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Allotanaupodus williamsi Zhang & Fan, 2007: 8 — Holotype female (NZAC), in Malaise traps, Kawau I., AK, New Zealand (36.42°S, 174.85°E)., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 51, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Zhang, Z. Q. & Fan, Q. H. (2007) Allotanaupodidae, a new family of early derivative Parasitengona (Acari, Prostigmata). Zootaxa, 1517, 1 - 52."]}

LIU, DONG, YI, TIAN-CI, XU, YUN, ZHANG, ZHI-QIANG (2013): new mite species described during 2007 to 2012 3663. Zootaxa 3663 (1): 1-102, DOI: 10.11646/zootaxa.3663.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3663.1.1

Nanotanaupodus gracehallae Zhang & Fan, 2007: 44 — Holotype male (NZAC), Waenga Bush, Lottin Point Road, BP, New Zealand.

Allotanaupodus winksi Zhang & Fan, 2007: 26 — Holotype male (NZAC), Mt. Messenger, TK, New Zealand., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 51, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Zhang, Z. Q. & Fan, Q. H. (2007) Allotanaupodidae, a new family of early derivative Parasitengona (Acari, Prostigmata). Zootaxa, 1517, 1 - 52."]}

Allotanaupodus orete Zhang & Fan, 2007: 20 — Holotype male (NZAC), pit trap, Te Puia Hut, Orete Forest, BP, New Zealand., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 51, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Zhang, Z. Q. & Fan, Q. H. (2007) Allotanaupodidae, a new family of early derivative Parasitengona (Acari, Prostigmata). Zootaxa, 1517, 1 - 52."]}

Nanotanaupodus andrei Zhang & Fan, 2007: 33 — Holotype male (NZAC), from litter, moss and rotten wood, Waituhi Saddle, TO, New Zealand., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 51, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Zhang, Z. Q. & Fan, Q. H. (2007) Allotanaupodidae, a new family of early derivative Parasitengona (Acari, Prostigmata). Zootaxa, 1517, 1 - 52."]}

Nanotanaupodus gracehallae Zhang & Fan, 2007: 44 — Holotype male (NZAC), Waenga Bush, Lottin Point Road, BP, New Zealand., Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 51, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/5630595, {"references":["Zhang, Z. Q. & Fan, Q. H. (2007) Allotanaupodidae, a new family of early derivative Parasitengona (Acari, Prostigmata). Zootaxa, 1517, 1 - 52."]}

Nanotanaupodus andrei sp. nov. (Figs 21���29) Description Adult female. Idiosoma elongate-ovoid; length 788 ��m, maximum width 490 ��m. Sejugal furrow clearly evident, hysteronotum almost three times as long as prodosum. Prodorsum. Almost half as long (200 ��m) as wide (370 ��m). Prodorsal plate longer (190 ��m) than wide (160 ��m), not reaching anterior end of propodosoma, with 13���14 lanceolate setae on each side, anteriormost pair 40 ��m long, posteriormost pair 32 long. Crista metopica linear, rod-like, 155 ��m long, widest anteriorly, ca 10 ��m. A pair of lanceolate setae posterolateral to prodorsal plate, each on its platelet and ca 40 ��m long. A pair of posterolateral plates each bearing 5 setae of ca 40 long ��m. Eyes absent. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D by evident furrow. C row with medial setae each on a platelet, but lateral setae clustered on a pair of plates; setae 30���40 ��m long, all lanceolate. D row with medial setae (30 ��m long) each on a platelet, a pair of sublateral plates each with three setae (35���40 ��m long) and a pair of lateral plates each with 6���9 setae (35���40 ��m long). E row with a pair of lateral plates each with lanceolate 11���12 setae (35���45 ��m long) and many small lanceolate setae each on a platelet between them. F and H rows both with similar setal and plate arrangements as in E. PS row with a pair of plates each bearing numerous lanceolate setae mostly over 40 ��m long. Idiosomal venter. Intercoxal area of leg I with eight short setae (20���30 ��m long). Intercoxal area of leg II with six short setae. Intercoxal area of leg III with nine pairs of setae and that of leg IV with three pairs. Coxal I with 14���15 setae, 1 a (30 ��m) not much longer than other secondary setae. Coxa II with 14 setae. Coxa III with 12 setae. Coxa IV with eight setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 100 ��m long. A pair of central valves each bearing six setae of similar size (shorter than 20 ��m). A pair of lateral valves each bearing 5���6 setae of similar size (shorter than 20 ��m long). Eugenital setae absent. Anal opening 93 ��m long, its valves each with 3 setae (over 20 ��m long). Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 505 ��m. Trochanter I with three long setulose setae and two short ones. Basifemur I 80 ��m. Telofemur I 77 ��m long, without solenidia. Genu I 82 ��m long, with numerous setulose setae and 1 solenidion. Tibia I 90 ��m long, with numerous setulose setae and four solenidia. Tarsus I over twice as long (165 ��m) as wide (70 ��m), with many setulose setae and solenidia. Leg II 290 ��m. Trochanter II with three long setae and two shorter setae. Basifemur II 60 ��m long, with nine setulose setae. Telofemur II 50 ��m long with 10���12 setulose setae. Genu II 50 ��m long with numerous setulose setae and one solenidion. Tibia II 50 ��m long, with numerous setulose setae and one solenidion. Tarsus II 78 ��m long with many strongly setulose setae, two subdistal solenidia and one basal solenidion. Leg III 340 ��m. Trochanter III with three long setulose setae and three shorter ones. Basifemur III 62 ��m long with 10 setulose setae. Telofemur III 50 ��m long with 10���12 setulose setae. Genu III 58 ��m long with numerous setulose setae and one solenidion. Tibia III 60 ��m long with numerous setulose setae and two solenidia. Tarsus III 92 ��m long with many strongly setoluse setae and one solenidion. Leg IV 450 ��m. Trochanter IV with six setulose setae. Basifemur IV 80 ��m long with eight setulose setae. Telofemur IV 64 ��m long with 12 setulose setae. Genu IV 80 ��m long with numerous setulose setae and two solenidia. Tibia IV 80 ��m long with numerous setulose setae and two solenidia. Tarsus IV 110 ��m long with many strongly setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae: or 1 20 ��m, or 2 20 ��m. Cheliceral base 85 ��m long, cheliceral blade 38 ��m long. Palp 200 ��m long. Palptrochanter with two smooth setae, ca 40 ��m long. Palpfemur with seven setae. Palpgenu with seven setae. Palptibia with five nude setae, an enlarged seta (15 ��m) near base of palptarsus and a strong terminal setae (i.e. tibial claw, 25 ��m long), ratio of tibial length excluding tibial claw and maximum width 1.9. Palptarsus with a basal solenidion (10 ��m long), four nude setiform setae, one short spinelike seta and three terminal eupathidia. Deutonymph. Idiosoma elongate-ovoid; length 600���645 ��m, maximum width 370���375 ��m. Sejugal furrow clearly evident in dorsomedial portion, hysteronotum almost three times as long as prodosum. Prodorsum. Almost half as long (167���175 ��m) than wide (310���322 ��m). Prodorsal plate longer (131 ��m) than wide (106 ��m), not reaching anterior end of propodosoma, with 7 lanceolate setae each side, anteriormost pair 30 ��m long, posteriormost pair 30 ��m long. Crista metopica linear, rod-like, 110���118 ��m long, widest anteriorly, ca 8 ��m. A pair of lanceolate setae posterolateral to prodorsal plate, each on its platelet and ca 32���35 ��m long. A pair of posterolateral plates each bearing three setae of ca 32���40 ��m long. Eyes absent. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D evident furrow. C row with medial setae each on a platelet, but lateral setae clustered on a pair of plates; setae 23���35 ��m long, all lanceolate. D row with medial setae (20���25 ��m long) each on a platelet, a pair of sublateral plates each with 2���3 setae (30 ��m long) and a pair of lateral plates each with 5���6 setae (30���35 ��m long). E row with a pair of lateral plates each with 5���6 lanceolate setae (30���38 ��m long) and numerous lanceolate setae on a platelet between them. F row with similar setal and plate arrangement as in E, but only several setae between plates. H row with similar setal and plate arrangement as in E, but only a few setae between plates. PS row with a pair of plates each bearing numerous lanceolate setae mostly over 40 ��m long. Idiosomal venter. Intercoxal area of leg I with four short setae. Intercoxal area of leg II with four short setae. Intercoxal area of leg III with four pairs of setae and that of leg IV with three pairs. Coxal I with 8���10 setae, 1 a (25 ��m) not much longer than other setae. Coxa II with 10���12 setae. Coxa III with 7���10 setae. Coxa IV with 4���5 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 60 ��m long. A pair of central valves each bearing three setae of similar size (10���12 ��m long). A pair of lateral valves each bearing 1���2 setae of similar size (12���15 ��m long). Eugenital setae absent. Anal opening 65 ��m long, its valves each without setae. Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 363���375 ��m. Trochanter I with 2 long well-setulose setae and 2 short, nearly smooth setae. Basifemur I 55 ���58 ��m. Telofemur I 50 ���55 ��m. Genu I 58 ���62 ��m long, with one solenidion (15 ��m long). Tibia I 62 ��m long, with numerous setulose setae and 4���5 solenidia (15 ��m long). Tarsus I over twice as long (120���125 ��m) as wide (50���58 ��m), with many setulose setae and solenidia. Leg II 225 ��m. Trochanter II with two long setae and one short seta. Basifemur II 42 ��m long with five setulose setae. Telofemur II 35 ��m long with six setulose setae. Genu II 37 ��m long, with numerous setulose setae and one solenidion near the base of segment. Tibia II 40 ��m long, with numerous setulose setae and a single solenidion near the base of segment. Tarsus II 66 ��m long, with many setulose setae and no solenidia. Leg III 265���270 ��m. Trochanter III with three long setulose setae and two shorter ones. Basifemur III 40 ��m long with seven setulose setae. Telofemur III 35 ��m long with six setulose setae. Genu III 40 ��m long with numerous setulose setae and one solenidion. Tibia III 43 ��m long with numerous setulose setae and two solenidia. Tarsus III 73 ��m long with many strongly setulose setae. Leg IV 310���325 ��m. Trochanter IV with four setulose setae and a shorter one. Basifemur IV 48 ��m long with three setulose setae. Telofemur IV 46 ��m long with 7 setulose setae. Genu IV 55 ��m long with numerous setulose setae and two solenidia. Tibia IV 60 ��m long with numerous setulose setae and two solenidia. Tarsus IV 86 ��m long with many strongly setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae or 1 15 ��m, or 2 20 ��m. Subcapitular setae absent. Cheliceral body 90 ��m long, cheliceral blade 30���33 ��m long. Palp 140���150 ��m long. Palptrochanter with one smooth seta, ca 30 long. Palpgenu with six or five setae. Palpgenu with four setae. Palptibia with three nude setae, an enlarged seta (10���12 ��m) long near the base of palptarsus and a strong terminal setae (i.e. tibial claw, 20���25 ��m long), ratio of tibial length excluding tibial claw and maximum width 1.3���1.5. Palptarsus with a basal solenidion (8 ��m long), four nude setiform setae, one short spine-like setae and three terminal eupathidia (15 ��m long). Etymology Named in memory of Marc Andr�� for his many contributions to systematics of Trombidiina. Type specimens Holotype male and paratype deutonymph (on the same slide ZQZ 2002 -01231A,B,C,D: C= holotype; D= paratype), New Zealand, TO, Waituhi Saddle, from litter, moss and rotten wood, 16 Nov. 1983, C.F. Butcher (litter 83 / 112). Deposited in NZAC., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 33-43, DOI: 10.5281/zenodo.177319

