Search

2. Sisyromyia angustivitta sp. nov. (Figs 2, 3) Type specimens. Holotype ♂ Australia, WA, Albany, 5 Oct 1951 IFB Common. Paratype ♂ Australia, WA, same data as holotype. Other specimen examined. Australia, WA, ♀ Stirling Rang Drive, Stirling Range Nat[ional] P[ar]k, 23 Sep 1981, ID Naumann & JC Cardale (WAM). Diagnosis. Large-sized fly, body with mostly reddish hairs. Wing membrane brown. Abdominal tergites 2–7 with an inconspicuous narrow medial stripe consisting of sparse, decumbent short pale yellow scales. Description. Male. Body length 11.9–12.0 mm, wing length 12.1–12.2 mm. Head. Head about 2.3 x wider than long, mostly blackish with thick pale pruinescence and covered in admixed yellow to golden scales and brown to black hairs. Eyes holoptic. Frons short, 1.6 x length of ocellar tubercle, upper narrow and black; lower half triangular, as long as upper half, with sparse pale pruinescence and golden scales admixed with black hairs. Ocellar tubercle slightly raised, blackish brown to black with grey pruinescence, with black hairs admixed with few golden scales. Face with thick pale pruinescence and golden scales admixed with long black hairs, parafacial area without black hairs. Gena with thick pale pruinescence and long yellow scales. Clypeus with thick pale pruinescence and otherwise bare. Occiput with thick pale pruinescence and golden to reddish scales admixed with some black hairs. Posterior eye margin slightly convex. Antennal scape and pedicel black with sparse pale pruinescence, scape with long black hairs admixed with long golden scales, pedicel with short black hairs; flagellum black without pruinescence, subapex with 2–3 long hairs. Scape 2.4 x as long as wide, and 3.0 x as long as pedicel, uniform from base to apex. Pedicel as long as wide. Flagellum 12.0 x as long as wide, 2.3 x as long as scape + pedicel, 3.1 x as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus, and with one long subapical hairs (Fig. 2c). Palpus long, just extending beyond oral cavity, yellow except apex brown, with pale yellow hairs except brown hairs apically, one-segmented, without palpal pit. Mouthparts slender, 3.9 x as long as eye length, 2.0 x as long as head length, labellum thin and filiform (Fig. 2h). Thorax. Integumental colour of scutum mostly black with thick pale pruinescence. Scutum covered with short golden to reddish hairs admixed with few black hairs, some golden to reddish hairs turning brown apically. Scutellum brown with sparse pale pruinescence, with golden to reddish hairs admixed with few black hairs. Pleura black with thick pale pruinescence, mostly covered in long pale yellow hairs, except anepisternum with golden hairs admixed with few reddish hairs, anepimeron, meron, laterotergite and mediotergite bare. Legs. Legs dark yellow. Femora with dense long yellow hairs and scales, admixed with few black hairs ventrally. Fore and mid femora with few anterior bristle on apical half; hind femur with one row of anteroventral bristles, and some separate bristles. Bristles and other hairs on legs brown. Fore tibia 2.4 x longer than fore basitarsus, mid tibia 2.7 x longer than mid basitarsus, hind tibia 2.7 x longer than hind basitarsus. Wings. Wing membrane uniformly infuscated. Cell r 5 open, M 1 close to R 5 ; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m nearly as long as crossvein r-m; cell cup open (Fig. 2d). Haltere stem and knob yellow. Abdomen. Integumental colour of tergites black mostly with sparse pale pruinescence. Tergite 1 with thick pale pruinescence and dense golden to reddish hairs; tergites 2–7 with long golden to reddish hairs, hairs near posterior margin of tergites turning reddish on apical half; tergites 2–7 with thin median stripe consisting of sparse, decumbent short pale yellow scales. Sternites black except posterior margin yellow, with thick pale pruinescence, covered with yellow hairs. Genitalia. Epandrium anterior margin slightly concave, posterior margin slightly convex (Fig. 3d). Lateral margin of gonocoxite strongly curved (Figs 3a,b); lateral ejaculatory process narrow; inner apex of gonocoxite long and sharp, outer apex of gonocoxite long and rounded; gap between inner apices of gonocoxites medium; phallus shorter than gonocoxite (Fig. 3c). Female. Body length 12.0 mm, wing length 12.1 mm. Very similar to male, except frons black with sparse pale pruinescence, 3.0 x as wide as ocellar tubercle, frons with short golden scales admixed with long black hairs (Fig. 2l). Abdominal tergites with reddish scales and hairs, admixed with black hairs; median stripe wider than male. Around 30 acanthophorite spines present on each side of tergite 9+10 (Fig. 3g). Remarks. Sisyromyia angustivitta sp. nov. is similar to Sisyromyia rutila, but the stripe on abdominal tergites 2–7 is narrow and inconspicuous, and the outer apex of gonocoxite is long. Distribution. Australia (WA). Etymology. The specific name refers to the inconspicuous narrow medial stripe on the abdominal tergites.

9. Sisyromyia thomsoni Evenhuis & Greathead, 1999 (Figs 20, 21, 29a) Bombylius scutellaris Thomson, 1869: 488. Type-locality: Australia (NSW); holotype, NHRS. [Preoccupied by Wiedemann, 1828.] Sisyromyia thomsoni Evenhuis & Greathead, 1999: 170 (new replacement name for Bombylius scutellaris Thomson, 1869). Other specimen examined. Australia, ACT, ♀ Gibraltar Creek, 18 Oct 1967, Z Liepa. NSW, ♂ Killara, 6 Aug 1934, D Waterhouse. ♂ Narrabeen, 10 Aug 1935, MF Day. ♂ Mungo, Myall Lakes, 23 Aug 1934, D Waterhouse. ♀ (2) Medlow, 4 Oct 1926. ♂ Sydney, Feb 1952. Qld., ♀ -28.828 151.978, North slopes, Girraween NP, 14 Aug 2009, MG Jefferies (QM). Diagnosis. Medium-sized fly, body with mostly brownish hairs. Wing membrane dark brown on anterior half and clear on posterior half. Male eyes holoptic. Gap between inner apices of gonocoxites narrow. Inner apex of gonocoxite short. Redescription. Male. Body length 10.3–11.5 mm, wing length 9.9–10.4 mm. Head. Head about 2.2 x wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to black hairs and scales. Eyes holoptic. Frons short, 1.8 x length of ocellar tubercle, upper narrow and black; lower half triangular, 0.8 x length of upper half, with thick pale pruinescence and pale yellow to brown scales admixed with few brown hairs. Ocellar tubercle slightly raised, blackish brown to black with thick grey pruinescence, with long black hairs. Face with thick pale pruinescence and golden scales admixed with some long black hairs; parafacial area with long white scales admixed with long yellow hairs. Gena with thick pale pruinescence and long white to yellow hairs. Clypeus with thick pale pruinescence and otherwise bare. Occiput with thick pale pruinescence and white to brown scales. Posterior eye margin slightly convex. Antennal scape and pedicel brown with thick pale pruinescence, scape with long black hairs admixed with few golden hairs laterally, pedicel with short black hairs; flagellum black without pruinescence, subapex with 3–5 long hairs. Scape 2.9 x as long as wide, and 2.5 x as long as pedicel, uniform from base to apex. Pedicel 1.3 x as long as wide. Flagellum 8.0 x as long as wide, 1.6 x as long as scape + pedicel, 2.4 x as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus (Fig. 20c). Palpus long, just extending beyond oral cavity, dark yellow with pale yellow hairs, except apex with black hairs, one-segmented, without palpal pit. Mouthparts slender, 3.1 x as long as eye length, 2.1 x as long as head length, labellum thin and filiform (Fig. 20h). Thorax. Integumental colour of scutum mostly black with thick brown pruinescence except two median wide white pruinescent stripes, postalar callus brown with sparse pale pruinescence. Scutum covered with short pale yellow to brown hairs, admixed with few short black hairs, lateral stripe consisting of short white to pale yellow scales, and posteriorly with few short white scales. Six brownish yellow notopleural setae present. Scutellum black with sparse pale pruinescence, anterior margin with an indistinct band consisting of short white scales, posterior margin with long brown hairs admixed with some long black hairs. Pleura black with thick pale pruinescence, mostly covered in long pale yellow hairs, except dorsal half of anepisternum with brown hairs, anepimeron, meron, laterotergite and mediotergite bare. Legs. Legs mostly brown, except tibiae yellow. Femora with long brown hairs. Mid femur with one anterior bristle on apical half; hind femur with one row of anteroventral bristles and one row of posteroventral bristles, more separate bristles on apical half. Bristles and other hairs on legs brown. Fore tibia 2.0 x longer than fore basitarsus, mid tibia 2.3 x longer than mid basitarsus, hind tibia 2.2 x longer than hind basitarsus. Wings. Wing membrane dark brown on anterior half and clear on posterior half, darker towards wing base. Cell r 5 open, M 1 close to R 5 ; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m nearly as long as crossvein r-m; cell cup open (Fig. 20d). Haltere stem brown, and knob yellow. Abdomen. Integumental colour of tergites black mostly with sparse pale pruinescence. Tergite 1 with thick pale pruinescence and dense brown hairs; tergites 2–7 with yellow to brown hairs, more pale hairs on anterior half, posterior half admixed with some black hairs, hairs darker and denser laterally, and with median stripe consisting of dense, decumbent short white scales; tergites 3–7 with pale yellow to white scales on lateral side, best viewed from posterodorsal side. Sternites black except posterior margin yellow, with thick pale pruinescence, covered with pale yellow to brown hairs, hairs darker laterally. Genitalia. Epandrium anterior margin slightly concave, posterior margin slightly convex (Fig. 21d). Lateral margin of gonocoxite nearly straight (Figs 21 ab); lateral ejaculatory process wide; inner apex of gonocoxite short and acute, outer apex of gonocoxite long and sharp; gap between inner apices of gonocoxites narrow; phallus shorter than gonocoxite (Fig. 21c). Female. Body length 7.4–11.1 mm, wing length 9.1–12.2 mm. Very similar to male, except frons black with thick pale pruinescence, 3.4 x as wide as ocellar tubercle, frons with short golden scales admixed with long black hairs (Fig. 20l). Body hairs paler than male. Around 30 acanthophorite spines present on each side of tergite 9+10 (Fig. 21g). Remarks. Bombylius scutellaris was first considered a synonym of Sisyromyia decorata by Roberts (1928: 429). However, this synonymy was not followed by Evenhuis & Greathead (1999). We compared the specimens from NSW and WA, and agree with Evenhuis & Greathead’s decision (also see ‘4. Sisyromyia decorata (Walker, 1849) ’ earlier). Therefore, Sisyromyia thomsoni is recognised as a valid species in the present revision. Sisyromyia thomsoni is similar to Sisyromyia decorata, but differs as follows: male eyes holoptic; gap between inner apices of gonocoxites narrow; inner apex of gonocoxite short. Distribution. Australia (ACT, NSW, Qld.)., Published as part of Li, Xuankun & Yeates, David K., 2019, Revision of the Australian bee fly genus Sisyromyia White, 1916 (Bombyliidae, Bombyliinae, Acrophthalmydini), pp. 201-244 in Zootaxa 4711 (2) on pages 229-232, DOI: 10.11646/zootaxa.4711.2.1, http://zenodo.org/record/3576849, {"references":["Evenhuis, N. L. & Greathead, D. J. (1999) World Catalog of Bee Flies (Diptera Bombyliidae). Backhuys Publishers, Leiden, 756 pp.","Thomson, C. G. (1869) Diptera. Species nova descripsit. In: Kongliga svenska fregatten Eugenies resa omkring jordan under befal af C. A. Virgin, aren 1851 - 1853. Vol. 2 (Zoologi). Section 1, (Insecta). P. A. Norstedt & Soner, Stockholm, pp. 443 - 614.","Wiedemann, C. R. W. (1828) Aussereuropaische zweiflugelige Insekten. Als Fortsetzung des Meigenschen Werkes. Erster Theil. Schulz, Hamm, xxxii + 608 pp.","Roberts, F. H. S. (1928) A revision of the Australian Bombyliidae (Diptera). Part II. The Proceedings of the Linnean Society of New South Wales, 53, 413 - 455.","Walker, F. (1849) List of the specimens of dipterous insects in the collection of the British Museum. Parts II-IV. Printed by order of the Trustees, London, 942 pp. [pp. 231 - 1172]"]}

