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Two types of secondary conidia and their formation are described from six species of Strongwellsea infecting hosts from Anthomyiidae, Muscidae and Fanniidae. We used a simple device allowing secondary conidia to be produced under very moist or comparatively dry conditions. Ellipsoid type secondary conidia, which are formed under very moist conditions, have never been reported before from the genus Strongwellsea, and they are unique for Entomophthorales; these are broadly ellipsoidal with a clearly pointed basal papilla and are actively discharged. Subglobose type secondary conidia are, for the first time, described from several species in the genus Strongwellsea; they are subglobose to almost bell-shaped with a flattened papilla and are actively discharged. Subglobose type secondary conidia are formed under more dry conditions. A general pattern of the formation of secondary conidia in Strongwellsea and the ecological roles of primary conidia and of the two types of secondary conidia are discussed.

A decaying cadaver is particularly attractive to necrophagous insects, more specifically Diptera. These are the first to lay their eggs on corpses. For the post mortem interval determination, the entomologist needs to know the precise time of the first egg-laying. The objective of this study is to determine the egg laying delay of these insects on a cadaver exposed in different conditions in the Guinean zone of Cote d'Ivoire. To do this, our work was been carried out in a natural environment at the National Agronomic Research Center. The experimental setup consisted of four types of wire mesh cages corresponding to the following cases: cadavers exposed to the open air or control cadavers, semi-immersed cadavers, cadavers wrapped in a shroud and suspended cadavers. Work on the site has been made from 29 October to 5 November 2019. The spawning period in species of Calliphoridae, was shorter on control and suspended cadavers. In Sarcophagidae, we noted a larviposition late on the suspended cadavers. Muscidae and Fanniidae, which intervened later on the decomposing corpses, were not observed on the suspended cadavers. These carcasses quickly dried out, no longer being able to provide nutrients essential for the proper development of larvae of species of these Diptera families. Depending on the accessibility of the corpses to necrophagous insects, the first egg-laying of Diptera were observed after 6 hours of exposure and the last after 174 hours. The exposure conditions of the corpses significantly influenced the time taken to lay the main necrophagous Diptera. As the first egg-laying of the flies occurs in the first moments after death, as long as the corpses is accessible, the results obtained during these experiments should be taken into account by the expert entomologist, in the estimation of the interval post-mortem upon discovery of a corpse.

Fannia barbata (STEIN, 1892) NÖ, Hainburg, Spitzerberg, 30.4.01 und 9.4.17. Neu für Niederösterreich., Published as part of Schlüsslmayr, Gerhard, 2018, Erstnachweise von Fliegen (Diptera) für Österreich und einige Bundesländer, pp. 765-800 in Linzer biologische Beiträge 50 (1) on page 781, DOI: 10.5281/zenodo.4004374

Background In the current study, 23 species within 18 genera and 13 families of order Diptera were taxonomically studied. These members were collected from, on, in, around and beneath the carcasses of rabbits and guinea pigs. Results The families which collected from the carcasses are as follows: Agromyzidae (one subfamily, one genus and only one species), Calliphoridae (three subfamilies, three genera and sex species), Dolichopodidae (one subfamily, one genus and only one species), Drosophilidae (one subfamily, one genus and only one species), Ephydridae (one subfamily, one genus and only one species), Fanniidae (one genus and only one species), Muscidae (two subfamilies, three genus and four species), Phoridae (one subfamily, one genus and only one species), Piophilidae (one subfamily, one genus and only one species), Psychodidae (one subfamily, one genus and only one species), Sarcophagidae (two subfamilies, two genera and three species), Sphaeroceridae (one subfamily, one genus and only one species), and finally Ulidiidae (one subfamily, one genus and only one species). Only seven of which (Calliphoridae, Fanniidae, Muscidae, Phoridae, Piophilidae, Sarcophagidae and Ulidiidae) are of forensic importance or necrophagous. Other families are the following: Drosophilidae, Psychodidae, Agromyzidae, Dolichopodidae, Ephydridae, and Sphaeroceridae are of minor importance as forensic indicators. Conclusions Taxonomic keys for the adults which collected from the carcasses were constructed. Also diagnosis, synonyms, photographs and the forensic importance for the taxa were provided.

Resumo. No semiárido, poucos são os estudos que abordam a dipterofauna necrófaga. Assim, o presente trabalho teve por objetivo realizar um levantamento da fauna de dípteros necrófagos e avaliar sua distribuição espaço-temporal em diferentes localidades dos municípios de Serra Talhada/PE e Triunfo/PE. Foram realizadas coletas, ao longo do período de setembro de 2013 (estação seca) e abril de 2014 (estação chuvosa), com auxilio de armadilhas aéreas contendo três tipos de iscas diferentes (peixe, frango e carne bovina) em quatro pontos de coletas, três em Serra Talhada (área rural, área urbana e lixão) e um em Triunfo (área rural). A partir de 1.215 indivíduos coletados foram identificadas 12 espécies distribuídas entre as famílias Calliphoridae, Muscidae, Fanniidae, Phoridae, Piophilidae, Stratiomyidae, Sarcophagidae, Tephritidae e Ulidiidae.Necrophagous dipterofauna diversity associated to different types of organic matter decomposed in two vegetation types of the semi-arid PernambucoAbstract. In the semi-arid region, there are few studies that address the necrophagous dipterofauna. Then, this study aimed to carry out a survey the fauna of necrophagous insects of Diptera order and assess its spatiotemporal distribution in different locations in the municipalities of Serra Talhada/PE and Triunfo PE. The insects have been collected during the September 2013 (dry season) and April 2014 (rainy season), using air traps with three different types of bait (fish, chicken and beef) in four collection points, three in Serra Talhada (rural area, urban area and dump city) and Triunfo (rural area). From 1,215 individuals collected were identified 12 species distributed among the families Calliphoridae, Muscidae, Fanniidae, Phoridae, Piophilidae, Stratiomyidae, Sarcophagidae, Tephritidae and Ulidiidae.

Insects associated with carrion can have parasitological importance as vectors of several pathogens and causal agents of myiasis to men and to domestic and wild animals. We tested the attractiveness of animal baits (chicken liver) at different stages of decomposition to necrophagous species of Diptera (Calliphoridae, Fanniidae, Muscidae, Phoridae and Sarcophagidae) in a rainforest fragment in Brazil. Five types of bait were used: fresh and decomposed at room temperature (26 °C) for 24, 48, 72 and 96 h. A positive correlation was detected between the time of decomposition and the abundance of Calliphoridae and Muscidae, whilst the abundance of adults of Phoridae decreased with the time of decomposition. Ten species of calliphorids were registered, of which Chrysomya albiceps, Chrysomya megacephala and Chloroprocta idioidea showed a positive significant correlation between abundance and decomposition. Specimens of Sarcophagidae and Fanniidae did not discriminate between fresh and highly decomposed baits. A strong female bias was registered for all species of Calliphoridae irrespective of the type of bait. The results reinforce the feasibility of using animal tissues as attractants to a wide diversity of dipterans of medical, parasitological and forensic importance in short-term surveys, especially using baits at intermediate stages of decomposition.

Necrodes littoralis (Linnaeus, 1758) (Coleoptera: Silphidae), also known as the "shore sexton beetle," is a common silphid beetle that visits and breeds on large vertebrate cadavers. This study describes, for the first time, the involvement of N. littoralis on human corpses based on a large dataset of 154 French forensic cases. Various parameters regarding corpse location, decomposition stages, and entomofauna were extracted from each file. Compared to all of the forensic entomology cases analyzed between 1990 and 2013 (1028), N. littoralis was observed, on average, in one case out of eight; most of these cases occurred during spring and summer (73.5%). More than 90% of the cases were located outdoors, especially in woodlands, bushes, and fields. The decomposition stage of the corpse varied among cases, with more than 50% in the advanced decomposition stage, 36% in the early decomposition stage, and less than 10% in the fresh, mummified, or skeletonized stages. Regarding other necrophagous species sampled with N. littoralis, Calliphorid flies were found in 94% of the cases and Fanniidae/Muscidae in 65% of the cases. Chrysomya albiceps, a heliophilic species mostly located in the Mediterranean area, was present in 34% of the cases (only 20% in the whole dataset). The most common coleopteran species were Necrobia spp. (Coleoptera: Cleridae) and Creophilus maxillosus (Coleoptera: Staphylinidae); these beetles were observed in 27% of the cases. The over-representation of these species is likely due to similar requirements regarding the climate and decomposition stage. As N. littoralis is frequently observed and tends to become more common, we conclude that the developmental data for this species would be a precious tool for forensic entomologists in Europe.

