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Genus Abacaecus n. gen. Type species. Abacaecus walterrossii n. sp. Differential diagnosis. A genus of the tribe Abacetini Chaudoir, 1873 (sensu Will & Park, 2008) distinguished by the combination of second antennomere eccentrically inserted on first, strongly transverse mentum, a parascutellar setigerous pore at base of stria 2, angular base of stria 1 (������scutellar stria������ auct.) absent, median lobe of aedeagus distally almost rectilinear in lateral view and ostium in dorsal position. It differs from most species of this tribe by body slender and parallel sided, pronotum narrow with markedly protruding fore angles, eyes strongly reduced and not functional. Description. The description of this monospecific new genus is coincident with the description of the type species, Abacaecus walterrossii n. sp. Remarks. The attribution of this new genus to the tribe Abacetini sensu Will & Park (2008) is based on a set of characters some of which can be regarded as synapomorphic for Abacetini sensu stricto, such as the eccentrically inserted second antennomere and the strongly transverse mentum, and some others which are recurrent in this tribe, although also present in other pterostichine-grade taxa, such as the single pair of deeply impressed linear basal impressions of the pronotum, the angular base of stria 1 (������scutellar stria������ auct.) absent, no dorsal puncture on the third elytral interval, the parascutellar elytral puncture at stria two. Etymology. The genus name results from crasis of the tribal name (Abacetini) with the Latin adjective ��� caecus���, meaning ���devoid of sight���. The name is masculine., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2020, Abacaecus n. gen. walterrossii n. sp., a notable hypogean carabid species from Sierra Leone, Western Africa (Coleoptera, Carabidae, Pterostichinae), pp. 435-446 in Zootaxa 4766 (3) on page 437, DOI: 10.11646/zootaxa.4766.3.3, http://zenodo.org/record/3765564, {"references":["Will, K. W. & Park, J. K. (2008) A new species of the Oriental Abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus. The Coleopterists Bulletin, 62 (2), 189 - 196. https: // doi. org / 10.1649 / 980.1"]}

Abacaecus walterrossii n. sp. (Figs. 1, 2, 3, 4) Type locality. Sierra Leone, Eastern Province, near Lalehun, Gola Rainforest National Park (7�� 39.595���N, 10�� 56.308���W). Type Series. Holotype ♂: Sierra Leone, Eastern Province, near Lalehun, Gola Rainforest National Park, 2.XI.2013, Walter Rossi legit (CGi). Differential diagnosis. A medium-sized (ABL: 9.56 mm) wingless abacetine with eyes strongly reduced, scarcely depigmented body, a single pair of supraorbital setae, mentum tooth with simple rounded apex, second antennomere a little eccentrically inserted on first, a scutellar setigerous pore at base of stria 2, deeply impressed elytral striae, discal setae on elytra absent, apex of each elytron forming a distinct tooth, prosternal intercoxal process lozenge shaped, dorsal face of tarsomeres smooth and with a distinct medial carina, depressed at sides, mentum tooth not exceeding level of epilobes, paramedial pits of mentum small, ostium long and wide, almost reaching basal bulb. It is easily distinguished from the unique other blind African species belonging to Pterostichinae (excluding Drimostomatina) recorded to date, Pterostillichus caecus, which is characterized by smaller size (ABL: 7.15 mm), bifid mentum tooth, second antennomere almost centrally inserted on first, absence of parascutellar setigerous pore at base of stria 2, elytral striae nearly obsolete, prosternal intercoxal process widely rounded, dorsal face of tarsomeres rugulose and without medial carina, paramedial pits of mentum large, ostium long, but narrow and not reaching basal bulb. Description. Habitus: ABL: 9.56 mm (HT ♂). Body parallel-sided, moderately shiny, concolorous reddish brown with head, antennae and tibiae a little darker (fig. 1). Female unknown. Head longer than wide, narrow in comparison with pronotum (fig. 3a); eyes strongly reduced, only a small vestigial depigmented area is visible, without any evidence of ommatidia (fig. 2a); a single pair of supraorbital setae; dorsum densely and delicately punctate; microsculpture not evident except at high magnification; frontal impressions deep, elongate divergent, ending at mid distance between clypeofrontal suture and supraorbital seta; tempora superficially wrinkled (fig. 2a); a marked supraorbital carina dilated above antennal insertion; labrum rectangular, with six setigerous punctures on anterior margin; clypeus trapezoid, rectilinear anteriorly, with two setigerous punctures closer to lateral margins than to anterior one.Antennae moderately long, densely pubescent from second fourth of segment 4, with terminal three articles surpassing base of pronotum; second antennomere a little eccentrically inserted on first; mandibles elongate, with well defined and glabrous scrobe and apex pointed and slightly hooked; mentum (fig. 2b) clearly broader than long, anterior margin shallowly emarginate; epilobes rather broad, laterally prominent; mentum tooth simple, widely rounded and not exceeding level of epilobes; mentum glabrous, except for two paramedial setae; paramedial pits distinct but small; suture between mentum and submentum present; submentum with two pairs of setae, one medial and one lateral; labial and maxillary palps fusiform; labial palpomere 2 with 2 large medial setae. Thorax: PL: 2.34 mm; PW: 2.83 mm. Pronotum densely and minutely punctate throughout, without any distinct microsculpture even at>50x magnification; form subquadrate, broader than long at middle, widest at anterior third; side margins hardly rounded at anterior half and delicately sinuate backwards; hind angles right angled and lacking of seta; posterior margin slightly concave between basal impressions, which are linear and markedly impressed (fig. 2c); anterior margin with front angles acute and markedly protruding (fig. 3a); a pair of lateral setae at anterior third; lateral marginal bead continuous and dilated before hind angles; anterior submarginal sulcus very superficial; medial longitudinal sulcus fine, ending well before apex and base. Elytra: EL: 5.37 mm; EW: 3.10 mm. Parallel-sided, moderately convex and flattened on disk, widest at base, with right-angled shoulders (fig. 2c); distinctly sinuate before apex. Surface moderately shiny; microsculpture not evident except at high magnification. Epipleura with distinct external plicae (���crossed epipleura���). Intervals moderately convex, minutely and sparsely punctate; striae deeply impressed for whole length, delicately punctate. Parascutellar stria connected to stria 1; basal section of stria 1 absent; scutellar setigerous pore present at base of stria 2 (fig. 2c). Basal margin incomplete, reaching level of stria 1. Each elytron ended apically with a distinct tooth (fig. 3b). Discal setigerous punctures absent; stria 7 with two setigerous punctures near apex; umbilicate series of 7 (humeral) + 4 (preapical) + 3 (apical) setigerous punctures in stria 8, widely interrupted between the humeral and preapical group. Hind wings absent. Ventral surface (thorax and abdomen): prosternum and proepisterna glabrous, minutely punctate. Mesosternum densely and rugosely punctate. Metepisterna short, delicately punctate; metepimera large, broadly rounded. Prosternal intercoxal process lozenge shaped, unbordered, blunt at apex (fig. 2e). Sterna shiny, glabrous except one pair subapical setae at apex of VII (HT ♂). Legs moderately slender. Dorsal face of femora with 3-4 short setae; posterior face with 1 seta in profemora, 2 in mesofemora and 0 in metafemora. Metatrochanters glabrous and slightly shorter of half length of metafemora. Protibial antennal clearing organ well developed, with 2 clip setae. Protibiae robust, slightly bent inwards at apical fifth, with a delicate longitudinal keel at inner face, a strong apical spur, 3 outer apical spines and a small apical tooth; mesotibiae with a dense group of finer setae just before middle at inner face; metatibiae longitudinally furrowed. Dorsal face of tarsomeres smooth, with a delicate medial carina, depressed at sides (fig. 2d). Protarsomeres 2-4 triangular, distinctly dilated and delicately bidentate at apex, slightly asymmetrical (HT ♂); mesotarsomeres (at a lesser extent bidentate at apex too) and metatarsomeres 2-4 less dilated and symmetrical; tarsomere 5 ventrally glabrous, dorsally with two setae; claws smooth. Male genitalia: median lobe of aedeagus with moderately enlarged basal bulb and medioapical part distally almost rectilinear in lateral view, slightly bent downwards at apex (fig. 3c); median lobe with spatulate apical lamella, almost truncate at apex in dorsal view (fig. 3d). Ostium long, reaching basal bulb. Right paramere elongate, narrowing towards apex and blunt at tip; left paramere oval, blunt at tip. Remarks. The strongly transverse mentum with anterior margin shallowly emarginate, as well as the second antennomere eccentrical (even if only moderately) with respect to first, both likely synapomorphic features for Abacetini (Will & Park, 2008), together with the presence of the parascutellar setigerous pore at base of stria 2, provide reasonable evidence to support the taxonomic arrangement of A. walterrossii n. sp. inside the tribe. In any case, the set of its morphological traits as well as its hypogean life habit, almost unique inside the tribe (see also remarks concerning Pterostillichus caecus), suggest it belongs to a separated phyletic lineage. Other features shown by A. walterrossii n. sp. occasionally appear in other species of Abacetini: for instance the apically toothed elytra (see Metabacetus vandoesburgi Straneo, 1948 from Java, in Will & Park, 2008), the carinate dorsal surface of tarsi (African species of the genus Abacetus) and the triangular and bidentate protarsomeres 2-4 (African genus Distrigidius, see Jeannel, 1948). At present knowledge, without any molecular support, it is really hard to speculate about the closest relatives of this species. The most evident adaptive trait to the hypogean life of Abacaecus is microphthalmy, but other morphological features are also consistent with this way of life, such as the elongate form of body, very different from what is seen in most Abacetini, or the markedly protruding front angles of pronotum, which are present in many other species adapted to the subterranean environment