo Social Europeo for the period 2014–2020 (Grant No. PD/011/2019). Daniel Argüeso was funded by Spanish Ministry of Science and Innovation through the EPICC Project (PID2019-105253RJ-I00) and the Beatriz Galindo Programme (BG20/00078).

In the Mediterranean Sea, coastal extreme sea levels are mainly caused by storm surges driven by atmospheric pressure and surface winds from extratropical cyclones. In addition, wind-waves generated by the same atmospheric perturbations may also contribute to coastal extremes through wave setup (a temporary rise in mean sea level due to waves breaking close to shore). This study investigates the spatial and temporal variability of coastal extreme sea levels within the Mediterranean basin, using a new ocean hindcast generated with a coupled hydrodynamic-wave model that simulates storm surges and wind-waves generation and propagation. The numerical simulation spans the period 1950-2020 and is run with high temporal sampling (1h) and at unprecedented spatial resolution that reaches 200 m along the coastlines. Coastal storm surges and wave heights have been extensively validated with available observations (tide gauges and waves buoys). Comparison to tide gauges shows an average RMSE of 0.05m (0.08m for extreme events) and linear correlation of 0.75 for the period covering 1980-2020. Similarly, comparison of simulated and observed significant wave height shows good agreement with RMSE lower than 0.25 m and coefficient correlation as high as 0.95. Preliminary results show that coastal extreme sea levels found are more likely to be located in regions with wide and shallow continental shelves, where mild slope favour the combined effects of wind and wave setup. The contribution of waves to coastal extreme sea levels has been explored with an uncoupled simulation and has been shown to be significant, reaching up to 120% of observed maximum sea levels.

Twenty-first century projections for the Mediterranean water properties have been analyzed using the largest ensemble of regional climate models (RCMs) available up to now, the Med-CORDEX ensemble. It is comprised by 25 simulations, 10 historical and 15 scenario projections, from which 11 are ocean–atmosphere coupled runs and 4 are ocean forced simulations. Three different emissions scenarios are considered: RCP8.5, RCP4.5 and RCP2.6. All the simulations agree in projecting a warming across the entire Mediterranean basin by the end of the century as a result of the decrease of heat losses to the atmosphere through the sea surface and an increase in the net heat input through the Strait of Gibraltar. The warming will affect the whole water column with higher anomalies in the upper layer. The temperature change projected by the end of the century ranges between 0.81 and 3.71 °C in the upper layer (0–150 m), between 0.82 and 2.97 °C in the intermediate layer (150–600 m) and between 0.15 and 0.18 °C in the deep layer (600 m—bottom). The intensity of the warming is strongly dependent on the choice of emission scenario and, in second order, on the choice of Global Circulation Model (GCM) used to force the RCM. On the other hand, the local structures reproduced by each simulation are mainly determined by the regional model and not by the scenario or the global model. The salinity also increases in all the simulation due to the increase of the freshwater deficit (i.e. the excess of evaporation over precipitation and river runoff) and the related increase in the net salt transport at the Gibraltar Strait. However, in the upper layer this process can be damped or enhanced depending upon the characteristics of the inflowing waters from the Atlantic. This, in turn, depends on the evolution of salinity in the Northeast Atlantic projected by the GCM. Thus a clear zonal gradient is found in most simulations with large positive salinity anomalies in the eastern basin and a freshening of the upper layer of the western basin in most simulations. The salinity changes projected for the whole basin range between 0 and 0.34 psu in the upper layer, between 0.08 and 0.37 psu in the intermediate layer and between − 0.05 and 0.33 in the deep layer. These changes in the temperature and salinity modify in turn the characteristics of the main water masses as the new waters become saltier, warmer and less dense along the twenty-first century. There is a model consensus that the intensity of the deep water formation in the Gulf of Lions is expected to decrease in the future. The rate of decrease remains however very uncertain depending on the scenario and model chosen. At the contrary, there is no model consensus concerning the change in the intensity of the deep water formation in the Adriatic Sea and in the Aegean Sea, although most models also point to a reduction., This study was funded by the Spanish Ministerio de Ciencia, Innovación y Universidades funded CLIFISH Project (CTM2015-66400-C3-2-R).

