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The southeastward extrusion of Indochina along the Ailao Shan-Red River shear zone (ARSZ) is one of two of the most prominent consequences of the India-Asia collision. This plate-scale extrusion has greatly changed Southeast Asian topography and drainage patterns and effected regional climate and biotic evolution. However, little is known about how Indochina was extruded toward the southeast over time. Here, we sampled 42 plant and animal clades (together encompassing 1,721 species) that are distributed across the ARSZ and are not expected to disperse across long distances. We first assess the possible role of climate on driving the phylogenetic separations observed across the ARSZ. We then investigate the temporal dynamics of the extrusion of Indochina through a multitaxon analysis. We show that the lineage divergences across the ARSZ were most likely associated with the Indochinese extrusion rather than climatic events. The lineage divergences began at ~53 Ma and increased sharply ~35 Ma, with two peaks at ~19 Ma and ~7 Ma, and one valley at ~13 Ma. Our results suggest a two-phase model for the extrusion of Indochina, and in each phase, the extrusion was subject to periods of acceleration and decrease, in agreement with the changes of the India-Asia convergence rate and angle from the early Eocene to the late Miocene. This study highlights that a multitaxon analysis can illuminate the timing of subtle historical events that may be difficult for geological data to pinpoint and can be used to explore other tectonic events. [ABSTRACT FROM AUTHOR]

Dinomyidae is a South American group of caviomorph rodents with a single extant species (Dinomys branickii), but with a rich fossil record including medium-sized to giant species, which reached its peak diversity during the Late Miocene. The dinomyids from the Cerro Azul Formation (late Middle Miocene to Early Pliocene) have been studied in detail from localities in La Pampa Province, Argentina. In this contribution, dinomyids from Chasicoan Stage/Age (Late Miocene) levels of the Cerro Azul Formation at Arroyo Chasicó locality (Buenos Aires Province, Argentina) are revised, including previously published material and unpublished ones. We confirm the presence of Tetrastylus sp. Diaphoromys gamayensis, and define two new taxa, Bondesiomys chasiquensis gen. et sp. nov. and Gyriabrus sokka sp. nov. The diversity of dinomyids from the Cerro Azul Formation at Arroyo Chasicó is higher than other Chasicoan faunas at the same latitude, identified in localities of La Pampa Province, since in Cerro Patagua there are not Dinomyidae records and in Cerro la Bota Tetrastylus sp. and Dinomyidae indet. were the only recorded taxa. [ABSTRACT FROM AUTHOR]

ABSTRACT The warm Miocene (23.0–5.3 Ma) is considered a model case for the global climate of the near future. In view of the current coral reef crisis, studies on Miocene reefs can contribute to predictions regarding the stress resilience of today's reefs against global warming. A Late Miocene (6.2–5.5 Ma) reef coral fauna from Hispaniola Island (Caribbean) characterised by ecologically tolerant taxa was originally interpreted to reflect cool‐eutrophic waters. However, the associated planktonic foraminifer assemblage and new TEX86H–based temperature reconstructions document an oligotrophic shallow‐water environment harbouring 30–31°C. Thus, in conjunction with unusually low vertical growth rates and erratic trace element signatures of massive reef corals, the coral fauna documents critically high temperatures and a response to heat stress rather than cool upwelling. The findings demonstrate the relevance of understanding shallow‐water ecosystems from past warm periods for developing scenarios of a future, warmer world. [ABSTRACT FROM AUTHOR]

