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The Nok culture in central Nigeria, dated 1500–1 cal bc, is known for its famous terracotta sculptures. We here present a study on > 11,000 botanical macro-remains from 50 sites, including 343 samples from Nok contexts and 22 samples dating between cal ad 100 and 400, after the end of the Nok culture. With 9,220 remains, pearl millet (Cenchrus americanus (L.) Morrone, syn. Pennisetum glaucum (L.) R.Br.) is dominant in the Nok samples, followed by cowpea (Vigna unguiculata (L.) Walp.), Canarium schweinfurthii Engl., Nauclea latifolia Sm. (syn. Sarcocephalus latifolius (Sm.) E.A.Bruce), wild fruit trees and wild Poaceae. The pearl millet remains consist exclusively of charred caryopses; chaff remains are completely absent. Because we studied all size fractions, including the small 0.5 mm fractions usually containing the involucres, bristles and husks, the absence of pearl millet chaff is real, excluding a methodological explanation, and distinguishes Nok from contemporary other West African sites. We propose that most excavated Nok sites were consumer sites where clean grain was brought in from outside and consumed in a ritual context connected with feasting. This is in line with the archaeological evidence from the larger excavated sites with stone-pot arrangements that are interpreted as ritual places related to mortuary practices. In addition to the known southward branches of pearl millet diffusion from the Sahara we propose a new, hitherto unknown branch directly from the central Sahara to the central Nigerian savannas.

The development of cultivated landscapes under the influence of food production has decisively shaped the composition and appearance of West African savannas as present today. With trees and shrubs forming an important constituent of such landscapes, and its composition characterizing different types of land-use, anthracology is a tool particularly well suited to trace developments and changes through human exploitation of the landscape. Within the West African savanna belts, regions shaped by prolonged annual floods, such as the Middle Senegal Valley, the Inland Niger Delta and the Chad Basin, offer particular ecological preconditions for land-use. We present here a charcoal study for the Chad Basin of northeast Nigeria, demonstrating the special development of a cultivated landscape in an alluvial context. Charcoal samples from stratified Late Stone Age and Iron Age sites within the southwestern Chad Basin, Nigeria were analyzed. Main focus is on the site Mege, spanning more than 2500 years of occupation history. The results are supplemented with and compared to data from other, earlier as well as contemporary sites, in similar or slightly different environments and representing different settlement types, in order to discern general trends. The results illustrate the exploitation of woody vegetation mainly from the clay soils. This is in stark contrast to the results from other regions within the savanna belt, where the vegetation on sandy soils was preferentially exploited for fuel wood. The vegetation changes evidenced through the charcoal assemblages are attributed to reduced flooding as a consequence of falling lake levels during the Late Holocene. Parklands, typical cultivated landscapes of West African savannas, are invisible in the charcoal spectra, and fallow species and useful trees are rarely present. We postulate that the ecological conditions of the floodplain environment have decisively paved a different, less obvious trajectory of cultivated landscape development between 2000 BC and AD 1500 and that the cultivated landscapes present in the region today largely result from the increasing human impact of the last 500 years.

The central Nigerian Nok Culture is known for its elaborate terracotta figurines and iron metallurgy of the first millennium BC. Since 2005, Goethe University Frankfurt has carried out comprehensiv...

West African cuisine has long been known for its distinct ingredients and flavours, often enhanced by the addition of a large and diverse range of plant foods. A traditional meal comprises a starchy staple cooked in a pot, served with a sauce prepared from vegetables, fish and/or meat, often accompanied by pulses. However, reconstructing the antiquity of the full range of plant use by ancient peoples, using archaeobotanical remains, in West Africa is challenging due to their somewhat fragile nature. Hence, there is a strong bias toward food plants that survive in charred condition, rendering invisible those that easily decompose, such as leafy plants and tubers. Here, we combine organic residue analysis of 458 prehistoric vessels, with archaeobotanical evidence from ten sites of the prehistoric Nok culture, Nigeria, spanning a period of around 1500 years, beginning around the middle of the second millennium BC and terminating in the last centuries BC. Our results reveal a range of highly diverse and complex lipid distributions denoting the preparation and processing of various plant types, including leafy vegetables or ‘greens’, cereals, pulses and underground storage organs, possibly yams. Here, we render previously unidentifiable leafy plant use visible and suggest an early origin for the plant component of modern-day West African cuisine.

