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Acutandra caterinoi Lingafelter & Tishechkin, new species (Figs. 3���4) Description. Male (Figs. 3 a���g). Color generally piceous with some areas including parts of venter, legs, palpi, and antennomeres, dark reddish-brown. Body length (end of elytra to base of mandibles) = 19.7 mm; body width (at humeri) = 6.3 mm. Width of head including eyes slightly narrower than pronotum at anterior angles. Mandibles (Fig. 3 f) relatively short, robust, elongate-triangular, dorsal face with distinct sharp ridge; finely and sparsely punctate, punctures smaller than those of head; apices bifurcate, left mandible with three large teeth along inner curvature, the largest being the basal-most, right mandible with two weak tooth-like projections. Length of mandible shorter than length of head (left mandible = 2.0 mm, right mandible = 1.7 mm). Dorsal surface of head weakly convex with a shallow median inverted T-shaped sulcus. Disc finely, rather densely and evenly punctured; most punctures of head larger, deeper, and denser than those of mandibles and pronotum. Punctures small and sparse across anterior half of hypostoma and gula from between posterior eye margin to just before anterior margin, this area with two deep transverse sulci, spaces between outer ends of these sulci and posterior eye margins flat, with small dense punctures. Anterolateral region of gula merging with gena and extending in lobe on either side by less than one-third the length of gula. Clypeus with broad triangular projection with sub-acute apex medially. Eye (Figs. 3 f���g) bean-shaped, about twice as long as wide, weakly protuberant laterally (intraocular distance 5.3 mm). Antenna (Figs. 3 c���d) 11-segmented; 3���10 subequal in length, 3 being the longest; apicoventral projections weak and indistinct; ventral sensory regions pronounced and divided by distinct, median longitudinal carina on segments 3���11. Antennal pubescence sparse, present only in apical areas of most antennomeres. Pronotum (Fig. 3 a) moderately convex, maximum width near anterior angles, equal to elytral width at humeri, then narrowing posteriorly. Pronotal length = 4.7 mm; pronotal width = 6.5 mm. Lateral margins complete and demarcated, dorsally visible and continuous through posterior margin. Posterolateral regions not projecting. Pronotal margin not delineated on anterior edge of pronotum. Pronotal disc with distinct anteromedial and medial longitudinal impression and fine, shallow, relatively dense punctures, only slightly smaller than those on head and elytra. Elytra (Fig. 3 a) parallel-sided to near apex and then rounded to suture. Elytron 2.11 X longer than wide; elytral length = 13.3 mm; elytral width = 6.3 mm. Rather dense, fine, shallow punctures, similar in size to those of head, evenly covering elytral surface. Margin delineated and visible from dorsal view except for small area around slightly projecting humeri, anterior two-thirds of lateral sides distinctly sulcate next to margin. Elytral disc with two weak and low, but distinct, longitudinal ridges in anterior two-thirds, inner one being slightly shorter than outer one, especially posteriorly. Prosternum (Fig. 3 b) sparsely punctate, glabrous, with declivous rounded intercoxal process slightly extending beyond procoxae; dorsolateral extensions closing external halves of the procoxal cavities posteriorly. Prosternal intercoxal process subequal in width to mesosternal intercoxal process (Fig. 3 b). Mesosternum asetose. Metasternum almost half length of elytron and slightly shorter than abdomen, glabrous, with small sparse punctures being larger and denser anteriorly and laterally. Metepisternum (Fig. 3 g) glabrous, punctation similar to that of adjacent metasternum. Punctation of ventrites (Fig. 3 b) similar to that of metasternum, slightly denser, especially at sides. Ventrite 5 length subequal to that of ventrite 4, straight at apex. Femora (Figs. 3b, g) nearly glabrous and very sparsely, shallowly punctate; each subequal to and 2.0���2.2X the greatest width of the associated tibia. Tibiae sparsely, shallowly punctate and nearly glabrous with setae primarily on ventral margins and apex; about 2.5 times as wide at apex as base; with complete, or nearly complete straight middle carina on anterior faces; apices each with two ventral spurs and one dorsal spine. Each tarsus approximately three-quarters length of its associated tibia; tarsomere 5 slightly longer than 1���4 combined on each tarsus. Female (Figs. 4 a���g) with proportions, coloration, punctation, and pubescence similar to that of male with differences noted as follows: overall size, slightly longer and broader than male (body length = 19.2���23.7 mm; body width = 6.3���7.5 mm). Head less robust; width including eyes slightly narrower than that of pronotum at anterior angles. Eyes less projecting than in male and slightly smaller. Median sulcus not T-shaped, just longitudinal. Mandible (Fig. 4 f) shaped as in male, broader basally, with weak and indistinct dorsal ridge; slightly shorter than in male (left mandible = 1.3���2.0 mm; right mandible = 1.3���1.7 mm); both mandibles with three teeth, much shorter, broader and blunter than in males. Lateral margins of pronotum (Fig. 4 a) rounded, widest point approximately at anterior third; pronotum dimensions similar to those of male (length 4.0���5.0 mm, width 5.7���7.0 mm), punctation distinctly denser, longitudinal median impression much less pronounced. Anterior gular/ hypostoma area (Fig. 4 b) without transverse sulci, its surface somewhat irregularly, densely punctate. Punctation of metasternite and abdominal ventrites much sparser than male, especially laterally. Terminal ventrite somewhat narrower and longer than in male. Ovipositor (Fig. 4 e) highly sclerotized with two dorsally projecting teeth, basal one wider, bicuspidate. Discussion. This new species belongs in the genus Acutandra due to the open procoxal cavities, sinuate anterior margin of the pronotum, acute apex of the labrum in both sexes, and modest mandibles that are not falciform in both sexes. Acutandra contains five species and is restricted to South America (Santos-Silva and Martins, 2010). No species of Acutandra have been recorded from Ecuador. In Santos-Silva and Martins���s (2010) key to species of Acutandra, A. caterinoi possesses the following character states taking it to couplet 4 with the species A. murrayi (Lameere) from east and south Brazil and A. araucana (Bosq) from Chile and Argentina: ventral antennal carinae present dividing the sensory regions of each antennomere; eyes small and not prominent; antennomeres III and IV approximately equal in length; dorsal sensory region of antennomere XI small in both sexes, but well-delimited. It is most similar to A. murrayi in having antennomeres 3 and 4 subequal in length and a distinctly delimited dorsal sensory region on antennomere XI. A new couplet is presented (4a) and the original couplet 4 is translated (4b) from Santos-Silva & Martins (2010): 4a. Posterolateral margin of pronotum weakly produced, lateral margin unevenly curved at junction with posterior margin; pronotum without or with very weak medial longitudinal impression; color most often reddish brown (Brazil, Chile, Argentina)...................................................................................................... 4b 4a���. Posterolateral margin of pronotum not produced, lateral margin evenly curved to posterior margin; pronotum with distinct anteromedial and longitudinal impressions; color piceous over dorsal surface (Ecuador)................................................................................. Acutandra caterinoi Lingafelter & Tishechkin, new species 4b Antennomeres 3 & 4 subequal in length; dorsal sensory area of antennomere XI well delimited and deep. (East and south Brazil).......................................................................... Acutandra murrayi (Lameere) 4b���. Antennomere 3 longer than 4; dorsal sensory area of antennomere XI poorly delimited and shallow. (Chile, Argentina)................................................................................... Acutandra araucana (Bosq) Etymology. We dedicate this species to Michael S. Caterino, a colleague and friend, co-collector of the type series, in recognition of his contributions to the study of the Neotropical beetles. Type material. Holotype male: " ECUADOR: Pichincha, Res. El Pahuma. 1,900���2,100m. 0.0264��N 78.6344��W. Hand colld. 28.v.-1.vi.2011. AT1324. M.S. Caterino & A.K. Tishechkin " (USNM). Paratypes: one female with same data as holotype (CUAC), four females with the same locality and collector data, but collected on 28���31 May 2011 at 2500 m under bark/in rotten wood (PUCE, SWLC, USFQ, USNM), Published as part of Lingafelter, Steven W. & Tishechkin, Alexey K., 2017, Two new species of Parandrinae (Coleoptera: Cerambycidae) in the genera Parandra and Acutandra from South America, pp. 401-410 in Zootaxa 4272 (3) on pages 406-410, DOI: 10.11646/zootaxa.4272.3.5, http://zenodo.org/record/892474, {"references":["Santos-Silva, A. & Martins, U. R. (2010) Subfamilia Parandrinae. In: Martins, U. R. (Org.), Cerambycidae Sul-americanos (Coleoptera) Taxonomia. Uol. 11. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 5 - 79."]}

Parandra (Tavandra) santossilvai Lingafelter & Tishechkin, new species (Figs. 1���2) Description. Male (Figs. 1 a���g). Color generally piceous with some areas including vertex of head, most of venter, tarsi, palpi, and antennomeres, dark reddish-brown. Body length (end of elytra to base of mandibles) = 23.5 mm; body width (at humeri) = 6.5 mm. Width of head including eyes slightly broader than pronotum at anterior angles. Mandibles (Fig. 1 f) pronounced and sickle-shaped; finely and sparsely punctate, punctures smaller than those on disc of head; apices bifurcate, with three or four small teeth along inner curvature, the largest at approximately apical third. Length of mandible greater than length of head (left mandible = 4.3 mm, right mandible = 3.8 mm). Dorsal surface of head mostly flat with a shallow median longitudinal sulcus. Finely and sparsely punctured along anterior margin, punctation becoming denser and deeper at sides, around eye lobes, and along posterior margin and occiput; most punctures of head larger, deeper, and denser than those of mandibles and pronotum. Punctures very large, deep, and contiguous across anterior half of hypostoma and gula from between posterior eye margin to just before anterior eye margin; sides of punctate region of hypostoma demarcated by distinct, raised carina. Anterolateral region of gula merging with gena and extending as a lobe on either side by almost half the length of gula. Clypeus with abrupt, narrow, truncate projection medially. Eye (Figs. 1 f���g) pyriform, less than twice as long as wide, strongly protuberant laterally (intraocular distance 6.7 mm) with posterior ocular edge very distinct. Antenna (Figs. 1 c���d) 11-segmented; 3���10 subequal in length; 5���10 with apicoventral projections (largest on 6���8); ventral sensory regions pronounced and divided by pronounced, median longitudinal carina on segments 3���11. Antennal pubescence sparse, longer and more concentrated at apex of most antennomeres. Pronotum (Fig. 1 a) somewhat flattened, maximum width at anterior two-thirds, equal to elytral width at humeri but slightly narrower than head width at eyes, then narrowing markedly on posterior one-third. Pronotal length = 5.0 mm; pronotal width = 6.5 mm. Lateral margins complete and demarcated, but not visible from dorsal view for small region at approximately anterior one-third. Lateral margin continuous with posterior margin in even curvature around posterolateral regions which are not projecting. Pronotal margin not well delineated anteriorly at middle. Pronotal disc with very fine, shallow, widely separate punctures, much smaller than those on head and subequal in size and distribution to those on elytra. Elytra (Fig. 1 a) parallel-sided to near apex and then rounded to suture. Elytron 2.26 X longer than wide; elytral length = 14.7 mm; elytral width = 6.5 mm. Sparse, fine, shallow punctures, similar in size and distribution to those of pronotum, scattered throughout surface. Margin delineated and visible from dorsal view except for small area around slightly projecting humeri. Prosternum (Fig. 1 b) very sparsely punctate, glabrous, with protuberant but rounded intercoxal process extending beyond procoxae, and with dorsolateral extensions that completely close the procoxal cavities posteriorly. Prosternal intercoxal process 1.5X wider than mesosternal intercoxal process (Fig. 1 b). Mesosternum with moderate, fine pubescence anterior to mesocoxae. Metasternum almost half length of elytron and slightly shorter than abdomen, glabrous except for anterolateral portion adjacent to mesocoxae and metepisterna. Metepisternum (Fig. 1 g) with scattered, appressed setae throughout, denser than on remainder of venter; sparsely, finely punctate, but much more densely than on adjacent metasternum. Ventrites (Fig. 1 b) 1���4 heavily punctate at sides and ventrite 5 punctate throughout. Ventrite 5, 1.3X length of ventrite 4, broadly rounded at apex. Parameres broad, separate, apically rounded and pubescent with golden setae. Femora (Figs. 1 b, g) nearly glabrous and very sparsely, shallowly punctate; each slightly shorter than and 1.75X the greatest width of the associated tibia. Tibiae sparsely, shallowly punctate and nearly glabrous with setae primarily on ventral margin and apex; over three times as wide at apex as at base; with complete, or nearly complete median carina on anterior face (nearly straight on pro- and metatibia; sinuate on mesotibia); apices each with two ventral spurs and one dorsal spine. Each tarsus approximately the length of its associated tibia; tarsomere 5 slightly longer than 1���4 combined on each tarsus. Female (Figs. 2 a���g) with proportions, coloration, punctation, and pubescence similar to male with differences noted as follows: overall size slightly shorter but as broad as male (body length = 22.2 mm; body width = 6.5 mm). Head less robust, width including eyes slightly narrower than that of pronotum at anterior angles. Punctation less pronounced around eyes than in male (Fig. 2 f). Eyes less projecting than in male and slightly smaller. Mandible (Fig. 2 f) subtriangular, not sickle-shaped as in male; much shorter than in male (left mandible = 2.3 mm; right mandible = 2.0 mm); equal to two thirds length of head; more coarsely punctured than in male, punctures approximately the same size and distribution as on head; apex not bifurcate as in male; with three adjacent teeth at apical third. Pronotum (Fig. 2 a) similar to male, slightly shorter (4.7 mm) and narrower (6.3 mm), with an indistinct, oval depression on either side of disc just behind midline. Punctation of gula/hypostoma (Fig. 2 b) much reduced compared to male, with punctures smaller and more sparsely distributed. Gular-genal margins less pronounced and projecting anteriorly by less than one-fourth length of punctate gular region. Terminal ventrite and tergite densely fringed with short, golden pubescence. Ovipositor (Fig. 2 e) highly sclerotized with three dorsally projecting teeth increasing in length posteriorly; lateral face rugose. Discussion. This new species is in the genus Parandra based on the distinctly closed procoxal cavities (Santos-Silva & Martins, 2010). The strongly projecting eyes and hypostoma with distinct raised, lateral carinae place this species in the subgenus Tavandra, although it superficially resembles a species in the subgenus Hesperandra, also from Bolivia (Yungas de Totora, 2100 m), Parandra (Hesperandra) conspicua (Tippmann, 1960) (Fig. 5). Santos-Silva and Martins (2010) characterize the subgenus Hesperandra as lacking a hypostomal carina, thus distinguishing it from the subgenus Tavandra. Using the key of Santos-Silva and Martins (2010), the following character states place this new species in couplet 5 with Parandra (Tavandra) brevicollis Lameere and Parandra (Tavandra) villei Lameere: metepisternum distinctly pubescent; metasternum with pubescence more sparse than on metepisternum and concentrated only around the mesocoxae. It is most similar to P. (T.) villei due to the relatively more slender body, very fine but distinct elytral punctures, and labrum with an acute medial process. To modify the existing key, a new couplet is presented (5a) and the original couplet 5 from Santos-Silva & Martins (2010) is translated as (5b): 5a. Color reddish-brown over most of dorsal surface; anterior genal projections weak, less than one-third length of punctate region of hypostoma........................................................................................ 5b 5a���. Color piceous over most of dorsal surface; anterior genal projections at least one-third length of punctate region of hypostoma........................................ Parandra (Tavandra) santossilvai Lingafelter & Tishechkin, new species 5b. Body robust; elytral punctation extremely fine and inconspicuous microscopic; apex of labrum truncate (Ecuador)............................................................................ Parandra (Tavandra) brevicollis Lameere 5b���. Body not robust; elytral punctation very fine with distinct pores; apex of labrum acute (Venezuela, Colombia, Ecuador, Peru)........................................................................ Parandra (T avandra) villei Lameere Etymology. It is our pleasure to name this species in honor of Antonio Santos-Silva, among the most productive and generous cerambycid workers. In particular, Antonio has made tremendous contributions toward the knowledge of Parandrinae, and we are pleased to acknowledge his efforts with this patronym. Type material. Holotype male: " Bolivia: Dept. Santa Cruz, Prov. Florida, Vicoquin Area above Achira, rd to Amboro, 18��07'S, 63��47'W, 2000m, 5���6 Feb. 2013. UV/MV lights, Lingafelter, Wappes, Garzon " (USNM). Paratype female: " Bolivia, S. Cruz Dept., Achira area, N. rd to Amboro on Achira ridge, 18��09'S, 63��48'W, Wappes, Bonaso, Lingafelter, Garzon" (ACMT)., Published as part of Lingafelter, Steven W. & Tishechkin, Alexey K., 2017, Two new species of Parandrinae (Coleoptera: Cerambycidae) in the genera Parandra and Acutandra from South America, pp. 401-410 in Zootaxa 4272 (3) on pages 403-406, DOI: 10.11646/zootaxa.4272.3.5, http://zenodo.org/record/892474, {"references":["Santos-Silva, A. & Martins, U. R. (2010) Subfamilia Parandrinae. In: Martins, U. R. (Org.), Cerambycidae Sul-americanos (Coleoptera) Taxonomia. Uol. 11. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 5 - 79."]}