Sisyromyia White, 1916 Sisyromyia White, 1916: 192, 197. Type species: Bombylius auratus Walker, 1849, by subsequent designation (Bezzi, 1924: 7). Diagnosis (modified from Li & Yeates, 2018). Flagellum one-segmented, subapex with 1–5 long hairs. Base of wing vein Cu bare. Cell r5 open; cell br nearly as long as cell bm, crossvein m-m located on base of cell dm; crossvein m-m long, nearly as long as crossvein r-m; cell cup open. Abdomen with pale median stripe, consisting of dense, decumbent short scales. Male epandrium nearly rectangular, posterior half with long hairs. Hypandrium membranous. Gonocoxite elongate, basal half slightly wider than apical half; division short. Ejaculatory apodeme small. Gonocoxal apodeme short and pointed anteroventrally. Anterior arm of aedeagal sheath short, not extending beyond gonocoxal margin. Lateral ejaculatory process strong. Dorsal bridge without lateral hollow. Gonostylus normal. Phallus strong, slightly curved. Female sand chamber present. Tergite 8 with median apodeme on posterior margin, anterior margin with dense long hairs. Normally 20 to 30 acanthophorite spines present on each side of tergite 9+10, slightly curved apically. Genital fork slender and straight, connected anteriorly. Sperm pump strong and short, clothed in longitudinal muscle, without lateral papillae. Sperm pump basal and apical collars small. Distal spermathecal duct short. Spermatheca large and flattened, basal bulb present. Distribution. ACT, NSW, Qld., SA, Tas., Vic., WA. Species included. Sisyromyia albisquama sp. nov., Sisyromyia angustivitta sp. nov., Sisyromyia aurata (Walker, 1849), Sisyromyia decorata (Walker, 1849), Sisyromyia elongata sp. nov., Sisyromyia limbata (Bigot, 1892), Sisyromyia maculipennis sp. nov., Sisyromyia rutila (Walker, 1849), Sisyromyia thomsoni Evenhuis & Greathead, 1999, Sisyromyia umbra sp. nov., Sisyromyia vittata sp. nov., Published as part of Li, Xuankun & Yeates, David K., 2019, Revision of the Australian bee fly genus Sisyromyia White, 1916 (Bombyliidae, Bombyliinae, Acrophthalmydini), pp. 201-244 in Zootaxa 4711 (2) on pages 202-203, DOI: 10.11646/zootaxa.4711.2.1, http://zenodo.org/record/3576849, {"references":["White, A. (1916) The Diptera-Brachycera of Tasmania. Part III. Families Asilidae, Bombyliidae, Empidae, Dolichopodidae, & Phoridae. Papers and Proceedings of the Royal Society of Tasmania, 1916, 148 - 266. https: // doi. org / 10.5962 / bhl. part. 28993","Walker, F. (1849) List of the specimens of dipterous insects in the collection of the British Museum. Parts II-IV. Printed by order of the Trustees, London, 942 pp. [pp. 231 - 1172]","Bezzi, M. (1924) The Bombyliidae of the Ethiopian Region based on material in the British Museum (Natural History). British Museum (Natural History), London, 390 pp.","Li, X. & Yeates, D. K. (2018) Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera: Bombyliidae) with description of four new genera. Invertebrate Systematics, 32 (2), 319 - 399. https: // doi. org / 10.1071 / IS 17039","Bigot, J. M. F. (1892) Dipteres nouveaux ou peu connus. 37 e partie. XLVI Bombylidi (mihi) 1 re partie. Annals of the Entomological Society of France, 61, 321 - 76.","Evenhuis, N. L. & Greathead, D. J. (1999) World Catalog of Bee Flies (Diptera Bombyliidae). Backhuys Publishers, Leiden, 756 pp."]}

6. Sisyromyia limbata (Bigot, 1892) (Figs 12–14) Sparnopolius limbatus Bigot, 1892: 369. Type-locality: “ Australie ”; holotype, BMNH. Type specimen. Holotype of Sparnopolius limbatus Bigot, ♂ (NHMUK 010921895) (Fig. 12). Other specimen examined. Australia, SA, ♂ Adelaide, 1949. WA, ♂ (13) ♀ (6), Mt. Singleton, 20 Jul 1983, W Williamson. ♂ (1) ♀ (1), Eradu, 8 Sep 1926, EW Ferguson. ♂ 2 mi. E. of Ghooli, 29 Sep 1956, JH Calaby. Diagnosis. Medium-sized fly, body with mostly brownish hairs. Wing membrane darker on anterior half and spotted. Anterior margin of scutellum with a band consisting of short white scales. Male eyes holoptic; frons long, 2.8 x length of ocellar tubercle. Gap between inner apices of gonocoxites wide. Outer margin of gonocoxite nearly straight. Redescription. Male. Body length 6.9–8.3 mm, wing length 8.0– 9.8 mm. Head. Head about 2.1 x wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to pale yellow scales and black hairs. Eyes holoptic. Frons long, 2.8 x length of ocellar tubercle, upper narrow and black; lower half triangular, 0.8 x length of upper half with sparse pale pruinescence and white scales admixed with black hairs. Ocellar tubercle slightly raised, blackish brown to black with grey pruinescence, with long black hairs admixed with few pale yellow scales. Face with thick pale pruinescence and long black hairs admixed with some pale yellow scales, parafacial area with white scales. Gena with thick pale pruinescence and long white scales. Clypeus with thick pale pruinescence and otherwise bare. Occiput with thick pale pruinescence and dense white to pale yellow scales, admixed with black hairs near dorsal eye margin. Posterior eye margin slightly convex. Antennal scape and pedicel black with thick pale pruinescence, scape with long black hairs admixed with some white scales laterally, pedicel with short black hairs and admixed with few white hairs ventrally; flagellum black without pruinescence, subapex with 3–5 long hairs. Scape 2.9 x as long as wide, and 2.6 x as long as pedicel, uniform from base to apex. Pedicel 1.1 x as long as wide. Flagellum 8.4 x as long as wide, 1.9 x as long as scape + pedicel, 2.7 x as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus (Fig. 13c). Palpus long, just extending beyond oral cavity, brown with white hairs, one-segmented, without palpal pit. Mouthparts slender, 3.7 x as long as eye length, 2.4 x as long as head length, labellum thin and filiform (Fig. 13h). Thorax. Integumental colour of scutum mostly black with thick pale pruinescence, brown pruinescence present on transverse suture, dorsocentral, and lateral margin. Scutum covered with short pale yellow hairs admixed with short black hairs and few golden scales, hairs denser anteriorly, lateral stripe consisting of long white scales. Four brownish yellow notopleural setae present. Scutellum black with sparse pale pruinescence, anterior half with a band consisting of short white scales, posterior half with long black hairs admixed with some golden hairs, black hairs denser on posterior margin. Pleura black with thick pale pruinescence, mostly covered in long pale yellow hairs, except dorsal half of anepisternum with brown hairs admixed with few black hairs, anepimeron, meron, laterotergite and mediotergite bare. Legs. Legs mostly dark yellow except femora brown. Femora covered in golden scales, ventral face with long brown hairs. Hind femur with one row of anteroventral bristles. Bristles and other hairs on legs brown. Fore tibia 2.2 x longer than fore basitarsus, mid tibia 2.7 x longer than mid basitarsus, hind tibia 2.4 x longer than hind basitarsus. Wings. Wing membrane slightly infuscated, with inconspicuous light markings on the area around base of vein R 4 , crossvein m-m, crossvein dm-cu and apex of cell bm and cell br. Cell r 5 open, M 1 close to R 5 ; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m nearly as long as crossvein r-m; cell cup open (Fig. 13d). Haltere stem and knob pale yellow. Abdomen. Integumental colour of tergites black mostly with sparse brown pruinescence. Tergite 1 with dense yellow hairs, admixed few long black hairs laterally; tergite 2 with long golden hairs on anterior half and laterally, and with long black hairs admixed with short golden scales on posterior half; tergites 3–7 with sparse long black hairs admixed with short golden scales, laterally with long white to pale yellow long scales; tergites 2–7 with median stripe consisting of dense, decumbent short white scales. Sternites black except posterior margin yellow, with thick pale pruinescence, covered with white to pale yellow hairs, admixed with few brown hairs. Genitalia. Epandrium anterior margin slightly concave, posterior margin slightly convex (Fig. 14d). Lateral margin of gonocoxite nearly straight (Figs 14a,b); lateral ejaculatory process wide; inner apex of gonocoxite short and acute, outer apex of gonocoxite long and sharp; gap between inner apices of gonocoxites wide; phallus shorter than gonocoxite (Fig. 14c). Female. Body length 7.4–8.2 mm, wing length 8.6–10.0 mm. Very similar to male, except frons black with thick pale pruinescence, 3.4 x as wide as ocellar tubercle, frons with long black hairs admixed with some short golden scales (Fig. 13l). Around 20 acanthophorite spines present on each side of tergite 9+10 (Fig. 14g). Distribution. Australia (SA, WA)., Published as part of Li, Xuankun & Yeates, David K., 2019, Revision of the Australian bee fly genus Sisyromyia White, 1916 (Bombyliidae, Bombyliinae, Acrophthalmydini), pp. 201-244 in Zootaxa 4711 (2) on pages 219-222, DOI: 10.11646/zootaxa.4711.2.1, http://zenodo.org/record/3576849, {"references":["Bigot, J. M. F. (1892) Dipteres nouveaux ou peu connus. 37 e partie. XLVI Bombylidi (mihi) 1 re partie. Annals of the Entomological Society of France, 61, 321 - 76."]}

