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Understanding the neurobiology of social bonding in non-human primates is a critical step in understanding the evolution of monogamy, as well as understanding the neural substrates for emotion and behavior. Coppery titi monkeys (Callicebus cupreus) form strong pair bonds, characterized by selective preference for their pair mate, mate-guarding, physiological and behavioral agitation upon separation, and social buffering. Mate-guarding, or the "maintenance" phase of pair bonding, is relatively under-studied in primates. In the current study, we used functional imaging to examine how male titi monkeys viewing their pair mate in close proximity to a stranger male would change regional cerebral glucose metabolism. We predicted that this situation would challenge the pair bond and induce "jealousy" in the males. Animals were injected with [18F]-fluorodeoxyglucose (FDG), returned to their cage for 30 min of conscious uptake, placed under anesthesia, and then scanned for 1 hour on a microPET P4 scanner. During the FDG uptake, males (n=8) had a view of either their female pair mate next to a stranger male ("jealousy" condition) or a stranger female next to a stranger male (control condition). Blood and cerebrospinal fluid samples were collected and assayed for testosterone, cortisol, oxytocin, and vasopressin. Positron emission tomography (PET) was co-registered with structural magnetic resonance imaging (MRI), and region of interest analysis was carried out. Bayesian multivariate multilevel analyses found that the right lateral septum (Pr(b>0)=93%), left posterior cingulate cortex (Pr(b>0)=99%), and left anterior cingulate (Pr(b>0)=96%) showed higher FDG uptake in the jealousy condition compared to the control condition, while the right medial amygdala (Pr(b>0)=85%) showed lower FDG uptake. Plasma testosterone and cortisol concentrations were higher during the jealousy condition. During the jealousy condition, duration of time spent looking across at the pair mate next to a stranger male was associated with higher plasma cortisol concentrations. The lateral septum has been shown to be involved in mate-guarding and mating-induced aggression in monogamous rodents, while the cingulate cortex has been linked to territoriality. These neural and physiological changes may underpin the emotion of jealousy, which can act in a monogamous species to preserve the long-term integrity of the pair.

Early experience and social context interact to alter the phenotype of complex social behaviors. These early experiences can also result in alterations to cortical organization and connections. Given the ability of the neuropeptide oxytocin (OT) to modulate social and reproductive behavior, OT is likely involved in these cortical processes. However, little is known about the distribution of OT and OT receptors (OTR) within the neocortex. Using autoradiographic and neuroanatomical techniques, we characterized the cortical distribution of OT receptors (OTR) in prairie voles, a socially monogamous rodent species. We found that OTR density was low in the primary sensory areas (including primary somatosensory and auditory regions) but was quite high in association regions (including temporal and parietal association areas, and prelimbic regions). In the primary motor area as well as the temporal and parietal association areas, we observed differences in OTR density across cortical layers. Specifically, cortical layers 2/3 and 5 exhibited greater OTR density than layer 4. Our results point to a role for OT in integrating sensory and motor in the prairie vole brain, providing a complementary mechanism for the modulation of social interactions. Given the ability of early social experience and developmental manipulations of OT to affect the brain and behavior, these results suggest a novel mechanism for how OT may influence cortical organization.

Pair bonding leads to increases in dopamine D1 receptor (D1R) binding in the nucleus accumbens of monogamous prairie voles. In the current study, we hypothesized that there is similar up-regulation of D1R in a monogamous primate, the titi monkey (Callicebus cupreus). Receptor binding of the D1R antagonist [11 C]-SCH23390 was measured in male titi monkeys using PET scans before and after pairing with a female. We found that within-subject analyses of pairing show significant increases in D1R binding in the lateral septum, but not the nucleus accumbens, caudate, putamen, or ventral pallidum. The lateral septum is involved in a number of processes that may contribute to social behavior, including motivation, affect, reward, and reinforcement. This region also plays a role in pair bonding and paternal behavior in voles. Our observations of changes in D1R in the lateral septum, but not the nucleus accumbens, suggest that there may be broadly similar dopaminergic mechanisms underlying pair bonding across mammalian species, but that the specific changes to neural circuitry differ. This study is the first research to demonstrate neuroplasticity of the dopamine system following pair bonding in a non-human primate; however, substantial variability in the response to pairing suggests the utility of further research on the topic.

Social bonds are necessary for many mammals to survive and reproduce successfully. These bonds (i.e. pair-bonds, friendships, filial bonds) are characterized by different periods of development, longevity and strength. Socially monogamous species display certain behaviors not seen in many other mammals, such as adult pair-bonding and male parenting. In our studies of prairie voles (Microtus ochrogaster) and titi monkeys (Callicebus cupreus), we have examined the neurohormonal basis of these bonds. Here, we discuss the evidence from voles that aspects of adolescent and adult social behavior are shaped by early experience, including changes to sensory systems and connections, neuropeptide systems such as oxytocin and vasopressin, and alterations in stress responses. We will compare this with what is known about these processes during development and adulthood in other mammalian species, both monogamous and non-monogamous, and how our current knowledge in voles can be used to understand the development of and variation in social bonds. Humans are endlessly fascinated by the variety of social relationships and family types displayed by animal species, including our own. Social relationships can be characterized by directionality (either uni- or bi-directional), longevity, developmental epoch (infant, juvenile or adult) and strength. Research on the neurobiology of social bonds in animals has focused primarily on 'socially monogamous' species, because of their long-term, strong adult affiliative bonds. In this Review, we attempt to understand how the ability and propensity to form these bonds (or lack thereof), as well as the display of social behaviors more generally, are transmitted both genomically and non-genomically via variation in parenting in monogamous and non-monogamous species.