Ecclisister bickhardti (Reichensperger, 1923) Cyclechinus bickhardti Reichensperger 1923: 248. Ecclisister bickhardti: Reichensperger 1935: 203–204; Mazur 1984: 304, 1997: 159, 2011: 116; Helava et al. 1985: 324. Zoobank Identifier. urn:lsid:zoobank.org:act: 9F196B10-F503-405E-B28D-C1AB32A23739. Type Material Examined. Lectotype (designated by Dégallier (1998), Fig. 1a–f). A male mounted on a point along with a small E. burchellii worker and labeled “St. Catarina Blumenau / Eciton burchelli / TYPUS / Cyclechinus bickhardti Reichensp. n. sp. / Coll. Reichensperger ” (FIMAK). Paralectotypes (designated by Dégallier (1998)). Two females mounted and labeled as lectotype but with “Cotype” instead of “ TYPUS ” and our “ Paralectotype ” instead of “ Lectotype ” labels (FIMAK). Additional Material Examined. BRAZIL: Santa Catarina. One specimen from same locality as of the type series members, 8.vi.[year unknown], with E. burchellii (NMMW)., Published as part of Tishechkin, Alexey K., Kronauer, Daniel J. C. & Beeren, Christoph Von, 2017, Taxonomic Review and Natural History Notes of the Army Ant-Associated Beetle Genus Ecclisister Reichensperger (Coleoptera: Histeridae: Haeteriinae), pp. 279-288 in The Coleopterists Bulletin 71 (2) on page 281, DOI: 10.1649/0010-065X-71.2.279, http://zenodo.org/record/5460674, {"references":["Reichensperger, A. 1923. Neue sudamerikanische Histeriden als Gaste von Wanderameisen und Termiten. I. Systematischer Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 13: 313 - 36.","Reichensperger, A. 1935. Beitrag zur Kenntnis der Myrmekophilenfauna Brasiliens und Costa Ricas III. (Col. Staphyl. Hist.). Arbeiten uber Morphologische und Taxonomische Entomologie aus Berlin - Dahlem 2: 188 - 218.","Mazur, S. 1984. A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54: 1 - 376.","Mazur, S. 1997. A world catalogue of the Histeridae (Coleoptera: Histeroidea). Genus. International Journal of Invertebrate Taxonomy. Supplement 7: 1 - 373.","Mazur, S. 2011. A Concise Catalogue of the Histeridae (Coleoptera). Warsaw University of Life Sciences SGGW Press, Warsaw, Poland.","Helava, J. V. T., H. F. Howden, and A. J. Ritchie. 1985. A review of the New World genera of the myrmecophilous and termitophilous subfamily Hetaeriinae (Coleoptera: Histeridae). Sociobiology 10: 127 - 386.","Degallier, N. 1998. Coleoptera Histeridae Hetaeriinae: description de nouveaux taxons, designation de lectotypes et notes taxonomiques. Bonner Zoologische Beitrage. 47: 345 - 379."]}

Quantifying the spatio-temporal distribution of arthropods in tropical rainforests represents a first step towards scrutinizing the global distribution of biodiversity on Earth. To date most studies have focused on narrow taxonomic groups or lack a design that allows partitioning of the components of diversity. Here, we consider an exceptionally large dataset (113,952 individuals representing 5,858 species), obtained from the San Lorenzo forest in Panama, where the phylogenetic breadth of arthropod taxa was surveyed using 14 protocols targeting the soil, litter, understory, lower and upper canopy habitats, replicated across seasons in 2003 and 2004. This dataset is used to explore the relative influence of horizontal, vertical and seasonal drivers of arthropod distribution in this forest. We considered arthropod abundance, observed and estimated species richness, additive decomposition of species richness, multiplicative partitioning of species diversity, variation in species composition, species turnover and guild structure as components of diversity. At the scale of our study (2km of distance, 40m in height and 400 days), the effects related to the vertical and seasonal dimensions were most important. Most adult arthropods were collected from the soil/litter or the upper canopy and species richness was highest in the canopy. We compared the distribution of arthropods and trees within our study system. Effects related to the seasonal dimension were stronger for arthropods than for trees. We conclude that: (1) models of beta diversity developed for tropical trees are unlikely to be applicable to tropical arthropods; (2) it is imperative that estimates of global biodiversity derived from mass collecting of arthropods in tropical rainforests embrace the strong vertical and seasonal partitioning observed here; and (3) given the high species turnover observed between seasons, global climate change may have severe consequences for rainforest arthropods.

Strigister Caterino, Tishechkin, and Proudfoot, new genus Type species: Strigister tecolotito Caterino, Tishechkin, and Proudfoot, new species Description. Size: Length 2.3–2.9 mm; width 2.0– 2.7 mm. Body: Body piceous, elongate oval, sides rounded, moderately strongly convex dorsoventrally. Head: Frons flat to weakly convex, with frontal stria complete along inner edge of eye, recurved dorsad, acute at middle; supraorbital stria absent; epistoma weakly depressed, shallowly emarginate apically; labrum about 4X as wide as long, emarginate apically; mandibles strongly bent mediad, with very strongly pointed apices, right mandible with simple, acute tooth at base of incisor edge, left mandible with weakly bifid tooth at base; submentum produced into base of oral cavity; mentum about 3X wider than long, emarginate apically, with numerous long setae; maxillary cardo bare, stipes with several long setae near lateral margin, ultimate labial and maxillary palpomeres narrowed, subulate apically; antennal scape slightly longer than funicle, sides uneven, slightly widened apically, bluntly dentate apicomedially; funicle widening to cupuliform 8 th antennomere; antennal club small, about length of apical 3 funicular antennomeres, increasingly setose toward apex, with single evident, weakly v-shaped annulus. Pronotum: Pronotal sides narrowed slightly from base to apex, disk without obvious gland openings, marginal stria complete along side, interrupted at anterior corner and behind head, submarginal stria complete along lateral and anterior margins; pronotal disk lacking prescutellar impression, variously sculptured. Elytra: Elytral disk with conspicuous ground punctation, secondary punctation denser apically; striae somewhat varied, but with most dorsal striae more or less complete. Prosternum: Prosternal keel truncate to weakly emarginate basally, with weak or no carinal striae basally, middle of keel interrupted by deep transverse incision, anterior to which the keel profile is depressed, with anterior carinal striae diverging from incision; prosternal lobe about two-thirds as long as keel, truncate to weakly emarginate apically, with very fine marginal stria. Mesoventrite: Mesoventral disk about 3X wider than long, anterior margin truncate to weakly produced, marginal stria complete, with varied anterolateral strioles. Metaventrite: Mesometaventral stria complete across front, continued by lateral metaventral stria toward metacoxa; postmesocoxal stria present, recurved anterad toward mesepimeron; metaventral disk weakly to moderately punctate. Abdomen: First abdominal ventrite with single complete lateral stria, disk moderately punctate, posterior margin moderately (females) to strongly (males) emarginate; ventrites 2–5 weakly punctate at sides; propygidium and pygidium very large, similar in midline length, variably punctate, lacking obvious gland openings or striae. Legs: Profemur weakly notched at inner apex; protibia with outer edge weakly rounded, weakly 4–5 dentate, with 2 small, apical spurs; protarsus with simple ventral spines in both sexes; mesotibia weakly curved, with 5–6 prominent marginal spines and a few submarginal spines along anterior surface; metatibia longer, more or less straight, with 4–5 marginal and a few weakly submarginal spines. Male genitalia: Accessory sclerites absent, T8 with deep, narrow basal emargination, sides weakly rounded to apex, with narrow, shallow apical emargination, ventrolateral apodemes weakly produced, widely separated beneath; S8 divided along midline, hinged to T8 near base, apices with few divergent setae, emarginate on inner edges and curved inward, with apical and midline membraneous velum; T9 with rather thick basal apodemes, sclerotized along their lower edges, ventromedial apodemes well-developed, strongly recurved proximad, T9 apex rather narrowly truncate; T10 elongate cordate, weakly desclerotized along midline; S9 elongate, narrow along most of stem base bulbous, apex narrowly T-shaped, with very weak median emargination and strong lateral and apical flanges; tegmen very narrow, elongate, parallel-sided in basal two-thirds, narrowed to weakly bifid apex, lacking ventral tooth or process, but with moderately well-defined median ventral keel; median lobe short, simple, extruded through ventral foramen one-third proximad apical narrowing; basal piece nearly one-half tegmen length. Female genitalia: T8 forming a single plate, apically narrowly desclerotized, with deep, arcuate, basal emargination, ventrolateral edges desclerotized/incised about midway from base to apex; S8 divided along midline with basal baculi narrowly attached at basolateral corners, basally convergent and subparallel in proximal half; S9 small, triangular, articulated with strap-shaped extension from apex of S8; T10 broad, desclerotized along midline; valviferae paddle-shaped, paddles about one-third total length; coxites rather stout, apically weakly tridentate, with distinct, articulated apical stylus; bursa copulatrix mostly membraneous, not strongly expanded, with paired sclerites above attachment of spermatheca; spermatheca weakly sclerotized, approximately spherical, with elongate, weakly spiraled spermathecal gland attached near its base. Diagnosis. This new genus can be easily recognized by several features unusual or unique among New World Exosternini. The single v-shaped annulus of the antennal club (Fig. 1) and the transversely incised prosternal keel (Fig. 2) are unique in Exosternini. The recurved and angulate frontal stria (Figs. 5–6), the truncate base of the prosternal keel (Figs. 7–8), the apically truncate prosternal lobe (Figs. 7–8), and the weakly bifidly toothed left mandible (Figs. 5–6) are found in few other New World Exosternini, and never in combination. Etymology. The name of this genus refers to the habits of the new species, showing an apparently exclusive but somewhat general preference for the cavity nests of true owls (family Strigidae), in combination with the common histerid ending –ister. The name is masculine., Published as part of Caterino, Michael S., Tishechkin, Alexey K. & Proudfoot, Glenn A., 2013, A New Genus and Species of North American Exosternini Associated with Cavity-Nesting Owls and a Reassignment ofPhelister simoniLewis (Coleoptera: Histeridae: Histerinae), pp. 557-565 in The Coleopterists Bulletin 67 (4) on pages 558-559, DOI: 10.1649/0010-065x-67.4.557

Key to the Central American species of Nymphister 1. Dorsal surface smooth and shiny; longitudinal discal striae of metaventrite absent, inner lateral striae of metaventrite represented only by recurrent arms.............................................. N. monotonus ( Reichensperger, 1935) - Dorsal surface with fine alutaceous microsulpture; longitudinal discal striae and inner lateral striae of metaventrite complete..................................................................................................... 2 2. Elytral surface smooth, without any punctures............................... N. simplicissimus Reichensperger, 1933 - Elytral surface with numerous shallow, circular, postobsolete (puncture rims broken and excavations level posteriorly) punctures, irregularly tracing course of 2–5 th dorsal and sutural striae................................ N. rettenmeyeri sp. n.

Trichoreninus neo Tishechkin & Mercado sp. n. (Figs. 3 & 4) Material Examined. Holotype male: " COSTA RICA: Heredia: La Selva Biol. Res. Stn.; 3 km S Puerto Viejo 80 m 10 ° 26 'N 84 ° 1 'W 2–5 -VI- 2001 S. Chatzimanolis ex. flight intercept trap CR 1 C01 0 11 / SM0747423 KUNHM- ENT [bar code label] / HOLOTYPE Trichoreninus neo sp. n. A. K. Tishechkin & A. Mercado C. des. 2012 " (SEMC). Paratypes (5): COSTA RICA: Puntarenas Province: Península de Osa, Rancho Quemado, collected on October 11 –28, 1993, leg. A. H. Gutiérrez (1 female: INBIO); Limon Province: Sector Cerro Cocori, Finca de E. Rojas, collected between May 28 and June 17, 1992, leg. E. Rojas (1 female: INBIO). PANAMA: Colón Province: Achiote, P.N. San Lorenzo, 9 ° 12 'N 79 ° 59 'W, flight intercept trap on June 11 –25, 2007, leg. A. Mercado (1 male: FMUP); San Lorenzo Forest, STRI crane site, 9 ° 17 'N 79 ° 58 'W, flight intercept trap, May 21 –24, 2004 by A. K. Tishechkin (1 male: FMUP); Darién Province: Cana Biological Station, 550 m, 79 ° 45 ' 18 ’’N 77 ° 41 ' 6 ’’W, flight intercept trap, on June 7 –9, 1996, leg. J. Ashe & R. Brooks (1 female: SEMC) Description: L: 2.28–2.35 mm; W: 1.87–1.91 mm; E/Pn L: 2.11–2.14; E/Pn W: 1.16–1.17; Pn W/L: 2.06–2.09; E L/W: 0.86–0.89; Pr/Py: 0.81–0.83; Sterna: 0.72 –. 75, 0.20–0.22, 0.53–0.55; Tibiae: 0.64 –.066, 0.73–0.76, 0.89 –093 (n= 6). Body oval in shape, convex dorsally, dark rufescent brown, surface with fine background punctuation throughout, without setae. Frons longer than wide, trapezoid, wider at posterior base, with prominent latero-marginal frontal carina and interrupted frontal stria; vertex with several large, deep, oval punctures arranged into two irregular, transverse rows. Labrum short, with row of small circular punctures along anterior margin, each bearing short seta, anterior margin of labrum projecting forward at the middle, convex on sides; mandibles long, robust, with curved long tips and a wide tooth on inner edge of each. Antenna with scape longer than wide, pyramidal, bearing a row of deep setose punctures along the edges; antennal funicle collectively shorter than scape; antennal clubs oval, with dense pubescence except for sclerotized areas in dorsal basal, outer lateral and outer ventral surfaces. Pronotum with posterior margin evenly arcuate, with a row of small shallow punctures along it; lateral margins almost straight, converging anteriorly towards prominent triangular anterior corners, anterior margin deeply emarginate behind the head; pronotal dics evenly convex, with irregularly spaced, large, deep, circular and oval punctures; marginal stria complete, thin and inconspicuous, outer lateral pronotal stria present, widely interrupted along anterior margin, occasionally also briefly interrupted around anterior corners, anterior portions are much more pronounced deeper and wider, than thin lateral portions; antennal cavities not visible in dorsal view. Prosternum with prosternal lobe sparsely punctate, long and wide, stria separating it from of the rest of prosternum irregularly punctate, indistinct; prosternal lobe with complete thin marginal stria (occasionally briefly interrupted medially), deep, large preapical foveae and deep lateral notches; prosternal keel rather narrow, flat, with scattered punctures, carinal striae convergent and united anteriorly, area between striae concave; lateral prosternal striae carinate, diverging anteriorly; posterior margin triangularly emarginate. Scutellum minute, triangural; elytra convex, widest around midpoint, lateral sides almost straight; all elytral striae deep, wide and sulcate, with small anostomosing punctures on bottoms of sulci; both outer and inner subhumeral striae complete, sometimes briefly interrupted around midpoint; 1 st– 3 rd dorsal and sutural striae complete, 4 th and 5 th striae variably abbreviate between midpoint and margin of anterior third, 4 th dorsal being on average longer than 5 th. Mesoventrite with medial triangular projection on anterior margin; marginal stria absent, disc of metaventrite with two narrow, deep, transverse depressions, which may be treated as fragments of extremely deepened transverse discal stria; meso-metaventral suture obsolete; metaventrite with median suture weak and inconspicuous, disc of metaventrite in both sexes mostly flat, with weak and narrow elevation along median suture in posterior third; disc usually with few (0–6) large, deep, elongate punctures, often positioned asymmetrically relative to midline; inner and outer lateral and longitudinal discal striae of metaventrite sulcate, similar to dorsal elytral striae, but impunctate, all but longitudinal discal striae complete, the latter are represented by a short fragments in posterior thirds of metaventrite, next to metacoxae; posterior ends of inner lateral striae straight, not approaching corresponding ends of longitudinal discal striae; recurrent arms of inner lateral striae present, long, widely separated from inner lateral striae; recurrent arms and outer lateral striae run at margins of depressions of lateral discs, where middle and hind legs rest in repose. First visible abdominal ventrite unmodified, flat, with a dense row of large, deep, postobsolete punctures along anterior margin, with paired long postmetacoxal striae laterally, inner ones being wide, sulcate and short, outer – thin and costate, with long recurrent arms. Femora elongate rectangular, profemora with punctures near anterior margin; meso- and metafemora smooth. Protibiae elongate oval, outer margin with 8–9 weak wide teeth topped with spines, both teeth and spines diminishing in size towards tibial apices and bases; apices rounded, rugose in texture, inner margin with row of setae, ventral surfaces with irregular rugose texture; meso- and metatibiae paddle-shaped, outer margins obtusely angulate at anterior third, smooth, each with a pair of spines at angles and near apices; inner margins with row of setae; tarsomeres 1–4 each bearing pair of ventral setae; pretarsal claws slightly curved. Propygidium hexagonal, marginal stria present along lateral and posterior margins. Pygidium trapezoid, convex, relatively short; surface with network of very five, transverse and circular strioles in addition to background tiny punctures (which are less pronounced than on propygidium); fine complete marginal stria present; discs in both sexes similar, no modifications present in females. Male genitalia as illustrated (Fig. 4). Etymology. This species is dedicated to the Neotropical Environment Option graduate studies program between McGill University and Smithsonian Tropical Research Institute, in which one of the authors (AMC) conducted her doctoral studies. The name is a noun in apposition. Distribution. Known from several localities in Costa Rica and Panama.