Archaeobotanical evidence from archaeological sites in northeast Burkina Faso dating to the first and second millennia AD has provided a useful insight into crop cultivation and the development of the West African savanna landscape. Nitrogen isotopic analysis of charred pearl millet grains from the same sites now provides the first opportunity to investigate how increased crop production and permanence of cultivated fields related to the intensity of household waste/manure application. Nitrogen isotope values of pearl millet grains increased during the first two millennia AD, indicating an intensification of manuring that would have enabled soil to stay fertile for longer, reducing the agricultural footprint of shifting cultivation. This may have been advantageous as population and settlement density increased, thereby increasing competition over land. The intensity of manure application in the second millennium AD at sites close to the Mare d’Oursi suggests that manure was likely sourced from outside the farming settlements, from livestock herded by nomadic pastoralists who would have been drawn to the mare for water. This is rare evidence for specialisation of sedentary farmers and pastoralists, demonstrating how the novel combination of fruit/seed, charcoal, faunal and isotopic evidence used in this study can enrich our knowledge of past lifeways in West Africa.

Many societal and environmental changes occurred between the 2nd millennium BC and the middle of the 2nd millennium AD in western Africa. Key amongst these were changes in land use due to the spread and development of agricultural strategies, which may have had widespread consequences for the climate, hydrology, biodiversity, and ecosystem services of the region. Quantification of these land-use influences and potential feedbacks between human and natural systems is controversial, however, in part because the archaeological and historical record is highly fragmented in time and space. To improve our understanding of how humans contributed to the development of African landscapes, we developed an atlas of land-use practices in western Africa for nine time-windows over the period 1800 BC–AD 1500. The maps are based on a broad synthesis of archaeological, archaeobotanical, archaeozoological, historical, linguistic, genetic, and ethnographic data, and present land use in 12 basic categories. The main differences between categories is the relative reliance on, and variety of, domesticated plant and animal species utilized, and the energy invested in cultivating or keeping them. The maps highlight the irregular and frequently non-linear trajectory of land-use change in the prehistory of western Africa. Representing an original attempt to produce rigorous spatial synthesis from diverse sources, the atlas will be useful for a range of studies of human–environment interactions in the past, and highlight major spatial and temporal gaps in data that may guide future field studies., Open access funding provided by Max Planck Society. This work was supported by Grants to JOK from the European Research Council (COEVOLVE, 313797), the Swiss National Science Foundation (ACACIA, CR10I2_146314), and the Fondation Herbette (2016-2-E-16). JMR was funded by the Funds for Research Training in Industry and Agriculture (National Funds for Scientific Research, Belgium) and the Leopold II Funds (Belgium). Data were also acquired in the framework of the Era-Net BiodivERsA CoForChange project, funded by the French National Research Agency and National Environment Research Council (UK).

Pits are common features at sites of the Nok Culture, which is known for its masterful terracotta sculptures and for early iron metallurgy in sub-Saharan Africa. If pits are stratified and well-dated, contained archaeological charcoal may serve as a proxy for vegetation change—especially for changes mediated by human impact. Recent investigations into charcoal samples from several pits at Pangwari, a Nok site that has been—not necessarily continuously—in use during the last 1.5 millennia BC, revealed that the taphonomy of these assemblages is not as straightforward as expected. Find distribution analysis documents that regularly only the lowermost parts of the pits are still in their original state. Later re-working of the original pit filling, erosion, and recent illicit digging for terracotta figurines challenge the interpretation of the charcoal data. Moreover, the distribution of 14C dates and of potsherds assigned to different pottery groups, as well as differences in the charcoal assemblages of some pits, hint towards divergent backfill histories. Some pits seem to have been refilled quite quickly, possibly even in a single event, while other pits might have remained open for a longer time. Additionally, pits likely were reused in later phases and the infill became mixed. For charcoal analysis, it is hence crucial to consider the backfill history of each pit based on the archaeological evidence.

