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Agathidium (Agathidium) pilulum sp. nov. (Figs 1–4, 13) Type locality. China, Hainan Island, Limushan Mts, 19°10.5–10.9′N, 109°44–45′E, 650– 900 m. Type material. Hඈඅඈඍඒඉൾ: J, “ CHINA: Hainan isl. [MF 19] / Limushan Mts, mountains / above first admin.centre / 19°10.5–10.9′N 109°44–45′E/ 650-900 m, 6.v.2011, Fikáček // sifting small accumula- / tions of moist leaf litter along / and on the trail in secondary / forest partly with Cyathea and / bamboo” (NMPC). Pൺඋൺඍඒඉൾඌ: 5 JJ 2 ♀♀, same data as holotype (NMPC, ZSPC). Description. Length of body in holotype 1.9 mm, head 0.3 mm, pronotum 0.7 mm, elytra 0.9 mm, antenna 0.6 mm, aedeagus 0.74 mm. Maximum width of head 0.8 mm, pronotum 1.2 mm, elytra 1.2 mm. Body short oval. Pronotum chestnut, elytra dark chestnut with feebly expressed reddish narrow strip along suture, head lighter than pronotum. Legs and antennae yellow-red with A9 and A10 a little darker. Ventral surface yellow-brown. Dorsum punctured, without microsculpture. Sutural striae absent; femoral lines developed, supraocular carina raised anteriorly. Head. Maximum width of head far before posterior margin of eyes. Eyes parabola slice-shaped. Supraocular carina low posteriorly, raised anteriorly. Subocular line not developed. Clypeus straight, feebly emarginate, clypeal line lacking (Fig. 13). Antennomere 3 distinctly longer than A2 (A3/A2 = 1.5). Ratio of lengths of A2–A11 (A2 = 1.0): 1.0-1.5-0.8-0.6-0.5-0.5-0.6-1.3-1.1-2.0. Ratio of widths of A2–A11 (A2 = 1.0): 1.0-0.5-0.7-0.8-1.0-1.2- 1.3-1.7-1.8-1.8. Ratio of W/L of A2–A11: 0.8-0.3-0.7- 1.0-1.5-1.8-1.6-1.0-1.2-0.7. Surface of head smooth with very fine very sparse punctures separated by more than 10 times their own diameters. Pronotum. Lateral outline with shortly rounded angle in lateral view (as in Fig. 3). Puncturation sparse and fine, similar to that on head with punctures separated by about 10 or even more times their own diameter. Elytra widest at lateral angles, then evenly roundly narrowed posteriorly. Elytral margin straight behind lateral angles in lateral view (Fig. 3). Lateral elytral channel as narrow behind lateral angles as in basal part. Puncturation similar to that on head but punctures a little stronger, separated by about 6–10 or more times their own diameter. Sutural striae not developed. Legs. Tarsi slim, all tibiae slim, metafemora simple, without specific characters. Tarsal formula: 5-5- 4 in male. Mesoventrite. Posterior part subconcave. Anterior part raised with longitudinal carina. Lateral lines not developed. Metaventrite moderately developed. Femoral lines developed, shortened for half of their possible length. Fovea located in middle of posterior margin of metaventrite equipped with bush of erect setae. Membranous wings developed. Genitalia. Basal part of median lobe narrow ring-shaped of type C. Shape of median lobe apex resembles skittle pin in dorsal view. Apical part of median lobe first contracted, emarginated laterally, then feebly widened. Operculum egg-shaped. Parameres narrowed apically; each with two short and fine setae (aedeagus as in Figs 1, 2). Variation and sexual dimorphism. Length of body is 1.8–2.0 mm in the type series. The ratio of A3/A2 varies between 1.3–1.6. Tarsal formula 5-4- 4 in female. Spermatheca with pear-shaped basal part and simply bowed distal thin part (Fig. 4). Length of spermatheca 0.13 mm. Differential diagnosis. Agathidium (Agathidium) pilulum sp. nov. as well as the species it is compared with bellow habitually belong to the A. madurense species group sensu AඇGൾඅංඇං (1993) due to raised anterior part of the supraocular carina. The new species stands closest to A. (Agathidium) occultum Angelini & De Marzo, 1998 from China, Yunnan (AඇGൾඅංඇං & Dൾ Mൺඋඓඈ 1998b) as both species are similar in the shape of body that is broadly oval, in the shape of straight, feebly emarginated clypeus, in the eyes that are parabola slice-shaped and in the raised anterolateral bead of head. Colour of the antenna is yellow-red with A9 and A10 a little darker in both species. Both compared species possess A3 distinctly longer than A2 and also slim tibiae. Sutural stria is missing in both species. Agathidium pilulum sp. nov. differs from A. occultum in the specific shape of its median lobe that is simply cylindrical with broadly rounded tip in A. occultum. Agathidium pilulum sp. nov. can be also compared to A. (A.) alesi Angelini & De Marzo, 1998 from Taiwan (AඇGൾඅංඇං & Dൾ Mൺඋඓඈ 1998a), that also belongs to the A. madurense species group, having similar skittle pin-shaped apical part of the median lobe. The new species differs from A. alesi in distinctly smaller body (1.8–2.0 mm), while the length of body is 3.0– 4.1 mm in A. alesi. Antenna is bicolored in A. pilulum sp. nov., antenna in A. alesi is unicolorous, light. Posterior femora are straight on their hind margin, while the same are toothed in A. alesi. Moreover, the apex of the tegmen is distinctly bent upwards and paramera is shorter in A. alesi compared with A. pilulum sp. nov. Spermatheca is similar in its shape in all the compared species; basal part of spermatheca is pear-shaped in A. pilulum sp. nov. while the same is subcylindrical in A. occultum and A. alesi. Etymology. The name of the new species draws the attention to the globular shape of the apex of the median lobe. Therefore the name is derived from the Latin word pilula (= globule in English). Noun in apposition., Published as part of Švec, Zdeněk, 2022, Leiodidae (Coleoptera) of the Hainan Island with new faunistic records from China and with notes on the unique body modifications in the genus Agathidium, pp. 155-164 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 62 (1) on pages 157-158, DOI: 10.37520/aemnp.2022.012, http://zenodo.org/record/7161302