The distribution and habitat preferences of necrophagous Diptera in northeastern Brazil is poorly known despite the medical and forensic relevance of species in the Families Calliphoridae, Sarcophagidae, and Muscidae. We performed a survey on the diversity of necrophagous species in 4 types of environments: rainforest, agroecosystem, beach, and urban areas. Adult flies were collected by using suspended traps containing decomposing animal tissue (chicken liver, sardine, or pork) as baits. A diverse assemblage of necrophagous Diptera was registered in all environments, consisting of 20 species from 7 families: Calliphoridae, Fanniidae, Muscidae, Phoridae, Piophilidae, Sarcophagidae, and Ulidiidae. Megaselia scalaris (Loew) (Phoridae), Chrysomya albiceps (Wiedemann) (Calliphoridae), and Tricharaea sp. (Muscidae) were the most abundant species. The rainforest fragment and the sugarcane plantation were the environments with the highest degree of species similarity. The type of bait did not significantly influence the number of species captured. The invasive species Chrysomya megacephala (Fabricius) and C. albiceps were present in high abundance in all environments, especially at the sandy beach, where they corresponded to 100% of all Calliphoridae specimens.

"Canyon flies" are flies within the Fannia benjamini Malloch species complex. These flies can be considerable pests to humans and animals due to their habit of feeding on animal body secretions such as tears, mucus, sweat, and blood. Adult "canyon flies" (Fannia conspicua Malloch) were collected from the coastal mountain community of La Habra Heights in Los Angeles County, CA, during late spring and early summer of 2011. Canyon flies were colonized in the laboratory and maintained for over two years. Larval growth, sex-specific adult emergence, and fecundity were evaluated quantitatively. At 25°C, male flies required a minimum of 22 d postoviposition to develop from egg to adult, with peak emergence occurring at 24-26 d; female flies required a minimum of 25 d postoviposition, with peak emergence occurring at 26-28 d. Female flies were capable of oviposition starting at 7 d postemergence, and produced a mean of 90.6±54.7 eggs over a 19-41-d life span. Canyon fly eggs were quite resistant to desiccation, with hatching rate of eggs reduced only after ≥12 wk of desiccation, and some successful hatching even when eggs were desiccated up to 28 wk. When immature flies were removed from their food source and subjected to continuous desiccation and starvation, flies at 3-d-old posthatching did not survive, and only 8% of flies at 5-d posthatching survived and completed development to the adult stage. Immature flies that were 7- or 11-d-old posthatching survived to adulthood in good numbers, having reached the mid to late L3 stage at the time of food removal. Exposure to desiccation in either the egg or larval stage had no effect on fecundity of surviving females. This extreme level of desiccation tolerance is likely an adaptation to increase survival in desert climates of the southwestern United States with long, hot summers and little precipitation, where Fannia conspicua are typically distributed.

Os dípteros necrófagos utilizam matéria orgânica em decomposição como fonte proteica em ao menos uma fase de sua vida. Deste modo o conhecimento taxonômico dessa fauna se torna importante para estudos forense, sanitário e médico. Os dípteros das famílias Calliphoridae, Muscidae e Fanniidae possuem espécies necrófagas associadas ao ambiente humano, estando entre os insetos de maior destaque nesse aspecto e até então nunca foram estudados na cidade de Feira de Santana, Bahia. O objetivo deste estudo foi realizar um levantamento da fauna de dípteros necrófagos das famílias Calliphoridae, Muscidae e Fanniidae de Feira de Santana, Bahia. Para isso, foram realizadas coletas mensais ao longo de um ano em três áreas da cidade com um total de cinco armadilhas iscadas (com 24 horas de exposição) por área, resultando em um subconjunto de 60 armadilhas/área e um total de 180 amostras. Foram coletados 4.818 dípteros, pertencentes a 20 espécies das três famílias estudadas: Calliphoridae (seis espécies), Fanniidae (três espécies) e Muscidae (11 espécies). A maior abundância das famílias ocorreu no outono e a menor no verão. Observou-se que treze das vinte espécies coletadas, possuem interesse forense e sanitário descritos na literatura. Taxonomic Survey and Seasonality of Calliphoridae, Muscidae and Fanniidae (Insecta: Diptera) in Feira de Santana, Bahia, Brazil Abstract. Taxonomic studies in necrphagous Diptera is relevant because of its importance for forensic, health and medical purposes, since they feed in decaying organic matter as protein source, at least in one stage of their lives. Synantropic Diptera, such us some species of Calliphoridae, Muscidae and Fannidae had not been studied in Feira de Santana, BA. The aim of this study was to do an inventory of necrophagous Diptera belonging to the families cited above. Five baited traps (24 hours of exposure) were placed in three sites in the town, once a month, resulting in a subset of 60 traps per site and a total of 180 samples. A total of 4.818 specimens were collected, belonging to 20 species: Calliphoridae (six species), Fannidae (three species) and Muscidae (11 species). The abundance of synantropic Diptera showed seasonality being the fall the season with higher abundance while in the summer the abundance was significant lower. Thirteen out of the twenty species collected were cited in the literature as being of forensic importance.

The harsh conditions of the Brazilian seasonally dry tropical forest known as Caatinga pose challenges to the insects specialized in the exploitation of ephemeral resources. We investigated the diversity and daily flight activity of dipterans associated with decomposing rat carcasses in a field experiment performed in the semi-arid region of Pernambuco State, Brazil. We also analyzed the temporal arrival of adult insects on the carcasses at three stages of decomposition: early, intermediate, and advanced. We collected 1173 individuals, of which Muscidae had the highest abundance (36.5%), followed by Sarcophagidae (28.1%), Calliphoridae (25.2%), and Fanniidae (10.2%). Chrysomya albiceps (Wiedemann, 1819) (Calliphoridae), Fannia pusio (Wiedemann, 1830) (Fanniidae), Atherigona orientalis (Schiner, 1868), and Musca domestica (Linnaeus, 1758) (Muscidae) were the most abundant species. The richness reached its maximum value on the second day of decomposition, with 18 species, decreasing to 8 species on the last day of decomposition (7 days). The ecological indices of diversity, dominance, and evenness varied little among the stages. There was an overlap of most species throughout the decomposition, although the overall abundance was higher at the intermediate stage for Calliphoridae, Muscidae, and Sarcophagidae. In accordance to previous studies, nocturnal flight was rare, as approximately 8% of insects were captured at night. Our results expand the knowledge on ecological and behavioral aspects of necrophagous flies under inhospitable environments, such as the dry season in the Caatinga. The dominance of the invasive species C. albiceps reinforced here illustrates its geographical expansion towards the countryside of Northeastern Brazil.

A historical review of the Fannia carbonaria-group is provided and three new species are described from China: F. fani Wang & Wu, sp. n., F. nitidiventris Wang & Zhang, sp. n. and F. submaculata Wang & Zhao, sp. n.. One species, Fannia norvegica Ringdahl, 1934, is recorded for the first time from China. Illustrations of male terminalia of these four species and a taxonomic key to the males of known species in the group are given. The F. carbonaria-group now includes 30 species distributed in the Holarctic Region and northern part of the Oriental Region.

Pseudoscorpions employ phoresy for dispersal, in which one animal attaches to another for transportation. This study documents the occurrence and phoresy by Americhernes aff. incertus (Pseudoscorpiones: Chernetidae) on the fly Fannia canicularis (Diptera: Fanniidae) in a fragment of semi-deciduous seasonal Atlantic Forest, in the state of Pernambuco, northeast Brazil. The pseudoscorpions were found attached to the ventral surface of six individual flies. To our knowledge, no previous studies have shown the presence of Americhernes aff. incertus in the northeast Atlantic Forest and its use of flies as a dispersal method.

