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Glyphiulus subgranulatus n. sp. (Figs 7-9) TYPE MATERIAL. — China. Yunnan Prov., Mengzi County, cave near footpath to plateau, 5.I.1989, leg. P. Beron, holotype ♂ (NMNHS); paratypes 7 ♂♂, 14 ♀♀, 11 juv. (NMNHS); 2 ♂♂, 2 ♀♀ (MNHN GA 035); 2 ♂♂, 2 ♀♀ (ZMUM); 1 ♂, 1 ♀ (ZMUC); 1 ♂, 1 ♀ (MCSNV); 1 ♂, 1 ♀ (SEM). — Yunnan Prov., Mengzi County, Pothole No. 2 (Ma Fa Tiao Dong), 6.I.1989, leg. P. Beron, paratype 1 ♀ (NMNHS). ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus. DIAGNOSIS. — Differs from the most similar congeners by the crests on the collum being underdeveloped (especially so anteriorly), the body segments somewhat compressed laterally, the anterior gonopods with unusually slender telopodites, etc. DESCRIPTION Length of adults of both sexes 19-28 mm, width 1.0- 1.3 mm; males usually a little shorter and more slender than females. Coloration much as in G. granulatus, variegate, yellow-brown, with a dark brown vertex, blackish ocellaria, mainly brown crests on collum and grey-brownish lateral longitudinal stripes running down from dark brown ozoporiferous tubercles; dorsum generally light, but a thin axial line mostly present due to darker median crests. Adults with 38-52p+5-2a+T in males, 39-63p+5- 1a+T in females; largest juvenile ♂ with 31p+3a+T; two smallest juveniles with 17 or 18 haplopodous segments after 6th, plus 2a+T. Holotype about 23 mm long, 1.0 mm wide, with 47p+2a+T. Other characters as in G. granulatus, including mentum either divided (n = 1) (Fig. 9B) or undivided (n = 1) (Fig. 7A), 11-18 ocelli, antennae (Figs 8A; 9A), the carinotaxy formulae (Fig. 7 B-F), claw, posterior gonopod structure (Fig. 9F, G), male legs 1 and 2 (Figs 8D; 9C, D), telson (Fig. 8B), etc., but body size much larger (see above); segments compressed laterally (Fig. 8C); crests on collum (7+m+7 at caudal margin) evidently reduced in size, almost obsolete anteromedially (Figs 7B; 8A); legs somewhat longer (about 4/5 as long as body height); anterior gonopods with telopodites more slender, while the coxosternum has much shorter distomedian outgrowths and acute distolateral corners (Fig. 9E). REMARKS In the absence of evident troglomorphic traits, this species can only be considered as troglophilic.

Genus Glyphiulus Gervais, 1847 TYPE SPECIES. — Iulus (recte: Julus) granulatus Gervais, 1847. TYPE LOCALITY. — Mascarene Islands. DIAGNOSIS. — Cambalopsidae with tergal crests evident at least on postcollar segments; similar crests often present also on collum, these not fragmented into more than two parts. Carinotaxy formulae of collum highly variable, often complex, but median crest, even if abbreviated anteriorly, usually clear. Carinotaxy formulae of postcollar segments usually 2(1)/2(1)+I/i+4/3+I/i+2(1)/2(1) (granulatus -group) or 2(1)/2(1)+I/i+3/3+I/i+2(1)/2(1) (javanicus -group), crests undivided or divided into three tubercles occurring only occasionally, same as three crests below I/i. Basalmost crests on ozoporiferous segments never hypertrophied, instead always being smallest, sometimes even abbreviated. Ozopores starting from segment 5, always lying on particularly prominent tubercles. Eyes usually present, though often reduced and sometimes unpigmented, especially so in cavernicoles. Mentum of gnathochilarium usually divided. Male legs 1 from little modified to strongly reduced, either with coxal outgrowths contiguous but virtually never completely fused medially (javanicus -group) or with leg remains (more rarely, retaining a few segments) widely separated due to a sternum devoid of median strictures (granulatus -group). Male legs 2 nearly normal, not incrassate, coxae somewhat modified, and prefemora abbreviated frontally, for accommodation of penes. Male legs 3 with strongly elongated coxae. Anterior gonopods usually with a shield-like coxosternum and a moveable, lateral, 1-segmented, often curved telopodite; flagella absent. Posterior gonopods compact, not elongated, often with apical flagelloid outgrowths. REMARKS The general distribution of the granulatus -group shows a quite coherent pattern, occupying a region between Sichuan, Hubei and Shanghai in the North, and central Vietnam in the South. This matches very closely the distribution of the javanicus -group (Fig. 27), which ranges from between Sichuan and Zhejiang in the North to Java in the South. In other words, the distribution of the granulatus -group is somewhat more restricted and falls entirely within that of the javanicus -group, the latter thus fully reflecting the general range of Glyphiulus. Even after this review, there is no doubt that these generally subtropical to tropical, highly montane and often karstic areas contain many new species of this widespread and large genus waiting to be found and described. The only outlying member of the javanicus -group appears to be G. javanicus itself.The strong isolation of this species from the remaining bulk, which is restricted to continental Southeast Asia (Fig. 27), might well represent an accidental introduction from the mainland. Carl (1911) explicitly indicated a sugar cane plantation as the place whence the type series of G. javanicus had come. The fact that the species of Glyphiulus are mostly highly local in distribution deserves attention. This can often be accounted for by their being troglobites, naturally restricted to one single cave or a limited group of caves. However, the impression that the majority of the species are scarcely more than troglophiles suggests that a wealthy, barely sampled fauna also exists epigeically. This undersampling is due to the fact that most of the expeditions to these often remote and poorly accessible places have been speleological, concentrating on the “mystique” of the underground world, thus giving the biased impression that cambalopsids in general, and Glyphiulus species in particular, are especially common in caves. Above-ground explorations will certainly reveal a far richer fauna of forest-dwelling species in the region. Summarising the poor state of current knowledge, we have still only touched the tip of the iceberg (Golovatch et al. 2007)., Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, pp. 417-456 in Zoosystema 29 (3) on pages 454-455, DOI: 10.5281/zenodo.5393630, {"references":["CARL J. 1911. - Drei neue Diplopoden des Genfer Museums. Revue suisse de Zoologie 19: 397 - 407.","GOLOVATCH S. I., GEOFFROY J. - J., MAURIES J. - P. & VAN DEN SPIEGEL D. 2007. - Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatusgroup. Zoosystema 29 (1): 7 - 49."]}

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: http://doi.org/10.5281/zenodo.5392079, {"references":["ATTEMS C. 1900. - Dr. Brauer's Myriopoden-Ausbeute auf den Seychellen im Jahre 1895. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 13: 133-171.","ATTEMS C. 1926. - Myriapoda in KUKENTHAL W. & KRUMBACH T. (eds), Handbuch der Zoologie, IV, 1, Progoneata, Chilopoda, Insecta. W. de Gruyter & Co., Berlin; Leipzig: 1-402.","ATTEMS C. 1938. - Die von Dr C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'Histoire naturelle n.s. 6 (2): 187-353.","ATTEMS C. 1953. - Myriopoden von Indochina. Gesammelt von Dr C. Dawydoff (1938-1939). Memoires du Museum national d'Histoire naturelle ser. A (Zoologie), 5 (3): 133-230.","BROLEMANN H. W. 1931. - Myriapodes recueillis par Mme Pruvot en Nouvelle-Caledonie et aux Loyalty. Archives de Zoologie experimentale et generale 72 (2): 275-316.","CARL J. 1911. - Drei neue Diplopoden des Genfer Museums. Revue suisse de Zoologie 19: 397-407.","CHEN J. X. & MENG W. X. 1991. - A new species of the genus Glyphiulus (Diplopoda, Spirostreptida, Cambalopsidae).Acta Zootaxonomica Sinica 16: 394- 397 (in Chinese, English summary).","CROSLAND M. W. J. 1994. - Millipede myrmecophile, Glyphiulus granulatus, found with the primitive ant Harpegnathos venator. Journal of Entomological Science 29: 141-142.","ENGHOFF H. 1993. - Haplopodous diplopods: a new type of millipede body construction discovered in cambalopsid juveniles (Diplopoda, Spirostreptida). Acta Zoologica 74 (3): 257-261.","ENGHOFF H. 2005. - The millipedes of Thailand (Diplopoda). Steenstrupia 29: 87-103.","ENGHOFF H., DOHLE W.&BLOWER J. G. 1993. - Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations.Zoological Journal of the Linnean Society 109: 103-234.","ENGHOFF H., GOLOVATCH S. I. & NGUYEN DUC ANH 2004. - A review of the millipede fauna of Vietnam (Diplopoda). Arthropoda Selecta 13 (1-2): 29-43.","GEOFFROY J.-J. & GOLOVATCH S. I. 2004. - Some polydesmidan millipedes from caves in southern China (Diplopoda: Polydesmida), with descriptions of four new species. Arthropoda Selecta 13 (1-2): 19-28.","GERVAIS P.1847. - Myriapodes, in WALCKENAER C. A. & G ERVAIS P. (eds), Histoire naturelle des Insectes Apteres, Tome 4e. Librairie encyclopedique de Roret, Paris: 1-623.","GOLOVATCH S. I., GEOFFROY J.-J. & MAURIES J.-P. 2006a. - Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92.","GOLOVATCH S. I., GEOFFROY J.-J. & MAURIES J.-P. 2006b. - Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema 28 (4): 887-915.","GOLOVATCH S. I., GEOFFROY J.-J. MAURIES J.-P. & VAN DEN SPIEGEL D. in press. - Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus - group. Zoosystema.","HOFFMAN R. L. 1977. - Diplopoda from Malayan caves collected by M. Pierre Strinati. Revue suisse de Zoologie 84 (3): 699-719.","HOFFMAN R. L. 1980. - Classification of the Diplopoda. Museum d'histoire naturelle, Geneve, 237 p. (dated 1979, published 1980).","JEEKEL C. A. W. 1963. - Diplopoda of Guiana (1-5), in GIJSKES D. C. & WAGENAAR HUMMELINCK P. (eds), Studies on the Fauna of Suriname and Other Guyanas 4. Martinus Nijhoff, The Hague: 1-157.","JEEKEL C. A. W. 1971. - Nomenclator generum et familiarum Diplopodorum. A list of the genus and family-group names in the class Diplopoda from the 10th edition of Linnaeus, 1758, to the end of 1957.Monografieen van de Nederlandse Entomologische Vereiniging 5: i-xii + 1-412 (dated 1970, published 1971).","JEEKEL C. A. W. 1985. - The distribution of the Diplocheta and the \"lost\" continent Pacifica (Diplopoda). Bijdragen tot de Dierkunde 55 (1): 100-112.","JEEKEL C. A. W. 2004. - A bibliographic catalogue of the \"Cambaloidea\" (Diplopoda, Spirostreptida). Myriapod Memoranda 7: 43-109.","KARSCH F.1881. - Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaften 54: 1-79.","KORSOS Z. 2004. - Checklist and bibliography of millipedes (Diplopoda) of Taiwan. Collection and Research (NMNS, Taichung) 17: 11-32.","KRABBE E. 1982. - Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Neue Folge 24: 1-476.","LOKSA I. 1960. - Einige neue Diplopoden- und Chilopodenarten aus chinesischen Hohlen. Acta Zoologica Academiae Scientiarum Hungaricae 6 (1-2): 137-148.","MAURIES J.-P. 1970. - Examen des types des genres Cambalomorpha et Cambalopsis Pocock, 1895. Essai de classification des Glyphiulinae Verhoeff, 1936 (Diplopoda, Cambalidea).Bulletin du Museum national d'Histoire naturelle ser. 2, 42 (3): 509-519.","MAURIES J.-P. 1977. - Le genre Glyphiulus Gervais, 1847, et sa place dans la classification des Cambalides, a propos de la description d'une nouvelle espece du Viet-Nam (Diplopoda, Iulida, Cambalidea). Bulletin du Museum national d'Histoire naturelle ser. 3, 431 (Zoologie 301): 243-250.","MAURIES J.-P. 1981. - Craspedosomida, Stemmiulida et Cambalida (Myriapoda: Diplopoda) de Sri Lanka (Ceylan). Entomologica Scandinavica, Supplementum 11: 33-62.","MAURIES J.-P. 1983. - Cambalides nouveaux et peu connus d'Asie, d'Amerique et d'Oceanie.I. Cambalidae et Cambalopsidae (Myriapoda, Diplopoda). Bulletin du Museum national d'Histoire naturelle 4e ser., sect. A, 5 (1): 247-276.","MAURIES J.-P. 1987. - Cambalides nouveaux et peu connus d'Asie, d'Amerique et d'Oceanie. II. Pseudonannolenidae, Choctellidae (Myriapoda, Diplopoda). Bulletin du Museum national d'Histoire naturelle 4e ser., sect. A, 9 (1): 169-199.","MAURIES J.-P. 1992. - Sur la vraie place du genre Protosilvestria Handschin dans la classification des Diplopodes Iuliformes, in MEYER E., THALER K. & SCHEDL W. (eds), Proceedings of the 8th International Congress of Myriapodology. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, Supplementum 10: 23-31.","MAURIES J.-P. 2005. - Guizhousoma latellai gen. n., sp. n., de Chine continentale, type d'une nouvelle famille de la superfamille des Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta 14 (1): 11-17.","MAURIES J.-P. & NGUYEN DUY- JACQUEMIN M. 1997. - Nouveaux Craspedosomides et Glyphiulides cavernicoles de Chine (Diplopoda). Memoires de Biospeologie 24: 46-62.","MURAKAMI Y.1975. - The cave myriapods of the Ryukyu Islands (I). Bulletin of the National Science Museum Series A (Zoology), 1 (2): 85-113.","PETERS W. 1864. - Eine Ubersicht der im Konigl. zoologischen Museum befindlichen Myriopoden aus der Familie der Polydesmi, so wie Beschreibungen einer neuen Gattung, Trachyjulus, der Juli und neuer Arten der Gattung Siphonophora. Monatsbericht der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1864: 529-551.","POCOCK R. I. 1893. - Report upon the Myriopoda of the \"Challenger\" Expedition, with remarks upon the fauna of Bermuda. Annals and Magazine of Natural History (6) 11: 121-142.","POCOCK R. I. 1894. - Chilopoda, Symphyla and Diplopoda from the Malay Archipelago,in WEBER M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien 3: 307-404.","POCOCK R. I. 1895. - Report upon the Chilopoda and Diplopoda obtained by P. W. Basset-Smith, Esq., Surgeon R. N., during the cruise in the Chinese seas of H.M.S. \"Penguin\", Commander W. U. Moore commanding.Annals and Magazine of Natural History (6) 15: 121-142.","STOEV P.2004. - The first troglomorphic species of the millipede genus Paracortina Wang & Zhang, 1993 from South Yunnan, China (Diplopoda: Callipodida: Paracortinidae). Zootaxa 441: 1-8.","STOEV P.&ENGHOFF H. 2005. - A new cave-dwelling millipede of the genus Bollmania Silvestri, 1896 from Yunnan, China, with remarks on the reduction of the second female leg-pair (Diplopoda: Callipodida: Caspiopetalidae). Journal of Natural History 39: 1875- 1891.","STOEV P. & GEOFFROY J.-J. 2004. - Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida). Acta Arachnologica 53 (2): 93-103.","VERHOEFF K. W. 1924. - Results of Dr. E. Mjoberg's scientific expeditions to Australia 1910-1913. 34. Myriapoda: Diplopoda. Arkiv for Zoologi 16 (5): 1-142.","VERHOEFF K. W.1936. - Zur Kenntnis der Glyphiuliden (Cambaloidea). 143. Diplopoden-Aufsatz. Zoologischer Anzeiger 113 (3/4): 49-62.","ZHANG C. Z. & LI Z. Y.1982. - Die Gattung Glyphiulus (Diplopoda: Cambalidea) von China. Sinozoologia 2: 85-93 (in Chinese, German summary)."]}

