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Cisplatin-based chemotherapy is the most recommended treatment for metastatic urothelial cancer (mUC). However, about 50% of patients are considered to be cisplatin ineligible. Anti-programmed cell death protein 1/programmed death-ligand 1 (PD-L1) therapies have, nevertheless, increased the options available to clinicians and are especially valuable for treating these patients. This study therefore tested the activity and safety of avelumab as first-line therapy for mUC.Patients with mUC who were ineligible for cisplatin-based chemotherapy were screened centrally for PD-L1 expression and only those with a tumour proportion score ≥ 5% were enrolled in the trial. The primary endpoint was 1-year overall survival (OS), and the secondary endpoints were median OS, median progression-free survival, overall response rate, duration of the response, safety and tolerability. All the survival rates were estimated with the Kaplan-Meier product-limit methodology and compared across groups using the log-rank test.A total of 198 patients were screened, with 71 (35.9%) whose PD-L1 expression was ≥5% enrolled in the study. The median age was 75 years, bladder cancer was the primary tumour in 73.2% of cases and 25.3% had liver metastases. The main reasons for the cisplatin ineligibility were a low rate of creatinine clearance (60 ml/min), present in 70.4% of patients, and an Eastern Cooperative Oncology Group performance status of 2, which affected 31%. The median OS was 10.0 months (95% confidence interval 5.5-14.5 months) and 43% of patients were alive at 1 year. A complete response was achieved in 8.5% of cases, and 15.5% had a partial response. Adverse any-grade and high-grade events occurred in 49.3% and 8.5% of patients, respectively. A grade 3 infusion reaction was the only high-grade treatment-related adverse event. No treatment-related deaths were reported.This ARIES trial confirmed the activity and safety of avelumab for treating mUC, adding a new therapy option to the armamentarium of checkpoint inhibitors already approved for platinum-ineligible, locally advanced/mUC.

Haploops similis Stephensen 1925 Type species collected by the R. V “ Ingolf”, one specimen, (sex?) length: 4.5 mm, Station 36: 61°50’N – 56°21’W, depth 2702 m, bottom: sediment type unknown. BIOICE material examined. Station 2314: nine specimens, 11September 1992, depth 156 m, 63° 42.16’N – 23°03.50’W; bottom:muddy sand.Station 2328: four specimens, 3 May 1993, depth 429m, 63°20.00’N – 10°57.00’W; bottom: sand. Station 2616: four specimens, 11 July 1994, depth 535 m, 67°11.38’N – 16°50.40’W; bottom: sandy silt. Station 2873: one specimen, 22 August 1996, depth 555 m, 64°37.41’N – 27°14.28’W; bottom: gravely sand. Station 2946: two specimens, 28 August 1996, depth 229 m, 65°47.90’N – 25°38.70’W; bottom: sandy silt. Station 3026: five specimens, 8 July 1997, depth 564 m, 63°53.96’N – 12°44.18’W; sediment type unknown. Station 3099: two specimens, 21August 1999, depth 229 m, 67°11.02’N – 21°45.68’W; bottom: gravely sandy silt. Station 3282: four specimens, 16 September 2001, depth 1808 m, 62°48.00’N – 16°14.80’W; bottom: sandy silt. Station 3531: two specimens, 9 September 2002, depth 1712 m, 62°43.26’N – 14°34.70’W; bottom: sandy silt. Station 3544: 16 specimens, 11September 2002, depth 1632 m, 61°33.005’N – 13°40.14’W; bottom: gravely sand. Station 3598: one specimen, 10 September 2003, depth 768 m, 62°17.37’N – 26°37.58’W; bottom: silty sand. All specimens of these stations are females. Diagnosis. Blind species; body without long dorsal setae on the pereon, pleon and urosome; A1 = 8–9/10 A2, A2 = 2/3 body; coxa 4 heart-shaped, Pereopod 7: basis narrow, anterior distal lobe of the carpus developed. Description. Head: Square shaped, without corneal lenses, anterior margin oblique and straight. Antenna 1 length is 8–9/10 of Antenna 2 length; Antenna 2 length is 2/3 body length. Pereon: without long dorsal setae. Gnathopod 1: coxa 1 roughly oval, distal margin rounded and fringed with long setae; basis long, slightly curved, approximately rectangular with many long setae, as long as ischium + merus + carpus, the merus, carpus and propodus bearing long setae; propodus oval. Gnathopod 2: coxa 2 approximately rectangular, distal margin straight with few setae; basis long, slightly curved, with many long setae, as long as ischium + merus + carpus; the merus, carpus and propodus bearing long setae; propodus oval. Pereopod 3: coxa 3 approximately rectangular, distal margin straight with few setae; basis, merus and carpus with long setae on the posterior margin; dactylus slender, curved and longer than propodus. Pereopod 4: coxa 4 heart-shaped, basis, merus and carpus with long setae on the posterior margin, and tuft of long setae on distal anterior margin of merus and propodus; dactylus slender, curved and shorter than propodus. Pereopod 5: coxa 5 rectangular, basis roughly rectangular with few setae on the anterior margin; carpus rectangular, anterior margin with few setae, posterior margin with rows of little spines, postero-distal lobe with short spines and a long one; propodus rectangular, longer than carpus, with few short setae on anterior margin, and long distal setae; dactylus short and curved. Pereopod 6: basis rounded with few short setae on the anterior margin; carpus rectangular, posterior margin with rows of little spines, postero-distal lobe ornamented with short spines and a very