Allotanaupodus orete sp. nov. (Figs 11–15) Description Adult male. Idiosoma ovoid; length 1500 µm, maximum width 950 µm. Sejugal furrow present. Hysteronotum about twice as long as prodorsum. Prodorsum. Much wider than long. Prodorsal plate poorly defined due to over-maceration of the specimen, about as long as wide, with 14 pairs of lanceolate setae each side, ca 60 µm long. Crista metopica linear, rod-like, 287 µm long, widest anteriorly, ca 30 µm. Four pairs of lanceolate setae posterolateral to prodorsal plate (one of them laterad of eyes), each on a platelet, ca 75 µm long. Prodorsal lateral plates absent. Eye plates 95 x 50 µm, diameter of eyes 32 µm. Hysterosomal dorsum. Hysterosomal segmentation obscured in this over-macerated specimen. C row with medial setae each on a platelet, but lateral setae clustered on a pair of large plates; setae 70–80 µm long, all slightly lanceolate. D–E row similar to C row. H row with setae over 100 µm long. Idiosomal venter. Intercoxal area of leg I with 14 short setae (ca 60 µm long; posterolateral pair the longest, 1 a over 100 µm long), intercoxal area of leg II with four pairs of setae. Intercoxal area of leg III with ca 13 pairs of setae and that of leg IV with numerous setae not separable from those anterior to genital area. Coxal I with 29–32 setae. Coxa II with 24–25 setae. Coxa III with 20–21 setae. Coxa IV with 20–23 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 230 µm long. A pair of central valves each bearing six setae (45 µm long). A pair of lateral valves each bearing 10 or 11 setae of similar size. Eugenital setae three pairs, all nude, setiform (ca 30 µm long). Anal opening 200 µm long, its valves each with three pairs of setae (45 µm long). Legs. Lengths of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 1465 µm. Trochanter I with 18–19 setulose setae. Basifemur I 190 µm, with numerous setulose setae. Telofemur I 235 µm with numerous setulose setae and one distal solenidion. Genu I 270 µm, with numerous setulose setae, two solenidia and one microseta (35 µm long). Tibia I 290 long, with numerous setulose setae and solenidia, and one microseta (33 µm long). Tarsus I over twice as long (380 µm) as wide (150 µm), with many setulose setae and solenidia. Leg II 920 µm. Trochanter II with 13–14 setae. Basifemur II 140 long, with numerous setulose setae and one subdistal solenidion. Telofemur II 145 µm long with numerous setulose setae and a distal solenidion. Genu II 156 µm long, with numerous setulose setae, two solenidia and one distal microseta (32 µm long). Tibia II 168 µm long, with numerous setulose setae and two solenidia. Tarsus II 225 µm long, with numerous setulose setae, four dorsal solenidia, one famulus (13 µm long), one ventral solenidion and two distal eupathidia. Leg III 1050 µm. Trochanter III with 20–21 setulose setae. Basifemur III 165 µm long with numerous setae. Telofemur III 145 µm long with numerous setulose setae and a distal solenidion. Genu III 180 µm long, with numerous setulose setae and three solenida. Tibia III 210 µm long, with numerous setulose setae and two solenidia. Tarsus III 260 µm long, with numerous setulose setae and two solenidia. Leg IV 1390 µm. Trochanter IV with 29 setae. Basifemur IV 200 µm long with numerous setae. Telofemur IV 180 µm long with numerous setae and a distal solenidion (40 µm long). Genu IV 250 µm long with numerous setulose setae and four solenidia. Tibia IV 295 µm long with numerous setulose setae and four solenidia. Tarsus IV 300 µm long with many setulose setae and 1 solenidion. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae and a pair of subcapitular setae. Adoral: setae or 1 40 µm, or 2 10 µm, or 3 40 µm. Subcapitular setae 60 µm. Cheliceral base 160 µm long, cheliceral blade 88 µm long. Palp 680 µm long. Palptrochanter with 2 setae, 80 µm and 100 µm long. Palpfemur with 11 setae. Palpgenu with 16 setae. Palptibia with 16 nude setae, a strong terminal seta (i.e. tibial claw, 45 µm long) and a ventral enlarged seta (35 µm) near base of tibial claw. Ratio of tibial length excluding tibial claw to maximum width 3.0. Palptarsus with a basal solenidion (18 µm long), two nude setiform setae, and six terminal eupathidia (22 µm long). Etymology After the name of the forest at the type locality; a noun in apposition. Type specimens Holotype male, New Zealand, BP: Orete Forest, Te Puia Hut, 19 Oct to 23 Nov. 1992, pit trap, G. Hall. Deposition of type NZAC. Remarks This new species is similar to A. williamsi sp. nov. Key differences between this species and A. williamsi sp. nov. and A. winksi sp. nov. are presented table 1.

A new family of early derivative Parasitengona (Acari: Prostigmata), Allotanaupodidae fam. nov., is described from New Zealand based on adults and deutonymphs of two new genera and five new species and a new superfamily Allotanaupodoidea is erected to accommodate it. The new family is characterized by the absence of prodorsal trichobothria and sensory areas, the presence of one or two pairs of plates with multiple setae on C to PS rows of dorsal hysterosoma, the presence of only two pairs of genital acetabula in adults, and short, distally inserted palptarsus on the palptibia. The family consists of two subfamilies (Allotanaupodinae subfam. nov. and Paratanaupodinae subfam. nov.), with the former endemic to New Zealand. Allotanaupodinae subfam. nov. has a single genus, Allotanaupodus gen. nov., which is represented by three new species from New Zealand: Allotanaupodus williamsi sp. nov. (type species) from Kawau I., Auckland, Allotanaupodus orete sp. nov. from Orete Forest, Te Puia Hut and Allotanaupodus winksi sp. nov. from Mt. Messenger, Taranaki. The eyeless Paratanaupodinae subfam. nov. consists of two genera: Nanotanaupodus gen. nov. and Paratanaupodus Andre & Lelievre-Farjon, 1960. The type genus Paratanaupodus Andre & Lelievre-Farjon, 1960 was previously placed in the Tanaupodidae and is represented by a single species, Paratanaupodus insensus Andre & Lelievre-Farjon, 1960, from South America. Nanotanaupodus gen. nov. is represented by two new species from New Zealand: Nanotanaupodus andrei sp. nov. (type species) from Waituhi Saddle and Nanotanaupodus gracehallae sp. nov. from Orete Forest, Te Puia Hut. A key to superfamilies of terrestrial Parasitengona (post-larval stages) is provided, along with keys to subfamilies, genera and species of the new family.