Glyphiulus percostulifer n. sp. (Figs 20; 21) TYPE MATERIAL. — Laos. Khammouan Prov., Ban Nakok (= Nakhok), Tham Thê, 104.49°E, 17.90°N, cave, hand collection, 11.II.1998, leg. A. Bedos & L. Deharveng (LAO-016), holotype ♂ (MNHN GA 051); paratypes 1 ♂, 1 ♀, 1 subad. ♂ (MNHN GA 051); 1 ♀ (ZMUM / SEM). ETYMOLOGY. — To emphasize the very close resemblance to G. costulifer n. sp., but with more strongly developed tergal crests. DIAGNOSIS. — Differs from congeners in the small size, the completely unpigmented tegument and ocelli, coupled with certain details of leg and gonopod structure (see also key below). DESCRIPTION Holotype with 34p+2a+T, about 14 mm long and 1.0 mm wide; adult paratypes with 36p+2a+T, about 14 mm long and 1.0 mm wide (♂), with 40p+1a+T or 38p+1a-T, about 17 mm long and 1.0 mm wide (♀); subadult ♂ with 31p+2a+T. Coloration uniformly pallid, ocelli not visible (Fig. 20A). Postcollar constriction, dorsal crests and carinotaxy pattern on postcollar terga (Fig. 20 A-E), segment’s texture and cross-section (Fig. 20 B-D), paraprocts and hypoproct (Fig. 20F), ventral flaps of male segment 7, male legs 2 and 3 (Fig. 21D, E), penes (Fig. 21D) generally as in G. costulifer n. sp., but antennae elongate, antennomeres 5-7 richer in sensilla (Fig. 21A), gnathochilarium (mentum divided, n = 2) (Fig. 21B) slightly even more oligotrichous; collum (Fig. 20A) with a nearly typical carinotaxy formula (I-IV+5a+pc+ma+pc+5a+IV-I); all tergal crests (Fig. 20 A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones about as high as wide; epiproct (Fig. 20E, F) upturned and broadly rounded in caudal part, with one median and two paramedian crests; legs slightly longer than body diameter (Fig. 20B), always with an accessory filament/spine at base of claw, which is usually slightly more than half as long as claw (Fig. 21 D-F); male leg 1 with normal, 5-segmented telopodites and large, paramedian, coxal processes (Fig. 21C); each anterior gonopod plate with a high and subsecuriform distomedial process (Fig. 21G); posterior gonopods with distally slightly plumose flagella (Fig. 21H). REMARKS Based on several troglomorphic traits, such as the completely unpigmented body and ocelli, as well as the elongated antennae and legs, this species seems to be troglobitic, although the small size may also be evidence of its being an inhabitant of the MSS (“milieu souterrain superficiel”).

Key to Pacidesmus species 1. Paraterga, even of anterior segments, remaining below level of dorsum. Sterna between male legs 6 and 7 with conspicuous paramedian tubercles. Outgrowth at base of gonopod acropodite short and unciform. Epigean species from Thailand. P. shelleyi ��� Paraterga more strongly elevated, especially on anterior segments. Sterna between male legs 6 and 7 usual, unmodified. Outgrowth at base of gonopod acropodite elongate. Cavernicoles from Guizhou, China..................................................................... 2 2. Paraterga of anterior postcollar segments strongly elevated above dorsum (Fig. 2). Caudal corners of paraterga narrowly rounded, but never pointed (Figs 1���7). Outgrowth l at base of a lamelliform; distal part of a bare (Figs 8, 11��� 12)...... P. martensi ��� Paraterga of anterior postcollar segments only barely elevated above dorsum. Caudal corners of most of paraterga pointed. Outgrowth l at base of a flagelliform, unciform terminally; a with a distinct tooth subapically ................................................ P. sinensis, Published as part of Golovatch, Sergei I. & Geoffroy, Jean-Jacques, 2006, Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae), pp. 363-368 in Zootaxa 1325 on page 368, DOI: 10.5281/zenodo.174082

Hyleoglomeris speophila n. sp. (Fig. 9) TYPE MATERIAL. — Vietnam. Hai Phong, Cat Ba Island, Phu Long, Thien Luong Cave, 29.IX.1998, leg. L. Deharveng, holotype ♂ (MNHN CC 157). ETYMOLOGY. — To emphasize cavernicoly. DIAGNOSIS. — Differs from congeners in some troglomorphic traits, combined with a relatively broad caudofemoral process of the telopod. DESCRIPTION Length 8.5 mm, width 4.1 mm. Coloration entirely pallid, unpigmented. Antennomere 6 about 2.0 times longer than wide, antennomere 7 very short; 6 poorly visible, unpigmented but convex ocelli on each side of head; Tömösváry's organ transverse-oval, about twice as wide as long. Collum with two transverse striae. Thoracic shield with a rather narrow hyposchism slightly surpassing the caudal tergal contour; 9 transverse striae, of which 6 start above the schism and 5 (neither the first nor the last one though) cross the dorsum. Male pygidium virtually not sinuate medially at caudal margin. Male leg 17 (Fig. 9A) with a high, rounded outer coxal lobe; telopodite 4-segmented. Male leg 18 (Fig. 9B) with a rounded syncoxital notch; telopodite 4-segmented. Telopods (Fig.9C, D) with a rather small, roundly subtrapeziform syncoxital lobe flanked by two setose horns crowned with an apical setoid. Prefemur and femur micropapillate nearly throughout. Caudomedial outgrowth of femur wide at base, that of tibia with a pilose tubercle at base.Tarsus narrowly rounded apically, almost pointed. REMARKS Based on morphological evidence, this new species resembles several epigean congeners from China or Indochina, such as H. maculata n. sp. (see above), H. electa in the sense of Attems (1938) and others, but none of these shows the troglomorphic features of complete depigmentation and elongate Tömösváry's organs. However, the ocelli are still discernible and neither the antennae nor the legs are particularly long. To prove that we face a troglobite, additional observations and material are necessary. In general, Vietnam supports numerous karst caves rich in troglobites (Deharveng et al. 2001), while Thien Luong Cave is known to mainly harbour troglobitic arthropods (Deharveng pers. comm.).