Unlike classical correlation, quantum entanglement cannot be freely shared among many parties. This restricted shareability of entanglement among multi-party systems is known as monogamy of entanglement, which is one of the most fundamental properties of entanglement. Here, we summarize recent theoretical progress in the field of monogamy of entanglement. We firstly review the standard CKW-type monogamy inequalities in terms of various entanglement measures. In particular, the squashed entanglement and one-way distillable entanglement are monogamous for arbitrary dimensional systems. We then introduce some generalized version of monogamy inequalities which extend and sharpen the traditional ones. We also consider the dual polygamy inequalities for multi-party systems. Moreover, we present two new definitions to define monogamy of entanglement. Finally, some challenges and future directions for monogamy of entanglement are highlighted.

We study monogamy and polygamy inequalities of unified entanglement in multipartite quantum systems. We first derive the monogamy inequality of unified-(q, s) entanglement for multi-qubit states under arbitrary bipartition, and then obtain the monogamy inequalities of the α th ( 0 ≤ α ≤ r 2 , r ≥ 2 ) power of entanglement of formation for tripartite states and their generalizations in multi-qubit quantum states. We also generalize the polygamy inequalities of unified-(q, s) entanglement for multi-qubit states under arbitrary bipartition. Moreover, we investigate polygamy inequalities of the β th ( β ≥ max { 1 , s } , 0 ≤ s ≤ s 0 , 0 ≤ s 0 ≤ 2 ) power of the entanglement of formation for 2 ⊗ 2 ⊗ 2 and n-qubit quantum systems. Finally, using detailed examples, we show that the results are tighter than previous studies. [ABSTRACT FROM AUTHOR]

Interest in attachment theory and polyamory, a form of consensual non-monogamy (CNM), has grown and evolved. However, romantic adult attachment is still understood within a monogamous construct, where insecurely attached individuals are stigmatized. The attachment literature describes those who exhibit dimensions of avoidant attachment as desiring multiple relationships with little emotional depth and commitment. However, empirical research illustrates that polyamorous individuals are predominantly securely or anxiously attached. Securely attached individuals are better able to communicate about intimate subjects, which often occurs in polyamory, while anxiously attached individuals thrive in an environment where intimacy is plentiful. This article provides an overview regarding attachment theory and polyamory and examines the contention that avoidantly attached individuals are attracted to CNM. While attachment theory is an empirically validated framework, modifications will be required in order to accommodate the polyamorous community. [ABSTRACT FROM AUTHOR]

The critically endangered indri (Indri indri) is the largest extant lemur species and its population size is projected to decline over the next three generations due to habitat loss, hunting and climate change. Accurate information on the demographic parameters driving the population dynamics of indri is urgently needed to help decision-making regarding the conservation of this iconic species. We monitored and followed the life histories of 68 individually recognizable indris in 10 family groups in the Maromizaha New Protected Area (Madagascar) for 12 years. We estimated age and sex-specific survival trajectories using a Bayesian hierarchical survival model and found that the survival curves for male and female indris show a similar pattern, consistent with what found typically in primates; i.e., a high infant mortality rate which declines with age in the juvenile phase and increases again for adults. Also, life expectancies at 2 years of age (e2), were found to be similar between the sexes (e2 females = 7.8 years; e2 males = 7.5 years). We suggest that the lack of strong differences in the survival patterns for male and female indris are related to the strictly monogamous mating system and the lack of sexual dimorphism in this species. Our study provides, for the first time, robust estimates for demographic parameters of indris and one of the very few datasets on survival trajectories available for primates.

Abstract Intraspecific competition plays an important role for territory acquisition and occupancy, in turn affecting individual fitness. Thus, understanding the drivers of intraspecific aggression can increase our understanding of population dynamics. Here, we investigated intraspecific aggression in Eurasian (Castor fiber) and North American (Castor canadensis) beavers that are both monogamous, territorial mammals. Combined, we examined tail scars from >1,000 beavers (>2,000 capture events) as part of two long‐term studies in Norway and the USA. We investigated the influence of landscape structure, population density, sex, age, and (for Eurasian beavers only) social status and group size on the number of tail scars caused by conspecifics. The number of tail scars was affected by population density in well‐connected landscape types (large lakes and rivers), but not in more isolated areas (ponds), where individuals generally had fewer tail scars. Further, the relationship of population density was not linear. In the North American beaver population occurring in large lakes, intraspecific aggression increased with population density. Conversely, in the saturated Eurasian beaver population, intraspecific aggression was in a negative relationship with population density (except at the highest densities), likely due to inverse density‐dependent intruder pressure via dispersers. Our findings emphasize that population density can affect intraspecific aggression depending on landscape structure, which might have important consequences for local patterns of dispersal, mate change, and territory occupancy, all of which can affect population dynamics.