Key to subfamilies and genera of Allotanaupodidae (adults and deutonymphs) 1 A pair of eyes present lateral to prodorsal sclerite; subcapitular setae present in addition to two or three pairs of adoral setae (in adults); telofemur of legs I–IV with a solenidion; palptibia elongate, more than three times as long as “tibial claw”...............................................................Allotanaupodinae subfam. nov........................................................................................................ (single genus Allotanaupodus gen. nov.) - Eyes absent; subcapitulum with only two pairs of adoral setae, subcapitular setae absent posterior to insertion of palptrochanter (in adults); telofemur of legs I–IV without any solenidia; palptibia not elongate, obviously less than three times as long as “tibial claw”.......................Paratanaupodinae subfam. nov.... 2 2 All dorsal hysterosomal setae on individual small platelets; palptibia neither elongate nor shortened, about twice as long as “tibial claw”; palptarsus with four terminal eupathidia; palptarsus without a thick and short spine-like seta; leg IV longer than leg I ................... Paratanaupodus André & Lelievre-Farjon, 1960 Only some dorsal hysterosomal setae on individual small platelets; C, D, E, F, H and PS rows each with at least 1 pair of plates with multiple setae; palptibia relatively short, obviously less than twice as long as “tibial claw”; palptarsus with three terminal eupathidia; palptarsus with a thick and short spine-like seta; leg I longer than leg IV...................................................................................... Nanotanaupodus gen. nov.

Nanotanaupodus gen. nov. Diagnosis Adults and deutonymphs: Paratanaupodinae with a pair of posterolateral plates on prodorsum, each bearing multiple setae. Palptibia not elongate, less than twice as long as tibial claw. Palptibia with a distal-ventral hypertrophied seta located near base of tarsus in addition to terminal tibial-claw. Palptarsus short and inserted ventrally near base of tibia. Palptarsus with a spine-like seta and a cluster of 3 eupathidia terminally. Larvae not known. Type species Nanotanaupodus andrei sp. nov. (by original designation). Etymology From Nano (small) and the root Tanaupodus; masculine in gender. Remarks This genus is most similar to Paratanaupodus. Both genera lack eyes and trichobothria. The two can be distinguished by a number of features listed in the key to genera above., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on page 33, DOI: 10.5281/zenodo.177319

Allotanaupodus winksi sp. nov. (Figs 16–20) Description Deutonymph. Idiosoma ovoid; length 750–950 µm, maximum width 525–650 µm. Sejugal furrow clearly evident. Hysteronotum only slightly longer than prodosum. Prodorsum. About half as long (280 µm) as wide (400 µm). Prodorsal plate longer (295 µm) than wide (220 µm), extending beyond anterior edge of propodosoma, with 7–9 pairs of lanceolate setae on each side, anteriormost pair 95–100 µm long. Crista metopica linear, rod-like, 205–220 µm long, widest anteriorly, ca 10–12 µm. 2–3 pairs of lanceolate setae posterolateral to prodorsal plate (one of them laterad of eyes), each on a platelet, ca 65–75 µm long. Prodorsal lateral plate absent. Eye plates 63– 65 x 33–39 µm, diameter 25–30 µm for anterior eyes and 20–25 for posterior eyes. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D by evident furrow. C row with a pair of sublateral plates each bearing 7–8 setae; other setae each on a platelet. D row with a pair of sublateral plates each bearing three pairs of setae; lateral and medial setae each on a platelet. E row with a pair of sublateral plates each bearing three setae; lateral and medial setae each on a platelet. F row with a pair of sublateral plates each bearing four setae, other setae each on a platelet. H segment with two pairs of plates each bearing three pairs of setae; a few lateral and medial setae each on a platelet. PS segment with a pair of terminal plates each bearing 5–6 setae. Idiosomal venter. Intercoxal area of leg I with 3–4 pairs of short setae (1 a over 50 µm long). Intercoxal area of leg II with three pairs of setae. Intercoxal area of leg III with 3–4 pairs of setae and that of leg IV with 4–5 setae. Coxal I with 9–12 setae. Coxa II with 6–9 setae. Coxa III with 5–8 setae. Coxa IV with 6–10 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 100 µm long; a pair of central valves each bearing 2–3 setae (25 µm long); a pair of lateral valves each bearing 1–3 setae of similar size. Eugenital setae absent. Anal opening 125–130 µm long, its valves each with 4–7 pairs of setae. Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 821–852 µm. Trochanter I with 6–7 setae. Basifemur I 95 –105 µm, with numerous setulose setae. Telofemur I 118–120 with numerous setulose setae and one subdistal solenidion. Genu I 140–150 µm long, with numerous setulose setae, two solenidia and one microseta (32 µm long). Tibia I 163–175 µm long with numerous setulose setae and solenidia, and 1 microseta (26 µm long). Tarsus I more than twice as long (235– 238 µm) as wide (90–100 µm), with many setulose setae and solenidia; famulus not visible. Leg II 520–540 µm. Trochanter II with 6 setae. Basifemur II 80 µm long with numerous setulose setae. Telofemur II 82 µm long with numerous setulose setae and a distal solenidion. Genu II 86 µm long, with numerous setulose setae, two solenidia and 1 microseta (20 µm long). Tibia II 90 µm long, with numerous setulose setae and two solenidia. Tarsus II 140 µm long, with numerous setulose setae, four dorsal solenidia, and two distal eupathidia. Leg III 535–635 µm. Trochanter III with 8 setae. Basifemur III 90 µm long with numerous setae. Telofemur III 82 µm long with numerous setulose setae and a distal solenidion. Genu III 105 µm long with numerous setulose setae and two solenidia. Tibia III 125 µm long with numerous setulose setae and 2 solenida. Tarsus III 160 µm long, with numerous setulose setae and two solenidia. Leg IV 660–758 µm. Trochanter IV with 7 setulose setae. Basifemur IV 130 µm long, with numerous setae. Telofemur IV 107 µm long, with numerous setae and 1 distal solenidion (28 µm long). Genu IV 155 µm long, with numerous setulose setae and 3 solenidia. Tibia IV 180 µm long with numerous setulose setae and three solenidia, of which the distal one is twice as long as others. Tarsus IV 195 µm long with many setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae: or 1 32–35 µm and or 2 25 µm. Cheliceral base 160–175 µm long, cheliceral blade 55–58 µm long. Palp 375–415 µm long. Palptrochanter with one smooth seta, ca 50 long. Palpfemur with seven setae. Palpgenu with eight setae. Palptibia with seven nude setae, an enlarged seta (20 µm) near base of palptarsus and a strong terminal setae (i.e. tibial claw, 32 – 23 µm long), ratio of tibial length excluding tibial claw and maximum width 2.2–2.4. Palptarsus with a basal solenidion (14 µm long), 3 nude setiform setae and five terminal eupathidia (20 µm long). Etymology Named after the collector, Chris Winks (invertebrate ecologist at Landcare Research, Auckland). Remarks This new species differs from the other two species by a number of differences in morphology (see table 1). The difference in the shape of prodorsal plate, for example, is not likely to be due to ontogeny as nymphs and adults of Nanotanaupodus andrei sp. nov. have similar dorsal plates.

Key to subfamilies and genera of Allotanaupodidae (adults and deutonymphs) 1 A pair of eyes present lateral to prodorsal sclerite; subcapitular setae present in addition to two or three pairs of adoral setae (in adults); telofemur of legs I���IV with a solenidion; palptibia elongate, more than three times as long as ���tibial claw���...............................................................Allotanaupodinae subfam. nov........................................................................................................ (single genus Allotanaupodus gen. nov.) - Eyes absent; subcapitulum with only two pairs of adoral setae, subcapitular setae absent posterior to insertion of palptrochanter (in adults); telofemur of legs I���IV without any solenidia; palptibia not elongate, obviously less than three times as long as ���tibial claw���.......................Paratanaupodinae subfam. nov.... 2 2 All dorsal hysterosomal setae on individual small platelets; palptibia neither elongate nor shortened, about twice as long as ���tibial claw���; palptarsus with four terminal eupathidia; palptarsus without a thick and short spine-like seta; leg IV longer than leg I ................... Paratanaupodus Andr�� & Lelievre-Farjon, 1960 Only some dorsal hysterosomal setae on individual small platelets; C, D, E, F, H and PS rows each with at least 1 pair of plates with multiple setae; palptibia relatively short, obviously less than twice as long as ���tibial claw���; palptarsus with three terminal eupathidia; palptarsus with a thick and short spine-like seta; leg I longer than leg IV...................................................................................... Nanotanaupodus gen. nov., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on page 6, DOI: 10.5281/zenodo.177319, {"references":["Andre, M. & Lelievre-Farjon, J. (1960) Un nouveau genre sud-Americain de Tanaupodinae (Thrombidion) (Paratanaupodus insensus n. gen., n. sp.). Acarologia, 2, 466 - 468."]}