Medicanes, for Mediterranean hurricanes, are mesoscale cyclones with morphological and physical characteristics similar to tropical cyclones. Although less intense, smaller, and rarer than their Atlantic counterparts, medicanes are very hazardous events threatening islands and continental coasts within the Mediterranean Sea. The latest strong episode, Medicane Ianos (September 2020), resulted in severe damages in Greece and several casualties. This work investigates the oceanic response to these extreme events along the Mediterranean coasts under present-day and future (late 21st century) climate conditions. To this end, a coupled hydrodynamic-wave model is used to simulate both storm surges and wind-waves generation and propagation in the Mediterranean Sea at high resolution (∼2 km) along the coastlines. A data set of thousands of medicanes synthetically generated from 20 global climate models and two atmospheric reanalyses is used to derive the atmospheric forcing fields. Regional coastal hazards assessment is performed for the present and future climates. For the first period, highest medicane-induced waves are found in the central and the southwest part of the western Mediterranean, while greatest storm surges are found in the Adriatic Sea and regions characterized by wide and gently sloping continental shelves. Results obtained for future changes show amplitudes generally smaller than the associated uncertainty due to limited agreement among models (especially for coastal elevation). Though, model consensus is reached (60–75%) and relative intensity change is significant (10–20%) at some locations (e.g., 1 m increase of medicane-induced significant wave height on average for south coasts of Sicilia)., This study was supported by the Project RTI2018-093941-B-C31 funded by MCIN/AEI/10.13039/501100011033/and by FEDER Una manera de hacer Europa. Tim Toomey acknowledges an FPI grant associated with the MOCCA project (Grant No. PRE2019-088046). Angel Amores was funded by the Conselleria d’Educació, Universitat i Recerca del Govern Balear through the Direcció General de Política Universitària i Recerca and by the Fondo Social Europeo for the period 2014–2020 (Grant No. PD/011/2019). Marta Marcos was supported by the Ministerio de Ciencia e Innovación and la Agencia Estatal de Investigación through Grant No. IED2019-000985-I. Romu Romero was funded by FEDER/Ministerio de Ciencia, Innovación y Universidades—Agencia Estatal de Investigación through the COASTEPS project (Grant No. CGL2017-82868-R).