long one; propodus a little longer than carpus, with few short setae on anterior margin, and long distal setae; dactylus short and curved. Pereopod 7: basis narrow, anterior and posterior margin slightly concave, lobe not deflected, rounded with few long setae, not reaching the ischium-merus joint; ischium quadrangular; merus rectangular with an antero-inferior lobe, anterior and posterior margin ornamented with few little spines and long postero-distal setae; carpus oval, with spines on the anterior and posterior margin, anterior distal lobe developed; propodus and dactylus short and narrow. Pleon: without long dorsal setae. Epimeral plate 1: anterior margin oblique and straight, ventral margin and posterior margin straight, corners rounded. Epimeral plate 2: inferior and posterior margins slightly convex, postero-inferior corner rounded. Epimeral plate 3: anterior margin straight, posterior margin oblique and straight, inferior one slightly convex, anterior corner rounded, posterior corner slightly acute. Urosome: carina small. Uropod 1: the rami are long and slender, of unequal length (2/3). Uropod 2: rami short, of equal size, and armed with a row of spines on each ramus. Uropod 3: peduncle short and strong; rami of unequal length, ornamented with numerous setae. Telson: triangular, apically rounded, cleft on 2/3 of the length; two setae at the apex of each lobe. Distribution: North Atlantic Ocean; wide bathymetric range species: this species was collected at 2702m depth (Stephensen,1925), between 100–2900 m (Mills,1971), between 156–1808 m, on sand, sandy silt, silty sand, muddy sand, gravely sand, gravely sandy silt (BIOICE samples), one specimen at 1024 m on bathyal mud in the south of the Bay of Biscay (Dauvin & Bellan-Santini,1996). Taxonomic remarks. H. similis belongs to the sub-group of blind species with a narrow Pereopod 7 basis and without dorsal tuft setae, which includes six other species: H. abyssorum, H. vallifera, H. lodo, H. dauvini, H. bjarnii, and H. faroensis. It differs from these species by having: — Antennae: Antenna 1= 8–9/10 Antenna 2, Antenna 2 = 2/3 body length; the antennae are longer than the body length for H. bjarnii; they are half body length for H. vallifera, H. dauvini, and H. faroensis; for H. lodo, A1= ped A2, A2 shorter than body (in Barnard, 1961). — Coxa 4 heart-shaped (also the case for H. bjarnii and H. vallifera); but square shaped for H. abyssorum, H. lodo, H. dauvini and H. faroensis. — Epimeral plates 3 with posterior margin oblique and straight (also for H. faroensis); slightly convex for H. bjarnii and H. dauvini; rounded for H. vallifera and H. lodo (not indicated by Chevreux 1908 for H. abyssorum). — Uropod 1: the rami are long and slender, of unequal length (2/3); the rami are subequal for H. bjarnii, H. vallifera; H. lodo and H. dauvini; they have the same size for H. abyssorum and H. faroensis., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 166-168, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Mills, E. L. (1971) Deep-Sea Amphipoda from the Western North Atlantic Ocean, the family Ampeliscidae. Limnology and Oceanography, 16, 357 - 386. https: // doi. org / 10.4319 / lo. 1971.16.2.0357","Dauvin, J. C. & Bellan-Santini, D. (1996) Ampeliscidae (Amphipoda) from the Bay of Biscay. Journal of Crustacean Biology, 16, 149 - 168. https: // doi. org / 10.2307 / 1548938","Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de l a Princesse Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13."]}

Haploops vallifera Stephensen 1925. (Figures 9–12) BIOFAR material examined. Station 124, one specimen, 26 July 1987, depth 600 m, 62°16.94’N – 09°38.93’W; sediment type unknown. Station 261, nine specimens, 14 May 1988, depth 1003 m, 61°35.57’N – 09°35.47’W; sediment type unknown. Station 731, 182 specimens, 29 September 1990, depth 1042 m, 60°29.70’N – 07°14.10’W; bottom: sand, gravel, cobbles and stones. Station 738, one specimen, 1 October 1990, depth 749 m, 62°19.30’N – 10°13.30’W; bottom: gravel, cobbles and stones. Male unknown. Diagnosis. Body with few small dorsal setae on the pereon and the pleon. Corneal lenses absent. Segments of dorsal side of mesosome, marked. Urosome segment 1 with a small rounded carena. Epimeral plate 3 rounded. The two pairs of antennae of equal length not longer than half of body. Description. (Fig. 9–12). Adult female (Fig. 9) length 6 mm Biofar 731. Head: nearly square, with a small rostrum, without corneal lenses, blind species, straight lateral lobe. Antenna 1 (Fig. 9, 10 A): nearly equal to half of the body length. Peduncle with article 3 Antenna 2 (Fig. 9, 10 B): Equal to antenna 1 in length. Peduncle segment 5> 4. Flagellum with 12 articles. Mandible (Fig. 10 C): molar strong, palp very long, article 2> article 3, artcles 2, 3 bear long setae, article 3 with a long apical seta. Lower lip bilobate (Fig. 10 F). Maxilla 1 (Fig. 10 E): inner plate conical, outer plate with spine teeth distally, last article of palp ended with some setae and 5 strong teeth. Maxilla 2 (Fig. 10 G): the two plates are subequal in length and apically densely setose. Maxilliped (Fig. 10 D): inner plate sub-rectangular with short setae on the lateral and distal part. Internal margin of the outer plate with numerous long setae and strong teeth. Palp elongate, longer than outer plate, articles 2 and 3 with long setae, article 3 distally enlarged, dactylus elongate slender with 4 setae at the inferior margin. Pereon: small