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Guiodytes cavicola Tian, 2013: 116 Type locality: Nongshui Cave, in Shangjia, Disu, Du���an County of northern Guangxi. Male genitalia (Figs. 2���3): Median lobe of aedeagus comparatively more elongate, moderately arcuate, ventral margin bisinuate, gradually contracted towards apex which is very narrow; basal orifice large; inner sac armed with a very long longitudinal copulatory piece; right paramere well developed, especially on base, left one narrower, shorter and very thin, each bearing 3���4 long setae apically. Additional material examined: 2 males and 1 female, Guangxi: Du���an County: Chengjiang: Ganwan: Nongzhong Cave I, 23 �� 56.644 N, 108 �� 10.072 E, 469 m, 2013 VI- 27, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin & Sunbin Huang; 1 male and 2 females, Guangxi: Du���an County: Longwan: Nongqu Cave I, 23 �� 56.021 N, 108 �� 10.962 E, 459 m, 2013 -VI- 27, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin & Sunbin Huang. All specimens are deposited in the insect collections of South China Agricultural University, Guangzhou, China (SCAU), except one male in Mus��um National d'Histoire Naturelle, Paris, France (MNHN). Distribution: Du���an County, northern Guangxi. Known from three limestone caves in Disu, Chengjiang and Longwan, respectively (Fig. 1)., Published as part of Tian, Mingyi, 2014, New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae), pp. 355-362 in Zootaxa 3861 (4) on page 357, DOI: 10.11646/zootaxa.3861.4.5, http://zenodo.org/record/225326, {"references":["Tian, M. Y. (2013) Occurrence of troglobitic clivinines in China (Insecta: Coleoptera: Carabidae). Journal of Cave and Karst Studies, 75 (2), 113 - 120."]}

Guiodytes deharvengi n. sp. (Figs. 1, 4��� 7) Holotype: male, Guangxi: Huanjiang Maonan Autonomous County: Shuiyuan Zhen: Sancai Cun: a cave named as Shuiku Dong, 24 �� 49.430 ���N, 108 ��05.504���E, altitude 280 m, 2012 -XII- 4, leg. Louis Deharveng, Anne Bedos, Arnaud Faille and Mingyi Tian, in SCAU; paratypes: 1 male and 2 females, ibid, in SCAU and MNHN, Paris, respectively. Length (from apex of left mandible to end of elytra): 9.2���9.5 mm; width: 2.5���2.8 mm. Habitus as in Fig. 4. Head stout, from apex of mandible longer than wide, HL/HW= 1.16���2.02, mean 1.18, or slightly longer than wide when measured from apex of clypeus, HL/HW= 0.81���0.84, mean 0.83; distinctly narrower than pronotum, HW/PW= 0.78���0.81, mean 0.8; clypeus (with wings) strongly transverse, convex, bisetose at base, widely and deeply emarginate at front, making clypeal wings very protruding anteriorly; supra-antennal plates well marked, clearly separated from clypeal wings by a small but obvious notch, nearly rounded, strongly convex, smooth and glabrous; frons and vertex convex; two pairs of supraorbital pores present, at behind of supra-antennal plates and at neck constriction, respectively, and both on supraorbital furrows; supraorbital furrows very deep and wide, parallel-sided at middle of frons, then strongly divergent afterwards, ended to posterior supraorbital pores; supraorbital carinae distinct; frontoclypeal sulcus indistinct; eyes completely disappeared; genae well developed, subparallel-sided, frontal portion strongly broadened laterally; neck constriction distinct, beginning at the level of hind supraorbital pores; labrum conspicuously transverse, distinctly narrower than clypeus excluding clypeal wings, 6 -setose, and ciliate on both sides, slightly emarginated at anterior margin; mandible short and stout, right mandible with a distinct median tooth; palps ensiform, glabrous, apical segments of both maxillary and labial palps longer than penultimate ones respectively; labial palpomere 2 bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pairs situated beneath mental tooth, the other at base near lateral margin; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, extended to scutellum; antennomeres 1 unisetose at subapex, antennomere 2 asetose, pubescent from antennomere 3; antennomeres 2 and 3 subequal in length; antennomeres 5���10 subcylindrical, somehow flat, evidently longer than wide. Pronotum peltate in form, much wider than head; disc smooth, moderately convex; slightly wider than long, PL/PW= 0.96���0.99, mean 0.97; apex deeply concave, beaded in median portion, fore angles strongly protruding, sharper than in G. cavicola; widest at about middle, gently and gradually narrowed forwards to apex, and strongly contracted towards basal angles; side with three conspicuous teethed projections before hind angle; hind angle obtuse; two lateral setiferous pores present, anterior seta at about apical quarter, posterior one at about basal third, just before anterior lateral projection; marginal channel wide and uneven, narrower before anterior seta, wider between anterior and posterior lateral setae, ended before posterior seta; base unbordered; median line clear and deep; basal transverse impression deep and wide, rugose, and faintly carinated posteriorly; anterior transversal line free from median line, very close to anterior margin. Peduncle short, base of scutellum visible and rounded. Elytra elongate ovate, wider than pronotum, EW/PW= 1.34���1.35, mean 1.35, much longer than wide, EL/ EW= 1.76���1.79, mean 1.78; strongly convex; widest at about middle, gently contracted anteriad and posteriad, base finely bisinuate; shoulders broadly obtuse, with two large serrated projections; apex pointed; side not parallelsided, distinctly crenulated from base to apical quarter; elytral striae punctate-striate, striae deep, intervals strongly convex; stria 1 with a small blunt tubercle at the base, just before scutellar pore; intervals 1���4 unbordered at base, other bordered at base; intervals 7 and 8 jointed before base, then jointed to interval 6 at base, all of them carinated near base; interval 3 with five foveolate setiferous pores at 8 /9, 2/7, 1/2, 2/ 3 and 5 / 6 from base respectively; marginal channel with uninterrupted series of small setiferous pores and several large pores which bearing much longer setae; scutellar striae and scutellar pores present; hind wings reduced. Underside of head covered with coarsely rugose and short wrinkles, proepisternum with dense isodiametric punctures, without wrinkle; pro-, meso- and metasterna smooth in middle portion, covered with small and sparse punctures at sides, pro-, meso- and meteepisterna coarsely punctured, epipleuron with a few coarser punctures near base; abdominal ventrites sparsely punctured at sides, each of ventrites IV���VI bearing a pair of paramedial setae, and ventrite VII of both male and female with two pairs of paramedial setae. Legs rather stout and short; fore leg stout, profemur simple, moderately dilated; protibia well developed, quadridentate on outer margin, dorsal carina and sulcus distinct; lateral upper spine elongate ensiform, blunt at apex, much longer and stouter than subapical spur; protarsi short, tarsomere 1 much longer than other, tarsomeres 2���4 flat and transverse; middle and hind legs slender. Male genitalia (Figs. 5���6). Weakly sclerotized; median lobe of aedeagus stouter than in G. cavicola, strongly arcuated, ventral margin unisinuate, gently enlarged towards apex which is very broad; basal orifice small; inner sac armed with longitudinal copulatory piece, apical one longer and broader; parameres similar to G. cavicola. Female genitalia (Fig. 7). Similar to G. cavicola Tian, 2009 though gonocoxite and gonosubcoxite are slightly more slender; bursa copulatrix wide and folded, basally narrower; spermathecal gland very long and large, spermathecal gland duct wide and short. Etymology. The name of this new species is dedicated to Prof. Dr. Louis Deharveng (MNHN, Paris), a wellknown biospeleologist. Remarks. G. deharvengi n. sp. is similar to G. cavicola, but it is easily distinguished from the latter by its depigmented and stouter body, as well as its broader pronotum. Furthermore, elytra interval 3 bearing five dorsal setiferous pores in G. deharvengi n. sp. (while four pores in G. cavicola). Distribution. China (Guangxi: Huangjiang County) (Fig. 1). Known only from Shuiku Cave, the type locality. In this cave are living also two blind trechines: Uenotrechus liboensis Deuve & Tian, 1999 (in Deuve et al., 1999) and Pilosaphaenops whitteni Tian, 2009., Published as part of Tian, Mingyi, 2014, New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae), pp. 355-362 in Zootaxa 3861 (4) on pages 357-361, DOI: 10.11646/zootaxa.3861.4.5, http://zenodo.org/record/225326, {"references":["Tian, M. Y. (2009) New records and new species of cavedwelling trechine beetles from Mulun Nature Reserve, northern Guangxi, China (Insecta: Coleoptera: Carabidae: Trechinae). Subterranean Biology, 7, 69 - 73.","Deuve, T. Tian, M. Y., Ran, J. C. (1999) Trois Caraboidea remarquables de la reserve nationale de Maolan, dans le Guizhou meridional, Chine (Coleoptera, Carabidae et Trechidae). Revue francaise d'Entomologie, New Series, 21, 131 - 138."]}