Made available in DSpace on 2021-06-25T11:54:14Z (GMT). No. of bitstreams: 0 Previous issue date: 2021-03-09 National Institute of Health (USA) Deutsche Forschungsgemeinschaft National Science Foundation (USA) Office of Polar Programs Agence Nationale de la Recherche Schweizerischer Nationalfonds zur Forderung der Wissenschaftlichen Forschung The study of sex determination and sex chromosome organization in nonmodel species has long been technically challenging, but new sequencing methodologies now enable precise and high-throughput identification of sex-specific genomic sequences. In particular, restriction site-associated DNA sequencing (RAD-Seq) is being extensively applied to explore sex determination systems in many plant and animal species. However, software specifically designed to search for and visualize sex-biased markers using RAD-Seq data is lacking. Here, we present RADSex, a computational analysis workflow designed to study the genetic basis of sex determination using RAD-Seq data. RADSex is simple to use, requires few computational resources, makes no prior assumptions about the type of sex-determination system or structure of the sex locus, and offers convenient visualization through a dedicated R package. To demonstrate the functionality of RADSex, we re-analysed a published data set of Japanese medaka, Oryzias latipes, where we uncovered a previously unknown Y chromosome polymorphism. We then used RADSex to analyse new RAD-Seq data sets from 15 fish species spanning multiple taxonomic orders. We identified the sex determination system and sex-specific markers in six of these species, five of which had no known sex-markers prior to this study. We show that RADSex greatly facilitates the study of sex determination systems in nonmodel species thanks to its speed of analyses, low resource usage, ease of application and visualization options. Furthermore, our analysis of new data sets from 15 species provides new insights on sex determination in fish. IINRAE, LPGP, Rennes, France Univ Lausanne, Dept Ecol & Evolut, Lausanne, Switzerland Swiss Inst Bioinformat, Lausanne, Switzerland Hunan Normal Univ, Coll Life Sci, State Key Lab Dev Biol Freshwater Fish, Changsha, Peoples R China Uppsala Univ, Dept Organismal Biol, Systemat Biol, Uppsala, Sweden Univ Montpellier, Inst Genom Fonct, INSERM, IGF,CNRS, Montpellier, France INRAE, Math & Informat Appl Toulouse, SIGENAE, Castanet Tolosan, France Univ Wurzburg, Physiol Chem, Bioctr, Wurzburg, Germany Texas State Univ, Xiphophorus Genet Stock Ctr, Dept Chem & Biochem, San Marcos, TX USA Univ Wurzburg, Dev Biochem, Bioctr, Wurzburg, Germany Univ Oregon, Inst Neurosci, Eugene, OR USA NOAA, Environm & Fisheries Sci Div, Northwest Fisheries Sci Ctr, Natl Marine Fisheries Serv, Seattle, WA USA Sophia Univ, Fac Sci & Technol, Dept Mat & Life Sci, Tokyo, Japan Northeastern Univ, Ctr Marine Sci, Dept Marine & Environm Sci, Nahant, MA 01908 USA Sao Paulo State Univ, Inst Biosci, Dept Struct & Funct Biol, Reprod & Mol Biol Grp, Botucatu, SP, Brazil Tokyo Univ Marine Sci & Technol, Dept Aquat Marine Biosci, Tokyo, Japan Inst Marine Res, Bergen, Norway Chinese Acad Sci, Inst Hydrobiol, Beijing, Peoples R China IGB, Leibniz Inst Freshwater Ecol & Inland Fishe, Berlin, Germany Michigan State Univ, Dept Integrat Biol, Ecol Evolut & Behav Program, E Lansing, MI 48824 USA Sao Paulo State Univ, Inst Biosci, Dept Struct & Funct Biol, Reprod & Mol Biol Grp, Botucatu, SP, Brazil National Institute of Health (USA): R01GM085318 National Institute of Health (USA): R35GM139635 Deutsche Forschungsgemeinschaft: SCHA 408/10-1 National Science Foundation (USA) Office of Polar Programs: OPP-1543383 National Science Foundation (USA) Office of Polar Programs: PLR-1247510 National Science Foundation (USA) Office of Polar Programs: PLR-1444167 Agence Nationale de la Recherche: SCHA 408/10-1 Agence Nationale de la Recherche: ANR-10-INBS-09 Schweizerischer Nationalfonds zur Forderung der Wissenschaftlichen Forschung: PP00P3_170664

Every year the Caribbean Sea faces the passage of powerful tropical cyclones that generate coastal extreme sea levels with potential strong and hazardous impacts. In this work we simulate the storm surges and wind-waves induced by a set of over 1000 tropical cyclones over the Caribbean Sea that are representative of the present-day climate and that have been extracted from a global database of synthetic hurricanes spanning a 10,000-year period. The atmospheric forcing fields, built from the synthetic tropical cyclones, are used to feed a fully coupled hydrodynamic-wave model with high resolution (~1 km) along the continental and island coasts. Given the large number of events modelled, the outputs allow detailed statistical analyses of the magnitude and mechanisms of coastal extreme sea levels as well as the identification of most exposed areas to both storm surges and large wind-waves.

Medicanes, for Mediterranean hurricanes, are mesoscale cyclones with morphological and physical characteristics similar to tropical cyclones. Although less intense, smaller and rarer than their Atlantic counterparts, Medicanes remain very hazardous events threatening islands and continental coasts within the Mediterranean Sea. The latest strong episode Medicane Ianos (September 2020), resulted in severe damages in Greece and several casualties. This work investigates the oceanic response to these extreme events along the Mediterranean coasts under present-day and future (21 st century) conditions. To this end, a coupled hydrodynamic-wave model is used to simulate both storm surges and wind-waves generation and propagation in the Mediterranean Sea at high resolution (~2 km) along the coastlines. A dataset of thousands of Medicanes synthetically generated from twenty global climate models and two reanalyses is used to derive the atmospheric forcing fields. Regional coastal risks assessment is performed for the present and future climate. We found increased coastal extreme sea levels in line to the reported changes in Medicane activity, with fewer events but of larger intensity projected by late 21 st century.