setae on the posterodorsal part of segment 6 and 7 (Fig. 9). Gnathopod 1 (Fig. 11A): longer than gnathopod 2. Coxa 1 as long as basis, distal margin rounded and fringed with long setae. Basis a little longer than carpus+propodus, setose. Merus, carpus and propodus bears long setae, dactylus curved with 5 short setae. Gnathopod 2 (Fig. 11B): Coxa 2 shorter than basis, triangular, distal margin pointed. Basis long with long setae, egal to merus+carpus+propodus. Carpus longer than propodus, roughtly rectangular with long setae mainly on the inferior margin. Propodus roughtly rectangular. Dactylus curved and ornamed with 5 setae. Pereopod 3 (Fig. 11C): Coxa 3 triangular, distal margin pointed. Basis rectangular, basis = ischium + merus + carpus, the margins of the different articles except dactylus, bearing long setae. Dactylus strong, curved, very long, longer than propodus. Pereopod 4 (Fig. 11D): Coxa 4 pointed, shorter and wider than coxa 3, posteriorly excavate, ventral margin straight with one seta on the distal part. Basis rectangular, longer than ischium + merus + carpus. Dactylus curved, as long as propodus. Long setae are present on the margins of all articles except dactylus. Pereopod 5 (Fig. 11E): coxa 5 bilobate, posterior margin rounded. Basis oval anterior margin fringed with short setae. Carpus sub-rectangular with a small postero-distal lobe, posterior and distal margins ornamented with 4 rows of spines. Propodus rectangular, longer than carpus. Dactylus very strong and curved. Pereopod 6 (Fig. 11F): Coxa 6 triangular, distal edge rounded. Basis pyriform with a small indentation at the posterodistal corner. Carpus rectangular with a small distal lobe, posterior and distal margins ornamented with 3 rows of spines. Pereopod 7 (Fig. 12A, B): coxa rectangular, distally rounded. Basis broad, width / length = 1/2, anterior and posterior margins with numerous long setae. Ischium quadrangular with 2 small strong spines on the posterior margin. Merus quadrangular but longer than width. Carpus shorter and narrower of merus with inferior corner elongate. Propodus and dactylus short. Pleon: Segment 1 and 2 with short setae (Fig. 9). Epimeral plate 1: short, ventral margin rounded. Epimeral plate 2 (Fig. 12 F): Rounded distal margin with 2 setae. Epimeral plate 3 (Fig. 12 F): anterior margin straight, anteroventral corner round, vental margin slightly convex, postero-ventral corner rounded, posterior margin convex with 4 setae. Urosome (Fig. 9): segment 1 with a small hump anteriorly at a high rounded dorsal carina. Uropod 1 (Fig. 12C): long, the rami unequal and slightly curved, rami inermous, peduncle longer than rami (peduncle/outer ramus/inner ramous = 40/36/30), ornamented with only one seta at the inner distal corner. Uropod 2 (Fig. 12D): rami subequal, equal to peduncle, one spine on the distal inner corner of the peduncle and 3 on the edge of the inner ramus. Uropod 3 (Fig. 12E): peduncle short and strong, rami foliaceous, longer than peduncle, inner ramus with 2 strong spines, outer ramus with long apical setae and some ones on the margin. Telson (Fig. 12 G): Triangular, rounded, slightly wide than long, cleft on 3/4 of the length, 2 or 3 long setae on each lobe. Known distribution: North Atlantic Ocean; wide bathymetric range species. BIOFAR material, this study, bathymetric range: 600–1042 m; the nature of the bottom is indicated for only two samples: sand, gravel, cobbles and stones. Offshore Iceland, 913–1960 m (Dauvin and Bellan-Santini,1990). Faroe Islands: 600-1098 m (Dauvin 1996), Iceland one station 1392 m (Bellan-Santini and Dauvin, 1997), south and west of Iceland, 13 stations from 285 to 1963 m (Dauvin et al., 2012). Taxonomic remarks. H. vallifera belongs to the sub-group of blind species with a narrow Pereopod 7 basis and without dorsal tuft setae which includes six other species: H. abyssorum, H. similis, H. lodo, H. dauvini, H. bjarnii, and H. faroensis. It differs from these species by having: — Dorsal side of pereon and urosome carinate. — Coxa 1 rectangular with the distal margin rounded. — Coxa 2 triangular, with the distal margin pointed., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 162-166, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Dauvin, J. C. & Bellan-Santini, D. (1990) An overview of the amphipod genus Haploops (Ampeliscidae). Journal of the Biological Association of the United Kingdom 70, 887 - 903. https: // doi. org / 10.1017 / S 0025315400059129","Bellan-Santini, D. & Dauvin, J. C. (1997) Ampeliscidae (Amphipoda) from Iceland with a description of a new species: Ampelisca islandica (Contribution to the BIOICE programme research). Journal of Natural History, 31, 1157 - 1173. https: // doi. org / 10.1080 / 00222939700770621","Dauvin, J. C., Alizier, S., Weppe, A. & Gudmundsson, G. (2012) Diversity and zoogeography of Icelandic deep-sea Ampeliscidae (Crustacea: amphipoda). Deep Sea Research I, 68, 12 - 23. https: // doi. org / 10.1016 / j. dsr. 2012.04.013"]}

(IF 1.02; Q2); International audience; Two new Haploops species are described from the North Atlantic Ocean: a blind species Haploops faroensis spec. nov. and Haploops truncata spec. nov. with a single pair of corneal lenses. In addition, Haploops vallifera Stephensen 1925 and Haploops similis Stephensen 1925, are re-described and the status of Haploops spinosa Shoemaker 1931, is reestablished as a valid species. A table is given of the 75 morphological characters of the studied species.

