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Avena breviaristata, collected only once (1882) in Algeria and never re-collected since, is a very mysterious grass species because unfortunately even the type specimen got lost 60--80 years ago. Morphological information on this species was thus based on a few published descriptions, which made it difficult, however, to correctly infer the genus affiliation of this morphologically odd species.Avena breviaristatabecame affiliated in the past with various oat-like genera (Avenula,Helictotrichon,Tricholemma). Due to the recent rediscovery of the type specimen at the P herbarium and the opportunity to study this specimen, we report here on the morphological characters ofA. breviaristataunderpinned by meaningful illustrations. They are discussed in comparison with the morphology of representative species of the above-mentioned genera. Uncommon characters of the spikelets (type of disarticulation of the rachilla, lemma structure, lodicules) and to some extent of the inflorescences, leaves and leaf sheaths support the inclusion ofA. breviaristatain the North African genusTricholemma. Considering biogeography,T. breviaristatumfrom the arid Hauts Plateaux in Algeria is a highly xeromorphic counterpart of the mesomorphic speciesT. jahandiezii, which is confined to higher altitudes of the rather humid Moyen Atlas in Morocco. This underlines the status ofTricholemmaas a relic endemic. Our morphological survey supports the classification ofAvenula(onlyA. pubescens) as separate fromHelictotrichons.str. andHelictochloa. Moreover, morphological evidence does not support an origin ofA. pubescensby intergeneric hybridization between the latter genera as hypothesized in some prior studies. Especially the glabrous palea, the special shape of the lodicules and the structure of the awn show no intermediacy. The complicated history of the type collection ofT. breviaristatumand the role of botanical authors are given.

Vertebrate Type Specimens Collected from Singapore: A Digitisation Workflow

The morphology of Clessidromma palmeri Jarzembowski et al. from mid-Cretaceous Burmese amber is revised based on a reexamination of the type specimen. Contrary to previous studies, our observation confirms that Clessidromma has open procoxal cavities. The characters such as platform on the ventral side of head, modified metacoxae, and ridges on head and pronotum are suggested to be diagnostic characters for Clessidromma. Clessidromma zengi Kirejtshuk from the same deposit is transferred into a new genus, as Kirejtomma zengi comb. nov.

Li, Yun, Chen, Chun, Chang, Yanlin (2021): Further notes on the genus Kuzicus Gorochov, 1993 (Orthoptera: Tettigoniidae Meconematinae) with description of one new species from China. Zootaxa 5067 (2): 273-278, DOI: https://doi.org/10.11646/zootaxa.5067.2.9

Chelonarium signatum Dalman, 1824 is known from northern and central South America. The original description of C. signatum was very brief and lacked description of much of the external morphology and the genitalia. We present a redescription of C. signatum based on examination of material from southern Paraguay, including descriptions of the male and female genitalia. Images are presented of the adult habitus and external morphology, along with notes on the reproductive biology and collection in Paraguay. The history of the type specimen is traced, and photographs of the specimen and its labels are provided. A lectotype is designated for the species.

The pygmy marmoset, the smallest of the anthropoid primates, has a broad distribution in Western Amazonia. Recent studies using molecular and morphological data have identified two distinct species separated by the Napo and Solimões-Amazonas rivers. However, reconciling this new biological evidence with current taxonomy, i.e., two subspecies, Cebuella pygmaea pygmaea (Spix, 1823) and Cebuella pygmaea niveiventris (Lönnberg, 1940), was problematic given the uncertainty as to whether Spix’s pygmy marmoset (Cebuella pygmaea pygmaea) was collected north or south of the Napo and Solimões-Amazonas rivers, making it unclear to which of the two newly revealed species the name pygmaea would apply. Here, we present the first molecular data from Spix’s type specimen of Cebuella pygmaea, as well as novel mitochondrial genomes from modern pygmy marmosets sampled near the type locality (Tabatinga) on both sides of the river. With these data, we can confirm the correct names of the two species identified, i.e., C. pygmaea for animals north of the Napo and Solimões-Amazonas rivers and C. niveiventris for animals south of these two rivers. Phylogenetic analyses of the novel genetic data placed into the context of cytochrome b gene sequences from across the range of pygmy marmosets further led us to re-evaluate the geographical distribution for the two Cebuella species. We dated the split of these two species to 2.54 million years ago. We discuss additional, more recent, subdivisions within each lineage, as well as potential contact zones between the two species in the headwaters of these rivers.