The pace of the sequencing and computational assembly of novel reference genomes is accelerating. Though DNA sequencing technologies and assembly software tools continue to improve, biological features of genomes such as repetitive sequence as well as molecular artifacts that often accompany sequencing library preparation can lead to fragmented or chimeric assemblies. If left uncorrected, defects like these trammel progress on understanding genome structure and function, or worse, positively mislead such research. Fortunately, integration of additional, independent streams of information, such as a genetic map – particularly a marker-dense map from RADseq, for example – and conserved orthologous gene order from related taxa can be used to scaffold together unlinked, disordered fragments and to restructure a reference genome where it is incorrectly joined. We present a tool set for automating these processes, one that additionally tracks any changes to the assembly and to the genetic map, and which allows the user to scrutinize these changes with the help of web-based, graphical visualizations. Chromonomer takes a user-defined reference genome, a map of genetic markers, and, optionally, conserved synteny information to construct an improved reference genome of chromosome models: a “chromonome”. We demonstrate Chromonomer’s performance on genome assemblies and genetic maps that have disparate characteristics and levels of quality.

Sexual selection results in sex-specific characters like the conspicuously pigmented extension of the ventral tip of the caudal fin - the “sword” - in males of several species of Xiphophorus fishes. To uncover the genetic architecture underlying sword formation and to identify genes that are associated with its development, we characterized the sword transcriptional profile and combined it with genetic mapping approaches. Results showed that the male ornament of swordtails develops from a sexually non-dimorphic prepattern of transcription factors in the caudal fin. Among genes that constitute the exclusive sword transcriptome only two are located in the genomic region associated with this trait; the chaperone, fkbp9, and the potassium channel, kcnh8 that in addition to its neural function performs a role known to affect fin growth. This indicates that during evolution of swordtails a brain gene has been recruited for an additional function in establishing a male ornament.

18 pages, 13 figures, supplementary material https://www.frontiersin.org/articles/10.3389/fmars.2020.00497/full#supplementary-material.-- Publicly available datasets were analyzed in this study. This data can be found here: http://marine.copernicus.eu/services-portfolio/access-to-products/, http://www.ba.ieo.es/ibamar, http://thredds.socib.es/thredds/catalog.html, Numerical modeling is a key tool to complement the current physical and biogeochemical observational datasets. It is essential for understanding the role of oceanographic processes on marine food webs and producing climate change projections of variables affecting key ecosystem functions. In this work, we evaluate the horizontal and vertical patterns of four state-of-the-art coupled physical–biogeochemical models, three of them already published. Two of the models include data assimilation, physical and/or biological, and two do not. Simulations are compared to the most exhaustive dataset of in situ observations in the North Western Mediterranean, built ad hoc for this work, comprising gliders and conventional CTD surveys and complemented with satellite observations. The analyses are performed both in the whole domain and in four subregions (Catalan Shelf, Ebro Delta, Mallorca Channel, and Ibiza Channel), characterized by a priori divergent primary production dynamics and driving mechanisms. Overall, existing models offer a reasonable representation of physical processes including stratification, surface temperature, and surface currents, but it is shown that relatively small differences among them can lead to large differences in the response of biogeochemical variables. Our results show that all models are able to reproduce the main seasonal patterns of primary production both at the upper layer and at the deep chlorophyll maximum (DCM), as well as the differential behavior of the four subregions. However, there are significant discrepancies in the local variability of the intensity of the winter mixing, phytoplankton blooms, or the intensity and depth of the DCM. All model runs show markedly contrasting patterns of interannual phytoplankton biomass in all four subregions. This lack of robustness should dissuade end users from using them to fill gaps in time series observations without assessing their appropriateness. Finally, we discuss the usability of these models for different applications in marine ecology, including fishery oceanography, This work was supported by the European Union’s Horizon 2020 Research and Innovation Program under grant agreement no. 678193 (CERES, Climate Change and European Aquatic Resources). ER-R is grateful for the funding from “Govern de les Illes Balears-Conselleria d’Innovació, Recerca i Turisme, Programa Vicenç Mut.”, With the funding support of the ‘Severo Ochoa Centre of Excellence’ accreditation (CEX2019-000928-S), of the Spanish Research Agency (AEI)

