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Anchoa helleri (Hubbs, 1921). Gulf Anchovy. To 10.3 cm (4.1 in) SL (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California). Bah��a Magdalena, southern Baja California (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California) and into Gulf of California (Robertson and Allen 2008). Nearshore; depth: about 0���4 m (12 ft) (min.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California; max.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 41, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Hubbs, C. L. (1921) Notes on Cottus asper and Cottus aleuticus. Copeia, 90, 7 - 8.","Robertson, D. R. & Allen, G. R. (2008) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 1.0 (2008). Smithsonian Tropical Research Institute, Balboa."]}

Unidentified opistognathid. Lissner and Dorsey (1986) observed from a manned submersible a jawfish they were unable to identify at Tanner��� Cortes Bank, southern California., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 150, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Lissner, A. L. & Dorsey, J. H. (1986) Deep-water biological assemblages of a hard-bottom bank-ridge complex of the southern California continental borderland. Bulletin of the Southern California Academy of Sciences, 85, 87 - 101."]}

Anchoa nasus (Kner & Steindachner, 1867). Bignose Anchovy or Longnose Anchovy. To 17 cm (6.7 in) TL (Robertson and Allen 2002). Bah��a Magdalena, southern Baja California (De La Cruz-Ag��ero et al. 1994) and throughout Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995) to Arica, northern Chile (Sielfeld et al. 2010). Pelagic and nearshore, tolerating some low salinity (Whitehead et al. 1994); depth: 0���142 m (466 ft) (min.: Robertson and Allen 2015; max.: Zeballos Flor et al. 1998). The maximum depth of capture was made by bottom trawl and it is possible this species was captured in midwater., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 42, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2002) Shorefishes of the tropical eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama.","De La Cruz-Aguero, J., Galvan-Magana, F., Abitia-Cardenas, L. A., Rodriguez-Romero, J. & Gutierrez-Sanchez, F. J. (1994) Systematic list of marine fishes from Bahia Magdalena, Baja California Sur (Mexico). Ciencias Marinas, 20, 17 - 31.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome.","Sielfeld, W., Laudien, J., Vargas, M. & Villegas, M. (2010) El Nino induced changes of the coastal fish fauna off northern Chile and implications for ichthyogeography. Revista de Biologia Marina Oceanografia, 45, S 1, 705 - 722. https: // doi. org / 10.4067 / s 0718 - 19572010000400014","Robertson, D. R. and Allen, G. R. (2015) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 2.0 (2008). Smithsonian Tropical Research Institute, Balboa. http: // biogeodb. stri. si. edu / sftep / en / pages","Zeballos Flor, J., Samame, M. & Romero, M. (1998) Estructura especiologica demersal observada durante el Crucero de Evaluacion de la Merluza entre Puerto Pizarro y Huarmey. Informe IMARPE, 138, 87 - 100."]}

Cetengraulis mysticetus (G��nther, 1867). Anchoveta. To about 22 cm (8.7 in) TL (Robertson and Allen 2002). Los Angeles, southern California (Miller and Lea 1972) to Callao, Peru (Beltr��n-Le��n and Rios Herrera 2000), including Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995) and Islas Gal��pagos (Grove and Lavenberg 1997). Marine and brackish waters (Fricke et al. 2020); depth: surface to 38 m (155 ft) (min.: Robertson and Allen 2002; max.: Personal communication: Los Angeles County Museum of Natural History Fish Collection, Los Angeles, California). The maximum depth capture was made by bottom trawl and it is possible this species was captured in midwater., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 42, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2002) Shorefishes of the tropical eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama.","Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Beltran-Leon, B. S. & Rios Herrera, R. (2000) Estadios Tempranos de Peces del Pacific Colombiano. Republica de Colombia. Instituto Nacional de Pesca y Acuicultura INPA.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome.","Grove, J. S. & Lavenberg, R. J. (1997) The Fishes of the Galapagos Islands. Stanford University Press, Stanford.","Fricke, R., Eschmeyer, W. N. & van der Laan, R. (Eds). 2020. Eschmeyer's Catalog of Fishes: Genera, Species, Reference. Available from: http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (accessed 17 April 2021)."]}

Lile nigrofasciata Castro-Aguirre, Ruiz-Campos, & Balart, 2002. Blackstripe Herring. To 13 cm (5.1 in) (Robertson and Allen 2008). Bah��a Magdalena, southern California and Gulf of California to northern Peru (Castro-Aguirre et al. 2002). Lagoons and other shallow littoral waters (Castro-Aguirre et al. 2002)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 44, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2008) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 1.0 (2008). Smithsonian Tropical Research Institute, Balboa.","Castro-Aguirre, J. L., Ruiz-Campos, G. & Balart, E. F. (2002) A new species of the genus Lile (Clupeiformes: Clupeidae) of the eastern tropical Pacific. Bulletin of the Southern California Academy of Sciences, 101, 1 - 12."]}

** Alosa sapidissima (Wilson, 1811). American Shad. To 76 cm (30 in) TL (Miller and Lea 1972). Native to Atlantic Ocean; intentionally introduced to Pacific Ocean, spread to Kamchatka, Russia to south-eastern Bering Sea and Gulf of Alaska (Mecklenburg et al. 2002) to Bah��a de Todos Santos, northern Baja California (Miller and Lea 1972). Depth: surface to 250 m (820 ft) (Allen and Smith 1988). While there are a few other, deeper, records (e.g., 440 m, 1,443 ft, Bradburn et al. 2011; 646 m, 2,119 ft, 881 m, 2,890 ft, and 1,151 m, 3,885 ft, NWFSC-FRAM), all are from bottom trawls and the fish could have entered the nets in the water column as the net were being deployed or retrieved. Anadromous., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 43, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Allen, M. J. & Smith, G. B. (1988) Atlas and zoogeography of common fishes in the Bering Sea and northeastern Pacific. NOAA Technical Report NMFS, 66.","Bradburn, M. J., Keller, A. A. & Horness, B. H. (2011) The 2003 to 2008 U. S. West Coast bottom trawl surveys of groundfish resources off Washington, Oregon, and California: estimates of distribution, abundance, length, and age composition. National Marine Fisheries Service Technical Memorandum, NMFS-NWFSC- 114."]}

Anchoa argentivittata (Regan, 1904). Silverstripe Anchovy. To 12.5 cm (4.9 in) TL (Robertson and Allen 2002). Central Baja California (29��32���N, 115��12���W) (Personal communication: Universidad National Autonoma de Mexico, Colleccion Nacional de Peces, Mexico City, Mexico) to Bah��a Magdalena, southern Baja California (Galv��n-Maga��a et al. 2000) and central-eastern Gulf of California to Peru, including Islas Gal��pagos (Robertson and Allen 2015). Depth: surface to 10 m (33 ft) (Robertson and Allen 2002)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 41, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2002) Shorefishes of the tropical eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama.","Galvan-Magana, F., Gutierrez-Sanchez, F., Abitia-Cardenas, L. A. & Rodriquez-Romero, J. (2000) The distribution and affinities of the shorefishes of the Baja California Sur lagoons. In: Munawar, M., Lawrence, S. G., Munawar, I. F. & Malle, D. F. (Eds.), Aquatic Ecosystems of Mexico: Status and Scope. Ecovision World Monograph Series. Backhuys Publisher, Leiden, The Netherlands, pp. 383 - 398.","Robertson, D. R. and Allen, G. R. (2015) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 2.0 (2008). Smithsonian Tropical Research Institute, Balboa. http: // biogeodb. stri. si. edu / sftep / en / pages"]}