Haploops truncata spec. nov. (Figures 5–8) Type material. HOLOTYPE. One female without oostegite. Length: 5.57 mm (Fig. 5). BIOICE: Station 3140; one specimen, 25 August 1999, depth: 768 m, 67°51.90’N – 22°14.89’W, bottom: sediment type unknown. Holotype IINH 42245 + slides IINH 42247. The specimen is deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2011: one specimen, 20 July 1991, depth: 768 m, 65°41.43’N – 11°16.77’W; sediment type unknown (IINH 42251). Station 2414: three specimens, 20 July 1991, depth 978 m, 65°35.01’N – 10°59.98’W; bottom: brown sandy, silt mixed with foraminifera (IINH 42250). Station 2897: eight specimens, 24 August 1996, depth 672 m, 65°29.44’N – 27°32.55’W; sediment type unknown (IINH 42249). Station 2898: one specimen, 24 August 1996, depth 672 m, 65°29.30’N – 27°32.70’W; sediment type unknown (IINH 42248). Station 3140: one specimen, 25 August 1999, depth 768 m, 67°51.90’N – 22°14.89’W; sediment type unknown (IINH 42245). Station 3501: two specimens, 31 August 2002, depth 829 m, 62°59.84’N – 20°30.25’W; sediment type unknown (IINH 42246). Male unknown. Ethymology. The species name refers to the shape of the head (square) with the lateral lobe truncate. Diagnosis. Pereon, Pleon and Urosome without long dorsal setae, Head square with anterior margin straight (lateral lobe truncate). One pair of superior corneal lenses. Antenna 1 length = 0.8 Antenna 2 length; Antenna 2 as long as body length (0.876). Coxa 4 external side bearing numerous short setae. Description. Holotype. Female without oostegites. Length: 5.57 mm (Fig. 5). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 6A): square with anterior margin straight, one pair of superior corneal lenses. Antenna 1 (Fig. 6B): shorter than antenna 2, peduncle of A1 with article 3 Antenna 2 (Fig. 6C): longer than antenna 1 (A1/A2 = 0.81). Peduncle segment 5 longer than 4 (23/19). Flagellum with 23 articles. Antenna 2 near the body length. The two antennae bearing long numerous setae (Fig. 6B, C). Upper lip and Lower lip bilobate (Fig. 6D, G). Mandible (Fig. 6H): molar strong; palp long, article 2 and 3 of same length; setae are present on the margin of each article, article 3 with very long apical setae. Maxilla 1 (Fig. 6E). the inner plate is conical shaped with two apical setae and some short hairs; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 6F): dense apically setae on the two plates. Maxilliped (Fig. 6J): inner plate sub-rectangular, elongated, with few distal setae; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, article 2 and 3 with long setae, article 3 triangular shaped, dactylus elongate and slender. Pereon: without long dorsal setae (Fig. 5). Gnathopod 1 (Fig. 7A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis long, slightly curved, approximately rectangular with some setae on the two margins, length longer than ischium + merus + carpus, the merus, carpus and propodus bearing long setae; propodus oval; dactylus slender and curved with small setae, dactylus length = 0.51 propodus length. Gnathopod 2 (Fig. 7B): coxa 2 approximately rectangular and short (coxa length / basis length= 65/85), distal margin slightly rounded with few short setae; basis long with some setae, basis longer than ischium + merus +carpus; carpus longer than propodus (carpus length / propodus length = 53/30), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 7C), dactylus length = 0.46 (17/37) propodus length. Pereopod 3 (Fig. 7D): coxa 3 triangular, distal margin rounded; basis rectangular, basis> ischium + merus + carpus. The margins of the different articles of pereopod 3, except dactylus, bearing few long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 1.35) (27/20). Pereopod 4 (Fig. 7E): coxa 4 square-shaped, external side bearing numerous short setae; anterior margin straight, length / width = 43/40, antero-ventral corner quadrate but not sharp, ventral margin straight with short setae, posterior concavity = 0.46 (17/37) length of the posterior length of the coxa 4, posterior hook broad, length = 17/42 width of coxa 4, posterior corner blunted; basis length> ischium + merus; the margins of the different articles of pereopod 4, except dactylus, bearing few long setae; dactylus slender, slightly curved and longer than propodus (dactylus length / propodus length = 1.125 (27/24). Pereopod 5 (Fig. 7F): coxa 5 rectangular, basis roughly rectangular with few setae on the anterior margin; carpus rectangular, anterior margin with few setae, posterior