The ocean component and coastal impacts of Storm Gloria, that hit the Western Mediterranean between January 20th and 23rd 2020 are investigated with a numerical simulation of the storm surges and wind-waves. Storm Gloria caused severe damages and beat several historical records such as significant wave height or 24-h accumulated precipitation. The storm surge developed along the eastern coasts of the Iberian Peninsula reached values up to 1 m, and were accompanied by wind-waves with significant wave height up to 8 m. Along the coasts of the Balearic Islands, the storm footprint was characterised by a negligible storm surge and the impacts were caused by large waves. The comparison to historical records reveals that Storm Gloria is one of the most intense among the events in the region during the last decades and that the waves direction was particularly unusual. Our simulation permits quantifying the role of the different forcings in generating the storm surge. Also, the high spatial grid resolution down to 30 m over the Ebro Delta, allows determining the extent of the flooding caused by the storm surge. We also simulate the overtopping caused by high wind waves that affected a rocky coast of high cliffs (~ 15 m) in the eastern coast of Mallorca Island.

A set of historical tide-gauge sea-level records from two locations in Santander (Northern Spain) and Alicante (Spanish Mediterranean coast) have been recovered from logbooks stored in national archives. Sea-level measurements have been digitised, quality-controlled and merged into three consistent sea-level time series (two in Alicante and one in Santander) using high precision levelling information. The historical sea-level record in Santander consists of a daily time series spanning the period 1876-1924 and it is further connected to the modern tide-gauge station nearby, ensuring datum continuity up to the present. The sea-level record in Alicante starts in 1870 with daily averaged values until the 1920s and hourly afterwards, and is still in operation, thus representing the longest tide-gauge sea-level time series in the Mediterranean Sea. The long-term consistency and reliability of the new records is discussed based on the comparison with nearby tide gauge time series.

The Maldives, located in the Indian Ocean, are the paradigm of low-lying coral-reef islands where adaptation to climate change is essential. Besides the mean sea level rise in this region, that is expected to be around 1 m by the end of the century according to the last IPCC report, these islands are exposed to one of the largest swells in terms of significant wave height and peak period. In this study we characterize, using the output of global wind-wave models forced by wind fields from the CMIP5 ensemble delivered by CSIRO, the present conditions and future projections of the waves around the Maldivian archipelago, as well as the return periods for the predominant swell directions. We then propagate extreme waves inside the domain using WaveWatch III model run onto a high-resolution grid (down to 500 m at the coast). Finally, we evaluate the coastal impacts of extreme swell waves in two strategic case study islands with different exposures and where land reclamation and different adaptation solutions have been/are being done. To do so, we propagate waves using SWASH model with rising mean sea level towards the shoreline and assess the flooding extend under different conditions. This kind of studies are essential to help the policy makers in defining the most accurate and appropriate adaptation strategies.