Anchoa exigua (Jordan & Gilbert, 1882). Slender Anchovy. To 7.5 cm (3 in) TL (Robertson and Allen 2002). Bah��a Playa Maria (28��56���N, 114��32���W), central Baja California (Knaggs et al. 1975) to southern Gulf of California (Whitehead et al. 1988) to Puerto Pizarro, Peru (Chirichigno and V��lez 1998). Pelagic, nearshore (Whitehead et al. 1988); depth: to 10 m (33 ft) (Robertson and Allen 2002). The author names for this species belong within parentheses, not without as sometimes seen; the species was classified by Jordan and Gilbert in the genus Stolephorus., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 41, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2002) Shorefishes of the tropical eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama.","Knaggs, E. H., Sunada, J. S. & Lea, R. N. (1975) Notes on some fishes collected off the outer coast of Baja California. California Fish and Game, 61, 56 - 59.","Whitehead, P. J. P., G. J. Nelson, G. J. & Wongratana, T. (1988) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 2. Engraulididae. FAO Fisheries Synopsis No. 125. FAO, Rome.","Chirichigno, F. N. & Velez D, J. (1998) Clave para identificaticar los peces marinos del Peru (segunda edicion, revisada y actualizada). Instituto de Mar de Peru Publicacion Especial."]}

Lile stolifera (Jordan & Gilbert, 1882). Pacific Piquintinga or Striped Herring. To 15.9 cm (6.3 in) TL (Coello et al. 2021). Bah��a Ballenas (26��40���N, 113��30���W), southern Baja California (Grove and Lavenberg 1994) to Puerto Pizarro, Peru (Chirichigno and V��lez 1974), including Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995) and Islas Gal��pagos (Grove and Lavenberg 1997). Marine, brackish, and fresh waters (Allen and Robertson 1994); depth: surface (nightlight; Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California), 1���3 m (3���10 ft) (min.: Gonz��lez-Acosta et al. 1999; max.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 44, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Coello, D., Herrera, M. & Zambrano, R. (2021) Length-weight relationship of 74 fish species caught in the continental coast of Ecuador. Journal of Applied Ichthyology, 37, 129 - 134.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome.","Grove, J. S. & Lavenberg, R. J. (1997) The Fishes of the Galapagos Islands. Stanford University Press, Stanford.","Allen, G. R. & Robertson, D. R. (1994) Fishes of the Tropical Eastern Pacific. University of Hawaii Press, Honolulu.","Gonzalez-Acosta, A. F., De La Cruz-Aguero, J. & Cota-Gomez, V. M. (1999) Extension of geographical distribution and first occurrence of fishes in the northwest of Mexico. Hidrobiologica, 9, 39 - 44."]}

Anchoa delicatissima (Girard, 1854). Slough Anchovy. To 12.9 cm (5.1 in) TL (Miller et al. 2008). Marina del Rey, southern California (Soule and Oguri 1987) to Bah��a Magdalena, southern Baja California (Miller and Lea 1972). Marine and brackish waters (Whitehead et al. 1988); depth: intertidal to 27 m (89 ft) (min.: Allen 1999; max.: City of San Diego 2020)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 41, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Miller, E. F. & Curtis, M. D. (2008) First occurrence of a Pacific crevalle jack, Caranx caninus, north of San Diego, California. Bulletin of the Southern California Academy of Sciences, 107, 41 - 43. https: // doi. org / 10.3160 / 0038 - 3872 (2008) 107 [41: fooapc] 2.0. co; 2","Soule, D. F. & Oguri, M. (1987) The marine Environment of Marina Del Rey. A report to the Department of Beaches and Harbors, County of Los Angeles. Harbor Environmental Projects, University of Southern California. Marine Studies of San Pedro Bay, California, Part 20 D.","Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Whitehead, P. J. P., G. J. Nelson, G. J. & Wongratana, T. (1988) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 2. Engraulididae. FAO Fisheries Synopsis No. 125. FAO, Rome.","Allen, L. G. (1999) Fisheries Inventory and Utilization of San Diego Bay, San Diego, California. Final report, sampling periods July 1994 to April 1999. Prepared for the United States Navy, Department of Defense, Agreement N 68711 - 94 - LT- 4033.","City of San Diego. 2020. Comprehensive Listing of Demersal Fish Species Collected by the City of San Diego's Ocean Monitoring Program. City of San Diego, Public Utilities Department, Environmental Monitoring and Technical Services Division, San Diego, CA. Updated August 2020."]}

Engraulis japonicus Temminck & Schlegel, 1846. Japanese Anchovy. To about 16 cm (6.3 in) SL (Lindberg and Legeza 1965). western Pacific Ocean north to south-eastern Kamchatka; one record from Alaska, south of western Aleutian Islands (Mecklenburg et al. 2002). Near surface, epipelagic and coastal, but also occurring to over 1,000 km (620 miles) from shore; depth: to 150 m (495 ft) (Mecklenburg et al. 2002)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 42, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Lindberg, G. U. & Legeza, M. I. (1965) Fishes of the Sea of Japan and the adjacent areas of the Sea of Okhotsk and the Yellow Sea. Part 2. Teleostomi. XII. Acipenseriformes - XXVIII. Polynemiformes. Handbook on the Identification of Animals, Zoological Institute of the Russian Academy (84) (Translation by the Israel Program for Scientific Translations, 1969.)"]}

Opisthopterus dovii (G��nther, 1868). Dove���s Longfin Herring or Pacific Longfin Herring. To 23 cm (9.1 in) (Robertson and Allen 2008). Punta Marquez, southern Baja California (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California) to Punta Malpelo, Peru (Chirichigno and V��lez 1998), including Gulf of California (Allen and Robertson 1994). Inshore waters and estuaries (Allen and Robertson 1994); depth: intertidal to 37 m (121 ft) (min.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California; max.: Personal communication: Los Angeles County Museum of Natural History Fish Collection, Los Angeles, California)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 43, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2008) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 1.0 (2008). Smithsonian Tropical Research Institute, Balboa.","Chirichigno, F. N. & Velez D, J. (1998) Clave para identificaticar los peces marinos del Peru (segunda edicion, revisada y actualizada). Instituto de Mar de Peru Publicacion Especial.","Allen, G. R. & Robertson, D. R. (1994) Fishes of the Tropical Eastern Pacific. University of Hawaii Press, Honolulu."]}