Allotanaupodus winksi sp. nov. (Figs 16���20) Description Deutonymph. Idiosoma ovoid; length 750���950 ��m, maximum width 525���650 ��m. Sejugal furrow clearly evident. Hysteronotum only slightly longer than prodosum. Prodorsum. About half as long (280 ��m) as wide (400 ��m). Prodorsal plate longer (295 ��m) than wide (220 ��m), extending beyond anterior edge of propodosoma, with 7���9 pairs of lanceolate setae on each side, anteriormost pair 95���100 ��m long. Crista metopica linear, rod-like, 205���220 ��m long, widest anteriorly, ca 10���12 ��m. 2���3 pairs of lanceolate setae posterolateral to prodorsal plate (one of them laterad of eyes), each on a platelet, ca 65���75 ��m long. Prodorsal lateral plate absent. Eye plates 63��� 65 x 33���39 ��m, diameter 25���30 ��m for anterior eyes and 20���25 for posterior eyes. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D by evident furrow. C row with a pair of sublateral plates each bearing 7���8 setae; other setae each on a platelet. D row with a pair of sublateral plates each bearing three pairs of setae; lateral and medial setae each on a platelet. E row with a pair of sublateral plates each bearing three setae; lateral and medial setae each on a platelet. F row with a pair of sublateral plates each bearing four setae, other setae each on a platelet. H segment with two pairs of plates each bearing three pairs of setae; a few lateral and medial setae each on a platelet. PS segment with a pair of terminal plates each bearing 5���6 setae. Idiosomal venter. Intercoxal area of leg I with 3���4 pairs of short setae (1 a over 50 ��m long). Intercoxal area of leg II with three pairs of setae. Intercoxal area of leg III with 3���4 pairs of setae and that of leg IV with 4���5 setae. Coxal I with 9���12 setae. Coxa II with 6���9 setae. Coxa III with 5���8 setae. Coxa IV with 6���10 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 100 ��m long; a pair of central valves each bearing 2���3 setae (25 ��m long); a pair of lateral valves each bearing 1���3 setae of similar size. Eugenital setae absent. Anal opening 125���130 ��m long, its valves each with 4���7 pairs of setae. Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 821���852 ��m. Trochanter I with 6���7 setae. Basifemur I 95 ���105 ��m, with numerous setulose setae. Telofemur I 118���120 with numerous setulose setae and one subdistal solenidion. Genu I 140���150 ��m long, with numerous setulose setae, two solenidia and one microseta (32 ��m long). Tibia I 163���175 ��m long with numerous setulose setae and solenidia, and 1 microseta (26 ��m long). Tarsus I more than twice as long (235��� 238 ��m) as wide (90���100 ��m), with many setulose setae and solenidia; famulus not visible. Leg II 520���540 ��m. Trochanter II with 6 setae. Basifemur II 80 ��m long with numerous setulose setae. Telofemur II 82 ��m long with numerous setulose setae and a distal solenidion. Genu II 86 ��m long, with numerous setulose setae, two solenidia and 1 microseta (20 ��m long). Tibia II 90 ��m long, with numerous setulose setae and two solenidia. Tarsus II 140 ��m long, with numerous setulose setae, four dorsal solenidia, and two distal eupathidia. Leg III 535���635 ��m. Trochanter III with 8 setae. Basifemur III 90 ��m long with numerous setae. Telofemur III 82 ��m long with numerous setulose setae and a distal solenidion. Genu III 105 ��m long with numerous setulose setae and two solenidia. Tibia III 125 ��m long with numerous setulose setae and 2 solenida. Tarsus III 160 ��m long, with numerous setulose setae and two solenidia. Leg IV 660���758 ��m. Trochanter IV with 7 setulose setae. Basifemur IV 130 ��m long, with numerous setae. Telofemur IV 107 ��m long, with numerous setae and 1 distal solenidion (28 ��m long). Genu IV 155 ��m long, with numerous setulose setae and 3 solenidia. Tibia IV 180 ��m long with numerous setulose setae and three solenidia, of which the distal one is twice as long as others. Tarsus IV 195 ��m long with many setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae: or 1 32���35 ��m and or 2 25 ��m. Cheliceral base 160���175 ��m long, cheliceral blade 55���58 ��m long. Palp 375���415 ��m long. Palptrochanter with one smooth seta, ca 50 long. Palpfemur with seven setae. Palpgenu with eight setae. Palptibia with seven nude setae, an enlarged seta (20 ��m) near base of palptarsus and a strong terminal setae (i.e. tibial claw, 32 ��� 23 ��m long), ratio of tibial length excluding tibial claw and maximum width 2.2���2.4. Palptarsus with a basal solenidion (14 ��m long), 3 nude setiform setae and five terminal eupathidia (20 ��m long). Etymology Named after the collector, Chris Winks (invertebrate ecologist at Landcare Research, Auckland). Remarks This new species differs from the other two species by a number of differences in morphology (see table 1). The difference in the shape of prodorsal plate, for example, is not likely to be due to ontogeny as nymphs and adults of Nanotanaupodus andrei sp. nov. have similar dorsal plates., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 26-32, DOI: 10.5281/zenodo.177319

Allotanaupodus orete sp. nov. (Figs 11���15) Description Adult male. Idiosoma ovoid; length 1500 ��m, maximum width 950 ��m. Sejugal furrow present. Hysteronotum about twice as long as prodorsum. Prodorsum. Much wider than long. Prodorsal plate poorly defined due to over-maceration of the specimen, about as long as wide, with 14 pairs of lanceolate setae each side, ca 60 ��m long. Crista metopica linear, rod-like, 287 ��m long, widest anteriorly, ca 30 ��m. Four pairs of lanceolate setae posterolateral to prodorsal plate (one of them laterad of eyes), each on a platelet, ca 75 ��m long. Prodorsal lateral plates absent. Eye plates 95 x 50 ��m, diameter of eyes 32 ��m. Hysterosomal dorsum. Hysterosomal segmentation obscured in this over-macerated specimen. C row with medial setae each on a platelet, but lateral setae clustered on a pair of large plates; setae 70���80 ��m long, all slightly lanceolate. D���E row similar to C row. H row with setae over 100 ��m long. Idiosomal venter. Intercoxal area of leg I with 14 short setae (ca 60 ��m long; posterolateral pair the longest, 1 a over 100 ��m long), intercoxal area of leg II with four pairs of setae. Intercoxal area of leg III with ca 13 pairs of setae and that of leg IV with numerous setae not separable from those anterior to genital area. Coxal I with 29���32 setae. Coxa II with 24���25 setae. Coxa III with 20���21 setae. Coxa IV with 20���23 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 230 ��m long. A pair of central valves each bearing six setae (45 ��m long). A pair of lateral valves each bearing 10 or 11 setae of similar size. Eugenital setae three pairs, all nude, setiform (ca 30 ��m long). Anal opening 200 ��m long, its valves each with three pairs of setae (45 ��m long). Legs. Lengths of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 1465 ��m. Trochanter I with 18���19 setulose setae. Basifemur I 190 ��m, with numerous setulose setae. Telofemur I 235 ��m with numerous setulose setae and one distal solenidion. Genu I 270 ��m, with numerous setulose setae, two solenidia and one microseta (35 ��m long). Tibia I 290 long, with numerous setulose setae and solenidia, and one microseta (33 ��m long). Tarsus I over twice as long (380 ��m) as wide (150 ��m), with many setulose setae and solenidia. Leg II 920 ��m. Trochanter II with 13���14 setae. Basifemur II 140 long, with numerous setulose setae and one subdistal solenidion. Telofemur II 145 ��m long with numerous setulose setae and a distal solenidion. Genu II 156 ��m long, with numerous setulose setae, two solenidia and one distal microseta (32 ��m long). Tibia II 168 ��m long, with numerous setulose setae and two solenidia. Tarsus II 225 ��m long, with numerous setulose setae, four dorsal solenidia, one famulus (13 ��m long), one ventral solenidion and two distal eupathidia. Leg III 1050 ��m. Trochanter III with 20���21 setulose setae. Basifemur III 165 ��m long with numerous setae. Telofemur III 145 ��m long with numerous setulose setae and a distal solenidion. Genu III 180 ��m long, with numerous setulose setae and three solenida. Tibia III 210 ��m long, with numerous setulose setae and two solenidia. Tarsus III 260 ��m long, with numerous setulose setae and two solenidia. Leg IV 1390 ��m. Trochanter IV with 29 setae. Basifemur IV 200 ��m long with numerous setae. Telofemur IV 180 ��m long with numerous setae and a distal solenidion (40 ��m long). Genu IV 250 ��m long with numerous setulose setae and four solenidia. Tibia IV 295 ��m long with numerous setulose setae and four solenidia. Tarsus IV 300 ��m long with many setulose setae and 1 solenidion. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae and a pair of subcapitular setae. Adoral: setae or 1 40 ��m, or 2 10 ��m, or 3 40 ��m. Subcapitular setae 60 ��m. Cheliceral base 160 ��m long, cheliceral blade 88 ��m long. Palp 680 ��m long. Palptrochanter with 2 setae, 80 ��m and 100 ��m long. Palpfemur with 11 setae. Palpgenu with 16 setae. Palptibia with 16 nude setae, a strong terminal seta (i.e. tibial claw, 45 ��m long) and a ventral enlarged seta (35 ��m) near base of tibial claw. Ratio of tibial length excluding tibial claw to maximum width 3.0. Palptarsus with a basal solenidion (18 ��m long), two nude setiform setae, and six terminal eupathidia (22 ��m long). Etymology After the name of the forest at the type locality; a noun in apposition. Type specimens Holotype male, New Zealand, BP: Orete Forest, Te Puia Hut, 19 Oct to 23 Nov. 1992, pit trap, G. Hall. Deposition of type NZAC. Remarks This new species is similar to A. williamsi sp. nov. Key differences between this species and A. williamsi sp. nov. and A. winksi sp. nov. are presented table 1., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 20-26, DOI: 10.5281/zenodo.177319