Naked mole-rats form colonies with a single reproductively active female surrounded by subordinate workers. Workers perform offspring care, construction and defense of the burrow system, and food supply. Such division of labor, called “cooperative breeding,” is strongly associated with the evolution of monogamous mating behavior, as seen in several mammalian lineages. This association is explained by the evolutionary theory of kin selection, according to which a subordinate adult may help to raise other’s offspring if they are in full sibling relationship. In conflict with this theory, the naked mole-rat is widely considered to be polyandrous, based on reports on multiple males contributing to a colony’s progeny. In order to resolve this contrast, we undertook an in-depth microsatellite-based kinship analysis on captive colonies. Four independent colonies comprising a total of 265 animals were genotyped using a panel of 73 newly established microsatellite markers. Our results show that each mole-rat colony contains a single monogamous breeder pair, which translates to a reproductive skew of 100% for both sexes. This finding, also in conjunction with previously published parental data, favors monogamy as the best-fitting model to describe naked mole-rat reproduction patterns. Polyandry or other polygamous reproduction models are disfavored and should be considered as exceptional. Overall, the empirical genetic data are in agreement with the kin selection theory.

Relatively little is known about serotonergic involvement in pair-bonding despite its putative role in regulating social behavior. Here we sought to determine if pharmacological elevation of serotonin 1A (5-HT1A) receptor activity would lead to changes in social behavior in pair-bonded male titi monkeys (Callicebus cupreus). Adult males in established heterosexual pairs were injected daily with the selective 5-HT1A agonist 8-OH-DPAT or saline for 15days using a within-subjects design. Social behavior with the female pair-mate was quantified, and plasma concentrations of oxytocin, vasopressin, and cortisol were measured. When treated with saline, subjects showed reduced plasma oxytocin concentrations, while 8-OH-DPAT treatment buffered this decrease. Treatment with 8-OH-DPAT also led to decreased plasma cortisol 15minutes post-injection and decreased social behavior directed toward the pair-mate including approaching, initiating contact, lipsmacking, and grooming. The reduction in affiliative behavior seen with increased activity at 5-HT1A receptors indicates a substantial role of serotonin activity in the expression of social behavior. In addition, results indicate that the effects of 5-HT1A agonism on social behavior in adulthood differ between rodents and primates.

Mate-guarding and territorial aggression (both intra- and inter-sexual) are behavioral components of social monogamy seen in male coppery titi monkeys (Callicebus cupreus) both in the field and in the laboratory. Methodology for studying these behaviors in captivity facilitates the translation of questions between field and laboratory. In this study, we tested whether exposure to a mirror would stimulate mate-guarding behavior in male titi monkeys, and whether this exposure was accompanied by hormonal changes. Eight males were exposed to a mirror condition (treatment) or the back of the mirror (control) for five sessions, and behavioral responses were filmed. Blood samples were taken to measure levels of cortisol, oxytocin, and vasopressin. Lipsmacks (P

Partner preference, or the selective social preference for a pair mate, is a key behavioral indicator of social monogamy. Standardized partner preference testing has been used extensively in rodents but a single test has not been standardized for primates. The goal of this study was to develop a partner preference test with socially monogamous titi monkeys (Callicebus cupreus) adapted from the widely used rodent test. In Experiment 1, we evaluated the test with pairs of titi monkeys (N = 12) in a three-chambered apparatus for 3 hr. The subject was placed in the middle chamber, with grated windows separating it from its partner on one side and an opposite sex stranger on the other side. Subjects spent a greater proportion of time in proximity to their partners' windows than the strangers', indicating a consistent preference for the partner over the stranger. Touching either window did not differ between partners and strangers, suggesting it was not a reliable measure of partner preference. Subjects chose their partner more than the stranger during catch and release sessions at the end of the test. In Experiment 2, we compared responses of females with current partners (N = 12) in the preference test with other relationship types representing former attachment bonds (N = 13) and no attachment bond (N = 8). Only females from established pair bonds spent significantly more time near their partner's window compared to the stranger's indicating that this measure of preference was unique to current partners. Other measures of preference did not differentiate behavior toward a current partner and other relationship types. This test reproduces behavioral patterns found in previous studies in titi monkeys highlighting the accuracy of this new partner preference test. This test can be used as a standardized measure of partner preference in titi monkeys to quantitatively study pair bonding and evaluate factors influencing partner preference.

This paper is the introduction to a special issue on "'Monogamy' in Primates: Variability, Trends, and Synthesis." The term "monogamy" has undergone redefinition over the years, and is now generally understood to refer to certain social characteristics rather than to genetic monogamy. However, even the term "social monogamy" is used loosely to refer to species which exhibit a spectrum of social structures, mating patterns, and breeding systems. Papers in this volume address key issues including whether or not our definitions of monogamy should change in order to better represent the social and mating behaviors that characterize wild primates; whether or not primate groups traditionally considered monogamous are actually so (by any definition); ways in which captive studies can contribute to our understanding of monogamy; and what selective pressures might have driven the evolution of monogamous and non-monogamous single female breeding systems.