In this paper we present an updated checklist of thallus-forming Laboulbeniomycetes (Ascomycota, Pezizomycotina), that is, the orders Herpomycetales and Laboulbeniales, from Belgium and the Netherlands. Two species are newly described based on morphology, molecular data (ITS, LSU ribosomal DNA) and ecology (host association). These areHesperomyces halyziaeonHalyzia sedecimguttata(Coleoptera, Coccinellidae) from both countries andLaboulbenia quarantenaeonBembidion biguttatum(Coleoptera, Carabidae) from Belgium. In addition, nine new country records are presented. For Belgium:Laboulbenia aubryionAmara aranea(Coleoptera, Carabidae) andRhachomyces spinosusonSyntomus foveatus(Coleoptera, Carabidae). For the Netherlands:Chitonomyces melanurusonLaccophilus minutus(Coleoptera, Dytiscidae),Euphoriomyces agathidiionAgathidium laevigatum(Coleoptera, Leiodidae),Laboulbenia fasciculataonOmophron limbatum(Coleoptera, Carabidae),Laboulbenia metabletionSyntomus foveatusandS. truncatellus(Coleoptera, Carabidae),Laboulbenia pseudomaseionPterostichus melanarius(Coleoptera, Carabidae),Rhachomyces canariensisonTrechus obtusus(Coleoptera, Carabidae), andStigmatomyces hydrelliaeonHydrellia albilabris(Diptera, Ephydridae). Finally, an identification key to 140 species of thallus-forming Laboulbeniomycetes in Belgium and the Netherlands is provided. Based on the combined data, we are able to identify mutual gaps that need to be filled as well as weigh the impact of chosen strategies (fieldwork, museum collections) and techniques in these neighboring countries. The aim of this work is to serve as a reference for studying Laboulbeniomycetes fungi in Europe.

A. grandidentatum Miller & Wheeler, 2005b: 114. Holotype male in CMNC. Type locality: 5.6 km SW Filo de Caballo, 2310 m, Guerrero State, Mexico. Distribution: Mexico: Guerrero State: known only from vicinity of type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. erythromelas Miller & Wheeler, 2005b: 97. Holotype male in EMEC. Type locality: 34 mi E Morelia, Michoacán State, Mexico. Distribution: Mexico: México, Michoacán, and Veracruz States.

A. rumsfeldi Miller & Wheeler, 2005b: 123. Holotype male in FMNH. Type locality: 2.5 mi N Tlanchinol, 5200 ft., Hidalgo State, Mexico. Distribution: Mexico: Hidalgo and Oaxaca States., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 68, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. multidentatum Miller & Wheeler, 2005b: 112. Holotype male in CMNC. Type locality: 3.5 mi S Suchixtepec, 8000’, Oaxaca State, Mexico. Distribution: Mexico: Oaxaca State: known only from vicinity of type locality.

A. mexicanum Hendrichs, 1979: 106. Holotype male not in MCMC (as indicated in original description), and not in UNAM (where part of that author’s specimens may be placed) [in Miller & Wheeler, 2005b: 164, specimens not seen] [six paratypes are in the National Collection of Insects, Mexico City, but lacked identification labels (José Luis Navarrete-Heredia, 2019, personal comm. to SBP)]. Original description states some specimens supposedly deposited in SBPC but this was never done. Type locality: Cerro Tlaloc, 3800 m, México State, Mexico. Distribution: Mexico: Ciudad de México and México States. The species is of uncertain status because specimens from the type series were not seen by Miller & Wheeler., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 69, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Hendrichs, S. J. (1979) Nuevo Agathidium de Mexico y sus relaciones zoogeographicas (Coleoptera: Leiodidae). Folia Entomologica Mexicana, 41, 103 - 114.","Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