Tian, Mingyi (2014): New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355-362, DOI: http://dx.doi.org/10.11646/zootaxa.3861.4.5

KEY TO THE SPECIES OF LIBOTRECHUSUÉNO 1. Body stouter; head with only the posterior supraorbital setae present; right mandible tridentate; pronotum with 2 pairs of lateral setae; hind pronotal angles obtuse; elytra with lateral margin distinctly serrate in basal 1/3; 3 rd elytral stria with 2 dorsal pores; scutellar stria present............... L. nishikawai 1′. Body more slender; head with only anterior supraorbital setae present; right mandible bidentate; pronotum with only the anterior pair of lateral setae, the posterior pair absent; hind pronotal angles nearly rectangular; elytra with lateral margins smooth, not serrate; 3 rd elytral stria with only 1 dorsal pore; scutellar stria absent............. L. duanensis, new species, Published as part of Lin, Wei & Tian, Mingyi, 2014, Supplemental Notes on the GenusLibotrechusUéno (Coleoptera: Carabidae: Trechinae), with Description of a New Species from Guangxi, Southern China, pp. 429-433 in The Coleopterists Bulletin 68 (3) on page 430, DOI: 10.1649/072.068.0314

Eutrichodesmus digitatus sp. n. Figs 1–2 Material: Holotype male (SCAU), China, Guangdong Prov., Qingyuan City, Jintan Town, Mi Dong, 0 5.2011, leg. Tian Mingyi (CHIgd 11 -LWX01). Paratypes: 1 male, 2 females (SCAU), same locality, together with holotype. Name: To emphasize the gonopod telopodite having a conspicuous, bifid, digitiform, distofemoral process at about midway. Diagnosis: Differs from other congeners by metaterga 2-13 supporting four transverse mixostictic rows of tubercles, and the gonopod telopodite with a conspicuous, bifid, digitiform, distofemoral process (dp) at about midway while the acropodite has several differentiated lobes. Description: Length of adults of both sexes ca 8.0- 8.5 mm, width 1.5-1.8 mm. Holotype ca 8.5 mm long and 1.8 mm wide. Coloration uniformly pallid (Fig. 1). Adults with 20 segments, body subcylindrical, conglobation complete, pattern of volvation typical, starting from segment 5. Head (Fig. 1 A) slightly transverse (wider than high), rather densely pilose, microgranular and microvillose just below antennae and on vertex, with a pair of rounded, paramedian knobs; isthmus between antennal sockets about as wide as antennal socket diameter. Axial suture conspicuous. Antennae (Fig. 1 A) rather short and clavate; antennomere 6 longer than 5 th, both with an evident dorso-apical pit containing a tight group of minute bacilliform sensilla; antennomere 8 with usual four sensory cones apically. Collum subtrapeziform, rather large, broader than head, not covering the head from above; with six transverse rows of round setigerous tubercles (Fig. 1 B). Prozona very finely alveolate, collum and metaterga covered with a cerotegument crust held by abundant microvilli; stricture between pro- and metazona broad and shallow, more finely alveolate-microgranular than prozona. Limbus microcrenulate, partly hidden by nearby abundant microvilli. Metaterga 2-13 with four transverse mixostictic rows of tubercles, following metaterga with more than four transverse rows of tubercles; three frontal rows highest and best expressed in anterior body part (Figs 1 B, D, E); each tubercle crowned with an abraded seta. Paraterga with evident shoulders anteriorly, strongly declivous, broad and usually trilobate laterally, evidently extending down below level of venter; paraterga 2 strongly enlarged, with three lobulations anterolaterally, schism and hyposchism both very small; paraterga 3 and 4 slightly shorter than others, usually bilobate laterally; paraterga 6-14 laterally with three, following paraterga with four, distinct lobulations, somewhat caudolaterally at base with 1 or 2 distinct lobulations. Pore formula normal, ozopores very indistinct, located near top of caudolateral lobulation. Pleurosternal ridges present only in ♂ segments 2 and 3. Epiproct short, also with differentiated tubercles, directed ventrocaudad, with usual four cones below tip. Hypoproct and paraprocts normal (Figs 1 C, D). Sterna usually with a deep, narrow, transverse depression between coxae, but sterna between ♂ coxae 5-7 and 9 much wider. Gonopod aperture transverse-oval, far from reaching lateral sides of segment 7 (Fig. 1 G). Legs rather long and slender, nearly reaching tips of paraterga; femur somewhat longer than tarsus; claw normal, simple, very slightly curved ventrad (Fig. 1 F). Gonopods (Figs 1 H, 2 A, B) relatively complex. Coxae large, microtuberculate and abundantly setose ventrolaterally. Telopodite a little longer than coxite, slender throughout, setose in its basal half, with a conspicuous, bifid, digitiform, distofemoral process (dp) at about midway. Acropodite with several differentiated lobes; seminal groove terminating subapically with a hairpad. Remarks: This pallid species is a typical “doratodesmid”, possibly a troglobite which is somewhat unusual in showing differentiated metatergal tubercles, coupled with rather long and slender legs.