Key to the Central American species of Trichoreninus 1. Dorsal surface with abundant distinct setae; inner subhumeral and dorsal elytral striae 1���2 distinctly keeled; outer and posterior parts of elytral surfaces with irregularly rugose microsulpture; disc of metaventrite without large punctures (Figs. 42���44 in Tishechkin 2007)................................................................... T. flohri ( Lewis, 1891) - Dorsal surface completely asetose; subhumeral and dorsal elytral striae not elevated, striate or sulcate; surfaces without irregularly rugose microsulpture; disc of metaventrite at least with a few large punctures................................. 2 2. Dorsal elytral striae sulcate, wide and deep; disc of metaventrite medially without or with a few (2���6) large deep circular punctures along midline, often asymmetrically positioned................................................ T. neo sp. n. - Dorsal elytral striae regular, narrow and punctatostriate; disc of metaventrite with numerous (more than 10) smaller, circular, relatively shallow punctures, regularly scattered over the most of disc surface...................................... 3 3. Body surface regularly covered with small dense background punctures; outer lateral striae of pronotum interrupted and/or abbreviated in posterior half, known from Belize and Honduras.................................... T. carltoni sp. n. - Body surface without distinct background punctuation, sometimes tiny background punctures present on pronotum and sterna, elytral surface smooth, alutaceous; outer lateral striae of pronotum complete, reach posterior angles, known from Nicaragua to Panama.................................................................. T. geminus (Reichensperger, 1935), Published as part of Tishechkin, Alexey K. & C��rdenas, Alida Mercado, 2012, Description of three new species of Nymphistrini (Coleoptera: Histeridae: Haeteriinae) from Central America, pp. 36-48 in Zootaxa 3500 on page 41, DOI: 10.5281/zenodo.208915, {"references":["Tishechkin, A. K. (2007) Phylogenetic revision of the genus Mesynodites (Coleoptera: Histeridae: Hetaeriinae) with descriptions of new tribes, genera and species. Sociobiology, 49, 1 - 167.","Lewis, G. (1891) On new species of Histeridae. The Annals and Magazine of Natural History, 6, 381 - 405.","Reichensperger, A. (1935) Beitrag zur Kenntnis attaphiler Histeriden aus Brasilien (Col.). Revista de Entomologia, 5, 25 - 32."]}

Trichoreninus carltoni Tishechkin & Mercado sp. n. (Figs. 5 & 6) Material Examined. Holotype male: " HONDURAS: Santa Barbara, La Fe, Finca La Roca, 5.3 km S. Pe��a Blanca 14 �� 57 ���N, 88 ��02���W, 740 m 19���21 VI 1994, Brooks, Ashe # 174, ex: flight intpt. trap / HOLOTYPE Trichoreninus carltoni sp. n. A. K. Tishechkin & A. Mercado C. des. 2012 " (SEMC). Paratypes (2): BELIZE: Orange Walk District, Rio Bravo Conservation Area, La Milpa Field Station, flight intercept trap, 25.iv��� 5.v. 1996, leg. C. E. Carlton (1 male: LSAM); same data except 15���25.v. 1997 (1 male: AKT). Description: L: 2.22���2.25; W: 1.91���1.94; E/Pn L: 2.19���2.20; E/Pn W: 1.14���1.16; Pn W/L: 2.21���2.23; E L/W: 0.86���0.88; Pr/Py: 0.86���0.88; Sterna: 0.63���0.64, 0.17���0.19, 0.58���0.60; Tibiae: 0.62���0.64, 0.71 ���072, 0.90���0.92; (n= 3). Body oval in shape, convex dorsally, dark rufescent brown, surface with fine background punctuation throughout, without setae. Frons longer than wide, trapezoid, wider at posterior base, with prominent lateromarginal frontal carina and interrupted frontal stria; vertex with numerous irregularly scattered, moderate, shallow, oval punctures. Labrum short, with row of small circular punctures along anterior margin, each bearing short seta, anterior margin of labrum projecting forward at the middle, convex on the sides; mandibles long, stout, with curved long tips and a wide tooth on inner edge of each. Antenna with scape longer than wide, pyramidal, bearing a row of deep setose punctures along the edges; antennal funicle collectively shorter than scape; antennal clubs oval, with dense pubescence except for sclerotized areas in dorsal basal, outer lateral and outer ventral surfaces. Pronotum with posterior margin evenly arcuate, with a row of tiny shallow punctures along it; lateral margins almost straight, converging anteriorly towards prominent triangular anterior corners, anterior margin deeply emarginate behind the head; pronotal disc evenly convex, with irregularly spaced, large, shallow, circular and oval punctures, irregularly anostomosing in anterior half, especially in antero-lateral corners; marginal stria complete, thin and inconspicuous, outer lateral pronotal stria present, widely interrupted along anterior margin, variably interrupted and/or abbreviate along posterior halves; antennal cavities not visible in dorsal view. Prosternum with prosternal lobe sparsely punctate, long and wide, stria separating it from of the rest of prosternum very fine and almost indistinct; prosternal lobe with complete thin marginal stria, small preapical foveae and deep lateral notches; prosternal keel rather narrow, flat, with scattered punctures, carinal striae convergent and united anteriorly, area between striae flat; lateral prosternal striae carinate, diverging anteriorly; posterior margin triangularly emarginate. Scutellum minute, triangural; elytra convex, widest at anterior third, narrowing almost straightly towards posterior ends; all elytral striae shallow and punctate; both outer and inner subhumeral striae complete, sometimes briefly interrupted around midpoint; 1 st��� 4 th dorsal and sutural striae complete, 5 th striae variably abbreviate in anterior third, anterior portions of 4 th dorsal and sutural striae in close proximity, but not united. Mesoventrite with medial triangular projection on anterior margin; marginal stria absent, disc of metaventrite with two fragments of deep transverse discal stria, each united to anterior ends of corresponding outer lateral striae of metaventrite and so looking as its continuation; meso-metaventral suture and median suture of metaventrite obsolete; disc of metaventrite (in males) with distinct, but shallow depression, occupying most of its surface; disc mostly covered with numerous (more than 14) large, shallow, circular punctures, few of such punctures also present on elevated parts of lateral discs; inner and outer lateral and longitudinal discal striae of metaventrite wide, sulcate, impunctate; outer lateral striae complete, inner lateral striae long, abbreviate in posterior fourths, longitudinal discal striae represented by short fragments, occasionally broken in two pieces, in posterior thirds of metaventrite, next to metacoxae; recurrent arms of inner lateral striae present, long, widely separated from inner lateral striae; recurrent arms and outer lateral striae run at margins of depressions of lateral discs, where middle and hind legs rest in repose. First visible abdominal ventrite unmodified, flat, with a row of moderate, deep, circular punctures along anterior margin, with paired long postmetacoxal striae laterally, inner ones being wide, sulcate and short, outer ��� thin and costate, with long recurrent arms. Femora elongate rectangular, profemora with punctures near anterior margin; meso- and metafemora smooth. Protibiae elongate-oval, outer margin with 7���8 weak wide teeth topped with spines, both teeth and spines diminishing in size towards tibial apices and bases; apices rounded, rugose in texture, inner margins with row of setae, ventral surfaces with irregular rugose texture; meso- and metatibiae paddle-shaped, outer margins weakly angulate at anterior third, smooth, each with a pair of spines at angles and near apices; inner margins with row of setae; tarsomeres 1���4 each bearing pair of ventral setae; pretarsal claws slightly curved. Propygidium hexagonal, with some larger shallow punctures, scattered among background punctuation; marginal stria present along most of lateral and posterior margins. Pygidium trapezoid, convex, relatively short, with network of very five, transverse and circular strioles in addition weakened background tiny punctuation; fine complete marginal stria present; disc modifications in females, if present, unknown. Male genitalia as illustrated (Fig. 6) Etymology. The species is named after Christopher E. Carlton (LSAM), collector of two type specimens, in appreciation of our long-term collaboration and his contributions to the knowledge of obscure New World beetles. Distribution. Known from two localities in Belize and Honduras., Published as part of Tishechkin, Alexey K. & C��rdenas, Alida Mercado, 2012, Description of three new species of Nymphistrini (Coleoptera: Histeridae: Haeteriinae) from Central America, pp. 36-48 in Zootaxa 3500 on pages 44-46, DOI: 10.5281/zenodo.208915

Nymphister rettenmeyeri Tishechkin & Mercado sp. n. (Figs. 1 & 2) Material Examined: Holotype male: " PANAMA: Panama Barro Colorado Isl. 09�� 11 'N. 79 �� 51 'W 16 July 1994. D. Banks ex: flight intercept trap / LSAM 0044245 / HOLOTYPE Nymphister rettenmeyeri sp. n. A. K. Tishechkin & A. Mercado C. des. 2012 " (SEMC). Paratypes (23): PANAMA: Panama Province: Barro Colorado Island, in a colony of Eciton hamatum on 6.vii. 1968, leg. R. L. Torgenson (1 female, WSU); Barro Colorado Island, in a colony of Eciton mexicanum, 2.v. 1967, leg. R. D. Akre (3 females, WSU); Barro Colorado Island, late statary refuse pile of Eciton burchelli, 7.ii. 1976, leg. A. F. Newton (1 female: FMNH); Barro Colorado Island, flight intercept traps on various dates in May 1981 and June 1983, leg. B. D. Gill (5 exs.: AKT, CMN and BDG); Barro Colorado Island, flight intercept trap, 3.vii. 1994, leg. D. Banks (1 female: SEMC); Col��n Province: Achiote, P.N. San Lorenzo, 9 �� 12 'N 79 �� 59 'W, flight intercept traps, various dates in May���June 2007 and January 2008, leg. A. Mercado (6 exs.: FMUP, AKT); San Lorenzo Forest, Smithsonian Tropical Research Institute canopy crane site, 9 �� 17 'N 79 �� 58 'W, flight intercept traps, 12���17.v. 2004, leg. A. K. Tishechkin (1 male, 1 female: FMUP); 14 km N junction of Escobal and Pi��a Roads, flight intercept trap, 2���11.vi. 1996, leg. J. Ashe & R. Brooks (1 female: SEMC). COSTA RICA: Puntarenas Province: Rincon de Osa, 8 �� 41.141 'N 83 �� 31.117 'W, flight intercept trap, 23���26.vi. 2001, leg. S. & J. Peck (1 female: SEMC); Estaci��n Quebrada Bonita, Reserva Biologica Carara, flight intercept trap on 1���28.ii. 1994, leg. R. M. Guzm��n (1 male: INBIO); Pen��nsula de Osa, Rancho Quemado, 1���30.vii. 1992, leg. F. Quesada (1 female: INBIO). Description: L: 1.58���1.63; W: 1.50���1.52; E/Pn L: 2.23���2.26; E/Pn W: 1.13���1.15; Pn W/L: 2.46���2.51; E L/W: 0.77���0.80; Pr/Py: 1.06���1.07; Sterna: 0.42���0.44, 0.10���0.12, 0.40���0.43; Tibiae: 0.53���0.56, 0.63���0.65, 0.71���0.74; (n= 10). Body round, convex dorsally, dark reddish brown, dark brown along the edges, body surface with fine alutaceous microsculpture throughout, without setae. Frons concave in the middle, with prominent latero���marginal frontal carina and interrupted frontal stria. Labrum narrow, asetose; mandibles wide, stout, with inwardly curved short tips. Antennae with scape thick, angulate, pyramide���shaped; antennal clubs oval, with dense pubescence except for sclerotized areas in dorsal basal, outer lateral and outer ventral surfaces. Pronotum with posterior margin obtusely angulate, with a row of small round deep punctures, lateral sides almost straight, narrowing anteriorly; pronotal disc convex, without punctures or setae, lateral margins narrowly explanate and slightly raised; lateral marginal stria complete, but inconspicuous; anterior margin emarginated behind the head; antennal cavities not visible in dorsal view. Prosternum with prosternal lobe short, separated of rest of prosternum by stria, with complete thin marginal stria and deep lateral notches; prosternal keel broad, flat, with two pairs of closely situated carinal striae along outer edges of the keel; lateral prosternal striae distinct, diverging anteriorly; posterior margin triangularly emarginated, depressed medially. Scutellum minute, triangular; elytra convex, widest at anterior third, narrowing arcuately towards posterior ends; both outer and inner subhumeral striae complete, thin and impunctate; 1 st dorsal striae present as long, shallow, narrow, but distinct impunctate sulci, the rest of dorsal and sutural striae each marked by a few irregularly spaced, shallow, large, postobsolete punctures. Mesoventrite with medial triangular projection on anterior margin; marginal stria complete, thin, disc of mesoventrite with transverse bisunuate discal stria separated in the middle into two long fragments; meso���metaventral suture very fine and inconspicuous; mesoventrite with median suture distinct and complete, discs of metaventrite in males with shallow, narrow median depression and slight median elevation in posterior third, evenly convex in females; inner and outer lateral and longitudinal discal striae of metaventrite complete, brief fragment of extra stria present anteriorly inward of each of longitudinal discal striae; posterior ends of inner lateral striae curved inwards, almost connected to corresponding ends of longitudinal discal striae; recurrent arms of inner lateral striae present, long, widely separated from inner lateral striae; recurrent arms and outer lateral striae run atop of sharp, low keels, margining depressions of lateral discs, where middle and hind legs rest in repose. First visible abdominal ventrite unmodified, weakly convex, with paired long postmetacoxal striae laterally. Pro-, meso- and metafemora elongate rectangular, without punctures, with complete transverse striae in anterior thirds, coxa flattened. Protibiae elongate oval, widest at the middle, outer margin with a few very weak shallow emarginations, almost straight, with 7���8 small spines along outer margin, apex rounded, rugose in texture, inner margin with row of setae; meso- and metatibiae paddle-shaped, outer margins obtusely angulate at anterior third, inner margins with row of setae; tarsomeres each with pair of setae on the ventral side, tarsal claws strongly bent near the base. Propygidium hexagonal, with shallow, irregular, anostomosing punctures creating rugose background, more pronounced anteriorly, with complete marginal stria. Pygidium convex, relatively short, trapezoid; background punctuation similar to those on propygidium, but finer; no marginal striae present; disc in both sexes medially with a pair of weak circular elevations, separated along midline. Male genitalia as illustrated (Fig. 2) Etymology. We dedicate this species to the memory Carl W. Rettenmeyer (1931���2009), an eminent specialist on army ants and their inquilines and collector of numerous haeteriine specimens. Distribution. Known from several localities in southern Costa Rica and west-central Panama., Published as part of Tishechkin, Alexey K. & C��rdenas, Alida Mercado, 2012, Description of three new species of Nymphistrini (Coleoptera: Histeridae: Haeteriinae) from Central America, pp. 36-48 in Zootaxa 3500 on pages 37-38, DOI: 10.5281/zenodo.208915