12. Dissodesma binghi (Evenhuis, 1989) comb. nov. (Figs 26���28) Bombylius brevirostris Macquart, 1850: 423 (119). Type locality: ��� Tasmanie ��� [error, probably = Australia (NSW)] [H in MNHN]. [Preoccupied by Olivier, 1789.] Systoechus eulabiatus Bigot, 1892: 366. Type locality: ���Van-Di��men��� [= Australia (Tas.)] [H in UNKN]. Sisyromyia binghi Evenhuis, 1989: 364 (new replacement name for Bombylius brevirostris Macquart, 1850). breverostris, incorrect original spelling of brevirostris (Macquart, 1840: pl. 7, fig. 7 [1841: pl. 7, fig. 7]). Type specimen. Holotype of Bombylius brevirostris Macquart, ♀ (MNHN ED9271) (Fig. 24). Other specimen examined. Australia, ACT, ♂ Gibraltar Peak, 9 Sep 1948. NSW, ♂ 5mi. S. Mendooran, 18 Mar 1972. G Daniels. ♂ (2) ♀ (3), Gundamain, Port Hacking, National Park, 1 Mar 1926, Mackerras. SA, ♀ Black Hill, near Montacute, Adelaide, 6 Mar 1949, HF Lower. Tas., ♀ Freycinet Nat [ional] P[ar]k, 28 Feb 1963, IFB Common & MS Upton. Redescription. Male. Body length 7.0��� 9.6 mm, wing length 7.3���10.0 mm. Head. Head about 2.0 x wider than long, mostly blackish with thick white pruinescence and covered in admixed white to pale yellow scales and hairs. Eyes holoptic. Frons long, 3.6 x length of ocellar tubercle, upper narrow and black; lower half triangular, 1.6 x length of upper half, with thick pale pruinescence and pale yellow hairs. Ocellar tubercle slightly raised, blackish brown to black with grey pruinescence, with long pale yellow hairs. Face with thick white pruinescence and long pale yellow scales, parafacial area with white scales. Gena with thick white pruinescence and long white hairs. Clypeus with thick pale pruinescence and otherwise bare. Occiput with thick white pruinescence and dense pale yellow scales. Posterior eye margin slightly concave. Antennal scape and pedicel black, scape with thick pale pruinescence and long pale yellow hairs, pedicel with sparse pale pruinescence and short pale yellow hairs; flagellum black without pruinescence. Scape 3.3 x as long as wide, and 3.6 x as long as pedicel, uniform from base to apex. Pedicel 1.1 x as long as wide. Flagellum 10.4 x as long as wide, 1.5 x as long as scape + pedicel, 1.9 x as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus (Fig. 25c). Palpus short, not extending beyond oral cavity, yellow with short black hairs, one-segmented, without palpal pit. Mouthparts slender, 3.9 x as long as eye length, 2.2 x as long as head length, labellum thin and filiform (Fig. 25h). Thorax. Integumental colour of scutum mostly black with thick grey pruinescence. Scutum covered with pale yellow hairs admixed with few black hairs near posterior margin, pale yellow hairs darker apically. Three yellow notopleural setae present. Scutellum black with sparse pale pruinescence, covered with long yellow hairs, posterior margin admixed with some strong black hairs. Pleura black with thick pale pruinescence, mostly covered in long pale yellow hairs, except anepimeron, meron, laterotergite and mediotergite bare. Legs. Legs mostly blackish brown except fore and mid femora dark yellow apically, fore and mid tibiae yellow. Femora covered in pale yellow scales, ventral face with long pale yellow hairs. Hind femur with one row of anteroventral yellow bristles. Mid and fore tibiae covered in pale yellow scales, anterior face admixed with some brown scales. Other bristles and hairs on legs brownish yellow. Fore tibia 1.7 x longer than fore basitarsus, mid tibia 2.4 x longer than mid basitarsus, hind tibia 2.2 x longer than hind basitarsus. Wings. Wing membrane slightly infuscated, darker towards wing base and along costa. Cell r 5 narrowly open; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m nearly as long as crossvein r-m; cell cup open (Fig. 25d). Haltere stem yellow, knob pale yellow. Abdomen. Integumental colour of tergites black mostly with sparse grey pruinescence. Tergite 1 with dense yellow hairs; tergites 2 to 4, and 7 to 8 with yellow hairs, admixed few long black hairs laterally and on posterior margin; tergites 5 and 6 admixed with dense long black hairs laterally. Sternites black except posterior margin yellow, with thick pale pruinescence, covered with pale yellow scales and hairs. Genitalia. Epandrium wide and short, anterior margin nearly straight, posterior margin strongly concave (Fig. 26d). Hypandrium present. Gonocoxal apex with slightly narrower than the base in dorsal view, ejaculatory apodeme small (Figs 26 ab); gonocoxal apodeme strong and incurved; lateral ejaculatory process strong; inner apex of gonocoxite acute, slender and elongate; outer apex of gonocoxite acute, normal length; dorsal bridge with lateral hollow; gonostylus normal; phallus strong, as long as gonocoxite (Fig. 26c). Female. Body length 5.2���8.4 mm, wing length 5.5���9.0 mm. Very similar to male, except frons black with thick pale pruinescence, 3.0 x as wide as ocellar tubercle, frons with short white to pale yellow scales on lower half, long pale yellow hairs on upper half, admixed with some long black hairs (Fig. 25k). Tergite 8 with thick hairs, 6 acanthophorite spines present on each side of tergite 9+10, slightly curved apically. Genital fork slender and straight, connect at apex. Sperm pump strong and long, with some lateral papillae; distal spermathecal duct of normal length, thickened in apical half; spermatheca small and nearly spherical, apex rounded (Fig. 26g). Remarks. Systoechus eulabiatus was first proposed as a junior synonym of Bombylius brevirostris by White (1916), and followed by Roberts (1928), but it was not followed by Evenhuis & Greathead (1999). The holotype of B. brevirostris was identified and examined by the senior author during his visit to MNHN, found in a very bad condition (Fig. 24). This holotype was examined by Evenhuis & Greathead (1999), but without a type label when Li visited MNHN in Sep 2018. The holotype was re-identified by Li, who added the holotype label and a label stating ��� Holotype identified by Xuankun Li Sep 2018 ��� (Fig. 24). The deposition of the holotype of S. eulabiatus is unknown, and we have not located it. By comparing the specimens from NSW and Tas., we agree with the arrangement by White (1916), and treat S. eulabiatus as a synonym of B. brevirostris. B. brevirostris was included in Sisyromyia when the genus was established (White, 1916), but the name was subsequently replaced by Sisyromyia binghi Evenhuis, 1989 due to preoccupation. Based on the genitalia characters of both male and female, this species belongs to the genus Dissodesma Bowden & Li, 2018 (Li and Yeates, 2018). Therefore, Dissodesma binghi is a new combination proposed in the present study. Distribution. Australia (ACT, NSW, SA, Tas.)., Published as part of Li, Xuankun & Yeates, David K., 2019, Revision of the Australian bee fly genus Sisyromyia White, 1916 (Bombyliidae, Bombyliinae, Acrophthalmydini), pp. 201-244 in Zootaxa 4711 (2) on pages 239-240, DOI: 10.11646/zootaxa.4711.2.1, http://zenodo.org/record/3576849, {"references":["Evenhuis, N. L. (1989) Family Bombyliidae. In: Evenhuis, N. L. (Ed.), Catalog of the Diptera of the Australasian and Oceanian Regions. Bishop Museum Press, Honolulu & E. J. Brill, Leiden, pp. 359 - 374.","Macquart, P. J. M. (1850) Dipteres exotiques nouveaux ou peu connus. 4 me supplement. Memoires de la Societe (Royale) des Science, de l'Agriculture et des Arts a Lille, 1849, 309 - 479.","Olivier, G. A. (1789) Insectes [part]. In: Encyclopedie methodique. Histoire naturelle. Tome Quatrieme. Pancoucke, Paris, pp. 45 - 331.","Bigot, J. M. F. (1892) Dipteres nouveaux ou peu connus. 37 e partie. XLVI Bombylidi (mihi) 1 re partie. Annals of the Entomological Society of France, 61, 321 - 76.","Macquart, P. J. M. (1840) Dipteres exotiques nouveaux ou peu connus. Tome deuxieme. - 1 re partie. N. E. Roret, Paris, pp. 5 - 135.","White, A. (1916) The Diptera-Brachycera of Tasmania. Part III. Families Asilidae, Bombyliidae, Empidae, Dolichopodidae, & Phoridae. Papers and Proceedings of the Royal Society of Tasmania, 1916, 148 - 266. https: // doi. org / 10.5962 / bhl. part. 28993","Roberts, F. H. S. (1928) A revision of the Australian Bombyliidae (Diptera). Part II. The Proceedings of the Linnean Society of New South Wales, 53, 413 - 455.","Evenhuis, N. L. & Greathead, D. J. (1999) World Catalog of Bee Flies (Diptera Bombyliidae). Backhuys Publishers, Leiden, 756 pp.","Li, X. & Yeates, D. K. (2018) Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera: Bombyliidae) with description of four new genera. Invertebrate Systematics, 32 (2), 319 - 399. https: // doi. org / 10.1071 / IS 17039"]}

Li, Xuankun, Yuan, David, Rodriguez, Juanita, Yeates, David K. (2019): Sand wasp (Hymenoptera: Crabronidae) parasites emerging from mud wasp nests (Hymenoptera: Sphecidae) - a reliable host record of Thraxan Yeates & Lambkin (Diptera: Bombyliidae: Anthracinae) with description of the pupal exuviae of three Thraxan species. Zootaxa 4609 (1): 149-159, DOI: https://doi.org/10.11646/zootaxa.4609.1.7

Thraxan luteus Yeates & Lambkin, 1998 (Fig. 2) Thraxan luteus Yeates & Lambkin, 1998. Type-locality: Australia (NT); Holotype ♂, ANIC. Specimens examined. Australia, Qld., ♂ Pupal exuvia, Cooktown, 12 October 1980, DH Colless, pupae in cells of?mud-wasp in termite-riddled wood. Description. Pupal exuvia. Length: 8.9 mm. Head width: 1.9 mm. Thorax width: 2.2 mm. Abdominal width: 2.6 mm, tapering to 0.5 mm at width of anal segment. Coloration. Predominantly pale yellow, spines dark brown to black. Head. Armed with 3 pairs of cephalic spines. One anterior antennal process (aap) present, base area with one dorsal long hair and one lateral long hair (Figs 2g, h). Two posterior antennal processes (pap) present, two pap about half length of aap (Figs 2g, h). One median facial hair (mfha) present, one frontal spine (fsp) present. Labral sheath (lsh) with a small ventral subapical process. Proboscidal sheath (prsh) short, about 0.5 length of lsh, slightly rugose laterally. Maxillary sheath (msh) smooth, extending and connecting well after prsh. Palpal sheath (pash) rugose, extending slightly over lsh (Figs 2c, d). One posterolateral facial hair (plfha) present. Thorax. Mostly smooth, one small posterior mesothoracic callosity (pmc) present. Thorax with one long hair located anterior of pmc, one long hair located above pmc, and two pairs of dorsal hairs. Wing sheath (wsh) and base half of leg sheaths (lesh) smooth, apex half of lesh rugose (Fig. 2d). Wsh reaching abdominal segment 3. Fore leg sheath (lesh 1) exceeding apex of wing sheath, reaching abdominal segments 3, mid leg heath (lesh 2) reaching abdominal segment 4, hind leg sheath (lesh 3) reaching abdominal segment 5. Abdomen. Eight abdominal segments visible, segments 8 and 9 fused (Fig. 2b). Abdominal tergite 1 with a row of sparse long hairs. Abdominal tergites 2 to 5 with well-developed chitinous rods with both apexes raised as spines, and with short spines on tergites 6 and 7; long hairs present laterally and between two chitinous rods or spines on tergites 2 to 7 (Fig. 2e). Tergite 8+9 with one weak dorsal spine, and one posterolateral callosity. Abdominal pleuras 2 to 7 each with a cambered row of long hairs, pleura 8+9 with four long hairs. Sternites 2 to 7 and 8+9 with a transverse row of hairs on posterior half. Anal segment smooth, some ventral small callosities present (Fig. 2j). Dorsal posterolateral process (dpp) small and fused, ventral posterolateral process (vpp) normal, acute with one inboard spine, dorsal area with one dorsal spine and one ventral spine (Figs 2i, j). Host. Based on the label: "mud-wasp"., Published as part of Li, Xuankun, Yuan, David, Rodriguez, Juanita & Yeates, David K., 2019, Sand wasp (Hymenoptera: Crabronidae) parasites emerging from mud wasp nests (Hymenoptera: Sphecidae) - a reliable host record of Thraxan Yeates & Lambkin (Diptera: Bombyliidae: Anthracinae) with description of the pupal exuviae of three Thraxan species, pp. 149-159 in Zootaxa 4609 (1) on page 152, DOI: 10.11646/zootaxa.4609.1.7, http://zenodo.org/record/3151146, {"references":["Yeates, D. K. & Lambkin, C. L. (1998) Cryptic species diversity and character congruence: review of the tribe Anthracini (Diptera: Bombyliidae) in Australia. Invertebrate Systematics, 12 (6), 977 - 1078. https: // doi. org / 10.1071 / IT 97019"]}

Thraxan misatulus Yeates & Lambkin, 1998 (Fig. 3) Thraxan misatulus Yeates & Lambkin, 1998. Type-locality: Australia (Qld); Holotype ♂, ANIC. Specimens examined. Australia, Qld., ♂ Pupal exuvia, bat cave. Boonah, October 1957, P.R. Wilkinson. Description. Pupal exuvia. Length: 17.9 mm. Head width: 3.0 mm. Thorax width: 4.1 mm. Abdominal width: 4.5 mm, tapering to 0.7 mm at width of anal segment. Coloration. Predominantly pale yellow, spines dark brown to black. Head. Armed with 3 pairs of cephalic spines. One anterior antennal process (aap) present, base area with one dorsal long hair and one lateral long hair (Figs 3g, h). Two posterior antennal processes (pap) present, two pap slightly shorter than aap (Figs 3g, h). One median facial hair (mfha) present, one frontal spine (fsp) present. Labral sheath (lsh) with a small ventral subapical process. Proboscidal sheath (prsh) short, about 0.5 length of lsh, slightly rugose laterally. Maxillary sheath (msh) smooth, extending and connecting well after prsh. Palpal sheath (pash) rugose, extending slightly over lsh (Figs 3c, d). One posterolateral facial hair (plfha) present. Thorax. Mostly smooth, one small posterior mesothoracic callosity (pmc) present. Thorax with one long hair located anterior of pmc, one long hair located above pmc, and two pairs of dorsal hairs. Wing sheath (wsh) and base half of leg sheaths (lesh) smooth, apex half of lesh rugose (Fig. 3d). Wsh reaching abdominal segment 3. Fore leg sheath (lesh 1) exceeding apex of wing sheath, reaching abdominal segments 3, mid leg heath (lesh 2) reaching abdominal segment 4, hind leg sheath (lesh 3) reaching abdominal segment 5. Abdomen. Eight abdominal segments visible, segments 8 and 9 fused (Fig. 3b). Abdominal tergite 1 with a row of sparse long hairs. Abdominal tergites 2 to 6 with well-developed chitinous rods with both apexes raised as spines on tergites 2 to 5, and posterior apex raised as spines on tergite 6, and with short spines on tergite 7; long hairs present laterally and between two chitinous rods or spines on tergites 2 to 7 (Fig. 3e). Tergite 8+9 with one dorsal spine, one lateral spine, and one posterolateral callosity. Abdominal pleuras 2 to 7 each with a cambered row of long hairs, pleura 8+9 with four long hairs. Sternites 2 to 7 and 8+9 with a transverse row of hairs on posterior half. Anal segment smooth, some ventral small callosities present (Fig. 3j). Dorsal posterolateral process (dpp) small and fused, ventral posterolateral process (vpp) elongate, acute with one inboard spine, dorsal area with one dorsal spine and two ventral spines (Figs 3i, j). Host. Based on the label: "ex mud cocoon of large psammocharid, with pupal case" (Hymenoptera: Pompilidae). Remarks. Because of the ���mud��� provenance of the host record, this pompilid species probably belongs to the tribe Ageniellini, which are the only Australian pompilids known to use pre-existing or build mud-nests (Elliot 2007). Currently the tribe includes three genera, Macromeris, Phanagenia and Auplopus. Macromeris includes a single species with large individuals (> 25mm) and the remaining two genera together include more than 20 species with smaller individuals (Macromeris violacea (Lepeletier) mud nest., Published as part of Li, Xuankun, Yuan, David, Rodriguez, Juanita & Yeates, David K., 2019, Sand wasp (Hymenoptera: Crabronidae) parasites emerging from mud wasp nests (Hymenoptera: Sphecidae) - a reliable host record of Thraxan Yeates & Lambkin (Diptera: Bombyliidae: Anthracinae) with description of the pupal exuviae of three Thraxan species, pp. 149-159 in Zootaxa 4609 (1) on pages 152-156, DOI: 10.11646/zootaxa.4609.1.7, http://zenodo.org/record/3151146, {"references":["Yeates, D. K. & Lambkin, C. L. (1998) Cryptic species diversity and character congruence: review of the tribe Anthracini (Diptera: Bombyliidae) in Australia. Invertebrate Systematics, 12 (6), 977 - 1078. https: // doi. org / 10.1071 / IT 97019","Elliot, M. G. (2007) Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa, 1428, 1 - 83."]}