muisca Grisales, Wolff & de Carvalho, 2012 a: 821. Type locality: Colombia, Cundinamarca, Mosquera, 2543 m. HT M (ICN), PT M (ICN, CEUA). Distr.: Colombia (Cundinamarca (Mosquera; Bogot��; Tolemaida), Boyac�� (Tunja))., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 a) Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia. Journal of Natural History, 46, 803 - 829."]}

uwa Grisales, Wolff & de Carvalho, 2012 a: 826. Type locality: Colombia, Santander, Piedecuesta, 2290-2390 m. HT M (CEUA). Distr.: Colombia (Santander (Piedecuesta))., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 a) Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia. Journal of Natural History, 46, 803 - 829."]}

tahami Grisales, Wolff & de Carvalho, 2012 a: 824. Type locality: Colombia, Antioquia, Medellin, 1550 m. HT M (CEUA), PT M (CEUA). Distr.: Colombia (Antioquia (Medellin, La Pintada)), Costa Rica., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 a) Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia. Journal of Natural History, 46, 803 - 829."]}

pusio Wiedemann, 1830: 437 (Anthomyia). Type locality: ���South America ���. LT M (ZMUC) (des. Pont 1977: 54), PLT (ZMUC). Distr. Colombia (Antioquia (Medellin, La Pintada), Caquet�� (Florencia), C��rdoba (Monteria), Magdalena (Santa Marta), Risaralda (Pereira), Santander (Bucaramanga)), Mexico, Guatemala, Panama, El Salvador, St. Vincent Island, Guadalupe Island, Puerto Rico, Cuba, Bahamas, Trinidad, Venezuela, Guyana, Brazil, Chile, Galapagos Islands, Easter Island; North America, Pacific, West Africa, Mediterranean. Refs.: S��guy, 1937: 113 (cat.); Pont, 1972: 6 (cat.); de Carvalho et al., 1993: 14 (cat.); de Carvalho et al., 2003: 17 (cat.); Grisales et al., 2012 b: 37 ���38 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 810, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Wiedemann, C. R. W. (1830) Aussereuropaische zweiflugelige Insekten. Part II. in der Schulzischen Buchhandlung, Hamm. 684 pp.","Pont, A. C. (1977) A revision of Australian Fanniidae (Diptera: Calyptrata). Australian Journal of Zoology, 51 (Supplement), 1 - 60. http: // dx. doi. org / 10.1071 / AJZS 051","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

abnormis Stein, 1900: 210 (Homalomyia). Type locality: Bolivia, S. Antonio and Songo. ST F destroyed (formerly HNHM). Distr.: Colombia (not specific locality), Venezuela, Peru, Bolivia. Refs.: Stein, 1918: 234 (distr.); Stein, 1919: 131 (cat.); S��guy, 1937: 164 (cat.); Pont, 1972: 3 (cat.); de Carvalho et al., 1993: 6 (cat.); de Carvalho et al., 2003: 7 (cat.); Grisales et al., 2012 b: 6 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1900) Vier neue aus Bolivia stammende Homalomyia-Arten des Ungarischen National Museums. Termeszettudomanyi Fuzetek, 23, 205 - 212.","Stein, P. (1918) Zur weitern Kenntnis aussereuropaischer Anthomyiden. Annales Historico-Naturales Musei Nationalis Hungarici, 16, 147 - 244.","Stein, P. (1919) Die Anthomyidengattungen der Welt, analytisch bearbeitet, nebst einem kritischsystematischen Verzeichnis aller aussereuropaischen Arten. Archiv fur Naturgeschichte, 83 [1917], 85 - 178.","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

flavicincta Stein, 1904: 453 (Homalomyia). Type locality: Peru, Vilcanota and Colombia, Cordillera. ST M (not found in ZMHB) ST M (destroyed in HNHM). Distr.: Colombia, Peru, Bolivia, Brazil. Refs.: S��guy, 1937: 167 (cat.); Pont, 1972: 4 (cat.); de Carvalho et al., 1993: 9 (cat.); de Carvalho et al., 2003: 11 (cat.); Grisales et al., 2012 b: 25 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1904) Die Amerikanischen Anthomyiden des Koniglichen Museums fur Naturkunde zu Berlin und des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 2, 414 - 495.","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

guane Grisales, Wolff & de Carvalho, 2012 a: 819. Type locality: Colombia, Santander, Piedecuesta, 2290-2390 m. HT M (CEUA). Distr.: Colombia (Santander (Piedecuesta)).

Grisales, Diana, De Carvalho, Claudio J. B. (2016): FAMILY FANNIIDAE. Zootaxa 4122 (1): 807-813, DOI: http://dx.doi.org/10.11646/zootaxa.4122.1.69

chitarera Grisales, Wolff & de Carvalho, 2012 a: 814. Type locality: Colombia, Norte de Santander, Pamplona, 2360 m. HT M (CEUA), PT M (CEUA). Distr.: Colombia (Antioquia (San Pedro de l��s Milagros), Cundinamarca (Facatativa), Norte de Santander (Pamplona))., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 808, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 a) Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia. Journal of Natural History, 46, 803 - 829."]}

trimaculata Stein, 1898: 176 (Homalomyia). Type locality: North America and Jamaica. ST M (ZMHB, MCZ, FMNH, USNM). Distr.: Colombia (Antioquia (Medellin)), Belize, Panama, Puerto Rico, Jamaica, Dominican Republic, Haiti, Santo Domingo, Ecuador, Peru, Venezuela, Brazil, Uruguay, Argentina. Refs.: de Carvalho et al., 1993: 16 (cat.); de Carvalho et al., 2003: 19 (cat.); Pont & Werner, 2006 (remarks of ST): 15; Grisales et al., 2012 b: 43 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 811, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1898) Nordamerikanische Anthomyiden. Beitrag zur Dipterenfauna der Vereinigten Staaten. Berliner entomologische Zeitschrift, 42 (1897), 161 - 288.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Pont, A. C. & Werner, D. (2006) The types of Fanniidae and Muscidae (Diptera) in the Museum fur Naturkunde, Humboldt- Universitat zu Berlin, Germany. Zoosystematics and Evolution, 82, 3 - 139. http: // dx. doi. org / 10.1002 / mmnz. 200600001","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

aburrae Grisales, Wolff & de Carvalho, 2012 a: 808. Type locality: Colombia, Antioquia, Copacabana, 1650 m. HT M (CEUA). Distr.: Colombia (Antioquia (Copacabana))., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 808, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 a) Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia. Journal of Natural History, 46, 803 - 829."]}

Genus Euryomma Stein Euryomma Stein, 1899: 19. Type species, Euryomma hispaniense Stein, 1899 = Euryomma peregrinum (Meigen, 1826). Refs.: de Carvalho et al., 2003: 5 (cat.)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 808, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1899) Euryomma, eine neue Gattung der Anthomyidengruppe Homalomyia. Entomologisches Nachrichtenblatt, 25, 19 - 22.","Meigen, J. W. (1826) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Tome 5. In Verlag der Schultz-Wundermann'schen Buchhandlung, Hamm 5, 412 pp.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32."]}

dodgei Seago, 1954: 4. Type locality: Panama, David. HT M (AMNH), PT (AMNH). Distr.: Colombia (Antioquia (Medellin), Magdalena (Santa Marta)), Panama, Cuba, Brazil. Refs.; Pont, 1972: 3 (cat.); de Carvalho et al., 1993: 9 (cat.); Carvalho et al., 2003: 11 (cat.); Grisales et al., 2012 b: 11 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 809, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Seago, J. M. (1954) The pusio Group of the genus Fannia Robineau - Desvoidy, with descriptions of the new species. American Museum Novittates, 1699, 1 - 14.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

obscurinervis Stein, 1900: 207 (Homalomyia). Type locality: Bolivia, Songo. ST M, F (destroyed HNHM). Distr.: Colombia (Amazonas (Leticia)), Mexico, Venezuela, Guyana Peru, Bolivia, Brazil, Paraguay. Refs.: Stein, 1900: 207; Stein, 1904: 458; Stein 1918: 235; S��guy, 1937: 172 (cat.); Pont, 1972: 5 (cat.); de Carvalho et al., 1993: 12 (cat.); de Carvalho et al., 2003: 14 (cat.); Couri & Winagraski, 2005: 645; Garcia-Amat, 2010: 399 (remarks); Grisales et al., 2012 b: 33 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 810, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1900) Vier neue aus Bolivia stammende Homalomyia-Arten des Ungarischen National Museums. Termeszettudomanyi Fuzetek, 23, 205 - 212.","Stein, P. (1904) Die Amerikanischen Anthomyiden des Koniglichen Museums fur Naturkunde zu Berlin und des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 2, 414 - 495.","Stein, P. (1918) Zur weitern Kenntnis aussereuropaischer Anthomyiden. Annales Historico-Naturales Musei Nationalis Hungarici, 16, 147 - 244.","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Couri, M. S. & Winagraski, E. (2005) New Fannia Robineau - Desvoidy from Amazonas, Brazil and new geographical record (Diptera, Fanniidae). Revista Brasileira de Zoologia, 22, 645 - 647. http: // dx. doi. org / 10.1590 / S 0101 - 81752005000300018","Garcia-Amat, E. C. A. (2010) Notes on necrophagous fies (Diptera: Calyptratae) associated to fish carrion in Colombian Amazon. Acta Amazonica, 40, 397 - 400. http: // dx. doi. org / 10.1590 / S 0044 - 59672010000200018","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

hirtifemur Stein, 1904: 457 (Homalomyia). Type locality: Colombia, Cordillera. ST M, F (ZMHB). Distr.: Colombia, Ecuador, Peru, Brazil, Argentina. Refs.: S��guy, 1937: 169 (cat.); Pont, 1972: 5 (cat.); de Carvalho et al., 1993: 11 (cat.); de Carvalho et al., 2003: 13 (cat.); Grisales et al., 2012 b: 27 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 810, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1904) Die Amerikanischen Anthomyiden des Koniglichen Museums fur Naturkunde zu Berlin und des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 2, 414 - 495.","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