Genus Trichoreninus Lewis, 1891 The current concept of the genus was established by Tishechkin (2007), who redescribed it, listed in it two Central and five South American species (two of which were retained formally, since they do not correspond to the generic diagnosis; however, their formal reassignment has not been performed yet) and included it in the key of the genera of Nymphistrini. At least ten undescribed species are known (Tishechkin, unpublished data), two of which are here described. The distribution of two described Central American Trichoreninus species has been summarized in Tishechkin (2007)., Published as part of Tishechkin, Alexey K. & C��rdenas, Alida Mercado, 2012, Description of three new species of Nymphistrini (Coleoptera: Histeridae: Haeteriinae) from Central America, pp. 36-48 in Zootaxa 3500 on page 41, DOI: 10.5281/zenodo.208915, {"references":["Lewis, G. (1891) On new species of Histeridae. The Annals and Magazine of Natural History, 6, 381 - 405.","Tishechkin, A. K. (2007) Phylogenetic revision of the genus Mesynodites (Coleoptera: Histeridae: Hetaeriinae) with descriptions of new tribes, genera and species. Sociobiology, 49, 1 - 167."]}

Renclasea mexicana Tishechkin & Caterino, n. sp. (Fig. 10) Material examined: Holotype male: " MEXICO: Hidalgo 14.5 NW Cardonal 13 July 1990, 1850 m Danoff- Burg, ex: from Neivamyrmex bivouac / HOLOTYPE Renclasea mexicana sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (SEMC). Diagnosis: Apart from its disjunct distribution, this species can be identified by the presence of distinct alutaceous microsculpture over almost entire dorsal and parts of ventral surfaces and presence of pair of small, shiny calluses near posterior margin of pronotum. Two other species with dorsal alutaceous microsculpture present lack the latter and have dorsal microsculpture much less widespread and present only on intervals and along apical margins on elytra. Description: L: 2.07; W: 1.64; E/Pn L: 1.95; E/Pn W: 1.17; Pn W/L: 1.82; E L/W: 0.91; Pr/Py: 0.99; Sterna: 0.45, 0.15, 0.46; Tibiae: 0.54, 0.62, 0.72 (n= 1). Body light reddish-brown, mostly covered with alutaceous microsculpture, smooth and asetose throughout except for a few setae on antennomeres 1���7. Vertex and most of frons covered with microsculpture; frons almost flat, clypeus depressed at middle between lateral carinae; labrum narrowly rectangular, weakly produced at the middle of apical margin. Prosternum covered with microsculpture, except for shiny smooth explanate lateral margins, a pair of slightly elevated elongate oval calluses at latero-medial parts of disc in its basal third and small opposing areas along basal margins. Prosternal sides convergent, much more strongly in anterior fourth, above antennal cavities, weakly outwardly sinuate, with the anterior angles narrowly rounded, almost rectangular; marginal stria present along lateral edge, interrupted at anterior angles; pronotal lateral sides narrowly flattened and slightly bent upwards; median angle of pronotal posterior margin about 100 ��. Prosternum with anterior margin of prosternal lobe weakly concave; prosternal keel slightly elevated and flattened, covered with fine microsculpture, without carinal striae, with short basal divergent fragments of lateral prosternal striae between procoxae. Scutellum short triangular, tiny, but distinct; elytra convex, widest at middle, completely covered by alutaceous microsculpture, slightly rugulose in inner posterior third; dorsal elytral striae 1���3 weakly marked on disc, abbreviated both anteriorly and posteriorly; sutural stria abbreviated in anterior fourth. Mesoventrite flat in males, mesoventral projection short, broadly triangular, its apex slightly elevated; mesometaventral suture 'curly bracket' (���{���)-shaped, thin and inconspicuous; disc of metaventrite in male in anterior half with shallow oval depression, not reaching anterior parts of outer metaventral striae laterally; weak wide elevation present near posterior margin of metaventrite. Lateral parts of meso- and metaventrite and abdominal ventrite 1 with fine alutaceous microsculpture. Propygidium weakly convex, microsculpture on its disc as on apices of elytra; marginal stria of propygidium complete; pygidium smooth, microsculptured in anterior part, weakly convex in males. Male genitalia as illustrated (Fig. 10), females are unknown. Etymology. The species epithet reflects its apparent endemicity to Mexico. Distribution. This species is only known from the type locality (Fig. 14)., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 12-13, DOI: 10.5281/zenodo.191863

Renclasea falli Tishechkin & Caterino, n. sp. (Figs. 5, 6, 7, 8) Material examined: Holotype male: "CA: Riverside Co. 33.7244 ��N, 116.8061 ��W, SBNF: San Jacinto R. v. 24.2005, M. Caterino at light / CA BEETLE PROJ, CBP0029649 / HOLOTYPE Renclasea falli sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (SBMNH). Paratype (1): female from Bautista Canyon, 12 mi SE Hemet, California collected on February 26, 1971 (CDFA); genitalia missing. Diagnosis: This species can be identified by the presence of distinct complete prosternal carinal striae (Fig. 6 A). In addition to that, it appears smaller and less robust than R. occidentalis and R. helavai, the only other species without alutaceous background microsculpture on dorsal surface. The female is one of only two species (as far as known) that have a mesosternal callus (Fig. 6 B), the other being R. helavai. In R. falli this callus is smooth, and only very finely punctate, whereas in R. helavai it is finely rugose. Description: L: 1.80; W: 1.41; E/Pn L: 1.75; E/Pn W: 1.15; Pn W/L: 1.64; E L/W: 0.93; Pr/Py: 1.04; Sterna: 0.41, 0.14, 0.42; Tibiae: 0.46, 0.54, 0.63 (n= 2). Body (Fig. 5) pale rufescent, shiny, smooth and asetose throughout except of a few setae on antennomeres 1���7. Frons and clypeus depressed at middle between lateral carina; labrum narrowly rectangular, with a small weak knob at the middle of apical margin. Prosternal sides convergent, much more strongly in anterior fifth; anterior margin straight above antennal cavities, with the anterior angles sharp, almost rectangular; marginal stria present along lateral edge, slightly extending around anterolateral corner; pronotal lateral sides narrowly flattened and slightly reflexed; median angle of pronotal posterior margin about 110 ��. Prosternum with anterior margin of prosternal lobe weakly concave; prosternal keel slightly elevated and weakly convex, carinal striae thin and long, converging anteriorly, their anterior ends in close proximity, but not united, almost reaching base of prosternal lobe. Scutellum short triangular, tiny, but distinct; elytra convex, widest at anterior third, with minute sparse background punctures, being denser and more conspicuous along sutural striae and at posterior fifth, where some merge into a weak background microsculpture; dorsal elytral striae 1���4 weakly marked on disc, abbreviated both anteriorly and posteriorly; sutural stria abbreviated in anterior eighth, its anterior end slightly bent outwards. Mesoventrite flat, with mesoventral projection short, triangular, its apex slightly elevated, in female with subcircular impressed callus (Fig. 6 A), its borders not closed posteriorly; mesometaventral suture widely Vshaped, thin and inconspicuous; disc of metaventrite in male in anterior two-thirds with shallow wide triangular depression, bordered laterally by anterior parts of outer striae of metaventrite, narrowing posteriorly; disc surface weakly raised posteriorly. Propygidium weakly convex, disc with fine microsculpture of merged short transverse striolets; marginal stria of propygidium widely interrupted along posterior margin; pygidium smooth, weakly convex in male. Female with striate pygidial ornament (Fig. 7 B). Male genitalia as illustrated (Fig. 8); female genitalia unknown. Etymology. We take pleasure in naming this new species for Henry Clinton Fall (1862-1939), one of the fathers of California coleopterology. Distribution. This species is known from two localities in western Riverside County, California (Fig. 14). The habitat at the type locality is dry, open, sandy riparian woodland, with western sycamores (Platanus occidentalis) and coast live oak (Quercus agrifolia) as the dominant trees, along with a large assortment of shrubs, including willows (Salix spp.), coyote brush (Baccharis sp.), Yucca sp. and others., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 6-8, DOI: 10.5281/zenodo.191863

Renclasea Tishechkin & Caterino, n. gen. Type species: Renclasea skelleyi n. sp. Description. Body elongate oval, convex dorsally (L: 1.6���2.3 mm). Body surface smooth and mostly shiny (Figs. 2 A, 5 A), sometimes with areas of alutaceous microsculpture dorsally (Fig. 1 B), mostly asetose, some species with sparse inconspicuous setae, especially on legs, antennal scape, labrum and clypeus. Head with latero-marginal frontal carina descending along inner edge of eyes (Fig. 2 A), then running obliquely forward to posterior margin of clypeus, interrupted along clypeolabral border. Frons concave between carinae, frontal stria runs on top of the carina and is interrupted in medial concavity; occipital stria thin, complete, continuous with branches of supraorbital stria. Labrum in the same plane as clypeus, small, rectangular, sometimes with small blunt knob in the middle of apical margin. Mandibles robust, short, with faces of bases unmodified. Antennal scape strongly expanded (Fig. 2 D), irregularly pyramidal, with all edges sharp. Antennal clubs with dense pubescence except for large sclerotized areas on dorsal, inner lateral and ventral surfaces. Pronotum convex, lateral sides weakly convergent anteriorly, moderately but conspicuously explanate. Pronotal basal margin obtusely angulate. Pronotal marginal stria complete (Fig. 1 A, 2 A), anterior stria broadly interrupted medially, present only around anterior angles. Anterior margin of pronotum with a pair of small glandular openings behind and slightly mediad eyes; lateral margin also with two elongated gland openings (Figs. 1 A). Scutellum triangular, minute and inconspicuous. Elytron with reduced set of thin, impunctate dorsal striae. Both subhumeral striae complete, carinate; sutural stria abbreviated anteriorly or completely absent, traces of dorsal striae 1���4 variably present, apical stria thin, complete, united with sutural (if present) and inner subhumeral striae. Elytro-epipleural border sharp (Fig. 2 C), angulate along outer subhumeral stria. Propygidium transversely hexagonal, with marginal stria complete, sometimes weakened along apical margin. Pygidium in females with ornament of thin striae represented by apical and/or basal transverse, and medial circular striations (Fig. 7). Prosternal lobe wide and short (Figs. 1 D, 2 B), with straight or slightly concave anterior margin. Anterior marginal stria of prosternal lobe complete, diverging from margin laterally; lateral foveae absent, lateral notches present. Prosternal keel relatively narrow, flat or weakly convex, with or without carinal striae, those if present narrowly separated, parallel (Fig. 6 A). Lateral marginal prosternal striae distinct and long. Mesoventrite narrow, its anterior margin produced medially as a weak triangular process (Figs. 1 D, 2 B, 6). Marginal lateral stria of mesoventrite present, discal marginal stria absent. Metaventral disc in males with weak median depression. Outer lateral stria of metaventrite present, complete. Inner lateral stria of metaventrite present only as long, double striated recurrent arm. First abdominal ventrite with lateral and postmetacoxal striae united near posterior edge of metacoxa, running thence as long, curved striae towards elytral margin. Legs relatively short; tibia wide, paddle-like. Protibia (Fig. 1 E) strongly expanded, with 14���17 short spines along outer margin and with 8���10 bristles along apical part of inner margin. Meso- and metatibia (Figs. 1 F���G) without teeth and spines on outer margins, on ventral side with complete marginal stria and two longitudinal striae closer to inner margin. Aedeagus with parameres long, with long basal fusion ventrally and dorsally. Penis aligned along the longitudinal paramere axis. Basal piece long, with shallow and wide dorsal and ventral apical emarginations, its posterior opening circular, in caudal position. Male sternite 8 with fused full-sized halves (although suture between halves distinct), pair of small velae and fringes of long setae along postero-lateral parts of apical margin. Dorsal apical parts of male sternite 8 with inward long narrow rectangular processes. Male tergite 8 with transverse anterior stria (TAS) and complete intra-TAS plate present and no transverse posterior suture. Intra-TAS plate not separated from the tergite body along TAS. Male sternite 9 with slightly expanded stickshaped ���handle���. Male tergite 9 with medium ventral apodemes, long basal projections, short pointed apical projections and medium sclerotized ventral processs. Halves of male tergite 9 separate; tergite 10 present, large, fitting into apical emargination of tergite 9, with apical margin widely and deeply acutely emarginated. Female sternite 8 with deeply emarginate apical margin, distinct basal bridge with angulate lateral parts and obtusely angulate basal angles. Female tergite 8 present as single plate or three narrowly separated plates. Coxites not connected. Female tergite 9 present as single transverse plate, its lateral margins close, but not connected to coxites. Female genital sclerites separate, elongate, strip-shaped. Etymology. The generic epithet represents the combination of hetaeriine generic names Euclasea and Reninus Lewis 1889, relatively closely related genera, some representatives of which resemble Renclasea species. The gender is feminine. Diagnosis. Despite the generalized appearance of Renclasea and history of taxonomic confusions surrounding the genus, it is relatively easily recognizable among Hetaeriinae, especially given its North American distribution. It runs to Euclasea in the key to North American Hetaeriinae (Kovarik & Caterino 2001) as well as in the most comprehensive key available for the entire subfamily (Helava et al. 1985). The latter key is not complete, but even given this fact and taking into consideration available undescribed hetaeriine material (Caterino & Tishechkin, unpublished), it would work well to identify Renclasea species in global hetaeriine context (keying them to Euclasea). The combination of convex elongate body shape, smooth shiny surfaces almost completely devoid of setae, explanate pronotal sides, strongly expanded pro-, but not meso- and metatibia and reduction of striae and punctures on elytra and metaventrite readily diagnose the genus. Some representatives of Euclasea (sensu Tishechkin 2007) may be quite similar externally, but none of them have explanate sides of pronotum and strongly expanded protibia. Some species of Reninus (those included earlier in the presently synonimized subgenus Brachylister Bickhardt, 1917) have similar general appearance and similar structure of male genitalia, but all of them have extremely expanded meso- and metatibia and complete set of discal and lateral striae on metaventrite., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 2-3, DOI: 10.5281/zenodo.191863, {"references":["Kovarik, P. W. & Caterino, M. S. (2001) Histeridae Gyllenhall, 1808. Pp. 212 - 227 in: Arnett, R. H. & M. C. Thomas (eds.). American beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton - London - New York - Washington, DC.","Helava, J. V. T., Howden, H. F. & Ritchie, A. J. (1985) A review of the New World genera of the myrmecophilous and termitophilous subfamily Hetaeriinae (Coleoptera: Histeridae). Sociobiology, 10, 127 - 386.","Tishechkin, A. K. (2007) Phylogenetic revision of the genus Mesynodites Reichardt, 1924 (Coleoptera: Histeridae: Hetaeriinae) with description of new tribes, genera and species. Sociobiology, 47, 1 - 167."]}