We describe a new species of Ampelopsideae (Vitaceae), Nekemias mucronata sp. nov., from the Rupelian of Cervera (Spain) and revise another fossil species, Ampelopsis hibschii, originally described from Germany. Comparison with extant Ampelopsideae suggests that the North American species Nekemias arborea is most similar to Nekemias mucronata while the East Mediterranean Ampelopsis orientalis is the closest living relative of A. hibschii. Our review of fossil data indicates that, during the Eocene, four species of Ampelopsideae occurred in Eurasia, that is, N. mucronata in the Czech Republic, A. hibschii in Kazakhstan, and two fossil species in the Far East (Ampelopsis cercidifolia and Ampelopsis protoheterophylla). In the Oligocene, a new species, Ampelopsis schischkinii, appeared in Kazakhstan; meanwhile, N. mucronata spread eastwards and southwards, and A. hibschii mainly grew in Central Europe. In the late Oligocene, N. mucronata became a relict in the Iberian Peninsula and Nekemias might have persisted in Western Eurasia until the latest Miocene (“Ampelopsis” abkhasica). The last occurrence of A. hibschii was in the Middle Miocene in Bulgaria, probably a refuge of humid temperate taxa, along with Ampelopsis aff. cordata. Carpological remains suggest that this lineage persisted in Europe at least until the Pleistocene. Our data confirm previous notions of the North Atlantic and Bering land bridges being important dispersal routes for Ampelopsideae. However, such dispersion probably occurred during the Paleogene rather than the Neogene, as previously suggested. A single species of Ampelopsideae, A. orientalis, has survived in Western Eurasia, which appears to have been linked to a biome shift. [ABSTRACT FROM AUTHOR]

Aeolian concentration of iron-titanium oxides (black sand) is a common phenomenon on coasts around the world but associated aeolian detrital gold is relatively rare. In contrast, Holocene and active dunes at Waipapa beach on the coast of the southern South Island of New Zealand host aeolian black sand layers with locally abundant finely particulate (<200 µm) gold and platinum, some of which have been mined historically. Aeolian concentration of black sand, gold and platinum is driven by strong prevailing westerly winds (gusts >100 km/h) on annual or decadal time scales. Holocene fluvial recycling of aeolian sands has been important in upgrading the black sand, gold, and platinum contents. Surf concentration of black sand on the beach is minor and merely feeds the aeolian system, and this differs from other beach gold deposits around the world, which are surf-dominated. Most of the Waipapa beach gold (>90%) has flaky morphology with abundant superficial transport-related deformation reflective of fluvial transport, probably down the nearby Mataura River (∼200 km transport) between Miocene and Holocene. Minor distinctive wind-sculpted toroidal gold, and the platinum, arrived from the west (∼300 km transport) via the Waiau River and Foveaux Strait. [ABSTRACT FROM AUTHOR]

The identification of additional Late Miocene (MN10, Late Vallesian) Decennatherium rex cranial remains at Batallones-10 (Torrejón de Velasco, Madrid Basin, Spain) provides valuable insights into the ontogenetic variability of this extinct giraffe. Most recovered specimens are adults, making the discovery of calves significant. The 2022 excavation yielded two juvenile hemimandibles, a maxilla, and a juvenile ossicone, likely from different individuals. The dental material represents one of the youngest specimens in the Batallones complex, with an estimated age of 4 months to 4.5 years, based on molar eruption stages seen in modern giraffes. The ossicone, resembling that of an adult male in shape but much smaller, mirrors the growth patterns in extant giraffes, where juvenile ossicones share adult proportions but are smaller. This ossicone was detached from the skull, typical for juveniles, as ossicones do not fuse with the cranial roof until later. This discovery broadens our understanding of giraffe ontogeny and includes the first giraffe calf ossicone found in the Iberian fossil record. [ABSTRACT FROM AUTHOR]

“Nubecularia bioherms” represent unique foraminifera dominated buildups which are restricted to the late Middle Miocene (late Serravallian) of the Paratethys. They occur in the Central Paratethys in the late Sarmatian and in the Eastern Paratethys in the Volhynian–Bessarabian, both regional stages of the respective Paratethyan areas. The generic assignment of the main constituent of the bioherms changed in time from Spirorbis to Nubecularia and Sinzowella. No clear diff erential diagnosis has been provided for the establishment of the genus Sinzowella by Cushman (1933) and the diff erence between both porcelaneous taxa was reported to be the agglutination of foreign particles in Nubecularia (Loeblich and Tappan 1964, 1987). Since in studied Central Paratethyan material agglutinated particles occur frequently in the test wall a separation of Sinzowella from Nubecularia is not justifi ed based on this character and Sinzowella is considered synonymous with Nubecularia. At the species level the situation is even more confusing since a variety of species (nodulus, novorossica, caespitosa), subspecies or/and “types” (e.g., nodulus, solitaria, deformis) have been described. All these taxa are considered to be subjective junior synonyms of Nubecularia nodulus which is the only Nubecularia species in the Central Paratethys since all described morphotypes co-occur even in a single rock thin section. [ABSTRACT FROM AUTHOR]