Africa with its impressive, deep history and ecosystem diversity continues to offer an ideal setting to expand our frontiers of understanding plants and people in the past. Early and unique interrelationships between humans and plants make Africa a critical reference point for interdisciplinary studies of cultural developments and environmental transformations.

Charcoal samples from final Stone Age sites and stratified iron age settlement mounds in northern Burkina Faso illustrate the development of the West African savanna as a cultural landscape between 2000 BC and modern times. In the second millennium BC, plant cultivation was only a minor activity of predominantly mobile groups and did not alter the vegetation significantly. In the Iron Age, after 0 BC/AD, large settlement mounds were formed by sedentary populations, and a shifting cultivation system with fields and fallows established on the dunes. Cultivation was practiced in park savannas, an agroforestry system with protected wild fruit trees interspersed in the fields. During the first millennium AD, extensive cultivation with long fallow periods was practised. With the beginning of the second millennium AD, fallow periods became shorter and the influence of cattle herding on the vegetation increased, changing the composition of the woody vegetation. Decreasing precipitation and anthropogenic pressure eventually led to the much lower species diversity of the modern woody vegetation.

Agriculture was introduced into the Central African rainforest from the drier West African savanna, in concert with a major climatic change that amplified seasonality just after 2500 BP. The savanna crop pearl millet (Pennisetum glaucum), dated to 2400–2200 BP, could only be cultivated due to the development of a distinct dry season. Increasing seasonality and the replacement of mature forests by pioneer formations is indicated by Trema orientalis in the pollen diagram of Nyabessan after 2400 BP. However, charcoal data do not point to the existence of savannas in South Cameroon during this period, but rather to a mosaic of mature and pioneer forests. The early rainforest farmers combined the cultivation of pearl millet with the exploitation of wild oil-containing tree fruits, such as oil palm and Canarium. The existence of pioneer formations that can be easily cut favoured the establishment of shifting cultivation. The archaeobotanical finds fit into a linguistic scenario of West-Bantu speakers making the cultivation of pearl millet one of their food production strategies before expanding further to the South. The reconstructed inherited pearl millet vocabulary for the early phases of Bantu language history provides strong circumstantial evidence for an overlap of the major stages of the Bantu expansion with the dispersal of food production.

Identification of archaeological or soil charcoal in a species-rich biome, such as the Central African rainforest, is challenging because of the large number of woody taxa with similar and overlapping wood anatomical features. Valid environmental or archaeological interpretations can only derive from reliable and transparent identifications that allow comparison of and referencing between different charcoal assemblages. The identification of 30 archaeological charcoal types from the site Dibamba in southern Cameroon serves as a starting point for a discussion on classification and naming. These 30 types are fully documented and illustrated in the Supplementary Online Material (SOM). The discussion underlines the basics of “good practice” of charcoal identification in a speciesrich tropical environment. The value of differential diagnosis is stressed, as is the importance of leaving identification levels on higher taxonomic level if necessary. We argue that the level of identification must be reflected in the name of the charcoal type. Names of charcoal types are written in small capitals to clearly distinguish them from botanical taxa with which they are not necessarily identical. The Dibamba charcoal assemblage offers the first and so far unique possibility to directly comprehend human impact on the structure and composition of West Central African rainforest over the last 3000 years. The paleoenvironmental significance of the results presented here will be subject of a forthcoming publication.