Renclasea skelleyi Tishechkin & Caterino, n. sp. (Figs. 1, 3, 4) Material examined: Holotype male: "FLORIDA: Levy Co. 4.0 mi. SW Archer 16 -V- 1991; P. E. Skelley blacklight on ground / HOLOTYPE Renclasea skelley sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (FSCA). Paratypes (6): female from Florida, Putnam Co., 2.5 mi NE Florahome, collected by P. Skelley on January 8, 1993 (FSCA); male from Florida, Marion Co., Lake Delancy collected at UV lights by J. Cicero on August 31, 1996 (FSCA); male from Florida, Marion Co., Ocala Nat. Forest, 2.5 mi N of 40, FS 97 collected at UV/MV lights by R. Morris and S. Lingafelter on July 26 ���27, 2002 (AKT); male and female from Georgia, Tattnall Co., 2���3 mi E of 147 along Ohoopee River near Reidsville, 32 �� 0.54 'N 82 �� 8.48 'W, collected at UV lights by R. Morris and E. H. Donaldson on May 7, 1998 (PWK); male from the same locality collected at UV lights by R. Morris on August 1, 1998 (AKT). Diagnosis: By the presence of alutaceous microsculpture in elytral intervals 1���4 and along the posterior margin of elytra this species resembles only R. helavai. Apart from its disjunct distribution, R. skelleyi can be distinguished from the latter by only weakly marked traces of dorsal striae 1���3, different pattern of female pygidial ornament (Fig. 7 A) and lack of median callus on mesoventrite in females. Description: L: 1.89; W: 1.49; E/Pn L: 1.81; E/Pn W: 1.19; Pn W/L: 1.66; E L/W: 0.92; Pr/Py: 1.15; Sterna: 0.47, 0.16, 0.44; Tibiae: 0.51, 0.60, 0.69 (n= 7). Body reddish-brown, shiny, except areas of alutaceous microsculpture on elytra (Fig. 1 B) and pygidium, smooth and asetose. Frons almost flat, clypeus depressed at middle between lateral carinae; labrum narrowly rectangular, weakly produced at the middle of apical margin. Prosternal sides convergent (Fig. 1 A), much more strongly in anterior fourth, above antennal cavities, weakly inwardly sinuate, with the anterior angles narrowly rounded, almost rectangular; marginal stria present along lateral edge, almost entire, variably narrowly interrupted between anterior angles and outer part of anterior emargination; pronotal lateral sides narrowly flattened and slightly reflexed; median angle of pronotal posterior margin about 100 ��. Prosternum (Fig. 1 D) with anterior margin of prosternal lobe weakly concave; prosternal keel slightly elevated and flattened, covered with fine microsculpture, without carinal striae. Scutellum elongate triangular, small; elytra (Fig. 1 B) convex, widest at middle, with minute sparse background punctures, these more conspicuous along sutural stria; dorsal elytral striae 2���3 weakly marked on disc, abbreviated both anteriorly and posteriorly, weaker traces of oblique humeral stria and dorsal stria 1 also present; sutural stria abbreviated in anterior fourth; elytral intervals with elongate band of fine alutaceous microsculpture, somewhat expanded and merging in posterior third. Mesoventrite (Fig. 1 C) flat in males, with somewhat depressed areas laterad of the mesoventral projection base in females; mesoventral projection short, triangular, its apex slightly elevated; mesometaventral suture 'curly bracket'-shaped, thin and inconspicuous; disc of metaventrite in males in anterior half with shallow wide triangular depression bordered laterally by anterior parts of outer striae of metaventrite; weak wide elevation present near posterior margin of metaventrite; disc of metaventrite in females weakly, evenly convex. Lateral parts of meso- and metaventrite with fine alutaceous microsculpture. Propygidium weakly convex, disc with fine dense shallow punctures merging locally into shallow transverse wrinkles; marginal stria of propygidium complete; pygidium smooth, with few small shallow punctures along anterior margin, weakly convex, with striate ornament in females (Fig. 7 A). Male and female genitalia as illustrated (Figs. 3, 4). Etymology. This species is dedicated to one of its collectors, Paul Skelley of FSCA, in appreciation of our long collaboration and his efforts in collecting rare and poorly known beetles in southeastern United States. Distribution. Known from several localities in northern Florida and southern Georgia (Fig. 14)., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 4-5, DOI: 10.5281/zenodo.191863

Renclasea cazieri Tishechkin & Caterino, n. sp. (Figs. 2, 13) Material examined: Holotype male: "Portal, Ariz. Cochise Co. 4700 ft. at light VI- 28-64 / Collectors Jean H. Puckle M. A. Mortenson M. A. Cazier / PROPERTY OF ARIZONA STATE UNIVERSITY / HOLOTYPE Renclasea cazieri sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (ASU). Diagnosis: This is one of three species without alutaceous background microsculpture on dorsal surface (Fig. 2 A). It can be easily distinguished from other species in this group by the total lack of traces of dorsal and sutural striae on elytra as well as the presence of sharp medial longitudial keel on meso- and anterior part of metaventrite (Fig. 2 B). Description: L: 2.34; W: 1.91; E/Pn L: 1.86; E/Pn W: 1.19; Pn W/L: 1.76; E L/W: 0.88; Pr/Py: 1.08; Sterna: 0.53, 0.22, 0.45; Tibiae: 0.52, 0.63, 0.78 (n= 1). Body rufescent, shiny, smooth, short inconspicuous setae present on antennae, frons, legs, prosternal lobe, lateral parts of sterna and elytra. Frons and clypeus (Fig. 2 E) weakly depressed at middle between lateral carinae; labrum narrowly rectangular, its apical margin weakly inwardly arcuate, unmodified. Prosternal sides convergent, much more strongly in anterior fourth, above antennal cavities, straight, with the anterior angles blunt, almost rectangular; marginal stria present along lateral edge, extending around anterolateral corner, interrupted at middle of anterior emargination of pronotum; pronotal lateral sides narrowly flattened and weakly reflexed; median angle of pronotal posterior margin about 110 ��. Prosternum with anterior margin of prosternal lobe almost straight; prosternal keel moderately elevated and narrow, its base in male not excavated; carinal striae absent, though with weak basal fragments marked between procoxae. Scutellum elongate triangular, small; elytra convex, widest around midpoint, smooth and shiny; any traces of dorsal elytral and sutural striae absent. Mesoventrite in males with sharply delimited median longitudinal keel (Fig. 2 B) extending onto anterior part of metaventrite; mesoventral projection short, broadly triangular, its apex slightly elevated, continuous with anterior part of longitudinal keel; mesometaventral suture obsolete; disc of metaventrite in males flat, in addition to posterior part of longitudinal keel of mesoventrite bears short, sharp tooth on midline around posterior third. Propygidium weakly convex, smooth; marginal stria of propygidium abbreviated along posterior margin; pygidium in males smooth, convex. Male genitalia as illustrated (Fig. 13); females are unknown. Etymology. We name this species in honor of Mont A. Cazier (1911���1995), collector of the type and founding director of the Southwest Research Station, who facilitated and inspired generations of entomological exploration in the American desert southwest. Distribution. This species is only known from the type locality, on the edge of the Chiricahua Mountains of southeastern Arizona (Fig. 14)., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 15-16, DOI: 10.5281/zenodo.191863

Renclasea helavai Tishechkin & Caterino, n. sp. (Figs. 6, 7, 9) Material examined: Holotype female: "Tucson, Ariz. Aug. 5 1935 Bryant 20 At light / SEM / Collection of the CALIFORNIA ACADEMY OF SCIENCES, San Francisco, Calif. / Euclasea n. sp. 2 Helava ' 82 / HOLOTYPE Renclasea helavai sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (CAS). Diagnosis: Among three species with the presence of dorsal alutaceous microsculpture (R. helavai, R. mexicana, R. skelleyi), this species can be identified by its distinct dorsal elytral elytra striae 1���4. The prominent circular medial callus of mesoventrite (in females; Fig. 6 B) is shared only with R. falli, in which it is smooth rather than faintly rugose. Description: L: 1.68; W: 1.33; E/Pn L: 1.95; E/Pn W: 1.20; Pn W/L: 1.73; E L/W: 0.94; Pr/Py: 1.00; Sterna: 0.39, 0.18, 0.42; Tibiae: 0.45, 0.55, 0.62 (n= 1). Body reddish-brown, shiny, except areas of alutaceous microsculpture on frons, pronotum, elytra and pygidia; smooth and asetose. Frons almost flat, microsculptured, clypeus smooth, depressed at middle between lateral carinae; labrum narrowly rectangular, its apical margin weakly inwardly arcuate. Prosternal sides convergent, much more strongly in anterior third, above antennal cavities, weakly inwardly sinuate, with the anterior angles narrowly rounded, almost rectangular; marginal stria present along lateral, but not anterior edge. Pronotal surface mostly covered with alutacecous microsculpture, except bands along narrowly flattened and reflexed lateral sides; median angle of pronotal posterior margin about 100 ��. Prosternum with anterior margin of prosternal lobe weakly convex; prosternal keel slightly elevated and flattened, without carinal striae, its base excavated to fit process of mesoventrite. Scutellum elongate triangular, tiny; elytra convex, widest at middle, smooth and shiny; dorsal elytral striae 1���4, distinct, not punctured, long, progressively shorter from stria 1 to 4, stria 1 almost reaching elytral apex and stria 4 abbreviated in posterior third; sutural stria complete; elytral intervals with elongate bands of fine alutaceous microsculpture, somewhat expanded and merging in posterior fifth. Mesoventrite in females with median part of disc flat and lateral sides depressed, flat median part most mostly occupied by circular low microsculptured callus (Fig. 6 B); mesoventral projection long, triangular; mesometaventral suture obsolete; disc of metaventrite in females weakly, evenly convex. Lateral parts of meso- and metaventrite with fine alutaceous microsculpture. Propygidium weakly convex, disc with fine, dense, shallow punctures merging locally into shallow transverse wrinkles; marginal stria of propygidium complete; pygidium in females weakly convex, with rugose sculpture and striate ornament. Female genitalia as illustrated (Fig. 9), males are not known. Etymology. This species is dedicated to Jussi Helava, who first recognized it as an undescribed, honoring his contributions to the taxonomy and systematics of Hetaeriinae. Distribution. This species is only known from the type locality (Fig. 14)., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 9-10, DOI: 10.5281/zenodo.191863

Key to the species of Renclasea 1. Prosternal keel with carinal striae present (Fig. 6 A) ................................................................................ R. falli n. sp. 1 '. Prosternal keel without carinal striae............................................................................................................................ 2 2. Substantial areas of dorsal body surface with alutaceous microsculpture, at least on elytral intervals and apical areas..................................................................................................................................................................................... 3 2 '. Dorsal body surface without alutaceous microsculpture.............................................................................................. 5 3. Dorsal striae inconspicuous, barely traceable; surface of pygidium smooth, shiny or finely microsculptured........... 4 3 '. Dorsal striae 1���4 distinct, almost complete, thin, impunctate and slightly costate, especially anteriorly; surface of pygidium rugose; disc of mesoventrite (in females) medially with circular flat microsculptured callus (Fig. 6 B)....... .............................................................................................................................................................. R. helavai n. sp. 4. Surface of frons, vertex, pronotum, most of elytra and propygidium with alutaceous microsculpture; northern Mexico (Hidalgo).................................................................................................................................... R. mexicana n. sp. 4 '. Dorsal alutaceous microsculpture present as longitudinal bands at interstrial intervals on elytra, these being somewhat expanded and merged at posterior fourth (Fig. 1 B); southeastern United States (Florida and Georgia)............... .............................................................................................................................................................. R. skelleyi n. sp. 5. Elytra with distinct traces of dorsal striae 1���3 and almost complete sutural striae; mesoventrite and anterior portion of metaventrite without medial longitudinal keel........................................................................ R. occidentalis n. sp. 5 ' Elytra without any traces of dorsal and sutural striae; mesoventrite and anterior portion of metaventrite (in males) with sharp medial longitudinal keel (Fig. 2 B)...................................................................................... R. cazieri n. sp., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 3-4, DOI: 10.5281/zenodo.191863

Renclasea occidentalis Tishechkin & Caterino, n. sp. (Figs. 7, 11, 12) Material examined: Holotype male: " NEW MEXICO: Catron Co.; Puebla Cmpgd S. of Reserve 9 -VI- 1987 Robert Gordon / HOLOTYPE Renclasea occidentalis sp. n. A. K. Tishechkin & M. S. Caterino des. 2008 " (USNM). Paratype (1): female from Arizona, Cochise Co., 5 mi N Benson collected by A. R. Hardy, F. G. Andrews and J. W. Smith on July 26, 1969 (CDFA). Diagnosis: This is one of two species without alutaceous background microsculpture on dorsal surface and presence of distinct traces of dorsal striae. It can be distinguished from smaller and less robust Renclasea falli by the absence of prosternal carinal striae. Description: L: 2.09; W: 1.60; E/Pn L: 1.76; E/Pn W: 1.16; Pn W/L: 1.60; E L/W: 0.94; Pr/Py: 1.07; Sterna: 0.46, 0.17, 0.47; Tibiae: 0.53, 0.62, 0.71 (n= 2). Body rufescent, shiny, smooth and asetose throughout except for a few setae on antennomeres 1���7. Frons and clypeus depressed at middle between lateral carinae; labrum narrowly rectangular, its apical margin straight, unmodified. Prosternal sides convergent, much more strongly in anterior fourth, above antennal cavities, weakly outwardly sinuate, with the anterior angles blunt, almost rectangular; marginal stria present along lateral edge, extending around anterolateral corner, interrupted at middle of anterior emargination of pronotum; pronotal lateral sides narrowly flattened and slightly reflexed; median angle of pronotal posterior margin about 110 ��. Prosternum with anterior margin of prosternal lobe almost straight; prosternal keel moderately elevated and flat, its base in male slightly excavated, carinal striae absent, weak basal fragments of lateral prosternal striae marked between procoxae in female specimen. Scutellum elongate triangular, small; elytra convex, widest at anterior third, with minute sparse background punctures, being denser and more conspicuous along sutural striae and at posterior fourth, where some merge into a weak background microsculpture; dorsal elytral striae 1���3 weakly marked on disc, abbreviated posteriorly; sutural stria abbreviated in anterior fifth. Mesoventrite flat in males, with low elevated area in median fourth incorporating metaventral projection in females, mesoventral projection short, wide triangular, its apex slightly elevated; mesometaventral suture 'curly bracket'-shaped, thin and inconspicuous; disc of metaventrite in male in anterior half with shallow oval depression, not reaching anterior parts of outer metaventral striae laterally; metaventral disc in females weakly, evenly convex. Propygidium weakly convex, disc with fine microsculpture of merged shallow dense punctures and short transverse striolets, alutaceous; marginal stria of propygidium complete; pygidium smooth, weakly convex, with striate ornament in females (Fig. 7 D). Male and female genitalia as illustrated (Figs. 11, 12, respectively). Etymology. The species epithet reflects its distribution in the American West. Distribution. Known from two localities in Arizona and New Mexico (Fig. 14). Remarks. Two female specimens from western Texas, both attracted to UV lights, from Fort Davis collected on May 30, 1959 by H. Howden and E. Becker (CMN) and from Randall Co., Palo Duro State Park at 34 �� 56.5 'N 101 �� 39.6 'W collected on May 15, 2002 by E. G. Riley (TAMU), are very similar to this species. Pygidial ornamentation and female genitalia of these Texas specimens are, however, sufficiently different from those of R. occidentalis n. sp., the most superficially similar (and potentially sympatric) species that we suspect them to represent an additional undescribed species. However, in this case we prefer to wait for the discovery of corresponding males from Texas before making a definite conclusion on their status., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2009, A new North American genus of Hetaeriinae (Coleoptera: Histeridae), with descriptions of six new species from the U. S. A. and Mexico, pp. 1-18 in Zootaxa 2311 on pages 13-14, DOI: 10.5281/zenodo.191863

