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Agricultural extensification refers to an expansive, low-input production strategy that is land rather than labour-limited. Here we present a robust method, using the archaeological proxies of cereal grain nitrogen isotope values and settlement size, to investigate the relationship between agricultural intensity and population size at Neolithic to Bronze/Iron Age settlement sites in northern Mesopotamia, the Aegean and south-west Germany. We conclude that urban form—in particular, density of occupation—as well as scale shaped the agroecological trajectories of early cities. Whereas high-density urbanism in northern Mesopotamia and the Aegean entailed radical agricultural extensification, lower density urbanism in south-west Germany afforded more intensive management of arable land. We relate these differing agricultural trajectories to long-term urban growth/collapse cycles in northern Mesopotamia and the Aegean, on the one hand, and to the volatility of early Iron Age elite power structures and urban centralisation in south-west Germany, on the other.

Around 10,000 y ago in southwest Asia, the cessation of a mobile lifestyle and the emergence of the first village communities during the Neolithic marked a fundamental change in human history. The first communities were small (tens to hundreds of individuals) but remained semisedentary. So-called megasites appeared soon after, occupied by thousands of more sedentary inhabitants. Accompanying this shift, the material culture and ancient ecological data indicate profound changes in economic and social behavior. A shift from residential to logistical mobility and increasing population size are clear and can be explained by either changes in fertility and/or aggregation of local groups. However, as sedentism increased, small early communities likely risked inbreeding without maintaining or establishing exogamous relationships typical of hunter-gatherers. Megasites, where large populations would have made endogamy sustainable, could have avoided this risk. To examine the role of kinship practices in the rise of megasites, we measured strontium and oxygen isotopes in tooth enamel from 99 individuals buried at Pınarbaşı, Boncuklu, and Çatalhöyük (Turkey) over 7,000 y. These sites are geographically proximate and, critically, span both early sedentary behaviors (Pınarbaşı and Boncuklu) and the rise of a local megasite (Çatalhöyük). Our data are consistent with the presence of only local individuals at Pınarbaşı and Boncuklu, whereas at Çatalhöyük, several nonlocals are present. The Çatalhöyük data stand in contrast to other megasites where bioarchaeological evidence has pointed to strict endogamy. These different kinship behaviors suggest that megasites may have arisen by employing unique, community-specific kinship practices.

Abbo and Gopher contend that we offer nothing new to the study of domestication in three recent papers (Bogaard et al., 2021; Allaby et al., 2021, Allaby et al., 2022b). They claim that we offer no “innovation, a new venue of research” and “use a new jargon to express old ideas.” They further claim as erroneous our key conclusions about domestication as: protracted, co-evolutionary, comprising multiple pathways of convergent evolution, and taking place at the landscape scale. Here we defend these recent contributions as genuine progress that builds on previous ideas and hypotheses through empirical illustration and a raft of new data. Combining new data with old and new theory, we develop frameworks that suggest future directions for research.

Key message We present a comprehensive survey of cytogenetic and genomic diversity of the GGAtAt genepool of wheat, thereby unlocking these plant genetic resources for wheat improvement. Abstract Wheat yields are stagnating around the world and new sources of genes for resistance or tolerances to abiotic traits are required. In this context, the tetraploid wheat wild relatives are among the key candidates for wheat improvement. Despite its potential huge value for wheat breeding, the tetraploid GGAtAt genepool is largely neglected. Understanding the population structure, native distribution range, intraspecific variation of the entire tetraploid GGAtAt genepool and its domestication history would further its use for wheat improvement. The paper provides the first comprehensive survey of genomic and cytogenetic diversity sampling the full breadth and depth of the tetraploid GGAtAt genepool. According to the results obtained, the extant GGAtAt genepool consists of three distinct lineages. We provide detailed insights into the cytogenetic composition of GGAtAt wheats, revealed group- and population-specific markers and show that chromosomal rearrangements play an important role in intraspecific diversity of T. araraticum. The origin and domestication history of the GGAtAt lineages is discussed in the context of state-of-the-art archaeobotanical finds. We shed new light on the complex evolutionary history of the GGAtAt wheat genepool and provide the basis for an increased use of the GGAtAt wheat genepool for wheat improvement. The findings have implications for our understanding of the origins of agriculture in southwest Asia.