A fungal palynomorph corresponding to the fossil-species Lirasporis intergranifer R. Potonié & S.C.D. Sah, from the Middle Miocene lignite mine in Tomisławice (central Poland), is considered as the new combination Cancellidium intergraniferum (R. Potonié & S.C.D. Sah) G. Worobiec & E. Worobiec, comb. nov. The fossil-genus Lirasporis is treated as a synonym of the modern genus Cancellidium Tubaki. Cancellidium intergraniferum from the Miocene of Poland represents the first fossil record of Cancellidium outside Asia, the first record of this genus from Europe, and the northernmost known fossil occurrence of Cancellidium. Living representatives of Cancellidium seem to prefer warm and humid climates and, being aero-aquatic hyphomycetes, are usually associated with decaying woody remains in aquatic environments. A similar ecology is suggested for the fossil C. intergraniferum, which could be a reliable fungal non-pollen palynomorph proxy for palaeoclimatic and palaeoenvironmental reconstructions. [ABSTRACT FROM AUTHOR]

In the present-day Indo-Pacific coral-reef settings two genera of alveolinoidean porcelaneous larger benthic foraminifera (LBF) occur, namely Alveolinella and Borelis. Alveolinella is represented by a single species, A. quoyi , whose northernmost record is in Okinawa-jima (central Ryukyu Islands, Japan). Although the Indo-Pacific area, and especially the Coral Triangle, is a biodiversity hotspot since the Early Miocene, in-depth investigation on fossil representatives of present-day LBF is limited to a few taxa. To help bridge this knowledge gap, the palaeobiogeographical dynamics of A. quoyi is assessed. Analysis of data from the palaeontological literature shows that its first appearance datum is from the Tortonian (Late Miocene) of East Kalimantan and Papua New Guinea. In the Pliocene–Pleistocene the Indonesian Throughflow constrained the species within the Central Indo-Pacific. Finally, during the Late Pliocene the northward migrants arrived in the shallow-water carbonate settings of Okinawa-jima where the species is still thriving. [ABSTRACT FROM AUTHOR]

The Guadix-Baza Basin (southern Spain) displays one of the best continental records from the Late Miocene to the Middle Pleistocene in western Europe, with many localities of fossil small mammals in a quite complete and continuous stratigraphic sequence. The Late Miocene continentalization of the Guadix-Baza Basin was firstly mentioned in the 1980s based on the presumed presence of Hipparion gromovae granatensis at the site of Abla (Almería province). Subsequent discoveries of late Turolian deposits confirmed the Late Miocene continentalization of the basin. The interest in the Abla site led to a second sampling campaign to look for microvertebrate fossils, which proved successful. In this paper, a complete taxonomic study of the novel microvertebrate fauna from Abla is presented, providing updated information on the age of the locality. Representatives of the vertebrate families Cyprinidae, Alytidae, Anguidae, Soricidae, Cricetidae, and Muridae have been identified. The presence of Stephanomys specimens exceeding the size of latest Miocene-earliest Pliocene species allows us to correlate the Abla site with the Early Pliocene (Ruscinian), a younger age than initially stated. A qualitative paleoecological interpretation based on the herpetofauna suggests the dominance of open environments with presence of permanent water bodies during the deposition of the Abla site, under warmer and more humid climate conditions than today in the Guadix-Baza Basin. A revision of the Hipparion remains from this locality showed that the sample is closer to Hipparion fissurae than to the initially assigned species. [ABSTRACT FROM AUTHOR]