Genus Agathidium Panzer Agathidium Panzer, 1797: no. 13. Type species: Tetratoma globosa Herbst [1792] (= Silpha seminulum Linnaeus [1758], syn. in ?? [already by Gyllenhal 1810: 573] (by monotypy). Note: Proposed as nomen protectum by provision of ICZN 1999: Art. 23.9 (Wheeler & Miller, 2005: 34). = Volvoxis Kugelann, 1794: 535 (not preoccupied by Volvox Linnaeus [1758], Chlorophyta). Type species: Tetratoma globosa Herbst (= Silpha seminulum Linnaeus [1758] by subsequent designation of Newton, 1998: 95 (10 species in original description). Note: Proposed as nomen oblitum by provision of ICZN 1999: Art. 23.9 (Wheeler & Miller, 2005: 34) due to lack of use as a valid name after 1899. = Agathidium Illiger, 1798: 81 (unnecessary replacement name for Volvoxis Kugelann [1794] (in part), incorrectly assumed to be preoccupied by Volvox Linnaeus [1758]). Type species: Tetratoma globosa Herbst [1792] (= Silpha seminulum Linnaeus [1785]), fixed by autotypy as replacement name for Volvoxis. Note: It is not clear if Illiger intended to propose a new generic name here; he did not refer the generic name to Panzer (1797), although he did refer to Panzer’s (1797: 13) species treatment of Agathidium globosum. The above treatment assumes it was an intentional proposal of a new name to replace Volvoxis. Note 1: Volvoxis Kugelann has priority for the name of this genus, but this name was rejected and replaced with Agathidium by Illiger (1798) in the mistaken belief that Volvoxis was preoccupied by Volvox Linnaeus, and this replacement name became generally accepted. However, Panzer (1797) had already used Agathidium in the same sense a year earlier, without explanation. Illiger (1798) made no mention of Panzer’s (1797) use of Agathidium so these have often been treated as independent names (e.g., Newton, 1998: 95 – 96), with Panzer’s name having priority and Illiger’s name being a junior homonym of it. All three names have the same type species as indicated above, and thus are objective synonyms. Wheeler & Miller (2005: 34) cited Article 23.9 of the Code (ICZN, 1999) to set aside priority and accept Agathidium Panzer as the valid name for the genus (nomen protectum) and treat Volvoxis as a forgotten name (nomen oblitum). Agathidium has been attributed to both Panzer and Illiger at various times, sometimes (by accident) even in the same publication (see Newton, 1998: 94, 95), but Panzer, 1797 is correct. Note 2: Revised in Hlisnikovský (1964a) and Miller & Wheeler (2005b) and Wheeler & Miller (2005). Note 3: Wheeler & Miller (2005) abandoned the use of subgenera and proposed a system of species groups, as used below, which may have some relationship to previously established subgenera. Most Mexican and Central American species are in the oniscoides species group. The rich species diversity of Mexico and Central America is obviously only partly known. Key: to species groups in Wheeler & Miller, 2005: 37. Distribution: 822 spp.—Holarctic, Africa, Oriental, northern Neotropical. Note 4— Species wrongly reported for the Neotropics: A. exiguum Melsheimer [1844] —The record of A. californicum Horn [1880] [= A. exiguum, Miller & Wheel- er, 2005b: 156 syn] for Guatemala (Quiché Mts., 7000 – 9000 ft.) in Matthews, 1887: 75 is a misidentification of an unknown different species (Miller & Wheeler, 2005b: 160). A. oniscoides Palisot de Beauvois [1805] —The record of Matthews, 1887: 74 for Mexico (Jacala, Hidalgo State) is in error (a misidentification, in Miller & Wheeler, 2005b: 151). Biology: Usually in forested habitats and on slime molds (Hatch, 1957; Russell, 1979; Newton, 1984; Wheeler, 1987, 1990)., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on pages 66-67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg.","ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth edition. London. International Trust for Zoological Nomenclature. Available from: http: // www. iczn. org / iczn / index. jsp (accessed 24 September 2019)","Wheeler, Q. D. & Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 2.0. CO; 2","Kugelann, J. G. (1794) Verzeichniss der in einigen Gegenden Preussens bis jetzt entdeckten Kaefer-Arten nebst kurzen Nachrichten von denselben. Neuestes Magazin fur die Liebhaber der Entomologie, 1, 513 - 582.","Newton, A. F. Jr. (1998) Phylogenetic problems, current classification and generic catalog of World Leiodidae (including Cholevidae). Museo Regionale di Scienze Naturali di Torino Atti, 8, 41 - 177.","Illiger, J. K. W. (1798) Verzeichniss der Kafer Preussens. J. J. Gebauer, Halle, xlii + 510 pp.","Hlisnikovsky, J. (1964 a) Monographische bearbeitung der gattung Agathidium Panzer (Coleoptera). Acta Entomologica Musei Nationalis Prague, Supplementum 5, 3 - 239.","Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2","Horn, G. H. (1880) Synopsis of the Silphidae of the United States with reference to the genera of other countries. Transactions of the American Entomological Society, 8, 219 - 319, pls. 5 - 7. https: // doi. org / 10.2307 / 25076393","Matthews, A. (1887 - 88) Fam. Silphidae. In: Biologia Centrali-Americana. Insects. Coleoptera. Vol. 2 (1). Taylor & Francis, London, pp. 72 - 101, pl. 3.","Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II. Staphyliniformia. University of Washington Publications in Biology, 16, i-ix + 1 - 384.","Russell, L. K. (1979) Beetles associated with slime molds (Mycetozoa) in Oregon and California (Coleoptera: Leiodidae, Sphindidae, Lathridiidae). Pan-Pacific Entomologist, 55, 1 - 9.","Newton, A. F. Jr. (1984) Mycophagy in Staphylinoidea (Coleoptera). In: Wheeler, Q. & Blackwell, M. (Eds.), Fungus-insect relationships: perspectives in ecology and evolution. Columbia University Press, New York, pp. 302 - 353.","Wheeler, Q. D. (1987) A new species of Agathidium associated with an \" epimycetic \" slime mold plasmodium on Pleurotus fungi (Coleoptera: Leiodidae-Myxomycetes: Physarales-Basidiomycetes: Tricholomataceae). Coleopterists Bulletin, 41, 395 - 403.","Wheeler, Q. D. (1990) Morphology and ontogeny of postembryonic larval Agathidium and Anisotoma (Coleoptera: Leiodidae). American Museum Novitates, 2986, 1 - 46."]}

A. chauliodoum Miller & Wheeler, 2005b: 145. Holotype male in CMNC. Type locality: 7.5 km S Purulha, 1730 m, Baja Verapaz Department, Guatemala. Distribution: Guatemala: Baja Verapaz Department: known only from vicinity of type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on pages 68-69, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. disgregum Miller & Wheeler, 2005b: 116. Holotype male in CNCI. Type locality: Cerro Potosí, 8500’, Nuevo León State, Mexico. Distribution: Mexico: Nuevo León State: known only from type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. lobosternum Miller & Wheeler, 2005b: 103. Holotype male in CMNC. Type locality: Tlanchinol, 1500 m, 43 km SW Huejutla, Hidalgo State, Mexico. Distribution: Mexico: Hidalgo State: known only from type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 68, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. tenangoense Miller & Wheeler, 2005b: 121. Holotype male in CMNC. Type locality: 10 km W La Capellania, Huehuetenango Department, Guatemala. Distribution: Guatemala: Huehuetenango Department: known only from type locality.

A. nimbosilva Miller & Wheeler, 2005b: 148. Holotype male in CMNC. Type locality: 7 km E Purulha, 1600 m, Baja Verapaz Department, Guatemala. Distribution: Guatemala: Baja Verapaz Department: known only from type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 69, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. oaxacaense Miller & Wheeler, 2005b: 117. Holotype male in CMNC. Type locality: 37 mi S Valle Nacional, 8500’, Oaxaca State, Mexico. Distribution: Mexico: Oaxaca State: known only from type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 68, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. skoliosternum Miller & Wheeler, 2005b: 96. Holotype male in CMNC. Type locality: 7 km E Huatusco, 1040 m, Veracruz State, Mexico. Distribution: Mexico: Puebla and Veracruz States., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 68, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. hidalgoense Miller & Wheeler, 2005b: 95. Holotype male in CMNC. Type locality: Tlanchinol, 1500 m, 43 km SW Huejutla, Hidalgo State, Mexico. Distribution: Mexico: Hidalgo State: known only from type locality.