Based on type specimen investigation, the status of Helops tristis Rossi, 1790 as the type species of the genus Dendarus Dejean, 1821 is confirmed. Lectotypes are designated for Helops tristis and Pandarus coarcticollis Mulsant, 1854 – both are classified in Dendarus. Pandarus libanicus Desbrochers des Loges, 1881 is recognised as a junior synonym of Dendarus calcaratus Baudi di Selve, 1881. Both taxa share the same locus typicus (Mount Lebanon in Lebanon) and are morphologically distinct from the Italian Dendarus lugens (Mulsant & Rey, 1854). Finally, D. lugens is considered as a synonym of D. (Dendarus) coarcticollis.

The genus Heraclia (Lepidoptera, Noctuidae, Agaristinae) and the species involved in the confused nomenclature and taxonomy of the type species, Phalaena euphemia Stoll, 1781 are investigated. The rediscovery of the type specimen of Noctua geryon Fabricius, 1781, a taxon that had previously been treated as the senior synonym of euphemia, justified the revival of the latter from synonymy with the former. The type material of euphemia could not be traced and a neotype is designated to fix the published name to a physical specimen. In addition to detailed diagnoses and re-descriptions, the revised taxonomy of Heraclia euphemia and Heraclia geryon is provided and the following new synonymies are proposed: Eusemia pallida Walker, 1854 and Eusemia niveosparsa Westwood, 1881 are synonymised with H. geryon and Eusemia nugatrix Westwood, 1881 is synonymised with H. euphemia. Distributional data for each of the species are presented.

Sematophyllum latifolium Brotherus (1911: 362), known only from the type collection, was originally described by Brotherus (1911) based on a collection made by C.C. Hosseus from Mt. Doi Suthep (Doi Sutäp), in northern Thailand. Pollawatn (2008) revised the family Sematophyllaceae s.l. in Thailand but did not see the type specimen of S. latifolium. During the study of Hosseus’s collections, however, we found two duplicates of type specimens of S. latifolium one located in H-BR and one in M. We found that several critical features of S. latifolium, such as the 1) irregular-pinnately branching habit with the erect flagelliform branches (Fig. 1A), 2) stem and branch leaves strongly differentiated (Fig. 1B−F), and 3) brotherelloid type alar cells often divided into larger hyaline cells towards leaf margins (Fig. 1G), were indistinguishable from those in the type material of Wijkia surcularis (Mitten 1859: 112) Crum (1971: 173), a common species found growing from India to Indochina (Gangulee 1980; Tan & Iwatsuki 1993; Tan & Jia 1999; Jia et al. 2005). Thus, we here propose S. latifolium as a new synonym of W. sucularis. In the protologue, Brotherus (1911) did not designate the holotype, therefore, it is necessary to select a lectotype for S. latifolium ((see Art. 9.11 of the Shenzhen Code (Turland et al. 2017)). We designate Hosseus’s collection (Hosseus s.n.) in H-BR (H) as the lectotype of the name S. latifolium.

Two species of Trichostema L. (Ajugoideae, Lamiaceae) from western states need nomenclatural clarification. Trichostema simulatum Jeps. occurs throughout northern California and southern Oregon. It was published by Willis Linn Jepson in his 1925 flora of California citing the type collection as “Klamathon, Copeland 3534, Type”. This is a conflation of the collector's name and the specimen number of the exsiccata West American Plants, in which the type specimen was distributed by C. F. Baker in 1903. Furthermore, an herbarium name, “Trichostema laetum Greene,” appears on most T. simulatum type material. This designation was never published and is a nomen nudum. Trichostema austromontanum F.H.Lewis occurs throughout the mountain ranges of southern California and Baja California. Originally published with two subspecies by Harlan Lewis in 1945, the three taxa have since been determined to be invalidly published because Lewis did not provide a species description, only subspecies descriptions. Herein, we provide a clarification on the type specimens of T. simulatum Jeps. as well as a valid publication of the species name T. austromontanum.

International audience; Reflectance Transforming Imaging (RTI) produces photographs in which the angle of the light can be changed at will, allowing to investigate remotely minute details of the 3D structure of sub-planar objects. Here we apply this technique to the type specimen of Gingkophyllum grassetii, a vegetative shoot with putative ginkgophyte affinities from the early Permian of Lodève (France). Gingkophyllum grassetii, the type species of the genus, was originally described by Saporta in 1875 and has never been illustrated other than by drawings representing only portions of the shoot. Using RTI, we describe and illustrate the specimen and discuss some key features, such as the arrangement of the leaves on the shoot or the presence of abundant secretory structures. Through an historical account, we show how the lack of illustration led to confusion about the morphology of Ginkgophyllum and we discuss the importance of reinvestigating type specimens of Paleozoic ginkgophyte taxa in order to clarify their taxonomy and better understand their evolutionary history. The sharing of RTI files is one example of approaches that can be further developed to improve remote access to specimens.