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema 28 (4): 887-915, DOI: http://doi.org/10.5281/zenodo.4525415, {"references":["ATTEMS C. 1897. - Myriopoden. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 23 (3): 473-536.","ATTEMS C. 1899. - Neues uber palaarktische Myriopoden. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 12: 286-336.","ATTEMS C. 1909. - Die Myriopoden der Vega-Expedition. Arkiv for Zoologi 5 (3): 1-84.","ATTEMS C. 1930.- Myriopoden der Kleinen Sunda-Inseln, gesammelt von der Expedition Dr Rensch. Mitteilungen aus dem Zoologischen Museum Berlin 16: 117-184.","ATTEMS C. 1938. - Die von Dr C. Dawydoff in Franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'Histoire naturelle n.s., 6 (2): 187-353.","ATTEMS C. 1953. - Myriopoden von Indochina. Gesammelt von Dr C. Dawydoff (1938-1939). Memoires du Museum national d'Histoire naturelle ser. A (Zoologie), 5 (3): 133-230.","BESSON J.-P., DEHARVENG L. & BREHIER F.2001. - Laos, in JUBERTHIE C. & DECU V. (eds), Encyclopaedia Biospeologica 3. Societe internationale de Biospeologie, Moulis: 1883-1889.","BROLEMANN H. W. 1896. - Sur quelques myriapodes de Chine. Memoires de la Societe zoologique de France 9: 349-362.","CARL J. 1912. - Die Diplopoden-Fauna von Celebes. Revue suisse de Zoologie 20 (4): 73-206.","CHAMBERLIN R. V. 1921. - New Chilopoda and Diplopoda from the East Indian region. Annals and Magazine of Natural History ser. 9, 7: 50-87.","CHAMBERLIN R. V. & WANG Y. M. 1953. - Records of millipeds (Diplopoda) from Japan and other oriental areas, with descriptions of new genera and species. American Museum Novitates 1621: 1-13.","CHEN Z., DECU V., JUBERTHIE C. & UENO S.-I. 2001. - Chine, in JUBERTHIE C. & DECU V. (eds), Encyclopaedia Biospeologica 3. Societe internationale de Biospeologie, Moulis: 1763-1781.","DEHARVENG L., LE CONG K. & BEDOS A. 2001. - Vietnam, in JUBERTHIE C. & DECU V. (eds), Encyclopaedia Biospeologica 3. Societe internationale de Biospeologie, Moulis: 2027-2037.","GOLOVATCH S. I. 1975. - [Two genera of Oniscomorpha, found in Transcaucasia, new to the USSR fauna and their zoogeographical connections]. Zoologicheskii Zhurnal 54 (10): 1566-1571 (in Russian, English summary).","GOLOVATCH S. I. 1976. - [New or poorly-known species of Glomeridae (Diplopoda, Oniscomorpha)]. Zoologicheskii Zhurnal 55 (6): 931-935 (in Russian, English summary).","GOLOVATCH S. I. 1978. - [Some East-Asiatic millipedes (Diplopoda) in the collection of the Zoological Institute of the Academy of Sciences of the USSR]. Entomologicheskoe Obozrenie 57 (3): 677-681 (in Russian, English title).","GOLOVATCH S. I. 1981. - Some East-Asiatic millipedes (Diplopoda) in the collection of the Institute of Zoology of the Polish Academy of Sciences. Annales Zoologici 36 (8): 161-168.","GOLOVATCH S. I. 1983. - On several new Glomeridae (Diplopoda) from Indochina. Annales Historico-Naturales Musei Nationalis Hungarici 75: 107-116.","GOLOVATCH S. I. 1987. - Diplopoda from the Nepal Himalayas. Glomeridae, additional Opisotretidae. Courier Forschungsinstitut Senckenberg 93: 219-228.","GOLOVATCH S. I. 1989. - Diplopoda of the Caucasus, 2. Glomeridae, with contributions to the fauna of Turkey.Senckenbergiana Biologica 69 (4/6): 421-440 (dated 1988, published 1989).","GOLOVATCH S. I. & ENGHOFF H. 2003. - Pill-millipedes of the Canary Islands: the Glomeris alluaudi -group (Diplopoda, Glomeridae). Vieraea 31: 9-25.","HAGA A. 1968. - [Japanese Millipedes]. Private publication, 11 p., 6 pls (in Japanese).","HOFFMAN R. L. 1980. - Classification of the Diplopoda. Museum d'Histoire naturelle, Geneve, 237 p. (dated 1979, published 1980).","JEEKEL C. A. W. 1970. - Nomenclator generum et familiarum Diplopodorum. A list of the genus and family-group names in the class Diplopoda from the 10th edition of Linnaeus, 1758, to the end of 1957.Monografieen van de Nederlandse Entomologische Vereiniging 5: i-xii, 1-412.","KIM T. H. & LIM K. Y. 1995a. - [Millipedes (Diplopoda) from Cheju Island], in Insects of Quelpart Islands. Chejudo Folklore and Natural History Museum, Cheju: 271-280 (in Korean, English abstract).","KIM T. H. & LIM K. Y. 1995b. - A check list of Diplopoda from Chejudo. Insecta Koreana Supplementum 5: 211-214.","KORSOS Z. 2004. - Checklist and bibliography of millipedes (Diplopoda) of Taiwan, R.O.C. Collection and Research (NMNS, Taichung) 17: 11-32.","LIM K. Y., KIM T. H. & KWAK J. S. 1992. - [Distribution of millipedes in relation to altitude and flora on Mt. Chiri]. Korean Journal of Ecology 15 (4): 329-335 (in Korean, English abstract).","LOHMANDER H. 1939. - Uber einige neue Diplopoden aus dem ostlichen Anatolien. Verhandlungen der Naturforschenden Gesellschaft in Basel 50: 126-150.","MAURIES J.-P. 1966. - Decouverte, par H. Coiffait, de representants des genres Speleoglomeris Silv. et Trichoblaniulus Verh. dans les grottes de Grece (Diplopoda). Annales de Speleologie 21 (3): 621-630.","MAURIES J.-P. 1971. - Diplopodes epiges et cavernicoles des Pyrenees espagnoles et des monts Cantabriques.VII. Glomerides. Essai de classification des Glomeroidea. Bulletin de la Societe d'Histoire naturelle de Toulouse 107 (3-4): 423-436.","MAURIES J.-P. 1984. - Deux especes nouvelles de Diplopodes cavernicoles des Cyclades : Hyleoglomeris beroni (Glomerida) et Syrioiulus andreevi (Iulida). Biologia Gallo-Hellenica 11 (1): 37-49.","MAURIES J.-P. 2005. - Essai de classification des Glomerida (Diplopoda), et description de deux nouveaux genres du nord-ouest de la peninsule iberique. Arthropoda Selecta 14 (3): 241-249.","MIKHALJOVA E. V. & KIM J. P. 1993. - Contribution to the millipede fauna of Korea (Diplopoda). Korean Arachnology 9 (1-2): 31-42.","MIKHALJOVA E. V. & LIM K. Y. 2000. - Millipede fauna (Diplopoda) of South Korea. Korean Journal of Systematic Zoology 16 (2): 147-157.","MIKHALJOVA E. V., GOLOVATCH S. I. & WYTWER J. 2000. - On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109-122.","MIYOSI Y. 1955. - [Beitrage zur Kenntnis japanischer Myriopoden. 14. Aufsatz: Uber eine neue Art von Lithobiomorpha und zwei neue Arten von Diplopoda]. Zoological Magazine 64 (8): 267-270 (in Japanese, German summary).","MIYOSI Y.1959. - [Uber japanische Diplopoden]. Arachnological Society of East Asia, Osaka, iii + 223 p., 19 pls (in Japanese, German title).","MURAKAMI Y.1975. - The cave myriapods of the Ryukyu Islands (I). Bulletin of the National Science Museum ser. A (Zoology), 1 (2): 85-113.","POCOCK R. I. 1894. - Chilopoda, Symphyla and Diplopoda from the Malay Archipelago,in WEBER M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien 3: 307-404.","READ H. J. & GOLOVATCH S. I. 1994. - A review of the Central Asian millipede fauna. Bulletin of the British Myriapod Group 10: 59-70.","SHINOHARA K. 1978. - [Myriapod-fauna of Saitama], in The fauna of Saitama Prefecture. Saitama Prefecture Board of Education, Urawa: 459-470 (in Japanese, English summary).","SHINOHARA K. 1981. - Chilopods and diplopods of the National Park for Nature Study in Tokyo, based on the 1978-1979 investigations. Miscellaneous Reports of the National Park for Nature Study 12: 89-91 (in Japanese, English title).","SILVESTRI F. 1895. - I chilopodi ed i diplopodi di Sumatra e delle isole Nias, Engano e Mentavei.Annali del Museo Civico di Storia Naturale di Genova ser. 2, 14 (34): 705-760.","SILVESTRI F. 1917. - Contributions to a knowledge of the Oriental Diplopoda Oniscomorpha, I. The family Glomeridae. Records of the Indian Museum 13 (3, 9): 103-151.","STOEV P.&ENGHOFF H. 2005. - A new cave-dwelling millipede of the genus Bollmania Silvestri, 1896 from Yunnan, China, with remarks on the reduction of the second female leg-pair (Diplopoda:Callipodida: Caspiopetalidae). Journal of Natural History 39: 1875-1891.","TAKAKUWA Y. 1954. - [The Diplopods of Japan and its Adjacent Areas]. Society for the Promotion of Science, Tokyo, vi + 255 p. (in Japanese, partly in German).","TAKANO M. 1981. -Taxonomical studies on the Japanese Myriapoda, II. A new species of the genus Hyleoglomeris (Diplopoda: Glomeridae) from Kumamoto Prefecture in Southwest Japan. Edaphologia 24: 35-38.","TAKASHIMA H. & HAGA A. 1956. - [A contribution towards the Japanese cave-dwelling species of the class Diplopoda]. Journal of the Yamashina Institute for Ornithology 8: 329-343, pls I-II (in Japanese with English notes on type localities).","VERHOEFF K.W. 1910.- Ueber Diplopoden. 41. Aufsatz: Indomalayische Glomeriden. Sitzungsberichte der Gesellschaft naturforschender Freunde 5: 240-249.","VERHOEFF K. W. 1912. - Uber Nesoglomeris n. g. J. Carl. Zoologischer Anzeiger 40 (4/5): 150-151.","VERHOEFF K. W. 1915. - Zur Kenntnis der Plesiocerata. (Uber Diplopoden, 82. Aufsatz). Zoologischer Anzeiger 46 (1): 16-48.","VERHOEFF K.W. 1929. - Zur Systematik, vergleichenden Morphologie und Geographie europaischer Diplopoden, zugleich ein zoogeographischer Beitrag. 111. Diplopoden-Aufsatz. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 57: 555-659.","VERHOEFF K. W. 1936. - Ueber Diplopoden aus Japan gesammelt von Herrn Y.Takakuwa. Transactions of the Sapporo Natural History Society 14 (3): 148-172.","VERHOEFF K. W. 1937. - Zur Kenntnis ostasiatischer Diplopoden. II. Zoologischer Anzeiger 119 (1/2): 33-40.","VERHOEFF K. W. 1942. - Ascospermophoren aus Japan und uber neue japanische Diplopoden. Zoologischer Anzeiger 137 (11/12): 201-217.","WANG D. & MAURIES J.-P. 1996. - Review and perspective study of myriapodology of China, in GEOFFROY J.-J., MAURIES J.-P. & NGUYEN DUY- JACQUEMIN M. (eds), Acta Myriapodologica. Memoires du Museum national d'Histoire naturelle 169: 81-99.","WANG Y. M. 1951. - Serica. Volume 1. The Myriapoda of the Philippine Islands. Edwards Brothers Publ., Ann Arbor, Michigan, 79 p.","WANG Y. M. 1961. - Serica 12. The Myriapoda of the Philippine Islands. A revision of the Myriapoda of the Philippine Islands.Quarterly Journal of theTaiwan Museum 14 (1-2): 89-140.","ZHANG F. & ZHANG C. Z. 1995. - [A new troglobitic species of glomerid millipeds from Yunnan ( Diplopoda, Glomerida, Glomeridae)]. Zoological Research 16 (1): 17-21 (in Chinese, English summary)."]}