Excavations at Knossos have uncovered faunal and archaeobotanical archives spanning the Neolithic and Bronze Age (7th–2nd millennia bce), during which one of Europe’s earliest known farming settlements developed into its first major urban settlement and centre of one of its oldest regional states. Through stable isotope (δ13C, δ15N) analysis of seeds and bones (as evidence for the growing conditions of cereal and pulse crops and for the types of forage consumed by livestock), land use and, ultimately, political economy are explored. Changing husbandry conditions overwrite any effects of long-term aridification. Early (7th–6th millennium bce) Knossian farmers grew intensively managed cereals and pulses (probably in rotation) that were closely integrated (as manured sources of forage) with livestock. Through the later Neolithic and Bronze Age, settlement growth accompanied more extensive cultivation (eventually with cereals and pulses not in rotation) and greater use of rough graze and, by goats, browse. Pasture on cultivated land remained central, however, to the maintenance of sheep, cattle, and pigs. Variable diet of early sheep suggests management at the household level, while thereafter progressive dietary divergence of sheep and goats implies their separate herding. Until the Old Palace phase (early 2nd millennium bce), urban growth was matched by increasingly extensive and probably distant cultivation and herding but somewhat more intensive conditions during the New and Final Palace phases (mid-2nd millennium bce) perhaps reflect greater reliance on surplus from prime land of previously rival centres that now came under Knossian control.

Andrew Sherratt’s ‘Water, soil and seasonality’, World Archaeology (1980), signposted a long-term debate surrounding early farming adaptations to riverine landscapes in western Asia and Europe. Recent research at Çatalhöyük in central Anatolia, a key case study in Sherratt’s ‘floodplain cultivation’ model, enables integrated, evidence-based assessment of the local hydrology and agroecology, and of farmers’ resilience over more than a millennium. In contrast to previous models, the agroecological niche at Çatalhöyük featured strategic planting of diverse crops across a range of hydrological conditions, within and beyond a broad ‘belt’ of small anastomosing river channels extending a kilometre from the site. Growing conditions likely depended on location relative to settlement, a nutrient-rich ‘hot spot’, with diminishing inputs of organic matter and mechanical disturbance away from the tell. This reconstruction contrasts with the original model of ‘floodplain cultivation’ and demonstrates the complexity with which agroecologies evolved through landscape affordances, creative cropping, and resilience.

Reconstructing the origins of plant cultivation in southwest Asia is crucial for understanding associated processes such as the emergence of sedentary communities and domesticated crops. Among the criteria archaeobotanists developed for identifying the earliest plant cultivation, the presence of potential arable weeds found in association with wild cereal and legume remains has been used as a basis for supporting models of prolonged wild plant cultivation before domesticated crops appear. However, the proposed weed floras mainly consist of genus-level identifications that do not differentiate between arable weeds and related species that characterise non-arable habitats. Here we test, for the first time, whether the potential arable weed taxa widely used to identify wild plant cultivation also occur in non-cultivated wild cereal populations. Based on modern survey data from the southern Levant we show that the proposed weed taxa characterise both grasslands and fields. Our findings, therefore, do not support the use of these taxa for reconstructing early cultivation. Instead, for future studies we suggest an approach based on the analysis of plant functional traits related to major agroecological variables such as fertility and disturbance, which has the potential to overcome some of the methodological problems.