penicillaris Stein, 1900: 205 (Homalomyia). Type locality: Bolivia, Songo. ST M, F (ZMHB) or destroyed (formerly HNHM). Distr.: Colombia, Peru, Bolivia, Brazil. Refs.: S��guy, 1937: 172 (cat.); Pont, 1972: 5 (cat.); de Carvalho et al., 1993: 13 (cat.); de Carvalho et al., 2003: 16 (cat.); Grisales et al., 2012: 34 (remarks)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 810, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Stein, P. (1900) Vier neue aus Bolivia stammende Homalomyia-Arten des Ungarischen National Museums. Termeszettudomanyi Fuzetek, 23, 205 - 212.","Seguy, E. (1937) Diptera Fam. Muscidae. In: Wytsman, P. (Ed.), Genera Insectorum, 205, pp. 1 - 604.","Pont, A. C. (1972) Family Muscidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Vol. 9 7. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, pp. 1 - 111.","de Carvalho, C. J. B., Pont, A. C., Couri, M. S. & Pamplona, D. (1993) Parte I. Fanniidae. In: de Carvalho, C. J. B. (Ed.), A Catalogue of the Fanniidae and Muscidae of the Neotropical region. Sociedade Brasileira de Entomologia, Sao Paulo, pp. 1 - 29.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32."]}

grandis Malloch, 1912: 3. Type-locality: Panama, Puerto Bello. HT M (USNM), PT M (USNM). Distr.: Colombia (Antioquia (San Roque)). Refs.: de Carvalho et al., 2003: 12 (cat.); Grisales et al., 2012 b: 25 (redescription)., Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2016, FAMILY FANNIIDAE, pp. 807-813 in Zootaxa 4122 (1) on page 810, DOI: 10.11646/zootaxa.4122.1.69, http://zenodo.org/record/267354, {"references":["Malloch, J. R. (1912) New Diptera from Panama. Smithsonian Miscellaneous Collections, 59, 1 - 8.","de Carvalho, C. J. B., Pont A. C., Couri, M. S. & Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32.","Grisales, D., Wolff, M. & de Carvalho, C. J. B. (2012 b) Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia. Zootaxa, 3591, 1 - 46."]}

Fly larvae living on dead corpses can be used to estimate post-mortem intervals. The identification of these flies is decisive in forensic casework and can be facilitated by using DNA barcodes provided that a representative and comprehensive reference library of DNA barcodes is available.We constructed a local (Belgium and France) reference library of 85 sequences of the COI DNA barcode fragment (mitochondrial cytochrome c oxidase subunit I gene), from 16 fly species of forensic interest (Calliphoridae, Muscidae, Fanniidae). This library was then used to evaluate the ability of two public libraries (GenBank and the Barcode of Life Data Systems – BOLD) to identify specimens from Belgian and French forensic cases. The public libraries indeed allow a correct identification of most specimens. Yet, some of the identifications remain ambiguous and some forensically important fly species are not, or insufficiently, represented in the reference libraries. Several search options offered by GenBank and BOLD can be used to further improve the identifications obtained from both libraries using DNA barcodes.

We present a forensic case associated with skeletonized human remains found inside a cistern in a coastal town located in the eastern Iberian Peninsula (Valencian Regional Government, Spain). In order to analyse the particular environmental conditions that occurred during oviposition and development of the collected insects, estimated temperatures at the crime scene were calculated by a predictive mathematical model. This model analyses the correlation between the variability of the internal temperature depending on the variability of the external ones. The amplitude of the temperature oscillations inside the tank and the containment of the enclosure is reduced by the presence of water. Such variation occurred within about 2 h due to the time required for heat exchange. The differential equations employed to model differences between outdoor and indoor temperatures were an essential tool which let us estimate the post-mortem interval (PMI) that was carried out by the study of the insect succession and the development time of the collected Diptera specimens under the adjusted temperatures. The presence of live larvae and pupae of Sarcophagidae and empty pupae of Calliphoridae, Sarcophagidae, Fanniidae, Muscidae, Phoridae and Piophilidae and the decomposition stage suggested the possibility that the remains were in the tank at least a year. We highlight the absence of Calliphora and Lucilia spp., and the first occurrence of the phorid Conicera similis in a human cadaver among the entomological evidence. R.L. and J.M.G.V. are thankful for support from MEC of Spain [Grant No. CTQ 2010-19232]. This study was supported by a grant for studies of Phoridae from the Balfour-Browne Trust Fund (University of Cambridge) and Calliphoridae and Sarcophagidae studies by Generalitat Valenciana [GV/2011/039] and Alicante University projects [GRE09-27].

The alimentary canal of Fannia canicularis was studied intensively for the first time, highlighting the description of morphological characteristics and providing detailed morphometric data. Two pairs of Malpighian tubules with considerably different length were found which has been rarely documented for Calyptratae flies before. Furthermore, stout cardia and sinewy crop of the samples were ob- served and supposed to be responsible for the energy needed to support the long- time swarming behavior of the species.

An overview is given of the species-group names in the families Fanniidae and Muscidae described by Paul Stein (1852–1921) together with their type specimens. Stein described 57 new species in the Fanniidae and 783 species in the Muscidae together with 5 replacement names, to which are added 6 unavailable species names (nomina nuda) in Fanniidae and 26 in Muscidae. A brief account of his life and career and of the various collections that he studied is given, followed by an alphabetical list of the species-group names within the Fanniidae and Muscidae. The type specimens (holotypes and syntypes) that have been located in museums and institutes throughout Europe and the United States are enumerated. The bibliography includes all the papers published by Stein on these families. The account concludes with three appendices: a systematic list of the species, a list of the species by zoogeographic region, and a list of the localities mentioned in Stein's papers. Four lectotypes are designated (Homalomyia carbonella Stein, 1895; Homalomyia lineata Stein, 1895; Fannia nigra Stein, 1920; Mydaea pallidicornis Stein in Becker, 1910), and two replacement names for junior homonyms are given (Neodexiopsis simplicissima nom. nov. for Coenosia simplex Stein, 1920, and Helina steini nom. nov. for Aricia punctata Stein, 1898, both from the Nearctic Region). (© 2013 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim)

Euryomma is the second largest genus within Fanniidae, but only two species, E. panamensis Chillcott, 1958 and E. rettenmeyeri Chillcott, 1958 have been recorded in Central America. In this paper we describe two new species of the genus found in Costa Rica, Euryomma chillcotti sp. nov. (Province of Puntarenas) and Euryomma steini sp. nov. (Provinces of Cartago and Limón). Euryomma tahami Grisales, Wolff & Carvalho, 2012 is recorded for the first time to Costa Rica. Identification key to all species found in Central America, descriptions and re-descriptions of species occurring in the region and illustrations of the terminalia of new species are presented. RESUMO: Euryomma é o segundo maior gênero dentro de Fanniidae, mas apenas duas espécies são conhecidas para América Central, E. panamensis Chillcott, 1958 e E. rettenmeyeri Chillcott, 1958. Nesta contribuição são descritas duas espécies novas do gênero encontradas na Costa Rica, Euryomma chillcotti sp. nov. (Província de Puntarenas) e Euryomma steini sp. nov. (Províncias de Cartago e Limón). Euryomma tahami Grisales, Wolff & Carvalho, 2012 é registrada pela primeira vez na Costa Rica. É apresentada uma chave de identificação para as espécies da América Central, descrições e redescrições das espécies que ocorrem na região, assim como ilustrações da terminália das espécies novas. COL0066697

A total of 17 species of the Fannia postica-group Chillcott, 1961 from the Palearctic and Oriental regions are reviewed herein, 2 of which are described from China as new: Fannia ningxiaensis Wang & Zhang, sp. n. and Fannia subaethiops Wang & Zhu, sp. n.. Fannia labidocerca Feng & Xue, 2006, originally placed in F. serena-group Chillcott, 1961, is moved to the postica-group and re-described. An identification key to the males of known species from these regions is provided.

A historical review of the Fannia carbonaria-group is provided and three new species are described from China: Fannia fani Wang & Wu, sp. n., Fannia nitidiventris Wang & Zhang, sp. n. and Fannia submaculata Wang & Zhao, sp. n.. One species, Fannia norvegica Ringdahl, 1934, is recorded for the first time from China. Illustrations of male terminalia of these four species and a taxonomic key to the males of known species in the group are given. The Fannia carbonaria-group now includes 30 species distributed in the Holarctic Region and northern part of the Oriental Region.