and are best interpreted as the result of convergent evolution. It is quite impressive, in this sense, the habitus resemblance of A. walterrossii n. sp. with the Iberian Pterostichini Molopina Zariquieya troglodytes Jeannel, 1924 and Z. boumortensis Faille, Fresneda & Bourdeau, 2011, Oscadytes rovirai Lagar, 1975 and Henrotius jordai (Reitter, 1914), all of them adapted to hypogean life and phylogenetically not closely related to Abacetini. Distribution and ecology. Geographical distribution: this species is recorded only from Gola Rainforest National Park, near Lalehun, Eastern Province, Sierra Leone, Africa. Life habits: the unique specimen was collected in the banks of a water stream (fig. 4) covered by lush vegetation during wet season (November); the strongly reduced eyes indicate an adaptation of this species to the hypogean environment. Etymology. This species is dedicated to its collector, Walter Rossi, a world renowned specialist in entomoparasitic fungi, as a token of our esteem and as a sign of gratitude for the gift to the authors of the specimen of the new species., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2020, Abacaecus n. gen. walterrossii n. sp., a notable hypogean carabid species from Sierra Leone, Western Africa (Coleoptera, Carabidae, Pterostichinae), pp. 435-446 in Zootaxa 4766 (3) on pages 437-442, DOI: 10.11646/zootaxa.4766.3.3, http://zenodo.org/record/3765564, {"references":["Will, K. W. & Park, J. K. (2008) A new species of the Oriental Abacetine genus Metabacetus Bates, 1892 (Coleoptera: Carabidae) and a key to the species of the genus. The Coleopterists Bulletin, 62 (2), 189 - 196. https: // doi. org / 10.1649 / 980.1","Jeannel, R. (1948) Faune de l'Empire Francais. X. Coleopteres Carabiques de la region malgache (Deuxieme partie). Office de la Recherche Scientifique Coloniale, Editions du Museum, Paris, 393 pp. [pp. 373 - 765]"]}

Pterostillichus caecus Straneo, 1949 (Figs. 5, 6) Additional description. In the original description given by Straneo (1949) some characters were ignored or, in our opinion, erroneously interpreted. Moreover, only a schematic drawing of habitus was provided by this author and therefore many important features were insufficiently illustrated, such as male genitalia. For this reason we consider as important to provide an integrative description of a few characters and their illustrations. Second antennomere almost centrically inserted on first (fig. 6a). Prosternal intercoxal process widely rounded (fig. 6e). Mentum glabrous, distincly broader than long, medial tooth with bifid apex; paramedial mental setae present; paramedial pits distinct, large (fig. 6b). Elytra with obsolete striae; lacking of a parascutellar pore at base (fig. 6c). Epipleura almost indistincly ���crossed��� before apex. Tarsomeres dorsally glabrous and delicately rugulose, without medial carina (fig. 6d). Aedeagus with median lobe distally subrectilinear in lateral view (fig. 5b); median lobe in dorsal view with a triangular apical lamella, moderately bent to right and blunt at apex (fig. 5c). Ostium long, in dorsal position, not reaching basal bulb. Remarks. The taxonomic position of P. caecus is unclear and its tribal affiliation remains problematic. It seems to belong to a phyletic lineage long separated from that of A. walterrossii n. sp. With the exception of some common characters that can be interpreted as adaptation to hypogean life and as a result of convergent evolution (microphthalmy, reduced pigmentation, markedly protruding fore angles of pronotum, presence of a single supraorbital seta, lack of seta at hind angles of pronotum), which are recurrent also in Pterostichini adapted to the subterranean environment, there are very few other characters shared between these two species. Straneo (1949) attributed P. caecus to Pterostichini sensu stricto (sensu Jeannel, 1942b) which are, as far as we know, absent in Africa (Jeannel, 1948). More probably it belongs, in our opinion, to a separated phyletic lineage within Pterostichinae, maybe related to Abacetini. The strongly transverse mentum, the angular base of stria 1 (������scutellar stria������ auct.) absent, and the morphology of aedeagus (incl. dorsal position of ostium) support the hypothesis of a relation with Abacetini; on the contrary, the second antennomere not eccentrically inserted (although some genera of Abacetini sensu lato possess the second antennomere more or less centrally inserted) and the lack of a parascutellar pore at base of elytra (even if this character is common to other pterostichine-grade taxa) contrast this hypothesis. Probably only molecular analysis will help to shed light on its taxonomic position and phylogenetic relationships., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2020, Abacaecus n. gen. walterrossii n. sp., a notable hypogean carabid species from Sierra Leone, Western Africa (Coleoptera, Carabidae, Pterostichinae), pp. 435-446 in Zootaxa 4766 (3) on pages 443-445, DOI: 10.11646/zootaxa.4766.3.3, http://zenodo.org/record/3765564, {"references":["Straneo, S. L. (1949) Due nuovi Pterostichini ciechi dell'Africa Orientale (Coleoptera: Carabidae). Bollettino della Societa Entomologica Italiana, Genova, 79 (3 - 4), 23 - 26.","Jeannel, R. (1942 b) Coleopteres Carabiques. Deuxieme partie. Faune de France 40, Paris, Librairie de la Faculte des Sciences, 1173 pp.","Jeannel, R. (1948) Faune de l'Empire Francais. X. Coleopteres Carabiques de la region malgache (Deuxieme partie). Office de la Recherche Scientifique Coloniale, Editions du Museum, Paris, 393 pp. [pp. 373 - 765]"]}

Trechisibus longipenis n. sp. (Figs. 11 a, 11b, 11c, 11d, 20) Type locality. Argentina, Neuquen, Pen. Quetrihu��. Diagnosis. A Trechisibus species 3.37���3.73 mm long, dark brown and shiny; legs, mouth parts and lateral border of elytra reddish brown; antennae slightly darkened from 3rd���4th antennomeres. Hind angles of pronotum acute and protruding backwards and outwards. Elytra oval, widest at middle, moderately depressed on disc, with 2 discal setigerous punctures (in addition to the preapical pore) on the 3rd stria, the first at about 1/6 from base and the second just before middle. T. longipenis n. sp. is distinguished from all other Trechisibus species distributed in southern Andes by the peculiar morphology of the median lobe of the aedeagus, which is extraordinarily long and slender (Fig. 11 b). Type series. HT ♂, Argentina, Neuquen, Pen. Quetrihu��, 12.II.1998, legit Riccardo Sciaky (CSc). PTT: 2 ♂♂ 1 ♀, same data as the holotype (CAl, CGi, CSc). Description. Habitus as in Fig. 11 a. Overall length of the HT ♂ (from labrum to apex of elytra) 3.68 mm (PTT ♂♂ 3.37���3.73, ♀ 3.68 mm). Body dark brown, obovate, shiny. Microsculpture markedly impressed on head, very superficial on pronotum and nearly absent on elytra. Legs, mouth parts, and lateral border of elytra reddish brown; antennae slightly darkened from 3rd���4th antennomeres. Brachypterous. Head moderately large, eyes small and convex; temples convex and delicately pubescent, hardly longer than eyes. Microsculpture in isodiametric meshes markedly impressed on whole upper surface of head. Clypeus with two apical setae on each side; labrum transverse, 6-setose, distinctly excavate at apex. Frons between eyes convex, depressed on vertex. Antennae moderately long, hardly reaching the basal 4th of elytra. Pronotum convex on disc, transverse (width/length = 1.31), with base as wide as anterior margin (Fig. 20). Microsculpture very superficial, nearly absent on disc and more visible near base and fore margin. Sides rounded for most of length, shortly sinuate before hind angles; basal margin sinuate at sides. Front angles feebly prominent; hind angles acute and protruding backwards and outwards. Mid longitudinal line distinctly impressed between the submarginal sulci, deepened towards base. A wide basal impression on each side, separated at middle by the wrinkled mid convexity of the base. Lateral border moderately wide on whole length; anterior margin unbordered, the posterior bordered only at sides. Two lateral setae on each side, one at hind angles (moved a little forwards from the angle) and one at about 2/3 from base. Elytra oval (length/width=1.35), widest at middle, feebly depressed on disc. Microsculpture in transverse meshes, very superficial, nearly absent on disc and hardly visible near base and apex. Shoulders rounded with humeral angle indistinct. The basal margin interrupted in correspondence with the 5th interval. A short scutellar stria is present, as well as a juxtascutellar pore near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border wider in the basal half, reddish brown and distinctly contrasting in colour with the darker elytra. Two discal setigerous punctures in small and superficial foveae on the 3rd stria, the 1st at basal 6th of elytra and the 2nd just before middle; the punctures divert the 3rd stria linearity; preapical pore at about middle of the recurrent striole. Umbilicate series of 4+2+2 punctures, with groups widely spaced from each other. The five inner striae distinctly impressed, the others obliterated; the 8th stria deeply impressed in the posterior half. Intervals nearly flat. Elytral apex broad and rounded. The recurrent striola moderately long and arcuate. Legs slender. Protibiae nearly straight and externally distinctly furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus extraordinarily slender, in lateral view tapered and bent upward towards apex (Fig. 11 b); apical blade long and, in dorsal view, spoon shaped and rounded at extreme apex (Fig. 11 c). The internal armature is made by a copulatory piece long and thin, wides at base (Figs. 11 b, 11c). Parameres long and slender, almost equally long, each provided with 4 apical setae (Fig. 11 b). Etymology. The specific epithet refers to the unusual slenderness of the aedeagus. Distribution and habitat. At present Trechisibus longipenis n. sp. is only recorded from the type locality. The specimens of the type series were collected at about 800 m a.s.l. Relationships. At the present state of knowledge, the phylogenetic affinities of Trechisibus brevis n. sp. are unclear, but despite the strong differences in the morphology of aedeagus, this species shares many external morphological features with Trechisibus striatus n. sp., Trechisibus sciakyi n. sp. and Trechisibus brevis n. sp., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 66-68, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Trechisibus