Anchoa mundeoloides (Breder, 1928). Northern Gulf Anchovy. To 15 cm (5.9 in) TL (Robertson and Allen 2002). Bah��a Sebastian de Vizcaino (27��51���N, 115��04.7���W), southern Baja California (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California) to Gulf of California to Chiapas, Mexico (Gonz��lez-Acosta et al. 2018); possibly Costa Rica (Fricke et al. 2020). Depth: surface to perhaps 27 m (90 ft) (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 42, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. & Allen, G. R. (2002) Shorefishes of the tropical eastern Pacific: an information system. CD-ROM. Smithsonian Tropical Research Institute, Balboa, Panama.","Gonzalez-Acosta, A. F., Rodiles-Hernandez, R. & Gonzalez-Diaz, A. A. (2018) Checklist of the marine and estuarine fishes of Chiapas, Mexico. Marine Biodiversity, 48, 1439 - 1454. https: // doi. org / 10.1007 / s 12526 - 016 - 0630 - y","Fricke, R., Eschmeyer, W. N. & van der Laan, R. (Eds). 2020. Eschmeyer's Catalog of Fishes: Genera, Species, Reference. Available from: http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (accessed 17 April 2021)."]}

Sardinops sagax (Jenyns, 1842). Pacific Sardine. To 41 cm (16.1 in) TL (Clemens and Wilby 1961). Indo-Pacific (Whitehead 1985); north Pacific from south of Japan to southern Kamchatka and Commander Islands (Mecklenburg et al. 2002), and Cross Sound, south-eastern Alaska (Bruce Wing, pers. comm. to M.L.) to Gulf of California (Galv��n-Maga��a et al. 1996), and Islas Gal��pagos (Grove and Lavenberg 1997). Pelagic; depth: surface (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California), intertidal to 383 m (1,256 ft) (min.: Carlisle et al. 1960; max.: Jacobsen Stout et al. 2016) or perhaps to 982 m (3,221 ft) (Bradburn et al. 2011), although this latter depth was based on a bottom trawl catch and may represent fish caught well above the bottom., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 44, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Clemens, W. A. & Wilby, G. V. (1961) Fishes of the Pacific Coast of Canada, Second Edition. Bulletin of the Fisheries Research Board of Canada, 68.","Whitehead, P. J. P. (1985) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 1. Chirocentridae, Clupeidae and Pristigasteridae. FAO Fisheries Synopsis No. 125, FAO, Rome.","Galvan-Magana, F., Abitia-Cardenas, L. A., Rodriguez-Romero, J., Perez-Espana, H. & Chavez-Ramos, H. (1996) Systematic list of the fishes from Cerralvo Island, Baja California Sur, Mexico. Ciencias Marinas, 22, 295 - 311.","Grove, J. S. & Lavenberg, R. J. (1997) The Fishes of the Galapagos Islands. Stanford University Press, Stanford.","Carlisle, J. G. Jr., Schott, J. W. & Abramson, N. J. (1960) The barred surfperch (Amphistichus argenteus Agassiz) in southern California. California Department of Fish and Game, Fish Bulletin, 109.","Jacobsen Stout, N., Kuhnz, L., Lundsten, L., Schlining, B., Schlining, K. & von Thun, S. (Eds.). (2016) T he Deep-Sea Guide (DSG). Monterey Bay Aquarium Research Institute. http: // dsg. mbari. org. Accessed 30 September 2016.","Bradburn, M. J., Keller, A. A. & Horness, B. H. (2011) The 2003 to 2008 U. S. West Coast bottom trawl surveys of groundfish resources off Washington, Oregon, and California: estimates of distribution, abundance, length, and age composition. National Marine Fisheries Service Technical Memorandum, NMFS-NWFSC- 114."]}

Clupea pallasii Valenciennes, 1847. Pacific Herring. To 46 cm (18 in) TL (Miller and Lea 1972). Korea and Japan to Arctic Ocean off Alaska to northern Baja California; Arctic Canada as far east as Queen Maud Gulf (Coad in Coad and Reist 2018). Scattered populations in Norwegian Barents Sea and Russian White and Pechora Seas (Whitehead 1985, J��rstad et al. 1994, J��rstad 2004, Laakkonen et al. 2015). This species hybridizes with Clupea harengus Linnaeus, 1758 in northern Europe (Laakkonen et al. 2015). Marine, brackish waters and, very occasionally, fresh waters (Moyle 2002); depth: surface to 250 m (820 ft) (Whitehead 1985). There are a number of deeper capture records (e.g., 723 m, 2,371 ft, Bradburn et al. 2011; 1,033 m, 3,388 ft, 1,068 m, 3,503 ft, NWFSC-FRAM) all recorded from bottom trawl catches. In these circumstances it is not possible to ascertain if these fish were caught on the bottom or in midwaters during net deployment or recovery. Almost always captured above 150 m (492 ft) (Whitehead 1985)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 43, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Coad, B. W. & Reist, J. D. (Eds.). (2018) Marine Fishes of Arctic Canada. University of Toronto Press, Toronto.","Whitehead, P. J. P. (1985) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 1. Chirocentridae, Clupeidae and Pristigasteridae. FAO Fisheries Synopsis No. 125, FAO, Rome.","Jorstad, K. E., Dahle, G. & Paulsen, O. I. (1994) Genetic comparison between Pacific herring (Clupea pallasi) and a Norwegian fjord stock of Atlantic herring (Clupea harengus). Canadian Journal of Fisheries and Aquatic Sciences, 51, 233 - 239.","Jorstad, K. E. (2004) Evidence for two highly differentiated herring groups at Goose Bank in the Barents Sea and the genetic relationship to Pacific herring, Clupea pallsi. Environmental Biology of Fishes, 69, 211 - 221. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0983 - 6 _ 17","Laakkonen, H. M., Strelkov, P., Lajus, D. L. & Vainola, R. (2015) Introgressive hybridization between the Atlantic and Pacific herrings (Clupea harengus and C. pallasii) in the north of Europe. Marine Biology, 162, 39 - 54. https: // doi. org / 10.1007 / s 00227 - 014 - 2564 - x","Moyle, P. B. (2002) Inland Fishes of California. Revised and Expanded. University of California Press, Berkeley.","Bradburn, M. J., Keller, A. A. & Horness, B. H. (2011) The 2003 to 2008 U. S. West Coast bottom trawl surveys of groundfish resources off Washington, Oregon, and California: estimates of distribution, abundance, length, and age composition. National Marine Fisheries Service Technical Memorandum, NMFS-NWFSC- 114."]}

Anchoa compressa (Girard, 1858). Deepbody Anchovy. To 16.5 cm (6.5 in) TL (Miller and Lea 1972). Morro Bay, central California (Miller and Lea 1972) to Bah��a Magdalena, southern Baja California (Whitehead et al. 1988). Depth: intertidal to perhaps 19 m (61 ft) (min.: Carlisle et al. 1960; max.: Allen et al. 2002)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 41, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Whitehead, P. J. P., G. J. Nelson, G. J. & Wongratana, T. (1988) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 2. Engraulididae. FAO Fisheries Synopsis No. 125. FAO, Rome.","Carlisle, J. G. Jr., Schott, J. W. & Abramson, N. J. (1960) The barred surfperch (Amphistichus argenteus Agassiz) in southern California. California Department of Fish and Game, Fish Bulletin, 109.","Allen, M. J., Groce, A. K., Diener, D., Brown, J., Steinert, S. A., Deets, G., Noblet, J. A., Moore, S. L., Diehl, D., Jarvis, E. T., Raco- Rands, V., Thomas, C., Ralph, Y., Gartman, R., Cadien, D., Weisberg, S. B. & Mikel, T. (2002) Southern California Bight 1998 Regional Monitoring Program: V. Demersal Fishes and Megabenthic Invertebrates. Southern California Coastal Water Research Project. Westminster, California."]}