Lipseuma josianae n. sp. (Figs 5; 6) TYPE MATERIAL. — Holotype: China, Hubei Prov., Banqiao County, Cave Chuan Dong Zi, Nos. 482, 485, 494, 9.VIII.1999, leg. J. & B. Lips, ♂ (MNHN DA155). Paratypes: same locality, 1 ♂ (MNHN DA155); 1 ♂, 2 juvs (MNHN DA155); ♀ (MNHN DA155); 2 juvs (MNHN DA155). ETYMOLOGY. — To honour Josiane Lips. DIAGNOSIS. — Differs from the only congener (L. bernardi n. gen., n. sp.) by the slightly larger size, as well as in certain details of leg and gonopod structure. DESCRIPTION Length 9.3-9.8 (♂) to 10.1 mm (♀); width of midbody segments, 0.75-0.8 (♂) to 0.9 mm (♀), maximum width of head, 0.95 mm (holotype). Vertical diameter of midbody segments, 0.75 (♂) to 0.88 mm (♀). Coloration entirely pallid. Head width >> collum> segment 2> 3> 4> 5> 6> 7> 8> segment 9 = 18; thereafter body very gradually tapering toward telson. Body with 28 segments (46 leg-pairs). Head sparsely setose, clypeolabral region slightly convex. Eyes completely reduced. Antennae very long and slender (Figs 5A; 6C), reaching beyond body segment 6 (♂) or 5 (♀) dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third; antennomere 8 with four unusually long and slender sensory cones. Gnathochilarium without promentum. Collum usual, obcordate in shape, with rudimentary paraterga. Tegument smooth, modestly shining, non-areolated, translucid. Metatergal setation 3 + 3, typical, macrochaetae very long, pointed, positioned on minute knobs; exterior macrochaeta longest, c. 0.7 times as short as body width and nearly three times longer than median or interior one; stricture between pro- and metazona very shallow, inconspicuous (Fig. 5B, C). Paraterga very small dorsolateral shoulders, rather regularly rounded in dorsal view (Fig. 5C), slightly better developed on anterior body third. CIX (15) ≈ 0.9; MIX (15) = 1.25; MA (15) ≈ 175°; PIX impossible to evaluate as paraterga too poorly developed. Axial suture distinct, usual; segment 7 inconspicuous, pleurite without ventral apophyses. Pleural crests evident. Limbus invisible. Legs generally very long and slender, c. 1.8 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes, but without papillae; male coxa 2 perforated by gonopore orifice. Tarsal papillae absent. Male legs 3-5 unmodified. Male legs 6 (Fig. 5D) exceptionally incrassate, with a distoventral process both on coxa and prefemur; acropodite strongly clasper-like, tarsus with two bulges subventrally and a modified claw. Male legs 7 (Fig. 5E) slightly incrassate, with evidently enlarged coxa and prefemur; coxa with a pointed lobe ventrally. Male legs 10 (Fig. 6A) and 11 normal, both pairs with coxal glands. Claws very long, sabre-shaped, simple. Anterior gonopods (leg-pair 8) unusually simple, small; coxites with traces of median division basally on caudal face, completely fused on front face; distally subsecuriform telopodites laterocaudal, with a few setae distally, nearly as high as the unpaired, linguiform, distally pilose syncolpocoxite, but somewhat higher than the voluminous angiocoxal sacs with a ridge on top; sternum plate-like, with each lateral bulge on front face supporting a strong seta (Fig. 5 F-H). Posterior gonopods (leg-pair 9) also very simple (Fig. 6B), placed on a plate-like sternum; coxae contiguous medially, each crowned with a simple, long, slender, erect process ventrally; telopodite reduced to a single voluminous segment filled with glandular material. Female legs 2 normal. Vulva (Fig. 6D, E) very broad; operculum small, devoid of outgrowths. REMARKS In one of the male paratypes, leg 6 is normal, not modified, whereas the anterior gonopod syncolpocoxite shows a small median notch apically, but there are no lateral setae on the coxite. Some of these traits are probably anomalous, likely neotenic., Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques & Mauriès, Jean-Paul, 2006, Four new Chordeumatida (Diplopoda) from caves in China, pp. 75-92 in Zoosystema 28 (1) on pages 86-90, DOI: 10.5281/zenodo.5393502

Lipseuma bernardi n. sp. (Figs 7; 8) TYPE MATERIAL. — Holotype: China, Sichuan Prov., Xin Long County, Three Eyes Cave, No. 438, 7.VIII.1999, leg. B. & J. Lips, ♂ (MNHN DA156). Paratypes: same locality, 1 ♂, incomplete (MNHN DA156); 1 ♂, subadult (MNHN DA156). ETYMOLOGY. — To honour Bernard Lips. DIAGNOSIS. — Differs from the only congener (L. josianae n. gen., n. sp.) by the slightly smaller size, as well as minor details of leg and gonopod structure. DESCRIPTION Length 7.2 (holotype) or 6.8 mm (subadult ♀); width of body segment 6, 0.72 mm, of midbody segments, 0.78 mm, maximum breadth of head, 0.9 mm (holotype). Antennae (Fig. 7A) with even more exposed, highly elongate, pointed apical cones on stalks. All peripheral characters like in L. josianae n. gen., n. sp. except as follows: exterior macrochaetae on midbody segments 0.6 mm long, i.e. nearly as wide as segment itself (Fig. 7B). Male legs 6 (Fig. 7C, D), 10 (Fig. 8A) and 11 virtually the same as in L. josianae n. gen., n. sp., but male legs 7 (Fig. 7E) with a characteristic protuberance on tarsus ventrally at about midway. Anterior gonopods (Fig. 7F, G) also virtually the same as in L. josianae n. gen., n. sp., but posterior gonopods (Fig. 8B) with traces of a second telopoditomere distally on caudal face. Subadult male (26 body segments) with small but evident gonopod promordia, legs 7 almost as strongly incrassate as in adult male, legs 7 also slightly enlarged. REMARKS Both Lipseuma n. gen. species are remarkable in being completely unpigmented and eyeless, as well as in bearing strongly elongated antennae, legs (especially tarsi) and exterior macrochaetae. All these features leave no doubt that we are dealing with troglobites. It is noteworthy that interspecific variation in Lipseuma n. gen. is very modest, in contrast to that observed in the other two kashmireumatid genera. In fact, L. josianae n. gen., n. sp. and L. bernardi n. gen., n. sp. are barely distinguishable. The very small differences noted between them suggest either a quite recent colonization of caves by their common ancestor or a fairly recent vicariance event with speciation within their respective cave systems. The latter scenario seems more plausible, given the profound troglomorphism of both species, which would require a geological time-scale to develop, along with the geographical proximity of Chuan Dong Zi and Three Eyes caves (both situated in the Xin Long geographical zone just at the border between Hubei and Sichuan provinces [Degouve et al. 1997]). In other words, the old geological age of the genus Lipseuma n. gen. seems unquestionable, whereas the isolation of both hitherto known congeners probably occurred only relatively recently., Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques & Mauriès, Jean-Paul, 2006, Four new Chordeumatida (Diplopoda) from caves in China, pp. 75-92 in Zoosystema 28 (1) on pages 90-91, DOI: 10.5281/zenodo.5393502, {"references":["DEGOUVE P., LIPS B. & LIPS J. 1997. - Speleologie au pays de l'homme sauvage no 3. Rapport de la 3 e expedition speleologique en Chine, A. K. L. 1997, province du Sichuan. Federation francaise de Speleologie, Paris, 130 p."]}

Pacidesmus martensi sp. n. Figs 1���12 Material examined. �� holotype (MNHN JC 306), China, Guizhou Prov., Qianxi County, Hong Lin Town, Ishui Luo Dong Cave (= Waterfall Cave), 16.XI. 2003, leg. L. Latella. ��� Paratypes: 2 ��, 4 ��, 1 juv. (MNHN JC 306), 1 ��, 1 �� (ZMUM), 1 ��, 1 �� (BDBFUL), 1 ��, 1 �� (ZMUC), Guizhou Prov., Dafang County, Yangzhamba Village, Hei Dong Cave (terrestrial), II. 2004, leg. S. Prevorčnik & B. Sket. Etymology. We take pleasure in dedicating this species to Prof. Dr Jochen Martens (Mainz University), a devoted student and collector of soil arthropods, birds and other animals in various parts of the world, including China. Diagnosis. Differs from congeners by the strongly elevated paraterga with narrowly rounded caudal corners, the poorly developed accessory seminal chamber, and the shape of the gonopod acropodite (see Key below). Description. Length of holotype about 21 mm, width of midbody pro�� and metazona 1.3 and 2.1 mm, respectively. Paratypes 17���22 mm, width of midbody pro�� and metazona 1.3���1.8 and 2.1���2.8 mm; �� usually slightly smaller than ��. Coloration entirely pallid, whitish, with pinkish tint in places. Head usual, densely setose; genae roundly subquadrate; epicranial groove evident but shallow. Antennae (Figs 1 ���2, 9) long and slender, weakly clavate; antennomeres 5 and 6 each with a distodorsal field of minute bacilliform sensilla; antennomere 7 dorsally with a small, but evident, claviform, transparent tubercle at base and a flat tubercle in distal quarter. Body polydesmoid, with 20 segments. Collum subequal in width to head and segments 2 and 3, with three indistinct transverse rows of very flat setigerous bosses; setae of front row slightly longer than others; paraterga broadly rounded, slightly elevated up to level of dorsum, devoid of evident indentations. Body parallel��sided on segments 7���17, thereafter abruptly tapering caudally. Paraterga of segment 2 elevated, not surpassing rear tergal contour. Subsequent paraterga slightly projecting caudad beyond rear tergal contour, evidently elevated above dorsum until about segment 10, onward about level to dorsum until segment 19, evidently below level of dorsum only on segment 19 (Figs 1���7). Caudal corners of all paraterga more or less narrowly rounded, never pointed. Tergal surface smooth, with three transverse setigerous rows of very flat bosses, forming a typical pattern (Fig. 5); setae minute, very short. Prozona very finely shagreened, metazona below paraterga microgranulate. Structure between pro�� and metazona nearly smooth, only sides with extremely faint transverse striolation. Limbus microdenticulate. Pore��bearing paraterga with four, poreless paraterga with three, minute lateral incisions. Pore formula normal, ozopores lying entirely dorsally, at bottom of a lanceolate groove near lateral margin between two caudolateral incisions (Figs 3, 5). Epiproct coniform (Figs 6���7), barely surpassing paraprocts caudally. Hypoproct distinctly trapeziform, with a pair of long setae at caudolateral corners. A distinct pleurosternal flap��shaped crest present only on segment 2 (Fig. 2), thereafter totally wanting. Sterna evidently setose, but without modifications. Gonopod aperture obcordate, with a small, rounded shelf caudomedially. Legs very long and slender (Figs 2, 4, 7, 10), especially so towards caudal body end, nearly twice as long as body height; telopodite devoid of sphaerotrichomes; claw very long and slender, distinctly curved (Fig. 10). Gonopods (Figs 8, 11��� 12) with typical, enlarged, subtriangular, medially fully coalesced coxae, each coxa bearing 2���3 macrosetae ventrally and a cannula dorsomedially. Telopodite elongate, relatively simple; acropodite (= branch lying distally of recurvature point of seminal groove, cf. Golovatch 1991; Djursvoll et al. 2001) split into two unequal parts: a longer, slender, distally curved endomere (a) and a shorter, lobuliform exomere (l). Endomere with a small denticle (d) frontally at base. Region near pulvillus, and that lying distolaterally of recurvature point of seminal groove, conspicuously papillate��spinulate, often with another small but clear denticle distally on mesal face. Accessory seminal chamber very small, inconspicuous. Orifice on pulvillus surrounded by spinules. Epigynal ridge low, slightly elevated and rounded only laterally. Vulvae usual, elongate, densely setose, with spiral canals at bottom of gutter. Remarks. Because this remarkable species shows several evident traits of troglomorphism, such as the completely unpigmented tegument and the elongate appendages, it can be considered as troglobitic with a fair degree of certainty. The cave Ishui Luo Dong supports two further diplopods, both presumably troglobitic and highly local: Guizhousoma latellai Mauri��s, 2005 (Chordeumatida, Guizhousomatidae) (Mauri��s 2005) and a Glyphiulus species from the granulatus ��group (Spirostreptida, Cambalopsidae) (Golovatch et al., in press). On the other hand, the cave Hei Dong harbours a tiny species of Doratodesmidae (Polydesmida), most probably troglobitic as well. The distance between the two caves being only about 20 air��km, both lying within the same karst massif situated between Dafang and Qianxi, it is hardly surprising that they contain the same species of Pacidesmus. In addition, Guizhousoma latellai has hitherto been encountered in as many as four of the massif���s caves (Mauri��s 2005). The discovery of P. martensi sp. n. in Guizhou ��� a species very closely related to and sympatric with P. sinensis, despite the latter���s poor original description (Loksa 1960)��� leaves no doubt whatever that they are congeneric. Therefore, the doubts expressed by Golovatch (1991) concerning the allocation of P. sinensis no longer apply., Published as part of Golovatch, Sergei I. & Geoffroy, Jean-Jacques, 2006, Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae), pp. 363-368 in Zootaxa 1325 on pages 364-367, DOI: 10.5281/zenodo.174082, {"references":["Golovatch, S. I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141 - 159.","Djursvoll, P., Golovatch, S. I., Johanson, K. A. & Meidell, B. (2001) Phylogenetic relationships within Polydesmus sensu lato (Diplopoda: Polydesmidae). Fragmenta Faunistica, 43 (Suppl.), 37 - 59 (for 2000).","Mauries, J. - P. (2005) Guizhousoma latellai gen. n., sp. n., de Chine continentale, type d'une nouvelle famille de la superfamille des Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta, 14 (1), 11 - 17.","Loksa, I. (1960) Einige neue Diplopoden- und Chilopodenarten aus chinesischen Hohlen. Acta Zoologica Academiae Scientiarum Hungaricae, 6 (1 - 2), 137 - 148."]}