Fannia papei sp. nov. (Figs. 6���9) Description. MALE. Body length 3.75���4.10 mm. Eye with short and distant hairs; facets slightly enlarged on anterior margin in upper part; fronto-orbital plate and parafacial with dense grayish-silvery pollinosity, frons almost as wide as the distance between two posterior ocelli at narrowest point; frontal vitta black, with brownishyellow pollinosity, the narrowest point narrower than the width of anterior ocellus; frontal setae eight or night, stout, nearly reaching ocellar triangle, orbital setae absent; postocular setae in one row, hair-like, occipital setae behind the postocular setae on vertex in one shorter row; parafacial bare, at middle about 1/3���2/3 as wide as the width of postpedicel; antenna black, postpedicel 1.70x to 2.00x as long as wide, arista black and short plumose, slightly swollen in basal part, the longest individual hair shorter than basal aristal width, epistoma not projecting beyond vibrissal angle, vibrissal angle beyond frontal angle in profile; subvibrissal setae in one row, lateral with one short row of setae; gena and genal dilation with fine black setae; gena with grayish-brown pollinosity, upper margin of gena without upcurved seta; proboscis short, prementum shining, its length 2.00x as long as its width; palpus black, claviform, as long as or slightly shorter than the length of prementum. Thorax ground-color black, notum with grayish-brown pollinosity, without distinct vitta; presutural acr triserial, hair-like, only prescutellar pairs slightly stout, the distance between two outer acr rows narrower than the distance between acr and dc rows, dc 2+2(3), ia 0+2, pra 1, about 1/2���2/3 as long as posterior notopleural seta; scutellum with widely spread fine setae in the dorsal except median parts; notopleuron without seta; basisternum, proepisternum, anepimeron, meron and katepimeron bare; katepisternal setae 1+1, katepisternum without ventral spine; anterior spiracle brownish, posterior one brown; calypters brown with yellow margin, the lower one distinctly projecting beyond the upper one. Wing dark brown; veins dark brown; tegula black; basicosta dark brown; costal spine inconspicuous; node of Rs bare on ventral and dorsal surfaces; vein M straight, slightly parallel to vein R4+5 distally; vein r-m straight, crossveins without obvious cloud; haltere brown. Legs black except fore legs brown in basal part; fore femur with complete and stout pv and pd rows, with several upcurved p rows, fore tibia with one apical d; mid coxa without any hook-like spine or spine-like seta, mid femur with long and sparse av row in basal half, becoming gradually shorter towards apex, bare in subapical part, three or four short setae in apical part, pv row complete, slightly biserial in middle part, p row slender, mid tibia slightly swollen in distal 1/3, with two or three ad and two pd, and with numerous slender setae on ventral surface, mid first tarsomere without basal tooth-like spine on ventral surface; hind coxa with one long seta on posterior surface, hind femur with one long av row and one short pv row, in distal 1/3 from the posteroventral to posterior surface bare, hind tibia with two av, one ad and two d. Abdomen long, depressed and flattened, with dense grayish-white pollinosity; syntergite 1+2 to tergite 4 each with one median inverted triangular vitta, tergite 5 with one narrow median stripe, each tergite with long marginal setae; sternite 1 with setae, sternite 5 profoundly indented on posterior margin and covered with setae (Fig. 8C); cercus elongate, concave on the median line, from lateral view slightly curved towards ventral (Figs. 8A, B), surstylus becoming slightly broader towards apex (Fig. 8B), bacilliform process absent. FEMALE. Body length 3.70���4.00 mm. Fronto-orbital plate and parafacial with dense brownish-gray pollinosity; frons at middle about 2/5 as wide as the distance between the width of head; frontal vitta at narrowest point about 6.00x as wide as the width of anterior ocellus; frontal setae five or six, upper orbital setae two. Thorax presutural acr biserial; calypters brownish-yellow. Wing basicosta brownish-yellow; costal spine conspicuous. Legs entirely black; mid femur with complete and hair-like av row, with short pv in basal half, mid tibia with one ad, two or three pd; hind femur with one av row in basal 1/3 (with three or four stout), pv inconspicuous, hind tibia with one av. Abdomen with thin gray pollinosity, without obviously mark; sternite 1 bare; terminalia see fig. 9. Other morphological characteristics are similar to those of the male. Remarks. This new species resembles F. nigriclara Zhang & Xue, 1993 but differs from it by having parafacial about 1/3���2/3 as wide as the width of postpedicel; palpus as long as or slightly shorter than prementum; presutural acr triserial; calypters brown; mid tibia with two or three ad; hind coxa with one long seta on posterior surface; hind tibia with one ad; male sternite 5 slightly narrower and cercus slightly straight. Type material examined. Holotype ♂, Beijing: Yanqing, Mt. Songshan, 12.VI.2010, Coll. D. Zhang (MBFU); paratypes 4♂, 2♀, the same data as holotype (MBFU). Etymology. The new species is named after Dr. Thomas Pape in honor of his outstanding work on Calyptratae. Distribution. Paleactic: China (Beijing)., Published as part of Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu & Wang, Rong-Rong, 2016, Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species, pp. 401-414 in Zootaxa 4079 (4) on pages 410-412, DOI: 10.11646/zootaxa.4079.4.1, http://zenodo.org/record/1050841, {"references":["Zhang, L. F. & Xue, W. Q. (1993) A new species of the genus Fannia from Liaoning, China (Diptera: Fanniidae). Acta Zootaxonomica Sinica, 18 (1), 85 - 86. [In Chinese with English abstract]"]}

Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu, Wang, Rong-Rong (2016): Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species. Zootaxa 4079 (4): 401-414, DOI: http://doi.org/10.11646/zootaxa.4079.4.1

Fannia dupla Nishida, 1974 (Fig. 4) Fannia dupla Nishida, 1974: 205. Type locality: Japan, Ishikawa, Kanazawa, Mt. I��zen. Fannia dupla: Pont 1977: 448; Pont 1986: 47; Nishida 1994: 82; Xue & Wang 1998: 816; Lin & Chen 1999: 108; Wang & Xue 2002: 56; Su & Wang 2004: 111. Redescription based on the specimens collected in Beijing, China. MALE. Body length 4.00���5.00 mm. Eye bare; facets slightly enlarged on anterior margin in upper part; fronto-orbital plate and parafacial with grayishsilvery pollinosity; frons slightly narrower than the distance between two posterior ocelli at narrowest point; frontal vitta black, linear at narrowest point; frontal setae twelve to fifteen, long, nearly reaching ocellar triangle; orbital setae absent; postocular setae in one row, short and neatly arranged, without occipital seta behind the postocular setae on vertex; parafacial bare, at middle about 2/5 as wide as the width of postpedicel; antenna brown, postpedicel about 2.00x longer than wide, arista black and short plumose, slightly swollen in basal part, the longest individual hair shorter than aristal base; epistoma about at the same level of vibrissal angle, vibrissal angle behind frontal angle in profile; subvibrissal setae in one row, lateral with three or four fine setae; gena and genal dilation with fine black setae, upper margin of gena without upcurved seta; prementum with thin grayish-yellow pollinosity, its length about 2.00x as long as its width; palpus black, claviform, slightly longer than the length of prementum. Thorax ground-color black, notum with dark brown pollinosity, without distinct vitta; presutural acr triserial, hairlike, only prescutellar pairs long, dc 2+3, ia 0+2, pra 2 (fine but distinct), the anterior one about 1/3 as long as posterior notopleural seta; notopleuron without seta; basisternum, proepisternum, anepimeron, meron and katepimeron bare; katepisternal setae 1+1, katepisternum with some slightly straight setae and without ventral spine; anterior spiracle yellowish, posterior one brown; calypters brownish with dark brown margin, the lower one distinctly projecting beyond the upper one. Wing brown; veins dark brown; tegula black; basicosta dark brown; costal spine inconspicuous; node of Rs bare on ventral and dorsal surfaces; vein M straight, slightly close to vein R4+5 distally; crossveins without obvious cloud; haltere yellowish-brown. Legs black; fore coxa without anterior spine on ventral surface, fore femur with complete pv row, fore tibia without ad and median p, with two median pv; mid coxa without hook-like spine or spine-like seta, mid femur with stout av row in basal part, becoming short and comb-like setae row in distal 1/3, with complete pv row, slightly biserial in median part, mid tibia with one ad, one or two pd, the longest one shorter than mid tibial width in distal part, mid first tarsomere without basal tooth-like spine on ventral surface; hind coxa with one long seta on posterior surface, hind femur with a short av row in basal 2/3, with two stout setae in distal 1/3, without pv row, hind tibia with three to five av, four or five ad and two pd. Abdomen oval and flattened, ground-color black, with dense grayish-brown pollinosity; syntergite 1+2 to tergite 4 each with one broad median inverted triangular vitta, tergite 5 with one narrow median stripe; sternite 1 bare; terminalia see Nishida (1974: 206). FEMALE. Unknown. Material examined. CHINA: Beijing: 2♂, Yanqing, Mt. Songshan, 31.V.2009, Coll. D. Zhang (MBFU). Distribution. Palearctic: China (Beijing, new record from mainland China), Japan; Oriental: China (Taiwan)., Published as part of Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu & Wang, Rong-Rong, 2016, Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species, pp. 401-414 in Zootaxa 4079 (4) on pages 406-408, DOI: 10.11646/zootaxa.4079.4.1, http://zenodo.org/record/1050841, {"references":["Nishida, K. (1974) Studies on the species of Fanniinae (Diptera: Muscidae) from Japan. II. One new species from Japan and Taiwan, and four newly recorded species of genus Fannia from Japan. Japanese Journal of Sanitary Zoology, 25 (3), 205 - 210.","Pont, A. C. (1977) Family Fanniidae. In: Delfinado, M. D. & Hardy, D. E. (Eds.), A catalog of Diptera of the Oriental Region. Vol. 3. Suborder Cyclorrhapha (excluding Division Aschiza). University of Hawaii, Honolulu, pp. 447 - 450.","Pont, A. C. (1986) Family Muscidae. In: Soos, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Scathophagidae - Hypodermatidae. Akademiai Kiado, Budapest, 11, pp. 41 - 57.","Nishida, K. (1994) The Fanniidae from Nepal (Diptera). Japanese Journal of Sanitary Zoology, Supplement 45, 81 - 97. [In English with Japanese abstract]","Xue, W. Q. & Wang, M. F. (1998) Fanniidae. In: Xue, W. Q. & Chao, C. M. (Eds.), Flies of China. Liaoning Science and Technology Press, Shenyang, pp. 809 - 835. [In Chinese with English abstract]","Lin, F. J. & Chen, J. S. (1999) Family Fanniidae. In: Lin, F. J & Chen, C. S. (Eds.), The name list of Taiwan Diptera. The Museum, Institute of Zoology, Academia Sinica Taipei, Taiwan, R. O. C., pp. 107 - 108. [In Chinese with English Title]","Wang, M. F. & Xue, W. Q. (2002) Taxonomic study on Fanniidae of China (Diptera: Cyclorrhapha). In: Li, D. M., Kang, L., Wu, J. W. & Zhang, R. Z. (Eds.), Advance in entomology: proceedings of the 2002 ' conference organized by Entomological Society of China. Science and Technology Press of China, Beijing, pp. 54 - 59. [In Chinese]"]}