Nanotanaupodus gracehallae sp. nov. (Figs 30���34) Adult male Idiosoma elongate, slightly narrowed between bases of legs III and IV; length 550 ��m, maximum width 315 ��m. Sejugal furrow clearly evident, hysteronotum about twice as long as prodosum (Fig. 30). Prodorsum. About two-thirds as long (160 ��m) as wide (250 ��m). Prodorsal plate much longer (185 ��m) than wide (125 ��m), extending anteriorly beyond anterior edge of propodosoma, with 27 lanceolate setae on each side, anteriormost pair 40 ��m long, posteriormost pair 32 ��m long. Crista metopica linear, rod-like, 140 ��m long, widest anteriorly, ca 7 ��m. A pair of lanceolate setae (40 ��m) posterolateral to prodorsal plate, each on a platelet. A pair of posterolateral plate each bearing 9 setae of ca 35���40 ��m long. Eyes absent. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae. C row with medial setae each on a platelet, but lateral setae clustered on a pair of large plates; setae 32���40 ��m long, all lanceolate. D row with medial setae (30���35 ��m long) each on a platelet, a pair of sublateral plates each with 3 setae (32���40 ��m long) and a pair of lateral plates each with 11���12 setae (35���40 ��m long). E row with a pair of lateral plates each with 8 lanceolate setae (35���40 ��m long) and numerous lanceolate setae each on a platelet between them (a couple of them have joined platelets) and also a few lateral to them above trochanter III. F row with similar setal and plate arrangement as in E, but only several setae between the pair of large plates. H row with a pair of lateral plates each with 18 setae and about 12 setae between them each on a platelet. PS row with a pair of large plates each bearing numerous lanceolate setae mostly 50 ��m long; the pair of plates ventrally touching each other. Idiosomal venter. Intercoxal area of leg I with 4 short setae (20���34 ��m long; posterolateral pair 1a 25 ��m long, on its own platelet). Intercoxal area of leg II with 3���4 pairs of setae on a pair of plates. Intercoxal area of leg III with 6 pairs of setae and that of leg IV with several numerous setae not separable from those anterior to genital area. Coxal I with 13���15 setae. Coxa II with 14 setae. Coxa III with 9 setae. Coxa IV with 8���9 setae. Opisthoventer with a large plate enclosing anus and genital area (240 ��m long and 180 ��m wide), with numerous short setiform setae similar to coxal setae. Genital opening 78 ��m long. A pair of central valves each bearing 7���8 setae, anterior and posterior ones setiform, but median 3 pairs thickened and branched distally. A pair of lateral valves each bearing 8���9 setae of similar size (up to 20 ��m long). Five pairs of eugenital setae, all nude, setiform, the posterior three pairs relatively close together (Fig. 33 B). Anal opening 85 ��m long, its valves each with a pair of setae. Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 494 ��m. Trochanter I with five relatively long setae with strong setulose and two shorter setae. Basifemur I 70 ��m. Telofemur I 75 ��m. Genu I 85 ��m long, with numerous setulose setae and one solenidion. Tibia I 88 ��m long, with numerous setulose setae and four solenidia. Tarsus I about twice as long (156 ��m) as wide (80 ��m), with many setulose setae and solenidia. Leg II 272 ��m. Trochanter II with three long setulose setae and two short nude setae. Basifemur II 52 ��m long with 10 setulose setae. Telofemur II 45 ��m long with 11 setulose setae. Genu II 50 ��m long, with numerous setulose setae and 1 solenidion. Tibia II 48 ��m long with numerous setulose setae and one solenidion. Tarsus II 85 ��m long, with many strongly setulose setae, two subdistal solenidia and one basal solenidion. Leg III 315 ��m. Trochanter III with seven setulose setae. Basifemur III 60 ��m long with eight setulose setae. Telofemur III 45 ��m long with 11 setulose setae. Genu III 55 ��m long with numerous setulose setae and one solenidion. Tibia III 53 ��m long with numerous setulose setae and two solenida. Tarsus III 90 ��m long, with many strongly setulose setae and two solenidia. Leg IV 405 ��m. Trochanter IV with seven setulose setae and one short nude seta. Basifemur IV 60 ��m long with six setulose setae and one short nude seta. Telofemur IV 80 ��m long with 11 setulose setae and two short nude setae. Genu IV 70 ��m long with numerous setulose setae and two solenidia. Tibia IV 74 ��m long with numerous setulose setae and two solenidia. Tarsus IV 110 ��m long with many strongly setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae: or 1 23 ��m, or 2 20 ��m. Subcapitular setae absent. Cheliceral body 100 ��m long, cheliceral blade 45 ��m long. Palp 170 ��m long. Palptrochanter with two smooth setae, ca 40 ��m long. Palpfemur with seven setae. Palpgenu with six setae. Palptibia with four nude setae, an enlarged seta (16 ��m) near base of palptarsus and a, Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 44-49, DOI: 10.5281/zenodo.177319

Allotanaupodus gen. nov. Diagnosis See that for the subfamily. Type species Allotanaupodus williamsi sp. nov. (by present designation). Etymology A different (Allo-) ��� Tanaupodus ���; masculine in gender. Remarks This genus is well represented by three new species from different areas of New Zealand. A fourth species is also recognized but the single specimen was mounted in such a position that a complete description is not attempted here., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 7-8, DOI: 10.5281/zenodo.177319