Simple Summary Elateroidea is one of the large superfamilies in the beetle suborder Polyphaga. Many adpression-type elateroid fossils were insufficiently described, which hinders the interpretation of their systematic position. Here, we figure and re-describe an elateroid fossil, Archaeolus funestus, from the Jurassic of South China. Our observations support that Archaeolus might be a member of the Throscidae family. Abstract The morphology of the Jurassic fossil Archaeolus funestus Lin, 1986, which was previously placed in the extinct click-beetle subfamily Protagrypninae (Coleoptera: Elateridae), is revised based on a re-examination of the type specimen. The validity of Protagrypninae is discussed and further questioned, partly based on the newly observed characters in A. funestus, including the surface sculpture of the mesoventrite. A possible Throscidae affinity of monotypic Archaeolus Lin, 1986, as suggested in a recent study, is further critically reviewed.

Dipluridae represent a small Mygalomorphae family of South American origin, the family includes two subfamilies Diplurinae and Masteriinae although the placement of the latter in Dipluridae is still under debate. The family has a predominantly South American distribution although the genus Masteria L. Koch, 1873 presents an interesting distribution with representatives found in Fiji, Micronesia, New Caledonia, New Guinea and Australia. This genus is diverse at the species level in tropical South America and the Caribbean but no species have been described from Ecuador to date. Ongoing field work as part of the BIO-GEEC Project—a consortium established by several Ecuadorian and German institutions—has resulted in the discovery of several new species from both lowlands and highland habitats in Ecuador. Herein we described seven new species of Masteria from Ecuador: M. jatunsacha n. sp. (male); M. machay n. sp. (female); M. chalupas n. sp. (male); M. papallacta n. sp. (male and female); M. pasochoa n. sp. (male and female); M. lasdamas n. sp. (male); and M. otongachi n. sp. (male). The type species of the genus, Masteria hirsuta L. Koch, 1873 from Fiji, is redescribed and re-illustrated, from the original type specimen.

Acanthocalyx (DC.) Tiegh. (1909: 199) is a small genus in Caprifoliaceae Jussieu (1789: 210), endemic to the Chinese Hengduan Mountains and adjacent montane regions, yet represents the family’s most characteristic alpine herbaceous perennials in the region. With only three species recognized (A. alba, A. delavayi, A. nepalensis), the taxonomy of Acanthocalyx has not been fully disentangled, and this is particularly due to problem related to the typification of A. delavayi (Franch.) Cannon & Cannon (1984: 14).

Archisargidae, an extinct family of the suborder Brachycera, are traditionally attributed to the infraorder Stratiomyomorpha (Oberprieler & Yeates, 2012). The type genus, Archisargus Rohdendorf, 1938, is a small group including only four species from two well-known localities, the Karatau Formation of Kazakhstan and the Jiulongshan Formation of China. Rohdendorf (1938) erected the genus based on the species Archisargus pulcher from the Karatau Formation of Kazakhstan, and assigned it to Stratiomyidae. Hennig (1954) questioned Rohdendorf’s assignment and pointed out that the characters of the Stratiomyidae were absent, but he didn’t propose an appropriate placement for this genus. Rohdendorf (1962) reconsidered the systematic position of the genus and established the new family Archisargidae based on the genus. In fact, the description of the Archisargus was not informative due to the incomplete preservation of the type specimen (only Sc and R were mentioned) (Rohdendorf, 1938). Mostovski (1997) re-described the type specimen of Archisargus pulcher, clarified the status of the genus, and described a second species, Archisargus maximus, which was also from Karatau, Kazakhstan. A decade later, Zhang et al. (2007) described two older species from the Middle Jurassic of Daohugou, China. The Daohugou locality is well-known for its diverse fossil insects, representing an important diversification center of insects during the Mesozoic Era (Ren et al., 2009; Wang et al., 2010, 2012; Gao et al., 2012; Gu et al., 2012; Meng et al., 2017; Ren et al., 2019). Herein, we describe a new species, Archisargus aequinervus sp. nov., from this locality. Additionally, a key to species of the genus is provided.