species from Peru The following new species are described., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on page 50, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Trechisibus striatus n. sp. (Figs. 8 a, 8b, 8c, 8d, 17) Type locality. Argentina, Neuquen, Pen. Quetrihu��. Diagnosis. A Trechisibus species 3.46���3.74 mm long, dark brown and shiny; legs, antennae, mouth parts and lateral border of elytra reddish brown. Hind angles of pronotum usually acute and protruding outwards. Elytra oval, slightly convex on disc, with 2 discal setigerous punctures (in addition to the preapical pore) on the 3rd stria, the first at about 1/4 from base and the second just behind middle. T. striatus n. sp. is distinguished from all other Trechisibus species distributed in Patagonian Argentina by the deeply impressed 1st���3rd elytral striae (Fig. 8 a) and by the peculiar morphology of the median lobe of aedeagus as well as of the copulatory piece (Fig. 8 b). Type Series. HT ♂, Argentina, Neuquen, Pen. Quetrihu��, 12.II.1998, legit Riccardo Sciaky (CSc). PTT: 4 ♀♀, same data as the holotype (CAl, CGi, CSc). Description. Habitus as in Fig. 8 a. Overall length of the HT ♂ (from labrum to apex of elytra) 3.74 mm. Body dark brown, obovate, shiny. Microsculpture very superficially impressed on pronotum and elytra in both sexes, markedly impressed on head. Antennae, mouth parts, legs and lateral border of elytra reddish brown. Brachypterous. Head moderately large, eyes large and convex; temples slightly convex and delicately pubescent, hardly shorter than eyes. A microsculpture in isodiametric meshes markedly impressed on whole upper surface of head. Clypeus with two apical setae on each side; labrum transverse, 6-setose, distinctly excavate at apex. Frons between eyes convex with a small depression on vertex. Antennae moderately long, reaching the basal 4th of elytra. Pronotum convex on disc, transverse (width/length = 1.46), with base slightly wider than anterior margin (Fig. 17). Disc almost shiny and microsculpture nearly absent except at sides and near fore margin. Sides rounded on whole length; basal margin almost rectilinear, slightly sinuate at sides. Front angles hardly prominent; hind angles usually acute and protruding outwards (seldom nearly right angles). Mid longitudinal line distinctly impressed, deepened and enlarged near base. A wide basal impression on each side, separated at middle by a wrinkled convexity of the base and by the enlarged mid longitudinal line. Lateral border narrow and only slightly broadened towards base; anterior and posterior margins bordered only at sides. Two lateral setae on each side, one at hind angles (moved a little forwards from the angle) and one at about 2/3 from base. Elytra oval (length/width=1.33), fairly convex and not depressed on disc. Microsculpture in transverse meshes, very superficial in both sexes and hardly visible only near base and at sides. Shoulders with humeral angle distinct. The basal margin interrupted in correspondence with the 5th stria. A short scutellar stria is present, as well as a juxtascutellar pore near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border narrow over the whole length, reddish brown and distinctly contrasting in colour with the darker elytra. Two discal setigerous punctures in small foveae on the 3rd stria, the 1st at basal 4th of elytra and the 2nd just behind middle; the punctures usually divert the 3rd stria linearity; preapical pore at about middle of the recurrent striole. Umbilicate series of 4+2+2 punctures, with a wide distance between the 1st and the 2nd group. 1st���3rd striae (and sometimes 4th) deeply impressed but vanishing at base and at apex, the 5th���7th very superficial and hardly visible, the 8th deeply impressed in the posterior half. Inner intervals feebly convex, the others flat. Elytral apex broad and rounded. The recurrent striola moderately long and arcuate. Legs slender. Protibiae straight and externally distinctly furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus moderately slender, in lateral view the inferior edge delicately sinuate towards apex (Fig. 8 b), which is short and, in dorsal view, widely blunt (Fig. 8 c). The internal armature consists of a cylindrical copulatory piece anteriorly provided with a longitudinal bundle of sclerotized teeth (Fig. 8 c). Parameres wide and short, almost equally long, each provided with 4 apical setae (Fig. 8 b). Etymology. The specific epithet refers to the deeply impressed inner elytral striae. Distribution and habitat. At present Trechisibus striatus n. sp. is only recorded from the type locality. The specimens of the type series were collected at about 800 meters a.s.l. Relationships. At the present state of knowledge, the phylogenetic affinities of Trechisibus striatus n. sp. are unclear. Despite the strong differences in the morphology of aedeagus, this species shows many similarities in the external morphology with Trechisibus sciakyi n. sp., Trechisibus brevis n. sp. and Trechisibus longipenis n. sp., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 61-63, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Trechisibus recuayi n. sp. (Figs. 3, 4 a, 4b, 4c, 4d, 14) Type locality. Peru, Dip. Ancash, Punta Olímpica 4,550 m, laguna. Diagnosis. A Trechisibus species 2.98–3.24 mm long, dark brown, moderately shiny; elytra with apex, marginal border and sutural interval lighter brown; legs, antennae and mouth parts reddish-brown. Hind angles of pronotum nearly right or scarcely obtuse. Elytra oval, not depressed on disc, with one setigerous puncture on 3rd stria at about 1/3 from base. Due to the peculiar median lobe of aedeagus with hook shaped apex, T. recuayi n. sp. may be confused with two other species living in Cordillera Blanca (Trechisibus lamasi Etonti & Mateu, 1992 and Trechisibus decensii Allegro, Giachino & Sciaky, 2008), which also share a very similar external morphology, but it the can be easily distinguished by the smaller size and by a shorter and more stumpy median lobe of aedeagus, with differently shaped copulatory piece (Fig. 5). Type Series. HT ♂, Peru, Dip. Ancash, Punta Olímpica, m 4,550, 5.VII.2008, legit G. Allegro (CAl). PTT: 1 ♀, same data as the holotype (CGi); 1 ♀, Peru, Dip. Ancash, Punta Olímpica, m 4,550, 6.XII.2005, legit G. Allegro (CAl). Description. Habitus as in Fig. 4 a. Overall length of the HT ♂ (from labrum to apex of elytra) 3.24 mm (PTT ♀♀ 2.98–3.17 mm). Body dark brown, obovate, moderately shiny. Microsculpture in transverse meshes superficially impressed on elytra and on disc of pronotum, more visible near base and apex of pronotum and on head. Antennae, tibiae, clypeus and labrum reddish, femora a little darker; palpi yellowish. Brachypterous. Head moderately large, eyes small and scarcely convex; temples oblique and delicately pubescent, as long as eyes. Microsculpture markedly impressed over all surface but even stronger (almost wrinkled) near clypeus and towards eyes, laterally to frontal impressions, which are arcuate and deep. Clypeus with two apical setae on each side, the inner one shorter and thinner; labrum transverse, 6-setose, excavate at apex. Frons between eyes convex. Antennae slender, reaching the basal fifth of elytra. Pronotum moderately convex on disc, transverse (width/length = 1.38), with base wider than anterior margin (Fig. 14). Disc shiny with very superficial microsculpture, which is distinct only near base and near front margin. Sides rounded in the fore half and subrectilinear in the basal half, shortly sinuate before base; basal margin excavate at middle and oblique near hind angles. Front angles scarcely prominent; hind angles marked, right or nearly obtuse. Mid longitudinal line distinctly impressed between the submarginal sulci. Lateral margins wide and broadened towards base; anterior and posterior margins bordered only at sides. Two lateral setae on each side, one at hind angles and one about 3/4 from base. Elytra oval (length/width=1.40), fairly convex and not depressed on disc. Microsculpture in transverse meshes, superficial in both sexes. Shoulders scarcely angulate with humeral angle distinct. The basal margin is interrupted in correspondence with the 6th interval. Scutellar stria absent. A juxtascutellar pore is present near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border narrow. One setigerous puncture on the 3rd stria at about 1/3 from base; preapical pore at about middle of the recurrent striole. Umbilicate series of 4+2+2 punctures, with the groups widely distanced from each other. 1st–5th striae very superficial and hardly visible, the others obliterated. Intervals flat. Elytral apex broad and rounded. The recurrent striola short and subrectilinear, without a remarkable apical carina. Legs slender. Protibiae nearly straight and externally superficially furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus short and stout (Fig. 4 b), in lateral view the inferior edge delicately bent downward before apex, which is short and, in dorsal view, sinuate at sides and blunt (Fig. 4 c). The internal armature is made by a robust copulatory piece with a dense cover of sclerotized teeth (Fig. 4 b). Parameres narrow and long, almost equally long and longer than half of the median lobe, each provided with 3 apical setae (Fig. 4 b). Etymology. Species name derived from the noun, in the genitive case, of the pre-columbian culture Recuay, that flourished in the Andean area of Cordillera Blanca from 200 BC to 600 AC. Distribution and habitat. At present Trechisibus recuayi n. sp. is only recorded from the type locality. The three specimens of the type series were collected under stones laying on humid soil near a lake (about 4,550 meters a.s.l.) below the Punta Olímpica Pass, in a glacial area characterized by coarse detritic sediments (Fig. 3). It lives in syntopy with Trechisibus huascarani n. sp. and Blennidus (Agraphoderus) huascarani Allegro, 2010, which are recorded from the same type locality. Relationships. Trechisibus recuayi n. sp. is probably a close relative of Trechisibus lamasi Etonti & Mateu, 1992 and Trechisibus decensii Allegro, Giachino & Sciaky, 2008, according to similarities in external morphology and in male genitalia (in particular all them have a hooked apex of the median lobe of aedeagus) (Fig. 5). These species live in Cordillera Blanca, all inhabiting humid soils. Due to the marked similarities in male genitalia, they can be included in the ‘ T. lamasi ’ species group.