Chlamydopsis caterinoi n. sp. (Figs 2-4) HOLOTYPE. — “ VANUATU: Santo I., Cumberland Peninsula, Saratsi Range at 14.9667°S 166.6560°E. 900m Flight intercept FL9C-13. 25-27 Nov 2006. A.K.Tishechkin AT791 / HOLOTYPE Chlamydopsis caterinoi sp. n. A.K.Tishechkin des. 2008”, ♂ pointmounted (MNHN). PARATYPES (2). — Same locality as the holotype, “Caterino DNA Voucher Extraction: MSC-1214 Species: Chlamydopsis Extraction Date: i.26.2007”, flight intercept trap, 10-11.XI.2006, A. K. Tishechkin coll., 1 ♂ (coll. AKT). — Saratsi Range, 14.9641°S, 166.6479°E, 600 m, flight intercept trap, 10-11.XI.2006, A. K. Tishechkin coll., 1 ♀ (MNHN). ETYMOLOGY. — Th e species is dedicated to my colleague and friend Mike Caterino of the Santa Barbara Museum of Natural History, in recognition of his great contributions to the taxonomy and systematics of Chlamydopsinae. DESCRIPTION L: 1.72; W: 1.22; E/Pn L: 2.06; E/Pn W: 1.24; Pn W/L: 1.62; E L/W: 1.02; Pr/Py: 0.80; sterna: 0.47, 0.09, 0.55; tibiae: 0.62, 0.65, 0.73. Body (Fig. 2A) elongate rectangular, dark rufescent brown, shining; most dorsal surfaces reticulostrigose, sparsely setose, setae faintly scale-like. Front of head (Fig. 3D), when retracted, slightly proclinate; frons with sides weakly rounded, about 1.2 times longer than wide, reticulately punctured, with sparse scattered scalelike setae; labrum about 2.3 times as wide as long, anterior margin obtusely triangular, disc shallowly reticulopunctate; mandibles strongly bent, with long narrow tips, with the same type of setae and smaller punctures on outer edges; maxillar palpi with four palpomeres, labial palpi with three palpomeres, mentum present as separate sclerite; antennal scape 2.2 times longer than wide, widest near basal third, disc faintly reticulate in proximal half, with scattered scale-like setae; antennal club of female about twothirds, that of male almost equals, scape length. Pronotum (Figs 2A; 3E) with posterior margin obtusely angulate, its sides approximately parallel in basal half, converging apically to one-half basal width; lateral pronotal margin slightly elevated, with several short, scale-like setae (absent from anterior margin); lateral and anterior pronotal margins with continuous, fine, deep groove just inside margin; pronotal disc moderately convex medially, flattened towards sides, entirely reticulostrigose, more elongately so posteromedially, except narrow smooth band along lateral elevation. Antennal cavities (Fig. 3E) broadly exposed from above, margined posteriorly with fine, almost continuous marginal stria, briefly interrupted at the middle, just anteriad of marginal groove of pronotal disc. Prosternum (Figs 2B; 3F) short, its anterior margin broadly, weakly emarginate, with fine marginal stria broadly interrupted in the middle; prosternal keel sharply rising in anterior two-thirds (reducing prosternal “depth” anteriorly), narrowed between procoxae, slightly emarginate posteriorly, bordered laterally by deeply impressed circumcoxal stria; anterior half of prosternal disc reticulostrigose, prosternal keel smooth. Scutellum (Figs 2A; 3A) distinct, small, triangular. Elytra (Figs 2A; 3A) with prominent humeral trichomes in basal third, with single broad, more or less flat, setose anterior elevation bearing anterior superficial stria along its outer edge, with a fringe of longer and denser setae along apex; apical edges of anterior and posterior trichome elevations meeting near their apices, both with dense, golden setal fringe beneath meeting point; posterior trichome elevations much smaller than anterior ones, conical, with a tuft of dense setae on inner edge near apex corresponding to a similar, but denser and larger tuft on inner edge of anterior elevation; epipleuron separated from elytral dorsum by finely impressed stria extending from posterolateral corner anteriorly, curving into opening of trichome; most of elytral dorsum posteriad of trichome elevation covered with dense longitudinal microsculpture consisting of short longitudinal ridges, sparsely setose, with reticulae more elongate across middle third of elytral dorsum; mediobasal depression and posterior sixth of elytral dorsum impunctate and glabrous; epipleuron mostly smooth, with reticulostrigose area confined to antero-dorsal corners; sutural striae continuous with marginal striae, abbreviated in basal third. Mesoventrite (Figs 2B; 3B) rather narrow, weakly projecting at middle, and extending forward around inner edge of mesocoxa; marginal stria obsolete at middle, visible mainly as oblique lateral fragments well mesal of mesocoxa; disc with small shallow punctures; mesometasternal suture and median metasternal sutures finely but deeply impressed; disc of metaventrite impunctate, smooth, asetose, with a few small punctures at sides and within mesotibial depressions, with complete arched lateral stria of metaventrite; first abdominal ventrite smooth, impunctate and asetose, with postcoxal stria deeply impressed, continuous across middle. Femora (Figs 2B; 3B) slightly widened apically, outer surface of profemur with few punctures along anterior margin near base, otherwise exposed surfaces of all femora impunctate, with fine sparse setae; protibia (Fig. 3C) broadly rounded, somewhat thickened just beyond middle, tarsal groove expanded, with single fovea near its midpoint (about one-third from apex of tibia), outer margin sinuous; posterior surface of tibia sparsely setose; posterior tibiae with outer edges rounded, mesotibia widest near middle, metatibia widest about two-thirds from base; tarsi slightly laterally compressed, about 0.7-0.8 times length of corresponding tibiae; tarsal claws simple, divergent, weakly arcuate, about 0.5 times length of corresponding apical tarsomere. Propygidium and pygidium (Fig. 3A) both weakly convex, both reticulopunctate, pygidium becoming smooth in apical half. Male genitalia as on Figure 4., Published as part of Tishechkin, Alexey K., 2009, Discovery of Chlamydopsinae (Insecta, Coleoptera, Histeridae) in Vanuatu with the description of eight new species from Espiritu Santo Island, pp. 661-690 in Zoosystema 31 (3) on pages 666-668, DOI: 10.5252/z2009n3a13, http://zenodo.org/record/5398808

Eucurtiopsis corbarai n. sp. (Fig. 5) HOLOTYPE. — “ VANUATU: Santo I., Cumberland Peninsula, Saratsi Range at 14.9626°S 166.6485°E. 600 m Flight intercept FL6B-5. 9-10 Nov 2006. A. K.Tishechkin AT648 / HOLOTYPE Eucurtiopsis corbarai sp. n. A. Tishechkin des. 2008”, ♀ point-mounted (MNHN). PARATYPES (3). — Same locality and date as the holotype, flight intercept trap, A. K. Tishechkin coll., 1 ♀ (coll. AKT). — Saratsi Range, 14.9657°S, 166.6521°E, 700 m, flight intercept trap, 30.XI-1.XII.2006, A. K.Tishechkin coll., 1 ♀ (MNHN). — Saratsi Range, 14.9667°S, 166.6560°E, 900 m, flight intercept trap, 16.XI.2006, A. K. Tishechkin coll., also bears the following label “DNA Extraction TAK-0003. May 2009 Baton Rouge”, 1 ♀ (MNHN). ETYMOLOGY. — Th e species honours Bruno Corbara of Université Blaise Pascal, Clermont-Ferrand, a true leader of the IBISCA Team, in appreciation of his inspiring enthusiasm for bringing people together to study poorly known tropical insect communities and great moments of joint fieldwork across the globe. DESCRIPTION L: 1.36; W: 0.90; E/Pn L: 1.93; E/Pn W: 1.56; Pn W/L: 1.18; E L/W: 1.05; Pr/Py: 0.89; Sterna: 0.42, 0.08, 0.37; Tibiae: 0.36, 0.42, 0.46. Body (Fig. 5A) elongate, rufescent brown, with antennae and legs somewhat paler, prothorax substantially narrower than elytra. Frons (Fig. 5D) 1.5 times longer than wide, sides weakly arcuate, incised at antennal bases, narrowed anteriorly, covered with deep, dense, mostly elongate punctures, with two parallel, longitudinal rows consisting of four irregular blunt tubercles arranged into indistinct ridges, each tubercle with a cluster of long branched setae; labrum weakly convex, semicircular, with few setae, punctation similar to that on frons; mandibles strongly bent, with long narrow tips, with few setae and small punctures on smooth outer edges; maxillar palpi with three palpomeres, labial palpi with two palpomeres, mentum apparently present as separate sclerite, separating suture present at least in laterobasal areas; antennal scape elongate triangular, 1.4 times longer than wide, with inner edge weakly inwardly arcuate, its surface more or less flat, with apical angle bluntly rounded, disc densely punctuate throughout with oval punctures, with conspicuous long branched setae; antennal funicle and club (of female) about one-half, and three-quarters length of scape, respectively. Pronotum (Fig. 5A, C) with posterior margin shallowly obtusely angular, with sides unmargined, straight, faintly narrowed anteriorly; antennal cavities partially visible from above, with pronotal margin not elevated above; medial portion of pronotal margin unelevated, weakly arcuate; marginal striae visible from above only near base, then abruptly descend downwards to meet supracoxal striae, ascend again anteriorly towards antennal sockets; pronotal dorsum strongly convex, densely and deeply punctate throughout, with eight parallel, longitudinal rows of blunt tubercles arranged into indistinct ridges, two outer ridges on each side much less regularly linearly arranged than discal ridges, each ridge consisting of four or five tubercles, clumps of elongate branched setae present on every tubercle, several single setae present along lateral sides of pronotum. Prosternum (Fig. 5B) with anterior margin broadly concave, marginal stria represented by series of elongate wrinkles; prosternal disc evenly convex, keel elevated between procoxae, flattened, slightly expanded posteriorly, emarginate at apex, disc densely punctate throughout, punctures primarily elongate, short branched setae scattered throughout the prothorax surface. Scutellum (Fig. 5A) tiny, sunk below the elytralpronotal plane, very poorly visible. Elytra (Fig. 5A, C) with sides weakly arcuate, widest around anterior trichome process; humeral trichome prominent, elevated, longitudinally oriented, anterior process rising almost vertically at base till transverse ridge, weakly obliquely rising thereafter, surface of its anterior part weakly convex with slight transverse concavity in the middle; posterior process less robust, evenly rising towards anterior one, its surface evenly convex; trichome with setae only along apical edges of processes,shallowly excavate beneath setose fringe, width of the fringe more than one-third of elytral width, trichome gap wide,about one-seventh of elytra length, fringe setae long and densely packed; dorsum of elytral disc with punctation occupying most of posterior half (although only few punctures present at lateral margins near posterior one-fourth) and a small cluster of punctures around scutellum with a single row of punctures extending to the middle of anterior elytral margin, no punctation present on and between trichome elevations except of mentioned prescutellar cluster; punctures deep and elongate, sparsely spaced, branched setae present throughout, being larger and clustered into separate clumps or looser groups in punctated areas and smaller and evenly distributed across smooth surfaces; sutural stria thin, but distinct, abbreviate in posterior onethird; epipleuron smooth and glabrous, with sparsely scattered short branched setae; marginal epipleural stria distinct, elevated above metafemur, continuous with complete marginal elytral stria; no traces of accessory epipleural stria present. Mesoventrite (Fig. 5B) wide, short, weakly convex, bluntly projecting at middle, no traces of marginal stria present, surface with dense, deep, elongate punctures; mesometaventral suture and median suture of metaventrite finely impressed, complete, but inconspicuous, lateral stiae of metaventrite present, elevated, enclosing largely impunctate depression for mesotibiae in repose; disc of metaventrite punctate throughout, primarily with circular punctures, with larger, denser and more elongate along anterior and lateral margins; first abdominal ventrite similarly punctate, punctures being on average larger and denser than on disc of metaventrite, with postmetacoxal line originating at metacoxa, extended directly posteriorly close to edge of sternite, curving laterad, terminating freely just before reaching epipleuron; surfaces of meso- and metaventrite and first abdominal ventrite with scattered short inconspicuous branched setae. Profemora (Fig. 5B, C) with dense punctures in basal two-thirds, becoming impunctate toward apex, with posterior margins obtusely angular in basal third, meso- and metafemora impunctate, their margins arcuate, metafemora much more robust than mesofemora; protibia with prominent angle at basal one-third of outer margins, meso- and metatibiae with outer margins more bluntly angulate, around midpoint; all tibiae longitudinally convex, meso- and metatibia with longitudinal sulci along inner edge; tarsi weakly compressed laterally about 0.6-0.7 times length of corresponding tibiae; tarsal claws simple, divergent, almost straight, about 0.4 times length of corresponding apical tarsomere. Propygidium twice as wide as midline length, weakly convex; pygidium nearly as long as wide, weakly convex; both with punctures and setae as on the posterior half of elytral disc., Published as part of Tishechkin, Alexey K., 2009, Discovery of Chlamydopsinae (Insecta, Coleoptera, Histeridae) in Vanuatu with the description of eight new species from Espiritu Santo Island, pp. 661-690 in Zoosystema 31 (3) on pages 668-670, DOI: 10.5252/z2009n3a13, http://zenodo.org/record/5398808

Eucurtiopsis pascali n. sp. (Figs 12; 13) HOLOTYPE. — “ VANUATU: Santo I., Cumberland Peninsula, Saratsi Range at 14.9638°S 166.6362°E. 500 m. Flight intercept FL5A-1. 27-28.xi 2006. A. K.Tishechkin AT796 / HOLOTYPE Eucurtiopsis pascali sp. n. A. Tishechkin des. 2008”, ♂ point-mounted (MNHN). PARATYPES (29). — Same locality as the holotype, flight intercept trap, 27.XI-1.XII.2006, A. K.Tishechkin coll., one also bears the following label “Caterino DNA Voucher Extraction: MSC-1213 Species: Eucurtiopsis Extraction Date: i.26.2007”, 11 ♂♂ (MNHN), 9 ♂♂ (coll. AKT [including DNA voucher]), 2 ♂♂ (LSAM). — Saratsi Range, 14.9626°S, 166.6485°E, 300 m, flight intercept trap, 23.XI-1.XII.2006, A. K.Tishechkin coll., one also bears the following label “Caterino DNA Voucher Extraction: MSC-1230 Species: Eucurtiopsis Extraction Date: ii.2.2007”, 6 ♂♂ (MNHN [including DNA voucher]), 1 ♂ (coll. AKT). ETYMOLOGY. — Th e species is named after Olivier Pascal of Pronatura International in recognition of his enthusiastic efforts in organization and support of biotic inventories and conservation of tropical forests and commemoration of our joint field experience in Panamá and Vanuatu. DESCRIPTION L: 1.87; W: 1.26; E/Pn L: 1.75; E/Pn W: 1.35; Pn W/L: 1.36; E L/W: 1.00; Pr/Py: 0.94; sterna: 0.61, 0.11, 0.47; tibiae: 0.56, 0.61, 0.61. Body (Fig. 12A) rectangular, robust, dark reddish brown, with antennae and legs somewhat paler, prothorax substantially narrower than elytra, entire body surface with scattered or clustered minute, appressed, inconspicuous scale-like setae. Frons (Fig. 12D) 1.2 times longer than wide, sides nearly straight, incised at antennal bases, narrowed anteriorly, covered with deep, small, sparse, circular punctures, with two parallel longitudinal rows consisting of three irregular blunt tubercles arranged into indistinct ridges, each tubercle with a cluster of setae; labrum weakly convex, semicircular, with few setae and punctures similar to that on frons; mandibles strongly bent, with long narrow tips, with few setae and deep small punctures on smooth outer edges; maxillar palpi with three palpomeres, labial palpi with two palpomeres, mentum present as separate sclerite; antennal scape elongate triangular, 1.4 times longer than wide, with inner edge weakly inwardly arcuate, its surface more or less flat, with apical angle bluntly rounded, outer edge with blunt strong angle, disc densely punctate throughout with circular, dense punctures, with setae present in anterior half; antennal funicle and club (of male) about two-thirds of, and same length as scape, respectively. Pronotum (Fig. 12A, C) with posterior margin shallowly obtusely angular, with sides unmargined, straight, faintly narrowed anteriorly; antennal cavities partially visible from above, with pronotal margin not elevated above; medial portion of pronotal margin unelevated, straight; marginal striae visible from above in basal third, then abruptly descending downwards to meet supracoxal striae, ascending again anteriorly towards antennal sockets; pronotal dorsum strongly convex, densely and deeply punctate throughout with small circular punctures, with eight parallel, longitudinal rows of blunt tubercles arranged into indistinct ridges, two outer ridges on each side much less regularly linearly arranged than discal ridges, each ridge consisting of 4-6 tubercles, clumps of short setae present on every tubercle, two similar tubercles present on each lateral lateral margin in median third. Prosternum (Fig. 12B) with anterior margin broadly concave, marginal stria complete; prosternal disc evenly convex, keel in the same plane as the rest of prothorax, flattened, slightly expanded posteriorly, emarginate at apex, disc sparsely punctate throughout, punctures circular, intervals raised creating honeycomb sculpture, intervals bearing setae throughout the prothorax surface. Scutellum (Fig. 12A) invisible. Elytra (Fig. 12A, C) with sides weakly arcuate, widest around trichome gaps; humeral trichome prominent, elevated, longitudinally oriented, both processes evenly rising obliquely, conical in shape, surface of anterior process deeply concave, of posterior one convex; trichome deeply excavate beneath setose fringe, with setae along all edges of trichome gap and narrow triangular inward expansion on elytral disc plane reaching middle of elytron width, width of the fringe about half of elytral width, trichome gap wide, about one-fifth of elytra length, fringe setae long and densely packed, short and even denser in inward extensions; dorsum of elytral disc strongly convex, with punctuation occupying most of posterior half reaching lateral margins and bases of posterior trichome elevations and narrow band of punctures along anterior margin, narrow band of punctures may connect those two areas along elytral suture, varying from four or five punctures wide band to total absence of punctutes; punctures small, deep and elongate, sparsely spaced, setae sparsely scattered throughout, being more abundant in anterior third; sutural stria fine, but distinct, complete; epipleuron smooth and glabrous, with sparsely scattered setae and few punctures along margin in posterior half; marginal epipleural stria distinct, elevated above metafemur, continuous with complete marginal elytral stria; no traces of accessory epipleural stria present. Mesoventrite (Fig. 12B) wide, short, weakly convex, bluntly projecting at middle, no traces of marginal stria present, surface with dense, shallow, circular punctures; mesometaventral suture complete, distinct, median suture of metaventrite finely impressed, abbreviated both anteriorly and posteriorly, lateral striae of metaventrite present, elevated, enclosing largely impunctate depression for mesotibiae in repose; disc of metaventrite punctate around margins, with large, ocellate, circular punctures, shallow longitudinal impression present (in males) along longitudinal suture; first abdominal ventrite similarly punctate, punctures being on average smaller and scattered throughout disc, with postmetacoxal line originating at metacoxa, extended directly posteriorly close to edge of sternite, curving laterad, terminating freely just before reaching epipleuron; surfaces of meso- and metaventrite and first abdominal ventrite with sparsely scattered setae. Profemora (Fig. 12B, C) with dense punctures, with posterior margins obtusely angular in basal third, meso- and metafemora with row of puntures along margins, those arcuate, metafemora much more robust than mesofemora; protibia with prominent angle around midpoint of outer margins, meso- and metatibiae with outer margins more bluntly angulate, around midpoint; all tibiae longitudinally convex, meso- and metatibia with longitudinal sulci along inner edge; tarsi weakly compressed laterally about 0.6-0.7 times length of corresponding tibiae; tarsal claws simple, divergent, almost straight, about 0.4 times length of corresponding apical tarsomere. Propygidium twice as wide as midline length, weakly convex; pygidium nearly as long as wide, weakly convex; both with punctures and setae as on the posterior half of elytral disc, but spaced more sparsely. Male genitalia as on Figure 13., Published as part of Tishechkin, Alexey K., 2009, Discovery of Chlamydopsinae (Insecta, Coleoptera, Histeridae) in Vanuatu with the description of eight new species from Espiritu Santo Island, pp. 661-690 in Zoosystema 31 (3) on pages 680-682, DOI: 10.5252/z2009n3a13, http://zenodo.org/record/5398808