Brachinus mindanaoensis Tian and Deuve new species. (Figs. 2, 6) Holotype. ♂, labeled ���Mindanao���, ���Ex-Musaeo Mniszech���, in MNHN; Paratype: 1 ♀, idem, in MNHN. Length: 13.5 ���15.0 mm; width: 5.2���5.5 mm. Habitus as in Fig. 2. Dark brown on head, pronotum and elytra, but antennomeres, palps, labrum, clypeus, tibiae and tarsi lighter, femora (except for tips) and frontal spots yellow to reddish brown. Microsculptural meshes densely isodiametric on head, pronotum and elytra. Head longer than wide (HL/HW= 1.1), anterior margin of labrum straight, bearing sexsetose, clypeus bisetose, with a few shorter setae aside, glabrous in median portion of clypeus, frons and vertex; eyes moderate for Brachinus; palps moderate, pubescent and subcylindrical; maxillary palpomere 4 normal at apex, labial palpomere 3 slightly dilated near apex, both obliquely truncate at apex; maxillary palpomere 4 much longer than palpomere 3 (1.4 times), labial palpomere 3 as long as 2; labial palpomere 2 plurisetose; ligula robust and wide, adnated to paraglossae posteriorly, both multisetose at apex; mentum without median tooth, basal foveae large, surrounded by numerous setae of mentum and submentum; palpiger bisetose at base. Antennae extending at about middle of elytra, comparative length ratio of antennomeres 1���5 as: 1.0, 0.7, 1.9, 1.3 and 1.1. Pronotum wider than long (PW/PL= 1.1), widest in anterior half, at about 1 / 3 of pronotum from apex, hind margin straight, fore margin widely and evenly emarginated, deeply sinuated laterally before hind angle, hind angle acute; disc moderately convex, surface finely striate, with a few setae located near to basal and apical margins. Elytra elongate-ovate, EL/EW= 8.5 / 5.1, shoulders broad, widening backwards till 1 / 3 of elytra from apex, not parallel-sided, widest at about apical 1 / 3 of elytra; elytral apex with membrane and fringed, depressions and costae well marked, depressions much wider than costae (nearly twice), rather flat, costae faintly convex, rather wide; sparsely pubescent on median portion, denser at apical and basal areas. Pro-episternum slightly tumescent, invisible from dorsal view; whole glabrous and smooth. Fore tibia with subapical spur externally. Male genitalia (Fig. 6): median lobe of aedeagus slender, strongly arcuate, evenly expanded at median portion. Remarks: Brachinus mindanaoensis is similar to B. bigutticeps Chaudoir (Fig. 4) for the body shape, but it differs from the latter with it dark colour, wide pronotum, aedeagus slender in contrast to the lighter body, narrow pronotum and stout aedeagus of B. bigutticeps (Fig. 7). Etymology: The specific name of the new species refers to its type locality. Distribution: The Philippines (Mindanao Island)., Published as part of Tian, Mingyi & Deuve, Thierry, 2013, Three new species of the genus Brachinus Weber, 1801 from the Philippines, with a modified key to species (Coleoptera: Carabidae: Brachininae), pp. 289-294 in Zootaxa 3737 (3) on pages 291-293, DOI: 10.11646/zootaxa.3737.3.7, http://zenodo.org/record/215916

Eutrichodesmus planatus sp. n. Figs 9���11 Material: Holotype male (SCAU), China, Guangxi Zhuang Autonomous Region, Hechi City, Liujia Town, Cave Zhenzhuyan, 22.11. 2011, leg. Tian Mingyi, Liu Weixin, Zheng Yuan & Sun Feifei (CHIgx 11 -LWX07). Name: To emphasize the metaterga with several transverse rows of flattened tubercles. Diagnosis: Differs from congeners by the metaterga behind the collum usually supporting three or four transverse mixostictic rows of tubercles, but the tubercles are distinctly flattened from segment 10 on towards the telson, whereas the gonopod telopodite has a conspicuous, denticulate, lateral, distofemoral process (dp) near midway and a biramous acropodite. Description: All characters as in E. digitatus sp. n., except as follows. Holotype ca 10 mm long and 2.5 mm wide. Coloration uniformly pallid. Conglobation pattern typical of ��� Doratodesmidae ���, but volvation apparently imperfect because of paraterga being too broad and mostly only slightly declivous, leaving small lacunae laterally even when body is maximally enrolled. Antennae rather long and slender (Fig. 9 A). Collum with several irregular transverse rows of round tubercles (Fig. 9 B). Metaterga 2-16 usually with three or four transverse mixostictic rows of tubercles, following metaterga with more than four transverse mixostictic rows of tubercles, first row being highest and best expressed in anterior part of body (Figs 9 B, E), but tubercles distinctly flattened from segment 10 on towards telson (Fig. 9 H). Paraterga slightly declivous, rather broad, paraterga 3-11 directed laterad, following ones subhorizontal, hardly extending below level of venter. Paraterga 2 strongly enlarged, with four lobulations anterolaterally; paraterga 3 and 4 slightly shorter than others, slightly differently lobulate; following paraterga laterally with three or four distinct lobulations, caudolaterally at base with a distinct lobulation. Sterna between ♂ coxae 6, 7 and 9 much wider. Legs rather long and slender, hardly reaching tips of paraterga. Gonopods (Figs 10���11) rather complex. Coxae subquadrate, large, microtuberculate and sparsely setose ventrolaterally. Telopodite much longer than coxite, clearly expanded just distal to a conspicuous, digitiform, simple, lateral, nearly midway, distofemoral process (dp), a mesal tooth (t) near dp base and, still more distally, a clearly biramus acropodite microdenticulate/fringed on both sides; seminal groove terminating in distal third on a clear-cut hairy pad. Remarks: This is another typical ���doratodesmid���, possibly a troglobite., Published as part of Liu, Weixin & Tian, Mingyi, 2013, Four new cavernicolous species of the millipede genus Eutrichodesmus Silvestri, 1910 from southern China (Diplopoda: Polydesmida: Haplodesmidae), pp. 281-291 in Zootaxa 3734 (2) on pages 289-290, DOI: 10.11646/zootaxa.3734.2.11, http://zenodo.org/record/218501

Liu, Weixin, Tian, Mingyi (2013): Four new cavernicolous species of the millipede genus Eutrichodesmus Silvestri, 1910 from southern China (Diplopoda: Polydesmida: Haplodesmidae). Zootaxa 3734 (2): 281-291, DOI: http://dx.doi.org/10.11646/zootaxa.3734.2.11