Thraxan sp. (Fig. 1) Specimens examined. Australia, ACT, ♀ Pupal exuvia, Black Mountain, Bike tunnel, 5 September 2018; Pupated 30 September; Emerged 22 October. ♀ Pupal exuvia, Australian National University campus, 1 October; Pupated 8 October; Emerged 4 November. Description. Pupal exuvia. Length: 13.0 mm. Head width: 2.2 mm. Thorax width: 2.9 mm. Abdominal width: 3.0 mm, tapering to 0.9 mm at width of anal segment. Coloration. Predominantly pale yellow, spines dark brown to black. Head. Armed with 3 pairs of cephalic spines. One anterior antennal process (aap) present, base area with one dorsal long hair and one lateral long hair (Figs 1g, h). Two posterior antennal processes (pap) present, two pap slightly shorter than aap (Figs 1g, h). One median facial hair (mfha) present, one frontal spine (fsp) present. Labral sheath (lsh) with a small ventral subapical process. Proboscidal sheath (prsh) short, about 0.5 length of lsh, slightly rugose laterally. Maxillary sheath (msh) smooth, extending and connecting well after prsh. Palpal sheath (pash) rugose, extending slightly over lsh (Figs 1c, d). One posterolateral facial hair (plfha) present. Thorax. Mostly smooth, one small posterior mesothoracic callosity (pmc) present. Thorax with one long hair located anterior of pmc, one long hair located above pmc, and two pairs of dorsal hairs. Wing sheath (wsh) and base half of leg sheaths (lesh) smooth, apex half of lesh rugose (Fig. 1d). Wsh reaching abdominal segment 3. Fore leg sheath (lesh 1) not exceeding apex of wing sheath, mid leg heath (lesh 2) reaching abdominal segment 3, hind leg sheath (lesh 3) reaching abdominal segment 4. Abdomen. Eight abdominal segments visible, segments 8 and 9 fused (Fig. 1b). Abdominal tergite 1 with a row of dense long hairs. Abdominal tergites 2 to 6 with well-developed chitinous rods with both apexes raised as spines on tergites 2 to 5, and posterior apex raised as spines on tergite 6, and with short spines on tergite 7; long hairs present laterally and between two chitinous rods or spines on tergites 2 to 7 (Fig. 1e). Tergite 8+9 with one dorsal spine, and one posterolateral callosity. Abdominal pleuras 2 to 7 each with a cambered row of long hairs, pleura 8+9 with two long hairs. Sternites 2 to 7 and 8+9 with a transverse row of hairs on posterior half. Anal segment smooth, some ventral small callosities present (Fig. 1j). Dorsal posterolateral process (dpp) small and fused, ventral posterolateral process (vpp) elongate, acute with one inboard spine, dorsal area with one dorsal spine and one ventral spine (Figs 1i, j). Host. Pison simillimum Smith (Hymenoptera: Crabronidae: Crabroninae)., Published as part of Li, Xuankun, Yuan, David, Rodriguez, Juanita & Yeates, David K., 2019, Sand wasp (Hymenoptera: Crabronidae) parasites emerging from mud wasp nests (Hymenoptera: Sphecidae) - a reliable host record of Thraxan Yeates & Lambkin (Diptera: Bombyliidae: Anthracinae) with description of the pupal exuviae of three Thraxan species, pp. 149-159 in Zootaxa 4609 (1) on pages 150-152, DOI: 10.11646/zootaxa.4609.1.7, http://zenodo.org/record/3151146

Thornhill, Andrew H., Crisp, Michael D., Külheim, Carsten, Lam, Kristy E., Nelson, Leigh A., Yeates, David K., Miller, Joseph T. (2019): A dated molecular perspective of eucalypt taxonomy, evolution and diversification. Australian Systematic Botany 32: 29-48, DOI: 10.1071/SB18015

Key to Australian genera of Bombyliini 1 M 1 ending at wing margin, cell r 5 open............................. 2 ��� M 1 ending at r 5 , cell r 5 closed............................................. 9 2 Palpus two-segmented......................................................... 3 ��� Palpus one-segmented......................................................... 4 3 Flagellum apex truncate, with dorsal apical stylus; anterior half of scutum strongly arched; abdominal hairs uniformly yellowish, without longitudinal pale scales in middle part, hairs relatively long laterally.................................. Eusurbus ��� Flagellum apex tapering, with apical stylus; anterior half of scutum slightly arched; abdominal hairs with a remarkable black and white pattern, longitudinal pale scales present in middle part, hairs long and brush-like laterally................................................................. Zentamyia 4 Cell br much longer than cell bm, crossvein r-m located on apical half of cell dm........................................................... 5 ��� Cell br nearly as long as cell bm, crossvein r-m located on basal half of cell dm............................................................ 7 5 Pedicel nearly as long as width, and scape ~3.0�� longer than pedicel; horizontal line dividing compound eye................................................................ Cryomyia (part) ��� Pedicel ~2.0�� longer than wide, and scape less than 2.0�� longer than pedicel; without horizontal line dividing compound eye............................................................................. 6 6 Pedicel 2.5�� longer than wide; wing brown towards costal third; female sperm pump shorter than 1.0�� length of genital fork; distal spermathecal duct longer than 2.0�� of genital length; sperm bulb with apical small conical point; basal bulb absent (Li & Yeates 2018a: fig. 50g).............................................................. Robertsmyia ��� Pedicel 1.8�� longer than wide; wing uniformly slightly infuscated, without distinct darker area towards costal third; female sperm pump longer than 2.5�� length of genital fork, strong and sinuous; distal spermathecal duct shorter than 1.0�� length of genital fork; sperm bulb flattened, apex spherical; basal bulb present (Li & Yeates 2018a: fig. 26g).................................................................... Laurella 7 Male eyes narrowly separated by 0.5�� width of ocellus; femora thickened; pulvillus and claw enlarged, longer than distitarsus; male genitalia enlarged; female genital fork enlarged.......................................... Paramonovius gen. nov. ��� Male eyes clearly holoptic, not separated; femora not thickened; pulvillus and claw not enlarged, shorter than distitarsus; male genitalia not enlarged; female genital fork not enlarged......................................................................... 8 8 Flagellum with subapical long hairs (Li & Yeates 2018a: fig. 6x); labellum broad and fleshy; abdomen with median longitudinal stripe, consisting of dense, decumbent, short scales; abdominal apex without long, brush-like hairs...................................................................... Sisyromyia ��� Flagellum with subapical setae and setula (Li & Yeates 2018a: fig. 21c) or bare (Li & Yeates 2018a: fig. 18c); labellum thin and filiform; abdomen with median longitudinal stripe, consisting of long scales; abdominal apex with long, brush-like hairs................................................... Dissodesma 9 Crossvein m-m much shorter than crossvein r-m, sometimes absent; M 1 + 2 present (Li & Yeates 2018a: fig. 7m)....... 10 ��� Crossvein m-m nearly as long as crossvein r-m; M 1 + 2 never present (Li & Yeates 2018a: fig. 7o)................................ 12 10 Compound eye with horizontal line............ Cryomyia (part) ��� Compound eye without horizontal line............................. 11 11 Second flagellomere long; base of wing vein Cu bare; thoracic scutum without pale hairs on lateral margin; abdominal tergites largely yellow with black medially; female without sand chamber...................................................... Eristalopsis ��� Second flagellomere short; base of wing vein Cu often with long hairs; thoracic scutum often with pale hairs on lateral margin; abdominal tergites blackish; female with sand chamber............................................................ Staurostichus 12 Cell br much longer than cell bm, crossvein r-m located on middle part of cell dm.......................................... Nigromyia ��� Cell br nearly as long as cell bm, crossvein r-m located on basal part of cell dm.......................................................... 13 13 Base of wing vein Cu with long hairs.............................. 14 ��� Base of wing vein Cu bare................................................ 15 14 Flagellum two-segmented; female eye separation less than 2.5�� of ocellar tubercle width, frons normal, less than 3.5�� width of frons......................................................... Mandella ��� Flagellum one-segmented; female eye separation more than 3.5�� of ocellar tubercle width, frons wide, over 3.5�� width of frons................................................................... Choristus 15 Flagellum elongate and slender; palpus short, without palpal pit; abdomen with brush-like long hairs................ Meomyia ��� Flagellum not elongate; palpus long, with lateral palpal pit; abdomen without brush-like long hairs........ Lambkinomyia, Published as part of Li, Xuankun & Yeates, David K, 2019, A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position, pp. 92-203 in Austral Entomology 58 on pages 93-94, DOI: 10.1111/aen.12361, http://zenodo.org/record/3265414, {"references":["Li X & Yeates DK. 2018 a. Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera: Bombyliidae) with description of four new genera. Invertebrate Systematics 32, 319 - 399."]}

Genus Paramonovius gen. nov. http://zoobank.org/ urn:lsid:zoobank.org:act: 4E3A2504-8599- 4322-8841-8D1217A566B7 Type Species. Paramonovius nightking sp.nov. (here designated). Diagnosis Medium-sized bombyliine (body length around 8���11 mm). Male eyes narrowly separated (Fig. 2e); female frons wide (Fig. 3c). Antennal flagellum one-segmented with apical stylus, subapex with 3���5 long hairs (Fig. 2c). Palpus short, one-segmented. Mouthparts long and slender, labellum thin and filiform (Fig. 2f). Scutum, scutellum, pleura and coxae densely clothed in hairs, except anepimeron, mero, laterotergite and mediotergite bare. Femora strong, with anterior blackish brown bristles. Claw and pulvillus enlarged, as long as distitarsus (Figs 2i and 3g). Wing with cell r 5 widely open; cell br nearly as long as cell bm, crossvein r-m arising at base of cell dm; cell cup open; alula well developed (Fig. 2d). Abdomen broad and short. Tergites with median stripe consisting of dense, decumbent short white scales. Male genitalia enlarged, epandrium posterolateral corner with thick long hairs (Fig. 4d); hypandrium fused with gonocoxite; gonocoxite basal half wide and apical half abrupt narrowed (Fig. 4c); anterior arm of aedeagal sheath long and wide, extending beyond gonocoxal margin. Female sand chamber present, tergite 8 fused and sclerotized. Acanthophorite spines present. Genital fork strong and broad, enlarged basally. Sperm pump strong and of normal length; common and basal spermathecal duct normal; apical spermathecal duct short; spermatheca elongate, cylindrical with apex spherical (Fig. 4f). Description Head. Head wide and relatively short, covered in admixed hairs and scales. Eyes narrowly separated in male, widely separated in female. Male eyes separated by 0.5�� width of ocellus, frons short (Fig. 2e). Female eyes widely separated, frons wide, around 3.0�� as wide as ocellar tubercle (Fig. 3c). Posterior eye margin slightly sinuous. Antennal scape elongate, about 3.5�� as long as wide; pedicel elongate, about 1.5�� as long as wide; flagellum one-segmented with apical stylus, subapex with 3���5 long hairs (Fig. 2c). Palpus short, not extending beyond oral cavity, onesegmented, without palpal pit. Mouthparts long and slender, labellum thin and filiform (Fig. 2f). Thorax. Scutum and scutellum with pale pruinescence and covered with hairs and scales. Five notopleural setae present. Postalar callus setae absent. Pleura with thick pale pruinescence, anepisternum, katepisternum, katepimeron and coxae densely covered with long scales, but anepimeron, mero, laterotergite and mediotergite bare (Figs 2b and 3b). Femora strong, with anterior blackish brown bristles. Claw and pulvillus enlarged, as long as distitarsus (Figs 2i and 3g). Wing with cell r 5 widely open; cell br nearly as long as cell bm, crossvein r-m arising base of cell dm; crossvein m-m much longer than crossvein r-m; cell cup open; alula well developed (Fig. 2d). Abdomen. Abdomen broad and short, tergites 3���7 compact, tergites 7���9 rotated, with epandrium and gonocoxite rotated 180��; tergites 2���6 with median stripe consisting of dense, decumbent short white scales. Male epandrium enlarged, nearly rectangular, anterior margin deeply concave, posterolateral corner with thick long hairs (Fig. 4d). Hypandrium fused with gonocoxite. Gonocoxite enlarged, basal half wide and apical half abruptly narrowed. Ejaculatory apodeme large (Fig. 4a,b); gonocoxal apodeme strong and pointed anteriorly; anterior arm of aedeagal sheath long and wide, extending beyond gonocoxal margin; lateral ejaculatory process strong; inner and outer apexes of gonocoxite sharp; gonostylus small and strongly incurved (Fig. 4c). Female sand chamber present. Tergite 8 fused and sclerotized. Around 30 acanthophorite spines present on each side. Genital fork strong and broad, enlarge basally. Sperm pump strong and about as long as basal spermathecal duct; common and basal spermathecal duct not elongate; apical spermathecal duct shorter than spermatheca; spermatheca elongate, cylindrical with apex spherical (Fig. 4f). Etymology This generic name is in honour of Dr. Sergei Jacques Paramonov for his significant contribution to Australian dipterology. Included species This genus is monotypic for Paramonovius nightking sp. nov., Published as part of Li, Xuankun & Yeates, David K, 2019, A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position, pp. 92-203 in Austral Entomology 58 on pages 94-96, DOI: 10.1111/aen.12361, http://zenodo.org/record/3265414