Key to the species of Hippeutister 1 Elytral disc bearing setae (Figs. 1 A, 1 C, 4 A, 4 C)....................................................................................... 2 - Elytral disc glabrous; epipleura may be setose............................................................................................ 3 2 Setae of elytron (in part) clearly marking the basal portions of elytral striae 1–3 as well as sutural stria, otherwise evenly distributed (Figs. 1 C, 4 C); dorsal setae borne in elevated pustules, somewhat flattened, faintly scale-like (Figs. 4 C, 5 C) .................................................................................. H. castaneus (Lewis) - Setae of elytron more randomly scattered, not obviously marking any elytral striae, and denser in posterior half (Figs. 1 A, 4 A); setal bases not elevated, setae finer, not at all scale-like ...... H. californicu s n. sp. 3 Elytral epipleuron and lateral pronotal margin bearing dense rows of setae (Figs. 5 B, 8 B) H. solisi n. sp. - Elytral epipleuron and lateral pronotal margin glabrous............................................................................. 4 4 Body moderately convex (Figs. 8 A–C); elytron with sutural stria marked by row of distinctly larger punctures; pronotum with few or no superficial wrinkles on surface................................................................... ............................................................................. H. plaumanni Reichensperger and H. amabilis (Wenzel) - Body strongly convex (Fig. 8 D); elytron densely punctate, but punctures in area of sutural stria not standing out; pronotum with distinct transverse to oblique wrinkles in anterior corners (Fig. 5 D)...................... .................................................................................................................................... H. manicatus (Lewis)

Hippeutister manicatus (Lewis, 1891) (Figs. 1 D, 4 D, 5 D, 6 F, 7 D, 8 D) Paratropus manicatus Lewis, 1891: 390. Lectotype male, hereby designated: " Mexico, A.G."/"G. Lewis Coll., B.M. 1926 - 369 "/" Paratropus manicatus Lewis Type "/" Hippeutister manicatus (Lewis) nov. comb. 1988, N. Degallier"/" LECTOTYPE Paratropus manicatus Lewis M. Caterino & A. Tishechkin des. 2008 ", BMNH. Hippeutister solenopsidis Reichensperger, 1935: 208. Lectotype, hereby designated: "La Caja, X. 1934 "/" LECTOTYPE Hippeutister solenopsidis Reichensperger M. Caterino & A. Tishechkin des. 2008 ", USNM. Paralectotype, hereby designated: "La Caja, X. 1934 "/" PARALECTOTYPE Hippeutister solenopsidis Reichensperger M. Caterino & A. Tishechkin des. 2008 ", USNM. New Synonomy. Hippeutister manicatus; Mazur, 1997: 153. Diagnosis: L: 2.94; W: 2.95; E/Pn L: 1.50; E/Pn W: 1.26; Pn W/L: 1.99; E L/W: 0.60; Pr/Py: 1.11; Sterna: 0.79, 0.26, 0.76; Tibiae: 0.80, 0.90, 1.03 (n= 3). This is probably the most readily recognized species of Hippeutister. It has a very distinctive body form, being very strongly convex dorsally (best seen in lateral view: Fig. 8 D), and more broadly rounded (Fig. 4 D) than any of the others. It is also the largest species of the genus, and is completely glabrous and thoroughly punctate, both dorsally (Fig. 4 D) and ventrally (Fig. 6 F). Its mesosternal projection is less strongly acute anteriorly, and the median-most pair of prosternal keel setae are strongly abbreviated from the base relative to most other species. In these last characters it is rather similar to H. plaumanni, from which it is nonetheless still easily separated by body shape alone. Distribution: This species is known from the type locality ('Mexico'; Lewis, 1891), and from Costa Rica (three localities in the provinces Limon and San Jose). Although not mounted with the beetle, and therefore unconfirmed, the original description and FIMAK specimens of this species indicate they were collected with Solenopsis geminata (Fabricius). Reichensperger (1935) mentioned that H. Schmidt reported to him that he found type specimens either 'riding large soldiers of Solenopsis, or sitting quietly [in the colony].' Material examined: 13 specimens (excluding type material) from Costa Rica; San Jose Province, Farm La Caja near San Jose, all collected by H. Schmidt: three in July 1936 (FMNH and USNM), one in October 1937 (USMN); San Jose Province, San Jose, all collected by H. Schmidt: five in September 1935 and four without date (all FIMAK); one from Limon Province, Hamburg Farm, collected on September 8, 1935 by F. Nevermann (USNM)., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 46-47, DOI: 10.5281/zenodo.184407, {"references":["Lewis, G. (1891) On new species of Histeridae. Annals and Magazine of Natural History, (6) 8, 380 - 405.","Reichensperger, A. (1935) Beitrag zur Kenntnis der Myrmekophilenfauna Brasiliens und Costa Ricas III. (Col. Staphyl. Hist.). Arbeiten uber Morphologische und Taxonomische Entomolgie aus Berlin-Dahlem, 2, 188 - 218.","Mazur, S. (1997) A World Catalogue of the Histeridae. Genus, International Journal of Invertebrate Taxonomy (Supplement), 1997, 1 - 373."]}

Hippeutister amabilis (Wenzel, 1938) (Fig. 1 E) Solenopsister amabilis Wenzel, 1938: 318; Blackwelder, 1944: 186. Diagnosis: L: 1.5mm; W: 1.25mm (measurements from Wenzel, 1938). Wenzel initially described this species in its own monotypic genus, having been unaware of Reichensperger's then recent description of two species of Hippeutister. Thus he offered no putatively diagnostic species level characters. Based on his description and illustrations the species is clearly correctly placed in Hippeutister. Like only H. manicatus and H. plaumanni, it is evidently completely glabrous, and it seems very similar to the latter in general, showing a distinct sutural row of enlarged punctures. However, Wenzel's figures suggest this species is more densely punctate than H. plaumanni or any of the others of the genus. Distribution: This species has been reported only from the type locality, Cordoba, Veracruz, Mexico where it was collected with Solenopsis xyloni McCook (Wenzel, 1938). Remarks: Unfortunately the type of this species appears lost. Having initially been collected by Charles Seevers and then described by Rupert Wenzel, both Field Museum (Chicago, IL) associates, that seems the only likely repository, although Wenzel did not specify it in his description. However, it has not been found there., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 49-50, DOI: 10.5281/zenodo.184407, {"references":["Wenzel, R. L. (1938) New and little known Neotropical Hetaeriomorphini (Coleoptera: Histeridae). Revista de Entomologia, 9, 317 - 321.","Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Bulletin of the USNM, 185 (1 - 6), 1 - 1492."]}

Hippeutister californicus Caterino & Tishechkin, n. sp. (Figs. 1 A, 2 A, 3 A, 3 C, 4 A, 5 A, 6 A���B, 7 A, 8 A) Holotype of undetermined sex: "CA: San Diego Co., 32.7517 ��N, 116.6641 ��W, Cleveland NF, Horsethief Cyn., i. 6.2005, M. Caterino"/"CA BEETLE PROJ, CBP0056475", SBMNH. Diagnosis: Apart from its disjunct distribution, this species can be identified by the fine setae on most body surfaces, especially on the elytral dorsum (Figs. 1 A, 4 A, 8 A). The only other species with dorsal (as opposed to epipleural) elytral setae is H. castaneus, in which they are coarse, fewer, and borne in prominent raised pustules (Fig. 4 C). In most other characters it is very similar to the other new species, H. solisi, not only in discrete characters, but also in size (being smaller than the other species of the genus). Hippeutister solisi has much denser setae along the pronotal and epipleural margins. Some very slight differences in mesosternal striae of the two may be significant, but with only a single specimen of each known, may not be reliable. Description: L: 2.46; W: 2.12; E/Pn L: 1.72; E/Pn W: 1.17; Pn W/L: 2.00; E L/W: 0.74; Pr/Py: 0.96; Sterna: 0.65, 0.25, 0.59; Tibiae: 0.69 0.69, 0.81 (n= 1). Body rounded (Fig. 1 A, 4 A), moderately convex (Fig. 8 A), narrowed anteriorly (Figs. 4 A, 5 A), pale rufescent and sparsely setose throughout. Frons (Fig. 7 A) with longitudinal carinae parallel on epistoma, bent laterad, then obliquely over antennal insertions, ending freely before meeting ocular carina along inner margin of eye; both of these carinae bearing moderately dense row of short fine setae; frons flat to depressed at middle, and distinctly depressed along inner margin of eye, surface sparsely setose, each seta arising from a fine puncture; labrum flat, apical margin weakly emarginate, bearing numerous setae near apex; mandibles with numerous setae on sides; antenna with scape pyramidal, with all edges sharp, weakly produced, and bearing marginal setae; antennal club nearly cylindrical (Fig. 3 B), slightly flattened, and obliquely truncate at the apex, the apical surface densely tomentose. Prothoracic sides strongly convergent (Fig. 5 A), weakly sinuate, with the anterior angles distinctly truncate; lateral margin bearing setae between elevated marginal and lateral striae; marginal stria not quite attaining base, extending around anterolateral corner, ending at anteromedian corner behind eye; lateral stria barely extending around basal corner, paralleling marginal stria, meeting it at anterolateral corner; anterior marginal stria present behind head, nearly meeting lateral marginal stria in anteromedial corner, replaced at middle behind head by a short, transverse striole; pronotal surface obliquely depressed on each side, glabrous, with scattered minute punctures, a few faint oblique wrinkles on each side, and several rather large but very shallow ocellate punctures, with their posterior edges effaced, in front of the scutellum. Prosternum broad, (Fig. 6 A) with prosternal lobe strongly deflexed, its anterior margin striate at middle and bearing fine marginal setae; prosternum otherwise glabrous. Scutellum distinct, shallowly longitudinally depressed; elytra convex, widest at middle, with setae sparsely scattered over entire surface, becoming more dense posteriorly (Fig. 4 A), each seta suberect, directed posterad, short, very weakly flattened, arising from a shallow but fairly broad puncture; elytral striae not marked apart from three distinct epipleural carinae (Fig. 8 A), one of which delimits the dorsal edge of the elytron and is complete from base to apex, the middle on the epipleural surface and also complete, and the outer-most extending from the humeral corner only about one third of the length of the epipleuron, where it meets the elytral margin. Meso- and metasterna (Fig. 6 B) with few fine setae laterally, depressed and coarsely punctate on and behind mesometasternal suture; mesosternal projection triangular, with fine marginal stria; two lateral metasternal striae present, one along anterior margin, one curving from anterior margin posterad along mesocoxa; metaventrite with three oblique lateral striae, two arising behind inner corner of mesosternal projection, one extending laterad to mesocoxa, thence forming postmesocoxal stria, the other extending straight to metacoxa, the third arising from the first at the inner edge of the mesocoxa and ending freely in front of metacoxa. First visible abdominal sternite with crenulate punctures along anterior margin, and two oblique postmetacoxal striae. Femora of all legs short, broad and subquadrate, with scattered fine setae on outer surfaces, the meso-and metafemora bearing anterior marginal and submarginal striae; protibiae broad, parallel sided in apical twothirds, with distinct marginal spines; meso- and metatibiae more rounded laterally, slightly narrowed to apex, lacking marginal spines, with complete marginal striae and bearing fine setae on outer face; tarsomeres 1���4 bearing pair of apicoventral setae; pretarsal claws straight. Propygidium flat across base, weakly convex along apical margin, bearing striae along anterolateral margins, disc with fine setose punctures throughout; pygidium (Fig. 3 C) smooth, convex, with fine setose punctures throughout. Distribution: This species is only known from the type locality (Fig. 2 A), in a hilly region covered with chaparral (shrubby sclerophyllous, xerophilous) vegetation, at about 2100ft elevation, just a few miles north of the Mexican border. This region of the California Floristic Province is rather unusual in receiving a significant portion of its annual precipitation during the summer months, possibly accounting for this more tropical genus's presence there. The type was collected on the underside of a rock with a colony of Solenopsis amblychila Wheeler during the early part of the winter rainy season. The type was collected in 2005 and the entire site burned in 2006 in the 'Horse Fire'. Attempts to recollect the species at the type locality in 2007 were unsuccessful, and no host ants were found. Remarks: DNA has been extracted from the head and prothorax of the type specimen, and they are mounted on a separate point from the remainder of the body. This DNA resides in the SBMNH tissue collection under extract #MSC- 1027., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 41-43, DOI: 10.5281/zenodo.184407