International audience; New stable carbon (δ13C) and nitrogen (δ15N) isotopic values of charred plant and bone collagen remains from 6th mill. BCE Halai, central Greece, together with datasets from 6th mill. BCE Kouphovouno, southern Greece, and later 6th/early 5th mill. BCE Makriyalos, northern Greece, demonstrate how early farming communities in mainland Greece adapted mixed farming strategies to distinct local environmental and cultural settings. Intra-site similarities or differences in δ13C and δ15N values of distinct crop species, along with the intra-species variabilities in stable isotopic values, are used to assess the cultivation choices that farmers at the three sites made to fulfill distinct economic goals. At Halai, farmers cultivated multiple crops under variable soil conditions, a strategy likely geared towards minimizing overall risk in a relatively arid coastal setting. At better-watered Kouphovouno, by contrast, farmers practiced strategic manuring to maximize the yield of free-threshing wheat, likely grown exclusively for human consumption and rotated with nitrogen-fixing pulses. At Makriyalos, the limited sample size of cereal remains suggests a lack of intensive manuring, in contrast to the two other sites. Assessment of which crops may have been consumed by livestock sheds further light on symbiotic relationships between crop cultivation and animal husbandry.

Broomcorn and foxtail millet are the only major domesticated plants indigenous to prehistoric Eurasia to rely on the C4photosynthetic pathway. Here we study the impact of animal manure (AM) on broomcorn millet (Panicum miliaceum) and foxtail millet (Setaria italica), grown in the Askov Long-Term Experiment, using unmanured soil and soil dressed with phosphorus plus potassium (PK) as reference treatments. Animal manure had a marked effect on yields and on the δ15N values of grains. For broomcorn millets grown on manured soil, the average δ15N value was 5.8‰. When grown on unmanured and PK-treated soils, the δ15N values were 0.4‰ and 0.2‰, respectively. For foxtail millet the δ15N values also differed between grains from unmanured (-1.0‰), PK (0.7‰) and manured (6.3‰) treatments. Thus, when compared to unmanured soil, the offset due to manure was 7.3‰ forSetariaand 5.3‰ for the twoPanicumvarieties. In accordance with previous studies on C3crops, our study suggests that δ15N values in charred millet grains recovered fromarchaeological sitescould provide a robust indicator of prehistoric manuring intensity.

In much of Europe, the advent of low-input cereal farming regimes betweenc.ad800 and 1200 enabled landowners—lords—to amass wealth by greatly expanding the amount of land under cultivation and exploiting the labour of others. Scientific analysis of plant remains and animal bones from archaeological contexts is generating the first direct evidence for the development of such low-input regimes. This article outlines the methods used by the FeedSax project to resolve key questions regarding the ‘cerealization’ of the medieval countryside and presents preliminary results using the town of Stafford as a worked example. These indicate an increase in the scale of cultivation in the Mid-Saxon period, while the Late Saxon period saw a shift to a low-input cultivation regime and probably an expansion onto heavier soils. Crop rotation appears to have been practised from at least the mid-tenth century.

Stable nitrogen (N) isotope analysis of bulk tissues is a technique for reconstructing the diets of organisms. However, bulk nitrogen isotope (δ15N) values can be influenced by a variety of metabolic and environmental factors that can confound accurate dietary reconstruction. Compound-specific isotope analyses of amino acids (CSIA-AA) have demonstrated the power of the approach in understanding how the δ15N values of bulk collagen are assembled from the constituent AAs. Furthermore, by connecting these AA δ15N values within a robust biochemical framework interpretation of diet and environment are greatly enhanced. Several new proxies have emerged, built around selected AAs; however, the interconnectedness of AA biosynthetic pathways means that patterning of δ15N values across a wider suite of collagen AAs will occur under different environmental or dietary influences. This work seeks to test this idea by situating CSIA-AA within a robust statistical framework using principal component analysis (PCA) and Bayesian statistics to increase the interpretability of a wider range of AA δ15N values in terms of reconstructing herbivore diet. The model was tested using wild and domestic herbivores from the Neolithic settlements of Çatalhöyük (Turkey), Makriyalos (Greece), and Vaihingen (Germany) as case studies. It was found that at Makriyalos there was a sharp separation between domesticated and wild herbivores, which was present to a lesser extent at Çatalhöyük and not observed at Vaihingen. The case studies presented in this work demonstrate that multivariate statistical treatment of CSIA-AA data can deliver new insights into herbivore diet, exceeding those achievable with the Bayesian model.