This paper primarily presents analyses from geophysical data and archaeological remains collected at one of a few known sites to the west of Lake Chad associated with fired-brick structures. It reports on previous fieldwork at the location, considers its alleged relationship with the early history of the Borno kingdom and then turns to present new data that provide fresh insights into the physical appearance, dating, material remains, economy and environment of the site. A brief discussion of the historical purpose of the fired-brick structures and the site itself concludes the paper. Cet article presente les dates geophysiques et trouvailles archeologiques recoltees dans un des rares sites a l'ouest du Lac Chad associe avec des structures en briques. Il contient tout d’abord un rapport court sur les travaux antecedents sur l'endroit, ensuite des considerations sur sa relation avec l'histoire du jeune royaume de Bornou et tourne enfin a la presentation et l’interpretation des nouvelles dates qui permettent une premiere approche a la physionomie du site, son datation, la culture materielle, l'economie et l’environnement. Nous concluons avec une discussion breve de la signification des structures en briques et du site Garu Kime lui-meme.

Since 2003, a joint research project of the universities of Frankfurt and Tubingen (Germany) has explored the changing interrelationship of environment and culture in the forest-savanna regions of West and Central Africa. This paper provides the first archaeological and archaeobotanical results of three field seasons in the rainforest of southern Cameroun. Excavations were carried out at Bwambe Hill in the vicinity of Kribi at the Atlantic coast as well as at Akonetye, Minyin and Abang Minko’o, all located in the hinterland near Ambam. At all sites a number of pit structures, which contained mostly ceramics, were excavated. In addition, at Akonetye two graves with rich ceramic and iron offerings were unearthed. They seem to be the oldest graves with iron objects yet known in Central Africa.A large body of archaeobotanical material was retrieved from the structures excavated (charcoal fragments, charred fruits and seeds, phytolith and starch samples). Of high importance is the presence of pearl millet (Pennisetum glaucum) at Bwambe Hill and Abang Minko’o in archaeological contexts dated to about 2200 bp. Charcoal and pollen data indicate that the ancient settlements were situated in a closed rainforest which was, however, massively disturbed and partly substituted by pioneer plant formations.

Leguminosae constitute an important proportion of the charcoal sampIes recovered at archaeological sites in the West African savannas. Identification of these fragments to a level below family or subfamily was problematic, because a comparative survey was missing. Therefore, the wood anatomy of 31 species (23 genera) of Mimosoideae and Caesalpinioideae growing in the Sudanian savannas of West Africa was examined. The species were grouped into 18 types according to wood anatomical structure. The types represent single species or genera (fourteen types), two genera (three types) or three genera (one type) . The following features are regarded as suitable for a reliable delimitation and identification . Heterocellular rays and storied structure allow for a first differentiation. Enlarged, non-bordered vessel-ray pitting , nonvestured vessel-pits, silica, axial canals, septate fibres and crystals in non-eharnbered ray cells are additional features characterizing few or single types. Types without these features are delimited less easily. Parenchyma distribution and ray width are, due to variability, not as reliable , but remain necessary features for identification. Types characterized by these features only may not always be recognized correctly. Quantitative features of the vessels are not regarded as helpful for the differentiation within the set of examined species. A table (Table 1) summarizes the results for easy reference.

Bayon et al . (Reports, 9 March 2012, p. 1219) interpreted unusually high aluminum-potassium ratio values in an Atlantic sediment core as indicating anthropogenic deforestation around 2500 years before the present (B.P.). We argue that there is no terrestrial evidence for forest destruction by humans and that the third millennium B.P. rainforest crisis can be clearly attributed mostly to climatic change.

In a paper by Garcin et al. in PNAS (1), it is assumed that a sharp increase in settlement activities in the Central African rainforest during the first millennium BC caused widespread deforestation between 2,600 and 2,020 cal y BP (the late Holocene rainforest crisis or LHRC) (2, 3). Archaeology was only marginally used in this new study by means of a newly compiled radiocarbon database containing 1,202 14C dates from 460 sites in Central Africa covering the past 10,500 y. Those dates must be critically evaluated, as not all of them are relevant for assessing the human factor in the LHRC. Not only the quantity of … [↵][1]1To whom correspondence should be addressed. Email: bernardolivier.clist{at}ugent.be. [1]: #xref-corresp-1-1