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: http://doi.org/10.5281/zenodo.5393502, {"references":["CHEN J. & ZHANG C. Z. 1990. - A cave-dwelling new species of the diplopod genus Epanerchodus from Guizhou Province (Polydesmida: Polydesmidae). Acta Zootaxonomica Sinica 15: 406-409 (in Chinese, English summary).","CHEN Z., DECU V., JUBERTHIE C. & UENO S.-I. 2003. - Chine, in JUBERTHIE C. & DECU V. (eds), Encyclopaedia Biospeologica 3. Societe Internationale de Biospeologie, Moulis: 1763-1781.","DEGOUVE P., LIPS B. & LIPS J. 1997. - Speleologie au pays de l'homme sauvage no 3. Rapport de la 3e expedition speleologique en Chine, A.K.L. 1997, province du Sichuan. Federation francaise de Speleologie, Paris, 130 p.","ENGHOFF H., DOHLE W. & BLOWER J. G. 1993. - Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations. Zoological Journal of the Linnean Society 109: 103-234.","GEOFFROY J.-J. & GOLOVATCH S. I. 2004. - Some polydesmidan millipedes from caves in southern China (Diplopoda: Polydesmida), with descriptions of four new species. Arthropoda Selecta 13 (1-2): 19-28.","GOLOVATCH S. I. 1983. - Contributions to the millipede fauna of Vietnam (Diplopoda) I. Chordeumatida. Acta Zoologica Academiae Scientiarum Hungaricae 29 (1-3): 123-127.","GOLOVATCH S. I. 1984. - Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Academiae Scientiarum Hungaricae 30 (1-2): 53-77.","GOLOVATCH S. I. 1991. - The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia 17 (4): 141- 159.","GOLOVATCH S. I. & ENGHOFF H. 1994. - Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45-71.","GOLOVATCH S. I. & HOFFMAN R. L. 1989. - Identity of Polydesmus hamatus Brandt, 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae). Tropical Zoology 2: 159-164.","GOLOVATCH S. I. & WYTWER J. 2003. - A new genus and species of the millipede family Altajellidae from eastern Kazakhstan, Central Asia (Diplopoda: Chordeumatida). Annales Zoologici 53 (3): 579-584.","LOKSA I. 1960. - Einige neue Diplopoden- und Chilopodenarten aus chinesischen Hohlen. Acta Zoologica Academiae Scientiarum Hungariae 6 (1-2): 135-148.","MAURIES J.-P. 1982. - Une famille nouvelle et deux genres nouveaux de Cleidogonoidea, avec notes sur la classification de la superfamille (Diplopoda, Craspedosomida). Steenstrupia 8 (6): 165-176.","MAURIES J.-P. 1988. - Myriapodes du Nepal. II. Diplopodes craspedosomides nouveaux de l'Himalaya et de la region indo-malaise (Craspedosomidea et Chordeumidea). Revue suisse de Zoologie 96 (1): 3-49.","MAURIES J.-P. & NGUYEN DUY- JACQUEMIN M. 1997. - Nouveaux craspedosomides et glyphiulides cavernicoles de Chine (Diplopoda). Memoires de Biospeologie 24: 49-62.","SHEAR W. A. 1979. - Diplopoda from the Nepal Himalayas. Chordeumida with comments on the Asian chordeumid fauna. Senckenbergiana biologica 60 (1-2): 115-130.","SHEAR W. A. 1987. - Chordeumatid Diplopoda from the Nepal Himalayas, II. Courier Forschungsinstitut Senckenberg 93: 229-240.","SHEAR W. A. 1999. - A new troglobitic milliped of the genus Nepalella from China (Diplopoda, Chordeumatida, Megalotylidae). Myriapodologica 6 (1): 1-10.","SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63-72.","SHEAR W. A., SHELLEY R. M. & HEATWOLE H. 2003. - Occurrence of the milliped Sinocallipus simplipodicus Zhang, 1993 in Laos, with reviews of the Southeast Asian and global callipodidan faunas, and remarks on the phylogenetic position of the order (Callipodida: Sinocallipodidea: Sinocallipodidae). Zootaxa 365: 1-20.","SPELDA J. 2001. - Review of the millipede genus Pterygophorosoma Verhoeff, 1897 (Diplopoda, Chordeumatida, Craspedosomatidae). Andrias 15: 29-48.","STOEV P. 2004. - The first troglomorphic species of the millipede genus Paracortina Wang & Zhang, 1993 from south Yunnan, China (Diplopoda: Callipodida: Paracortinidae). Zootaxa 441: 1-8.","STOEV P. & GEOFFROY J.-J. 2004. - Review of the millipede family Paracortinidae Wang & Zhang, 1993 (Diplopoda: Callipodida). Acta Arachnologica 53 (2): 93-103.","WANG D. & MAURIES J.-P. 1996. - Review and perspective study of myriapodology of China, in GEOFFROY J.-J., MAURIES J.-P., NGUYEN DUY- JACQUEMIN M. (eds), Acta Myriapodologica. Memoires du Museum national d'Histoire naturelle 169: 81-99.","WANG D. & ZHANG C. Z. 1993a. - A new family of millipeds (Diplopoda: Callipodida) from southwestern China. Memoirs of the Peking Natural History Museum 53: 375-390 (in Chinese, English summary).","WANG D. & ZHANG C. Z. 1993b. - Diplopoda in caves of Yunnan I. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida), in SING LINHUA & TING HUAIYUAN (eds), Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205-219 (in Chinese, English summary).","ZHANG C. Z. 1993. - Diplopoda from Yunnan caves II. Contribution to the study of a new cavernous taxon of the nematophoran millipeds (Diplopoda: Coelocheta: Callipodida). Proceedings of the XIth International Congress of Speleology: 128-130.","ZHANG C. Z. & CHEN Z. 1983. - A new species of the genus Epanerchodus (Diplopoda: Polydesmidae) from Zhejiang Province. Acta Zhejiang Teacher's College (Nature Sciences) 6: 87-89 (in Chinese, English summary)."]}

Hyleoglomeris montana Golovatch, 1983 MATERIAL EXAMINED. — Thailand. Doi Inthanon National Park, 2000-2540 m, 13.XI.1984, leg. P. Beron & S. Andreev, 1 ♂, 1 ♀, 1 juv. (NMNHS); 1 ♀ (MNHN CC162). REMARKS This new material,which is strictly topotypical, allows a slight improvement to the original description (Golovatch 1983) in that the background coloration of the adults is not always blackish but gray-brown. The pattern is exactly as described earlier, but the axial stripe and the pale spots on all terga are slightly larger or wider., Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques & Mauriès, Jean-Paul, 2006, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia, pp. 887-915 in Zoosystema 28 (4) on page 894, DOI: 10.5281/zenodo.4525415, {"references":["GOLOVATCH S. I. 1983. - On several new Glomeridae (Diplopoda) from Indochina. Annales Historico-Naturales Musei Nationalis Hungarici 75: 107 - 116."]}

Golovatch, Sergei I., Geoffroy, Jean-Jacques (2006): Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae). Zootaxa 1325: 363-368, DOI: 10.5281/zenodo.174082