Twenty-first century projections for the Mediterranean water properties have been analyzed using the largest ensemble of regional climate models (RCMs) available up to now, the Med-CORDEX ensemble. It is comprised by 25 simulations, 10 historical and 15 scenario projections, from which 11 are ocean–atmosphere coupled runs and 4 are ocean forced simulations. Three different emissions scenarios are considered: RCP8.5, RCP4.5 and RCP2.6. All the simulations agree in projecting a warming across the entire Mediterranean basin by the end of the century as a result of the decrease of heat losses to the atmosphere through the sea surface and an increase in the net heat input through the Strait of Gibraltar. The warming will affect the whole water column with higher anomalies in the upper layer. The temperature change projected by the end of the century ranges between 0.81 and 3.71 °C in the upper layer (0–150 m), between 0.82 and 2.97 °C in the intermediate layer (150–600 m) and between 0.15 and 0.18 °C in the deep layer (600 m—bottom). The intensity of the warming is strongly dependent on the choice of emission scenario and, in second order, on the choice of Global Circulation Model (GCM) used to force the RCM. On the other hand, the local structures reproduced by each simulation are mainly determined by the regional model and not by the scenario or the global model. The salinity also increases in all the simulation due to the increase of the freshwater deficit (i.e. the excess of evaporation over precipitation and river runoff) and the related increase in the net salt transport at the Gibraltar Strait. However, in the upper layer this process can be damped or enhanced depending upon the characteristics of the inflowing waters from the Atlantic. This, in turn, depends on the evolution of salinity in the Northeast Atlantic projected by the GCM. Thus a clear zonal gradient is found in most simulations with large positive salinity anomalies in the eastern basin and a freshening of the upper layer of the western basin in most simulations. The salinity changes projected for the whole basin range between 0 and 0.34 psu in the upper layer, between 0.08 and 0.37 psu in the intermediate layer and between − 0.05 and 0.33 in the deep layer. These changes in the temperature and salinity modify in turn the characteristics of the main water masses as the new waters become saltier, warmer and less dense along the twenty-first century. There is a model consensus that the intensity of the deep water formation in the Gulf of Lions is expected to decrease in the future. The rate of decrease remains however very uncertain depending on the scenario and model chosen. At the contrary, there is no model consensus concerning the change in the intensity of the deep water formation in the Adriatic Sea and in the Aegean Sea, although most models also point to a reduction.

Pure and Applied Geophysics (PAGEOPH) is one of the leading journals in the field of geophysics. The first issue was published in 1939; thus, the journal is celebrating its 80th anniversary in 2018. The aim of this paper is to provide a complete lifetime overview of the academic structure of the journal using bibliometric indicators. This analysis includes key factors such as the most cited articles, leading authors, originating institutions and countries, publication and citation structures, and the most commonly used keywords. The bibliometric data used to conduct this analysis comes from the Scopus database. Additionally, the visualization of similarities (VOS)viewer software is used to create a graphic map of some of the bibliometric results. The graphical analysis uses co-citation, bibliographic coupling and co-occurrence of keywords. The results indicate that PAGEOPH is a leading journal in the areas in which it is indexed, with publications from a wide range of authors, institutions, and countries around the world.

Remotely generated swell waves are the dominant contributor of the coastal wind-wave climate along most of the world coastlines. In this work we describe the characteristics of swells from a coastal perspective. We identify the main regions of formation of swell waves at present and during the late twenty-first century under the RCP8.5 emissions/climate change scenario. We have applied an algorithm that allows one to unequivocally differentiate the swell component from the local wind-waves for a global wave hindcast and for eight CMIP5 state-of-the-art wave model climate projections. We have identified four different regions of swell formation, two in each hemisphere, with the Southern Ocean being by far the main region of swell generation. By the end of this century, the number of swell events generated in the Northern Hemisphere is expected to decrease while the opposite is projected to occur in the Southern Hemisphere. The increase in the Southern Hemisphere is directly associated with a poleward movement and intensification of the wind belts projected by atmospheric climate models., This study was supported by the ERA4CS INSeaPTION project (grant number: 690462 and PCIN-2017-038) funded by the Spanish Ministerio de Economía, Industria y Competitividad - Agencia Estatal de Investigación.

Good knowledge of mesoscale eddy properties and their spatial and temporal distribution in the world ocean is important for an accurate estimate of their role in heat, salt and/or momentum transports. Composites of satellite sea level anomaly, used to evaluate internal eddy structure, commonly produce, at larger radii, a series of rings of alternating sign. We suggest that these secondary rings are not a part of the dynamical structure of the "central" eddy but are a rectified signature of the surrounding eddies. We show that statistical distribution of eddy polarities is not random but that an eddy of given polarity tends to be surrounded by eddies of opposite polarity. This observed spatial eddy polarity distribution, which is hypothesized to be a result of eddy self- organization, is not only responsible for the secondary rings observed in the composite estimates but also causes a reduction of around 20% of the eddy composite amplitude and an increment of around 10% in the maximum rotational velocity, while it has negligible effect on the estimated eddy scale.