We explore the monogamy of multipartite quantum discord. In the article [Quantum Science and Technology 6, 4, 045028], Guo et al. show that quantum discord for multiqubit systems is monogamous provided that it does not increase under discard of subsystems. we illustrate that the above-mentioned preconditions are valid for a family of multiqubit states. Based on the analytical expressions of quantum discord and geometric discord of the family of states obtained in the articles [Phys. Rev. A 104, 012428] and [arXiv:], we investigate the dynamic behavior of these under the local decoherence channel, and show that the quantum discord of even partite systems have frozen phenomenon while the odd partite systems does not exist. For geometric discord, this family of states has the frozen phenomenon under local decoherence conditions. The results show that compound noises are not necessary for sudden changes in quantum correlation, and one qubit of the quantum noise is sufficient. The research of these non-loss conditions is of great significance for understanding the evolution of quantum systems in the environment. [ABSTRACT FROM AUTHOR]

Female mate choice can involve a complex assessment process which is hypothesized to have evolved in order for females to maximize their reproductive success. In this laboratory experiment, we examined the mate choices of female convict cichlids (Amatitlania nigrofasciata) (who have an extended period of mate assessment and benefit from relatively long phases of biparental care) when presented a dichotomous choice between males that they had previously observed winning or losing a fight or novel males. When winner and loser males were presented to females together, females did not show a general preference for either male, but spent substantial time alone, away from both males. When the winner male was paired with a novel male, females showed an evident preference for the winner male. Females also showed an eventual preference for the loser male when paired with a novel male, but in this case, the female spent initially more time with the novel male before shifting to the loser male. In spite of losing their initial competition, loser males were attractive to females, more so than novel males, illustrating that participation in a fight observed by a female is beneficial to the male, even if he is not ultimately victorious. [ABSTRACT FROM AUTHOR]

We investigated factors leading to variation in social complexity or 'social systems' among plateau pika family groups within a contiguous local population across 2 years. Plateau pikas are small, diurnal, nonhibernating, sexually monomorphic lagomorphs that occupy family home ranges on open alpine meadows on the Qinghai-Tibetan Plateau. Expression of the social organization, social structure, mating system and parental care system in plateau pikas did not follow expectations from traditional ecological or evolutionary explanations. Variability in plateau pika family group size and the transitions of group size between years allowed us to investigate potential advantages and disadvantages of group living. Evidence that group living served to protect pikas against predation was weak. Although social huddling could have minimized thermoregulatory costs during the extremely cold Tibetan winters, there was no correlation of overwinter survivorship among pika families of different sizes. There was no apparent group-living benefit with regard to foraging, and the occurrence of cohesive social families on the flat, continuous meadow contradicts the hypothesis that sociality is related to patchiness of critical resources. Cost of maintaining burrows appeared unrelated to group size. Most interactions between pikas occurred within family groups and were affiliative (99% of adult interactions; 97% of adult–juvenile interactions), and most interactions between adult males of different family groups were aggressive (96% of interactions). Matings were primarily within families (88% of copulations). Pikas also possess a complex vocal repertoire that enhanced interactions within social families. Demographic constraints associated with variable overwinter survivorship appeared to be the dominant precondition that produced a given family size and mating system type, coupled with selective dispersal by some pikas before the start of the breeding season. Paternal care enhanced juvenile survival, and thus led to an equalization of reproductive success among adults in families with different mating combinations. • Plateau pikas live in families occupying exclusive burrow systems and home ranges. • Pika family group size varied from one to five adults within and between years. • Pikas failed to adhere to conventional explanations for establishment of group size. • Pika families exhibited multiple social mating system types. • Pika families did not adhere to explanations for establishment of mating system types. [ABSTRACT FROM AUTHOR]

We study the monogamy and polygamy inequalities of quantum correlations in arbitrary dimensional multipartite quantum systems. We first derive the monogamy inequality of the α th 0 ≤ α ≤ r 2 , r ≥ 2 power of concurrence for any 2 ⊗ 2 ⊗ 2n− 2 tripartite states and generalize it to the n-qubit quantum states. In addition to concurrence, we show that the monogamy relations are satisfied by other quantum correlation measures such as entanglement of formation. Moreover, the polygamy inequality of the β th (β ≤ 0) power of concurrence and the β th (β ≥ s, 0 ≤ s ≤ 1) power of the negativity are presented for 2 ⊗ 2 ⊗ 2n− 2. We then obtain the polygamy inequalities of quantum correlations for multipartite states. Finally, our results are shown to be tighter than previous studies using detailed examples. [ABSTRACT FROM AUTHOR]