International audience; Process philosophy offers a metaphysical foundation for domestication studies. This grounding is especially important given the European colonialist origin of 'domestication' as a term and 19th century cultural project. We explore the potential of process archaeology for deep-time investigation of domestication relationships, drawing attention to the variable pace of domestication as an ongoing process within and across taxa; the nature of domestication 'syndromes' and 'pathways' as general hypotheses about process; the importance of cooperation as well as competition among humans and other organisms; the significance of non-human agency; and the ubiquity of hybrid communities that resist the simple wild/domestic dichotomy.

© 2019, Springer-Verlag GmbH Germany, part of Springer Nature. The nitrogen isotope compositions of charred wheat and barley grains reflect manuring intensity and have been used to reconstruct past manuring practices at archaeological sites across Europe and western Asia. To assess whether this analytical method can be applied to a staple crop in the West African Sahel, the nitrogen isotope values of Pennisetum glaucum grains in this region were determined and the effect of charring ascertained. Pennisetum glaucum ears were collected from fields in northeast Senegal, where the fertilisation histories of the plots (manure and/or household waste) were known. The nitrogen isotope values of these millet grains provide an insight into the values to expect for P. glaucum grains grown with low to moderate addition of manure/household waste in a semi-arid climate. Charring of P. glaucum grains by heating at 215–260 °C for 4–24 h increases their nitrogen isotope values by a maximum of 0.34‰. In light of these modern data, the nitrogen isotope values of millet grains recovered from the archaeological settlement mound of Tongo Maaré Diabal, Mali, can be interpreted as evidence for modest levels of manure/household waste input throughout the occupation of the site from cal ad 500–1150. This study demonstrates the potential for nitrogen isotope values of P. glaucum grains to shed light on past farming practices in West Africa.

‘New glume wheat’ (NGW) is an archaeobotanical type increasingly recognised at Neolithic–Bronze Age sites across Europe and Western Asia. NGW has been recognised via aDNA and morphological analyses of chaff remains as a member of theTriticum timopheeviiwheat group, recent cultivation of which is known only from western Georgia. This study combines geometric morphometric (GMM) analysis of NGW grains with updated results from a parallel study of chaff dehiscence, to assess the taxonomic classification and domestication status of NGW from the Neolithic East Mound at Çatalhöyük (central Anatolia). Results confirm close comparability of NGW with modern wheats from the groupT. timopheevii, in a form which has remained remarkably similar over thousands of years. Furthermore, the analysis suggests that NGW was undergoing selection for domestication traits in terms of shattering behaviour and grain form during the 1150-year East Mound sequence. These findings are interpreted in the context of substantial archaeobotanical evidence for a broad-spectrum plant strategy at Çatalhöyük which mitigated the risk of resource failure and supported experimentation in cropping. Possible cultural and practical incentives are considered for investment in the crop, made despite the availability of a fully-domesticated glume wheat (emmer) with similar growing and processing requirements. Alongside this, the study demonstrates the sensitivity of GMM to differences between and within wheat species, with methodological findings that can inform future studies.