Allotanaupodus williamsi sp. nov. (Figs 1���10) Description Adult female (Figs 1���5): Idiosoma ovoid; a relatively large allotanaupodid mite, length 1250���1300 ��m, maximum width 900���910 ��m. Sejugal furrow clearly evident, hysteronotum slightly more than twice as long as prodorsum (Fig. 1). Prodorsum (Fig. 4 A ). About half as long (410 ��m) as wide (750 ��m). Prodorsal plate longer (410 ��m) than wide (350 ��m), not extending beyond anterior edge of propodosoma, with 11���12 pairs of lanceolate setae on each side, anteriormost pair 106���107 ��m long. Crista metopica linear, rod-like, 275���300 ��m long, widest anteriorly, ca 20���23 ��m. 2���3 pairs of lanceolate setae posterolateral to prodorsal plate (two of them laterad of eyes), each on a platelet; these setae ca 75 ��m long. Lateral prodorsal plate absent. Eye plates 85��� 90 x 54���55 ��m, diameter 31���33 ��m for anterior eyes and 26���28 ��m for posterior eyes. Hysterosomal dorsum. Hysterosoma with some indication of segmentation, each with large lateral plates bearing several to numerous setae; C and D separated by an evident furrow. C row with a pair of sublateral plates each bearing 10���12 setae; other setae each on a platelet; a few of these lateral to sublateral plates. D row with a pair of sublateral plates each bearing 3���5 pairs of setae; lateral and medial setae of the row each on a platelet, except a pair of submedian plates, each bearing two setae. E row with a pair of sublateral plate each bearing 4���5 setae; lateral and medial setae each on a platelet. F row with a pair of sublateral plate each bearing 7���9 setae; other setae of this row each on a platelet, except a pair of submedian plates near posterior margin of this row, each bearing two setae. H row with two pairs of plates each bearing 5���6 pairs of setae; a few lateral and medial setae each on a platelet. PS row with a pair of terminal plates each bearing several pairs of setae. Idiosomal venter (Fig. 2). Intercoxal area of leg I normally with 6 pairs of short setae (1 a the longest, over 100 long); intercoxal area of leg II with about 7 pairs of setae. Intercoxal area of leg III with numerous pairs of setae and that of leg IV with numerous setae not separable from those anterior to genital area. Between coxa II and coxa III on each side a plate of numerous setae. Coxal I with 22���26 setae. Coxa II with 17���20 setae. Coxa III with 16���18 setae. Coxa IV with 16���19 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 230���245 ��m long. A pair of central valves each bearing 7���9 setae (ca 50 ��m long). A pair of lateral valves each bearing 11 setae of similar size. Eugenital setae absent. Anal opening 168���185 ��m long, its valves each with three pairs of setae. Legs (Fig. 5). Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 1330���1333 ��m. Trochanter I with numerous setulose setae. Basifemur I 155���190 ��m, Telofemur I 208���225 ��m. Genu I 250���258 ��m long, with numerous setulose setae, three solenidia and one microseta (35 ��m long). Tibia I 280 ��m long, with numerous setulose setae and solenidia, and one microseta (44 ��m long). Tarsus I more than twice as long (335���350 ��m) as wide (150 ��m), with many setulose setae and solenidia; famulus ca. 20 ��m long. Leg II 830���890 ��m. Trochanter II with 12���13 setae. Basifemur II 140 ��m long with numerous setulose setae. Telofemur II 136 ��m long with numerous setulose setae and a distal solenidion. Genu II 146 ��m long, with numerous setulose setae, one solenidion and one microseta (30 ��m long). Tibia II 157 ��m long, with numerous setulose setae and two solenidia. Tarsus II 235 ��m long, with numerous setulose setae, 3 dorsal solenidia, one famulus (13 ��m long) and two distal eupathidia. Leg III 960���985 ��m. Trochanter III with 13���17 setae. Basifemur III 140 ��m long with numerous setae. Telofemur III 136 ��m long with numerous setulose setae and a distal solenidion. Genu III 164 ��m long with numerous setulose setae and three solenidia. Tibia III 195 ��m long with numerous setulose setae and two solenida. Tarsus III 233 ��m long, with numerous setulose setae and three solenidia. Leg IV 1280���1350 ��m. Trochanter IV with 17���22 setae. Basifemur IV 200 ��m long with numerous setae. Telofemur IV 176 ��m long with numerous setae and 1 distal solenidion (45 ��m long). Genu IV 230 ��m long with numerous setulose setae and 4 solenidia. Tibia IV 306 ��m long with numerous setulose setae and four solenidia, of which the two distal ones are twice as long as others. Tarsus IV 282 ��m long with many setulose setae and three solenidia. Gnathosoma (Fig. 3). Subcapitulum with two pairs of adoral setae and one pair of subcapitular setae (Fig. 3 B). Adoral setae: or 1 37���40 ��m, or 2 35���40 ��m. Subcapitular setae 70 ��m. Cheliceral base 225 ��m long, cheliceral blade 65 ��m long (Fig. 3 A). Palp 488���570 ��m long (Fig. 3 C). Palptrochanter with two setae, 75 and 100 ��m long, respectively. Palpfemur with 11 setae. Palpgenu with 12���15 setae. Palptibia with 14 nude setae, a strong terminal seta (i.e. tibial claw, 33 ��m long) and a ventral enlarged seta (27 ��m long) near base of tibial claw, ratio of tibial length excluding tibial claw and maximum tibial width 2.3���2.5. Palptarsus with a basal solenidion (20 ��m long), two nude setiform setae, and six terminal eupathidia (18���21 ��m long). Adult male (Figs 6���10). Idiosoma ovoid; length 1250���1450 ��m, maximum width 860���950 ��m. Sejugal furrow clearly evident, hysteronotum about three times as long as prodorsum (Fig. 6). Prodorsum. About half as long (385 ��m) as wide (725 ��m). Prodorsal plate longer (375 ��m) than wide (275 ��m), not reaching anterior end of propodosoma, with 12���14 pairs of lanceolate setae on each side, anteriormost pair 100���125 ��m long (Fig. 9 A). Crista metopica linear, rod���like, 275���325 ��m long, widest anteriorly, ca 15���22 ��m. 3���9 pairs of lanceolate setae posterolateral to prodorsal plate (one or two of them laterad of eyes), each on a platelet, ca 75���80 ��m long. Prodorsal lateral plates absent. Eye plates 75��� 95 x 50���60 ��m, diameter 32���37 ��m for anterior eyes and 25���30 for posterior eyes. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D a clearly evident furrow. C row with a pair of sublateral plates each bearing 7 setae; other setae each on a platelet. D row with a pair of sublateral plates each bearing 3���5 pairs of setae; lateral and medial setae each on a platelet (except a single medial pair together on one plate). E row with a pair of sublateral plates each bearing 5���7 setae, lateral and medial setae each on a platelet. F row with a pair of sublateral plate each bearing 8���9 pairs of setae, lateral and medial setae each on a platelet. H segment with two pairs of plates each bearing 5���6 pairs of setae; a few other setae each on a platelet. PS segment with a pair of terminal plates each bearing several pairs of setae. Idiosomal venter (Fig. 7). Intercoxal area of leg I with 9���10 pairs of short setae (1 a over 100 ��m long), intercoxal area of leg II with 5���9 pairs of setae. Intercoxal area of leg III with numerous pairs of setae and that of leg IV with numerous setae not separable from those anterior to genital area. Coxal I with 22���26 setae. Coxa II with 17���22 setae. Coxa III with 16���20 setae. Coxa IV with 17���24 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 190��� 250 ��m. A pair of central valves each bearing 7���9 setae. A pair of lateral valves each bearing 7���15 setae of similar size. With three or five pairs of eugenital setae, all nude, setiform. Anal opening 145���195 ��m long, its valves each with 1���4 pairs of setae. Legs (Fig. 10). Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 1075���1445 ��m. Trochanter I with 12���15 setae. Basifemur I 140���175 ��m. Telofemur I 185���215 ��m. Genu I 190���265 ��m long, with numerous setulose setae, two solenidia and a microseta (32 ��m long). Tibia I 200���350 ��m long with numerous setulose setae and solenidia, and one microseta (35 ��m long). Tarsus I more than twice as long (255���360 ��m) as wide (95���163 ��m), with many setulose setae and solenidia; famulus peglike, 3 ��m long. Leg II 760���830 ��m. Trochanter II with 9���15 setae. Basifemur II 130 ��m long with numerous setulose setae. Telofemur II 130 ��m long with numerous setulose setae and a distal solenidion. Genu II 135 ��m long, with numerous setulose setae, one solenidion and one microseta (22 ��m long). Tibia II 143 ��m long, with numerous setulose setae and two solenidia. Tarsus II 205 ��m long, with numerous setulose setae, five dorsal solenidia, one famulus (15 ��m long) and two distal eupathidia. Leg III 900���1085 ��m. Trochanter III with 16���20 setae. Basifemur III 130 ��m long with numerous setae. Telofemur III 130 ��m long with numerous setulose setae and a distal solenidion. Genu III 160 ��m long with numerous setulose setae and three solenidia. Tibia III 180 ��m long with numerous setulose setae and two solenida. Tarsus III 230 ��m long, with numerous setulose setae and two solenidia. Leg IV 1200���1375 ��m. Trochanter IV with 19���22 setae. Basifemur IV 175 ��m long with numerous setae. Telofemur IV 160 ��m long with numerous setae and 1 distal solenidion (48 ��m long). Genu IV 220 ��m long with numerous setulose setae and 4 solenidia. Tibia IV 295 ��m long with numerous setulose setae and four solenidia, of which two distal ones are twice as long as others. Tarsus IV 270 ��m long with many setulose setae and one solenidion (30 ��m long). Gnathosoma (Fig. 8). Subcapitulum with two pairs of large adoral setae (in addition, one minute pair also seen in some specimens) and one pair of subcapitular setae (in the holotype, the left side has abnormally three setae). Adoral setae: or 1 35 ��m, or 2 35 ��m. Cheliceral base 208���232 ��m long, cheliceral blade 65���70 ��m long (Fig. 8 A). Palp 490���597 ��m long (Fig. 8 C). Palptrochanter with two setae, 75 and 100 ��m long. Palpfemur with 9��� 11 setae. Palpgenu with 13���15 setae. Palptibia with 14���18 nude setae, a strong terminal seta (i.e. tibial claw, 35 ��m long) and a ventral enlarged seta (25 ��m) near base of tibial claw. Ratio of tibial length excluding tibial claw to maximum width 2.2���2.8. Palptarsus with a basal solenidion (15 ��m long), two nude setiform setae, and six terminal eupathidia (22 ��m long). Etymology Named after the collector, D. Williams. Type specimens Holotype female New Zealand, AK: Kawau I. (36.42 ��S, 174.85 ��E), 15 June to 10 August 1992, Malaise trap, D. Williams. Allotype male, one paratype one and two paratype females, same data. Four paratype males, same data except collected during 13 October to 12 November 1992. Biology The feeding habit of this species is unknown. All nine specimens were caught in Malaise traps; presumably they are free-living predators of small arthropods in the soil. The winter sample (15 June to 10 August) includes three males and two females. The spring sample (12 October to 12 November) includes four males only. Remarks This species has the largest series of specimens and multiple males and females are represented. Some variation among specimens was observed in the female holotype, the C row has a median plate with three setae, whereas the other two specimens have all median setae of the C row on a single small platelet., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 8-20, DOI: 10.5281/zenodo.177319