Eutrichodesmus spinatus sp. n. Figs 6���8 Material: Holotype male (SCAU), China, Hunan Prov., Sidu Town, Sidu Caves, 28.06. 2011. leg. Tian Mingyi, Gao Qi & Sun Feifei (CHIhn 11 -LWX01). Paratypes: 1 male, 2 females (SCAU), same locality, together with holotype. Name: To emphasize the gonopod telopodite carrying a conspicuous, spinigerous, lateral, distofemoral process (dp). Diagnosis: Differs from congeners by the volvation being apparently imperfect because the paraterga are mostly only slightly declivous; the metaterga behind collum with three transverse isostictic rows of large rounded tubercles; the gonopod telopodite with a conspicuous, spinigerous, lateral, distofemoral process (dp) at about midway while the acropodite is simple and very strongly curved. Description: All characters as in E. digitatus sp. n., except as follows. Length of adults of both sexes ca 6.5 -8.0 mm, width 1.2-1.5 mm. Holotype ca 7.5 mm long and 1.5 mm wide. Coloration uniformly pallid (Fig. 6 A). Conglobation pattern typical of ��� Doratodesmidae ���, but volvation apparently imperfect because of paraterga mostly being only slightly declivous, leaving small lacunae laterally even when body is maximally enrolled. Antennae (Fig. 3 C) rather short and stout. Collum with five regular transverse rows of large round tubercles (Fig. 6 B). Limbus not microcrenulate (Fig. 6 D). Dorsum behind collum with three transverse isostictic rows of large rounded tubercles, each row with about 10 tubercles, but dorsum only with few setae. Paratergal surface coarse, without tubercles; paraterga slightly declivous, broad, 3 - or 4 -lobate laterally, nearly parallel to level of venter, caudolaterally at base with one distinct impression, thus somewhat interrupting the contour of a convex dorsum (Figs 6 A-B). Paraterga 2 strongly enlarged, with three lobulations anterolaterally. Paraterga 3 and 4 slightly shorter than others, each with three lobulations anterolaterally. Pleurosternal ridges wanting. Sterna between ♂ coxae 6, 7 and 9 much wider. Legs rather long and relatively slender, reaching tips of paraterga; femora and tarsi longest, subequal in length; some setae with microdenticulations (Fig. 6 E). Gonopods (Figs 3 F-H) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally. Telopodite longer than coxite, slender throughout, setose in its basal half, with a conspicuous, spinigerous, lateral, distofemoral process (dp) at about midway. Acropodite very strongly curved ventrad; seminal groove terminating subapically with a hairpad. Remarks: This is a typical ���doratodesmid���, possibly a troglobite., Published as part of Liu, Weixin & Tian, Mingyi, 2013, Four new cavernicolous species of the millipede genus Eutrichodesmus Silvestri, 1910 from southern China (Diplopoda: Polydesmida: Haplodesmidae), pp. 281-291 in Zootaxa 3734 (2) on pages 285-287, DOI: 10.11646/zootaxa.3734.2.11, http://zenodo.org/record/218501

Eutrichodesmus similis Golovatch, Geoffroy, Mauri��s & VandenSpiegel, 2009 Material: 3 ♂, 2 ♀ (IZAS), 10 ♂, 6 ♀, 7 juv. (SCAU), China, Guangxi, Huanjiang County, Mulun Town, Xia Dong. 0 3.11. 2009, leg. Louis Deharveng, Anne Bedos & Tain Mingyi (CHIgx09-074). 3 ♂, 2 ♀, 2 juv. (SCAU), same Town, Number 3 Dong. 26.07. 2008, leg. Lin Wei & Xue Zhihong (CHIgx08-LWX02). Remarks: The above material is in good agreement with the original description by Golovatch et al. (2009). This species has originally been recorded from two caves (Gui Dong 2 and Shenlong Dong) in Huanjiang County, Guangxi, China (Golovatch et al. 2009), but Shenlong Dong is actually the cave Xialan Dong away from Gui Dong 2 about 40 km. The new samples come from two other caves which are away from Gui Dong 2 about 4 km., Published as part of Liu, Weixin & Tian, Mingyi, 2013, Four new cavernicolous species of the millipede genus Eutrichodesmus Silvestri, 1910 from southern China (Diplopoda: Polydesmida: Haplodesmidae), pp. 281-291 in Zootaxa 3734 (2) on page 290, DOI: 10.11646/zootaxa.3734.2.11, http://zenodo.org/record/218501

Tian, Mingyi, Deuve, Thierry (2013): Three new species of the genus Brachinus Weber, 1801 from the Philippines, with a modified key to species (Coleoptera: Carabidae: Brachininae). Zootaxa 3737 (3): 289-294, DOI: http://dx.doi.org/10.11646/zootaxa.3737.3.7

Key to the Philippine species of Brachinus (modified from Tian & Deuve, 2007) 1 Bicolor, head and pronotum yellow, elytra dark brown, body smaller (5.5 mm in length)............ B. luzonicus Chaudoir - Unicolor, head, pronotum and elytra dark brown, body larger (more than 10.0 mm in length).......................... 2 2 Head with yellow or red spots between eyes, mentum with deep basal pits or furrows, legs with femora dark brown at apex...................................................................................................... 3 - Head without spot between eyes, mentum without basal pits or furrows, legs with entire femora yellowish..................................................................................................... B. piceus Chaudoir 3 Light brown, spots between eyes large and yellow, frontal impressions convergent backwards, mentum with two basal furrows, but without fovea, elytral costae obscure........................................... B. philippinensis Tian & Deuve - Dark brown, spots between eyes small and dark brown, frontal impressions paralleled backwards, mentum with two basal well-defined foveae, elytral costae more or less distinct....................................................... 4 4 Labial palpiger unisetose............................................................... B. palawanensis n. sp. - Labial palpiger bisetose................................................................................ 5 5 Apical palps of mouthparts dilated, antennae longer, extending over middle of elytra......... B. solidipalpis Tian & Deuve - Apical palps of mouthparts normal, antennae shorter, extending at about middle of elytra........................... 6 6 Elytra broad, with shoulders square......................................................... B. grootaerti n. sp. - Elytra slender, with shoulders broad.................................................... B. mindanaoensis n. sp., Published as part of Tian, Mingyi & Deuve, Thierry, 2013, Three new species of the genus Brachinus Weber, 1801 from the Philippines, with a modified key to species (Coleoptera: Carabidae: Brachininae), pp. 289-294 in Zootaxa 3737 (3) on pages 293-294, DOI: 10.11646/zootaxa.3737.3.7, http://zenodo.org/record/215916