Trechisibus parvulus n. sp. (Figs. 12 a, 12b, 12c, 12d, 21) Type locality. Argentina, Neuquen, Pen. Quetrihu��. Diagnosis. A Trechisibus species 2.53���2.78 mm long, brown or reddish brown, shiny, with pronotum paler than elytra; legs, antennae, mouth parts and lateral border of elytra yellowish. Hind angles of pronotum obtuse. Elytra oval, widest at middle, moderately depressed on disc, with 2 discal setigerous punctures (in addition to the preapical pore) on the 3rd stria, the first at about 1/3 from base and the second just behind middle. T. parvulus n. sp. is distinguished from all other Trechisibus species distributed in the Patagonian Andes by its small size and by the peculiar morphology of the median lobe of aedeagus. Type Series. HT ♂, Argentina, Neuquen, Pen. Quetrihu��, 12.II.1998, legit Riccardo Sciaky (CSc). PTT: 6 ♂♂, same data as the holotype (CAl, CGi, CSc). Description. Habitus as in Fig. 12 a. Overall length of the HT ♂ (from labrum to apex of elytra) 2.67 mm (PTT ♂♂ 2.53���2.78). Body brown or reddish brown with pronotum paler than elytra, obovate, shiny. Microsculpture distinctly impressed only on fore half of head, nearly absent on pronotum and elytra. Legs, antennae, mouth parts and lateral border of elytra yellowish. Brachypterous. Head moderately large, eyes small and convex; temples convex and delicately pubescent, hardly longer than eyes. A microsculpture in isodiametric meshes impressed on fore half of head, on vertex and neck less distinct. Clypeus with two apical setae on each side; labrum transverse, 6-setose, distinctly excavate at apex. Frons between eyes convex depressed on vertex. Antennae relatively short, hardly reaching the basal 5th of elytra. Pronotum convex on disc, transverse (width/length = 1.34), with base much narrower than anterior margin (Fig. 21). Microsculpture very superficial, nearly absent on disc and hardly visible near base and fore margin. Sides rounded for most of length, shortly sinuate before hind angles; basal margin almost rectilinear. Front angles feebly prominent; hind angles obtuse. Mid longitudinal line distinctly impressed between the submarginal sulci, deepened towards base. A wide basal impression on each side, separated at middle by the superficially wrinkled convexity of the base. Lateral border moderately wide on whole length; anterior and posterior margins unbordered. Two lateral setae on each side, one at hind angles (moved distinctly forwards from the angle) and one at about 2/3 from base. Elytra oval (length/width=1.35), widest at middle, feebly depressed on disc. Microsculpture absent on disc and hardly visible at base and apex. Shoulders rounded with humeral angle indistinct. The basal margin interrupted in correspondence with the 5th interval. A short scutellar stria is present, as well as a juxtascutellar pore near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border moderately wide on whole length, reddish brown and distinctly contrasting in colour with the darker elytra. Two discal setigerous punctures in small and superficial foveae on the 3rd stria, the 1st at basal 3rd of elytra and the 2nd just behind middle; the punctures divert the 3rd stria linearity; preapical pore at about middle of the recurrent striole. Umbilicate series of 4+2+2 punctures, with groups widely spaced from each other. The four inner striae distinctly impressed only on disc, obliterated towards base and apex as well as all the other striae; the 8th stria deeply impressed in the posterior half. Intervals nearly flat. Elytral apex broad and rounded. The recurrent striola moderately long and arcuate. Legs relatively short. Protibiae nearly straight and externally superficially furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus tiny, ventrally rectilinear in lateral view and tapered towards apex (Fig. 12 b); apical blade short and, in dorsal view, rounded at apex (Fig. 12 c). A simple copulatory piece, short and triangular in dorsal view (Figs. 12 c). Parameres short, the right one shorter than the left, each provided with 4 apical setae (Fig. 12 b). Etymology. The specific epithet refers to the small size of this species. Distribution and habitat. At present Trechisibus parvulus n. sp. is only recorded from the type locality. The specimens of the type series were collected at about 800 m a.s.l. Relationships. At the present state of knowledge, the phylogenetic affinities of Trechisibus parvulus n. sp. are unclear, but due to the marked differences in the morphology of the habitus and aedeagus, this species probably belongs to a different lineage with respect to the other species collected at the same locality and here described, namely Trechisibus striatus n. sp., Trechisibus sciakyi n. sp., Trechisibus brevis n. sp. and Trechisibus longipenis n. sp. The Trechisibus species so far recorded from Argentina, with their respective regional distribution, are, Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 68-70, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Trechisibus curtii n. sp. (Figs. 6 a, 6b, 6c, 15) Type locality. Peru, Hu��nuco, Quebrada Queulacocha, 3,800 m. Diagnosis. A Trechisibus species 3.59 mm long, piceous black, moderately shiny; legs, antennae and mouth parts brown. Hind angles of pronotum acute and protruding outwards. Elytra oval, not depressed on disc, with four discal setigerous punctures (in addition to the preapical pore), the 1st���3rd on the 3rd stria and the 4th in the 3th interval, more or less regularly spaced; moreover, a single setigerous puncture is also present on the 5th stria, at about 1/4 from base (moved a little forwards with respect to the 2nd puncture of the 3rd stria). T. curtii n. sp. is easily distinguished from all other already known Trechisibus species by its peculiar elytral chaetotaxy: by comparison, the only other species showing a single setigerous pore on the 5th stria is Trechisibus hornensis (Fairmaire, 1885), but it only shows 2 discal pores on the 3rd. Type Series. HT ♂, Peru, Dept. Hu��nuco, Quebrada Queulacocha, m 3,800, VI.1981, legit M. Curti (CAl). Description. Habitus as in Fig. 6 a. Overall length of the HT ♂ (from labrum to apex of elytra) 3.59 mm. Body piceous black, obovate, moderately shiny. Microsculpture in transverse meshes superficially impressed on elytra, more distinct near base of pronotum and on head; disc of pronotum smooth and shiny. Antennae, legs, clypeus and labrum dark brown, tarsi and palpi lighter. Brachypterous. Head large with respect to pronotum, eyes large and convex; temples convex and delicately pubescent, a little shorter than eyes. Microsculpture superficially impressed on vertex, but very marked (almost wrinkled) towards eyes, laterally to frontal impressions, which are arcuate and deep. Clypeus with two apical setae in a deep depression on each side; labrum transverse, 6-setose, distinctly excavate at apex. Frons between eyes convex. Antennae short, hardly reaching the base of elytra. Pronotum moderately convex on disc, transverse (width/length = 1.34), with base hardly wider than anterior margin (Fig. 15). Disc shiny without any apparent microsculpture, which is distinct only near base. Sides rounded in the fore half and subrectilinear in the basal half, shortly but markedly sinuate before base; basal margin rectilinear. Front angles not prominent; hind angles acute and protruding outwards. Mid longitudinal line distinctly impressed between the submarginal sulci. Lateral margins wide and broadened towards base; anterior and posterior margins bordered only at sides. Two lateral setae on each side, one at hind angles and one at about 3/4 from base. Elytra oval (length/width=1.41), fairly convex and not depressed on disc. Microsculpture in transverse meshes, superficial in both sexes. Shoulders rounded with humeral angle indistinct. The basal margin interrupted in correspondence with the 5th interval. A short scutellar stria is present, as well as a juxtascutellar pore near base, in correspondence with the junction of the 1st and 2nd stria. Sides moderately rounded; lateral border narrow but broader at base. Four setigerous punctures in small foveae, the 1st���3rd on the 3rd stria and the 4th in the 3th interval, more or less regularly spaced; a single setigerous puncture is also present on the 5th stria, at about 1/4 from base (moved slightly forwards with respect to the 2nd puncture of the 3rd stria); preapical pore at about middle of the recurrent striole. Umbilicate series of 3+2+2 punctures, with the groups widely distanced from each other. Striae 1st���4th very superficial and hardly visible, the others obliterated. Intervals flat. Elytral apex broad and rounded. The recurrent striola long and arcuate, with a remarkable apical carina. Legs short. Protibiae delicately arcuate towards apex and externally superficially furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus slender, obliquely inserted on the basal bulb (Fig. 6 b), in lateral view the inferior edge nearly straight; in dorsal view, the median lobe is slender and parallel-sided, with apex short and broadly rounded (Fig. 6 c). The internal armature is made by a hardly sclerotized copulatory piece (Fig. 6 b). Parameres narrow and long, the left one longer than the right and as long