Purpose: This study aimed to investigate the sexual satisfaction and mental health indicators in polygamous and monogamous women in Turkey. The study included 108 female participants: 36 monogamous wives and 72 polygamous wives. Materials and Methods: The participants were evaluated with the Symptom Check List (SCL-90-R) and GolombokRust Inventory of Sexual Satisfaction (GRISS). Results: There were significant differences between the groups regarding the total GRISS score and Dissatisfaction, Sensuality, Vaginismus, and Anorgasmia subscale scores. In comparing the SCL-90 scores with ANOVA, the senior wife group had significantly higher scores in the Depression subscales. Monogamic wives had higher satisfaction from their sexual lives and had less frequent psychiatric symptoms than polygamic wives. Conclusion: It is important to raise awareness about the polygamic family structure, draw attention to often neglected issues such as sexual health, and improve the health services provided to this group. [ABSTRACT FROM AUTHOR]

Saudi Arabia has long been portrayed as a “hub of polygyny,” a practice of marriage in which a man marries several women simultaneously. However, my recent research among Saudi male youths suggests that this practice is waning nowadays. A younger generation in the Kingdom seems to prefer monogamy—a man marrying one wife—to polygamy for several essential reasons, factors, and arguments. Based on interviews and conversations with several young Saudis as well as outcomes of survey findings, this article tries to debunk the myth of polygyny in the Kingdom and attempts to understand the rationales and logic behind monogamy choice among male (and female) youths of contemporary Saudi Arabia. The article also briefly highlights the practice of polygyny and monogamy in multiple societies outside Saudi Arabia to compare and gain knowledge on various practices of marriage. Seen from another viewpoint, the article is a reflection of a modern-day fruitful socio-cultural development and change in Saudi Arabia that have received enthusiastic and public acclaim across the globe. [ABSTRACT FROM AUTHOR]

Given the historical entrenchment of racialised stereotypes of Black women and Black men as sexually promiscuous, we wondered whether consensual nonmonogamy (CNM) among Black partners would be seen as favourably as among white partners. We also wondered if Black participants would perceive different relationship types differently from white participants. We pursued these questions in a vignette study featuring heterosexual couples coded as Black or as white and engaged in three different relationship types: monogamy, nonconsensual nonmonogamy (NCNM) and CNM. To facilitate comparisons across race*gender intersections, we used a sample comprising equivalent numbers of Black women, white women, Black men and white men aged 18–40. Contrary to expectations, analyses did not offer evidence of a racialised sexual double standard insofar as participant perceptions of relationship quality did not differ when considering a Black couple or a white couple. Indicating the persistence of mononormativity, participants across race*gender subsamples perceived monogamous relationships to be of higher quality, regardless of the vignette couple's race. We also found Black women, Black men and white women perceived CNM more favourably than NCNM, while there was no differentiation between CNM and NCNM among white men. [ABSTRACT FROM AUTHOR]

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The critically endangered indri (Indri indri) is the largest extant lemur species and its population size is projected to decline over the next three generations due to habitat loss, hunting and climate change. Accurate information on the demographic parameters driving the population dynamics of indri is urgently needed to help decision-making regarding the conservation of this iconic species. We monitored and followed the life histories of 68 individually recognizable indris in 10 family groups in the Maromizaha New Protected Area (Madagascar) for 12 years. We estimated age and sex-specific survival trajectories using a Bayesian hierarchical survival model and found that the survival curves for male and female indris show a similar pattern, consistent with what found typically in primates; i.e., a high infant mortality rate which declines with age in the juvenile phase and increases again for adults. Also, life expectancies at 2 years of age (e2), were found to be similar between the sexes (e2 females = 7.8 years; e2 males = 7.5 years). We suggest that the lack of strong differences in the survival patterns for male and female indris are related to the strictly monogamous mating system and the lack of sexual dimorphism in this species. Our study provides, for the first time, robust estimates for demographic parameters of indris and one of the very few datasets on survival trajectories available for primates. [ABSTRACT FROM AUTHOR]

Abstract: One form of innovation proposed by Islamic banks to support the needs of the community is The Sharia Gold-pawning Service. However, worries arise for the Rahn service (gold pawning) since the service is not beneficial in gaining profits. By this service, Islamic banks only gain profits from the maintenance service of goods that are pledged as collateral. The problem occurs as the profit is taken from the percentage of the amount of the loan provided by a Sharia bank to its customer which is considered to have no cost differential as the practices undertaken in conventional banking. Therefore, this is an interesting topic to discuss among academics related to the amount and calculation of maintenance services that customers must pay to the bank. If maintenance services are determined based on the percentage of the financing or loan’s amount, then there is a credit interest value on the financing that must be returned by the customer as a collateral for maintenance services calculated from the amount of the loan. Thus, this study is intended to analyze the concept of covenant and its application in financing gold pawning in Islamic banks. The objects of this study are those concerning the Islamic Commercial Bank, including Bank Syariah Mandiri and Bank Jabar Banten Syariah. Abstrak: Di dalam Undang-undang Nomor 1 tahun 1974 Pasal 3 ayat 1 dan 2, memberi­kan penjelasan tentang asas perkawinan adalah monogami akan tetapi bersifat terbuka, atau dalam KHI menggunakan istilah poligami terbatas sampai dengan empat isteri, akan tetapi dengan catatan kemampuan memberi nafkah dan berbuat adil terhadap isteri-isteri dan anak-anaknya. Rumah tangga yang dibangun berdasarkan poligami sebagian masyarakat menjadi hal yang kontroversi, akan tetapi normatifitas hukum telah disahkan dan itu sebuah realitas sosial terjadi. Metode yang digunakan dalam penelitian ini dengan mengumpulkan data berupa pustaka pustaka (literature research), adapun langkah-langkah dalam penelitian ini dengan membaca literatur berupa buku, artikel, peraturan perundang-undangan serta fatwa. Data tersebut di analisis dengan metode deskriftif kualitatif guna memperoleh hasil penelitian dengan berpijak pada teori yang terkait dengan penelitian Hasil penelitian menunjukan bahwa tidak ada perempuan yang rela di poligami. Pada kasus poligami perempuan merupakan obyek kekerasan yang terbungkus rapi dalam kehidupan rumahtangga.