Brachinus grootaerti Tian and Deuve new species. (Figs. 1, 5) Holotype. ♂, labeled ���Coll. I. R. Sc. N. B, the Philippines, C. Palawan, 5 km E. of Napsan, cleared primary forest, 0 9.42. 256 N/118.31.034 E, 1���2. Dec. 2007, J. H. Lourens leg.��� and ���Coll. IRSNB, Achat J. H. Lourens, I. G.: 30.852 ���, in RISNB, Brusssels. Length: 13.0 mm; Width: 5.4 mm. Habitus as in Fig. 1. Medium sized; head, pronotum, elytra, tips of femora and tibiae black; tarsi, antennomeres 1���4 and ventral side of body dark brown; frontal spots, palps and other antennomeres brown; head and pronotum finely striate, with minute punctures. Head, pronotum and elytra with isodiametric microsculptural meshes. Head longer than wide (HL/HW= 1.02), with sparse, long setae; anterior margin of labrum straight, bearing sexsetose, clypeus with two long setae and a few short ones at base, glabrous medially; frons and vertex glabrous, eyes large and prominent; palps pubescent and subcylindrical; maxillary palpomere 4 longer than 3; labial palpomere 3 slightly longer than 2; ligula robust and wide, adnated to paraglossae posteriorly, both ligula and paraglossae multisetose at apex; mentum without median tooth, basal foveae large, rounded, smooth, surrounded by numerous mental and submental setae; both mentum and submentum multisetose; palpiger with two setae at base. Antennae extending at about middle of elytra; comparative length ratio of antennomeres 1���5 as 1, 0.4, 1.6, 1.1 and 0.9; antennomere 3 slightly longer than antennomeres 1 and 2 combined, antennomere 4 longer than 5. Pronotum longer than wide, PW/PL= 1.09, widest at about apical one fourth, fore margin with distinct median emargination, hind margin straight, lateral margin strongly sinuate before hind angle, hind angle acute and sharp; pronotal disc convex, weakly punctate, faintly wrinkled, with lateral setae at about middle and frontal portion. Elytra elongate ovate, EL/EW= 1.50, widest in the middle, pubescent; shoulders wide and square, not parallelsided; apex membrane and fringed; both costae and depressions faint, depressions wider than costae, rather flat, costae faintly convex. Pro-episternum slightly tumescent, invisible from dorsal view, sparsely setose. Fore tibia with subapical spur externally. Male genitalia (Fig. 5): median lobe of aedeagus slender, distinctly arcuate in middle, preapical portion sinuate, expanded at apex; apical portion broad, apical lamella shorter, wider than long, truncate at tip. Female unknown Remarks: Brachinus grootaerti n. sp. is the first species of the genus from Palawan Island. It is quite similar to B. philippinensis Tian & Deuve, but pronotum narrower, elytra broader, with shoulders distinctly square (broad in B. philippinensis); It also likes B. bigutticeps Chaudoir (Fig. 4) in appearance, but body is stouter, elytra and pronotum wider, and pro-episternum invisible from above (slightly visible from above in B. bigutticeps). Etymology: The name of this new species is in honor of Dr. Patrick Grootaert, head of Department of Entomology, IRSNB. Distribution: The Philippines (Palawan Island)., Published as part of Tian, Mingyi & Deuve, Thierry, 2013, Three new species of the genus Brachinus Weber, 1801 from the Philippines, with a modified key to species (Coleoptera: Carabidae: Brachininae), pp. 289-294 in Zootaxa 3737 (3) on pages 290-291, DOI: 10.11646/zootaxa.3737.3.7, http://zenodo.org/record/215916

Brachinus palawanensis Tian and Deuve new species. (Fig. 3) Holotype. ♀, labeled ���Coll. I. R. Sc. N. B, the Philippines, C. Palawan, 3 km N. of Nagtabon, dry coastal forest, 0 9.57. 0 15 N /118.39.275 E, 220 m, 12 Dec. 2007, J. H. Lourens leg.���, in RISNB, Brussels. Length: 12.5 mm; Width: 5.5 mm. Medium sized; head, pronotum, elytra, tips of femora and tibiae black; tarsi, antennomeres 1���4 and underside surface dark brown; frontal spots, palps and other antennomeres brown; head and pronotum finely striate and impunctate. Microsculptural meshes more or less isodiametric on head, pronotum and elytra. Head longer than wide (HL/HW= 1.1), covered with sparse setae; anterior margin of labrum straight, bearing sexsetose,, clypeus with two long setae, and a few short ones, glabrous in median portion; frons and vertex glabrous, eyes large and prominent; palps moderate, pubescent and subcylindrical, apical joints slightly dilated near apex; maxillary palpomere 4 longer than 3; labial palpomere 3 as long as 2; ligula robust and wide, adnated to paraglossae posteriorly, both multisetose at apex; mentum without median tooth, basal foveae large, nearly rounded, smooth, surrounded by numerous mental and submental setae; both mentum and submentum multisetose; palpiger unisetose at base. Antennae extending beyond middle of elytra; comparative length ratio of antennomeres 1���5 as: 1.0, 0.4, 2.2, 1.2 and 1.1; antennomere 3 longer than antennomeres 1 and 2 combined, antennomere 4 longer than 5. Pronotum wider than long (PW/PL= 1.1), widest in the apical portion, frontal margin distinctly emarginated in the middle, hind margin almost straight even slightly bisinuate, lateral margin deeply sinuate before hind angle, hind angle acute; disc moderately convex, faintly striate on surface, impunctate, with two marginal setae, one in the middle, the other at the front. Elytra elongate oval, EL/EW= 1.4, pubescent; shoulders broad, gradually widening backwards till middle; apex membranous and fringed; both costae and depressions faint, depressions wider than costae, rather flat, costae faintly convex. Pro-episternum slightly tumescent, invisible from dorsal view; sparsely setose. Fore tibia with subapical spur externally. Male: Unknown. Remarks: This new species is peculiar for its stout body. It differs from Brachinus palawanensis by its broader elytra and unisetosed palpiger. Etymology: The name of this new species refers to its type locality. Distribution: The Philippines (Palawan Island)., Published as part of Tian, Mingyi & Deuve, Thierry, 2013, Three new species of the genus Brachinus Weber, 1801 from the Philippines, with a modified key to species (Coleoptera: Carabidae: Brachininae), pp. 289-294 in Zootaxa 3737 (3) on page 293, DOI: 10.11646/zootaxa.3737.3.7, http://zenodo.org/record/215916