as half of the median lobe, each provided with 3���4 apical setae (Fig. 6 b). Etymology. This species is dedicated to its collector, Marc Curti, a talented French amateur entomologist, as a token of our esteem. Distribution and habitat. At present Trechisibus curtii n. sp. is only recorded from the type locality. The unique specimen of the type series was collected at 3,800 meters a.s.l., presumably in an Andean grassland habitat. Relationships. Trechisibus curtii n. sp. does not show evident affinities with any other known Trechisibus species, and therefore is not yet included in any species group., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 56-58, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Trechisibus huascarani n. sp. (Figs. 1, 2 a, 2b, 2c, 2d, 3, 13) Type locality. Peru, Dip. Ancash, Punta Ol��mpica 4,550 m, laguna. Diagnosis. A Trechisibus species 3.92���4.37 mm long, moderately shiny, brown with head a little darker; legs and antennae reddish-brown. Hind angles of pronotum obtuse. Elytra oval-elongate, depressed on disc, with 4���5 setae on the 3rd stria (in addition to the preapical seta) and 2���3 on 5th. It can easily be distinguished from T. amesi Etonti & Mateu, 1992, a very similar species recorded from the Cordillera Blanca with more than three discal setae on the 3rd elytral stria, by the presence of 2���3 setae on the 5th stria as well (absent in T. amesi) and by the shape of the median lobe of the aedeagus, which is not inferiorly bulging before apex in lateral view (distinctly bulging in T. amesi). Trechisibus curtii n. sp., a species recorded from Hu��nuco also showing more than three discal setae on the 3rd elytral stria, is very different from T. huascarani n. sp. in external morphology and morphology of the aedeagus. Type Series. HT ♂, Peru, Dip. Ancash, Punta Ol��mpica, m 4,550, 5.VII.2008, legit G. Allegro (CAl). PTT: 8 ♂♂ 5 ♀♀, same data as the holotype (CAl, CCa, CGi, CSc, BMNH). Description. Habitus as in Figs. 1 and 2 a. Overall length of the HT ♂ (from labrum to apex of elytra) 4.03 mm (PTT ♂♂ 3.92���4.37, ♀♀ 4.02���4.30 mm). Elytra brown, head black-brown and pronotum reddish-brown, moderately shiny. Microsculpture in transverse meshes superficially impressed on elytra, on pronotum visible only near hind angles. Antennae, tibiae and mouthparts reddish, femora a little darker. Brachypterous. Head large and stout, eyes small and moderately convex; temples convex and delicately pubescent, as long as eyes. Clypeus with two apical setae on each side, the inner one shorter and thinner; labrum transverse, 6-setose, excavate at apex. Frontal impressions arcuate and deeply impressed. Frons between eyes convex, smooth and shiny. Antennae long, reaching the basal fourth of elytra. Pronotum moderately convex on disc and explanate near hind angles, transverse (width/length = 1.41), with base as wide as the anterior margin (Fig. 13). Disc smooth and shiny, the microsculpture distinct only near hind angles, in the explanate area from the basal impression, which is wide and impunctate, to the lateral margin. Sides rounded in the fore half and linear in the basal half, with maximum width at about 3/4 from base; base subrectilinear. Front angles not prominent; hind angles marked and obtuse. Mid longitudinal line distinctly impressed between the submarginal sulci. Lateral margins wide and broadened towards base; anterior and posterior margins bordered only at sides. Two lateral setae on each side, one at hind angles and one about 4/5 from base. Elytra oval (length/width=1.40), fairly convex but depressed on disc. Microsculpture in transverse meshes, superficial in both sexes. Shoulders broadly rounded. The basal margin interrupted in correspondence with the 5th interval. Scutellar stria absent. A juxtascutellar pore is present near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border narrow. 4���5 setigerous punctures on the 3rd stria in addition to the preapical puncture, the 1st at basal 6th, the others more or less regularly spaced; 2���3 setigerous punctures also on the 5th stria, usually in the basal half, but sometimes the 2nd or 3rd just behind middle. Umbilicate series of 4+2+2 punctures, with the groups widely distanced from each other. Only striae 1st���5th superficially impressed and obliterated at base and towards apex (the 1st more impressed than 2nd���5th), the others hardly visible. Intervals flat, the 1st narrower than the followings. Elytral apex rounded, with sutural angles of elytra divergent. The recurrent stria long and distinctly impressed, with a remarkable apical carina. Legs slender. Protibiae nearly straight and not externally furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus stout (Fig. 2 b), in lateral view the inferior edge rectilinear towards apex, which is thin, short and, in dorsal view, broadly rounded (Fig. 2 c). The internal armature is peculiar as it is made by two overlapping sclerotized pieces, one in upper and one in lower position, long and sinuous, joining at apex (Fig. 2 b). Parameres narrow and long, the left one longer than the right and as long as half of the median lobe, each provided with 5���6 apical setae (Fig. 2 d). Etymology. Species name derived from the noun, in the genitive case, of Mt. Huascar��n, the highest mountain in Peru (6,768 m), dominating the Huascar��n National Park which includes the type locality. Distribution and habitat. At present Trechisibus huascarani n. sp. is only recorded from the type locality. All specimens of the type series were collected under stones laying on humid soil near a lake (about 4,550 meters a.s.l.) below the Punta Ol��mpica Pass, in a glacial area characterized by coarse detritic sediments (Fig. 3). It lives in syntopy with Trechisibus recuayi n. sp. and Blennidus (Agraphoderus) huascarani Allegro, 2010, which are described from the same type locality. Relationships. Trechisibus huascarani n. sp. is actually very similar to T. amesi, both in external morphology and in male genitalia (in particular in the copulatory piece), although these species can be easily distinguished by the characters mentioned here above in the diagnosis. The subgenus Trechisibiontes Etonti & Mateu, 1992, created for T. amesi on account of the presence of four or more setae on the third elytral stria, was recently synonymized with Trechisibiellus (Avon 2007). Both species live in Cordillera Blanca, and they also share the habitat of high altitude glacial areas, as also T. amesi is recorded from the vicinities of a glacier (Pastoruri) at 4,850���4,950 meters a.s.l. Mainly due to the marked similarities in male genitalia, T. amesi and T. huascarani n. sp. probably deserve a their own species group. Despite the presence of additional discal setae on the 5th elytral stria, which are also present in T. hornensis (Fairmaire, 1855) and T. curtii n. sp., T. huascarani n. sp. does not seem closely related to these species on account of marked differences in external morphology as well as morphology of the aedeagus., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 50-54, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618, {"references":["Etonti, M. & Mateu, J. (1992) Sette nuovi Trechinae del Peru (Coleoptera, Carabidae). Bollettino del Museo civico di Storia naturale di Venezia, 41, 109 - 130.","Allegro, G. (2010) A peculiar new species of Blennidus Motschulsky subgenus Agraphoderus Bates from the Andes of the Cordillera Blanca (Peru) (Coleoptera: Carabidae: Pterostichini). Studies and Reports, Taxonomical Series, 6 (1 - 2), 1 - 7.","Avon, C. (2007) Nouvelle monographie des Trechinae (Coleoptera Carabidae) 4. Tribu des Homaloderini (2 eme serie), Genre Trechisibus. Les Comptes-Rendus du Laboratoire d'Entomologie Faune Hypogee et Endogee, Nice, 367 - 600."]}

Trechisibus brevis n. sp. (Figs. 10 a, 10b, 10c, 10d, 19) Type locality. Argentina, Neuquen, Pen. Quetrihu��. Diagnosis. A Trechisibus species 3.42 mm long, dark brown and shiny; legs, antennae, mouth parts and lateral border of elytra reddish brown. Hind angles of pronotum acute and protruding backwards and outwards. Elytra short oval, obovate, widest behind middle, moderately depressed on disc, with 2 discal setigerous punctures (in addition to the preapical pore) on the 3rd stria, the first at about 1/5 from base and the second just behind middle. T. brevis n. sp. is distinguished from all other Trechisibus species distributed in Patagonia by the body short and obovate (Fig. 10 a) and by the peculiar morphology of the median lobe of aedeagus, provided with a distinct ventral swelling in lateral view (Fig. 10 b). Type Series. HT ♂, Argentina, Neuquen, Pen. Quetrihu��, 12.II.1998, legit Riccardo Sciaky (CAl). Description. Habitus as in Fig. 10 a. Overall length of the HT ♂ (from labrum to apex of elytra) 3.42 mm. Body dark brown, obovate, shiny. Microsculpture markedly impressed on head, very superficial on pronotum and nearly absent on elytra. Legs, mouth parts, antennae and lateral border of elytra reddish brown. Brachypterous. Head moderately large, eyes small and convex; temples hardly convex and delicately pubescent, as long as eyes. A microsculpture in isodiametric meshes markedly impressed on whole upper surface of head. Clypeus with