A. iota Miller & Wheeler, 2005b: 69. Holotype male in AMNH. Type locality: 14 mi W Xilitla, 4800 ft., San Luis Potosí State, Mexico. Distribution: Guatemala: Baja Verapaz Department; Mexico: Guerrero, Michoacán, Oaxaca, and San Luis Potosí States., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. minutum Peck & Cook, 2014: 13. Holotype male in WIBF. Type locality: Parque Nacional Los Haitises, W of Sabana de la Mar, Hato Mayor Province, Dominican Republic. Distribution: Dominican Republic: Hato Mayor Province: known only from type locality.

A. aztec Miller & Wheeler, 2005b: 127. Holotype male in CMNC. Type locality: 24 km N Xicotepec de Juarez, 1070 m, Puebla State, Mexico. Distribution: Mexico: Chiapas, Oaxaca, Puebla, and Veracruz States., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. popocatepetlae Miller & Wheeler, 2005b: 105. Holotype male in CNCI. Type locality: Volcan Popocatepetl, km 10, 10000’, México State, Mexico. Distribution: Mexico: México State: known only from type locality., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 68, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. bituberculatum Miller & Wheeler, 2005b: 91. Holotype male in CMNC. Type locality: Cerro Hultepec, 2700 m, ca. 5 km W San Cristobal, Chiapas State, Mexico. Distribution: Mexico: Chiapas State., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 67, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2"]}

A. difficile Matthews, 1887: 76; Hlisnikovský, 1964a: 232; Miller & Wheeler, 2005b: 163 (lectotype designation). No information about types given in original description; Lectotype female (designated in Miller & Wheeler, 2005b; who mentioned number of specimens not given in original description, one single specimen available for study) in BMNH. Type locality: Cerro de Plumas [= Cerro de Palmas, in Veracruz State, according to Selander & Vaurie (1962), but as Mexico State in Miller & Wheeler (2005b), without explanation], Mexico. Distribution: Mexico: Veracruz State: known only from type locality. Note: The species is of uncertain status because the lectotype is a female., Published as part of Peck, Stewart B., Gnaspini, Pedro & Newton, Alfred F., 2020, Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (" Latin America ": Mexico, the West Indies, and Central and South America), pp. 1-114 in Zootaxa 4741 (1) on page 69, DOI: 10.11646/zootaxa.4741.1.1, http://zenodo.org/record/3772899, {"references":["Matthews, A. (1887 - 88) Fam. Silphidae. In: Biologia Centrali-Americana. Insects. Coleoptera. Vol. 2 (1). Taylor & Francis, London, pp. 72 - 101, pl. 3.","Hlisnikovsky, J. (1964 a) Monographische bearbeitung der gattung Agathidium Panzer (Coleoptera). Acta Entomologica Musei Nationalis Prague, Supplementum 5, 3 - 239.","Miller, K. B. & Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 2.0. CO; 2","Selander, R. B. & Vaurie, P. (1962) A gazetteer to accompany the \" Insecta \" volumes of the \" Biologia Centrali-Americana \". American Museum Novitates, 2099, 1 - 70."]}