margin with two rows of little spines, distal margin with short spines and a long one; propodus rectangular, longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 0.23 (12/51). Pereopod 6 (Fig. 8A): coxa 6 roughly trapezoidal shaped, posterior margin rounded; basis rounded with few short setae and spines on the anterior margin; carpus rectangular, anterior margin with few short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a very long one; propodus a little longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 11/40. Pereopod 7 (Fig. 8B): coxa 7 roughly rectangular; basis narrow (length without lobe / width = 80/35 = 2.28), anterior and posterior margin slightly concave, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with an antero-inferior lobe ending with a spine, posterior margin ornamented with few little spines; carpus pyriform (width / length = 17/20), with strong spines on the anterior and posterior margin (propodus length / carpus length = 15/20; propodus + dactylus / carpus = 23/20); propodus narrow, width / length = 5/15 (propodus width / carpus width = 5/17), with short apical setae; dactylus rectangular, short and narrow (length / width = 7/1) with two very small apical setae (dactylus length / propodus length = 8/15). Pleon (Fig. 5): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 8C): anterior margin oblique and straight, ventral margin rounded, posterior margin slightly convex. Epimeral plate 2 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, postero-inferior corner rounded. Epimeral plate 3 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, corners rounded. Urosome (Fig. 5): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 8D): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the ramus, with one spine on the distal margin. length rami / length peduncle = 54–56 / 64. Uropod 2 (Fig. 8E): shorter than Uropod 1; rami triangular and short, inner ramus shorter than the outer one, with a row of spines on each ramus, outer ramus length / peduncle length = 78 /102; peduncle rectangular, robust with a strong spines on the distal margin. Uropod 3 (Fig. 8F): peduncle short and strong (length / width = 58/35); rami of unequal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 90–100/58); inner ramus with three spines and some apical setae, outer ramus with apical long setae and also some long setae on the outer margin. Telson (Fig. 8G): triangular, apically rounded, cleft on 25/35 of the length; one seta present on the apical part of each lobe. Distribution: North Atlantic Ocean; this species was collected between 672 m and 978 m. The nature of the bottom is indicated for only one sample: brown sandy silt mixed with foraminifera. Taxonomic remarks. H. truncata is probably a non mature female, but the original characters in the sub-group with superior corneal lenses justify its description as a new species. H. truncata belongs to the sub-group with the superior pair of corneal lenses clearly visible; the inferior pair is absent; a narrow Pereopod 7 basis, and the absence of dorsal tuft, this sub-group includes six other species: H. tubicola, H. spinosa, H. descansa, H. fundiensis, H. oonah, and H. antennata. H. truncata differs from the other neighbouring species by the following characters: — Head square with anterior margin straight (oblique in H. tubicola; H. spinosa; H. fundiensis; H. antennata). — Antennae subequal, Antenna 2 near the body length; in H. tubicola, the antennae are subequal and reach 2/3 of the body length; they reach 1/3 of the body length for H. fundiensis; in H. spinosa the Antenna 2 length is include between 1/3 and 1/2 of body length. For the two other species, H. descansa and H. oonah, the Antenna 1 is half the length of Antenna 2; whereas in H. antennata, the Antenna 1 is longer than Antenna 2. — Coxa 4 square-shaped, external side bearing numerous short setae. — Epimeral plate 3 square shaped; the posterior margin is oblique or slightly curved in the other species (except for H. descansa). — Uropod 1 long, with rami slender, curved, and of equal length; they are of unequal length for H. tubicola, H. descansa, H. fundiensis, H. oonah and H. antennata (except H. spinosa with rami of equal length, and especially as Uropod 2 rami are strongly armed). In the introduction we indicate that it is necessary to supplement the description of two species described by Stephensen in 1925: Haploops vallifera and Haploops similis. This is the object of the following section.