Allotanaupodidae fam. nov. Diagnosis The same as that for the superfamily. Type genus Allotanaupodus gen. nov. (by present designation). Etymology Derived from the type genus. Remarks This new family, as the name implies, is most similar to another early derivative family Tanaupodidae of the superfamily Tanaupodoidea. It is also similar to Johnstonianidae (placed by Welbourn, 1991 in his Trombiculoidea) in some aspects. Allotanaupodidae fam. nov. can not be placed in any of the four superfamilies of Trombidiina sensu Welbourn (1991), although it is a member of Trombidiina without question. Thus it is provisionally placed in its own superfamily within Trombidiina. This new family is characterized by a number of interesting features, some shared with other early derivative Parasitengona and others unique. A detailed analysis is presented below. Prodorsal trichobothria in the adults and deutonymphs. In Erythraeina, the Erythraeoidea has two pairs of trichobothria within well-developed sensory areas and the Calyptostomatoidea has one pair. Most Trombidiina have a single pair of trichobothria within a pair of well-developed sensory areas. Some Johnstonianidae (e.g. Diplothrombium) have two pairs of trichobothria within well-developed sensory areas. Allotanaupodidae fam. nov. is unique among terrestrial Parasitengona in the complete loss of both trichobothria and associated sensory areas. In early derivative terrestrial Parasitengona, the presence of two pairs of trichobothria is assumed to be the plesiomorphic condition (the outgroup Anystidae, for example, has two pairs). Newell (1957) noted the tendency toward the loss of the anterior pair of trichobothria in some Johnstonianidae (and also Tanaupodidae). In Allotanaupodidae fam. nov., the posterior pair of the trichobothria is also lost. It should be noted that the complete loss of trichobothria is also seen in most water mites, with few exceptions (e.g. Hydrophantoidea and Stygothrombidoidea). Prodorsal plate. The broad prodorsal plate in adults and deutonymphs in this family resembles those present in most larval Trombidiina. This plesiomorphic condition is shared by adults and deutonymphs of other early derivative groups such as Tanaupodidae (e.g. Lassenia) and Johnstonianidae (e.g. Johnstoniana) (Newell 1957). In most derived Trombidiina, the prodorsum is dominated by the crista metopica and the prodorsal plate is often restricted to areas close to the crista. Crista metopica. Most Trombidiina have well-developed crista metopica with associated sensory areas. The absence of a well-developed crista metopica is seen in some Trombellidae, some Johnstonianidae and Audyanidae. The crista metopica in the new family is a simple linear rod. A similar condition is seen in the early derivative Tanaupodidae, although the latter has a pair of trichobothria. Hysterosomal dorsal setae and plates. The simple setae arising from small plates in the new family resemble those seen in larval Trombidiina and in the adults and deutonymphs of some other early derivative Trombidiina (Tanaupodidae and Johnstonianidae). The new family differs from the last two families in the presence of at least one pair of plates in each of the C to H rows with multiple setae (although this is not seen in Paratanaupodus). Number of genital acetabula in adults. The number of genital acetabula is typically one pair in protonymphs, two pairs in deutonymphs and three pairs in tritonymphs/adults in the Acariformes (Grandjean- Oudemans rule). The number of genital acetabula is quite variable in water mites (Cook 1974) and also shows some variation in Erythraeina and Trombidiina. In Erythraeina, the adults of Calyptostomatoidea retain the deutonymphal condition of having two pairs of genital acetabula. In Trombidiina, parallel evolution is seen in Allotanaupodidae fam. nov. and some Trombellidae such as Neonothrothrombium and Durenia (Robaux 1968) and Charadracarus of Johnstonianidae (Newell 1960). Palptarsus. The palptarsus in the new family is short, its length similar to its diameter or slightly longer. In most other Trombidiina, the palptarsus is elongate (much longer than diameter). A correlated character is the subdistal insertion of the palptarsus on the ventral surface of the tibia in the new family. In other Trombidiina, the palptarsus is inserted proximally or near the middle on palptibia, and the elongate palptarsus and the distal part of the palptibia (including the tibial claw) form a thumb-claw complex. In the new family, the palptarsus is short and it thus can only form a thumb-claw complex by subdistal insertion. The above analysis shows that the monophyly of Allotanaupodidae fam. nov. is supported by the following synapomorphies in adults/deutonymphs: (1) absence of trichobothria and sensory areas, (2) the presence of one or two pairs of plates with multiple setae on C to PS rows of dorsal hysterosoma, (3) the presence of only two pairs of genital acetabula in adults, and (4) palptarsus distally inserted on the palptibia.

Allotanaupodidae fam. nov. Diagnosis The same as that for the superfamily. Type genus Allotanaupodus gen. nov. (by present designation). Etymology Derived from the type genus. Remarks This new family, as the name implies, is most similar to another early derivative family Tanaupodidae of the superfamily Tanaupodoidea. It is also similar to Johnstonianidae (placed by Welbourn, 1991 in his Trombiculoidea) in some aspects. Allotanaupodidae fam. nov. can not be placed in any of the four superfamilies of Trombidiina sensu Welbourn (1991), although it is a member of Trombidiina without question. Thus it is provisionally placed in its own superfamily within Trombidiina. This new family is characterized by a number of interesting features, some shared with other early derivative Parasitengona and others unique. A detailed analysis is presented below. Prodorsal trichobothria in the adults and deutonymphs. In Erythraeina, the Erythraeoidea has two pairs of trichobothria within well-developed sensory areas and the Calyptostomatoidea has one pair. Most Trombidiina have a single pair of trichobothria within a pair of well-developed sensory areas. Some Johnstonianidae (e.g. Diplothrombium) have two pairs of trichobothria within well-developed sensory areas. Allotanaupodidae fam. nov. is unique among terrestrial Parasitengona in the complete loss of both trichobothria and associated sensory areas. In early derivative terrestrial Parasitengona, the presence of two pairs of trichobothria is assumed to be the plesiomorphic condition (the outgroup Anystidae, for example, has two pairs). Newell (1957) noted the tendency toward the loss of the anterior pair of trichobothria in some Johnstonianidae (and also Tanaupodidae). In Allotanaupodidae fam. nov., the posterior pair of the trichobothria is also lost. It should be noted that the complete loss of trichobothria is also seen in most water mites, with few exceptions (e.g. Hydrophantoidea and Stygothrombidoidea). Prodorsal plate. The broad prodorsal plate in adults and deutonymphs in this family resembles those present in most larval Trombidiina. This plesiomorphic condition is shared by adults and deutonymphs of other early derivative groups such as Tanaupodidae (e.g. Lassenia) and Johnstonianidae (e.g. Johnstoniana) (Newell 1957). In most derived Trombidiina, the prodorsum is dominated by the crista metopica and the prodorsal plate is often restricted to areas close to the crista. Crista metopica. Most Trombidiina have well-developed crista metopica with associated sensory areas. The absence of a well-developed crista metopica is seen in some Trombellidae, some Johnstonianidae and Audyanidae. The crista metopica in the new family is a simple linear rod. A similar condition is seen in the early derivative Tanaupodidae, although the latter has a pair of trichobothria. Hysterosomal dorsal setae and plates. The simple setae arising from small plates in the new family resemble those seen in larval Trombidiina and in the adults and deutonymphs of some other early derivative Trombidiina (Tanaupodidae and Johnstonianidae). The new family differs from the last two families in the presence of at least one pair of plates in each of the C to H rows with multiple setae (although this is not seen in Paratanaupodus). Number of genital acetabula in adults. The number of genital acetabula is typically one pair in protonymphs, two pairs in deutonymphs and three pairs in tritonymphs/adults in the Acariformes (Grandjean- Oudemans rule). The number of genital acetabula is quite variable in water mites (Cook 1974) and also shows some variation in Erythraeina and Trombidiina. In Erythraeina, the adults of Calyptostomatoidea retain the deutonymphal condition of having two pairs of genital acetabula. In Trombidiina, parallel evolution is seen in Allotanaupodidae fam. nov. and some Trombellidae such as Neonothrothrombium and Durenia (Robaux 1968) and Charadracarus of Johnstonianidae (Newell 1960). Palptarsus. The palptarsus in the new family is short, its length similar to its diameter or slightly longer. In most other Trombidiina, the palptarsus is elongate (much longer than diameter). A correlated character is the subdistal insertion of the palptarsus on the ventral surface of the tibia in the new family. In other Trombidiina, the palptarsus is inserted proximally or near the middle on palptibia, and the elongate palptarsus and the distal part of the palptibia (including the tibial claw) form a thumb-claw complex. In the new family, the palptarsus is short and it thus can only form a thumb-claw complex by subdistal insertion. The above analysis shows that the monophyly of Allotanaupodidae fam. nov. is supported by the following synapomorphies in adults/deutonymphs: (1) absence of trichobothria and sensory areas, (2) the presence of one or two pairs of plates with multiple setae on C to PS rows of dorsal hysterosoma, (3) the presence of only two pairs of genital acetabula in adults, and (4) palptarsus distally inserted on the palptibia., Published as part of Zhang, Zhi-Qiang & Fan, Qing-Hai, 2007, Allotanaupodidae, a new family of early derivative Parasitengona (Acari: Prostigmata), pp. 1-52 in Zootaxa 1517 on pages 5-6, DOI: 10.5281/zenodo.177319, {"references":["Welbourn, W. C. (1991) Phylogenetic studies of the terrestrial Parasitengona. in: F. Dusbabek & V. Bukva (eds.) Modern acarology, Vol. 2. Academia, Prague, and SPB Academic Publishing, The Hague. pp. 163 - 170","Newell, IM (1957) Studies on the Johnstonianidae (Acari, Parasitengona). Pacific Science, 11, 396 - 466.","Cook, D. R. (1974) Water mite genera and subgenera. Memoir of the American Entomological Institute, 21, 1 - 860.","Robaux, P. (1968) Thrombidiidae d'Amerique du Sud I-Tanaupodinae, Johnstonianidae, Thrombellini (Acarina- Thrombidiidae). Acarologia, 10, 450 - 466.","Newell, I. M. (1960) Charadracarus new genus, Charadravarinae new subfamily (Acari, Johnstonianidae), and the status of Typhlothrombium Berlese, 1910. Pacific Science, 14, 156 - 172."]}