** Dorosoma petenense (G��nther, 1867). Threadfin Shad. To 33 cm (13 in) TL (Moyle 2002). Native to the southeastern United States and south to Belize. Introduced to West Coast. Off Oregon (Eschmeyer and Herald 1983) to San Diego Bay, southern California (Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California) and Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995). Shallow fresh, brackish, and marine waters., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 43, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Moyle, P. B. (2002) Inland Fishes of California. Revised and Expanded. University of California Press, Berkeley.","Eschmeyer, W. N. & Herald, E. S. (1983) A Field Guide to Pacific Coast Fishes of North America from the Gulf of Alaska to Baja California. Houghton Mifflin, Boston.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome."]}

Engraulis mordax Girard, 1854. Northern Anchovy. To 24.8 cm (9.7 in) TL (Whitehead et al. 1988). One record off Yakutat, eastern Gulf of Alaska (Mecklenburg et al. 2002); Haida Gwaii, British Columbia to Cabo San Lucas, southern Baja California (Miller and Lea 1972), and Gulf of California (Hammann and Cisneros-Mata 1989). Depth: surface (M.L., pers. obs.), intertidal to 528 m (1,732 ft) (min.: Chotkowski 1994; max: Jacobsen Stout et al. 2016). We note that, as of March 2021, the NWFSC-FRAM database, based on bottom trawl surveys, has six records deeper than 528 m: 529 m, 1,735 ft, 543 m, 1,781 ft, 571 m, 873 ft, 601 m, 1,981 ft, 806 m, 2,644 ft, and 945 m, 3,100 ft), thus while these may represent catches made in the water column during net deployment or retrieval, the species may enter waters deeper than the current published maximum., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 42, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Whitehead, P. J. P., G. J. Nelson, G. J. & Wongratana, T. (1988) FAO Species Catalogue. Volume 7. Clupeoid Fishes of the World (Suborder Clupeoidei). Part 2. Engraulididae. FAO Fisheries Synopsis No. 125. FAO, Rome.","Miller, D. J. & Lea, R. N. (1972) Guide to the coastal marine fishes of California. California Department of Fish and Game Fish Bulletin, 157.","Hammann, M. G. & Cisneros-Mata, M. A. (1989) Range extension and commercial captures of the northern anchovy, Engraulis mordax Girard, in the Gulf of California, Mexico. California Fish and Game, 75, 49 - 53.","Chotkowski, M. A. (1994) The Behavioral Ecology and Population Dynamics of the Intertidal Fishes of the Northeast Pacific. Ph. D. Thesis, University of California, Los Angeles.","Jacobsen Stout, N., Kuhnz, L., Lundsten, L., Schlining, B., Schlining, K. & von Thun, S. (Eds.). (2016) T he Deep-Sea Guide (DSG). Monterey Bay Aquarium Research Institute. http: // dsg. mbari. org. Accessed 30 September 2016."]}

Opisthonema libertate (G��nther, 1867). Deepbody Thread Herring or Pacific Thread Herring. To about 30 cm (11.8 in) TL (Amezcua Linares 1996). Port Hueneme, southern California (Lea and Rosenblatt 2000) to Chile (Peque��o 1l989), including Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995) and Islas Gal��pagos (Grove and Lavenberg 1997). Marine and brackish waters (Fricke et al. 2020); depth: surface (nightlight; Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California), intertidal to at least 70 m (3���230 ft) (min.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California; max.: Velasco and Thiel 2002)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 44, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Amezcua Linares, F. (1996) Peces Demersales de la Platforma Continental del Pacifico Central de Mexico. Instituto de Ciencias del Mar y Limnologica, Universidad Nacional Autonoma de Mexico.","Lea, R. N. & Rosenblatt, R. H. (2000) Observations on fishes associated with the 1997 - 98 El Nino off California. CalCOFI Reports, 41, 117 - 129.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome.","Grove, J. S. & Lavenberg, R. J. (1997) The Fishes of the Galapagos Islands. Stanford University Press, Stanford.","Fricke, R., Eschmeyer, W. N. & van der Laan, R. (Eds). 2020. Eschmeyer's Catalog of Fishes: Genera, Species, Reference. Available from: http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (accessed 17 April 2021).","Velasco, A. & Thiel, R. (2002) A bilingual field manual for the identification of juvenile fish over soft bottom off the Pacific Coast of Colombia a review from literature. Archive of Fishery and Marine Research, 50, 55 - 118."]}

Opisthonema bulleri (Regan, 1904). Slender Thread Herring. To 24 cm (9.4 in) TL (Robertson and Allen 2015). Several specimens taken south-east of Punta Abreojos, southern Baja California (Jason Masters, photographs to M.L., tentatively confirmed by Ross Robertson); Gulf of California to northern Peru (Robertson and Allen 2015). Depth: intertidal to perhaps 10 m (33 ft) (min.: Personal communication: Scripps Institution of Oceanography Fish Collection, La Jolla, California; max.: Robertson and Allen 2015)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 44, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Robertson, D. R. and Allen, G. R. (2015) Shorefishes of the Tropical Eastern Pacific: an Information System. Version 2.0 (2008). Smithsonian Tropical Research Institute, Balboa. http: // biogeodb. stri. si. edu / sftep / en / pages"]}

Pliosteostoma lutipinnis (Jordan & Gilbert, 1882). Yellowfin Herring. To 25 cm (9.8 in) TL (Jim��nez Prado and B��arez 2004). Southern Baja California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995) to Colombia (Allen and Robertson 1994), including Gulf of California (Whitehead and Rodr��guez-S��nchez in Fischer et al. 1995). Coastal, including estuaries; depth: to 50 m (164 ft) (Amezcua Linares 1996)., Published as part of Love, Milton S., Bizzarro, Joseph J., Cornthwaite, Maria, Frable, Benjamin W. & Maslenikov, Katherine P., 2021, Checklist of marine and estuarine fishes from the Alaska-Yukon Border, Beaufort Sea, to Cabo San Lucas, Mexico, pp. 1-285 in Zootaxa 5053 (1) on page 43, DOI: 10.11646/zootaxa.5053.1.1, http://zenodo.org/record/5578008, {"references":["Jimenez Prado, P. & Bearez, P. (2004) Peces marinos del Ecuador continental / marine fishes of continental Ecuador. SIMBIOE / NAZCA / IFEA TOMO II, Quito.","Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. & Niem, V. H. (1995) Guia FAO para la identificacion para los fines de la pesca. Pacifico centro-oriental. Volume II, Vertebrados, Parte 1. Volume III, Vertebrados, Parte 2. FAO, Rome.","Allen, G. R. & Robertson, D. R. (1994) Fishes of the Tropical Eastern Pacific. University of Hawaii Press, Honolulu.","Amezcua Linares, F. (1996) Peces Demersales de la Platforma Continental del Pacifico Central de Mexico. Instituto de Ciencias del Mar y Limnologica, Universidad Nacional Autonoma de Mexico."]}