We present a comprehensive survey of cytogenetic and genomic diversity of the GGA

The integration of stable isotope analysis of crop remains with the ecological analysis of archaeological weed flora allows for a detailed reconstruction of crop husbandry and arable growing conditions. This paper presents the first such study for developed Neolithic and Chalcolithic Central Anatolia, exploring crop husbandry at the large (8 ha) early 6th millennium BC site of Catalhoyuk West and the last phases of the preceding Neolithic Catalhoyuk East, located on the western end of the Konya Plain, and the small 4th millennium sites of Camlibel Tarlasi (0.25 ha), located in the loop of the Kizilirmak river, and Kurucay Hoyuk (~1 ha), located on the shore of Lake Burdur. The results reveal the mosaic nature of Catalhoyuk cultivation; the complex ecosystem, naturally 15N enriched, was additionally anthropogenically improved, while inhabitants also chose to cultivate barley in drier conditions to the wheats and pulses. The smaller, Late Chalcolithic site of Camlibel Tarlasi cultivated each individual crop species in variable conditions instead of selecting a specific species for cultivation in drier or wetter conditions. Each single species was cultivated in conditions with a wide range of water availabilities while receiving low-level manure application. Variation in water availability is possibly a consequence of the variable topography surrounding the site, with the hills potentially used for cultivation, providing a wide range of conditions due to their slope. Functional weed ecology complements the isotopic data, indicating the wider range of cultivation intensities surrounding Catalhoyuk West, compared with more consistent, narrower ranges of growing conditions at Camlibel Tarlasi and Kurucay Hoyuk. Settlement date, site size and environment seem to be key contributing factors shaping the method used to cultivate crops at these Chalcolithic sites. The larger, earlier community of Catalhoyuk West, located in a complex riverine ecosystem around the dryland anabranching channels of the Carsamba river, cultivated species in different conditions and maintained a range of crop husbandry intensities, potentially reflecting differential access to limited resources. The smaller Late Chalcolithic sites, with communities consisting of one household to several households and located in less complex ecosystems with higher precipitation, maintained more stable crop growing conditions and management intensities. This is plausibly a consequence of equal access and sharing of critical resources for crop cultivation, such as labour, manure and land, coupled with the adequate precipitation rates.

Wheat yields are stagnating around the world and new sources of genes for resistance or tolerances to abiotic traits are required. In this context, the tetraploid wheat wild relatives are among the key candidates for wheat improvement. Despite of its potential huge value for wheat breeding, the tetraploid GGAtAt genepool is largely neglected. Understanding the population structure, native distribution range, intraspecific variation of the entire tetraploid GGAtAt genepool and its domestication history would further its use for wheat improvement. We report the first comprehensive survey of genomic and cytogenetic diversity sampling the full breadth and depth of the tetraploid GGAtAt genepool. We show that the extant GGAtAt genepool consists of three distinct lineages. We provide detailed insights into the cytogenetic composition of GGAtAt wheats, revealed group-, and population-specific markers and show that chromosomal rearrangements play an important role in intraspecific diversity of T. araraticum. We discuss the origin and domestication history of the GGAtAt lineages in the context of state-of-the-art archaeobotanical finds. We shed new light on the complex evolutionary history of the GGAtAt wheat genepool. We provide the basis for an increased use of the GGAtAt wheat genepool for wheat improvement. The findings have implications for our understanding of the origins of agriculture in southwest Asia.

Specialized and systematic underwater fieldwork at the prehistoric site of Ploča Mičov Grad at Gradište (North Macedonia) on the eastern shore of Lake Ohrid was undertaken in 2018 and 2019. It has substantiated the archeological site's outstanding preservation condition, and furthermore proven the numerous construction timbers’ suitability for dendrochronological analysis. Dendrochronological analysis on archaeological timbers was applied, combined with radiocarbon dating. Bayesian radiocarbon modeling allowed to ‘wiggle match’ the dendrochronological mean curves, i.e. allowed the precise chronological anchoring of ‘floating’ tree-ring sequences. Furthermore, radiocarbon dates of plant remains from the site's main archaeological layer are statistically evaluated. Based on the new findings, the strikingly high density of wooden piles at the site can be attributed to several construction phases of Neolithic (middle of 5th millennium BC) and Bronze Age (2nd millennium BC: 1800, 1400 and 1300 BCE) settlements. Intense settlement activity is furthermore evidenced by a cultural layer of mainly organic material under the lakebed up to 1.7 m in thickness, which accumulated during the Neolithic occupation of the bay in the middle of the 5th millennium BC. The presented research enables precise absolute dating of a series of settlement phases at Ploča Mičov Grad from the Neolithic and the Bronze Age, and hence provides important reference points for an absolute chronological framework for the prehistory of the southwestern Balkans. The investigations underline the potential of future research on waterlogged prehistoric settlements in the region., Journal of Archaeological Science: Reports, 38