Fannia hirticeps (Stein, 1892) (Fig. 5) Homalomyia hirticeps Stein, 1892: 70. Type locality: Germany, near Genthin. Fannia hirticeps: Hennig 1955: 21; Chillcott 1961: 96; Zhang et al. 1984: 3; Pont 1986: 48; Wang & Wu 1996: 66; Rozkošný et al. 1997: 22; Xue & Wang 1998: 815; Wu & Wang 2002: 563; Su & Wang 2004: 111; Wang et al. 2004a: 34; Wang & Wang 2006: 63; Xue et al. 2006: 496; Domínguez & Roig-Juñent 2008: 577. Redescription based on the specimens collected in Beijing, China. MALE. Body length about 5.00 mm. Eye with long and dense brownish hairs; facets slightly enlarged on anterior margin in upper part; fronto-orbital plate and parafacial with dense grayish-silvery pollinosity; frons about as wide as the distance between two posterior ocelli at narrowest point; frontal vitta black, with grayish-white pollinosity, linear at narrowest point; frontal setae six to eight, stout, situated on the lower 3/4 of frons, orbital setae absent; postocular setae short, in one row, with some occipital setae behind the postocular setae on vertex; parafacial bare, at middle about 0.75x to 1.00x as wide as the width of postpedicel; antenna black, postpedicel about 2.00x longer than wide, arista black, bare or short plumose, slightly swollen in basal part; epistoma about at the same level of vibrissal angle, vibrissal angle behind frontal angle in profile; subvibrissal setae in one row, lateral with one or two rows of short setae; gena and genal dilation with fine black setae, upper margin of gena without upcurved seta; prementum with thin grayish-yellow pollinosity, its length about 2.00x as long as its width; palpus black, claviform, as long as the length of prementum. Thorax ground-color black, notum with dense grayish-yellow pollinosity, without distinct vitta; presutural acr biserial, hair-like, only prescutellar pairs long, dc 2+3, ia 0+2, pra 1, stout, about 2/3 as long as the length of posterior notopleural seta; notopleuron without seta; basisternum, proepisternum, anepimeron, meron and katepimeron bare; katepisternal setae 1+1, katepisternum with some setae and without ventral spine; anterior spiracle yellowish, posterior one brown; calypters yellowish, the lower one equal in size to the upper one. Wing brown; veins dark brown; tegula black; basicosta yellowish-brown; costal spine inconspicuous; node of Rs bare on ventral and dorsal surfaces; vein M straight, slightly close to vein R4+5 distally; crossveins without cloud; haltere brownish-yellow and dark drown at basal part. Legs black; fore coxa without anterior spine on ventral surface, fore femur with complete pv row, fore tibia with one apical d, each segment of fore tarsi with one pair of slender sensory setae; mid coxa without any hook-like spine or spine-like seta, mid femur with stout and sparse av row in basal part, becoming shorter and denser towards apex, with a gap in subapical part, with three or four comb-like setae in distal part, with stout pv row, slightly biserial in median part, p row long and sparse, mid tibia slightly swollen in distal 2/3, with two or three ad and two pd, and with numerous slender setae on ventral surface, the longest one slightly shorter than mid tibial width in distal part, mid first tarsomere with a basal tooth-like spine on ventral surface, the length of the spine about 1.50x to 2.00x as long as tibial width in basal part; hind coxa bare on posterior surface, hind femur slightly curved, with a complete av row, becoming stout and slightly upcurved in distal 3/4, the length of the longest av more than 2.00x of femur width in distal half, with one row of slender setae in posterior surface in basal part, becoming to posterodorsal surface in median part, and to posteroventral surface in distal part, several setae with tip curved in distal part, hind tibia with one stout median av, one ad, and one median a. Abdomen long and flattened, ground-color black, with dense greenish-gray pollinosity; syntergite 1+2 to tergite 4 each with one dark broad median triangular vitta, tergite 5 with one dark median vitta in basal part, each tergite with long lateral marginal setae; sternite 1 with four to six long setae on each side; terminalia see Chillcott (1961: 30, 34, 172). FEMALE. Body length about 3.90 mm. Eye facets equal large; fronto-orbital plate and parafacial with dense grayish pollinosity; frons at middle about 3/7 as wide as the distance between the width of head; orbital setae two; frontal setae five; at middle parafacial about 3/5 as wide as the width of postpedicel. Mid tibia without slender seta on ventral surface; hind tibia with two d, without a. Abdomen entirely black. Other morphological characteristics are similar to those of the male. Material examined. CHINA: Beijing: 10♂, Zhongguancun, 10−11.IV.1972, [collector unknown] (NZMC); 4♂, Yanqing, Mt. Songshan, 12.IV.2010, Coll. D. Zhang (MBFU); 21♂, 1♀, Haidian, Campus of Beijing Forestry University, 27.IV.2010, Coll. D. Zhang (MBFU); 12♂, Haidian, Campus of Beijing Forestry University, 12.IV.2012, Coll. M. Zhang & M.X. Zhang (MBFU); 24♂, Haidian, Campus of Beijing Forestry University, 13.IV.2012, Coll. Y.Z. Yang, Y.O. Chen & X.H. Liu (MBFU); 1♂, Haidian, Campus of Beijing Forestry University, 8.IV.2013, Coll. D. Zhang (MBFU); 7♂, Haidian, Campus of Beijing Forestry University, 15.IV.2013, Coll. X.H. Liu (MBFU); 29♂, Haidian, Campus of Beijing Forestry University, 20−24.IV.2013, Coll. M. Zhang (MBFU); 5♂, Haidian, Campus of Beijing Forestry University, 3.IV.2014, [collector unknown] (MBFU); 27♂, Haidian, Campus of Beijing Forestry University, 6.IV.2014, Coll. W. Li & H.J. Wan (MBFU). Distribution. Palearctic: China (Beijing, Shandong, Shanxi), Germany; Nearctic: USA (Alaska)., Published as part of Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu & Wang, Rong-Rong, 2016, Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species, pp. 401-414 in Zootaxa 4079 (4) on pages 408-410, DOI: 10.11646/zootaxa.4079.4.1, http://zenodo.org/record/1050841, {"references":["Stein, P. (1892) Drei neue merkwurdige Homalomyia-Arten. Ein dipterologischer Beitrag. Wiener entomogische Zeitung, 11, 69 - 77.","Hennig, W. (1955) Muscidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region, 63 b. Schweizerbart, Stuttgart, pp. 1 - 99.","Chillcott, J. G. (1961) A revision of the Nearctic species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, Supplement 14, 92, [1960], 1 - 295. http: // dx. doi. org / 10.4039 / entm 9214 fv","Zhang, W. Z., Wang, C. J., Zhao, B., Jing, T., Wu, S. T. & Wang, M. F. (1984) A revision of flies from Shanxi Province. II. Sanitation and Epidemic Prevention of Shanxi, Supplement, 47, 1 - 34. [In Chinese]","Pont, A. C. (1986) Family Muscidae. In: Soos, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Scathophagidae - Hypodermatidae. Akademiai Kiado, Budapest, 11, pp. 41 - 57.","Wang, M. F. & Wu, Y. X. (1996) A study on the classification of Fanniidae in Shanxi Province. Shanxi Journal of Preventive Medicine, 5 (2), 65 - 67. [In Chinese]","Rozkosny, R., Gregor, F. & Pont, A. C. (1997) The European Fanniidae (Diptera). Acta Scientiarum naturalium Academiae scientiarum Bohemicae - Brno, 31 (2), 1 - 80.","Xue, W. Q. & Wang, M. F. (1998) Fanniidae. In: Xue, W. Q. & Chao, C. M. (Eds.), Flies of China. Liaoning Science and Technology Press, Shenyang, pp. 809 - 835. [In Chinese with English abstract]","Wu, Y. X. & Wang, M. F. (2002) Taxonomic study on Fanniidae of Shanxi (Diptera: Cyclorrhapha). In: Li, D. M., Kang, L., Wu, J. W. & Zhang, R. Z. (Eds.), Advance in entomology: proceedings of the 2002 ' conference organized by Entomological Society of China. Science and Technology Press of China, Beijing, pp. 562 - 564. [In Chinese]","Wang, M. F., Xue, W. Q. & Cao, X. F. (2004 a) Studies on the Family Fanniidae from the Subregion Loess Plateau of China (Diptera: Fanniidae). Chinese Journal of Vector Biology and Control, 15 (1), 33 - 35. [In Chinese with English abstract]","Wang, M. F. & Wang, R. R. (2006) A review of the hirticeps - group of Fannia Robineau-Desvoidy (Diptera: Fanniidae), with the description of one new species from China. Zootaxa, 1295, 61 - 68.","Xue, J., Zhan, C. D. & Li, L. (2006) The catalogue and distribution of important families and genera of Calyptratae (Diptera) in Shandong Province. Chinese Journal of Vector Biology and Control, 17 (6), 495 - 498. [In Chinese]","Dominguez, M. C. & Roig-Junent, S. A. (2008) A phylogeny of the family Fanniidae Schnabl (Insecta: Diptera: Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia. Invertebrate Systematics, 22, 563 - 587. http: // dx. doi. org / 10.1071 / IS 08003"]}