Li, Xuankun, Yeates, David K (2019): A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position. Austral Entomology 58: 92-203, DOI: 10.1111/aen.12361

Paramonovius nightking sp. nov (Figs 2���4) http://zoobank.org/ urn:lsid:zoobank.org:pub: 936DFF93- AAA3-460F-AD5D-8EC342CB3132 Material examined Holotype ♂ AUSTRALIA Western Australia, Wandoo National Park, Kent Road, York, -31.96443 116.51224, Scattered Banksia with low open shrubs on white sand. On Leucopogon oxycedrus, 19 May 2012, J. and F. Hort (WAM: E97796). Paratypes 7♂, 2♀: AUSTRALIA Western Australia, Wandoo National Park, Kent Road, York, -32.1180 116.60557, Jarrah open woodland with low open mixed shrubs. Feeding on Leucopogon oxycedrus, 19 May 2012, J. and F. Hort (WAM: E97797 ��� E97798, E97801 ��� E97807). Other material 2♂ AUSTRALIA Western Australia, Flynn State Forest, Kent Road, York, -31.96443 116.51224, Scattered Banksia with low open shrubs on white sand. On Leucopogon oxycedrus, 19 May 2012, J. and F. Hort (WAM: E97799 ��� E97800). Diagnosis Body mostly covered with thick pale pruinescence. Scape and pedicel brownish yellow, flagellum black. Scutum with large black square markings in each corner and one thin black stripe in the midline. Wing membrane infuscated on anterior half, with inconspicuous lighter markings. Description Male Body length 8.7���10.9 mm, wing length 9.1���11.5 mm. Head. Head about 2.2�� wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to black hairs and scales. Eye narrowly separate by 0.5�� width of ocellus. Frons short, 2.0�� length of ocellar tubercle, with thick pale pruinescence and few golden scales. Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with long black hairs turning dark yellow at base. Face with thick pale pruinescence and long black hairs except turning dark yellow at base, parafacial area with long white scales admixed with a few long hairs. Gena with thick pale pruinescence and long white scales admixed with few long hairs. Clypeus with sparse pale pruinescence and otherwise bare. Occiput with dense white hairs and scales, dorsal central area admixed with fine black hairs and strong yellow hairs. Posterior eye margin slightly convex. Antennal scape and pedicel brownish yellow with thick pale pruinescence, scape with long black hairs turning dark yellow at base, pedicel with short white to pale yellow hairs; flagellum black without pruinescence, subapex with 3���5 long hairs. Scape 3.4�� as long as wide, and 2.8�� as long as pedicel, uniform from base to apex. Pedicel 1.6�� as long as wide. Flagellum 12.0�� as long as wide, 1.9�� as long as scape + pedicel, 2.6�� as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus (Fig. 2c). Palpus short, not extending beyond oral cavity, black with blackish brown hairs, one-segmented, without palpal pit. Mouthparts slender, 3.2�� as long as eye length, 4.8�� as long as head length, labellum thin and filiform (Fig. 2f). Thorax. Integumental colour of scutum mostly black with dense pale pruinescence except postalar callus brownish yellow, scutum with large black square markings in each corner and one thin black stripe in the midline. Scutum covered with short white and golden scales, admixed with some long black hairs. Five brownish yellow notopleural setae present. Postalar callus with long black hairs. Scutellum brownish yellow with sparse pale pruinescence, with short white scales on anterior margin, dense short black scales and long black hairs on posterior margin, admixed with short golden scales. Pleura black with thick pale pruinescence, anepisternum with long white scales except anterior half with strong yellow hairs, katepisternum, katepimeron and dorsal margin of metepisternum with long white scales, coxae with dense long white scales, anepimeron, meron, laterotergite and mediotergite bare. Legs. Legs mostly brownish yellow. Fore and mid femora with black scales on anterior face, hind femur with black scales on dorsal half, ventral half of femora with white to pale yellow hairs. Femora strong, with anterior blackish brown bristles. Other hairs and bristles on legs brown. Fore tibia 2.2�� longer than fore basitarsus, mid tibia 2.5�� longer than mid basitarsus, hind tibia 2.0�� longer than hind basitarsus. Claw and pulvillus enlarged, as long as distitarsus (Figs 2i and 3g). Wings. Wing membrane infuscated on anterior half, with inconspicuous light markings on the area around base of vein R 4 , crossvein r-m, crossvein dm-cu and apex of cell bm. Cell r 5 widely open; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m long, 2.0�� as long as crossvein r-m; cell cup open (Fig. 2d). Haltere stem yellow, knob dark yellow. Abdomen. Integumental colour of tergites black with thick grey pruinescence. Tergite 1 with pale yellow hairs admixed with long black hairs on posterior margin; tergite 2 covered with white scales admixed with brown to black hairs on posterior half and short golden scales on posterior margin; tergites 3 to 7 compact, with golden scales admixed with black hairs, and with thick short white scales posterolaterally; tergites 2���6 with median stripe consisting of dense, decumbent short white scales. Tergites 7���9 rotated, with epandrium and gonocoxite rotated 180��. Genitalia. Epandrium nearly rectangular, anterior margin deeply concave, posterolateral corner with thick long hairs (Fig. 4d). Hypandrium fused with gonocoxite. Gonocoxite elongate, basal half wide and apical half abrupt narrowed. Ejaculatory apodeme large (Fig. 4a,b); gonocoxal apodeme strong and pointed anteriorly; anterior arm of aedeagal sheath long and wide, extending beyond gonocoxal margin; lateral ejaculatory process strong; inner apex of gonocoxite sharp; outer apex of gonocoxite sharp; dorsal bridge without lateral hollow; aedeagal sheath with acute projection on subapex; gonostylus small and strongly incurved; phallus long and strong (Fig. 4c). Female Body length 10.8���11.0 mm, wing length 11.5���11.6 mm. Very similar to male, except frons black with thick pale pruinescence, 3.0�� as wide as ocellar tubercle, frons with long brown to black hairs admixed with short golden scales (Fig. 3c). Tergite 8 fused and sclerotized, with some sparse hairs. Around 30 acanthophorite spines present on each side, slightly curved apically. Genital fork strong and broad, enlarged basally. Sperm pump strong and about as long as basal spermathecal duct, clothed in longitudinal muscle, with lateral papillae; sperm pump basal collars present and apical collars absent; common and basal spermathecal duct not elongate; apical spermathecal duct shorter than spermatheca; spermatheca elongate, cylindrical with apex spherical (Fig. 4f). Remarks Wing with short stump vein sometimes present on crossvein m-cua. Etymology This species is named after the Night King in the American fantasy drama Game of Thrones, because all the specimens were collected in winter and the fly is mostly covered in thick pale pruinescence. The specific name is treated as a noun in apposition. Distribution Endemic to WA., Published as part of Li, Xuankun & Yeates, David K, 2019, A new genus and species of an unusual Australian winter bee fly (Diptera: Bombyliidae) with discussion on its phylogenetic position, pp. 92-203 in Austral Entomology 58 on pages 96-99, DOI: 10.1111/aen.12361, http://zenodo.org/record/3265414

Thevenetimyia mimula Hall, 1969 Fig. 18 Thevenetimyia mimula Hall, 1969: 61. Type-locality: not given [= Australia]; holotype, ANIC. Types and other specimens examined. Holotype ♂, Australia, Macl [eay] M[useum], now in ANIC 29-009851 (Fig. 18). QLD, 1♂, Rockhampton, Macl [eay] M[useum] [hind leg and abdomen missing], now in ANIC 29-038919. Diagnosis. Medium-sized Thevenetimyia, wing membrane hyaline, without infuscated marks; costa with two rows of small spines; scutum with two rows of short, thick spines and some adjacent, separate spines; three postalar setae present; abdominal tergites 5 to 7 with lateral dense long black hairs. Redescription. Male. Body length 12.5 mm, wing length 10.7–11.8 mm. Head. Head about 1.5× wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to blackish brown hairs. Eye widely connected, by 1.0× length of ocellar tubercle. Frons slender and long, upper triangular section small, lower triangular section black with thin pale pruinescence, middle narrow part about 0.7× length of frons. Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with blackish brown hairs. Face with thick pale pruinescence and with brown hairs, parafacial area blackish brown and bare. Gena with thick pale pruinescence and long white hairs. Clypeus swollen, with thick pale pruinescence and long blackish brown hairs. Occiput with long white hairs except dorsal margin admixed with brown to black hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral hairs of scape longer, hairs on pedicel short. Scape 3.5× as long as wide, and 4.0× as long as pedicel, uniform from base to apex. Pedicel 0.9× as long as wide. Flagellum 6.0× as long as wide, 1.2× as long as scape+pedicel (1.5× as long as scape), conical and slightly laterally compressed, one-segmented with apical style (Fig. 18c). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, twosegmented, with palpal pit. Mouthparts slender, 4.3× as long as eye length (2.6× as long as head length), labella thin and filiform (Fig. 18f). Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, pruinescence more dense around margins. Scutum and scutellum with admixed white and black hairs and white long scales. Three notopleural setae present, 3 postalar setae present. Posterior margin of scutellum with strong hairs. Pre- and postalar setae and strong hairs on postalar callus and scutellum pale yellow. Scutum with two rows of short, thick spines and some adjacent, separate spines. Pleura and coxae black with thick grey pruinescence, anepisternum with admixed long white hairs and scales, katepisternum and metepisternum with moderate long white scales, and mediotergite with dense white to pale yellow hairs and setae, anepimeron, meron and laterotergite bare. Legs. Legs mostly black. Femora mostly covered with white scales except on apex, ventral face of basal half with long white hairs; fore and mid femora covered with brownish yellow scales. Hind femur with a row of short ventral bristles. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 2.1× longer than fore basitarsus, mid tibia 2.1× longer than mid basitarsus. [Hind tibia and tarsus broken]. Wings. Wing membrane hyaline, without infuscated marks. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising a little over half way from the base of cell dm; crossvein m-m long, and 2.0× as long as crossvein r-m; cell cup open (Fig. 18d). Costa with two rows of short, thick spines. Haltere stem pale yellow, knob black. Abdomen. Integumental colour of tergites black with thin pale pruinescence, except tergite 1 and lateral margin of tergites 2 and 3 with thick pruinescence. Tergites mostly covered with admixed white and black hairs. Tergites 1 to 4 and 8 with dense long white hairs on lateral margin, tergites 5 to 7 with dense long black hairs on lateral margin. Genitalia. Genitalia have been dissected and are poorly preserved. See description in Hall (1969: p61; p84, figure 48). Female. Unknown. Remarks. Thevenetimyia mimula Hall, 1969 could be separated easily from other Australian Thevenetimyia by having a hyaline wing membrane, three postalar setae present, and abdominal tergites with dense long white and black hairs laterally. However, it is considered to be rare (only two specimens been documented so far) and mysterious (type-locality unknown for 49 years). We found a second specimen identified as Thevenetimyia mimula in ANIC. This specimen also with the label “Macl[eay]. M[useum].”, with another label “Rockhampton Queensland ”. Therefore, we are able to confirm the distribution of this species. Unfortunately, this specimen’s abdomen is missing, and the male genitalia of the holotype has been dissected but poorly preserved, so, we are unable to redescribe and photograph it. The female of this species is still unknown. Distribution. QLD., Published as part of Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos José Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), pp. 331-375 in Records of the Australian Museum 70 (3) on pages 356-358, DOI: 10.3853/j.2201-4349.70.2018.1678, http://zenodo.org/record/5237395, {"references":["Hall, J. C. 1969. Review of the subfamily Cylleniinae with a world revision of the genus Thevenetimyia Bigot (Eclimus auct.) (Diptera: Bombyliidae). University of California Publications in Entomology 56: 1 - 85."]}