Brachinus solidipalpis Tian & Deuve, new species (Figs. 1, 5, 9���10) Holotype. Male, labeled ���Museum Paris, Manille, Leotaud, 1843 ���, from Manila, the capital city of the Philippines, deposited in MNHN. Paratypes: 4 males, 2 females, same data as holotype, all deposited in MNHN except 1 male in SCAU. Diagnosis: Large sized, black, legs with femur (except apex) darker, mouthparts with palpi stout, mentum with two large pits at base, labial palpiger with three setae; head slightly wider than pronotum, lateral margin of pronotum deeply sinuate before hind angle, hind angle acute, proepisternum faintly visible from above; elytra distinctly carinate. Length: 14.0 mm; width: 5.5 mm. Habitus as in Fig. 1. Dark brown, but antennomeres 5���11, palpi, labrum, frontal impressions, coxae, trochanters, femora except apex, and median portions of lateral sides of abdominal ventrites II���VI reddish brown. Head with a pair of small red spots between eyes. Head slightly longer than wide (HL/LW= 1.01), labrum straight at front margin, sexsetose, clypeus sexsetose, glabrous in median portion, frons and vertex glabrous, eyes moderate for Brachinus; frontal impressions well marked, large and nearly paralleled backwards; palpi stout, pubescent, maxillary palpomere 4 slightly dilated at apex, labial palpomere 3 dilated at apex; maxillary palpomere 4 longer than palpomere 3, labial palpomere 3 subequal to palpomere 2; labial palpomere 2 plurisetose; ligula robust, pubescent at apex, adnated to paraglossae posteriorly; mentum without median tooth, with two large and deep pits at base, rounded by long setae located on mentum and submentum; mentum and submentum plurisetose, labial palpiger with three setae. Antennae extending to middle of elytra, antennomere 3 longer than antennomeres 1 and 2 combined, antennomeres 4 and 5 subequal to each other. Pronotum transverse (PW/PL= 1.2), widest at apical one-fourth, hind margin slightly oblique, lateral margin deeply and suddenly sinuate from about basal one-third, hind angle acutangular, rather sharp; disc moderately convex, faintly wrinkled, with a few setae on basal and apical portion, two setae near margin (Fig. 5). Elytra elongate-ovate (EL/EW= 1.73), nearly parallel-sided, widest at one-third from apex, apex with membrane and fringe, without setae, shoulders slightly prominent; depressions wider than costae, costae convex, slightly carinated; without pubescence on median portion. Proepisternum slightly tumescent, faintly visible from above; glabrous on anterior half, but with several setae near anterior suture, fine punctured on posterior. Fore tibia with subapical spur externally located, fore tarsomeres 1���3 slightly dilated, sponge setae diagnosed symmetric in male. Meso- and meta-sterna and median portions of abdominal ventrites with long setae, but with short setae at sides. Microsculpture meshes densely isodiametric on head, pronotum and elytra. Male genitalia: Median lobe of the aedeagus long and slender, arcuate, apex gently and gradually constricted, blunt; dorsally apical lamella rather long, broad at apex; virga absent (Figs. 9���10). Remarks: This new species is easily recognized from other Brachinus representatives in Philippines. It is also distinguishable within the genus Brachinus in Southeast Asia by the following characters combined: (1) black in coloration; (2) mouthparts with palpi stout; (3) antennomere 4 subequal to 5; (4) proepisternum faintly visible from above; and (5) the shape of aedeagus. Etymology: The specific epithet derives from Latin adjective solidus, a, um (firm, dense, solid) and noun palpus, i, m (the soft palm of the hand, feeler) and refers to the stout palpi of this new species. Distribution: The Philippines (Manila). Known only from the type locality., Published as part of Tian, Mingyi & Deuve, Thierry, 2007, Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines, pp. 15-22 in Zootaxa 1546 on pages 16-18, DOI: 10.5281/zenodo.178011

Brachinus philippinensis Tian & Deuve, new species (Figs. 2, 6) Holotype. Female, labeled “ Philippines, Ch. Semper, Tuga, 21 / 30. 4. 62 ”, “Museum Paris, 1953, Coll. R. Oberthür”, from Tawi-Tawi Island, southwest Philippines, deposited in MNHN. Diagnosis: Large sized, rust brown, legs with femur (except apex) darker, mouthparts with palpi slender, mentum without pit, but deep furrows, labial palpiger bisetose; head as wide as pronotum; lateral margin of pronotum deeply sinuate before hind angle, hind angle acute, proepisternum faintly visible from above; elytra indistinctly carinate. Length: 16.0 mm; width: 6.5 mm. Habitus as in Fig. 2. Rust brown, but frontal impressions, antennomeres 5–11, legs with femur (except apex), coxae, trochanters, and gula yellow, mouthparts palpi, apex of femur, antennomeres 1–4, elytral epipleuron and ventral surface light brown, lateral sides of abdominal ventrites II–VI and whole ventrites VII dark brown; head with a pair of large and yellow spots between eyes. Head longer than wide (HL/HW= 1.12), labrum slightly rounded at front margin, sexsetose, clypeus with shorter additional setae; mandibular scrobe with several setae on upper margin, glabrous on median portion; eyes small, frons and vertex convex, frontal impressions deep and wide, convergent backwards; with several setae along sides of eyes, neck well marked, with coarse transversal wrinkles, glabrous on frons and vertex; ligula broad, pubescent at apex, adnated to paraglossae posteriorly; palpi pubescent, maxillary palpomere 4 slightly dilated, labial palpomere 3 strongly dilated; labial palpomere 2 bisetose; mentum without tooth, with two deep furrows at base, and several setae on median portion, two setae on either side, submentum with a row of long setae; labial palpiger bisetose. Antennae extending to middle of elytra, pubescent from apex of antennomere 2, antennomere 3 longer than antennomeres 1 and 2 combined, antennomere 1 with several long setae. Pronotum (Fig. 6) wider than long (PW/PL= 1.13), widest at about one-sixth from apex, lateral margin deeply sinuate near hind angle, with a pair of lateral setae; basal transverse impression distinct; surface rather glabrous, distinctly wrinkled and punctured at base, median line clear; proepisternum slightly tumescent and faintly visible from above. Elytra elongate-ovate (EL/EW= 1.48), convex, without basal border, sides broad, not paralleled each other, widest at one-third from apex, shoulders slightly prominent, apex truncate, membrane narrow, with fringe of short setae and few sparsely long setae; depressions flat, wide, pubescent (indistinct on median portion), costae slightly convex, but not carinate, glabrous. Fore tibia with subapical spur externally located. Meso- and meta-sterna glabrous, abdominal ventrites densely setose and punctate. Proepisternum glabrous, slightly tumescent, faintly visible from above. Microsculpture meshes densely isodiametric on head, pronotum and elytra. Male: Unknown. Remarks: This new species is similar to Chinese species B. chinensis Chaudoir, but elytral costae not evidently carinate, pronotum deeply sinuated near hind angle, proepisternum slightly tumescent and visible from above, and median portion of elytra without pubescence. Etymology: The specific epithet, philippinensis, is an adjective derived from the type locality of this species, the Philippines. Distribution: The Philippines (Tawi-Tawi Island).