Agathidium mexicanum Hendrichs, 1979: Types rediscovered JOS�� L. NAVARRETE-HEREDIA 1, LILIANA P. SANDOVAL 1 & SANTIAGO ZARAGOZA-CABALLERO 2 1 Centro de Estudios en Zoolog��a, CUCBA, Universidad de Guadalajara, Apdo. Postal 134, 45100, Zapopan, Jalisco, M��xico; 2 Instituto de Biolog��a, Universidad Nacional Aut��noma de M��xico, 04510 Coyoac��n, Ciudad de M��xico, M��xico Agathidium mexicanum Hendrichs, 1979: 106 is the single Mexican species of Leiodidae described by a Mexican entomologist: Chemical Engineer Jorge Hendrichs. As an amateur coleopterist he made a beetle collection of 18,400 specimens, adequately labeled by his wife and daughter. Most of the specimens are now in the entomological collection of the Instituto de Biolog��a, UNAM. He published 12 papers, one of them devoted to the description of one leiodid species: Agathidium mexicanum Hendrichs, 1979. Type specimens of A. mexicanum were collected in Estado de Mexico and Distrito Federal (now Ciudad de M��xico) localities (type locality: Cerro Tlaloc, Estado de M��xico). The type series as described in the original description was constituted by 13 specimens: Holotype (male), allotype (female) and, 11 paratypes (6 males and 5 females). For specific details, data of the material examined is fully included (Hendrichs 1979: 110): ��� ESTADO DE M��XICO: Cerro Tlaloc 3800 m, 29-I-72, holotipo ♂, alotipo ♀ y 2 paratipos ♀ ♀; R��o Fr��o: 3300 m, 30- I-72, 2 paratipos ♂ ♂ y 1 paratipo ♀; Jalatlaco: 3400 m, 29.-XII-77, 1 paratipo ♂ y 1 paratipo ♀; Lagunas de Zempoala: 3100 m, 16-IV-76, 1 paratipo ♀. DISTRITO FEDERAL: Desierto de los Leones, 3300 m, 2-XII-72, 1 paratipo ♂; Agua de Leones, 3300m, 13-IV-74, 1 paratipo ♂; Puerto de las Cruces, 3100m, 24-IX-72, 1 paratipo ♂. Todos colectados por Jorge Hendrichs S. Y Jorge Hendrichs N.���. ���El holotipo y un paratipo ♀ se depositan en el Museo de Historia Natural de la Ciudad de M��xico. Dos paratipos en la colecci��n del Dr. S. B. Peck, el alotipo y resto de los paratipos en la colecci��n del autor��� Holotype and one paratype female to be deposited in the entomological collection of the Museo de Historia Natural de la Ciudad de M��xico. Two paratypes in the personal collection of Dr. S. B. Peck, allotype and remaining paratypes in the author collection (Hendrichs 1979: 110). Agathidium mexicanum is cited in the synopsis of Mexican Leiodidae by Peck (2000: 449). Later it is cited again in Miller and Wheeler (2005: 164) with a comment that the holotype ���is not in either Universidad Nacional Aut��noma de Mexico (UNAM) or the Museo de Historia Natural de la Ciudad de Mexico ���. Zaragoza-Caballero and P��rez-Hern��ndez (2017) published a catalog of Coleoptera types in the entomological collection at UNAM (CNIN) and include some leiodid species, but A. mexicanum is not mentioned. In a recent visit to the Entomological Collection of the Institute of Biology at Universidad Nacional Aut��noma de M��xico we searched again in the CNIN collection looking for unidentified leiodids. After a close examination we found ten Agathidium specimens without identification labels. All the specimens were part of the Hendrichs collection. Later we visited the entomological collection of the Museo de Historia Natural de la Ciudad de M��xico. Biol. Mar��a E. D��az Batres, curator of the collection for many years, told us that no types of the species were deposited in that collection. There are type specimens but of other leiodid species. She suspected that types will be in the entomological collection of the Institute of Ecology where some specimens of the J. Hendrichs collection are deposited. We asked Dr. C. Deloya and M.C. Leonardo Delgado, coleopterists of the Institute of Ecology, about such specimens, and no specimens were found. Finally, we asked Dr. S. Peck if he received the paratypes. He confirmed that no paratypes had been deposited in his collection. After a careful and detailed examination of the specimens found at CNIN we conclude that the ten specimens are part of the type series of Agathidium mexicanum (three specimens are lost) but without type labels. These specimens are clearly an Agathidium species and label data are the same as label data of the type series of A. mexicanum cited in the original description (Hendrichs, 1979). So, the type series is constituted as (data from labels; information in bold is not included in the original publication): ��� Tlaloc, Mex., 29.I.72, 3800 m, bajo tronco, Col. J. Hendrichs/ macho 1/ Ex Colecci��n Jorge Hendrichs S./ H. As the type series indicates that holotype is a male from Tlaloc and three paratype females from the same locality, this specimens we assume correspond to the holotype. So, a holotype label was added to the specimen. Remaining specimens were recognized as paratypes and labels were attached to each one. ��� Tlaloc, Mex., 29.I.72, 3800 m, bajo corteza, Col. J. Hendrichs/ hembra 2/ Ex Colecci��n Jorge Hendrichs S./ H. ��� Tlaloc, Mex., 29.I.72, 3800 m, bajo corteza, Col. J. Hendrichs/ hembra 3/ Ex Colecci��n Jorge Hendrichs S. /H. One specimen of this series is lost. Original locality cited as Cerro Tlaloc. ��� R��o Fr��o, Mex, 30.I.72, 3300m, Kiefernrinde [pine bark], Col. J. Hendrichs S./ Macho/ Ex Colecci��n Jorge Hendrichs S./ H. One adult plus a pronotum and one elytrum. ��� R��o Fr��o, Mex, 30.I.72, 3300m, Kiefernrinde, Col. J. Hendrichs S./ Hembra 8/ Ex Colecci��n Jorge Hendrichs S. ��� 8 km al este de Jalatlaco, Mex, 3400m, corteza de pino, Col. J. Hendrichs S./ one male and one female specimens/ Ex Colecci��n Jorge Hendrichs S. In the original paper, some specific data were omitted as is the case of the specimens from R��o Fr��o where Kiefernrinde [pine bark], Col. J. Hendrichs S./ Macho/ Ex Colecci��n Jorge Hendrichs S./ H. were omitted. We assume that the specific locality data of these specimens is the same as that mentioned in the paper just as Jalatlaco. Specimens were in the same lot and sex corresponds to the sex cited for the paratypes. So, we assume that these specimens are also specimens of the type series. ��� The specimen from Lagunas de Zempoala is lost. ��� The specimen from Desierto de los Leones is lost. ��� Agua de Leones, Desierto, D.F, 13.IV.74, 3300 m, trampa c. cebo, Col. J. Hendrichs S./ macho 6/ Ex Colecci��n Jorge Hendrichs S./ H. ��� Las Cruces, D.F., 24.IX.72,3100 m, Kiefernrinde, Col. J. Hendrichs S./ Macho 5/ Ex Colecci��n Jorge Hendrichs S. /H. The original locality cited as Puerto de las Cruces. Specimens are deposited at the Entomological Collection of the Institute of Biology (CNIN) in the type specimen collection. Specimens from Jalatlaco are in the Entomological Collection of the Centro de Estudios en Zoolog��a, Universidad de Guadalajara., Published as part of Navarrete-Heredia, Jos�� L., Sandoval, Liliana P. & Zaragoza-Caballero, Santiago, 2020, Agathidium mexicanum Hendrichs, 1979: Types rediscovered, pp. 295-296 in Zootaxa 4731 (2) on pages 295-296, DOI: 10.11646/zootaxa.4731.2.11, http://zenodo.org/record/3754822, {"references":["Hendrichs, S. J. (1979) Nuevo Agathidium de Mexico y sus relaciones zoogeograficas (Coleoptera: Leiodidae). Folia Entomologica Mexicana, 41, 103 - 114.","Peck, S. B. (2000) Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: hacia unas sintesis de su conocimiento. In: Llorente Bousquets, J., Gonzalez Soriano, E. & Papavero, N. (Eds.), Leiodidae (Coleoptera). Universidad Nacional Autonoma de Mexico, Mexico City, pp. 439 - 452.","Miller, K. B. & Wheeler, Q. D. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 % 3 C 0001: SBOTGA % 3 E 2.0. CO; 2","Zaragoza-Caballero, S. & Perez-Hernandez, C. X. (2017) An annotated catalogue of the Coleoptera types deposited in the National Insect Collection (CNIN) of the National Autonomous University of Mexico. Zootaxa, 4288 (1), 1 - 128. https: // doi. org / 10.11646 / zootaxa. 4288.1.1"]}