Half of all vertebrate species share a series of chromosome fusions that preceded the teleost genome duplication (TGD), but we do not understand the causative evolutionary mechanisms. The “Robertsonian-translocation hypothesis” suggests a regular fusion of each ancestral acro- or telocentric chromosome to just one other by centromere fusions, thus halving the karyotype. An alternative “genome-stirring hypothesis” posits haphazard and repeated fusions, inversions, and reciprocal and nonreciprocal translocations. To study large-scale karyotype reduction, we investigated the decrease of chromosome numbers in Antarctic notothenioid fish. Most notothenioids have 24 haploid chromosomes, but bullhead notothen (Notothenia coriiceps) has 11. To understand mechanisms, we made a RAD-tag meiotic map with ∼10,000 polymorphic markers. Comparative genomics aligned about a thousand orthologs of platyfish and stickleback genes along bullhead chromosomes. Results revealed that 9 of 11 bullhead chromosomes arose by fusion of just two ancestral chromosomes and two others by fusion of three ancestral chromosomes. All markers from each ancestral chromosome remained contiguous, implying no inversions across fusion borders. Karyotype comparisons support a history of: (1) Robertsonian fusions of 22 ancestral chromosomes in pairs to yield 11 fused plus two small unfused chromosomes, like N. angustata; (2) fusion of one of the remaining two ancestral chromosomes to a preexisting fused pair, giving 12 chromosomes like N. rossii; and (3) fusion of the remaining ancestral chromosome to another fused pair, giving 11 chromosomes in N. coriiceps. These results raise the question of what selective forces promoted the systematic fusion of chromosomes in pairs and the suppression of pericentric inversions in this lineage, and provide a model for chromosome fusions in stem teleosts.

A persistent signature of coherent mesoscale eddies in sea surface salinity (SSS) is revealed by analyzing the relationship between satellite SSS and sea surface height (SSH) variability in an eddy-following reference frame. Our analysis focuses on mid-ocean eddies in two representative regions, the southern Indian Ocean and the North Atlantic subtropical gyre. The resulting composite averages reveal a clear signature of mesoscale eddies in satellite SSS with typical SSS anomalies of 0.03-0.05 psu. The spatial structure of eddy-induced SSS perturbations can be characterized as a superposition of a dipole structure, arising from horizontal advection of the background SSS gradient by eddy velocity field, and a monopole structure related to the eddy core. The observed relationships between SSS and SSH anomalies are used to provide a regional assessment of the role of mesoscale eddies in the ocean freshwater transport in the North Atlantic subtropical gyre. This article is protected by copyright. All rights reserved.