During a recent survey, Oxygyne specimens were discovered in three lowland evergreen forests on Okinawa Island, Japan. After morphological investigation, we identified the specimens as O. shinzatoi. Intriguingly, the colour of the specimens from one of the localities was significantly different from that of specimens collected from both the type and two of the new localities. Furthermore, morphological analysis also revealed clear differences between specimens collected from the type locality, including the type specimen, and previous descriptions of O. shinzatoi. Therefore, an amended description of O. shinzatoi is provided, based on the type and newly collected specimens. Phylogenetic relationships of Oxygyne from Japan and western Central Africa are also discussed, with consideration of the characteristics of newly discovered colour variation.

The first genetic study of the holotype of the Gansu short-tailed shrew, Blarinella griselda Thomas, 1912, is presented. The mitochondrial analysis demonstrated that the type specimen of B. griselda is close to several recently collected specimens from southern Gansu, northern Sichuan and Shaanxi, which are highly distinct from the two species of Asiatic short-tailed shrews of southern Sichuan, Yunnan, and Vietnam, B. quadraticauda and B. wardi. Our analysis of four nuclear genes supported the placement of B. griselda as sister to B. quadraticauda / B. wardi, with the level of divergence between these two clades corresponding to that among genera of Soricinae. A new generic name, Parablarinella, is proposed for the Gansu short-tailed shrew. Karyotypes of Parablarinella griselda (2n = 49, NFa = 50) and B. quadraticauda (2n = 49, NFa = 62) from southern Gansu are described. The tribal affinities of Blarinellini and Blarinini are discussed.

Reinstatement of Myelophycus caespitosus Kjellman (Ectocarpales s.l., Phaeophyceae) described from western Japan is proposed based on the comparisons of DNA sequences of Japanese Myelophycus specimens, the type specimen of Chordaria simplex and lectotypified specimen of Myelophycus caespitosus. In the genetic analyses using mitochondrial cox1 and cox3, chloroplast atpB, psbA and rbcL DNA sequences, the specimens morphologically referable to M. simplex formed two distinct clades (clade‐1 and clade‐2) supported by high statistical values. Clade‐1 was distributed on the western coast of Japan (Honshu, Shikoku and Kyushu) from the Kii Peninsula to Tsushima and the Pacific coast of Aomori, northeastern Honshu, and clade‐2 on the Pacific coast of central Honshu from Sanriku to the Kii Peninsula. Based on the cox3 DNA sequences and the location of the type locality at Shimoda, Izu Peninsula, clade‐2 was concluded to correspond to true M. simplex. There were no recognizable differences in the representative morphological features (height and diameter of sporophytes and gametophytes, number of cells comprising cortical, subcortical and medullary layers, thickness of plurilocular gametangia, or length and diameter of unilocular zoidangia) between the specimens included in the two taxa. Analyses of the specimens of the two taxa, including sympatric populations on the Kii Peninsula, using a nuclear genetic marker ocm3 did not suggest any genetic exchanges between the two taxa. On the other hand, cox3 gene sequence of the voucher specimen of Myelophycus caespitosus collected from Goto, Kyushu, western Japan housed in the UPS herbarium was included in clade‐1. In conclusion, we propose the reinstatement of M. caespitosus and to lectotypify the specimen in UPS. Korean Myelophycus specimens reported from Cheju Island and Wando were considered to belong to M. caespitosus based on the reported rbcL sequences.

The mineral ‘oboyerite’, first described in 1979 from the Grand Central mine, Tombstone, Cochise County, Arizona, USA, has been re-examined. The type specimen from the Natural History Museum, London and a specimen from the Natural History Museum of Los Angeles County (traceable to S. A Williams, who first described ‘oboyerite’) were analysed in this study. The discreditation of ‘oboyerite’ as a valid mineral species has been approved by the Commission on New Minerals, Nomenclature and Classification of the International Mineralogical Association (Proposal 19-D). Single-crystal X-ray diffraction, powder X-ray diffraction, electron probe microanalysis and scanning electron microscopy were all employed to show that ‘oboyerite’ is formed of at least two distinct phases, including the lead–tellurium oxysalt minerals ottoite and plumbotellurite. During the course of the discreditation, plumbotellurite was confirmed to be identical to the synthetic compound α-Pb2+Te4+O3. Previously, in some mineralogical literature plumbotellurite was described as orthorhombic with no known crystal structure.