Agathidium mexicanum Hendrichs, 1979: 106 is the single Mexican species of Leiodidae described by a Mexican entomologist: Chemical Engineer Jorge Hendrichs. As an amateur coleopterist he made a beetle collection of 18,400 specimens, adequately labeled by his wife and daughter. Most of the specimens are now in the entomological collection of the Instituto de Biología, UNAM. He published 12 papers, one of them devoted to the description of one leiodid species: Agathidium mexicanum Hendrichs, 1979. Type specimens of A. mexicanum were collected in Estado de Mexico and Distrito Federal (now Ciudad de México) localities (type locality: Cerro Tlaloc, Estado de México). The type series as described in the original description was constituted by 13 specimens: Holotype (male), allotype (female) and, 11 paratypes (6 males and 5 females). For specific details, data of the material examined is fully included (Hendrichs 1979: 110)

Agathidium arcticum (Thomson 1862) (Leiodidae) Dies ist eine Art mit fragmentarischer Verbreitung ausserhalb Nordeuropas. Der Arealtyp von Agathidium arcticum wird üblicherweise als boreoalpin benannt, in Mitteleuropa sind nur Funde aus den Alpen bekannt geworden. In Frankreich und Mittelengland finden sich Reliktvorkommen allerdings in mässiger bis mittlerer Höhenlage, und die Verhältnisse an der südlichen Grenze des geschlossenen Vorkommens in Skandinavien entsprechen etwa dem mittleren Montanbereich in Mitteleuropa. Der Käfer gilt als fakultativ xylobiont, wurde an Myzelien und Fruchtkörpern verschiedener Pilze und Myxomyceten hauptsächlich an Totholz gefunden (Wheeler & Miller 2005)., Published as part of Huber, Barbara, Gubelmann, Päivi & Büche, Boris, 2019, Vielfalt der Totholzkäferfauna in obersubalpinen Lärchen-Arvenwäldern im Avers (Graubünden), pp. 111-128 in Entomo Helvetica 12 on page 118, DOI: 10.5169/seals-985866, http://zenodo.org/record/8072444, {"references":["Wheeler Q. D. & Miller K. B. 2005. Slime-mold beetles of the genus Agathidium Panzer in North and Central America: Coleoptera, Leiodidae. Part 1. Bulletin of the American Museum of Natural History 290: 1 - 95."]}

Se relata la primera captura de Agathidium ibericum desde su descripción original y la manera de distinguirlo de la especie más cercana Agathidium convexum Sharp. The first capture of Agathidium ibericum since its original description and how to distinguish it from the nearest species Agathidium convexum Sharp is showed. Los muestreos fueron financiados por el proyecto del Ministerio de Ciencia e Innovación, Spain (CGL2011-23658) dirigido por la Dr. Micó.