Este artículo contiene 11 páginas, 4 figuras, 1 tabla., A mass mortality event is devastating the populations of the endemic bivalve Pinna nobilis in the Mediterranean Sea from early autumn 2016. A newly described Haplosporidian endoparasite (Haplosporidium pinnae) is the most probable cause of this ecological catastrophe placing one of the largest bivalves of the world on the brink of extinction. As a pivotal step towards Pinna nobilis conservation, this contribution combines scientists and citizens’ data to address the fast- and vastdispersion and prevalence outbreaks of the pathogen. Therefore, the potential role of currents on parasite expansion was addressed by means of drift simulations of virtual particles in a high-resolution regional currents model. A generalized additive model was implemented to test if environmental factors could modulate the infection of Pinna nobilis populations. The results strongly suggest that the parasite has probably dispersed regionally by surface currents, and that the disease expression seems to be closely related to temperatures above 13.5 °C and to a salinity range between 36.5–39.7 psu. The most likely spread of the disease along the Mediterranean basin associated with scattered survival spots and very few survivors (potentially resistant individuals), point to a challenging scenario for conservation of the emblematic Pinna nobilis, which will require fast and strategic management measures and should make use of the essential role citizen science projects can play., This study was financed by the special action project “NACRAS – Evaluación de la magnitud de un evento de mortalidad combinando censos científicos ampliados con observaciones de ciencia ciudadana” cofunded by the Regional Government and Direcció General d’Innovació i Recerca of the Balearic Islands and the European Social Fund 2014–2020. Salinity and temperature data at Alfacs Bay was provided from the “Programa de Seguimiento de la calidad de las aguas, moluscos y fitoplancton tóxico en las zonas de producción de marisco del litoral catalán (PSQAM) de la DGPAM.” The authors acknowledge the MEDCLIC project, funded by “La Caixa” Foundation, contributing to the development of the WMOP hydrodynamic model. Maite Vázquez- Luis and Miguel Cabanellas-Reboredo were supported by a postdoctoral contract co-funded by the Regional Government of the Balearic Islands and the European Social Fund 2014–2020. Also, Vázquez-Luis was also supported by a postdoctoral contract Juan de la Cierva-Incorporación (IJCI-2016-29329) of Ministerio de Ciencia, Innovación y Universidades. Elvira Álvarez was supported by a Personal Técnico de Apoyo contract (PTA2015-10829-I) funded by the Spanish Ministry of Economy and Competiveness. Iris E. Hendriks was supported by Ramon y Cajal Fellowship RYC-2014-14970, cofunded by the Conselleria d’Innovació, Recerca i Turisme of the Balearic Government (Pla de ciència, tecnologia, innovació i emprenedoria 2013-2017) and the Spanish Ministry of Economy, Industry and Competitiveness. Jose Rafael Garcia-March and Jose Tena were supported by the project BF/HEM 15-1662

The dependence between extreme storm surges and wind waves is assessed statistically along the global coasts using the outputs of two numerical models consistently forced with the same atmospheric fields. We show that 55% of the world coastlines face compound storm surge wave extremes. Hence, for a given level of probability, neglecting these dependencies leads to underestimating extreme coastal water levels. Dependencies are dominant in midlatitudes and are likely underestimated in the tropics due to limited representation of tropical cyclones. Furthermore, we show that in half of the areas with dependence, the estimated probability of occurrence of coastal extreme water levels increases significantly when it is accounted for. Translated in terms of return periods, this means that along 30% of global coastlines, extreme water levels expected at most once in a century without considering dependence between storm surges and waves become a 1 in 50-year event., This study was supported by the ERA4CS INSeaPTION project (grants 690462 and PCIN‐2017‐038).

Intestinal tract development is a coordinated process involving signaling among the progenitors and developing cells from all three germ layers. Development of endoderm-derived intestinal epithelium has been shown to depend on epigenetic modifications, but whether that is also the case for intestinal tract cell types from other germ layers remains unclear. We found that functional loss of a DNA methylation machinery component, ubiquitin-like protein containing PHD and RING finger domains 1 (uhrf1), leads to reduced numbers of ectoderm-derived enteric neurons and severe disruption of mesoderm-derived intestinal smooth muscle. Genetic chimeras revealed that Uhrf1 functions both cell-autonomously in enteric neuron precursors and cell-non-autonomously in surrounding intestinal cells, consistent with what is known about signaling interactions between these cell types that promote one another's development. Uhrf1 recruits the DNA methyltransferase Dnmt1 to unmethylated DNA during replication. Dnmt1 is also expressed in enteric neurons and smooth muscle progenitors. dnmt1 mutants have fewer enteric neurons and disrupted intestinal smooth muscle compared to wildtypes. Because dnmt1;uhrf1 double mutants have a similar phenotype to dnmt1 and uhrf1 single mutants, Dnmt1 and Uhrf1 must function together during enteric neuron and intestinal muscle development. This work shows that genes controlling epigenetic modifications are important to coordinate intestinal tract development, provides the first demonstration that these genes influence development of the ENS, and advances uhrf1 and dnmt1 as potential new Hirschsprung disease candidates.