A crushed, partial skull of a juvenile tapir was found with isolated post cranial elements and other fauna in Russells Reserve Cave (RRC), Bath County, Virginia. The cranium is undated but more than likely of late Pleistocene age. It possesses a complete deciduous premolar series and first permanent molars. The M2s have not yet erupted but are visible in the crypt. The RRC specimen is slightly older, ontogenetically, than the type specimen of Tapirus excelsus Simpson (= T. vereoensis) which is probably one of the youngest specimens known. Based upon cranial characters, primarily the lack of a dorsal flange on the maxilla, presence of an interparietal bone, anteriorly ascending cheek tooth row, and lack of sagittal crest, the RRC specimen is referred to the extinct species, Tapirus vereoensis. The sizes of the teeth and morphology of P1 and P2 of the RRC specimen are also consistent with an assignment to T. veroensis. Tapirus pinchaque, the modern woolly tapir, is adapted to cold Andean environments; cladistics and paleoecological studies suggest that cold adaptation may have evolved a second time in T. veroensis.

The present paper describes Trichomerium bhatii, a new species in the family Trichomeriaceae (Chaetothyriales, Ascomycota). This taxon was isolated as epiphyte from dead bark of Prunus cerasoides collected from North-Western Himalayan region of India. The identity of isolate was based on asexual-morphs, cultural characteristics and phylogenetic analyses of partial nuclear ribosomal 28S large subunit and complete internal transcribed spacer rDNA sequence data with high statistical support value. The new taxon has turned out to be distinct from other known species in the genus showing 95.26% (624/655 bp) identity with closely related T. gloeosporum MFLUCC10-0087 on the basis of rDNA internal transcribed spacer region. A brief description of asexual-morph of T. bhatii with morphological and molecular data is provided. To the best of our knowledge this is the first report on this genus from India. The type specimen is deposited in the Ajrekar Mycological Herbarium (as voucher culture) and ex type cultures are deposited in Indian Type Culture Collection (ITCC-WDCM 430), New Delhi, and National Fungal Culture Collection of India (NFCCI-WDCM 932).

International audience

Application of the name Eucalyptus obtusiflora DC. (Myrtaceae). Telopea #: #–# (2019). The name E. obtusiflora has been applied to both an eastern Australian species and to a distantly-related species from Western Australia. We have re-examined the literature and type material of E. obtusiflora and conclude that the holotype was most likely collected on the west coast of Western Australia, possibly as part of the Baudin Expedition, and was certainly not collected by Sieber, with whom the type specimen has previously been associated.

The type speciesPlacenticeras warthiKossmat, 1895 described from the Early Cretaceous (Albian) of South India is crucial for understanding the true affinity of the genusHypengonocerasSpath, 1922. The species description was based on a holotype and was influenced by the erroneous and misleading hand sketches of the species published in the literature. We recently inspected the type specimen archived in Geological Survey of India, Kolkata, along with one additional specimen kept in Indian Museum, Kolkata, and redescribe them in the present study.

The holotype of the pinnotherid, Pinnotheres socius Lanchester, 1902, is extant and deposited in the University Museum of Zoology, Cambridge. The taxonomy of this pea crab is treated in this note and the specimen is illustrated by photographs and line drawings of the third maxilliped and pereiopods. Although the ovigerous type specimen is damaged, it is sufficiently intact to identify the specimen and synonymize the species with Nepinnotheres cardii (Bürger, 1895), described from the Philippines.

Enteromiuspallidus was described by Smith in 1841 without a designated type specimen for the species. Herein, we designate a specimen from the Baakens River system as a neotype for E.pallidus and provide a thorough description for this species to facilitate ongoing taxonomic revisions of southern African Enteromius. Enteromiuspallidus can be distinguished from the other minnows in the “goldie barb group” by having an incomplete lateral line, lack of distinct chevron or tubular markings around lateral line pores, absence of a distinct lateral stripe, absence of wavy parallel lines along scale rows and lack of black pigmentation around the borders of the scales. We provide mtDNA COI sequences for the neotype and an additional specimen from the Baakens River as DNA barcodes of types and topotypes are a fundamental requirement for further taxonomic studies.

A worldwide taxonomic revision and the relevant lectotypifications of Zoegea L. (Asteraceae) are presented. The recognized taxa, the relevant synonyms, type details, images of the type specimen, examined specimen lists, and a taxonomic key are provided for this genus. The geographic distribution of all recognized taxa is presented and tabulated. Zoegea leptaurea L. subsp. leptaurea for the floras of Iran and Saudi Arabia and also Z. purpurea Fresen. for the flora of Iraq are reported as new here.