Thevenetimyia major Li and Yeates, sp. nov. Figs 16–17 Holotype ♂, Australia, QLD, 30 km W of Alpha, T. M. S. Hanlon, ex dead standing Acacia, collected 30 Aug 1998, emerged 16 Nov 1999, AM K.364586. Paratype 1♂, NT, Koongarra, 15 km E of Mt Cahill, 12°52'S 132°50'E, 6 Oct 1975, A. Allwood & T. Angeles, ANIC 29-041486. Diagnosis. Large-sized Thevenetimyia, wing membrane with large brown mark on middle area, but remaining wing apex hyaline, base mostly infuscated; vein R 4 arising far from base, 0.8 from crossvein r-m; costa without small spines; hind femur with a row of short black bristles; abdominal tergites 5 to 8 with dense lateral long black hairs. Description. Male. Body length 14.6–16.7 mm, wing length 12.7–c. 12.9 mm. Head. Head about 1.7× wider than long, mostly blackish with thin pale pruinescence and covered in admixed white, pale yellow and black hairs. Eye narrowly separated, by 0.2× width of ocellus. Frons slender and long, upper triangular section small, lower triangular section black, middle narrow part about 0.7× length of frons. Ocellar tubercle slightly raised, black with grey pruinescence, with black hairs. Face with thin pale pruinescence, lateral area of antenna with some black hairs, parafacial area black and bare. Gena with thick pale pruinescence and long white hairs. Clypeus swollen, with thin pale pruinescence and blackish brown hairs. Occiput mostly with pale yellow hairs, except ventral area with white hairs and dorsal margin with black hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral face of scape admixed with white hairs. Scape 3.2× as long as wide, and 3.4× as long as pedicel, uniform from base to apex. Pedicel 1.1× as long as wide. Flagellum 8.3× as long as wide, 1.4× as long as scape+pedicel (1.8× as long as scape), conical and slightly laterally compressed, onesegmented with apical style (Fig. 16c). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, 3.5× as long as eye length (2.0× as long as head length), labella thin and filiform (Fig. 16f). Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, pruinescence more dense around margins. Scutum with short black hairs, anterior half and lateral margins admixed with dense long white scales, posterior half with long golden scales. Scutellum covered with admixed white and yellow scales and black hairs, posterior margin with black setae. Three pale yellow notopleural setae, and six pale yellow postalar setae present. Pleura and coxae black with thin pale pruinescence, anepisternum and katepisternum with admixed white and black hairs, dorsal half of metepisternum with long white scales, mediotergite with dense white hairs and setae, anepimeron, meron and laterotergite bare. Legs. Legs black, mostly covered with black scales, except basal half of hind femur and posterior face of mid femur and tibia admixed with white scales, ventral face of fore and mid femora with short black hairs, hind femur with a row of short black bristles. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 1.6× longer than fore basitarsus, mid tibia 1.7× longer than mid basitarsus, hind tibia 1.9× longer than hind basitarsus. Wings. Wing membrane with large brown mark in median area, but wing apex hyaline, base mostly infuscated. Cell r 1 , base 0.7 of cell r 2+3 , base 0.55 of cell r 5 , base 0.1 of cell m 1 , base 0.9 of cell m 2 , cell cua 1 and cell dm, apex 0.3 of cell cup, apex 0.4 of cell br and apex 0.2 of cell bm all covered with brown infuscation, except cell cua 1 and cell dm with small middle clear area. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising half way from the base of cell dm; crossvein m-m long, and 2.0× as long as crossvein r-m; cell cup open (Fig. 16d). Vein R 4 arising 0.8 from crossvein r-m. [Wing apex broken]. Costa without small spines. Haltere stem dark yellow, knob blackish brown. Abdomen. Integumental colour of tergites dark brown with thin pale pruinescence, except lateral margin with thick pruinescence. Tergite 1 with dense long white hairs. Tergites 2 to 8 mostly covered with black scales and short black hairs, tergite 2 admixed with long white hairs and posterior margin with white scales, tergites 3 to 4 with long white hairs laterally, posterolateral with dense white scales, tergites 5 to 8 with dense long black hairs laterally, tergite 5 with dense white scales on dorsal area. Sternites with black integumental colour except posterior margins yellow, mostly with thick grey pruinescence and white hairs, except sternites 5 to 8 covered with black hairs. Genitalia. Epandrium wide and short, posterior margin slightly convex (Fig. 17d). Hypandrium present. Gonocoxal apex slightly narrower than the base in dorsal view, ejaculatory apodeme large (Figs 17a, b); gonocoxal apodeme strong and incurved; lateral ejaculatory process strong, 3.7× longer than wide, apex expanded; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite rounded; dorsal bridge with long lateral hollow (Fig. 17c); gonostylus slender and pointed dorsally, 3.0× longer than wide. Female. Unknown. Remarks. Thevenetimyia major Li and Yeates, sp. nov. differs markedly from congeners by having a large body size, a large brown mark on middle area of wing, and a vein R 4 arising far from base, 0.8 from crossvein r-m. The paratype is without clear area in the middle of brown infuscation. Distribution. QLD, NT. Etymology. This specific name refers to the large body size of the species., Published as part of Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos José Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), pp. 331-375 in Records of the Australian Museum 70 (3) on pages 353-356, DOI: 10.3853/j.2201-4349.70.2018.1678, http://zenodo.org/record/5237395

Thevenetimyia furvicostata (Roberts, 1929) Figs 6���9 Eclimus furvicostatus Roberts, 1929: 578. Type-locality: Australia (QLD); holotype, QM. Thevenetimyia furvicostata (Roberts, 1929).��� Hall, 1969: 36. Types and other specimens examined. Holotype ♂, Australia, QLD, Westwood, Nov 1927, A. P. Dodd, QM D3526 (Fig. 6), and 1♂ paratype, QLD, Stanthorpe, 27 Jan 1927, [Eclimus furvicostatus Rob. R. W. Crosskey det. 1962/ 7279/ Press by Com Inst Ent BM 1962-1], NHMUK. QLD, 1♂, Carnarvon NP, Top Moffatt Camp, 25��04'08"S 148��03'03"E, 28 Nov 1997, J. Skevington, C. Lambkin, S. Evans, MV Light, ANIC 29-038915; 1♂, Davies Ck NP nr Mareeba, 16��56'S 145��32'E, 6 Jan 1992, AM K.364583; 5♂♂, 16 km N Boonah, SEQ 27��54'S 152��41'E, 19 Dec 1996, C. J. Burwell, S. Evans, QM; 1♂, Mt Moffatt NP, 3 km SE Park Headquarters, 740 m, 25��04'39"S 148��00'30"E, 20 Nov 1995, SCQ, C. J. Burwell, QM; 1♀, Chimneys, Mt Moffatt NP, C.Qld 25��06'S 147��52'E, 1 Dec 1997, C. Lambkin, S. Evans, J. Skevington, QM. Diagnosis. Medium-sized Thevenetimyia, wing membrane dark brown anterior ⅓, and posterior hyaline, with distinct margin; costa with two rows of small spines; scutum with two rows of distinctly short, thick spines and some adjacent, separate spines; ejaculatory apodeme relatively small. Redescription. Male. Body length 10.6���12.0 mm, wing length 8.4���9.8 mm. Head. Head about 1.4�� wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to blackish brown hairs. Eye narrowly separated, by 0.1�� width of ocellus. Frons slender and long, upper triangular section small, lower triangular section black, middle narrow part about 0.6�� length of frons. Ocellar tubercle slightly raised, black to dark brown with grey pruinescence, with blackish brown hairs. Face with thick pale pruinescence and with few brown hairs on laterodorsal area, parafacial area brown and bare. Gena with thick pale pruinescence and long white hairs. Clypeus swollen, with thick pale pruinescence and long white hairs. Occiput with long white hairs except dorsal margin admixed with brown to black hairs. Posterior eye margin slightly sinuous. Antennae blackish brown; scape and pedicel with thin pale pruinescence and black hairs, ventral hairs of scape slightly longer, hairs on pedicel short. Scape 3.5�� as long as wide, and 3.0�� as long as pedicel, uniform from base to apex. Pedicel 1.1�� as long as wide. Flagellum 4.6�� as long as wide, 1.1�� as long as scape+pedicel (1.5�� as long as scape), conical and slightly laterally compressed, one-segmented with apical style (Fig. 7c). Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, 3.0�� as long as eye length (2.0�� as long as head length), labella thin and filiform (Fig. 7f). Thorax. Integumental colour of scutum and scutellum mostly black with sparse grey pruinescence, pruinescence more dense around margins. Scutum and scutellum with admixed long white scales and white and blackish brown hairs. Three notopleural setae present. Postalar callus and posterior margin of scutellum with strong hairs. Notopleural setae and strong hairs on postalar callus and scutellum pale yellow. Scutum with two rows of distinct short, thick spines and some adjacent, separate spines. Pleura and coxae black with thick grey pruinescence, anepisternum and dorsal half of katepisternum with long white hairs, metepisternum with dense long white scales, and mediotergite with dense white hairs and setae, anepimeron, meron and laterotergite bare. Legs. Legs mostly dark brown, except hind femur basal half and fore and mid tibiae yellow. Femora and tibiae mostly covered with brownish-yellow scales, except femora and posterior face of fore and mid tibiae admixed with white scales. Other hairs and bristles on legs short and blackish brown to black. Fore tibia 2.2�� longer than fore basitarsus, mid tibia 2.0�� longer than mid basitarsus, hind tibia 1.9�� longer than hind basitarsus. Wings. Wing membrane dark brown anterior 1/3, and posterior hyaline. Cell br with base 0.4 slightly infuscated, dark area along vein M, and covered crossvein r-m, a small hyaline area in posterior margin of cell r 2+3 between crossvein r-m and vein R 4+5 branching. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising a little over half way from the base of cell dm; crossvein m-m long, and 2.0�� as long as crossvein r-m; cell cup open (Fig. 7d). Costa with two rows of short, thick spines. Haltere stem yellow, knob blackish brown. Abdomen. Integumental colour of tergites black with thin pale pruinescence, except lateral margin dark yellow. Tergites mostly covered with black scales and short black hairs. Tergite 1 covered with long white scales and hairs. Tergites 2 to 5 admixed with white scales laterally, and black scales on dorsal area. Sternites with dark brown integumental colour except posterior margins yellow, with thick grey pruinescence and white hairs. Genitalia. Epandrium wide and short, posterior margin slightly convex (Fig. 9d). Hypandrium present, constricted in the middle. Gonocoxal apex slightly narrower than the base in dorsal view, ejaculatory apodeme relatively small (Figs 9a, b); gonocoxal apodeme strong and incurved; lateral ejaculatory process strong, 2.0�� longer than wide, apex not expanded; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite rounded; dorsal bridge with long lateral hollow (Fig. 9c); gonostylus slender and pointed dorsally, 3.0�� longer than wide. Female. Body length 8.0���12.0 mm, wing length 7.0��� 9.5 mm, from Roberts (1929). Very similar to male, except frons 2.5�� as wide as ocellar tubercle, with short black hairs on dorsal half (Fig. 8c). Costa without small spines (Fig. 8e). Posterior margin of tergite 7 with slender dorsal median apodeme. Tergite 7 with ventral spines. Tergite 8 with a row of sparse hairs, 12 acanthophorite spines present, lateral spines long, spines in middle short, apex of spines expanded. Furca broad and connected at apex. Sperm pump strong and nearly as long as furca, not clothed in longitudinal muscle or lateral papillae; distal spermathecal duct short but strong; spermatheca elongate, cylindrical (Fig. 9g). Remarks. Thevenetimyia furvicostata (Roberts, 1929) is redescribed and the images of external and internal characters are provided for the male. The allotype of Thevenetimyia furvicostata labelled: ��� ♀, QLD, Stanthorpe, Queensland, 27th January 1927.��� (Roberts, 1929), is not in QM and is probably lost. Although Robert (1929) did not indicate where he deposit the paratype, PFMR found a specimen in NHMUK with the same label data. Despite this specimen has not a paratype label, it is probably the paratype originally designated by Roberts (1929). Distribution. QLD., Published as part of Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos Jos�� Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), pp. 331-375 in Records of the Australian Museum 70 (3) on pages 342-346, DOI: 10.3853/j.2201-4349.70.2018.1678, http://zenodo.org/record/5237395, {"references":["Roberts, F. H. S. 1929. A revision of the Australian Bombyliidae (Diptera). III. Proceeding of the Linnean Society of New South Wales 54: 553 - 583.","Hall, J. C. 1969. Review of the subfamily Cylleniinae with a world revision of the genus Thevenetimyia Bigot (Eclimus auct.) (Diptera: Bombyliidae). University of California Publications in Entomology 56: 1 - 85."]}