two apical setae on each side; labrum transverse, 6-setose, distinctly excavate at apex. Frons between eyes convex without depression on vertex. Antennae moderately long, hardly reaching the basal 4th of elytra. Pronotum convex on disc, transverse (width/length = 1.36), with base as wide as the anterior margin (Fig. 19). Disc almost shiny and microsculpture very superficial but visible on the whole pronotum. Sides rounded for most of their length, shortly sinuate before hind angles; basal margin sinuate at sides. Front angles feebly prominent; hind angles acute and protruding backwards and outwards. Mid longitudinal line distinctly impressed between the submarginal sulci, deepened towards base. A wide basal impression on each side, separated at middle by the wrinkled convexity of the base. Lateral border narrow on whole length; anterior margin unbordered, the posterior bordered only at sides. Two lateral setae on each side, one at hind angles (moved a little forwards from the angle) and one at about 2/3 from base. Elytra short oval (length/width=1.27), widest behind middle, feebly depressed on disc. Microsculpture in transverse meshes very superficial, nearly absent on disc. Shoulders rounded with humeral angle indistinct. The basal margin interrupted in correspondence with the 5th stria. A short scutellar stria is present, as well as a juxtascutellar pore near base, in correspondence with the 2nd stria. Sides moderately rounded; lateral border narrow in the fore half, posteriorly widened, reddish brown and distinctly contrasting in colour with the darker elytra. Two discal setigerous punctures in small and superficial foveae on the 3rd stria, the 1st at basal 5th of elytra and the 2nd just behind middle; the punctures divert the 3rd stria linearity; preapical pore at about middle of the recurrent striole. Umbilicate series of 4+2+2 punctures, with groups widely spaced from each other. The five inner striae moderately impressed, the others obliterated; the 8th stria deeply impressed in the posterior half. Intervals nearly flat. Elytral apex broad and rounded. The recurrent striola moderately long and arcuate. Legs slender. Protibiae nearly straight and externally distinctly furrowed. Tarsi pubescent on the upper side. The 1st and 2nd male protarsomeres asymmetrically dilated. Median lobe of aedeagus slender in lateral view, provided with a distinct ventral swelling and hardly bent downward at apex (Fig. 10 b); apical blade short and, in dorsal view, widely triangular, blunt at the extreme apex (Fig. 10 c). Little can be said about the internal armature, as the only known specimen (HT) shows it evaginate; a bundle of sclerotized scales is present (Figs. 10 b, 10c). Parameres wide and short, almost equally long, each provided with 4 apical setae (Fig. 10 b). Etymology. The specific epithet refers to the short oval shape of the body. Distribution and habitat. At present Trechisibus brevis n. sp. is only recorded from the type locality. The specimens of the type series were collected at about 800 m a.s.l. Relationships. At the present state of knowledge, the phylogenetic affinities of Trechisibus brevis n. sp. are unclear, but despite the strong differences in the morphology of the aedeagus, this species shares many external morphological features with Trechisibus striatus n. sp., Trechisibus sciakyi n. sp. and Trechisibus longipenis n. sp., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2016, Nine new Trechisibus species from Peru and Argentina (Coleoptera: Carabidae: Trechinae), pp. 49-72 in Zootaxa 4193 (1) on pages 65-66, DOI: 10.11646/zootaxa.4193.1.2, http://zenodo.org/record/166618

Blennidus (Agraphoderus) straneoi Allegro & Giachino, 2011 (Figs. 43, 81, 117) Blennidus straneoi Allegro & Giachino, 2011 a: 303 Type locality. Peru, Callanga. Examined material. HT ♂ (CSt at MSNM); 1 PT ♀ (CAl) (complete TS). Note. Species belonging to the ��� B. jelskii species group��� together with four other species (B. jelskii, B. abramalagae, B. etontii, B. procerus) distinguished by the distinctive morphology of aedeagus (Allegro & Giachino 2011 a). Distribution. The type locality probably does not refer to the town near Lima but to the ancient archaeological site in the Cuzco Dept. (southern Peru) (Allegro & Giachino 2011 a); in analogy with the species included in the same species group, Blennidus straneoi is probably restricted to the Andes of southern Peru., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2015, Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae), pp. 1-48 in Zootaxa 4000 (1) on page 16, DOI: 10.11646/zootaxa.4000.1.1, http://zenodo.org/record/289219, {"references":["Allegro, G. & Giachino, P. M. (2011 a) Studies on the subgenus Agraphoderus Bates of Blennidus Motschulsky from Peru: the jelskii species-group (Coleoptera, Carabidae, Pterostichini). Animal Biodiversity and Conservation, 34 (2), 295 - 308."]}

Key to the species of Blennidus subgenus Agraphoderus recorded from Peru (To cope with a possible misinterpretation at couplet 27, Blennidus anxius is keyed twice) 1. Pronotum flattened on disc and markedly restricted at base, with sides rectilinear or slightly sinuate in posterior third; hind angles of pronotum obtuse, with blunt apex (Fig. 62). Elytra subdepressed. Aedeagus curved in dorsal view and with apical blade sharply pointed (Fig. 98). Dept. Ancash, Cordillera blanca. 8.3–10.3 mm...... Blennidus huascarani Allegro, 2010. - Species dorsally convex. The above characters are not all present together....................................... 2. 2. Male mesotibiae with an inner spine or an acutely pointed swelling in subapical position........................... 3. - Male mesotibie without an inner subapical spine, at most with an obtuse swelling................................. 5. 3. Male metatibiae with a marked subapical swelling. Hind angles of pronotum widely rounded (Fig. 54). Male dorsally with strong metallic lustre. Dept. Cajamarca. 9.5–9.8 mm.............................. Blennidus bellesi (Straneo, 1993) - Male metatibiae subapically not swollen. Hind angles of pronotum obtuse but more or less indicated.................. 4. 4. Hind angles of pronotum with pointed vertex (Fig. 68). Elytra subparallel sided or slightly dilated backwards, with rounded apex. The subapical inner spine of male mesotibiae short. Aedeagus short and markedly curved downwards at apex in lateral view (Fig. 104). Dept. Cuzco. 8.5–9.5 mm.................................. Blennidus mesotibialis (Straneo, 1993) - Hind angles of pronotum with blunt vertex (Fig. 69). Elytra oval, with pointed apex. The subapical inner spine of male mesotibia long and acuminate. Aedeagus slender, scarcely curved downwards at apex in lateral view (Fig. 105). Dept. Cajamarca. 9.5 mm............................................................... Blennidus negrei (Straneo, 1993) 5. Pronotum usually disc-shaped, with hind angles more or less rounded (Figs. 56, 72, 76, 78). Species distributed in northern Peru (Dept. Cajamarca)............................................................................... 6. - Pronotum of different shape, or species distributed in central or southern Peru.................................... 9. 6. The 1 st to 5 th elytral striae all equally impressed, with moderately convex intervals. Hind angles of pronotum rounded or hardly evident (Fig. 76). Larger size (9.2–10.9 mm).......................... Blennidus pinguis (Allegro & Giachino, 2011) - The 1 st elytral stria more impressed than 2 nd to 5 th, which are more or less obsolete, with flat intervals. Hind angles of pronotum rounded or widely obtuse. Smaller size ( 7. Pronotum with base as wide as anterior margin, with sides strongly convex and hind angles rounded (Fig. 56). 9.0 mm. Dept. Cajamarca............................................................... Blennidus crassus (Straneo, 1993) - Pronotum with base definitely narrower than anterior margin, with sides less convex and hind angles widely obtuse but evident (Figs. 72, 78)........................................................................................ 8. 8. Larger size (8.5 –9.0 mm). Umbilicate pores at sides of elytra widely interrupted at middle. Sides of pronotum linear or slightly convex just before hind angles (Fig. 72). Sterna IV–VI with a series of evident punctures at sides. Aedeagus in dorsal view markedly arcuate (Fig. 108)................................................ Blennidus orbicollis (Straneo, 1993) - Smaller size (7.5 –8.0 mm). Umbilicate pores at sides of elytra not widely interrupted at middle. Sides of pronotum slightly sinuate just before hind angles (Fig. 78). Sterna IV–VI without punctures at sides. Aedeagus in dorsal view moderately arcuate (Fig. 114).................................................. Blennidus pseudolaevis ( Allegro & Giachino, 2011) 9. Male fore tarsi scarcely dilated, without scaly pubescence underneath. Head very large. Pronotum clearly transverse (W/ L> 1.4) (Fig. 63). The 1 st elytral stria definitely more impressed than others. Large body size (11–13 mm). Dept. Pasco (Cerro de Pasco)............................................................ Blennidus inca (Tschitschérine, 1898) -.. Male fore-tarsi normally dilated, with scaly pubescence underneath. The other above characters not all present together 10. 