Abstract Ungulates exhibit diverse mating systems that range from monogamous pair territories to highly polygynous leks. We review mating systems and behaviors across ungulates and offer a new approach synthesizing how interacting factors may shape those mating systems. Variability exists in mating systems among and within species of ungulates and likely is affected by predation risk, availability of resources (food and mates), habitat structure, and sociality. Ungulate mating systems may be labile as a consequence of the varying strength of those interacting factors. In addition, degree of polygyny and sexual dimorphism in size are associated with the evolution of mating systems. Neither male–male combat nor paternal care, however, can completely explain differences in sexual size dimorphism for ungulates, a necessary component in understanding the development of some mating systems. Whatever the evolutionary pathway, sexual segregation limits paternal care allowing more intense male–male competition. Selection of habitat structure, because it modifies risk of predation, is a major determinant of sociality for ungulates. Likewise, ruggedness and steepness of terrain limit the types of mating systems that can occur because of limitations in group size and cohesiveness, as well as the ability of males to herd even small groups of females effectively. The quality and defensibility of resources affect mating systems, as does the defensibility of females. Population density of females also may be a critical determinant of the types of mating systems that develop. Size of groups likewise constrains the types of mating tactics that males can employ. Our aim was to use those relationships to create a broad conceptual model that predicts how various environmental and social factors interact to structure mating systems in ungulates. This model provides a useful framework for future tests of the roles of both ecological and social conditions in influencing the social systems of ungulates.

Quantum entanglement is known to be monogamous, i.e. it obeys strong constraints on how the entanglement can be distributed among multipartite systems. Almost all the entanglement monotones so far are shown to be monogamous. We explore here a family of entanglement monotones with the reduced functions are concave but not strictly concave and show that they are not monogamous. They are defined by four kinds of the ‘partial-norm’ of the reduced state, which we call them partial-norm of entanglement , minimal partial-norm of entanglement, reinforced minimal partial-norm of entanglement, and partial negativity , respectively. This indicates that, the previous axiomatic definition of the entanglement monotone needs supplemental agreement that the reduced function should be strictly concave since such a strict concavity can make sure that the corresponding convex-roof extended entanglement monotone is monogamous. Here, the reduced function of an entanglement monotone refers to the corresponding function on the reduced state for the measure on bipartite pure states.

The Scandinavian medieval ballad "Olav Liljekrans" (TSB A 63) tells the story of a knight whose fate is determined by a meeting with the elves on the night before his marriage celebration. In this article the ballad is read in a social and historical context. It is argued that the ballad's main theme is Olav's lack of faithfulness to his coming bride, reflecting the transition in Scandinavia, as well as in Europe, from a pre-Christian marriage customs accepting polygamy, to an ecclesiastical marriage, only accepting monogamy.

There is great diversity in social behavior across the animal kingdom. Understanding the factors responsible for this diversity can help inform theory about how sociality evolves and is maintained. The Australian Tree Skink (Egernia striolata) exhibits inter- and intra-population variability in sociality and is therefore a good system for informing models of social evolution. Here, we conducted a multi-year study of a Tree Skink population to describe intra-population variation in the social organization and mating system of this species. Skinks aggregated in small groups of 2–5 individuals, and these aggregations were typically associated with shared shelter sites (crevices and hollows within rocks and trees). Aggregations were typically made up of one or more adult females and, often, one male and/or juvenile(s). Social network and spatial overlap analyses showed that social associations were strongly biased toward kin. Tree skinks also exhibited high site fidelity regardless of age or sex. There were high levels of genetic monogamy observed with most females (87%) and males (68%) only breeding with a single partner. Our results indicate that Tree Skinks reside in small family groups and are monogamous, which corresponds with existing research across populations. Similar to previous work, our study area consisted of discrete habitat patches (i.e., rock outcrops, trees, or both), which likely limits offspring dispersal and promotes social tolerance between parents and their offspring. Our study clearly demonstrates that there is intra-population variability in Tree Skink social behavior, but it also provides evidence that there is a high degree of inter-population consistency in sociality across their geographic range. We also highlight promising possible avenues for future research, specifically discussing the importance of studying the nature and extent of Tree Skink parental care and quantifying the fitness outcomes of kin-based sociality in this species, which are topics that will further our understanding of the mechanisms underlying variation in vertebrate social behavior.