Stock assessment is necessary to understand the status of fishery stocks. However, for the data-poor fishery, it is very challenging to assess the stock status. The length-based Bayesian biomass (LBB) technique is one of the most powerful methods to assess the data-poor fisheries resources that need simple length frequency (LF) data. Addressing the present gap, this study aimed to assess the stock status of three sardines (Sardinella fimbriata, Dussumieria acuta, and D. elopsoides) in the Bay of Bengal (BoB), Bangladesh using the LBB method. The estimated relative biomass for S. fimbriata was B/B0 &lt, BMSY/B0, indicating the overfished biomass, while the assessed B/B0 &gt, BMSY/B0 for D. acuta and D. elopsoides indicates healthy biomass. Additionally, for S. fimbriata, the length at first landing was smaller than the optimum length at first landing (Lc &lt, Lc_opt), indicating an overfishing status, but a safe fishing status was assessed for D. acuta and D. elopsoides (Lc &gt, Lc_opt). Therefore, increasing the mesh size of fishing gears may help to ensure the long-term viability of sardine populations in the BoB, Bangladesh.

The complete mitogenome of Sardinella zunasi was determined by next-generation sequencing. The S. zunasi mitogenome was a circular 16,307 bp molecule that contained 13 protein-coding genes, 22 tRNA genes, 2 rRNA genes, and one control region (D-loop). The gene arrangement was consistent with other Sardinella mitogenomes. The phylogenetic relationships of 29 Clupeoidei species based on 13 protein-coding genes from the available mitogenomes were analyzed. Sardinella zunasi clustered with Sardinella among Clupeidae, suggesting a closer relationship with this genus. These results will be useful for understanding the phylogenetic relationships, taxonomic classification, and phylogeography of the genus Sardinella relative to other genera of Clupeoidei.

In this study, the complete mitochondrial genome of Ilisha striatula is presented, and we also discussed its mitochondrial characteristics. The full length of the mitochondrial genome was 16,847 bp...

Hata, Harutaka, Motomura, Hiroyuki (2021): First Japanese Specimen-based Records of Sardinella gibbosa (Teleostei: Clupeiformes: Clupeidae) from Okinawa Island. Species Diversity 26: 197-204, DOI: 10.12782/specdiv.26.197

Despite being one of the richest Neogene deposits of marine fossil vertebrates along the Pacific coast of South America, no detailed study of bony fishes from the Bahia Inglesa Formation has been carried out. Through the revision of historical collections of the Caldera Paleontological Museum, we describe 27 fossil remains of bony fishes. Fossils described here are distributed in 13 taxa. Among these, Sardinops sp. cf. S. sagax (Clupeidae) represents the first fossil record of the genus for Chile, Labrodon sp. (Labridae) and Stelliferinae (Sciaenidae) are the first respective records for the southeast Pacific Ocean, Sarda sp. (Scombridae) represents the first fossil record for South America, Semicossyphus sp. (Labridae) and Anisotremus sp. (Haemulidae) are the first respective records for the Southern Hemisphere. Gymnosarda sp. (Scombridae) is described for the first time for the Neogene. We performed a paleobathymetric analysis of fossil fishes, including elasmobranchs, to understand the paleoenvironment followed by a similarity analysis that compares the faunal composition of the Bahia Inglesa Fm. with other fossil assemblages for the Pacific of South America. Our results show that the fish fauna from the Bahia Inglesa Fm. likely represents a depth range of 98 to 382 m with a mean depth range of 252 m in an uppermost bathyal environment. The assemblage from the Bahia Inglesa Fm. shows low taxonomic similarity levels with other Neogene geological units from the southeast Pacific Ocean. The study represents the first paleoecological and paleobiogeographic analysis of the fossil fish fauna from the Southeast Pacific. Key words. Atacama region. Clupeiformes. Eastern Pacific. Paleoenviroments. Perciformes. Ophidiiformes.

The new anchovy Stolephorus grandis n. sp., described on the basis of 10 specimens collected from Papua, Indonesia, and Australia, closely resembles Stolephorus mercurius Hata, Lavoué & Motomura, 2021, Stolephorus multibranchus Wongratana, 1987, and Stolephorus rex Jordan & Seale, 1926, all having double pigmented lines on the dorsum from the occiput to the dorsal-fin origin, a long maxilla (posterior tip just reaching or slightly beyond the posterior margin of preopercle), and lacking a predorsal scute. However, the new species clearly differs from the others in having fewer gill rakers (35–39 total gill rakers on the first gill arch in S. grandis vs. > 38 in the other species), a greater number of vertebrae (total vertebrae 42–43 vs. fewer than 41), longer caudal peduncle (21.9–23.7% SL vs. < 20.8%), and the depressed pelvic fin not reaching posteriorly to vertical through the dorsal fin-origin (vs. reaching beyond level of dorsal-fin origin).