Nanotanaupodus andrei sp. nov. (Figs 21–29) Description Adult female. Idiosoma elongate-ovoid; length 788 µm, maximum width 490 µm. Sejugal furrow clearly evident, hysteronotum almost three times as long as prodosum. Prodorsum. Almost half as long (200 µm) as wide (370 µm). Prodorsal plate longer (190 µm) than wide (160 µm), not reaching anterior end of propodosoma, with 13–14 lanceolate setae on each side, anteriormost pair 40 µm long, posteriormost pair 32 long. Crista metopica linear, rod-like, 155 µm long, widest anteriorly, ca 10 µm. A pair of lanceolate setae posterolateral to prodorsal plate, each on its platelet and ca 40 µm long. A pair of posterolateral plates each bearing 5 setae of ca 40 long µm. Eyes absent. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D by evident furrow. C row with medial setae each on a platelet, but lateral setae clustered on a pair of plates; setae 30–40 µm long, all lanceolate. D row with medial setae (30 µm long) each on a platelet, a pair of sublateral plates each with three setae (35–40 µm long) and a pair of lateral plates each with 6–9 setae (35–40 µm long). E row with a pair of lateral plates each with lanceolate 11–12 setae (35–45 µm long) and many small lanceolate setae each on a platelet between them. F and H rows both with similar setal and plate arrangements as in E. PS row with a pair of plates each bearing numerous lanceolate setae mostly over 40 µm long. Idiosomal venter. Intercoxal area of leg I with eight short setae (20–30 µm long). Intercoxal area of leg II with six short setae. Intercoxal area of leg III with nine pairs of setae and that of leg IV with three pairs. Coxal I with 14–15 setae, 1 a (30 µm) not much longer than other secondary setae. Coxa II with 14 setae. Coxa III with 12 setae. Coxa IV with eight setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 100 µm long. A pair of central valves each bearing six setae of similar size (shorter than 20 µm). A pair of lateral valves each bearing 5–6 setae of similar size (shorter than 20 µm long). Eugenital setae absent. Anal opening 93 µm long, its valves each with 3 setae (over 20 µm long). Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 505 µm. Trochanter I with three long setulose setae and two short ones. Basifemur I 80 µm. Telofemur I 77 µm long, without solenidia. Genu I 82 µm long, with numerous setulose setae and 1 solenidion. Tibia I 90 µm long, with numerous setulose setae and four solenidia. Tarsus I over twice as long (165 µm) as wide (70 µm), with many setulose setae and solenidia. Leg II 290 µm. Trochanter II with three long setae and two shorter setae. Basifemur II 60 µm long, with nine setulose setae. Telofemur II 50 µm long with 10–12 setulose setae. Genu II 50 µm long with numerous setulose setae and one solenidion. Tibia II 50 µm long, with numerous setulose setae and one solenidion. Tarsus II 78 µm long with many strongly setulose setae, two subdistal solenidia and one basal solenidion. Leg III 340 µm. Trochanter III with three long setulose setae and three shorter ones. Basifemur III 62 µm long with 10 setulose setae. Telofemur III 50 µm long with 10–12 setulose setae. Genu III 58 µm long with numerous setulose setae and one solenidion. Tibia III 60 µm long with numerous setulose setae and two solenidia. Tarsus III 92 µm long with many strongly setoluse setae and one solenidion. Leg IV 450 µm. Trochanter IV with six setulose setae. Basifemur IV 80 µm long with eight setulose setae. Telofemur IV 64 µm long with 12 setulose setae. Genu IV 80 µm long with numerous setulose setae and two solenidia. Tibia IV 80 µm long with numerous setulose setae and two solenidia. Tarsus IV 110 µm long with many strongly setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae: or 1 20 µm, or 2 20 µm. Cheliceral base 85 µm long, cheliceral blade 38 µm long. Palp 200 µm long. Palptrochanter with two smooth setae, ca 40 µm long. Palpfemur with seven setae. Palpgenu with seven setae. Palptibia with five nude setae, an enlarged seta (15 µm) near base of palptarsus and a strong terminal setae (i.e. tibial claw, 25 µm long), ratio of tibial length excluding tibial claw and maximum width 1.9. Palptarsus with a basal solenidion (10 µm long), four nude setiform setae, one short spinelike seta and three terminal eupathidia. Deutonymph. Idiosoma elongate-ovoid; length 600–645 µm, maximum width 370–375 µm. Sejugal furrow clearly evident in dorsomedial portion, hysteronotum almost three times as long as prodosum. Prodorsum. Almost half as long (167–175 µm) than wide (310–322 µm). Prodorsal plate longer (131 µm) than wide (106 µm), not reaching anterior end of propodosoma, with 7 lanceolate setae each side, anteriormost pair 30 µm long, posteriormost pair 30 µm long. Crista metopica linear, rod-like, 110–118 µm long, widest anteriorly, ca 8 µm. A pair of lanceolate setae posterolateral to prodorsal plate, each on its platelet and ca 32–35 µm long. A pair of posterolateral plates each bearing three setae of ca 32–40 µm long. Eyes absent. Hysterosomal dorsum. Hysterosoma with traces of segmentation, each with large lateral plates bearing several to numerous setae, division between C and D evident furrow. C row with medial setae each on a platelet, but lateral setae clustered on a pair of plates; setae 23–35 µm long, all lanceolate. D row with medial setae (20–25 µm long) each on a platelet, a pair of sublateral plates each with 2–3 setae (30 µm long) and a pair of lateral plates each with 5–6 setae (30–35 µm long). E row with a pair of lateral plates each with 5–6 lanceolate setae (30–38 µm long) and numerous lanceolate setae on a platelet between them. F row with similar setal and plate arrangement as in E, but only several setae between plates. H row with similar setal and plate arrangement as in E, but only a few setae between plates. PS row with a pair of plates each bearing numerous lanceolate setae mostly over 40 µm long. Idiosomal venter. Intercoxal area of leg I with four short setae. Intercoxal area of leg II with four short setae. Intercoxal area of leg III with four pairs of setae and that of leg IV with three pairs. Coxal I with 8–10 setae, 1 a (25 µm) not much longer than other setae. Coxa II with 10–12 setae. Coxa III with 7–10 setae. Coxa IV with 4–5 setae. Opisthoventer with numerous short setiform setae similar to coxal setae. Genital opening 60 µm long. A pair of central valves each bearing three setae of similar size (10–12 µm long). A pair of lateral valves each bearing 1–2 setae of similar size (12–15 µm long). Eugenital setae absent. Anal opening 65 µm long, its valves each without setae. Legs. Length of legs in decreasing order: leg I> leg IV> leg III> leg II. Leg I 363–375 µm. Trochanter I with 2 long well-setulose setae and 2 short, nearly smooth setae. Basifemur I 55 –58 µm. Telofemur I 50 –55 µm. Genu I 58 –62 µm long, with one solenidion (15 µm long). Tibia I 62 µm long, with numerous setulose setae and 4–5 solenidia (15 µm long). Tarsus I over twice as long (120–125 µm) as wide (50–58 µm), with many setulose setae and solenidia. Leg II 225 µm. Trochanter II with two long setae and one short seta. Basifemur II 42 µm long with five setulose setae. Telofemur II 35 µm long with six setulose setae. Genu II 37 µm long, with numerous setulose setae and one solenidion near the base of segment. Tibia II 40 µm long, with numerous setulose setae and a single solenidion near the base of segment. Tarsus II 66 µm long, with many setulose setae and no solenidia. Leg III 265–270 µm. Trochanter III with three long setulose setae and two shorter ones. Basifemur III 40 µm long with seven setulose setae. Telofemur III 35 µm long with six setulose setae. Genu III 40 µm long with numerous setulose setae and one solenidion. Tibia III 43 µm long with numerous setulose setae and two solenidia. Tarsus III 73 µm long with many strongly setulose setae. Leg IV 310–325 µm. Trochanter IV with four setulose setae and a shorter one. Basifemur IV 48 µm long with three setulose setae. Telofemur IV 46 µm long with 7 setulose setae. Genu IV 55 µm long with numerous setulose setae and two solenidia. Tibia IV 60 µm long with numerous setulose setae and two solenidia. Tarsus IV 86 µm long with many strongly setulose setae. Gnathosoma. Subcapitulum smooth with two pairs of adoral setae. Adoral setae or 1 15 µm, or 2 20 µm. Subcapitular setae absent. Cheliceral body 90 µm long, cheliceral blade 30–33 µm long. Palp 140–150 µm long. Palptrochanter with one smooth seta, ca 30 long. Palpgenu with six or five setae. Palpgenu with four setae. Palptibia with three nude setae, an enlarged seta (10–12 µm) long near the base of palptarsus and a strong terminal setae (i.e. tibial claw, 20–25 µm long), ratio of tibial length excluding tibial claw and maximum width 1.3–1.5. Palptarsus with a basal solenidion (8 µm long), four nude setiform setae, one short spine-like setae and three terminal eupathidia (15 µm long). Etymology Named in memory of Marc André for his many contributions to systematics of Trombidiina. Type specimens Holotype male and paratype deutonymph (on the same slide ZQZ 2002 -01231A,B,C,D: C= holotype; D= paratype), New Zealand, TO, Waituhi Saddle, from litter, moss and rotten wood, 16 Nov. 1983, C.F. Butcher (litter 83 / 112). Deposited in NZAC.