KEY TO THE SPECIES OF NEPALELLA S HEAR, 1979 1. Adults with 28 body segments (27 pleurotergites in terms of Enghoff et al. 1993)................................................................................................... N. phulcokia Mauriès, 1988 — Adults with 30 body segments.................................................................................... 2 2. Body length ≥ 30 mm, width 2.8-3.0 mm.................................................................. 3 — Body length ≤ 25 mm................................................................................................ 5 3. Body particularly large, ≥ 40 mm long; tegument and ocelli pallid; tergal macrochaetae medium-sized (Fig. 1 D-F); troglobite from Yunnan Prov., China........ N. grandis n. sp. — Body 30-33 mm long, tegument and ocelli distinctly pigmented; macrochaetae short (Fig. 1 A-C)................................................................................................................ 4 4. Each eye patch with 15-17 ocelli; troglophile from Sichuan, China.... N. marmorata n. sp. — Each eye patch with 25 ocelli; epigean in Yunnan, China.......... N. magna Shear, 2002 5. Body tegument pallid, but eye patches and antennae pigmented; body 21-23 mm long, 2.6-2.7 mm wide due to paraterga being well developed, in the form of distinct dorsolateral keels; tergal setae long; male legs 3-7 not enlarged; Myanmar... N. pallida Mauriès, 1988 — Body either entirely pallid (cavernicole) or distinctly pigmented; body width ≤ 2.3 mm, paraterga largely poorly developed, like indistinct dorsolateral swellings; tergal setae medium-sized at most; male legs 3-7 very often crassate.................................................. 6 6. Body length ≥ 18 mm, width ≥ 1.9 mm; coloration uniformly brown, male coxa 10 without distinct processes........................................................................................... 7 — Never all these three characters combined.................................................................. 8 7. Posterior gonopods (male legs 9) with colpocoxites divided distally into three branches; only male femur 4 with a rounded gibbosity ventrally; Thailand.................................................. N. taiensis Mauriès, 1988 (N. t. taiensis and N. t. inthanonae Mauriès, 1988) — Posterior gonopods (male legs 9) with colpocoxites not divided distally; male femora 3 and 4 each with a fungiform protuberance ventrally; Nepal......... N. gunsa Shear, 1987 8. Body entirely pallid; ocelli 9, reduced, only slightly pigmented and widely separated; male legs 3-7 not modified; cave in Guizhou Prov., China.................... N. caeca Shear, 1999 — Body pigmented, ocelli dark and never reduced,> 20 in number, always forming a compact triangular eye patch on each side of head; male legs 3-7 most often crassate........ 9 9. Body c. 16 mm long and 2.2 mm wide due to relatively well developed, shoulder-shaped paraterga; male femur 4 with a distal knob subtending a distal depression on ventral side; Yunnan.................................................................................. N. griswoldi Shear, 2002 — Body width usually ≤ 1.9 mm; paraterga modestly to poorly developed; male femur 4 either unmodified or otherwise modified.................................................................. 10 10. Tergal setae short and blunt; male legs 3-7 crassate, but without further modifications; posterior gonopod telopodite relatively strongly reduced, much shorter than colpocoxites; Yunnan.................................................................................... N. pianma Shear, 2002 — Tergal setae short to medium-sized, but always acute; at least some of male legs 3-7 usually with modifications; telopodite of posterior gonopods hypertrophied, (sub)equal in height to colpocoxite................................................................................................ 11 11. Body width 1.8-2.0 mm; male legs 3-7 with tarsal papillae and dorsally inflated prefemora; Yunnan.............................................................................. N. kavanaughi Shear, 2002 — Body width usually ≤ 1.9 mm; tarsal papillae largely absent; male prefemora 3-7 not enlarged dorsally.......................................................................................................... 12 12. Tarsal papillae present on most of male legs; male prefemur 11 with a long, digitiform, parabasal process; Vietnam.......................................... N. vietnamica Golovatch, 1983 — Tarsal papillae absent; male prefemur 11 devoid of processes.................................... 13 13. Claw simple; male coxa 10 with a long process distoventrally; male 11 coxa at most with one small process distoventrally, usually devoid of any; Nepal.................................. 14 — Claw complex, with both a minute accessory claw dorsally and a long setoid filament ventrally at base; male coxae devoid of processes, somewhat reduced and fused with sternite into a coxosternum subtending voluminous coxal glands, male coxa 11 with two small processes distoventrally; Myanmar.................................... N. birmanica Mauriès, 1988 14. Coloration ochraceous, with four dark, brown, longitudinal stripes.......................... 15 — General coloration ochraceous to brownish, with spots, or metazona dark............... 18 15. Colpocoxites of posterior gonopods divided into three branches or lobes................. 16 — Colpocoxites of posterior gonopods poorly divided distally into only two short branches................................................................................................................................. 17 16. Larger: 16-17 mm long, 1.8-1.9 mm wide; colpocoxite of posterior gonopods divided into three lobes; male coxa 10 with a C-shaped process... N. tragsindola Mauriès, 1988 — Smaller: 10-12 mm long, 1.0- 1.3 mm wide; colpocoxite of posterior gonopods divided into two lobes and a slender acuminate branch (solenomere?); male coxa 10 with a coniform process topped by a rounded, microgranolate bulge... N. gairiensis Mauriès, 1988 17. Larger: 17 mm long, 1.6 mm wide (♂); both branches of colpocoxite very short and erect; male coxa 10 with a bifid process................................... N. ringmoensis Mauriès, 1988 — Smaller: 11-14 mm long, 1.3-1.5 mm wide (♂, ♀); internal branch of colpocoxite directed medially; male coxa 10 with a subtruncate process.......... N. deharvengi Mauriès, 1988 18. Male coxa 10 with a straight, apically truncate process; male prefemora 3-7 each with a distoventral knob; male coxa 11 without gland, but with a small distomedial process................................................................................................... N. thodunga Shear, 1979 — Male coxa 10 with a curved, apically acuminate process; male prefemora 3-7 either unmodified or only third and fourth with distoventral knobs; male coxa 11 at most with a small gland, devoid of any processes......................................................................... 19 19. Male coxa 10 with a strong unciform process directed caudally; male femora 3-7 each with a fungiform protuberance at midway ventrally............. N. taplejunga Shear, 1987 — Male coxa 10 with process directed laterad; male femora 3-7 unmodified................. 20 20. Larger: c. 14 mm long, 1.4-1.5 mm wide; tergal setae medium-sized; male prefemora 3 and 4 with distoventral knobs.......................... N. khumbua Shear, 1979 (type species) — Smaller: c. 10 mm long, 1.0 mm wide; tergal setae short; male legs 3 and 4 without such modifications....................................................................... N. jaljalae Mauriès, 1988, Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques & Mauriès, Jean-Paul, 2006, Four new Chordeumatida (Diplopoda) from caves in China, pp. 75-92 in Zoosystema 28 (1) on pages 83-85, DOI: 10.5281/zenodo.5393502, {"references":["ENGHOFF H., DOHLE W. & BLOWER J. G. 1993. - Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations. Zoological Journal of the Linnean Society 109: 103 - 234.","MAURIES J. - P. 1988. - Myriapodes du Nepal. II. Diplopodes craspedosomides nouveaux de l'Himalaya et de la region indo-malaise (Craspedosomidea et Chordeumidea). Revue suisse de Zoologie 96 (1): 3 - 49.","SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63 - 72.","SHEAR W. A. 1987. - Chordeumatid Diplopoda from the Nepal Himalayas, II. Courier Forschungsinstitut Senckenberg 93: 229 - 240.","SHEAR W. A. 1999. - A new troglobitic milliped of the genus Nepalella from China (Diplopoda, Chordeumatida, Megalotylidae). Myriapodologica 6 (1): 1 - 10.","GOLOVATCH S. I. 1983. - Contributions to the millipede fauna of Vietnam (Diplopoda) I. Chordeumatida. Acta Zoologica Academiae Scientiarum Hungaricae 29 (1 - 3): 123 - 127.","SHEAR W. A. 1979. - Diplopoda from the Nepal Himalayas. Chordeumida with comments on the Asian chordeumid fauna. Senckenbergiana biologica 60 (1 - 2): 115 - 130."]}

Scutigerina malagassa (de Saussure et Zehntner, 1902) Scutigera malagassa de Saussure et Zehntner, 1902: 295, Pl. XII, fig. 1 ��� 1 c, Pl. XIV, fig. 19. Type locality: Madagascar: Sikora. Syntypes: females, MHNG. New record: MADAGASCAR: damaged ad. F, legs detached, Tananarive, 29.ix. 1897. Coll. Br��lemann, 1902 ��� CMXIV. General distribution: Madagascar: Sikora; Antananarivo; P��rinet, Sahamaloto; Andranoboka Cave near Majunga; Maroantsetra, Ambohitsitondrona., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 10, DOI: 10.5281/zenodo.158015

Pselliodes aff. vagans (Verhoeff, 1905) Pselliophora vagans Verhoeff, 1905 b: 95 ���97. Type locality: Unteres Kongogebiet: Chinchoxo (Tschinschoscho), n��rdlich der Kongom��ndung. Holotype: male, Nos 3893 a���e, Prep. Nos 2617���2621, ZMB. Material examined: 2 specimens ��� Pselliophora sp.��� J.��M. Demange det. 8 / 69 Africa, Guinea, Nimba 7 Cb, Environs base cote Keoulenta, Savane �� Andropogon, 5.viii. 1951, M. Lamotte leg. Remarks: The African and Neotropical family Pselliodidae has never been comprehensively revised, and is still a great challenge for taxonomists interested in the systematics of the Scutigeromorpha. Two African morphotypes, P. rugosus (East Africa) and P. vagans (West Africa), can easily be distinguished solely on the basis of their coloration. Our specimens, although badly damaged, are more like the latter, hitherto known from Angola (Chinchoxo; Landana, Enclave de Cabinda), Gabon (Moanda) and Zaire (Eala, Luali, N���Gongo Bata)., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 10, DOI: 10.5281/zenodo.158015, {"references":["Verhoeff, K. W. (1905 b) Uber Scutigeriden. 5. Aufsatz. Zoologischer Anzeiger, 29, 73 - 103; (4), 105 - 119."]}

Ballonemella jeanneli (Ribaut, 1914) Ballonema Jeanneli Ribaut, 1914: 6, fig. 1���3. Type locality: Afrique orientale allemande, R��gion c��ti��re, grotte A du Kulumuzi, niveau de la mer, st. no. 74. Holotype: ad. F, MNHN, Q 24. General distribution: Tanzania: A du Kulumuzi Cave near Tanga., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 4, DOI: 10.5281/zenodo.158015

Scutigera linceci (Wood, 1867) Cermatia Linceci Wood, 1867: 42 ���43. Type locality: USA: Texas. Syntypes: whereabouts unknown. Material examined: The MNHN houses specimens identified by H.W. Brolemann from Costa Rica (Cuesta del Tablazo, 1 500 m) and Guatemala. General distribution: Costa Rica. Cuba. El Salvador. Granada. Guatemala. Honduras. Mexico. Nicaragua. Panama. USA (Texas & Arizona)., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 4, DOI: 10.5281/zenodo.158015

Thereuopoda clunifera (Wood, 1862) Cermatia clunifera Wood, 1862: 10. Type locality: China: Hong Kong. Holotype: one specimen, sex unknown, No 255, USNM. Material examined: The MNHN houses specimens from Japan (Nakao��d�� Cave, Akiyoshi Plateau and Tokyo) identified by M. W��rmli. General distribution: China. Japan. Taiwan. Java? Remarks: It is very likely that Thereuopoda clunifera is a junior synonym or, at the best, a subspecies of T. longicornis (Fabricius, 1793) (cf. W��rmli, 1979). The records from Java ��� Batavia and Buitenzorg (Chamberlin, 1939; 1944), and Salak (Attems, 1903) are most probably based on specimens of typical T. longicornis., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 7, DOI: 10.5281/zenodo.158015, {"references":["Wurmli, M. (1979) Taxonomic problems in the genus Thereuopoda (Chilopoda: Scutigeromorpha: Scutigeridae): the role of postmaturational moultings. In: Camatini, M. (Ed.). Myriapod biology. Academic Press, London, New York, etc., 39 - 48","Chamberlin, R. V. (1939) On a collection of chilopods from the East Indies. Bulletin of the University of Utah, 29 (12), Biological Series, 5 (2), 1 - 19.","Chamberlin, R. V. (1944) Some chilopods from the Indo-Australian Archipelago. Notulae Naturae, Academy of Natural Sciences of Philadelphia, 147, 1 - 14.","Attems, C. G. (1903) Beitrage zur Myriopodenkunde. Zoologische Jahrbucher, Abteilung fur Systematik, 18, 63 - 154, t. 5 - 11."]}

Parascutigera nubila Ribaut, 1923 Parascutigera nubila Ribaut, 1923: 3, 20, figs 1���4. Type locality: Nouvelle��Cal��donie: Mont Canala. Types: 2 males, 3 females, syntypes, No 296 a, NMB. New records: NOUVELLE��CAL��DONIE: ad. M, St. 210. Barrage de la Neaoua (Ouen Sieu), 165 �� 32 ��� 35 ���E; 21 �� 21 ��� 57 ���S, 500 m, For��t humide, 20.xii. 1984, S. Tillier & P. Bouchet leg. General distribution: New Caledonia (Tchalabel; vall��e de la Tiouaka; Kon��; Mont Canala; Mt. Dogni; Barrage de la Neaoua). Remarks: A re��description of this species can be found in W��rmli (1974) who studied material from Mt. Dogni., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 8, DOI: 10.5281/zenodo.158015, {"references":["Wurmli, M. (1974) Ergebnisse der Osterreichischen Neukaledonien-Expedition 1965. Chilopoden. Annalen des naturhistorischen Museum in Wien, 78, 523 - 533."]}