10. Pronotum widely flattened before hind angles, which are angulately obtuse and clearly raised (Fig. 80). Elytra with subparallel sides and striae all equally impressed. 9.5 mm. Unknown distribution............ Blennidus refleximargo (Straneo, 1993) - Pronotum with different shape......................................................................... 11. 11. Elytra with a single discal setigerous pore on 3 rd interval and with marked green or bluish metallic lustre.............. 12. - Elytra with at least two discal setigerous pores on 3 rd interval. Brown or black species, at most with slight metallic lustre. 14. 12. Elytra with green-bluish lustre. Striae superficially impressed but evident; the 1 st, as well as the 8 th and 9 th, more impressed than others. Sides of pronotum regularly rounded with a very shortly sinuation before basal angles, which are obtusely angulate (Fig. 84). 9 mm. Unknown distribution.................................... Blennidus tenenbaumi (Lutshnik, 1927) - Elytra with marked green-olive lustre................................................................... 13. 13. Pronotum with basal angles right and acutely pointed (Fig. 71). Head with frontal furrows very faint. Elytral striae all very superficial. 9 mm. Unknown distribution............................... Blennidus olivaceus (Tschitschérine, 1897) - Pronotum with basal angles obtuse, with a small blunt tooth at vertex. Head with frontal furrows impressed. The 1 st elytral stria, as well the 7 th– 9 th, more impressed than others. 9 mm. Unknown distribution.................................................................................................... Blennidus idioderus (Tschitschérine, 1898) 14. Lateral margins of pronotum with two close setigerous pores on anterior half. Posterior angles of pronotum rounded (Fig. 60) 8.5 mm. Unknown distribution...................................... Blennidus euphaenops (Tschitschérine, 1898) - Lateral margins of pronotum with a single setigerous pore on anterior half. Posterior angles of pronotum rounded or angulate.................................................................................................. 15. 15. Species ranging in the mountains of Cordillera Blanca (Dept. Ancash). Only the 1 st elytral stria well impressed, the others superficial or obliterated. Male mesotibiae preapically swollen. Seta on posterior angles of pronotum not moved forewards.................................................................................................... 16. - Species recorded from other Andean areas of Peru. The above characters not all present together.................... 17. 16. Posterior angles of pronotum almost rounded (Fig. 86). Larger size (8.5–9.4 mm). Median lobe of aedeagus stouter (Fig. 119)........................................................................ Blennidus unistria (Straneo, 1993) - Posterior angles of pronotum obtuse but clearly indicated by a small tooth (Fig. 61). Smaller size (7.4–9.2 mm). Median lobe of aedeagus slender (Fig. 97).................................... Blennidus fitzcarraldi (Giachino & Allegro, 2011) 17. Species ranging in the Andes of southern Peru. Elytral striae all equally impressed (at most the 1 st slightly more impressed). Male mesotibiae not preapically swollen (sometimes metatibiae preapically swollen). Seta on posterior angles of pronotum not moved forewards................................................................................... 18. - Species recorded from other Andean areas of Peru. The above characters not all present together.................... 23. 18. Sides of pronotum sinuate at the base or posteriorly nearly straight; male metatibiae not preapically swollen........... 19. - Sides of pronotum rounded over entire length; male metatibiae preapically swollen or not.......................... 22. 19. Impressed elytral striae with feebly convex intervals in ♂. Distinctly pear-shaped elytra. Aedeagus with median lobe regularly curved and not depressed ventrally (Fig. 100). Species from the Puno area. 7.7–7.8 mm................................................................................................... Blennidus jelskii (Tschitschérine, 1897) - Superficial elytral striae with flat intervals in both sexes. Elytra not pear-shaped. Aedeagus with median lobe angulately inserted on the basal bulb and ventrally somewhat depressed. Species from the Cuzco area........................ 20. 20. Sides of pronotum markedly sinuate at base (Fig. 51). A faint metallic lustre is usually present in ♂. In lateral view, median lobe of aedeagus larger and distally nearly straight (Fig. 89). Relatively larger (8.0–9.0 mm)..................................................................................... Blennidus abramalagae (Allegro & Giachino, 2011) - Sides of pronotum weakly sinuate or nearly straight at base. Metallic lustre usually absent. Median lobe of aedeagus smaller and angulately bent downwards in the distal part (Figs. 113, 117). Relatively smaller (6.6–8.8 mm).................. 21. 21. Habitus more slender (length/width of elytra = 1.50–1.57). Elytra less convex, with apical declivity less marked. Aedeagus smaller even in the largest specimens (Fig. 113). 6.6–8.8 mm............ Blennidus procerus (Allegro & Giachino, 2011) - Habitus oval (length/width of elytra = 1.45). Elytra more convex, with marked apical declivity. Aedeagus more slender and larger (Fig. 117). 7.7–8.7 mm...................................... Blennidus straneoi (Allegro & Giachino, 2011) 22. Elytral striae definitely impressed, with slightly convex intervals. Male metatibiae preapically not swollen. Median lobe of aedeagus distally rounded in dorsal view (Fig. 91). Larger size (9.5 mm). Species from the Puno area.................................................................................. Blennidus aulacostigma (Tschitschérine, 1897) - Elytral striae superficial, with flat intervals. Male metatibiae preapically swollen. Median lobe of aedeagus distally pointed in dorsal view (Fig. 95). Smaller size (7.6–9.3 mm). Species from the Cuzco area.................................................................................................... Blennidus etontii (Allegro & Giachino, 2011) 23. Hind angles of pronotum more or less rounded, always blunt at apex, and postangular seta distinctly moved forewards... 24. - Hind angles of pronotum rounded or angulate; if rounded the postangular seta not moved forewards................. 27. 24. The 1 st elytral stria distinctly more impressed than others; intervals flat. Elytra oval shaped, narrow at base and dilated backwards; shoulders rounded. Male mesotibiae not preapically swollen. The apical blade of aedeagus slightly bent to left in dorsal view (Fig. 118). Ticlio, Abra de Anticona..................................... Blennidus ticlianus (Straneo, 1993) - Elytral striae well impressed and intervals subconvex; the 1 st stria only slightly more impressed than others. Male mesotibiae preapically swollen. The apical blade of aedeagus not laterally bent in dorsal view. Species from Depts. Pasco and Huanuco.................................................................................................... 25. 25. Male metatibiae with a marked subapical swelling. Depts. Pasco and Huanuco.. Blennidus languens (Tschitschérine, 1898) - Male metatibiae without marked subapical swelling........................................................ 26. 26. Larger size (10–12 mm). Elytra oval shaped, narrower at base and dilated backwards. Front angles of pronotum not prominent (Fig. 74).............................................................. Blennidus pascoensis (Straneo, 1954) - Smaller size (9–9.5 mm). Elytra shorter, with subparallel sides, not distinctly dilated backwards. Front angles of pronotum prominent (Fig. 55)........................................................... Blennidus bombonensis n. sp. 27. Hind angles of pronotum broadly or narrowly rounded...................................................... 28. - Hind angles of pronotum from almost right (sometimes blunt at vertex) to obtuse, but always angulate................ 34. 28. Larger size (> 11 mm). Elytral striae more or less equally impressed. Posterior angles of pronotum broadly rounded (Fig. 67). Depts. Junin and Huancavelica............................................... Blennidus mateui (Straneo, 1993) - Smaller size ( 29. Elytra short-oval, relatively wide (L/W= 1.40). Sides of pronotum arcuate and hind angles broadly obtuse (Fig. 82). 7.1 mm. Dept. La Libertad........................................................ Blennidus sublaevis (Straneo, 1993) - Elytra more slender (L/W= 1.50). Body larger (> 7.5 mm).................................................... 30. 30. Smaller (7.5–8.2 mm). Apical blade of aedeagus short and almost triangular in dorsal view......................... 31. - Larger (9.1–10.2 mm). Apical blade of aedeagus longer, widely truncate at apex and bent to the right in dorsal view (Figs. 88, 102).............................................................................................. 33. 