Stolephorus grandis n. sp. [New English name: Papuan Double-lined Anchovy] Figure 1; Tables 1���3 urn:lsid:zoobank.org:pub: 8C56370C-17C9-483A-973E-777C6553BB93 Holotype. RMNH. PISC. 24670, 102.9 mm SL, Merauke, Papua, Indonesia, 5 Apr. 1955. Paratypes. 9 specimens, 59.2���116.1 mm SL: AMS-I. 22777-006, 59.2 mm SL, south end of Ellis Beach, north of Queensland, Cairns, Australia (16��43���48���S, 145��39���00���E); RMNH. PISC. 84311, 116.1 mm SL, collected with the holotype; KAUM ���I. 156158, 82.6 mm SL, NSMT-P 140585, 80.5 mm SL, NTM S. 10071-001, 5 specimens, 77.7���88.0 mm SL, Van Diemen Gulf, Northern Territory, Australia. Diagnosis. A species of Stolephorus with the following combination of characters: maxilla rather short, 16.7��� 18.2% SL (mean 17.5%), its posterior tip slightly short of, or reaching to, or slightly beyond posterior margin of preopercle; mandible short, 14.3���15.3% SL (14.7%); posterior margin of preopercle convexly rounded (not indented); no predorsal scute; prepelvic scutes 3���6 (modally 4); pelvic scute without spine; gill rakers 14���16 (16) in upper series on first gill arch, 21���23 (23) in lower series, 35���39 (39) in total; gill rakers 10���12 (11) in upper series on second gill arch, 18���20 (20) in lower series, 28���32 (30) in total; gill rakers 8���9 (9) in upper series on third gill arch, 10���12 (11) in lower series, 18���21 (20) in total; gill rakers 7 or 8 (8) in upper series on fourth gill arch, 9���10 (9) in lower, 16���18 (17) in total; gill rakers 3���5 (5) on posterior face of third gill arch; transverse scales 8; pseudobranchial filaments 20���27 (22); paired dark patches on parietal and occipital regions; double dark lines on dorsum from occipital region to dorsal-fin origin but absent behind dorsal-fin base; no black spots on suborbital area and tip of lower jaw; head rather short, 22.0���24.0% SL (mean 23.0%), postorbital length 11.5���12.4% SL (11.9%); caudal peduncle long, 21.9���23.7% SL (22.8%); pelvic fin relatively short, 8.3���9.0% SL (8.6%), depressed pelvic fin not reaching posteriorly to vertical through dorsal-fin origin; distance from snout to pectoral-fin insertion 25.0���27.0% SL (25.6%); distance from snout to pelvic-fin insertion 42.8���44.4% (43.3%); dorsal-fin base short, 12.6���14.8% SL (13.9%); anal-fin base short, 16.0���17.8% SL (16.9%); third dorsal-fin ray short, 14.8���15.7% SL (15.3%); third analfin ray short, 11.9���13.8% (12.5%); vertebral count 21 or 22 (modally 21) + 21 or 22 (21) = 42 or 43 (42). Description. Data for holotype presented first, followed by data for non-type specimens in parentheses (if different). Counts and measurements, expressed as percentages of SL or HL, given in Tables 1 and 2. Body laterally compressed, elongate, deepest at dorsal-fin origin. Dorsal profile of head and body slightly convex from snout tip to dorsal-fin origin, gently lowering to uppermost point of caudal-fin base. Ventral profile of head and body slightly convex from lower-jaw tip to pelvic-fin insertion, thereafter slowly rising to lowermost point of caudal-fin base. Abdomen somewhat rounded, covered with four (three to six) spine-like scutes. Pelvic scute without spine. Postpelvic and predorsal scutes absent. Anus just anterior to anal-fin origin. Snout tip rounded; snout length less than eye diameter. Mouth large, inferior, ventral to body axis, extending backward beyond posterior margin of eye. Maxilla rather long, its posterior tip pointed, just reaching (short of or slightly beyond) to posterior margin of preopercle. Lower jaw slender. Single row of conical teeth on each jaw and palatines. Several distinct conical teeth on vomer. Teeth patch on pterygoid. A few fine teeth on upper edge of hyoid. Several rows of conical teeth on upper edges of basihyal and basibranchial. Eye large, round, covered with adipose eyelid, positioned laterally on head dorsal to horizontal through pectoral-fin insertion, visible in dorsal view. Pupil round. Orbit elliptical. Nostrils close to each other, anterior to orbit. Posterior margin of preopercle convexly rounded (not indented). Subopercle and opercle with smoothly rounded posterior margins. Gill membrane without serrations. Interorbital space flat, width less than eye diameter. Pseudobranchial filaments present, length of longest filament shorter than eye diameter. Gill rakers long, slender, rough, visible from side of head when mouth opened. Single row of asperities on anterior surface of gill rakers. Isthmus muscle long, reaching anteriorly to posterior margin of gill membranes. Urohyal hidden by isthmus muscle (not visible without dissection). Gill membrane on each side joined distally, most of isthmus muscle exposed (not covered by gill membrane). Scales cycloid, thin, deciduous, except for prepelvic scutes. Head scales absent. Lateral line absent. Fins scaleless, except for broad triangular sheath of scales on caudal fin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray, slightly posterior to middle of body. Dorsal and anal fins with three anteriormost rays unbranched and closely spaced. First dorsal- and anal-fin rays minute. Anal-fin origin just below base of twelfth (ninth to twelfth) dorsal-fin ray. Posterior tip of depressed anal fin not reaching caudal-fin base. Uppermost pectoral-fin ray unbranched, inserted below midline of body. Posterior tip of pectoral fin not reaching to pelvic-fin insertion. Dorsal, ventral, and posterior margins of pectoral fin nearly linear. Pelvic fin shorter than pectoral fin, insertion anterior to vertical through dorsal-fin origin. Posterior tip of depressed pelvic fin not reaching to vertical through dorsal-fin origin. Caudal fin forked, posterior tips pointed. Abbreviations: D���P1 (distance between dorsal-fin origin and pectoral-fin insertion); D���P2 (distance between dorsal-fin origin and pelvic-fin insertion); D���A (distance between origins of dorsal and anal fins); P1���P2 (distance between insertions of pectoral and pelvic fins); P2���A (distance between pelvic-fin insertion and anal-fin origin). Coloration of preserved specimens. Body uniformly pale. Silver longitudinal band running from just behind upper opercular margin to caudal-fin base. Cheek and opercle silver. Scattered melanophores on snout tip, absent on suborbital area. Paired dark patches on parietal and occipital regions. Double dark lines on dorsum from occipital area to dorsal-fin origin. Scale pockets on dorsum with black posterior margin. All fins whitish, semi-transparent. Melanophores distributed along fin rays of dorsal and caudal fins. Caudal fin with black posterior margin. Distribution. Currently known only from the southern coast off Merauke, Papua, Indonesia to the northern coast of Australia (from Van Diemen Gulf, Northern Territory to Cairns, Queensland) (Fig. 2). Etymology. The specific name ���grandis��� refers to the larger body of the species, compared with similar species. Remarks. The new species is assignable to the genus Stolephorus, as defined by Whitehead et al. (1988) and Wongratana et al. (1999), in having a long isthmus muscle reaching anteriorly to the posterior margin of the gill membrane, the urohyal covered by the isthmus muscle, and prepelvic scutes, in addition to the absence of postpelvic scutes. Stolephorus grandis is easily distinguishable from all congeners, except for Stolephorus mercurius Hata, Lavou�� & Motomura, 2021, Stolephorus multibranchus Wongratana, 1987, and Stolephorus rex Jordan & Seale, 1926, in having double pigmented lines on the dorsum from the occipital area to the dorsal-fin origin, a long maxilla (posterior tip just reaching or slightly beyond the preopercle posterior margin), and lacking a predorsal scute (Whitehead et al., 1988; Wongratana et al., 1999; Kimura et al., 2009; Hata & Motomura, 2018a ���d, 2021; Hata et al., 2019, 2020a, b, 2021; Gangan et al., 2020). Moreover, the new species differs from the latter three species in having fewer gill rakers [total gill rakers on first, second, third, and fourth gill arch 35���39, 28���32, 18���21, and 16���18, respectively in S. grandis vs. more than 38, 31, 20, and 15 (more than 18 in S. mercurius and S. multibranchus), respectively in the other three; Table 1; Fig. 3], more total vertebrae (42 or 43 vs. 40 or fewer; Tables 1, 3), a longer caudal peduncle (21.9���23.7% SL vs. shorter than 20.8%), and the depressed pelvic fin not reaching posteriorly to vertical through the dorsal-fin origin (vs. reaching posteriorly beyond vertical through dorsal-fin origin). Furthermore, the relative length of the pelvic fin separates S. grandis, S. mercurius, and S. multibranchus (pelvic-fin lengths 8.3���9.0% SL, 9.1���11.1%, and 10.0���11.9%, respectively). Lengths of both jaws and the head also separate S. grandis, S. multibranchus, and S. rex (lengths of maxilla, mandible, and head 16.7���18.2% SL, 14.3���15.3% SL, and 22.0���24.0% SL, respectively, in S. grandis vs. 18.0���21.1%, 16.0���18.5%, and 24.8���26.7%, respectively, in S. multibranchus, and 19.1���23.1%, 16.1���18.3%, and 24.3���27.6%, respectively, in S. rex). Additionally, S. grandis is further distinguished from S. rex by its shorter postorbital length (11.5���12.4% SL in S. grandis vs. 12.6���14.9% in S. rex) (Hata & Motomura, 2018c; Hata et al., 2021; this study), and from S. multibranchus by its shorter dorsal- (12.6���14.8% SL vs. 14.0���16.1%) and anal-fin bases (16.0���17.8% SL vs. 18.3���21.3%) (Table 2; Fig. 4; Hata & Motomura, 2018c; Hata et al., 2021; this study). Additionally, the other three species (S. mercurius, S. multibranchus, and S. rex) have never been recorded from New Guinea and Australia, where the new species occurs. Comparative material examined. Stolephorus mercurius (28 specimens, 62.3���103.6 mm SL) and Stolephorus rex (45 specimens, 57.3���104.4 mm SL): listed in Hata et al. (2021). Stolephorus multibranchus (17 specimens, 45.9���71.2 mm SL): listed in Hata & Motomura (2018c)., Published as part of Hata, Harutaka & Motomura, Hiroyuki, 2021, Stolephorus grandis, a new anchovy (Teleostei: Clupeiformes: Engraulidae) from New Guinea and Australia, pp. 481-489 in Zootaxa 5004 (3) on pages 482-488, DOI: 10.11646/zootaxa.5004.3.5, http://zenodo.org/record/5120816, {"references":["Whitehead, P. J. P., Nelson, G. J. & Wongratana, T. (1988) FAO species catalogue Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolfherrings. Part 2 - Engraulididae. FAO Fisheries Synopsis, No. 125, 7 (2), i - viii + 305 - 579.","Wongratana, T., Munroe, T. A. & Nizinski, M. S. (1999) Order Clupeiformes. Engraulidae. Anchovies. In: Carpenter, K. E. & Niem, V. H. (Eds.), FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 3. Batoid Fishes, Chimaeras and Bony Fishes Part 1 (Elopidae to Linophrynidae). Food and Agriculture Organization of the United Nations, Rome, pp. 1698 - 1753.","Hata, H., Lavoue, S. & Motomura, H. (2021) Taxonomic status of nominal species of the anchovy genus Stolephorus previously regarded as synonyms of Stolephorus commersonnii Lacepede 1803 and Stolephorus indicus (van Hasselt 1823), and descriptions of three new species (Clupeiformes: Engraulidae). Ichthyological Research, 68, 327 - 372. https: // doi. org / 10.1007 / s 10228 - 020 - 00792 - 0","Wongratana, T. (1987) Two new species of anchovies of the genus Stolephorus (Engraulidae), with a key to species of Engraulis, Encrasicholina, and Stolephorus. American Museum Novitates, 2876, 1 - 8.","Jordan, D. S. & Seale, A. (1926) Review of the Engraulidae, with descriptions of new and rare species. Bulletin of the Museum of Comparative Zoology, 67 (11), 355 - 418.","Kimura, S., Hori, K. & Shibukawa, K. (2009) A new anchovy, Stolephorus teguhi (Clupeiformes: Engraulidae), from North Sulawesi, Indonesia. Ichthyological Research, 56 (3), 292 - 295. https: // doi. org / 10.1007 / s 10228 - 009 - 0103 - 4","Hata, H. & Motomura, H. (2018 a) Stolephorus continentalis, a new anchovy from the northwestern South China Sea, and redescription of Stolephorus chinensis (Gunther 1880) (Clupeiformes: Engraulidae). Ichthyological Research, 65 (3), 374 - 382. https: // doi. org / 10.1007 / s 10228 - 018 - 0621 - z","Hata, H., Lavoue, S. & Motomura, H. (2019) Taxonomic status of seven nominal species of the anchovy genus Stolephorus described by Delsman (1931), Hardenberg (1933), and Dutt and Babu Rao (1959), with redescriptions of Stolephorus tri (Bleeker 1852) and Stolephorus waitei Jordan and Seale 1926 (Clupeiformes: Engraulidae). Ichthyological Research, 67 (1), 7 - 38. https: // doi. org / 10.1007 / s 10228 - 019 - 00697 - 7","Hata, H., Lavoue, S. & Motomura, H. (2020 a) Stolephorus babarani, a new species of anchovy (Teleostei: Clupeiformes: Engraulidae) from Panay Island, central Philippines. Zootaxa, 4718 (4), 509 - 520. https: // doi. org / 10.11646 / zootaxa. 4718.4.5","Gangan, S. S., Pavan-Kumar, A., Jahageerdar, S. & Jaiswar, A. K. (2020) A new species of Stolephorus (Clupeiformes: Engraulidae) from the Bay of Bengal, India. Zootaxa, 4743 (4), 561 - 574. https: // doi. org / 10.11646 / zootaxa. 4743.4.6","Hata, H. & Motomura, H. (2018 c) Additional specimens of the poorly known anchovy Stolephorus multibranchus (Clupeiformes: Engraulidae) from Kosrae, Caroline Islands. Biogeography, 20, 78 - 84. https: // doi. org / 10.11358 / biogeo. 20.78"]}