Fannia carbonaria (Meigen, 1826) (Figs. 2, 3) Anthomyia carbonaria Meigen, 1826: 154. Type locality: Germany, Stolberg. Fannia carbonaria: Stein 1915: 28; Hennig 1955: 23; Chillcott 1961: 124; Pont 1977: 448; Pont 1986: 46; Rozkošný et al. 1997: 27; Xue & Wang 1998: 818; Lin & Chen 1999: 108; Wang & Xue 2002: 55; Su & Wang 2004: 110; Wang et al. 2004b: 136; Domínguez & Roig-Juñent 2008: 577; Wang et al. 2009: 38. Redescription based on the specimens collected in Beijing, China. MALE. Body length about 4.60 mm. Eye bare; facets slightly enlarged on anterior margin in upper part; upper part of fronto-orbital plate with grayish-drown pollinosity, lower part of fronto-orbital plate and parafacial with grayish-silvery pollinosity; frons slightly narrower than the distance between two posterior ocelli at narrowest point; frontal vitta black, the narrowest point narrower than the width of anterior ocellus; frontal setae eight or nine, stout, reaching ocellar triangle, situated on the lower 5/6 of fronto-orbital plate; postocular setae in one row, without occipital seta behind the postocular setae on vertex; parafacial bare, at middle about 3/4 as wide as the width of postpedicel; antenna brown, postpedicel 2.00x longer than wide, arista black and short plumose, slightly swollen in basal part, the longest individual hair slightly shorter than aristal base; epistoma not projecting beyond vibrissal angle, vibrissal angle behind frontal angle in profile; subvibrissal setae in one row, lateral with one short row of setae; gena and genal dilation with fine black setae, upper margin of gena without upcurved seta; proboscis stout, prementum with thin grayish-yellow pollinosity; palpus black, claviform, longer than the length of prementum. Thorax ground-color black, notum slightly shining, with thinly brownish pollinosity, without distinct vitta; presutural acr triserial, prescutellar pairs slightly stronger, the distance between two outer acr rows narrower than the distance between acr and dc rows, dc 2+3, ia 0+2, pra 1, stout, about 3/4 as long as posterior notopleural seta; notopleuron without seta; basisternum, proepisternum, anepimeron, meron and katepimeron bare; katepisternal setae 1+1, katepisternum with some setae and without ventral spine; anterior spiracle small and brown, posterior one dark brown; calypters yellowish, the lower one slightly projecting beyond the upper one. Wing brownish; veins dark brown; tegula black; basicosta brownishyellow; costal spine inconspicuous; node of Rs bare on ventral and dorsal surfaces; vein M straight, slightly parallel to vein R4+5 distally; crossveins without obvious cloud; haltere orange. Legs entirely black; fore coxa without anterior spine on ventral surface, fore femur with complete pv row, short and weak in basal 1/3, fore tibia without ad and median p; mid coxa without any hook-like spine or spine-like seta, mid femur with complete av row, becoming gradually shorter and denser towards apex, pv row complete, in one row, p row slender, four or five setae strong in distal part, mid tibia slightly swollen in distal part, with two ad, one or two pd, and with numerous slender setae on ventral surface, the longest one about half as long as mid tibial width in distal part, mid first tarsomere without basal tooth-like spine on ventral surface; hind coxa bare on posterior surface, hind femur only with three or four long av in distal 2/5, without distinct pv row, hind tibia slightly curved, with two av, two ad, with one long median p. Abdomen oval and flattened, ground-color black, with thin dark brown pollinosity; syntergite 1+2 to tergite 4 each with one inconspicuous median inverted triangular vitta, tergite 5 without distinct stripe; sternite 1 with three slender setae in lateral part; terminalia see Chillcott (1961: 70). FEMALE. Unknown from China, description see Hennig (1955: 36). Type material examined. Lectotype ♀, Germany [probably Stolberg], designated by A.C. Pont (MNHN). Material examined. CHINA: Beijing: 1♂, Yanqing, Mt. Songshan, 31.V.2009, Coll. D. Zhang (MBFU). Distribution. Palearctic: China (Beijing, new record from mainland China), Austria, Belgium, Britain, Czech, Denmark, Finland, Germany, Hungary, Japan, Norway, Poland, Rumania, Slovakia, Sweden, Switzerland; Nearctic: Canada (British Columbia), USA (Washington); Oriental: China (Taiwan)., Published as part of Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu & Wang, Rong-Rong, 2016, Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species, pp. 401-414 in Zootaxa 4079 (4) on pages 403-405, DOI: 10.11646/zootaxa.4079.4.1, http://zenodo.org/record/1050841, {"references":["Meigen, J. W. (1826) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Vol. 5. Hamm, 412 pp.","Stein, P. (1915) H. Sauter's Formosa-Ausbeute. Anthomyidae (Diptera). Supplementa Entomologica, 4, 13 - 56.","Hennig, W. (1955) Muscidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region, 63 b. Schweizerbart, Stuttgart, pp. 1 - 99.","Chillcott, J. G. (1961) A revision of the Nearctic species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, Supplement 14, 92, [1960], 1 - 295. http: // dx. doi. org / 10.4039 / entm 9214 fv","Pont, A. C. (1977) Family Fanniidae. In: Delfinado, M. D. & Hardy, D. E. (Eds.), A catalog of Diptera of the Oriental Region. Vol. 3. Suborder Cyclorrhapha (excluding Division Aschiza). University of Hawaii, Honolulu, pp. 447 - 450.","Pont, A. C. (1986) Family Muscidae. In: Soos, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Scathophagidae - Hypodermatidae. Akademiai Kiado, Budapest, 11, pp. 41 - 57.","Rozkosny, R., Gregor, F. & Pont, A. C. (1997) The European Fanniidae (Diptera). Acta Scientiarum naturalium Academiae scientiarum Bohemicae - Brno, 31 (2), 1 - 80.","Xue, W. Q. & Wang, M. F. (1998) Fanniidae. In: Xue, W. Q. & Chao, C. M. (Eds.), Flies of China. Liaoning Science and Technology Press, Shenyang, pp. 809 - 835. [In Chinese with English abstract]","Lin, F. J. & Chen, J. S. (1999) Family Fanniidae. In: Lin, F. J & Chen, C. S. (Eds.), The name list of Taiwan Diptera. The Museum, Institute of Zoology, Academia Sinica Taipei, Taiwan, R. O. C., pp. 107 - 108. [In Chinese with English Title]","Wang, M. F. & Xue, W. Q. (2002) Taxonomic study on Fanniidae of China (Diptera: Cyclorrhapha). In: Li, D. M., Kang, L., Wu, J. W. & Zhang, R. Z. (Eds.), Advance in entomology: proceedings of the 2002 ' conference organized by Entomological Society of China. Science and Technology Press of China, Beijing, pp. 54 - 59. [In Chinese]","Wang, M. F., Xue, W. Q. & Su, L. X. (2004 b) Notes on carbonaria species group of genus Fannia Robineau-Desvoidy (Diptera: Fanniidae), in China. Entomologia Sinica, 11 (2), 135 - 142. [In English with Chinese abstract] http: // dx. doi. org / 10.1111 / j. 1744 - 7917.2004. tb 00187. x","Dominguez, M. C. & Roig-Junent, S. A. (2008) A phylogeny of the family Fanniidae Schnabl (Insecta: Diptera: Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia. Invertebrate Systematics, 22, 563 - 587. http: // dx. doi. org / 10.1071 / IS 08003","Wang, M. F., Zhang, D., Zheng, S. & Zhang, C. T. (2009) A review of the carbonaria - subgroup of Fannia Robineau-Desvoidy (Diptera: Fanniidae), with descriptions of two new species from China. Zootaxa, 2204, 37 - 47."]}