Key to species of Haploops 1 Corneal lenses present................................................................................. 2 - Corneal lenses absent................................................................................. 17 2 Two pairs of corneal lenses............................................................................. 3 - Only one pair of corneal lenses.......................................................................... 8 3 Pereopod 7 basis broad................................................................................. 4 - Pereopod 7 basis narrow................................................................................ 5 4 P7 carpus with anterior distal lobe; coxa1 inferior margin straight.......................... Haploops laevis Hoek, 1882 - P7 carpus without anterior distal lobe; coxa1 inferior margin rounded............................................................................................ Haploops quebecoisis Bellan-Santini Kaïm-Malka & Dauvin, 2018 5 Inferior lenses in the middle of the lobe head................................... Haploops sibirica Gurjanova, 1929 - Inferior lenses near or on the margin head.................................................................. 6 6 Inferior lenses near the margin head.......................................... Haploops dellavallei Chevreux, 1900 - Inferior lenses on the margin head........................................................................ 7 7 Inferior lenses on the margin, antennae = body length............................. Haploops nirae Kaïm-Malka, 1976 - Inferior lenses on the internal side of the margin, antennae = 1/2 body length........... Haploops tenuis Kanneworff, 1966 8 Only inferior lenses present (superior absent)............................................................... 9 - Only superior lenses present (inferior absent).............................................................. 10 9 Long setae on Md, back, telson, A1 Haploops longiseta Kaïm-Malka, 2010 - Few long dorsal setae, A1> A 2.............................................. Haploops proxima Chevreux, 1919 10 Pereopod 7 basis broad, A 2> body................................ Haploops gascogni Dauvin & Bellan-Santini, 1996 - Pereopod 7 basis narrow............................................................................... 11 11 Head truncated straight................................................................................ 12 - Head transversely truncated............................................................................ 15 12 A1> A 2.............................................................. Haploops antennata Kaïm-Malka, 2012 - A1 Haploops descansa Barnard, 1961 - A1 length> A2 peduncle.............................................................................. 14 14 A1 length a little more A 2 peduncle; coxa 1 oval; U1 rami unequal length.......... Haploops oonah Lowry & Poore, 1985 - A1= 2/3 A2, A2 = body; coxa 1 triangular; outer side coxa 4 setose; U1 rami equal length.... Haploops truncata spec. nov. 15 A 2 = 1/2 body; corneal lenses great size......................................... Haploops tubicola Liljborg, 1856 - A 2 = 1/3 body; corneal lenses small size.................................................................. 16 16. U1–2 strongly armed....................................................... Haploops spinosa Shoemaker 1931 - U1–2 weakly armed............................................. Haploops fundiensis Wildish & Dickinson, 1982 17 Pereopod 7 basis narrow............................................................................... 18 - Pereopod 7 basis broad................................................................................ 24 18 P7 basis, merus and carpus, rectangular and slender (male)....................... Haploops abyssorum Chevreux, 1908 - P7 basis, merus and carpus, not rectangular and slender...................................................... 19 19 Coxa 4 heart-shaped.................................................................................. 20 - Coxa 4 not heart-shaped............................................................................... 22 20 Dorsal side of the body carinate, coxa 2 acute................................... Haploops vallifera Stephensen,1925 - Dorsal side of the body not carinate...................................................................... 21 21 A1= 8–9/10 A 2, A2= 2/3 body............................................... Haploops similis Stephensen, 1925 - Antennae longer than the body length..................... Haploops bjarnii Bellan-Santini Kaïm-Malka & Dauvin, 2018 22 A1= A 2 peduncle, coxa1 oval.................................................... Haploops lodo Barnard, 1961 - Antennae Haploops dauvini Peart, 2018 - A1=A2=1/2 body length......................................................... Haploops faroensis spec. nov. 24 Long dorsal tuft of setae absent......................................................................... 25 - Long dorsal tuft of setae present........................................................................ 26 25 P7 basis lobe exceeding ischium distal edge; A1Haploops antarctica Bellan-Santini & Dauvin, 2008 - P7 basis lobe not exceeding ischium distal edge; A1=A2; coxa 4 heart-shaped............ Haploops kaimmalkai Peart, 2018 26 Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe triangular................... Haploops setosa Boeck, 1870 - Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe rounded............................................27 27 Urosome 1 high upturned carina.............................................. Haploops carinata Liljeborg, 1856 - Urosome 1 carina straight.............................................................................. 