When the first rice farmers expanded into Southeast Asia from the north about 4,000 y ago, they interacted with hunter-gatherer communities with an ancestry in the region of at least 50 millennia. Rigorously dated prehistoric sites in the upper Mun Valley of Northeast Thailand have revealed a 12-phase sequence beginning with the first farmers followed by the adoption of bronze and then iron metallurgy leading on to the rise of early states. On the basis of the burial rituals involving interment with a wide range of mortuary offerings and associated practices, we identify, by computing the values of the Gini coefficient, at least two periods of intensified social inequality. The first occurred during the initial Bronze Age that, we suggest, reflected restricted elite ownership of exotic valuables within an exchange choke point. The second occurred during the later Iron Age when increased aridity stimulated an agricultural revolution that rapidly led to the first state societies in mainland Southeast Asia.

Background and AimsPlants depend fundamentally on establishment from seed. However, protocols in trait-based ecology currently estimate seed size but not seed number. This can be rectified. For annuals, seed number should simply be a positive function of vegetative biomass and a negative function of seed size.MethodsUsing published values of comparative seed number as the ‘gold standard’ and a large functional database, comparative seed yield and number per plant and per m2 were predicted by multiple regression. Subsequently, ecological variation in each was explored for English and Spanish habitats, newly calculated C-S-R strategies and changed abundance in the British flora.Key ResultsAs predicted, comparative seed mass yield per plant was consistently a positive function of plant size and competitive ability, and largely independent of seed size. Regressions estimating comparative seed number included, additionally, seed size as a negative function. Relationships differed numerically between regions, habitats and C-S-R strategies. Moreover, some species differed in life history over their geographical range. Comparative seed yield per m2 was positively correlated with FAO crop yield, and increasing British annuals produced numerous seeds. Nevertheless, predicted values must be viewed as comparative rather than absolute: they varied according to the ‘gold standard’ predictor used. Moreover, regressions estimating comparative seed yield per m2 achieved low precision.ConclusionsFor the first time, estimates of comparative seed yield and number for >800 annuals and their predictor equations have been produced and the ecological importance of these regenerative traits has been illustrated. ‘Regenerative trait-based ecology’ remains in its infancy, with work needed on determinate vs. indeterminate flowering (‘bet-hedging’), C-S-R methodologies, phylogeny, comparative seed yield per m2 and changing life history. Nevertheless, this has been a positive start and readers are invited to use estimates for >800 annuals, in the Supplementary data, to help advance ‘regenerative trait-based ecology’ to the next level.

This paper investigates agricultural management choices of farmers at the Neolithic site of Kouphvouno, southern Greece. Previous stable isotopic analysis of charred plant remains and bone animal collagen showed that throughout the Neolithic occupation of this site, farmers employed species-specific strategies to cultivate crops and herd domestic animals. Additional analyses of charred plant remain carried out in this study (including einkorn, a cereal species not measured before) expanding our understanding of the diversity and flexibility of early crop cultivation on a local scale. Furthermore, sequential tooth enamel carbonate isotopic analyses are used to assess the seasonal dietary and grazing patterns of domestic sheep and goat, providing a more nuanced picture of the roles of these animals in the subsistence economy of this community. The results show that the species-specific cultivation system was dictated by the crops’ ecological adaptations. Based on a small number of individuals available for analysis, the findings suggest that animal management was also likely driven by cultural choices, and involved foddering of goats managed for milk and local grazing of sheep managed for meat.