Brachinus piceus Chaudoir (Figs. 3, 7, 11���12) Brachinus piceus Chaudoir, 1876: 53; Andrewes, 1930: 51; Csiki, 1933: 1625. Diagnosis: Medium sized, dark brown, legs yellow, mouthparts with palpi slender, mentum without pit or furrow, labial palpiger unisetose; head as wide as pronotum, lateral margins of pronotum deeply sinuate before hind angles, proepisternum invisible from above; elytra distinctly carinate, costae much narrower than depressions. Length: 10.0 mm; width: 4.0 mm. Habitus as in Fig. 3. Whole body black, except yellowish on antennae, legs and palpi. Head longer than wide (HL/HW= 1.18); labrum straight, sexsetose, clypeus quadrisetose, the additional two setae on inner portion; mandibular scrobe plurisetose, frons and vertex convex, frontal impressions punctured and setose, compound eyes rather small; palpi subcylindrical, pubescent; ligula dilated, plurisetose, adnated to paraglossae, labial palpomere 2 plurisetose, mentum without tooth and pit, with four setae, submentum with three pairs of setae; labial palpiger with one seta. Antennae extending to middle of elytra, antennomere 1 with several long setae, pubescent from antennomere 2; antennomere 3 much longer than the 1 and 2 combined. Pronotum quadrate (Fig. 7), as long as wide, disc moderately convex, lateral margin long and gently sinuate before hind angle, hind angle nearly rectangular, not sharp. Disc wrinkled, sparsely punctured and pubescent. Elytra without shoulders, elongate-ovate (EL/EW= 1.63), sides not paralleled, widest at about apical onethird; depressions much wider than costae, punctures well marked, pubescent; pubescent distinct on base, edge and apex, but indistinct on median portion; costae distinctly carinate, very narrow and glabrous; apex of elytra truncated, with narrow membrane, fringe distinct, without long setae. Fore tibia with subapical spur located externally; fore tarsomeres 1���3 of male slightly dilated, sponge setae diagnosed symmetric. Abdominal ventrites densely pubescent except proepisternum sparsely punctured and shortly setose. Male genitalia: Median lobe of aedeagus bent ventrally, apical lamella long, slightly constricted at subapex, apex broad; virga absent (Figs. 11���12). Female: Unknown. Material studied: Known only from the unique male holotype, labeled ��� piceus Chaudoir, I. Philippines ���, in MNHN. Distribution: The Philippines., Published as part of Tian, Mingyi & Deuve, Thierry, 2007, Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines, pp. 15-22 in Zootaxa 1546 on pages 19-20, DOI: 10.5281/zenodo.178011, {"references":["Chaudoir, M. de. (1876) Monographie des Brachynides. Annales de la Socit Entomologique de Belgique, 19, 11 - 104.","Andrewes, H. E. (1930) Catalogue of Indian Insects, 18, Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp.","Csiki, E. (1933) Carabidae III: Harpalinae VIII. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, part 126, Junk, Berlin, pp. 15991933."]}

Brachinus luzonicus Chaudoir (Figs. 4, 8, 13���14) Brachinus luzonicus Chaudoir, 1876: 68; Andrewes, 1930: 50; Csiki, 1933: 1623. Diagnosis: Small sized, head and pronotum yellow, elytra dark brown, legs yellow; mouthparts with palpi slender, mentum without pit or furrow, labial palpiger unisetose; head slightly wider than pronotum, lateral margin of pronotum shallowly sinuate before hind angle, proepisternum invisible from above; elytra indistinctly carinate, costae as wide as depressions. Length: 5.5 mm; width: 2.6 mm. Habitus as in Fig. 4. Head, pronotum, antennae, ventral surface of thoraces, median portions of abdominal ventrites II���VI, epipleuron of elytra, and palpi yellow to yellowish red, elytra, abdominal ventrites II���VI at sides and VII light dark brown. Head slightly longer than wide (HL/HW= 1.10), frontal impressions shallow; compound eyes quite large, frons flat, vertex slightly convex, neck distinct; frons and vertex smooth and glabrous, neck pubescent; labrum with anterior margin emarginated, sexsetose; clypeus bisetose, impunctate; ligula large, plurisetose, adnate to paraglossae; palpi subcylindrical, middle more dilated than apices; mentum without tooth and setae, with two pairs of setae, submentum with three pairs of setae; labial palpiger unisetose. Antennae long, extending to mid of elytra, pubescent on whole joints, but antennomere 1 with only longer setae; antennomere 3 slightly shorter than antennomeres 1 and 2 combined. Pronotum (Fig. 8) transverse (PW/PL= 1.07), slightly convex, widest at apical one-fourth, lateral margin deeply sinuate before hind angle; hind angle rectangular, somewhat pointed; pubescent throughout, with fine wrinkles or striae; hind impression faint. Elytra convex, elongate-ovate (EL/EW= 1.35), shoulder square, widest at a little behind of middle, pubescent mainly on depressions, costae slightly wider than depressions, convex, but not carinate, slightly setose; apical membrane presented. Fore tibia with subapical spur externally located, fore tarsomeres 1���3 slightly dilated in male, with two rows of spongy-like setae symmetric on ventral surface. Lower surface densely punctured and pubescent, except proepisternum and mesosternum glabrous. Male genitalia: Median lobe of aedeagus stout, gently bent ventrally, apical lamella narrow, apex rounded; virga absent (Figs. 1314). Materials studied: The holotype, a female, labeled luzonicus Chaud., Luzon, Dr. Semper, Thorey and Ex Musaeo Chaudoir; 1 male and 1 female, with labels Philippines, Ch. Semper and Museum Paris, 1952, Coll. R. Oberthr, deposited in MNHN and SCAU, respectively. Distribution: The Philippines (Luzon)., Published as part of Tian, Mingyi & Deuve, Thierry, 2007, Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines, pp. 15-22 in Zootaxa 1546 on pages 20-21, DOI: 10.5281/zenodo.178011, {"references":["Chaudoir, M. de. (1876) Monographie des Brachynides. Annales de la Socit Entomologique de Belgique, 19, 11 - 104.","Andrewes, H. E. (1930) Catalogue of Indian Insects, 18, Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp.","Csiki, E. (1933) Carabidae III: Harpalinae VIII. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, part 126, Junk, Berlin, pp. 15991933."]}