Hippeutister solisi Caterino & Tishechkin, n. sp. (Figs. 1 B, 2 B, 3 B, 3 D, 4 B, 5 B, 6 C, 7 B, 8 B) Type material: Holotype female: ���Cano Negro, 20m, R.N.V.S. Cano Negro, Prov. Alajuela, Costa Rica, 4 a 17 dic 1992, K. Flores, L-N 319100, 450200 ���/��� INBIO CRI 000911827���, INBIO. Diagnosis: This species is most similar to the other new species, H. californicus, being only slightly larger in size, similar in body shape, and in being somewhat setose. This species is easily distinguished by lacking dorsal setae on the pronotum and elytra, but with dense setae on the margins (the epipleuron in the latter) of both (Fig. 8 B). Description: L: 2.83; W: 2.49; E/Pn L: 1.76; E/Pn W: 1.19; Pn W/L: 2.03; E L/W: 0.73; Pr/Py: 1.00; Sterna: 0.78, 0.28, 0.62; Tibiae: 0.72, 0.75, 0.84 (n= 1). Body rufescent (Fig. 1 B), moderately convex (Fig. 8 B), rounded (Fig. 4 B), widest just behind humeri. Frons (Fig. 7 B) with about 40 fine setose punctures randomly scattered, these slightly more dense near vertex, and almost absent from epistoma; epistomal carinae well developed, sinuate over antennal bases, extending obliquely to ocular carina, both bearing denser fringe of setae; clypeolabral suture not evident; labrum truncate, bearing few discal and apical setae; mandibles finely setose on lateral surfaces. Prothorax (Fig. 5 B) widest at base, converging strongly and weakly sinuately to obliquely trunctate anterior angles; pronotal marginal stria present, extending to anterior corner; lateral pronotal stria close to marginal, extending around anterior angle along anterior margin, interrupted briefly behind head where a short tangential discal stria replaces it; lateral margin of pronotum with short, dense fringe of setae between marginal and lateral striae; disc of pronotum obliquely depressed along sides, faintly explanate, glabrous, smooth, with faint, shallow but broad punctures in front of scutellum. Prosternal keel (Fig. 6 C) broad, bearing a few setae; prosternal lobe short, not produced at middle, with moderately dense setose punctures near anterior margin. Elytra convex, lacking dorsal striae (Fig. 4 B), though with very faint oblique impressions near base, disc glabrous, with broad, shallow punctures near scutellum, and finer, denser ones extending posterad along suture; epipleuron margined dorsally by complete carinate stria and with less prominent subdorsal and lateral striae, the entire epipleuron finelly setose, especially on and immediately below striae (Fig. 8 B). Mesometasternal suture only faintly indicated at middle (Fig. 6 C); mesosternum projecting strongly into triangularly incised base of prosternal keel, with fine marginal stria continuous with complicated array of several oblique lateral mesometasternal striae; metasternal disc with numerous rather deep setose pits anteromedially, becoming finer and sparser posteriorly; lateral portions of metasternal disc with more numerous but finer setose punctures lining and between the lateral striae. First visible abdominal sternite with sparse moderately deep setose pits particularly near anterior margin, with finer but denser setose punctures laterally, between and mediad the two oblique lateral striae. Profemora (Fig. 8 B) very broad, nearly square, margined along anterior and posterior edges, with setose punctures on ventral (posterior) face; protibia broad, rounded, subangulate near base, with series of fine spines along apical two-thirds of margin; meso- and metafemora short, with anterior and posterior margins rounded, with marginal and anterior submarginal striae on ventral (anterior) face, finely setose throughout; meso- and metatibiae more rounded laterally, slightly narrowed to apex, lacking marginal spines, with complete marginal striae and bearing fine setae on outer face; tarsomeres 1���4 bearing pair of apicoventral setae; pretarsal claws straight. Propygidium flat to weakly impressed across base, notably convex along apical margin, bearing striae along anterolateral margins extending just around anterolateral corners, disc with fine setose punctures throughout; pygidium (Fig. 3 D) faintly depressed along posterolateral margins, with fine setose punctures throughout. Distribution: This species is only known from the type locality, in Alajuela province, north central Costa Rica (Fig. 2 B). There is no host data associated with the specimen. Remarks: We name this species for Angel Sol��s, of the Instituto Nacional de Biodiversidad in Costa Rica, in recognition of his efforts to advance knowledge and conservation of Costa Rican Coleoptera., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 43-45, DOI: 10.5281/zenodo.184407

Hippeutister plaumanni Reichensperger, 1936 (Figs. 1 E, 6 G) Hippeutister plaumanni Reichensperger, 1936: 226. Lectotype female, hereby designated: "F. Plaumann, Nova Teutonia Brasilien, 7.35 [July 1935]"/" Solenopsis saevissima "/" TYPUS "/" Hippeutister plaumanni Reichensp. "/"LECTO- TYPE Hippeutister plaumanni Reichensperger M. Caterino & A. Tishechkin des. 2008 ", FIMAK; Paralectotype, probably female: same data as lectotype, FIMAK. Diagnosis: L: 2.74; W: 2.40; E/Pn L: 1.59; E/Pn W: 2.85; Pn W/L: 0.79; E L/W: 0.70; Pr/Py: 1.00; Sterna: 0.75, 0.28, 0.69; Tibiae: 0.72, 0.78, 0.87 (n= 1). H. plaumanni and H. amabilis appear very similar, and as the latter is known from no extant specimens, a confident diagnosis separating these two is difficult to make. Both appear to be completely glabrous, like only H. manicatus, but they are much less strongly convex, and easily separated from it. The pronotum, elytra, frons and venter of H. amabilis are illustrated as densely though finely punctate, and are described as 'rather sparsely, finely punctate' (pronotum) or 'finely, rather shallowly, though not sparsely punctate' (elytra). Wenzel further describes the punctation of the pronotum as 'finely ocellate' under high magnification, though he offers some disclaimer that the punctation is 'not [..] quite so obvious on the sides of the pronotum [..] as shown in the figure'. In all cases, the punctation of H. plaumanni is so fine as to be almost invisible, except in the shared vague depressions along the elytral suture, and small groups of conspicuous punctures along the median metasternal suture, and these differences are probably real and diagnostic. The mesoventrite of H. plaumanni is rather more bluntly than acutely triangular in front, and is strongly depressed (Fig. 6 G). As in H. manicatus, the median-most pair of prosternal striae is strongly abbreviated from the base. Distribution: This species is only known from topotypical material, from Nova Teutonia, in the state of Santa Catarina, Brazil as a guest of Solenopsis saevissima (Smith). Material examined: five specimens (excluding type material), all collected by F. Plaumann in Nova Teutonia, one on July 30, 1952, one on August 3, 1952 (both FMNH), and two on August 9, 1953 (FMNH and USMN)., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 47-49, DOI: 10.5281/zenodo.184407, {"references":["Reichensperger, A. (1936) Beitrag zur Kenntnis der Myrmekophilen- und Termitophilenfauna Brasiliens und Costa Ricas. IV. (Col. Hist. Staphyl. Pseplaph.). Revista de Entomologia, 6, 222 - 242."]}

Hippeutister castaneus (Lewis, 1891) (Figs. 1 C, 4 C, 5 C, 6 D���E, 7 C, 8 C) Paratropus castaneus Lewis, 1891: 391. Lectotype, hereby designated: ��� Mexique ���/ ���G. Lewis Coll. B.M. 1926 - 369.���/ ��� Syntype ���/ ��� Type ���/ ��� Paratropus castaneus Lewis Type ���/" LECTOTYPE Paratropus castaneus Lewis M. Caterino & A. Tishechkin des. 2008 ", BMNH; Paralectotype: ��� Mexique ���/ ���G. Lewis Coll. B.M. 1926 - 369.���/ ��� Syntype ���/ ��� Type ���/ ��� Paratropus castaneus Lewis Co-type���/ " PARALECTOTYPE Paratropus castaneus Lewis M. Caterino & A. Tishechkin des. 2008 ", BMNH. Diagnosis: L: 2.88; W: 2.62; E/Pn L: 1.68; E/Pn W: 1.19; Pn W/L: 2.04; E L/W: 0.69; Pr/Py: 0.97; Sterna: 0.84, 0.27, 0.70; Tibiae: 0.67, 0.80, 0.88 (n= 3). This species is easily recognized by the unique setose 'pustules', enlarged, slightly raised pits bearing conspicuous, faintly scale-like setae. These are observed on essentially all body surfaces, but especially the frons, pronotum and elytra. In addition to being coarser, these setae are fewer and sparser than in the only other dorsally setose species, H. californicus. Distribution: This species is known from the unspecified type locality in Mexico (Lewis, 1891), and from recent material from northwestern Belize. No specimens of the species have been associated with host ants. Material: one female (excluding type material) from Belize: Orange Walk District, Rio Bravo Conservation Area, 2 nd Logging Rd. off main Rd. to Archeological Site, collected with flight intercept traps by P.W. Kovarik on July 18 ���20, 1996 (PWK)., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on pages 45-46, DOI: 10.5281/zenodo.184407, {"references":["Lewis, G. (1891) On new species of Histeridae. Annals and Magazine of Natural History, (6) 8, 380 - 405."]}

Hippeutister Reichensperger, 1935 Solenopsister Wenzel, 1938: 318; Blackwelder, 1944: 186. Type species: Solenopsister amabilis. Type species: Hippeutister solenopsidis Reichensperger, 1935: 208 (= H. manicatus Lewis), by monotypy. Diagnosis: Hippeutister is easily recognizable among Hetaeriinae, and almost undoubtedly monophyletic. It is most easily recognized by the broad, weakly convex prosternum with strongly divergent carinal striae, its deeply, triangularly emarginate prosternal keel, and by the correspondingly broad mesosternal projection (Fig. 6). A number of other features are more unusual among neotropical Hetaeriinae, and are also useful in recognizing members of Hippeutister. They have a cylindrical, obliquely truncate antennal club (Fig. 3 B), the frons is longitudinally depressed along inner edge of eye, resulting in a distinct ocular carina (Fig. 7), the prosternum has an extra, third pair of carinal striae (Fig. 6), the pronotum exhibits oblique depressions extending forward from posterior corners (Figs. 4, 5), the elytra lack striae (though they may be weakly marked by punctures or setae), and the dorsum of the elytra is margined by a strong epipleural carina (Figs. 4, 8). It is most similar to members of the genus Poneralister Bruch, which are identical in characters of the frontal carinae. The pro- and mesosternal characters of Hippeutister separate the two genera easily., Published as part of Caterino, Michael S. & Tishechkin, Alexey K., 2008, A review of Hippeutister Reichensperger with new species from California and Costa Rica (Coleoptera: Histeridae: Hetaeriinae), pp. 39-52 in Zootaxa 1895 on page 41, DOI: 10.5281/zenodo.184407, {"references":["Reichensperger, A. (1935) Beitrag zur Kenntnis der Myrmekophilenfauna Brasiliens und Costa Ricas III. (Col. Staphyl. Hist.). Arbeiten uber Morphologische und Taxonomische Entomolgie aus Berlin-Dahlem, 2, 188 - 218.","Wenzel, R. L. (1938) New and little known Neotropical Hetaeriomorphini (Coleoptera: Histeridae). Revista de Entomologia, 9, 317 - 321.","Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Bulletin of the USNM, 185 (1 - 6), 1 - 1492."]}

Eucurtiopsis ashei Tishechkin & Caterino, sp. n. (Figs. 1–3, 7) Holotype. Female labeled "PHILLIPINES: Naga City BGY. Panicuason Camarines Sur Feb 1992. D. General / HOLOTYPE Eucurtiopsis ashei sp. n. A. Tishechkin & M. Caterino des. 2006" (SEMC). Description. L: 1.45 mm; W: 0.89 mm; E/Pn L: 1.95; E/Pn W: 1.47; Pn W/L: 1.33; E L/W: 1.15; Pr/Py: 0.92; Sterna: 0.50, 0.08, 0.37; Tibiae: 0.42, 0.40, 0.54. Body dark rufescent brown, with elytra and legs slightly lighter, most of body surfaces covered with erect yellow setae. Frons slightly longer than wide, with sides subparallel, bordered by costate marginal striae, not indented at antennal insertions, with large deep punctures and fine alutaceous background microsculpture; labrum short, arcuate, with the same type of microsculpture; mandibles strongly bent, with long narrow tips, without setae; antennal scape about half as broad as long, arcuate, widest at about midpoint, punctuation and microsculpture similar to those of frons, but punctures smaller; antennal funicle (of female) collectively slightly shorter than scape; antennal club (of female) oval, as long as funicle, glabrous in basal fifth, otherwise densely covered with setae. Pronotum (Fig. 1) with posterior margin obtusely angulate, sides widest at about midpoint, bisinuate; marginal striae visible from above in basal half, then abruptly descend downwards to meet supracoxal striae; anterior margin straight; most of pronotal disk occupied by bifid anteromedian elevation, abruptly lowering towards anterior margin and laterally and smoothly descending towards basal margin; disk surface smooth, completely covered with large, dense, deep punctures and long erect setae; antennal cavities largely open above. Scutellum not visible. Elytra (Figs. 1, 3) with conspicuous small humeral trichomes; anterior and posterior elevations completely separated by deep transverse incision; outer corners of both their apices slightly projected closing incision externally; anterior elevation with anteromedian groove indistinct, marked as shallow broad sulcus along the middle of elevation, its apex straight, bearing dense tight fringe of short setae; posterior elevation also with straight anterior edge bearing slightly wider corresponding fringe of setae; posterior elevation broadening posteriorly and descending rather abruptly to elytral middle; elytral disk in posterior half (Fig. 3) with obtuse conical elevation gradually descending towards elytral apices; elytral disc moderately transversely depressed in anterior half between trichomes; elytral surface with numerous setae throughout, shorter, stouter and denser on anterior depression and trichome elevations; punctures of elytral disc small and sparse, less conspicuous on anterior depression and much denser on trichome elevations; elytral marginal stria complete, departing conspicuously from margin above metathoracic leg; epipleuron (Fig. 3) smooth and glossy, bearing numerous upward pointing setae. Prosternum (Fig. 2) long; prosternal leg depression margined by raised carina; prosternal keel almost straight, with the marginal stria distinct, thin and carinate; prosternal disk punctate throughout with irregularly spaced deep large, mostly elongate punctures; posterior part of prosternum shortly ascending and overhanging anterior part of mesoventrite; mesoventrite short, about six times as wide as median length; straight at middle, anteriorly bordered by a thin carinate stria; mesoventral disc slightly concave, with row of large deep punctures; mesepimeron prominent, impunctate, with dense alutaceous microsculpture; mesometaventral suture well impressed, continuous at side with stria delimiting the mesothoracic leg depression (lateral metaventral stria); metaventral disk glossy and smooth, with scattered small punctures and few fine setae; median metaventral suture distinct, well impressed, complete from anterior to posterior margin; posterior margin with transverse stria going laterally along anterior margin of metacoxa, than angling towards metepisternum, near which it is arched continuously with lateral metaventral stria; 1st abdominal ventrite similar in texture and setation to metaventrite, with raised stria delimiting depression for reception of metathoracic leg. Femora (Figs. 2, 3) rather stout, especially pro- and metafemora, edges of profemora weakly arcuate, edges of meso- and metafemora ascending posteriad, all margined along anterior and posterior sides; protibia smoothly angulate about one-third from base, arcuate to narrow rounded apex; meso- and metatibia with marginal angulation less pronounced, almost symmetrically rounded, widest near middle, mesotibia clearly narrower than metatibia; tarsi slightly laterally compressed, about 0.6–0.7 times length of corresponding tibiae; tarsal claws simple, divergent, weakly arcuate, about 0.3 times length of corresponding apical tarsomere. Propygidium not carinate transversely, with disc slightly elevated, deeply punctate, punctures separated by about their widths, intervening spaces weakly alutaceous; pygidium similarly textured and punctured, but with punctures smaller and becoming progressively sparser towards apex. Right valvifer and coxite as figured (Fig. 7). Etymology. The species is dedicated to the memory of the late J. S. Ashe, curator of the Natural History Museum, University of Kansas, in appreciation of our long-term cooperative relationship and his contributions to collecting tropical inquilinous beetles., Published as part of Tishechkin, Alexey K. & Caterino, Michael S., 2007, Description of the first Chlamydopsinae (Coleoptera: Histeridae) from the Philippines, pp. 39-44 in Zootaxa 1527 (1) on pages 40-41, DOI: 10.11646/zootaxa.1527.1.4, http://zenodo.org/record/5087577