The discovery of a large underground silo complex with spectacular intact grain stores at the Late Bronze Age Hittite capital of Hattusha in Turkey provides a unique snapshot of the mobilisation of crop production by the Hittite state. A combination of primary archaeobotanical analysis, crop stable isotope determinations and functional weed ecology reveals new insights into Hittite cultivation strategies, featuring a range of relatively low-input, extensive production regimes for hulled wheats and hulled barley. Taxation of extensively produced grain in the sixteenth century BC reveals how an ancient state sought to sustain itself, providing wider implications for the politics and ecology of territorially expansive states in Western Asia and beyond.

The analysis of microbotanical remains (starch grains and phytoliths) from food-related domestic contexts and artefacts has the potential to provide insights into daily plant preparation and consumption activities. In particular, pottery vessels offer an unparalleled comparative framework for the study of food preparation and consumption, since pottery is used for a variety of domestic food-related activities, including storage, processing and serving. This study illustrates the potential of microbotanical remains to provide a holistic approach to the plant food preparation and consumption cycle—from storage to disposal—through the analysis of starch grains and phytoliths from Late Minoan storage, cooking and serving vessels recently uncovered at the Knossian ‘neighbourhood’ of Lower Gypsades. The results show that starch grains and phytoliths are more abundant and diverse in cooking vessels, presumably reflecting a higher deposition of microbotanical remains as a result of the disturbance caused by cooking. The results further offer insights into Minoan storage and cooking practices at Gypsades, suggesting that cereals were stored de-husked in pithoi and, possibly, in palm leaf baskets, and later cooked together with non-staple plants. The virtual absence of microbotanical remains from serving vessels may be due to depositional factors and/or the fact that some of the vessels analysed in this study were used for serving/consuming liquids or animal products, and not plants., The project was made possible with funds provided by INSTAP (EH, GA, AB); Semple Fund, Department of Classics, University of Cincinnati (EH); ERC Grant No. 312785 (AB); Fell Fund, University of Oxford (AB); the British Academy Grant No. SG152915 (AB); and the BSA. While working on this manuscript, JJGG was funded by a Marie Skłodowska-Curie Individual Fellowship (MSCA 2015, European Commission Grant No. 704867) and a Juan de la Cierva Incorporación 2018 Fellowship (Spanish Ministry of Science, Innovation and Universities), and ET is currently a F.S.R. Research Fellow funded by the Universitè Catholique de Louvain.

Theories developed and validated using ethnographic and historical resources are often difficult to examine using sparse or fragmentary archaeological material. However, a number of statistical techniques make it possible to integrate data from ethnographic, historical, and archaeological resources into a single analytical framework. This article introduces Material Correlates Analysis (MCA)—a new method of filling gaps in the archaeological data using a strategic combination of data collection, multidimensional scaling, principal component analysis, and generalized liner modeling. Generalized liner modeling is a particularly useful tool in formal inferential statistics for comparing a priori classified groups of historical and/or ethnographic (known) cases with archaeological (unknown) ones on the basis of relevant variables. MCA allows us to overcome the inherent material culture limitations regarding data on key variables by using available historical or ethnographic evidence to make statistically testable inferences regarding archaeological data. Using the Modes of Religiosity theory as an example, we demonstrate how major gaps in the evidentiary record can be overcome using the techniques we outline. Specifically, we use the MCA approach to ascertain whether the agricultural transition in southwest Asia was associated with a shift from an imagistic to an increasingly doctrinal mode of religiosity.