28 28 Antenna length> body length, antennae very strong...... Haploops islandica Kaïm-Malka,Bellan-Santini & Dauvin, 2016 - Antenna length =1/2 body length............................................... Haploops robusta G.O. Sars, 1891, Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on page 173, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Hoek, P. P. C. (1882) Die Crustaceen, gesammelt waehrend der Fahrten des \" Willem Barents \" in den Jahren 1878 und 1879, Niederlandisches Archiv fur Zoologie, Supplement band I, 7, 61.","Gurjanova, E. F. (1929) Neue Forrnen arktischer Isopoden und Amphipoden. Zoologischer Anzeiger, 81, 309 - 317.","Chevreux, E. (1900) Amphipodes provenant des campagnes de l'Hirondelle (1885 - 1888). Resultats des Campagnes Scientifiques Accomplies par le Prince Albert I. Monaco. 16, 195 pp.","Kaim-Malka, R. A. (1976) Revision des Haploops (Crustacea, Amphipoda) de Mediterranee. Bolletino del Museo Civico di Storia Naturale, Verona, 3, 269 - 308.","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207.","Kaim-Malka, R. A. (2010) Haploops longiseta, a new species from the Atlantic Ocean (Crustacea, Gammaridea, Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 6]. Zootaxa, 2356 (1), 57 - 68. https: // doi. org / 10.11646 / zootaxa. 2356.1.3","Chevreux, E. (1919) Note preliminaire sur les Amphipodes recueillis par les expeditions du \" Travailleur \" et du \" Talisman \" (1880 - 1883). Bulletin du Museum d'Histoire Naturelle, 25, 574 - 580. https: // doi. org / 10.5962 / bhl. part. 7932","Dauvin, J. C. & Bellan-Santini, D. (1996) Ampeliscidae (Amphipoda) from the Bay of Biscay. Journal of Crustacean Biology, 16, 149 - 168. https: // doi. org / 10.2307 / 1548938","Kaim-Malka, R. A. (2012) Haploops antennata, a new species from the North Atlantic Ocean (Crustacea: Gammaridea: Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 7]. Zootaxa, 3320 (1), 36 - 46. https: // doi. org / 10.11646 / zootaxa. 3320.1.2","Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Lowry, J. K. & Poore, G. C. B. (1985) The ampeliscid amphipods of south-eastern Australia (Crustacea). Records of the Australian Museum, 36, 259 - 298. https: // doi. org / 10.3853 / j. 0067 - 1975.36.1985.348","Shoemaker, C. R. (1931) The Stegocephalid and Ampeliscid Amphipod Crustaceans of Newfoundland, Nova Scotia, and New Brunswick in t he United States National Museum. Proceedings of the United States National Museum, 79 (2888), 1 - 18, 6 figs. https: // doi. org / 10.5479 / si. 00963801.79 - 2888.1","Wildish, D. J. & Dickinson, J. J. (1982) A new species of Haploops (Amphipoda, Ampeliscidae) from the Bay of Fundy. Canadian Journal of Zoology, 60, 962 - 967. https: // doi. org / 10.1139 / z 82 - 133","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de l a Princesse Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13.","Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Peart, R. A. (2018) Ampeliscidae (Crustacea, Amphipoda) from the IceAGE expeditions. Zookeys, 731, 145 - 173, 2 tabs. https: // doi. org / 10.3897 / zookeys. 731.19948","Boeck, A. (1870) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Videnskabs-Selskabet i Christiania 1870, pp. 83 - 280. https: // doi. org / 10.5962 / bhl. title. 2056","Liljeborg, W. (1856) Om Hafs-Crustaceer vid Kullaberg i Skane. Ofversigt af Kongliga Vetenskaps - Akademien Forhandlingar, Stockholm, 12, 117 - 138.","Kaim-Malka, R., Bellan-Santini, D. & Dauvin, J. C. (2016) On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8]. Zootaxa, 4179 (1), 42 - 76. https: // doi. org / 10.11646 / zootaxa. 4179.1.2","Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway with short descriptions and figures of all the species, 1, 191 - 196, pls. 67 - 68."]}

Haploops faroensis spec. nov. (Figures 1–4) Type material. HOLOTYPE. One adult female with oostegites not completely developed. Length: 9.18 mm (Fig.1). BIOFAR: Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’ W; bottom: clay and silt, sand, gravel. Holotype + slides (NHMD – 873179). The specimen is deposited in the Natural History Museum of Denmark (Danish Statens Naturhistoriske Museum). BIOFAR material. The specimens are deposited in the Natural History Museum of Denmark. Station 269, five specimens, 15 May 1988, depth: 510 m, 62° 49.84’N – 08° 15.55’ W; bottom: mud, sand, cobbles and stones (NHMD-873180). Station 271, four specimens, 16 May 1988, depth: 559 m, 62° 52.30’N – 08° 09.24’W; bottom: soft bottom, foraminifers (NHMD-873181). Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’W; bottom: clay and silt, sand, gravel (NHMD-873179). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2673, nine specimens, 15 July 1994, depth: 227– 222 m, 66° 50.20’N – 16°15.74’W; bottom: sponge spicules. (IINH 42254). Station 3522, five specimens, 7 September 2002, depth: 1940 m, 62°31.14’N – 17° 09.87’W, bottom: sediment type unknown (IINH 42253). Station 3544, one specimen, 11 September 2002, depth: 1635– 1632 m, 61°33.00’N – 13°40.14’W, bottom: gravely sand. (IINH 42252). Male unknown. Etymology. The species name refers to the area of the Faroe Islands where the species was collected. Diagnosis. Blind species. Pereon, Pleon and Urosome without long dorsal setae. Antenna 1 and Antenna 2 have the same length. Antennae half length of the body. Pereopod 7 basis narrow. Description. Holotype. Adult female, 9.18 mm (Fig.1). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 2A): rectangular, with a short rostrum pointed, without corneal lenses, blind species. Anterior margin oblique and straight. Antenna 1 (Fig. 2B): as long as antenna 2, half length of the body, peduncle of A1 with article 3 Antenna 2 (Fig. 2C): as long as antenna 1. Peduncle segment 5 longer than 4 (24/20). Flagellum with 17 articles. The two antennae bearing long setae. Upper lip, and Lower lip bilobate. Mandible (Fig. 2D): molar strong; palp long, article 3 longer than article 2 (3/2 = 40/35), setae are present on the margin of each article, article 3 with 3 long apical setae. Maxilla 1 (Fig. 2G): the inner plate is conical shaped with one long apical seta; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 2H): dense apically setae on the two plates. Maxilliped (Fig. 2E): inner plate sub-rectangular, elongated; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, articles 