The Dotted Gizzard Shad (Konosirus punctatus) was one of the most important commercial fish species in China, Japan and Korea. In this study, the complete mitochondrial genome of K. punctatus was presented. The full length of the mitochondrial genome was 16,705 bp, including 13 protein-coding genes (PCGs), two ribosomal RNAs, 22 transfer RNA genes, one non-coding control region (CR) and one origin of replication on the light-strand. The total nucleotide composition of mitochondrial DNA was 25.79%A, 25.09%T, 29.05%C, 20.08%G, and AT was 50.88%. The mitochondrial genome provides an important resource for solving taxonomic problems and studying molecular evolution.

Sardinella dayi Regan 1917, the taxonomic status of which has long been uncertain, is redescribed on the basis of 27 specimens, including the holotype. Although Whitehead (1985) regarded the nominal species as a junior synonym of Sardinella jussieui (Valenciennes 1847), the former differs from S. jussieui in having higher counts of gill rakers on the first gill arch (59–77 + 92–118 = 155–190 vs. 49–60 + 88–106 = 137–158 in S. jussieui), a longer head (26.3–29.6% SL vs. 22.7–24.9%), upper jaw (10.9–12.4% SL vs. 9.6–10.7%), and lower jaw (11.5–13.1% SL vs. 10.3–11.7%), a shorter anal-fin base (13.5–16.2% SL vs. 16.7–18.5%), and body scales with numerous pores (pores absent in S. jussieui). Redescriptions of both species are provided.

Sardine is the common name for several small-sized pelagic species from Clupeiformes, representing a resource of great importance in the global fishery. Great efforts have been made to utilise these species as dried, smoked, and restructured fish products. However, in most of these products, it is quite challenging to identify the individual species as the external features are lost during processing, paving the way for species mislabelling. In this study, DNA barcoding (max, using about 650 bp, described as FDB; mini, of about 192 bp, described as MDB) was used for species identification of 139 specimens taken from 48 sardine products (canned and dried seasoning) randomly collected from local markets in Nanjing, China. Moreover, species specific primers were designed for Sardina pilchardus, with the aim to screen the species of S. pilchardus in mixed products. Results highlighted a success rate of amplification from 38.1% for FDB to 97.9% for MDB. Only one sample failed the Sanger-sequencing, and species-specific real time PCR confirmed the existence of S. pilchardus in the product. A maximum species identity in the range of 98-100% was obtained for all readable sequences and 11 species/genera were identified, belonging to 5 orders (Scorpaeniformes, Perciformes, Clupeiformes, Aulopiformes, Scombriformes). Significant legislative and managerial shortcomings and incentives to facilitate the market access of certain species, together with public indifference, represent the main reasons for fish fraud in China.