Analyses of house-size distributions in the Old and New World showed that wealth disparities increased with the domestication of plants and animals and with increased sociopolitical scale. Beneath headlines about booms and busts and other economic disturbance lies a deeper problem: wealth inequality. But what is its history, and what are the larger social factors that determine the disparate distribution of wealth? Timothy Kohler and colleagues look at the evolution of inequality worldwide since the Neolithic era, around 11,000 years ago, using house size as a proxy for calculating the Gini coefficient, a measure of wealth inequality. The study shows that, as may be expected, wealth inequality has generally increased. Unexpectedly however, inequality increased far more in the Old World of Europe and Asia than in the New (North and Central America). Even in highly urban New World sites, house sizes are generally similar. There are no enormous palaces, which one expects in Old World urban contexts. The authors suggest that the inherent wealth provided by large domesticated animals could explain the imbalance. Horses, for example, allowed people to ride around acquiring wealth from others. How wealth is distributed among households provides insight into the fundamental characters of societies and the opportunities they afford for social mobility1,2. However, economic inequality has been hard to study in ancient societies for which we do not have written records3,4, which adds to the challenge of placing current wealth disparities into a long-term perspective. Although various archaeological proxies for wealth, such as burial goods5,6 or exotic or expensive-to-manufacture goods in household assemblages7, have been proposed, the first is not clearly connected with households, and the second is confounded by abandonment mode and other factors. As a result, numerous questions remain concerning the growth of wealth disparities, including their connection to the development of domesticated plants and animals and to increases in sociopolitical scale8. Here we show that wealth disparities generally increased with the domestication of plants and animals and with increased sociopolitical scale, using Gini coefficients computed over the single consistent proxy of house-size distributions. However, unexpected differences in the responses of societies to these factors in North America and Mesoamerica, and in Eurasia, became evident after the end of the Neolithic period. We argue that the generally higher wealth disparities identified in post-Neolithic Eurasia were initially due to the greater availability of large mammals that could be domesticated, because they allowed more profitable agricultural extensification9, and also eventually led to the development of a mounted warrior elite able to expand polities (political units that cohere via identity, ability to mobilize resources, or governance) to sizes that were not possible in North America and Mesoamerica before the arrival of Europeans10,11. We anticipate that this analysis will stimulate other work to enlarge this sample to include societies in South America, Africa, South Asia and Oceania that were under-sampled or not included in this study.

Summary Plant allometries help us to understand resource allocation in plants and provide insight into how communities are structured. For woody species, a triangular allometric relationship between seed size and leaf size occurs in which all combinations are all possible, except for species with big seeds and small leaves (Cornelissen 1999). This relationship is thought to be a consequence of between habitat variation in abiotic conditions. In this study, we tested if the triangular relationship between seed mass and leaf area holds for annual species, and if variation in soil productivity and light (measured as Ellenberg indicator values: EIVs) are driving this relationship. We show that the triangular relationship also holds for annuals, which suggests that the allometric combinations between leaf area and seed mass are conserved across life-forms. The triangular relationship was driven by between-habitat variation in soil productivity. This means that as soil productivity increases, plants with bigger leaves could have either big or small seeds. However, in low soil productivity habitats, plants are constrained in their options, and plants with small leaves can only have small seeds. This article is protected by copyright. All rights reserved.

We use stable isotope analysis of crop, faunal and human remains to investigate agricultural strategies and diet at EBA-LBA Archontiko and MBA-LBA Thessaloniki Toumba. Crop production strategies varied between settlements, phases and species; flexibility is also apparent within the crop stores of individual houses. Escalating manuring intensity at LBA Thessaloniki Toumba coincides with large co-residential ‘blocks’ geared towards hoarding of agricultural surpluses, spectacularly preserved by fire at nearby LBA Assiros Toumba. Faunal isotope values reflect a range of feeding strategies, including probable herding of cattle on C4-rich coastal salt marshes, evident at Archontiko through to the LBA alongside bulk cockle harvesting. Palaeodietary analysis of LBA humans at Thessaloniki Toumba indicates that C3 crops represent the only plausible staples. Millet was a minor food but may have played a particular role in the sub-adult diet. Meat probably featured in supra-household food sharing and hospitality, associated with Mycenaean-style tableware in the LBA.