Agathidium (Agathidium) denticulatum Hlisnikovsk��, 1972 Agathidium (s.str.) denticulatum Hlisnikovsk��, 1972: 837. Type locality Turkey, Antakya, Osmaniye-Zorhun, 500 m. Distribution in Iran. Guilan, Nav's Valley, 1800 m, 6.VIII.1973 (leg. Vit, CV) (Angelini 1988). General distribution. Iran, Israel, Lebanon, Russia (South European territories, Caucasus), Syria, Turkey (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Neoceble) plagiatum (Gyllenhal, 1810) Anisotoma plagiatum Gyllenhal, 1810: 575. Type locality: [Sweden:] in silva Kinnensi Westrogothiae. Agathidium (Cyphoceble) besucheti Hlisnikovsk��, 1972: 829. Type locality: Turky, Bolu, Emalik, alt. 950 m. Agathidium (Neoceble) garganicum Fiori, 1915: 71. Type locality: Italy, Puglia, Gargano, Foresta Umbra. Agathidium hellenicum Reitter, 1884: 116. Type locality: Greece, Morea, Kumani. Agathidium (Saccoceble) hummleri Reitter, 1908: 243. Type locality: Italy, Calabria, Aspromonte. Agathidium (Neoceble) roubali Hlisnikovsk��, 1964a: 101. Type locality: Russia, Caucasus, Krasnaja Poljana, Sotschi. Agathidium (Neoceble) signatipenne Hlisnikovsk��, 1964a: 73. Type locality: Russia, Caucasus, Fischt, north Sotschi. Agathidium striatum Krynicki, 1829: 199. Type locality: Charkovia. Agathidium vittatum Motschoulsky, 1839: 53. Type locality: Montagnes du Caucase, Sabui. Distribution in Iran. MaZandaran, road to Tchorteh, 800 m, 3 eXX., 5.VIII.1974; MaZandaran, road to Tchorteh, 1000���1300 m, 1 eXX., 8.VII.1973; Guilan, south Asalem, 300���600 m, 1 eX., 30.IV.1973; Guilan, road to Here- Abad, 1600 m, 10.VIII.1973 (leg. Senglet, CA-MGE; MG); Guilan, Nav's Valley, 1800 m, 6.VIII.1973 (leg. Vit, CV) (Angelini 1988). General distribution. Europe (not in Great Britain, The Netherlands, Denmark, Albania), Armenia, AZerbaijan, Georgia, Iran, Turkey (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Gyllenhal, L. (1810) Insecta Svecica descripta. Classis I. Coleoptera sive Eleuterata. Tomus 1. Pars II. L. J. Leverentz, Scaris, xix + 660 pp.","Fiori, A. (1915) Appunti sulla fauna coleotterologica dell'Italia meridionale e della Sicilia. Rivista Coleotterologica italiana, 13 (4 - 9), 57 - 84.","Reitter, E. (1884) Bestimmungs-Tabellen der Europaischen Coleopteren XII. Necrophaga (Platypsyllidae, Leptinidae, Silphidae, Anisotomidae und Clambidae). Verhandlungen des naturforschenden Vereines in Brunn, 23, 3 - 122.","Reitter, E. (1908) Acht neue Coleopteren aus Europa, Kleinasien und Turkestan. Wiener entomologische Zeitung, 27 (8), 243 - 248.","Hlisnikovsky, J. (1964 a) Monographische Bearbeitung der Gattung Agathidium Panzer (Coleoptera). Acta faunistica Entomologica Musei nationalis Pragae, 5 (Supplementum), 1 - 255.","Krynicki, J. (1829) Literae cel. professoris J. Krynicki directori datae, Charkovia 28 Julii 1829. Bulletin de la Societe imperiale des naturalistes de Moscou, 1, 187 - 199.","Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Agathidium) mequignoni Roubal, 1911 Agathidium mequignoni Roubal, 1911: 49. Type locality: Russia, Krasnodar, Krasnaja Poljana. Agathidium (s.str.) melichari Hlisnikovsk��, 1964a: 205. Type locality: South European Russia, Caucasus, Teberda. Agathidium (s.str.) orientale Hlisnikovsk��, 1964a: 212. Type locality: Turkmenistan, Terski-tau, Issyk-Kul. Distribution in Iran. Guilan, Nav's Valley, 1800 m, 4 eXX, 6.VIII.1973 (leg. Vit, CA-MGE; CV); Guilan, south Asalem, 300���600 m, 1 eX., 30.VI.1973 (leg. Senglet, MG); MaZandaran, Nowshahr, Caspian Sea, 1 eX., VIII- IX.1961 (leg. Klapperich, CA-MGE); Northern Khorasan, Kopet-Dagh, Siaret, 1160 m, 1 eX. (leg. Hauser, CA- MGE); Golestan, Gole Lovae, 750���1400 m, 3.V.1970 (leg. Wittmer & Bothmer, NMB; MTO) (Angelini 1988). General distribution. Armenia, AZerbaijan, China (Beijing), Georgia, Iran, South Korea, KyrgyZstan, Russia (South European Territories; Siberia), Taiwan, Turkey (Angelini 1995; 2004; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Roubal, J. (1911) Koleopterologicke vysledky me cesty na Kavkaz v cervenci r. 1910. Casopis ceskoslovenske Spolecnosti Entomologicke, 8, 48 - 50.","Hlisnikovsky, J. (1964 a) Monographische Bearbeitung der Gattung Agathidium Panzer (Coleoptera). Acta faunistica Entomologica Musei nationalis Pragae, 5 (Supplementum), 1 - 255.","Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Angelini, F. (2004) Chiavi dicotomiche e catalogo degli Agathidium Panzer dell'Asia sudorientale e della Cina (Coleoptera, Leiodidae, Agathidiini). Annali del Museo civico di storia naturale Giacomo Doria, Genova, 95, 135 - 577.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Neoceble) marginatum Sturm, 1807 Agathidium marginatum Sturm, 1807: 62. Type locality: Oesterreich. Agathidium (Neoceble) graecum Hlisnikovsk��, 1968: 193. Type locality: Greece, Thessalien, Volo. Agathidium (Neoceble) mongolicum Hlisnikovsk��, 1967b: 245. Type locality: Mongolia, Central aimak, SO von Somon Bajanzogt, 1600 m. Agathidium pumilum Hardy, 1852: 244. Type locality: Great Britain, Ravensworth. North Sunderland, Hartlepool; Scotland, Berwickshire and Aberdeenshire. Agathidium (Neoceble) reticulatum Hlisnikovsk��, 1964a: 108. Type loc.: Iran, Kopet-Dag, Siaret, 1160 m. Agathidium (Neoceble) turkestanicum Hlisnikovsk��, 1964a: 106. Type locality: Turkmenistan, Aksu, 1067 m. Distribution in Iran. Northern Khorasan, Kopet-Dagh, 1160 m, VI.1899 (leg. Hauser, NMP, holotype of reticulatum Hlisnikovsk��, 1964) (Angelini 1988). General distribution. Europe (not in Portugal, Corsica, Sardinia, Albania, Crete), Algeria, Armenia, Cyprus, Iran, KaZakhstan, KyrgyZstan, Mongolia, Russia (south European territories, Siberia), Syria, Turkey (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Sturm, J. (1807) Deutschlands Fauna in Abbildung nach der Natur mit Beschreibungen. V. Abtheilung. Die Insecten. Zweites Banden. Kafer. J. Sturm, Nurnberg, 279 pp.","Hlisnikovsky, J. (1968) Uber neue Arten der Gattung Agathidium Panz. (Col. Liodidae). Entomologische Arbeiten aus dem Museum G. Frey, 19, 193 - 198.","Hlisnikovsky, J. (1967 b) 89. Agathidiini. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Coleoptera). Reichenbachia, 9 (27), 237 - 248.","Hardy, J. (1852) Hardy, J. & Thomas, J. B. A catalogue of the insects of Northumberland and Durham. Transactions of the Tyneside naturalist's field club, 2 (2), 165 - 287.","Hlisnikovsky, J. (1964 a) Monographische Bearbeitung der Gattung Agathidium Panzer (Coleoptera). Acta faunistica Entomologica Musei nationalis Pragae, 5 (Supplementum), 1 - 255.","Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Neoceble) clavulum Reitter, 1888 Agathidium clavulum Reitter, 1888: 156. Type locality: Russia, Carachaevo-Cherkessy. Agathidium (Neoceble) roubalianum Hlisnikovsk��, 1964a: 112. Type locality: Russia, Caucasus, Krasnaja Poljana. Distribution in Iran. MaZandaran, road to Tchorteh, 800 m, 2 eXX., 5.VIII.1974 (leg. Senglet, CA-MGE; MG); Guilan, Nav's Valley, 1500 m, 3 eXX., 6.VIII.1973 (leg. Vit, CV) (Angelini 1988). General distribution. AZerbaijan, Georgia, Iran, Russia (South European territories), Turkey (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Reitter, E. (1888) Coleopteren aus Circassien, gesammelt von Hans Leder im Jahre 1887. III. Theil. Wiener entomologische Zeitung, 7 (4), 143 - 156.","Hlisnikovsky, J. (1964 a) Monographische Bearbeitung der Gattung Agathidium Panzer (Coleoptera). Acta faunistica Entomologica Musei nationalis Pragae, 5 (Supplementum), 1 - 255.","Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Agathidium) laevigatum laevigatum Erichson, 1845 Agathidium laevigatum Erichson, 1845: 98. Type locality: Austria. Agathidium affine Stephens, 1929: 182. Type locality: Great Britain. Agathidium helveticum Hlisnikovsk��, 1974: 110. Type locality: Switzerland, Gen��ve, Mategnin. Agathidium (s.str.) laevigatum kostelniki Hlisnikovsk��, 1964a: 218. Type locality: South European Russia, Caucasus, Teberda. Agathidium (s.str.) languidum Hlisnikovsk��, 1967b: 247. Type locality: Mongolei, Central aimak, 126 km N von Ulan-Baator, am Wege in 1100 m H��he. Agathidium (s.str.) laevigatum sibiricum Hlisnikovsk��, 1964a: 218. Type locality: Russia, Ost-Sibirien, Irkut-Quellgebiet. Agathidium (s.str.) teberdense Hlisnikovsk��, 1964a: 214. Type locality: South European Russia, Caucasus, Teberda. Distribution in Iran. Guilan, Asalem, 450 m, 10.VI.1975 (leg. Senglet, MG) (Angelini 1988). General distribution. Europe from Iberian peninsula north to Fennoscandia, Armenia, AZerbaijan, Cyprus, Georgia, Iran, Israel, Mongolia, Russia (European and Asian territories), Turkey, Himalaya (north India, Sikkim, Nepal and Bhutan) (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Erichson, W. F. (1845) [I., II. Lieferungen]. In: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung: Coleoptera; dritter Band. Nicholaische Verlags-Buchhandlung, Berlin, pp. 1 - 320","Hlisnikovsky, J. (1974) Beitrag zur Kenntnis der Agathidiini der Schweiz. Mitteilungen der schweizerischen entomologischen Gesellschaft, 47 (1 - 2), 109 - 112.","Hlisnikovsky, J. (1964 a) Monographische Bearbeitung der Gattung Agathidium Panzer (Coleoptera). Acta faunistica Entomologica Musei nationalis Pragae, 5 (Supplementum), 1 - 255.","Hlisnikovsky, J. (1967 b) 89. Agathidiini. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Coleoptera). Reichenbachia, 9 (27), 237 - 248.","Angelini, F. & De Marzo, L. (1988) Anisotomini del Mediterraneo orientale e Iran con descrizione di una nuova specie (Coleoptera, Leiodidae). Revue suisse de Zoologie, Geneve, 95, 277 - 288.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Agathidium (Agathidium) atrum (Paykull, 1798) Sphaeridium atrum Paykull, 1798: 67. Type locality: Su��de, Westrogothia. Agathidium rufipes Stephens, 1832: 404. Type locality: Great Britain, Edinburg. Distribution in Iran. Guilan (IZW) (Angelini 1995). General distribution. Europe, Armenia, AZerbaijan, Iran, Georgia, Russia, Siberia, Turkey, (Angelini 1995; Perreau 2015)., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Paykull, G. (1798) Fauna svecica. Insecta. Tomus I. Joh. F. Edman, Upsaliae, 10 + 358 pp.","Stephens, J. F. (1832) Illustrations of British Entomology; or, a synopsis of indigeneous Insects: containing their generic and specific distinctions; with an account of their metamophoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V. Baldwin and Cradock, London, 448 pp.","Angelini, F. (1995) Revisione tassonomica delle specie paleartiche del genere Agathidium Panzer (Coleoptera, Leiodidae, Agathidiini). Monografie di Museo Regionale di Scienze Naturali, Torino, 18, 1 - 484.","Perreau, M. (2015) Leiodidae. In: Lobl, I. & Lobl, D. (Eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 180 - 290"]}