Genus Thevenetimyia Bigot Thevenemyia Bigot, 1875 a: 196 [1875b: clxxiv]. Type species: Thevenetimyia californica Bigot, 1875, by monotypy. [Unavailable name; incorrect original spelling.] Thevenetimyia Bigot, 1875 a: 196 [1875b: clxxiv] (justified emendation of Thevenetimyia [by Bigot, 1892: 339]). Type species: Thevenetimyia californica Bigot, 1875, automatic. Epibates Osten Sacken, 1877: 268. Type species: Epibates funestus Osten Sacken, 1877, by subsequent designation (Coquillett, 1910b: 538). Diagnosis. Hall (1969) summarized as ���Quite similar to Eclimus Loew. The chief differences lie in the slightly swollen occiput, the smooth integument, partial development of the alula, the narrowly open anal cell and the absence of transverse abdominal depressions in Thevene[ti]myia.��� Based on the Australian species, we further diagnose this genus as follows: Mostly medium-sized, slender Bombyliinae. Male eyes mostly connected or narrowly separated (Figs 1e, 4e, 7e, 10e, 13e, 16e, 18e, 20d, 23e, 27e). Female frons about 2.5�� as wide as ocellar tubercle (Figs 2d, 8c, 14c, 21c, 24c, 26e). Clypeus swollen. Flagellum one-segmented with apical style. Palp thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, labella thin and filiform (Figs 1f, 2e, 4f, 7f, 8d, 10f, 13f, 14d, 16f, 18f, 20e, 21d, 23f, 24d, 27f). Scutum and scutellum with admixed hairs and scales. Three to four notopleural setae present, postalar seta normally absent. Scutum sometimes with small short spines (Fig. 31d). Anepisternum, katepisternum, metepisternum, and mediotergite with hairs and/or setae, anepimeron, meron and laterotergite bare. Hind femora sometimes with short ventral bristles. Wing membrane various, from hyaline to mostly infuscated to having distinct dark marking on otherwise transparent wings. Female wing colour lighter in some species. Cell r 5 widely open; cell br much longer than cell bm; crossvein m-m long; cell cup narrowly open (Figs 1d, 2c, 4d, 7d, 8e, 10d, 13d, 16d, 18d, 19c, 21e, 23d, 26d, 27d). Male costa often with two rows of small spines (Fig. 31b), absent in females (Fig. 31a). Abdomen slender, tergites often with pale scales on posterior margin, apex tergites sometimes with dense long lateral hairs. Male epandrium wide and short, posterior margin slightly convex (Figs 3d, 5d, 9d, 11d, 15d, 17d, 22d, 25d, 28d). Hypandrium normally present. Gonocoxal apex slightly narrower than the base in dorsal view; gonocoxal apodeme strong and incurved; inner apex of gonocoxite rounded, elongate and broad; outer apex of gonocoxite normal length, rounded; gonostylus slender and pointed dorsally (Figs 3a,b; 5a,b; 9a,b; 11a,b; 15a,b; 17a,b; 22a,b; 25a,b; 28a,b). Female posterior margin of tergite 7 with slender dorsal median apodeme. Tergite 7 with ventral spines. Tergite 8 with a row of sparse hairs, acanthophorite spines present, lateral spines long, spines in middle short, apex of spines expanded. Furca connected at apex. Sperm pump strong and nearly as long as furca, not clothed in longitudinal muscle; distal spermathecal duct short but strong; spermatheca elongate, cylindrical (Figs 3g, 9g, 15g, 25g, 28g). Distribution. Nearctic (Canada, Mexico, USA), Palaearctic (Algeria, Greece, Iran, Japan, Tunisia, Turkey), Afrotropical (Mauritania, Madagascar), Neotropical (Mexico [Oaxaca]) and Australian (all Australian states and territories except TAS). Australian species. Thevenetimyia australiensis Hall, 1969; Thevenetimyia fergusoni Li and Rodrigues, sp. nov.; Thevenetimyia furvicostata (Roberts, 1929); Thevenetimyia infuscata Li and Yeates, sp. nov.; Thevenetimyia longipalpis (Hardy, 1921); Thevenetimyia major Li and Yeates, sp. nov.; Thevenetimyia mimula Hall, 1969; Thevenetimyia nigrapicalis (Roberts, 1929); Thevenetimyia nuri Rodrigues and Lamas, sp. nov.; Thevenetimyia tenta Hall, 1969., Published as part of Li, Xuankun, Rodrigues, Paula Fernanda Motta, Lamas, Carlos Jos�� Einicker & Yeates, David K., 2018, A Review of the Australian Species of Thevenetimyia Bigot, 1875 (Bombyliidae, Bombyliinae, Eclimini), with Description of Four New Species and the Pupal Case of T. longipalpis (Hardy), pp. 331-375 in Records of the Australian Museum 70 (3) on page 333, DOI: 10.3853/j.2201-4349.70.2018.1678, http://zenodo.org/record/5237395, {"references":["Bigot, J. M. F. 1875. Dipteres nouveaux provenant du Chili. Annals of the Entomological Society of France 5 (3): 277 - 308.","Hall, J. C. 1969. Review of the subfamily Cylleniinae with a world revision of the genus Thevenetimyia Bigot (Eclimus auct.) (Diptera: Bombyliidae). University of California Publications in Entomology 56: 1 - 85.","Roberts, F. H. S. 1929. A revision of the Australian Bombyliidae (Diptera). III. Proceeding of the Linnean Society of New South Wales 54: 553 - 583.","Hardy, G. H. H. 1921. Australian Bombyliidae and Cyrtidae (Dipt). Papers and Proceedings of the Royal Society of Tasmania 1921: 41 - 83."]}

2. Marmasoma hortorum sp. nov. (Figs 1���3) Type specimens. Holotype ♂ (WAM), Australia, WA, 12km S Billabong Rdhse. [= Roadhouse], 9 Sep 1996, James O'Grady (WAM: E97791). Paratype ♀ (1), Australia, WA, same data as holotype (WAM: E97792). Other specimens examined. Australia, WA, ♂ (2) ♀ (1), 12km S Billabong Rdhse., 12 Sep 1996, James O'Grady (WAM: ♂ E97790, E97793; ♀ E97789). ♂ (1) ♀ (1), 12km S Billabong Rdhse., 10 Sep 1996, James O'Grady (WAM: ♂ E97795; ♀ E97794). ♂ (5) ♀ (1), Buntine, -29.9812 116.5691, Euc. trees + low shrubs, 11 Sep 2016, J & F Hort (WAM: ♂ E97783 ��� E97784, E97786 ��� E97788; ♀ E97785). ♂ (1), Northampton, -27.7862 114.8348, Low shrubs + yellow sand, 16 Sep 2016, J & F Hort (WAM: E97782). ♀ (1), 11km SW Mt Leake, - 25.47 119.10, 16 Aug 1984, TF Houston & BP Hanich (WAM: E97781). ♂ (1), Mullewa, Sep, LJ Newman (WDFA: 73092). SA, ♂ (1) ♀ (1), 62km N of Pt. Augusta, 2 Sep 1987, MG Jefferies (ANIC No. 29- 041417, 29- 041418). Diagnosis. Wing with pale light markings. Body covered with mostly white to pale yellow scales. Male distiphallus straight, pointed posteriorly, subapex with long hairs. Female common spermathecal duct swollen. Description. Male. Body length 10.0��� 11.8 mm, wing length 7.4���9.0 mm. Head. Head about 1.5 x wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to pale yellow hairs and scales. Eye widely connected, by 2.8 x length of ocellar tubercle. Frons slender and long, upper triangular section small, lower triangular section black with thin pale pruinescence, middle narrow part about 0.6 x length of frons. Ocellar tubercle slightly raised, brown to blackish brown with grey pruinescence, with yellow hairs. Face with sparse pale pruinescence and otherwise bare, parafacial area blackish brown and bare. Gena with sparse pale pruinescence and long white to pale yellow hairs. Clypeus with sparse pale pruinescence and bare. Occiput with short white scales on eye margin and dense long white hairs admixed with some black bristles. Posterior eye margin slightly sinuous. Antennae black; pedicel with thin pale pruinescence, with short black hairs admixed with few yellow scales; scape with short black hairs and with long yellow ventral hairs. Scape 3.5 x as long as wide, and 3.5 x as long as pedicel, uniform from base to apex. Pedicel 1.1 x as long as wide. Flagellum 8.9 x as long as wide, 1.5 x as long as scape + pedicel (3.5 x as long as scape), conical and slightly laterally compressed, three-segmented with apical stylus (Fig. 1c). Palpus thin and long, extending beyond oral cavity, black with blackish brown hairs, two-segmented, with palpal pit. Mouthparts slender, 1.9 x as long as eye length (1.2 x as long as head length), labella thin and filiform (Fig. 1f). Thorax. Integumental colour of scutum and scutellum mostly black with dense grey pruinescence, scutum with two black stripes, either side of midline. Scutum anterior half mostly with white scales admixed with long yellow and black hairs and some black scales, posterior half of scutum and scutellum with admixed white and black scales and long black hairs. Three notopleural setae present. Postalar callus and posterior margin of scutellum with strong hairs. Notopleural setae and strong hairs on postalar callus and scutellum black. Pleura black with thick grey pruinescence, anepisternum with long white scales admixed with strong yellow hairs, dorsal half of katepisternum with long white scales, metepisternum with moderately long white scales, laterotergite and mediotergite with dense long white hairs and strong yellow hairs, anepimeron and meron bare. Legs. Legs black. Coxae with thick grey pruinescence. Femora mostly covered with white scales, except anterior and posterior faces of fore femur with brown scales. Tibiae covered with admixed white and brown scales. Mid femur with some long subapical black bristles, hind femur with two rows of long anteroventral black bristles. Other hairs and bristles on legs long and black. Fore tibia 2.2 x longer than fore basitarsus, mid tibia 2.5 x longer than mid basitarsus, hind tibia 2.3 x longer than hind basitarsus. Wing. Wing membrane mostly hyaline, except the area around base of vein R 4 , crossvein r-m, crossvein dm-cu, and apex of cell bm with light brown spot. Cell r 5 widely open; cell br much longer than cell bm, crossvein r-m arising apical half of cell dm; crossvein m-m long, and 1.5 x as long as crossvein r-m; cell cup open (Fig. 1d). Halter stem dark yellow, knob dark yellow. Abdomen. Integumental colour of tergites black with thick grey pruinescence. Tergites mostly covered with pale yellow and black scales, pale yellow scales denser on lateral and posterior margin, black scales denser on central area. Tergite 1 with black hairs on posterior margin, tergites 4���8 with black ventrolateral hairs. Sternites with dark brown integumental colour, with white scales admixed with black hairs on posterior margin. Genitalia. Epandrium nearly rectangular, anterior margin with a deep notch, posterior margin slightly concave, anteroventral corner with long stem (Fig. 3d). Hypandrium absent. Gonocoxite slender and elongate. Gonocoxal apex as wide as the base in dorsal view, ejaculatory apodeme small (Figs 3a, b); gonocoxal apodeme strong and incurved; lateral ejaculatory process small; inner apex of gonocoxite rounded; outer apex of gonocoxite rounded; dorsal bridge without lateral hollow; gonostylus bifid apically, 1.5 x longer than wide; phallus long and strongly curved, subapex with long hairs (Fig. 3c). Female. Body length 9.3���11.0 mm, wing length 7.1���9.1 mm. Very similar to male, except frons black with thick pale pruinescence, 2.5 x as wide as ocellar tubercle, frons with pale yellow hairs admixed with yellow and black bristles (Fig. 2c). Tergite 9+10 with some sparse hairs, 12 acanthophorite spines present, lateral spines long, spines in middle short. Furca straight and broad. Sperm pump strong and nearly as long as furca, clothed in longitudinal muscle, without lateral papillae; sperm pump apical and basal collars absent (Fig. 3j); common spermathecal duct swollen (Fig. 3i); spermatheca elongate, cylindrical with apex slightly swelled (Fig. 3g). Remarks. Marmasoma hortorum sp. nov. is similar to Marmasoma sumptuosum, but differs as follows: wing markings light, inconspicuous; body covered with mostly white to pale yellow scales; male distiphallus straight, pointed posteriorly, subapex with long hairs; female common spermathecal duct swollen. Distribution. Australia (SA, WA). Etymology. This species is named in honor of Fred and Jean Hort for their important contributions to photographing and collecting Diptera specimens from Western Australia., Published as part of Li, Xuankun & Yeates, David K., 2018, One hundred years of solitude ended: A second species of Marmasoma White, 1916 (Diptera, Bombyliidae, Bombyliinae, Eclimini) from Australia, pp. 34-42 in Zootaxa 4420 (1) on pages 34-42, DOI: 10.11646/zootaxa.4420.1.2, http://zenodo.org/record/1253131