Scutigera tonsoris W��rmli, 1977 Scutigera tonsoris W��rmli, 1977: 128 ���130, Abb. 9���16. Type locality: Spain: Sevilla. Holotype: subadult F, ZMUC. Paratype: subadult M, MWC. New records: ALGERIA: ad. M, Bougie and its environs, Duruy, 1902 leg.; TUNI�� SIA: ad. F, Sfax, 15.vi��� 11.vii. 1955, P. B��d�� leg. General distribution: Algeria (Bona and Bougie). Spain (Sevilla and Almaden de la Plata). Tunisia (Khroumirie and Sfax)., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 4, DOI: 10.5281/zenodo.158015

Thereuonema tuberculata (Wood, 1862) Cermatia tuberculata Wood, 1862: 12. Type locality: China: Hong Kong. Holotype: sex unknown, No 254, USNM. Material examined: The MNHN houses specimens identified by H.W. Brolemann and P. Stoev from China (W of Shanghai), Korea (Tsenongsan near Pyongyang) and Japan (Geji��geji). New records: CHINA: ad. F, Manjuria, Ourga �� Tsitsikhar, Chaffanjon leg.; KOREA: 2 subad. MM, Seoul, 06.ix. 1973, P. Jonet Coll. General distribution: China. Japan. Korea. Taiwan. Quelpart Island., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 6, DOI: 10.5281/zenodo.158015

Madagassophora hova (de Saussure et Zehntner, 1902) Scutigera hova de Saussure et Zehntner, 1902: 294, Pl. XIV, fig. 17; Pl. XII, fig. 2. Type locality: Madagascar: Sikora. Syntypes: several juveniles, MHNG. New records: MADAGASCAR: ad. M, R.C.P. 225, Massif de l'Andrangitra 1970��� 1971, and a second label RCP��mad�� 268; ad. F, juv., R.C.P., Massif de l'Andrangitra 1970��� 1971, 228 Mad.; ad. F, Manjakatompo, P. Tananarive, 10.xi. 1963, P. Remy leg.; ad. M & F, Manjakatompo, N 238, 04.vii. 1957, P. Remy leg.; ad. F, Manjakatompo, R��gion Station, vi. 1957, P. Remy leg.; ad. F, Manjakatompo, near laboratory, 03.vii. 1957, P. Remy leg. General distribution: Madagascar: Manjakatompo Forest near Ankaratra; Tsiafajavona, 2 500 m; Ambatofinandrahana, Itremo Summit; 26 km E of Antananarivo; Manjakandriana, Angavokely., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 11, DOI: 10.5281/zenodo.158015

Thereuopoda longicornis (Fabricius, 1793) Scolopendra longicornis Fabricius, 1793: 390. Type locality: Tranquebar. Type: whereabouts unknown. Material examined: The MNHN houses specimens identified by C. Attems, H.W. Brolemann and M. W��rmli from the following localities: CAMBODIA: Bokkor; Province Kampot: region Kompong: Trach and Tuk Meas Caves; VIETNAM: Tonkin: Caverne de Luc��an��Chau; Tonkin: Grotte de Than��moi; Tonkin: Hagiang; Bana, 1 400 m; Hu�� (Annam); Ca��da (Nhatrang); Hongay; LAOS: Province du Cammon: v. Na Kay Khia, alt. 140 m; v. Tham, alt. 140 m; INDIA: Sambalpur; INDONESIA: Borneo: Pantianak. UNKNOWN COUNTRIES: Keam; Pathang; Caudora [?]; Lue He; Cochinchine, Cap St�� Jacques, Capt. M. Modoit leg., 1908; Malacca Peninsula, Erington de la Croc et Chap�� leg., 1900; (c��te or.) Pelambang, forest Nirou. Along with the above material the types of Scutigera feae Pocock, 1891 (ad. M, Scutigera feae Pocock, Birmanie, Palon, Viaggio in Birmania di L. Fea, 1885 ��� 89) and S. birmanica Pocock, 1891 (ad. & subad. MM, Pocock Birmania, Bhamo, Viaggio in Birmania di L. Fea, 1885 ��� 89) are also present in the collection. New records: VIETNAM: ad. F, Dong Nai Province, Dinh Quan, Tunnel de lave 1, 23.xii. 1994, by hand, Deharveng, Tam & Bedos leg.; INDONESIA: ad. M, Malukku, Halmahera Island, Sagea, Batu Lubang Cave, 28.vii. 1988, by hand, Deharveng & Bedos leg.; ad. M & F, Java, 2428 �� 78 de la Savani��re. General distribution: Asia: Bhutan. Cambodia. China. India. Indonesia (Borneo, Halmahera, Java, Lombok, Nias, Sumatra, Sumbawa). Japan. Laos. Malaysia. Mauritius. Myanmar. Nepal. New Ireland. Nicobar Islands. Philippines. Quemoy (Kin��Men) Island. Singapore. Sri Lanka. Taiwan. Thailand. Vietnam. Australia. Remarks: The collection contains material of Scutigera feae and S. birmanica, both bearing the label ���Exempl. Type ���. Seemingly, this material is part of the type series represented by numerous specimens of different sex and age (cf. Pocock, 1891), and currently stored in the collections of the ZMH (Weidner, 1960), ZMUC (W��rmli, 1979) and perhaps also the BMNH., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 7, DOI: 10.5281/zenodo.158015, {"references":["Pocock, R. I. (1891) On the Myriapoda of Burma - Part 2. Report upon the Chilopoda collected by Sig. L. Fea and Mr. E. W. Oates. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 10, 401 - 432.","Weidner, H. (1960) Die entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg. III Teil. Chilopoda und Progoneata. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut, 58, 57 - 104.","Wurmli, M. (1979) Taxonomic problems in the genus Thereuopoda (Chilopoda: Scutigeromorpha: Scutigeridae): the role of postmaturational moultings. In: Camatini, M. (Ed.). Myriapod biology. Academic Press, London, New York, etc., 39 - 48"]}

Thereuonema microstoma (Meinert, 1886) New Combination! Scutigera microstoma Meinert, 1886 b: 173. Thereuonema syriaca Verhoeff, 1905 b: 88 ���90 New Synonymy! Type locality: a place 70 miles from Amballa and Koolloo, India. Lectotype: male in ZMUC (present designation); Paralectotypes: female in ZMUC; several specimens of different sex and age in MCZ. Material examined: The MNHN houses specimens identified by H.W. Brolemann, H. Ribaut and C. Attems from the following localities: SYRIA: Doummar; Kouta��f��; Ata��be, E of Damas; A��n��Fidj��, Djebel Kasioun; ERITREA: Saganeiti (sub Scutigera aethiopica Silvestri); as well as the holotype (adult male) of Thereuonema tropicalis Ribaut, 1907 from Baltchi, Ethiopia. The latter was recognised as a junior synonym of Thereuonema syriaca by W��rmli (1975). General distribution: Asia: India (East). Iran. Iraq. Israel. Jordan. Lebanon. Pakistan. Palestine. Saudi Arabia. Syria. Turkey (East Turkey, Kurdistan region). Yemen. Africa: Egypt. Eritrea. Ethiopia. Libya. Sudan. Remarks: Meinert (1886 b) described Scutigera microstoma from two localities ��� Amballa and Koolloo (=? Kulu) in NW India, both lying close to the border with Pakistan. Recently, Stoev (2002) noted that this species is identical to Thereuonema syriaca Ve rhoeff, 1905, the latter being widespread from the Near East and eastern Africa to Pakistan and NE India (W��rmli, 1975; Stoev, 2002). We use this opportunity to elucidate the taxonomic status of microstoma and to show that it is a valid species of Thereuonema, conspecific with Verhoeff���s T. syriaca. The syntypes from the collections of the ZMUC and the MCZ were studied (courtesy to Prof. H. Enghoff and Mrs. Laura Leibensperger) and compared with non type material of syriaca from Turkey, Yemen, Iran and Egypt. In spite of the different coloration in the type material, undoubtedly resulting from long preservation in alcohol, we could not find any further differences in the setal ornamentation, shape of terga, second maxillae and female gonopods. Hence, the following new combination and synonymy: Thereuonema microstoma (Meinert, 1886) New Combination = Thereuonema syriaca Verhoeff, 1905 New Synonym. This taxonomic alteration is in agreement with the article 23.1. of the International Code of Zoological Nomenclature (1999). The name T. syriaca has been cited only in 14 scientific papers published in the period 1905���2003. We cannot predict how long this name will be in use, as there is at least one older candidate to replace it, namely Cermatia rubrilineata Newport, 1844 (not rubrolineata after Newport (1845) and all subsequent authors), a possibility suggested by Haase (1887)., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on pages 5-6, DOI: 10.5281/zenodo.158015, {"references":["Meinert, F. (1886 b) Myriapoda Musei Cantabrigiensis, Mass. Part I. Chilopoda. Proceedings of the American Philosophical Society, 23, 161 - 233.","Verhoeff, K. W. (1905 b) Uber Scutigeriden. 5. Aufsatz. Zoologischer Anzeiger, 29, 73 - 103; (4), 105 - 119.","Wurmli, M. (1975) Revision der Hundertfusser-Gattung Thereuonema (Chilopoda: Scutigeridae). Entomologica Germanica, 2 (2), 189 - 196.","Stoev, P. (2002) The scutigeromorphs (Chilopoda: Scutigeromorpha) in the collection of the National Museum of Natural History, Sofia. Historia naturalis bulgarica, 15, 79 - 85.","ICZN (International Comission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Forth Edition. International Trust for Zoological Nomeclature, London, 1 - 306 + i - xxix.","Newport, G. (1844) A List of the species of Myriapoda, Order Chilopoda, contained in the cabinets of the British Museum, with synoptic descriptions of forty-seven new Species. Annals and Magazine of Natural History, 13, 94 - 101.","Newport, G. (1845) Monograph of the class Myriapoda, Order Chilopoda; with observations on the general arrangement of the Articulata. Transactions of the Linnean Society of London, 19, 265 - 302 (1844), 349 - 439 (1845).","Haase, E. (1887) Die Indisch-Australischen Myriapoden. I. Chilopoden. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden, 5, 118 pp."]}