Genus Agathidium Panzer, 1797 Volvoxis Kugelann, 1794: 535, type species: Tetratoma globosa Herbst, 1791 (= Agathidium seminulum Linn��, 1758) (Nomen oblitum). Agathidium Panzer, 1797: 13, type species: Tetratoma globosa Herbst, 1791 (= Agathidium seminulum Linn��, 1758) (Nomen protectum). Sphaeroliodes Portevin, 1905: 419, type species: Sphaeroliodes rufescens Portevin, 1905., Published as part of Perreau, Michel, Ghahari, Hassan & Angelini, Fernando, 2017, A checklist of Leiodidae (Coleoptera: Staphylinoidea) of Iran in Zootaxa 4286 (1), DOI: 10.11646/zootaxa.4286.1.3, http://zenodo.org/record/828324, {"references":["Panzer, G. W. (1797) Fauna insectorum germanicae initia oder Deutschlands Insekten. Larhang. Vierter lahrgang. Heft 37. Felssecker, Nurnberg, 24 pp., 24 pls.","Kugelann, J. G. (1794) Verzeichnis der in einigen Gegenden Preussens bis jetzt entdeckten Kafer-Arten nebst kurzen Nachrichten von denselben, in Schneider. Neuestes Magazin fur die Liebhaber der Entomologie, 1 (5), 513 - 582.","Herbst, J. F. W. (1791) Natursystem aller bekannten in- und auslandischen Insekten, als eine Fortsetzung der von Buffonschen Naturgeschichte. Der Kafer vierter Theil. J. Pauli, Berlin, xi + 346 pp.","Portevin, G. (1905) Troisieme note sur les Silphides du Museum. Bulletin du Museum national d'Histoire naturelle de Paris, 11 (6), 418 - 424."]}