2 and 3 with long setae, article 3 oval shape, dactylus elongate and slender (Fig. 2F). Pereon: without long dorsal setae (Fig. 1). Gnathopod 1 (Fig. 3A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis as long as ischium + merus + carpus, more broad on the distal part than proximal one, with some setae on the two margins, the merus, carpus and propodus bearing long setae; propodus oval; dactylus strongly curved, (Fig. 3B), dactylus length = 0.6 propodus length. Gnathopod 2 (Fig. 3C): coxa 2 approximately rectangular and short (coxa length / basis length = 30/33), distal margin straight with few short setae; basis long, rectangular, with some long setae, basis = ischium + merus + carpus; carpus longer than propodus (carpus length / propodus length = 26/15), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 3D), dactylus length = 1/3 propodus length. Pereopod 3 (Fig. 3E): coxa 3 rectangular, distal margin rounded; basis rectangular, basis longer than ischium + merus + carpus. The margins of the different articles of pereopod 3, except ischium and dactylus, bearing few long setae; dactylus slender, curved and as long as propodus (dactylus length / propodus length = 1). Pereopod 4 (Fig. 3F): coxa 4 square-shaped, anterior margin straight, length / width = 40/34, antero-ventral corner rounded, ventral margin slightly curved with short setae, posterior concavity 1/2 length of the posterior length of the coxa 4, posterior hook broad, length = 12/34 width of coxa 4, posterior corner rounded; basis longer than ischium + merus + carpus; the margins of the different articles of pereopod 4, except dactylus, bearing many long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 21/17). Pereopod 5 (Fig. 3G): coxa 5 roughly rectangular, bi-lobated; basis oval with few short setae on the anterior margin; carpus rectangular, anterior margin with two short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long seta; propodus rectangular a little longer than carpus, with few short setae on the anterior margin, and long distal setae; dactylus slender and curved, dactylus length / propodus length = 12/40 (0.3). Pereopod 6 (Fig. 4A): coxa 6 roughly rectangular, posterior margin rounded; basis rounded with few short setae on the anterior margin; carpus rectangular, anterior margin with few short spines, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long one; propodus rectangular a little shorter than carpus, with few short setae, and a long distal seta; dactylus slender and curved, dactylus length / propodus length = 8/21. Pereopod 7 (Fig. 4B): coxa 7 roughly rectangular, posterior margin rounded; basis narrow (length without lobe / width = 40/20), anterior margin slightly concave, numerous long setae on posterior margin, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with spines on the margins and long setae on the posterior one; carpus rectangular (width / length = 8/12), with small spines on the anterior and posterior margins and one spine on the posterodistal corners (propodus length / carpus length = 4/12; propodus + dactylus / carpus = 8/12); propodus narrow, width / length = 2/4 (propodus width / carpus width = 2/8), with a short apical seta; dactylus rectangular, short and narrow (length / width = 3/1) with two very small apical setae (dactylus length / propodus length = 3/4). Pleon (Fig. 1): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 4G): anterior margin oblique and straight, ventral margin and posterior one straight, inferior corners rounded. Epimeral plate 2 (Fig. 4G): rounded. Epimeral plate 3 (Fig. 4G): anterior margin straight, antero-ventral corner rounded, ventral margin straight, postero-ventral corner acute, posterior margin oblique and straight. Urosome (Fig. 4H): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 4C): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the rami, with 1 spine on the distal margin. length rami / length peduncle = 38 / 52. Uropod 2 (Fig. 4D): shorter than uropod 1; rami triangular, inner ramus longer than the outer one, with a row of five little spines on inner ramus and only two spines on outer, outer ramus length / peduncle length = 36 /41; peduncle rectangular, robust with spines on the inner margin. Uropod 3 (Fig. 4E): peduncle short and strong (length / width = 25/15); rami of equal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 35/25); inner ramus with three spines and some apical setae, outer ramus with an apical tuft of long setae and also some long setae on the outer margin. Telson (Fig. 4F): rectangular, apically rounded, cleft on 33/45 of the length; one spine and one seta are present on the apical part of each lobe. Distribution: North Atlantic Ocean; wide bathymetric range species: 222–1940 m, present mainly in soft sediments: mud, sand, gravely sand, clay and silt, gravel, cobbles and stones, foraminifers, sponge spicules. Taxonomic remarks. H. faroensis spec. nov. belongs to the sub-group of blind species with a narrow pereopod 7 basis and without dorsal tuft seta e. This sub-group includes six other species: H. abyssorum, H. similis, H. vallifera, H. lodo, H. dauvini and H. bjarnii. It differs from these species in having: — Antennae of the same length and as long as half body length; Antenna 2 = 2/3 body length for H. similis; the antennae are longer than the body length for H. bjarnii; for H. lodo, A1= ped A2, A2 shorter than body (in Barnard 1961) (they are half body length for H. vallifera, H. dauvini and H. faroensis), — Coxa 4 square-shaped, heart-shaped for H. bjarnii, H. vallifera, H. similis, but also square- shaped for H. abyssorum, H. lodo and H. dauvini. — Epimeral plate 3 with posterior margin oblique and straight (also for H. similis); slightly convex for H. bjarnii, H. dauvini, rounded for H. vallifera, H. lodo (not indicated by Chevreux 1908 for H. abyssorum). — Uropod 1: rami of same size (also for H. abyssorum), they are of unequal length (2/3 for H. similis), the rami are subequal for H. bjarnii, H. vallifera, H. lodo and H. dauvini.