This study presents an extensive review of published and unpublished occurrence records of fish species in the Rio Negro drainage system within the Brazilian territory. The data was gathered from two main sources: 1) litterature compilations of species occurrence records, including original descriptions and revisionary studies; and 2) specimens verification at the INPA fish collection. The results reveal a rich and diversified ichthyofauna, with 1,165 species distributed in 17 orders (+ two incertae sedis), 56 families, and 389 genera. A large portion of the fish fauna (54.3% of the species) is composed of small-sized fishes < 10 cm in standard length. The main groups are Characiformes (454 species; 39.0%), Siluriformes (416; 35.7%), Gymnotiformes (105; 9.0%), and Cichliformes (102; 8.8%). The species composition differs between the main aquatic environments, such as: main channel (159 species), lakes (296), tributary rivers (596), small streams (234), seasonal beaches (186), and rapids (41). Part of the ichthyofauna is shared with adjacent basins, such as the Orinoco, rivers of the Guiana Shield, lower Solimões/Amazonas and upper Amazonas, which contributes to the remarkable ichthyofaunal diversity of the basin. A high rate of species endemism was observed in Characidae (24), Loricariidae (18), Cichlidae (18) and Callichthyidae (18), totalling 156 species (13.4%) endemic to the basin. An estimation of the species richness for the Rio Negro basin, considering 23 published references, resulted in 1,466 and 1,759 species (Jackknife 1 and 2, respectively), which seems reasonable when considering the large number of morphotypes left out of the present list and the low sampling effort in many areas of the basin. The results presented herein provide an additional tool for environmental managers and decision makers for conservation purposes of one of the richest and most well-preserved sub-basins of the Rio Amazonas system.

Thrissina cultella sp. nov., and Thrissina serena sp. nov., are described from the Bay of Bengal and northwestern Indian Ocean, respectively. Although both species resemble T. vitrirostris (Gilchrist and Thompson 1908), which is redescribed from both type and non-type specimens, in having a long upper jaw (posterior tip reaching to pectoral-fin insertion) and similar numbers of gill rakers and ventral scutes, the two new species differ in having fewer transverse scales (9–10 in both vs. 11–12 in T. vitrirostris). Thrissina cultella differs from T. serena in having a slender body [24.4–26.9% of standard length (SL) vs. 26.6–29.4% SL in T. serena], shorter head (25.2–27.1% SL vs. 26.2–27.3%) and pectoral fin (17.5–19.1% SL vs. 19.5–21.3%), and longer pelvic fin (9.3–10.5% SL vs. 7.8–8.7%).

Sardinella tawilis, the only known freshwater sardinella in the world, is endemic to Taal Lake, Philippines. Previous studies found the Taiwan sardinella, S. hualiensis, to be morphologically very similar to S. tawilis and identified it as the marine sister species of S. tawilis. In this study, DNA barcoding using the mitochondrial cytochrome c oxidase I (COI) gene was carried out to analyze species demarcation in the Sardinella genus, focusing primarily on the relationship between S. tawilis and S. hualiensis. The neighbour-joining (NJ) tree that was constructed using Kimura 2-parameter (K2P) model showed a single clade for the two species with 100% bootstrap support. K2P interspecific genetic divergence ranged from 0% to 0.522%, which is clearly below the suggested 3–3.5% cutoff for species discrimination. Recombination activating gene 1 (RAG1), mitochondrial control region (CR), cytochrome b, 16S rRNA, and S7 markers were used to further validate the results. Sardinella tawilis and S. hualiensis clustered together with a bootstrap support of 99–100% in each of the NJ trees. Low interspecific genetic distances between S. tawilis and S. hualiensis for all the markers except CR could be attributed to incipient allopatric speciation.

The occurrence of a Lessepsian migrant, the Golani round herring, Etrumeus golanii DiBattista, Randall et Bowen, 2012, is reported here for the first time from the Alboran Sea (south-west Mediterranean). On May 2018, twenty individuals were caught by a commercial purse-seiner off Fnideq (Morocco), but five females and two males, were kept by the fisherman for further analyses. This new finding confirms the expansion of this Lessepsian migrant throughout the Mediterranean Sea.

A new sardine, Sardinellapacificasp. n., is described on the basis of 21 specimens collected from the Philippines. The new species closely resembles Sardinellafimbriata (Valenciennes, 1847), both species having lateral scales with centrally discontinuous striae, a dark spot on the dorsal-fin origin, more than 70 lower gill rakers on the first gill arch, the pelvic fin with eight rays, and 17 or 18 prepelvic and 12 or 13 postpelvic scutes. However, the new species is distinguished from the latter by lower counts of lateral scales, pseudobranchial filaments, and postpelvic scutes (38–41, 14–19 and 12–13, respectively vs. 44–46, 19–22 and 13–14), and a shorter lower jaw (10.4–11.6% of standard length vs. 11.1–12.2%). Sardinellapacificasp. n. is known only from the Philippines, whereas S.fimbriata is restricted to the Indian Ocean, although previously considered to be an Indo-West Pacific species, distributed from India to the Philippines.

The basic biology and ecology of the South African east coast round herring Etrumeus wongratanai was investigated from samples of fish collected between 2013 and 2016. This species is short-lived and reaches a maximum of 3 years of age, with rapid growth in its first year of life. It reproduces from June to December (austral summer) and condition factor was lowest in May through to August and increased from September, probably reflecting the physiological strain before and during spawning. Fish larvae were the most important food items consumed during summer, whereas eucalanid copepods were the most important prey at other times of the year. Stable-isotope data suggest that there are gradual changes in the trophic level with increasing fish size, δ15 N and δ13 C values also differed between seasons. The results obtained here are compared with those of other Etrumeus species, regionally and globally.

Three species of digenetic trematodes of the Family Hemiuridae Looss, 1899, Subfamily Aphanurinae Skrjabin & Guschanskaja, 1954 with their prevalence, intensity of infection and mean abundance are reported Aphanuroides lethrini, Aphanurus stossichii and Aphanurus tuberculatus. A total of 26 specimens of Thryssa malabarica (Bloch, 1795) were examined for digenean infection. Prevalence of infection of A. lethrini, A. stossichii and A. tuberculatus were 11.53%, 69.23% and 15.38%, respectively, the intensity of infection were 1.6, 3.1 and 2.5 and the mean abundance were 0.19, 2.15 and 0.38, respectively. The prevalence, intensity and mean abundance of infection were at the maximum level in A. stossichii infection. Recovery of these parasites from T. malabarica represents a new host record and the first report from Kerala. The characteristic features of the recovered digeneans are discussed.