Cybocephalus liui Tian, n. sp. Holotype: ��, Tongbiguan Natural Reserve, Yingjiang County, West Yunnan, 2005 ��VI�� 23, Liu Jingxian leg. Paratypes: 1 �� 2 ��, same data as holotype. All type materials are deposited in the insect collection of South China Agricultural University, Guangzhou. Description Medium��sized, strongly shiny, surface punctured and setose, antennae with club (antennomeres 9���11) well defined and asymmetric; legs slender, fore tibiae with long setae along outer margins, males with metallic sheen on head and pronotum. Length (not including the bent head): 1.4���1.5 mm; width: 1.0 mm. Habiti as in Figs. 1 and 3. Elongate oval, strongly convex, with prothorax and head bent, head concealed dorsally. Body dark brown, antennae, palpi of mouthparts and legs yellow to yellowish brown. Male with head, including base of mandible and anterior median portion of prothorax, bluish metallic sheen. Female similar to male, but without metallic sheen. Head bent, convex, wider than long; frons distinctly protruding anteriorly, truncate at front margin, clypeus small, labrum concealed by clypeus; mandibles stout, sharp at apices, teeth developed, left mandible with a single tooth, right one with two; base of mandibles with several long setae; eyes large, somewhat elliptical, pointed in front but broadly rounded behind; in dorsal view the inner margins of eyes broadly arcuate interiorly while the outer ones slightly convex; apical joints of labial palpi subcylindrical, apical joints of maxillary palpi thick; genal corner pointed (Fig. 2); antennae rather slender, well defined, 11 ��segmented, club strong, asymmetrical; antennomere 3 much longer than others, almost twice as long as antenommere 4; antennomeres 4���7 subequal in length, each longer than antennomere 8; antennomeres 2 to 8 each with a pair of setae, scape asetose; antennomere 8 shortest, longer than width; club segments setose, apical margin of antennomere 11 nearly truncate (Fig. 4). Pronotum strongly transverse, two times wider than length, markedly convex, lateral margins broadly lobed; median portion of anterior margin slightly convex; whole posterior margin broadly convex. Elytra strongly convex, as long as combined width, lateral portions lobed and almost vertical, apical margins broadly rounded; suture line a little convex in lateral view. Scutellum rather large and wide. Legs slender, fore tibia with apex slightly dilated; inner margin slightly sinuate at midlength, outer margin almost straight, subapical margin obliquely truncate, outer angle broadly rounded, apical margin narrow and nearly straight; long setae arranged along the outer margin (Fig. 5). Male genitalia. Structure of male genitalia normal for Cybocephalus, the basal plate (Fig. 6) broadly flat, roundly but distinctly protruded at apex. Penis stout, pointed at apex (Figs. 7���8). Remarks This new taxon is allied to C. chinensis Yu, but is easily separated from the latter by its fore tibia with outer angle broadly rounded, not angulate; the basal plate a little narrower, more protruded at apex, and setose around margins., Published as part of Tian, Mingyi, 2006, Checklist of the genus Cybocephalus Erichson (Coleoptera: Cybocephalidae) of China, with description of a new species from Yunnan Province, pp. 61-68 in Zootaxa 1202 on pages 64-67, DOI: 10.5281/zenodo.172323

Neoorthogonius orientalis Tian & Deuve, new species The following description will serve as a description for both the new genus and new species. Description. Male: length: 9.0 mm. Breadth: 3.5 mm. Small species of Orthogoniini, yellowish brown in coloration, but antennae, palpi of mouthparts, legs, and underside lighter. Moderately shiny, but elytra dull (Fig. 1). Head. Somewhat quadrate, short and wide, HW/HL ¼ 1.13, slightly narrower than pronotum; compound eyes large, prominent, epistomal suture clear, frontal impressions small, frons and vertex flat, faintly rugulose and punctate, neck moderate; clypeus bisetose, mandibles well developed, hooked at apex; paraglossae membranous, not adnate to ligula, extending forward; ligula small and narrow, bisetose at apex; labial palpomere 2 with three setae in the inner side (two normal and one additional), labial palpomere 2 longer than 3; maxillary palpomeres 3 and 4 subequal in length. Antennae (Fig. 2). Moderate in length, reaching beyond base of elytron, pubescent from antennomere 4; antennomeres 5–10 sub-moniliform; antennomere 1 twice as long as antennomere 2, slightly longer than antennomere 11; antennomeres 3 and 4 subequal, antennomere 4 longer than antennomere 5. Pronotum (Fig. 4). Quadrate, narrow, slightly wider than head, PW/PL ¼ 1.3, both fore and hind angles broadly rectangular, fore and hind margins bordered, lateral marginal bead very narrow, almost same wide throughout, but a little wider near hind angle, disc slightly convex, basal foveae distinct, wide, and moderately deep, fore transversal impression short and rather wide, hind impression indistinct. Elytra. Unbordered at base, elongate-ovate, EL/EW ¼ 1.55, surface less convex, humerus quadrate, parallel at sides, slightly contracted in middle portion; apex somewhat obliquely truncate, with outer angle rounded and inner angle nearly rectangular (Fig. 5); striate-punctate, rather deep, intervals convex, even and odd intervals subequal in width, interval 3 with three setiferous pores, basal pore closed to stria 3, middle and apical pores to stria 2. Legs. Moderate, femora strongly dilated, especially hind femora extraordinarily and laterally dilated (Fig. 6); fore, middle and hind femora with 2, 5 and 4 setae at frontal margins, respectively; middle and hind coxae glabrous; fore tibiae stout, with outer apical angles very sharp, outer margin distinctly serrate (Fig. 3); hind tibia with apical spurs short, broadly rounded; tarsomere 5 unisetose in dorsolateral aspect; hind tarsomere 1 as long as 2, tarsomere 3 longer than 4, tarsomere 4 truncate in frontal margin, not emarginate; all claws pectinate. Ventral. The prosternal process unbordered at apex, abdominal ventrites 4–6 with cilia in the middle of hind margins. Abdominal ventrite 7 not emarginate at apex in male. Microsculptures. Indistinct on head and pronotum, isiodiametric meshes on elytra. Male genitalia (Figs. 7–8). Base portion of aedeagus very large, stout, gradually narrowed towards apex; apical lamella small and rather short, with dorsal opening throughout; parameres normal for Orthogonius. Female. Unknown. Remarks. The single specimen is deposited in Bates' collection (belonging to Oberthür's collection), MNHN, Paris together with other Orthogoniini materials but without identification label. It is the smallest species in Orthogoniini, and easy to separate from other members of the tribe by its peculiar shape and the size. Holotype. Male, labeled '' Bombay' ' only, from Maharashtra State, India, deposited in MNHN, Paris. Etymology. This new taxon is named after the type locality, the oriental region. Distribution. Bombay, India., Published as part of Tian, Mingyi & Deuve, Thierry, 2006, Neoorthogonius orientalis, New Genus And New Species Of Orthogoniini (Coleoptera: Caraboidea) From India, pp. 89-94 in The Coleopterists Bulletin 60 (2) on pages 91-93, DOI: 10.1649/753.1, http://zenodo.org/record/4924224