Key to the known males of the family Fanniidae in Beijing, China 1 Arista long plumose; katepisternal setae 1+2 (Piezura Rondani); frons much broader than the distance between two posterior ocelli at narrowest point; legs yellow, except tarsi and apical part of femora black; abdomen ground-color reddish-yellow.......................................................................... Piezura shanxiensis Xue, Wang & Wu - Arista short plumose or nearly bare; katepisternal setae 1+1 (Fannia Robineau-Desvoidy)............................ 2 2 Abdomen spatulate in shape, becoming narrow basad (Fig. 1A); mid femur with two stout thorn-like av in subbasal part (Fig. 1B)....................................................................... Fannia subpellucens (Zetterstedt) - Abdomen not spatulate in shape; mid femur without any stout thorn-like av in subbasal part.......................... 3 3 Mid coxa with hook-like spines on lower margin (Fig. 1C, D, E)................................................ 4 - Mid coxa without any hook-like spine on lower margin, or only with stout and straight spine.......................... 6 4 Mid tibia with a tubercle on ventral surface (Fig. 1C); mid coxa with three hook-like spines (Fig. 1C)................................................................................................... Fannia scalaris (Fabricius) - Mid tibia without tubercle on ventral surface; mid coxa with one hook-like spine (Fig. 1D, E)......................... 5 5 Lower margin of katepisternum without stout spine-like seta; fore tibia yellow at the basal 2/3 and with a long setae cluster on the posteroventral surface (Fig. 1F).................................................. Fannia manicata (Meigen) - Lower margin of katepisternum with a stout spine-like seta (Fig. 1D); fore tibia entirely black and without long setae cluster on the posteroventral surface............................................................. Fannia fuscula (Fall��n) 6 Mid first tarsomere with a basal tooth-like spine (Fig. 1G, H; Fig. 5E)............................................ 7 - Mid first tarsomere without basal tooth-like spine............................................................9 7 Fore tibia with a row of slender pv (Fig. 1I).......................................... Fannia spathiophora Malloch - Fore tibia without pv row............................................................................... 8 8 Eyes bare; fore first tarsomere with slender and dense light brown setae on ventral surface (Fig. 1J); with two setae clusters between fore and mid coxae (Fig. 1K); cercus with long setae cluster (Fig. 1L)................... Fannia barbata (Stein) - Eyes with fine brownish hairs; fore first tarsomere without slender and dense seta on ventral surface; without setae cluster between fore and mid coxae; cercus without setae cluster.................................... Fannia hirticeps (Stein) 9 Hind coxa on posterior surface with setae................................................................. 10 - Hind coxa on posterior surface bare...................................................................... 15 10 Abdominal tergites 3 and 4 each with three black spots.................................. Fannia leucosticta (Meigen) - Abdominal syntergite 1+2 to tergite 4 with median stripes or triangular vittas even without any mark.................. 11 11 Mid tibia with two or more pd..........................................................................12 - Mid tibia with one pd................................................................................. 13 12 Hind tibia with three to six av and four to six ad; mid femur without notch in distal area; mid tibia without slender seta on ven- tral surface........................................................................... Fannia dupla Nishida - Hind tibia with two av and one ad; mid femur with a notch in distal area; mid tibia with numerous slender setae on ventral sur- face (Fig. 6E)........................................................................ Fannia papei sp. nov. 13 Lateral part of abdominal tergites yellow, with thin pollinosity.......................... Fannia canicularis (Linnaeus) - Abdomen black, with dense greyish-blue pollinosity......................................................... 14 14 Abdominal syntergite 1+2 to tergite 4 each with one inconspicuous median inverted triangular vitta, tergite 5 with one narrow median stripe; hind femur with two long av in distal 1/3 (Fig. 1N); hind tibia with one row of p (Fig. 1M), and with one row of ad, irregular in length (Fig. 1N).................................................... Fannia lepida (Wiedemann) - Abdominal syntergite 1+2 to tergite 5 each with one median longitudinal stripes; hind femur with four long av on distal 1/3; hind tibia without p and with one ad........................................................ Fannia prisca Stein 15 Hind femur with a setae cluster on posteroventral surface in distal 1/3 (Fig. 1O)......... Fannia metallipennis (Zetterstedt) - Hind femur without setae cluster on posteroventral surface in distal 1/3.......................................... 16 16 Hind tarsomeres stout (Fig. 2E); pra 1; thorax and abdomen ground-color gloss black (Fig. 2A, B); mid femur without notch in distal area...................................................................... Fannia carbonaria (Meigen) - Hind tarsomeres not stout; pra 2; thorax and abdomen with light gray pollinosity; mid femur with a notch in distal area........................................................................................ Fannia postica (Stein), Published as part of Zhang, Dong, Li, Wei, Zhang, Ming, Wang, Ming-Fu & Wang, Rong-Rong, 2016, Fanniidae (Insecta, Diptera) from Beijing, China, with key and description of one new species, pp. 401-414 in Zootaxa 4079 (4) on pages 402-403, DOI: 10.11646/zootaxa.4079.4.1, http://zenodo.org/record/1050841

Fanniidae (Insecta: Diptera) is a relatively small family (ca. 350 spp.) with fve genera, of which Fannia Robineau-Desvoidy, 1830 and Euryomma Stein, 1899 have Neotropical distributions. Some of these species are almost always found in forensic studies. Forensically relevant species have been neglected, despite recent forensic studies that suggest their importance for estimating post-mortem interval (PMI). Thus, current and updated keys to identify adults or larvae on carcasses are unavailable for the most important species. While immature stages are important in estimating PMI, evidence suggests that adults (Fanniidae as well as other families) may also be useful for this purpose. Here we provide a key to males of the species of Fanniidae (found on corpses and other decomposing organic matter) with a checklist of species that have been used in forensics in the Neotropical region. The key comprises all 38 species of Fannia and Euryomma that have already been successfully used in forensics, and species that are potentially useful for estimating PMI. These records were found after reviews of the literature and data from entomological collections. Photographs and illustrations of the main characters in the key are provided Fil: Grisales, Diana. Universidad de Antioquía. Instituto de Biología. Grupo de Entomología; Colombia Fil: Lecheta, Melise C.. Universidade Federal do Paraná. Departamento de Zoologia; Brasil Fil: Aballay, Fernando Hernán. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Mendoza. Instituto Argentino de Investigaciones de las Zonas Áridas. Provincia de Mendoza. Instituto Argentino de Investigaciones de las Zonas Áridas. Universidad Nacional de Cuyo. Instituto Argentino de Investigaciones de las Zonas Áridas; Argentina Fil: Carvalho, Claudio J. B. de. Universidade Federal do Paraná. Departamento de Zoologia; Brasil