31. Body dark brown. Legs reddish-brown. Elytral striae 2–5 generally superficial, distinctly less impressed than 1 st. Dept. Junin..................................................................... Blennidus vereshaginae (Straneo, 1993) - Body piceous black, with concolorous legs. Striae 2–5 complete and distinct, although less impressed than 1 st.......... 32. 32. Posterior angles of pronotum rounded (Fig. 83). Unknown distribution........... Blennidus tardus (Tschitschérine, 1898) - Posterior angles of pronotum obtuse with blunt vertex (Fig. 52). Dept. Lima...... Blennidus anxius (Tschitschérine, 1898) 33. Males usually with bronze lustre. Aedeagus not exceptionally slender (Fig. 88). Dept. Huancavelica................................................................................... Blennidus abditus (Giachino & Allegro, 2011) - Males usually without bronze lustre. Aedeagus exceptionally long and slender, very distinctive (Fig. 102). Dept. Junin............................................................................. Blennidus longiloba (Straneo, 1993) 34. Sides of pronotum markedly convex, widest at anterior third (Figs. 58, 73). The 1 st elytral stria definitely more impressed than others, which are very superficial; intervals flat........................................................... 35 - Sides of pronotum moderately convex, sometimes linear towards base, widest at middle or just beyond middle. The 1 st elytral stria hardly more impressed than others, which are complete and distinct; intervals flat or subconvex................. 36. 35. Smaller size (7 mm). Sides of pronotum less rounded in basal half, with very short sinuation at base (Fig. 58). Unknown distribution.............................................................. Blennidus egens (Tschitschérine, 1898) - Larger size (8.4 mm). Sides of pronotum evenly rounded on whole length (Fig. 73). Dept. Junin........................................................................................ Blennidus pachycerus (Tschitschérine, 1897) 36. Pronotum subquadrangular with sides hardly convex, almost linear or subsinuate before hind angles (Figs. 57, 70, 75)... 37. - Pronotum subquadrangular or transverse, with sides more convex and usually evenly rounded before hind angles....... 39. 37. Elytra short-oval, dull even in males due to a strong microsculpture; striae distinctly punctate and very superficial in posterior third. 8–9 mm. Depts. Pasco and Junin......................................... Blennidus curtatus (Straneo, 1993) - Elytra oval-elongate, moderately shiny in males; striae impunctate and well impressed even in posterior third.......... 38. 38. Larger size (8.5–9 mm). Sides of pronotum linear (or slightly convex) in basal half (Fig. 70). Elytra angulate at shoulders. Median lobe of aedeagus shorter and stouter (Fig. 106). Depts. Junin-Pasco......... Blennidus nigritulus (Straneo, 1993) - Smaller size (7.6 mm). Sides of pronotum with short sinuation before hind angles (Fig. 75). Elytra rounded at shoulders. Median lobe of aedeagus more slender (Fig. 111). Unknown distribution.... Blennidus phaenogonus (Tschitschérine, 1898) 39. Smaller size (8.2 mm). Fore angles of pronotum completely obliterated (Fig. 52). Median lobe of aedeagus hardly curved downwards at apex in lateral view (Fig. 90). Dept. Lima...................... Blennidus anxius (Tschitschérine, 1898) - Larger size (8.5–9 mm). Fore angles of pronotum more or less prominent (Fig. 79). Median lobe of aedeagus distinctly curved downwards at apex in lateral view (Fig. 115). Dept. Junin...................... Blennidus rectangulus (Straneo, 1993)

Blennidus refleximargo group sensu novo Diagnosis. A single species of unknown distribution, distinguished by pronotum widely flattened before hind angles, which are clearly raised: B. (Agraphoderus) refleximargo (Straneo, 1993), Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2015, Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae), pp. 1-48 in Zootaxa 4000 (1) on page 21, DOI: 10.11646/zootaxa.4000.1.1, http://zenodo.org/record/289219, {"references":["Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale \" G. Doria \", 89, 351 - 399."]}

Blennidus (Agraphoderus) nigritulus (Straneo, 1993) (Figs. 32, 70, 106) Ogmopleura nigritula Straneo, 1993: 383 Blennidus nigritulus (Straneo, 1993): Lorenz, 2005 a: 263 Blennidus nigritulus (Straneo, 1993): Lorenz, 2005 b: 681 Blennidus nigritulus (Straneo, 1993): Giachino & Allegro, 2011: 23 Type locality. The ♂ HT is labelled, in Straneo���s own hand, ���loc. 14 ���, which likely means, by analogy with all other specimens of TS and according to Straneo (1993), ��� Perou, 30 km S Cerro de Pasco, 4150 m ���. Examined material. HT ♂ (CMa at MRSN); 4 PTT ♂ and 3 PTT ♀ (CMa, CSt); 1 ♂ (CMa). One PT ♂ in CMa actually belongs to B. curtatus. One specimen of B. nigritulus is included in the TS of angularis Straneo, 1985 (= B. rectangulus syn. nov.) (CMa) and 6 others in the TS of B. rectangulus (Straneo, 1993) (CMa, CSt). Note. This species is mainly distinguished by the distinctive morphology of aedeagus, as it is very similar in external morphology to a few species from central Peru (B. rectangulus, B. euphaenops, B. vereshaginae, etc.). An aedeagus of B. nigritulus is attached on separate cards by two specimens of B. inca (CMa, CSt), due to an evident label inversion. Distribution. All the recorded specimens are from the same area (30 Km South of Cerro de Pasco), which is probably a border area between Depts. Pasco and Junin. Habitat. The altitude data reported on the labels (4,150 m a.s.l.) suggest a preference of this species for the Andean grassland., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2015, Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae), pp. 1-48 in Zootaxa 4000 (1) on page 13, DOI: 10.11646/zootaxa.4000.1.1, http://zenodo.org/record/289219, {"references":["Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale \" G. Doria \", 89, 351 - 399.","Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp.","Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp.","Straneo, S. L. (1985) On the genus Sierrobius Straneo, 1951 (Coleoptera: Carabidae: Pterostichini). Annals of Carnegie Museum, 54 (7), 233 - 245."]}

Blennidus (Agraphoderus) jelskii (Tschitsch��rine, 1897) (Figs. 26, 64, 100) Feronia jelskii Tschitsch��rine, 1897: 290 Feronia jelskii Tschitsch��rine, 1897: Tschitsch��rine, 1898: 146 (note) Ogmopleura jelskii (Tschitsch��rine, 1897): Straneo & Vereshagina, 1991: 203 Ogmopleura jelskii (Tschitsch��rine, 1897): Straneo, 1993: 379 Blennidus jelskii (Tschitsch��rine, 1897): Lorenz, 2005 a: 263 Blennidus jelskii (Tschitsch��rine, 1897): Lorenz, 2005 b: 539 Blennidus jelskii (Tschitsch��rine, 1897): Allegro & Giachino, 2011 a: 297 Type locality. Peru, Puno. Examined material. LT ♂ (CTs at ZIRA), 1 PLT ♀ (CSt) (complete TS). Note. This species belongs to a species group (B. jelskii species group) including 5 species distributed in the Andes of southern Peru and distinguished by the peculiar morphology of aedeagus (Allegro & Giachino 2011 a). Distribution. This species is only known from the type specimens, which were collected by Jelski in the Andean District of Puno (southern Peru)., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2015, Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae), pp. 1-48 in Zootaxa 4000 (1) on page 11, DOI: 10.11646/zootaxa.4000.1.1, http://zenodo.org/record/289219, {"references":["Tschitscherine, T. (1897) Materiaux pour servir a l'etude des Feroniens. III. Horae Societatis Entomologicae Rossicae, 30, 260 - 351.","Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224.","Straneo, S. L. & Vereshagina, T. (1991) Sui tipi del genere Ogmopleura Tschitscherine descritti da Tschitscherine nel 1896 e 1898. Bollettino della Societa entomologica italiana, Genova, 122 (3), 195 - 204.","Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale \" G. Doria \", 89, 351 - 399.","Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp.","Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp.","Allegro, G. & Giachino, P. M. (2011 a) Studies on the subgenus Agraphoderus Bates of Blennidus Motschulsky from Peru: the jelskii species-group (Coleoptera, Carabidae, Pterostichini). Animal Biodiversity and Conservation, 34 (2), 295 - 308."]}

Blennidus (Agraphoderus) refleximargo (Straneo, 1993) (Figs. 42, 80, 116) Ogmopleura refleximargo Straneo, 1993: 384 Blennidus refleximargo (Straneo, 1993): Lorenz, 2005 a: 263 Blennidus refleximargo (Straneo, 1993): Lorenz, 2005 b: 801 Type area. Peru. Examined material. HT ♂ (Oberth��r Collection at MNHNP). Note. Species known from the HT specimen only. The name was attributed in litteris by Chaudoir. Easily distinguished from all other species by a wide depression before the hind angles of pronotum., Published as part of Allegro, Gianni & Giachino, Pier Mauro, 2015, Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae), pp. 1-48 in Zootaxa 4000 (1) on page 16, DOI: 10.11646/zootaxa.4000.1.1, http://zenodo.org/record/289219, {"references":["Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale \" G. Doria \", 89, 351 - 399.","Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp.","Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp."]}