A re-description of the little-known holothurian species Holothuria (Lessonothuria) lineata Ludwig, 1875 is given. It is based on the single recovered type specimen and an individual recently collected on Glorioso Islands, near Madagascar. A key to separate three closely related and commonly confused species, i.e., Holothuria (Lessonothuria) pardalis Selenka, 1867, Holothuria (Lessenothuria) verrucosa Selenka, 1867 and Holothuria (Lessonothuria) insignis Ludwig, 1875, is presented.

The land area of Laos is composed of a large variety of undisturbed habitats, such as high mountainous areas, huge limestone karsts and the lower Mekong Basin. Therefore, Laos is expected to have a high species diversity, especially for the land snails. However, with respect to research on malacology, Laos is probably the least well-researched area for land snail diversity in Indochina (including Laos) over the past few centuries. The handful of species lists have never been systematically revised from the colonial period to the present, so these classifications are outdated. Herein we present the first comprehensive annotated checklist with an up-to-date systematic framework of the land snail fauna in Laos based on both field investigations and literature surveys. This annotated checklist is collectively composed of 231 nominal species (62 ‘prosobranch’ and 169 heterobranches), of which 221 nominal species are illustrated. The type specimens of 143 species from several museum collections and/or 144 species of newly collected specimens are illustrated. There are 58 species recorded as new to the malacofauna of the country, and two new replacement names are proposed asHemiplectalanxangnicaInkhavilay and Panha,nomen novum(Ariophantidae) andChloritiskhammouanensisInkhavilay and Panha,nomen novum(Camaenidae). Four recently described species of the genusAmphidromusfrom Laos, “thakhekensis”, “richgoldbergi”, “attapeuensis” and “phuonglinhae” are synonymized with previously described species. In addition, thirteen nominal species are listed as uncertain records that may or may not occur in Laos. This annotated checklist may inspire malacologists to carry on systematic research in this region.

Polygonatum is widely distributed in the northern hemisphere, especially in eastern Asia. There has been no comprehensive taxonomic study of Taiwanese taxa for some time, and researchers could not agree on a consistent treatment of the genus. Therefore, we revised the genus by literature review and type specimen examination along with comparison of morphology, karyotype and pollen characteristics. Only one species P.arisanense Hayata was recognized in this study. P.chingshuishanianum Ying and P.formosanum (Hayata) Masamune & Simada are regarded as varieties of P.arisanense and are here presented as two new combinations, P.arisanensevar.chingshuishanianum and P.arisanensevar.formosanum.

Species delimitation remains a challenge worldwide, but especially in biodiversity hotspots such as the Amazon. Here, we use an integrative taxonomic approach that combines data from morphology, phylogenomics, and leaf spectroscopy to clarify the species limits within the Protium heptaphyllum species complex, which includes subsp. cordatum, subsp. heptaphyllum, and subsp. ulei. Molecular phylogeny indicates that populations of subsp. cordatum do not belong to the P. heptaphyllum clade, while morphology and near-infrared spectroscopy data provide additional support for the recognition of a separate taxon. Protium cordatum (Burseraceae) is reinstated at species rank and described in detail. As circumscribed here, P. cordatum is endemic to white-sand savannas located in the Faro and Tucuruí Districts, Pará State, Brazil, whereas P. heptaphyllum is a dominant and widespread plant lineage found in Amazonia, the Cerrado, and the Brazilian Atlantic Forest. We present an identification key to P. cordatum and closely related lineages and a detailed taxonomic description of P. cordatum, including habitat and distribution, a list and images of diagnostic features. This study demonstrates the importance of using multiple tools to characterize and distinguish plant species in highly diverse tropical regions. © 2019 International Association for Plant Taxonomy.

The taxonomic status of Pseudopodaaureola (He & Hu, 2000) is revised based on the re-examination of the type specimen and fresh material collected from the type locality. The cheliceral dentition, with a large denticle field between two anterior and three posterior teeth, the male palps with simple RTA arising distally, narrow, not filiform embolus and sheet-like membranous conductor, and the female epigyne with a visible median septum indicate that the species does not belong to Pseudopoda or to the originally assigned genus, Heteropoda. Based on these characters, the species is here transferred to Gnathopalystes. The male is redescribed and the female is described and illustrated for the first time.