One of the fundamental traits of quantum entanglement is the restricted shareability among multipartite quantum systems, namely monogamy of entanglement, while it is well known that monogamy inequalities are always satisfied by entanglement measures with convexity. Here we present a measure of entanglement, logarithmic convex-roof extended negativity (LCREN) satisfying important characteristics of an entanglement measure, and investigate the monogamy relation for logarithmic negativity and LCREN both without convexity. We show exactly that the αth power of logarithmic negativity, and a newly defined good measure of entanglement, LCREN, obey a class of general monogamy inequalities in multiqubit systems, 2⊗2⊗3systems and 2⊗2⊗2nsystems for α≥4ln2. We provide a class of general polygamy inequalities of multiqubit systems in terms of logarithmic convex-roof extended negativity of assistance (LCRENoA) for 0≤β≤2. Given that the logarithmic negativity and LCREN are not convex, these results are surprising. Using the power of the logarithmic negativity and LCREN, we further establish a class of tight monogamy inequalities of multiqubit systems, 2⊗2⊗3systems and 2⊗2⊗2nsystems in terms of the αth power of logarithmic negativity and LCREN for α≥4ln2. We also show that the βth power of LCRENoA obeys a class of tight polygamy inequalities of multiqubit systems for 0≤β≤2.

Mating systems, influenced by the social and ecological environment and individual attributes, are fundamental components of animal social organisation, impacting behaviour, animal distribution, ecosystem processes, individual reproductive success, and population dynamics. Bats are of particular interest for studies of mating systems as they are thought to exhibit a greater diversity in mating systems than any other mammalian order, and thus make great models for improving our fundamental understanding of causes and consequences of social organisation. Here, we review the current knowledge of bat mating systems. Our analyses show that research on bat mating systems has not kept pace with research on bats in general and that traditional typologies do not accommodate the mating system of several species. Therefore, we propose an alternative, functional framework to categorise mating systems of bats and by extension of other taxa. We argue that mating systems can be classified according to a male reproductive skew continuum, with an increasing skew from monogamy to true lekking. We include an additional category of lek-like mating system along the continuum to account for previous trans-categorical cases that have the appearance of resource defence but are functionally akin to a lek. The new framework has a total of seven categories: promiscuity, monogamy, female defence polygyny, resource defence polygyny, a lek-like mating system, exploded classical lek, and clustered classical lek. Applying this framework to bats reveals that lek mating systems are more prevalent in bats than previously recognised. It is our aim that this review and the proposed framework provide a greater understanding of bat mating systems particularly and provoke research into the factors that shape mating systems across animal taxa more generally., Competing Interests: The authors declare no conflict of interest., (© 2024 The Author(s). Ecology and Evolution published by John Wiley & Sons Ltd.)

The quality of romantic relationships can predict health consequences related to aging. DNA methylation-based biomarkers of aging accurately estimate chronological age. We developed several highly accurate epigenetic aging clocks, based on highly conserved mammalian CpGs, for the socially monogamous prairie vole (Microtus ochrogaster). In addition, our dual-species human-vole clock accurately measured relative age and illustrates high species conservation of epigenetic aging effects. Next, we assessed how pair bonding impacts epigenetic aging. We did not find evidence that pair-bonded voles exhibit accelerated or decelerated epigenetic aging effects in blood, ear, liver, or brain tissue. Our epigenome wide association study identified CpGs in five genes strongly associated with pair bonding: Foxp4, Phf2, Mms22l, Foxb1, and Eif1ad. Overall, we present accurate DNA methylation-based estimators of age for a species of great interest to researchers studying monogamy in animals. We did not find any evidence that sex-naive animals age differently from pair-bonded animals., (© 2024. The Author(s).)

Genes involved in reproduction often evolve rapidly at the sequence level due to postcopulatory sexual selection (PCSS) driven by male-male competition and male-female sexual conflict, but the impact of PCSS on gene expression has been under-explored. Further, though multiple tissues contribute to male reproductive success, most studies have focused on the testes. To explore the influence of mating system variation on reproductive tract gene expression in natural populations, we captured adult males from monogamous Peromyscus californicus and polygynandrous P. boylii and P. maniculatus. We generated RNAseq libraries, quantified gene expression in the testis, seminal vesicle, epididymis, and liver, and identified 3627 mating system-associated differentially expressed genes (MS-DEGs), where expression shifted in the same direction in P. maniculatus and P. boylii relative to P. californicus. Gene expression variation was most strongly associated with mating behaviour in the seminal vesicles, where 89% of differentially expressed genes were MS-DEGs, including the key seminal fluid proteins Svs2 and Pate4. We also used published rodent genomes to test for positive and relaxed selection on Peromyscus-expressed genes. Though we did not observe more overlap than expected by chance between MS-DEGs and positively selected genes, 203 MS-DEGs showed evidence of positive selection. Fourteen reproductive genes were under tree-wide positive selection but convergent relaxed selection in P. californicus and Microtus ochrogaster, a distantly related monogamous species. Changes in transcript abundance and gene sequence evolution in association with mating behaviour suggest that male mice may respond to sexual selection intensity by altering aspects of sperm motility, sperm-egg binding and copulatory plug formation., (© 2024 The Author(s). Molecular Ecology published by John Wiley & Sons Ltd.)