1. Marmosoma sumptuosum White, 1916 Marmasoma sumptuosa White, 1916: 189���190, lectotype male (BMNH) designated by Rodrigues et al., 2017. Hardy, 1921: 63, 83, plate XVI; Tonnoir, 1927: 102; Roberts, 1929: 579. Marmasoma sumptuosum Hull, 1973: 239, 241, 471, 490, 514, 519, 553; Theodor, 1983: 85; Yeates, 1994: 21, 31, 125; Evenhuis & Greathead, 1999: 209; Evenhuis & Greathead, 2015: 219; Rodrigues et al., 2017: 185. Diagnosis. Wing with conspicuous dark spots. Body covered with mostly dark yellow scales. Male distiphallus Sshaped, curved dorsally, with serrated projections on dorsal and ventral surfaces of apical third. Female common spermathecal duct normal, not swollen. Specimens examined. Australia, ACT, ♂ (4), Blundell's, 6 Jan 1961, SJ Paramonov (ANIC No. 29- 041327 ��� 29- 041330). ♂ (3), Blundell's, 7 Nov 1947, SJ Paramonov (ANIC No. 29- 041331 ���29- 041333). ♂ (1), Mt. Gingera, 5500 ft., 10 Jan 1952, IFB Common (ANIC No. 29- 041334). ♂ (2), Mt. Gingera, 16 Jan 1952, KR Norris (ANIC No. 29- 041335, 29- 041336). ♂ (1), Mt. Gingera, 31 Jan 1952, SJ Paramonov (ANIC No. 29- 041337). ♀ (1), Mt. Gingera, 3 Jan 1952, SJ Paramonov (ANIC No. 29- 041338). ♂ (1), Paddy's Rv., 3 Nov 1948, SJ Paramonov (ANIC No. 29- 041339). ♀ (1), Lee's Springs, 3 Jan 1951, SJ Paramonov (ANIC No. 29- 041340). ♀ (1), Lee Spring, 13 Nov 1938, AL Tonnoir (ANIC No. 29- 041341). ♀ (1), Bendora, 7 Jan 1952, KR Norris (ANIC No. 29- 041342). ♂ (1), Mt. Gingera, 4 Feb 1952, Z Liepa (ANIC No. 29- 041343). ♀ (1), Mt. Gingera, 1650 m, 12 Jan 1985, MG Jefferies (ANIC No. 29- 041344). ♂ (2) ♀ (2), Nr. Bendora Hut Brindabella Ra., 35.25S 148.48E, 20 Jan 1997, MG Jefferies (ANIC No. 29- 041345 ���29- 041348). ♂ (9) ♀ (1), Yaouk Trail, South of Cotter Hut, Namadgi National Park, Open Eucalyptus forest, Hand Net. 35��39'19''S 148��49'44''E. 10 Dec 13, D Yeates, C Manchester, K Musemann (ANIC No. 29- 041349 ���29- 041358). NSW, ♂ (3), Tinderry Mountains, roadside puddles, netted, Tinderry Rd, 35��44'04.91''S 149��17'04.97''E, 1217 m, 21 Dec 2014, DJ Ferguson (ANIC No. 29- 041359 ���29- 041361). ♂ (1), Snowy-Thredbo Junction, 8 Nov 1961, DH Colless (ANIC No. 29- 041362). ♂ (1), Alpine Ck., Snowy Mt., Hwy., 2 Feb 1965, Z Liepa (ANIC No. 29- 041363). ♀ (1), Round Hill Nature Res, 5-24 Oct 2003, malaise across dry gully tall Callitrus / Eremopila forest, C Lambkin, N Starick, 184 m, ANIC Bulk Sample 2164, 33��02'45''S 146��14'33''E (GPS) (ANIC No. 29- 041364). ♂ (1), Kiandra, Aline Ck., 9 1933, HJ Willings (ANIC No. 29- 041365). ♂ (4), Tinderry Nat. Res., intersection East Tinderry & Horse Flat Fire Trails, 13.5 km, ENE Michelago, hand net near creek, 19 Dec 2004, 1029 m, C Lambkin, N Starick, A Zwick, 35��40'45''S 149��18'31''E GPS (ANIC No. 29- 041366 ���29- 041369). ♂ (1), South Bliack Range 9 km E. Hoskinstown SE NSW, 1130 m, Netted, 2 Jan 2006, DJ Ferguson (ANIC No. 29- 041370). ♂ (1), South Bliack Range 14.5 km, 1130 m, 16 Nov 2004, DJ Ferguson (ANIC No. 29- 041371). ♂ (1), Kiandra, 7 Jan 1984, MG Jefferies (ANIC No. 29- 041372). ♀ (1), National Park, 27 Sep 1925, Mackerres (ANIC No. 29- 041373). SA, ♂ (1) ♀ (3), Kangaroo Is, Bales Bay, 36.00S 137.22E, 26 Oct 1993, DK Yeates (ANIC No. 29- 041374 ���29- 041377). VIC, ♀ (1), Martin's Ck, East Gippsland, 9 Jan 1999, S Thiele (ANIC). ♂ (1), Frankston, 28 Sep 1918, CE Cole (NMV). ♀ (1), Beech Forest, 11- 19 Jan 1932, FE Wilson (NMV). ♂ (1), Dandenong Ranges, Nov 1898 (NMV). ♂ (1) ♀ (1), Gippsland (NMV). ♀ (1), Noorinbee, 12 Nov 1969, A Neboiss (NMV). ♂ (6) ♀ (4), Dandenong Ranges, 6 Nov 1898 (NMV). ♂ (1), Cement CK, 23 Feb 1957, A Neboiss (NMV). ♂ (1), Lake Mountain, 16 Jan 1965, A Neboiss (NMV). ♀ (1), Lake Mountain, 17 Jan 1965, A Neboiss (NMV). TAS, ♂ (6) ♀ (7), Meredith R. 12 mls., from Corinna, 6 Jan 1954, TG Campbell (ANIC No. 29- 041379 ���29- 041391). ♂ (1) ♀ (6), Meredith R. 12 mls., from Corinna, 5 Jan 1954, TG Campbell (ANIC No. 29- 041392 ���29- 041398). ♂ (4) ♀ (3), Meredith R. 12 mls., from Corinna, 4 Jan 1954, TG Campbell (ANIC No. 29- 041399 ���29- 041405). ♂ (3) ♀ (6), Cradle Mountain NP 1 km N Dove Lake, 7 Jan 1992, ME Irwin, BJ Irwin (ANIC No. 29- 041406 ���29- 041414). ♀ (2), Franklin-Gordon Wild River National Park, Donaghy's Hill, 42��11'52''S 145��55'55''E, 7 Jan 1999, J & A Skevington, M Matheison (ANIC No. 29- 041415, 29- 041416). ♀ (1), Tyenna, 11 Dec 1914, CE Cole (NMV). ♂ (1), Tyenna, 20 Dec 1917, CE Cole (NMV). ♂ (1), Tyenna, 21 Dec 1917, CE Cole (NMV). Distribution. Australia (ACT, NSW, SA (Kangaroo Island), TAS, VIC). Remarks. Hull (1973: 239) was aware that the genus occurred in Western Australia, but misidentified his Western Australian specimens as M. sumptuosum. An unverified record of this species from Queensland (Evenhuis & Greathead, 1999) was followed in the literature by Rodrigues et al. (2017)., Published as part of Li, Xuankun & Yeates, David K., 2018, One hundred years of solitude ended: A second species of Marmasoma White, 1916 (Diptera, Bombyliidae, Bombyliinae, Eclimini) from Australia, pp. 34-42 in Zootaxa 4420 (1) on pages 34-42, DOI: 10.11646/zootaxa.4420.1.2, http://zenodo.org/record/1253131, {"references":["White, A. (1916) The Diptera-Brachycera of Tasmania. Part III. Families Asilidae, Bombyliidae, Empidae, Dolichopodidae, & Phoridae. Papers and Proceedings of the Royal Society of Tasmania, 1916, 148 - 266. https: // doi. org / 10.5962 / bhl. part. 28993","Rodrigues, P. F. M., McAlister, E. & Lamas, C. J. E. (2017) Review of Marmasoma White with lectotype designation of M. sumptuosum White and an identification key to the Australasian / Oceanian genera of Ecliminae (Diptera, Bombyliidae). Zootaxa, 4232 (2), 185 - 196. https: // doi. org / 10.11646 / zootaxa. 4232.2.3","Hardy, G. H. H. (1921) Australian Bombyliidae and Cyrtidae (Dipt.). Papers & Proceedings of the Royal Tasmanian Society, 1921, 41 - 83.","Tonnoir, A. L. (1927) Descriptions of new and remarkable New Zealand Diptera. Records of the Canterbury Museum, 3, 101 - 112.","Roberts, F. H. S. (1929) A revision of the Australian Bombyliidae (Diptera). III. The Proceedings of the Linnean Society of New South Wales, 54, 553 - 83.","Hull, F. M. (1973) Bee flies of the world. The genera of the family Bombyliidae. Smithsonian Institution Press, Washington DC, 687 pp. https: // doi. org / 10.5962 / bhl. title. 48406","Theodor, O. (1983) The Genitalia of Bombyliidae (Diptera). Israel Academy of Sciences and Humanities, Jerusalem, 275 pp.","Yeates, D. K. (1994) The cladistics and classification of the Bombyliidae (Diptera: Asiloidea). Bulletin of the American Museum of Natural History, 219, 1 - 191.","Evenhuis, N. L. & Greathead, D. J. (1999) World Catalog of Bee Flies (Diptera Bombyliidae). Backhuys Publishers, Leiden, 756 pp.","Evenhuis N. L. & Greathead D. J. (2015) World Catalog of bee flies (Diptera: Bombyliidae). Netherlands: Backhuys Publishers Leiden. Available from: http: // hbs. bishopmuseum. org / bombcat / bombcat-revised 2015. pdf (Accessed 10 December 2017)"]}

Marmasoma White, 1916 Marmasoma White, 1916: 184, 188. Type species: Marmasoma sumptuosum White, 1916, by monotypy. Diagnosis (modified from Rodrigues et al., 2017). Scape with numerous very long hairs, longer on ventral surface and laterally; face flat, not projecting in lateral view; occiput tumid, with bristles. Thorax postpronotal lobe and antepronotum with long and strong bristles; anepisternum with short scales admixed with strong hairs on dorsal margin; laterotergite and mediotergite with long and strong hairs and bristles. Abdomen decumbent with apex raised in male; tergites covered in scales. Male epandrium anterior margin with a deep notch, anteroventral corner with long stem; hypandrium absent; gonocoxite slender and elongate; gonocoxal apex as wide as the base in dorsal view, ejaculatory apodeme small; gonocoxal apodeme strong and incurved; lateral ejaculatory process small; inner and outer apexes of gonocoxite rounded; dorsal bridge without lateral hollow; gonostylus bifid apically; phallus long and strongly curved. Female sand chamber present; tergites 9 and 10 fused; acanthophorite spines not in a row; furca broad; sperm pump strong; sperm pump apical and basal collars absent; spermatheca elongate, cylindrical with apex slightly swollen. Distribution. ACT, NSW, SA, VIC, TAS, WA. Species included. Marmasoma sumptuosum and Marmasoma hortorum sp. nov., Published as part of Li, Xuankun & Yeates, David K., 2018, One hundred years of solitude ended: A second species of Marmasoma White, 1916 (Diptera, Bombyliidae, Bombyliinae, Eclimini) from Australia, pp. 34-42 in Zootaxa 4420 (1) on pages 34-42, DOI: 10.11646/zootaxa.4420.1.2, http://zenodo.org/record/1253131, {"references":["White, A. (1916) The Diptera-Brachycera of Tasmania. Part III. Families Asilidae, Bombyliidae, Empidae, Dolichopodidae, & Phoridae. Papers and Proceedings of the Royal Society of Tasmania, 1916, 148 - 266. https: // doi. org / 10.5962 / bhl. part. 28993","Rodrigues, P. F. M., McAlister, E. & Lamas, C. J. E. (2017) Review of Marmasoma White with lectotype designation of M. sumptuosum White and an identification key to the Australasian / Oceanian genera of Ecliminae (Diptera, Bombyliidae). Zootaxa, 4232 (2), 185 - 196. https: // doi. org / 10.11646 / zootaxa. 4232.2.3"]}