Pselliodes rugosus (Newport, 1844) New Combination! Cermatia rugosa Newport, 1844: 95. Type locality: Africa. Type: number and sex unknown, BMNH. Material examined: The MNHN houses material from East Africa identified by H. Ribaut. New record: MOZAMBIQUE: ad. M, Zambezi, Nova Choufanga near Chemba, P. Lesne leg., 1929. General distribution: Kenya: Merifano. ���near some pools of water a little to the west of Shebeli River���. Mozambique: Nova Choufanga near Chemba. Remarks: Newport (1844) described Cermatia rugosa from Africa without specifying the exact locality. Since then this species has been reported only by Gray (1844), Newport (1845, 1856), Gervais (1847) and Pocock (1896, 1899). Its identity is not as yet entirely clear, although it certainly belongs to the genus Pselliodes Chamberlin, 1921, and perhaps is identical with some (if not all) of the species described by Verhoeff (1905 b) from East Africa and almost indistinguishable from each other. However, whether it is a good species can only be established after a re��examination of the types of Verhoeff and Newport, fortunately these are still preserved in the collections of the ZMB and the BMNH. We formalize here the following new combination: Cermatia rugosa Newport, 1844 = Pselliodes rugosus (Newport, 1844). It should be noted that Scutigera rugosa reported by Porath (1871) from South Africa, a record repeated by Attems (1928) is not identical with Pselliodes rugosus (Newport, 1844) (cf. Pocock, 1893). Having access to the types of Newport, and also to new material from the same region of Africa (Cape Town), Pocock recognised that his specimens and Porath���s differed significantly from Newport���s Cermatia rugosa, and described them as a new species, Scutigera planiceps. Its type locality (South Africa: Cape Town: Simon���s Bay) is, at present, part of the range of only two scutigeromorphs ��� the widespread subcosmopolitan synantrope of European origin Scutigera coleoptrata, and the autochthonous Scutigerina weberi Silvestri, 1903. The original description of Scutigera planiceps excludes the possibility that it is conspecific with coleoptrata, but shows it to have the external characters of Scutigerina weberi. However, in order to formalize this synonymy the type of S. planiceps, currently preserved in the collection of the BMNH (specimen 10 mm long, most legs missing but antennae still attached, the registered number BM1891.5.18.1, Paul Hillyard, in lit.), should be studied., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 9, DOI: 10.5281/zenodo.158015, {"references":["Newport, G. (1844) A List of the species of Myriapoda, Order Chilopoda, contained in the cabinets of the British Museum, with synoptic descriptions of forty-seven new Species. Annals and Magazine of Natural History, 13, 94 - 101.","Gray, J. E. (1844) List of the specimens of Myriapoda in the collection of the British Museum. Newman, London, v - vi + 1 - 16.","Newport, G. (1845) Monograph of the class Myriapoda, Order Chilopoda; with observations on the general arrangement of the Articulata. Transactions of the Linnean Society of London, 19, 265 - 302 (1844), 349 - 439 (1845).","Newport, G. (1856) Catalogue of the Myriapoda in the collection of the British Museum. Part I. Chilopoda. Taylor and Francis, London, 1 - 96.","Gervais, P. (1847) Chilopodes. Schizotarses. In: Walckenaer, B. & Gervais, P. Histoire naturelle des Insectes. Apteres. Tome Quatrieme. Librairie Encyclopedique de Roret, Paris, 213 - 226.","Pocock, R. I. (1896) On the scorpions, centipedes, and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History, Series 6, 17, 425 - 444, Pl. XVIII.","Pocock, R. I. (1899) Solifugae, Scorpiones, Chilopoda and Diplopoda. Appendix C to A. Donaldson Smith's \" Through unknown African Countries \", 392 - 407.","Verhoeff, K. W. (1905 b) Uber Scutigeriden. 5. Aufsatz. Zoologischer Anzeiger, 29, 73 - 103; (4), 105 - 119.","Porath, C. O. von (1871) Myriopoda Africae australis, in Museo Regio Holmiensi asservata, recensuit. Pars. I. Chilopoda. Ofversigt af Kunglika Vetenskaps-Akademiens Forhandlingar, Stockholm, 9, 1135 - 1167.","Attems, C. G. (1928) The Myriopoda of South Africa. Annals of the South African Museum, 26, 1 - 431, 84 figs.","Pocock, R. I. (1893) Report upon the Myriapoda of the \" Challenger \" Expedition, with Remarks upon the Fauna of Bermuda. Annals and Magazine of Natural History, Series 6, 11, 121 - 142, Pl. IX."]}

Family Scutigeridae Gervais, 1837 Scutigera coleoptrata (Linnaeus, 1758) Scolopendra coleoptrata Linnaeus, 1758: 638. Type locality: ���Habitat in Hispania ���. Type: whereabouts unknown. Material examined: The MNHN houses specimens identified by H. Lucas, H.W. Brolemann, J.��M. Demange, M. W��rmli and J.��J. Geoffroy from the following localities: ALGERIA: Bona; Miliana; ARGENTINA: Buenos Aires; CANARY ISLANDS: Grand Canaria Island: Las Palmas; Tenerife Island; CROATIA: Abbazia; FRANCE: Alpes��Maritimes: Menton, Borigo, La Caunet��de��Cannes; Ard��che: Grotte de la Vierge, Grotte de Tourange; Bouches��du��Rh��ne: La Crau pr��s d'Arles; Calvados: Honfleur; Charente: Le Plantier; Cher: Ainay��le��Vieil, Bourges, Osmery, St��Amand��Montrond; Dr��me: Le Po��t�� Laval; Essonne: Epinay��sous��S��nart; Etampes, ���B��ou de Vitte���; Gard, Pont du Gard; Gers; Hauts��de��Seine: Meudon, Sable des Carri��res; Loire��Atlantique: Le Pouliguen; Ni��vre: Corbigny; Pyr��n��es��Atlantiques: Pau (in houses); Hautes��Pyr��n��es: near grotte de Doulas St��P����de��Bigorre; Pyr��n��es��Orientales: Thuis, Banyuls��sur��Mer (Laboratory), Terge��de�� Sorede; Seine: Paris and suburbs; Var: French Riviera Coast; Vaucluse: Avignon; Vend��e: Saint Michel��en��Lherm; [unknown department]: Ruine, Bourdeaux; [unknown department]: Ruine la Beconne; [unknown department]: Ruissol; IRAN: Perse: Mechedissav; ITALY: Romagna: Santa Sofia, Meldola (Forl��); Lombardia: San Giovanni, Cremona; Milano; Alzate; LEBANON: Broumana; MADEIRA: Rabasal; MONACO; MOROCCO: Tanger; ROMANIA: Dobroudja: Macin Iglitza, Mangalia; SPAIN: Lomo de Bas��Anglais; Rozaelo de Calatrava; SOUTH AFRICA: " Colonie du Cap " [now Cape Province]: East London, Steynsburg; TUNISIA: Tunis; USA: New York City. New record: ALGERIA: 2 ad. FF, Bougie and its environs, Duruy leg., 1902. General distribution: Europe: Albania. Austria. Azerbaijan. Bosnia and Herzegovina. Bulgaria. Channel Isles (introduction?). Croatia (incl. Cherso Is). Czech Republic (Morava). France (South, incl. Corsica, synanthropic in Middle and North). Georgia. Germany (South). Greece (incl. Aegean and Ionian islands). Hungary (South). Italy (incl. Sicily, Sardinia). Malta. Monaco. Montenegro. Portugal (incl. Madeira). Republic of Macedonia. Romania. Russia. Salvage Islands. Serbia. Slovak Republic. Slovenia. Spain (incl. Balearic and Canary islands). Switzerland. Ukraine. Asia: Iran. Iraq. Israel. Jordan. Lebanon. Palestine. Syria. Turkey. Turkmenistan. Africa: Algeria. Egypt. Libya. Morocco. Tunisia. Introduced: Angola. Argentina. Bermudas: St. David���s Is. Cameroon. Canada. Cape Verde Islands. Denmark. Kenya. Mexico. Mozambique. Republic of South Africa. Saint Helena Island. Taiwan. Tanzania. The Netherlands. UK: England, Scotland, and Ireland. Uruguay. USA. Vietnam. Zimbabwe. Probably Introduced: Azores., Published as part of Stoev, Pavel & Geoffroy, Jean-Jacques, 2004, An annotated catalogue of the scutigeromorph centipedes in the collection of the Mus��um National d'Histoire Naturelle, Paris (France) (Chilopoda: Scutigeromorpha), pp. 1-12 in Zootaxa 635 on page 3, DOI: 10.5281/zenodo.158015

International audience; Chilopode lithobiomorphe troglobie d'Espagne, Lithobzus drescoi Demange, 1958, présente un développement remarquable des pattes locomotrices et des antennes, traits caractéristiques d'une adaptation aux milieux souterrains profonds (troglomorphie).

Mauriès, Jean-Paul, Geoffroy, Jean-Jacques (2000): Nouvelle description, classification, répartition et variations morphologiques interpopulations d'un diplopode troglobie du sud-est du Brésil (Diplopoda, Polydesmida, Chelodesmidae). Zoosystema 22 (1): 153-168, DOI: http://doi.org/10.5281/zenodo.5399263, {"references":["MZUSP 814. 1942. Dreifuss/Pavan coll. Areias de Baixo. 1 . Forme \" type \". Une preparation dessinee.","MZUSP 815. 1942. Dreifuss/Pavan coll. Areias de Baixo. 1 . Forme \" type \".","MZUSP 816. 1942. Dreifuss/Pavan coll. Areias de Baixo. 2 . Forme \" type \".","MZUSP 817. 3344. 19. II.1951. Schubart coll. Areias de Cima. 4 . Forme \" acutus \". Une preparation, un dessin.","MZUSP 818. 1942. Dreifuss/Pavan coll. Areias de Baixo. 1 (lectotype) disseque pour preparations. Forme \" type \".","MZUSP 819. 1942. Dreifuss/Pavan coll. Areias de Baixo. 1 , une preparation dessinee. Forme \" type \".","MZUSP 820. 1942. Dreifuss/Pavan coll. Areias de Baixo. 4 dissequees. Forme \" type \".","MZUSP 821. 3344. Areias de Cima. 19. II.1951. 3 . Forme \" acutus \".","MZUSP 822. 3344. Grutas de Areias de Cima. 19. II.1951. 1 , preparation microscopique. Forme \" acutus \".","MZUSP 823. 3344. Areias de Cima. 19. II.1951. 3 . Forme \" acutus \".","MZUSP 824. Areias de Cima. 19. II.1951. Schubart coll. 1 , 2 . Forme \" acutus \".","MZUSP. Caverna Agua Quente (Ressurgencia das Areias). ?. ?.1979. N. S. Cesar Jr coll. 1 , une preparation. Forme \" type \".","MZUSP. Caverna Areias de Cima. 10.XI.1982. E. Trajano coll. 1 . Forme \" acutus \".","MZUSP. Caverna Areias de Cima. 24.VIII.1983. N. M. Godoy coll. 4 , une preparation non dessinee. Forme \" acutus \".","MZUSP. Caverna Areias de Cima. 24.VIII.1983. N. M. Godoy coll. 4 . Forme \" acutus \".","MZUSP. Caverna Agua Quente (Ressurgencia das Areias), Bairro da Serra. 22.X.1983. N. M. Godoy coll. 2 , 2 , 1 preparation. Forme \" type \".","MZUSP. Caverna Alambari de Cima. 23.?.1984. E. Trajano coll. 1 , 1 . Forme \" alambari \".","MZUSP. Caverna Alambari de Cima. 8.IX.1990. M. C. Chamani et J. P. M. Seino coll. 1 . Forme \" alambari \".","MZUSP. Ressurgencia das Areias (= Agua Quente Cave). 29.X.1997. E. H. Santos coll. 2 , une preparation. Forme \" type \"."]}

International audience; Organisé par J.-J. Geoffroy, J.-P. Mauriès et M. Nguyen Duy-Jacquemin, cette réunion s'est tenue à Paris du 25 au 31 juillet 1993, au Muséum national d'histoire naturelle et à l'université Pierre-et- Marie-Curie (Paris-VI), qui ont apporté leur soutien à l'organisation de la manifestation. Elle a rassemblé 85 participants et reçu les contributions de 129 chercheurs du monde entier ; 37 nations ont en effet été représentées. Il s'agit de la plus forte participation enregistrée lors d'un congrès international de myriapodologie depuis la création du C I M (Centre international de myriapodologie) à Paris en 1968. La coopération internationale, soutenue par le ministère des Affaires étrangères et par celui de la Recherche, y a tenu une grande place.