Field studies of the factors that influence reproductive success are crucial to life history theory. We investigate annual fledgling production by the monogamous, cooperatively breeding Red-cockaded Woodpecker (Dryobates borealis) in relation to breeder age, experience, and group size at Brosnan Forest, South Carolina, and evaluate several associated hypotheses. Productivity in both female and male breeders increased nonlinearly with age, peaked at 6 and 7 years, respectively, and then declined. We tested if the early increase in productivity with age was associated with age, per se (general skills), or breeding experience. We found for both males and females that the productivity of novice breeders increased significantly with age, indicating age effects independent of prior experience, but that experienced breeders were no more productive than novices the same age. We conclude that the basic skills that accrue with age, but not prior breeding experience, explain increasing success among young birds. The probability of a group nesting increased nonlinearly with female and male age, reaching 100% among females [greater than or equal to] 7 and males [greater than or equal to] 11 years old. We found support for both the constraint and restraint hypotheses for young males, but only for the constraint hypothesis among young females. Declining productivity with senescence counters the restraint hypothesis's prediction of increased effort late in life, while the maximal propensity to nest among older birds supports it. Differential survival failed to explain increasing productivity with age. Holding breeder ages constant, we found a linear effect of group size on annual fledgling production, with productivity increasing by 12% with each added helper through the largest group sizes. This suggests (1) no evidence of net compensation as group size increased, (2) no evidence of negative feedback from reproductive or resource competition in large groups, and (3) inclusive fitness is a selective force in helping. Received 3 December 2019. Accepted 7 December 2020. Key words: age effects, breeding age hypothesis, breeding experience hypothesis, constraint hypothesis, cooperative breeding, helper effects, restraint hypothesis. Edad reproductiva, tamano de grupo y produccion de poiluelos en el carpintero Dryobates borealis que anida cooperativamente Los estudios de campo de los factores que influencian el exito reproductivo son cruciales para la teoria de historias de vida. Investigamos la produccion anual de poiluelos de Dryobates borealis, un carpintero monogamo que anida cooperativamente, en relacion con la edad reproductiva, experiencia y tamano de grupo en Brosnan Forest, South Carolina, y evaluamos varias hipotesis asociadas. La productividad en ambos, machos y hembras reproductivos, se incremento no-linealmente con la edad, tuvo su pico a 6 y 7 anos, respectivamente y posteriormente declino. Sometimos a prueba si el incremento temprano en productividad con la edad estaba relacionada con la edad per se (capacidades generales) o con la experiencia reproductiva. Encontramos que para ambos, machos y hembras, la productividad de los novatos anidando se incremento significativamente con la edad, indicando que los efectos de la edad son independientes de la experiencia previa, si bien los reproductivos experimentados no fueron mas productivos que los novatos de la misma edad. Concluimos que las habilidades basicas que se adquieren con la edad, mas no por su experiencia reproductiva previa, explican el incremento en exito reproductivo entre aves jovenes. La probabilidad de un grupo nidificante se incremento de manera no-lineal con la edad de hembras y machos, alcanzando 100% entre hembras de [greater than or equal to]7 y machos de [greater than or equal to]11 anos. Encontramos soporte para ambas hipotesis, limitaciones externas y autocontrol, para los machos jovenes, aunque solo para la hipotesis de limitaciones externas para hembras jovenes. El declive en la productividad con la senescencia se contrapone a la prediccion del autocontrol, que supone un incremento en el esfuerzo reproductivo a edades avanzadas, mientras que las aves mas viejas son mas propensas a anidar y la apoyan. La sobrevivencia diferencial no explica el incremento de la productividad con la edad. Al mantener las edades de anidacion constantes, encontramos un efecto lineal del tamano del grupo en la produccion anual de polluelos emancipados, con incrementos de productividad de 12% con la adicion de cada ayudante hasta los grupos mas grandes. Esto sugiere que (1) no hay evidencia de una compensacion neta con el incremento del tamano del grupo, (2) no hay evidencia de retroalimentacion negativa como resultado de la competencia reproductiva o por recursos en grupos grandes y (3) la ventaja reproductiva inclusiva es una fuerza selectiva en la anidacion cooperativa. Palabras clave: efecto de ayudantes, efecto de la edad, hipotesis de la edad reproductiva, hipotesis de la experiencia reproductiva, hipotesis de las limitaciones externas, hipotesis de autocontrol, reproduccion cooperativa., Long-term empirical studies of the factors that influence reproductive success are essential to the development of life history theory. Breeder age is one such factor that has received much attention, [...]