The predation-driven Mesozoic marine revolution (MMR) is believed to have induced a dramatic change in the bathymetric distribution of many shallow marine invertebrates since the late Mesozoic. For instance, stalked crinoids – isocrinids (Isocrinida) have undergone a striking decline in shallow-sea environments and today they are restricted to deep-sea settings (below 100 m depth). However, the timing and synchronicity of this shift are a matter of debate. A delayed onset of MMR and/or shifts to a retrograde, low-predation community structure during the Paleogene in the Southern Ocean were invoked. In particular, recent data from the Southern Hemisphere suggest that the environmental restriction of isocrinids to the deep-sea settings may have occurred at the end of the Eocene around Antarctica and Australia, and later in the early Miocene in New Zealand. Here, we report the anomalous occurrence of the isocrinids in shallow nearshore marine facies from the middle Miocene of Poland (Northern Hemisphere, Central Paratethys). Thus, globally, this is the youngest record of shallow-sea stalked crinoids. This finding suggests that some relict stalked crinoids may have been able to live in the shallow-water environments by the middle Miocene, and further confirms that the depth restriction of isocrinids to offshore environments was not synchronous on a global scale. [ABSTRACT FROM AUTHOR]

Nannofossils and planktonic foraminifers from the Lower Miocene Alkun Formation of central and western Ciscaucasia were studied in the following sections: the stratotype section on the Alkunka River, the Republic of Ingushetia; stratotype of the Caucasian Regional Stage on the Kuban River, Karachay-Cherkessia; Lower Miocene reference section on the Belaya River, Adygea; and in the sections of the Alkun Formation stratotype area on the Fiagdon and Mairamadag rivers, North Ossetia. Planktonic foraminifers in the Alkun Formation were studied for the first time. The available results made it possible to reveal the composition and structure of nannofossil and planktonic foraminiferal assemblages from the Alkun sediments in Ciscaucasia, to refine their stratigraphic and correlation potential and to recognize bionomic conditions of the Alkun basin in the studied area. As a result of the nannoplankton study the Cyclicargolithus floridanus Beds were recognized in the Alkun Formation and, taking into account the occurrence of the zonal species Triquetrorhabdulus carinatus along with Tr. milowii, they are correlated with the upper (Miocene) part of the Aquitanian Zone NN1. Planktonic foraminiferal data permitted to recognize the Streptochilus pristinum Beds. [ABSTRACT FROM AUTHOR]

New fossil remains of Rhinocerotidae, dated from the early Late Miocene, were discovered from the lowermost part of the Irrawaddy Formation in the Tebingan area, Magway Region, central Myanmar. The Tebingan rhinoceros fossils, consisting of isolated teeth, maxillae, and mandibles, are assigned to three taxa: Brachypotherium perimense defined by a constricted protocone, a well-developed parastyle and broad upper teeth, to 'Brachypotherium' fatehjangense with a flat ectoloph, crochet, and the absence of tubercle at the entrance of the median valley. Other specimens were assigned to Rhinoceros sp. due to having a developed parastyle, crochet and a strong anterior cingulum and the absence of a constricted protocone and ante-crochet. The evolutionary history of Rhinocerotidae is poorly known in Southeast Asia, and few rhinoceros' species have been identified from the Irrawaddy Formation. The newly identified Rhinoceros specimens from the early Late Miocene Tebingan area are the oldest fossil records for Rhinoceros in Southeast Asia. The Tebingan mammal fauna is similar to the Nagri and Dhok Pathan faunas of the lower/middle Siwaliks of the Indian subcontinent, indicating the faunal exchange between these two regions in the late Neogene. [ABSTRACT FROM AUTHOR]

Palaeotragus is a common genus of the Giraffidae that was widely distributed in the Neogene strata of Eurasia. Early studies usually placed small giraffids in the Palaeotragus clade, leading to considerable controversy over the validity and systematic classification of this genus. Palaeotragus microdon and Palaeotragus cf. coelophrys are two representative species of Palaeotragus in China, their dental specimens are mainly distinguished by size, causing problems in their actual identification. In this study, several mandibles of a small giraffid found in Linxia Basin, Gansu Province, China (MN 10–13), are studied and identified as Palaeotragus cf. coelophrys. Based on a detailed comparison, the size and morphological variability of the Chinese Palaeotragus dental material are discussed, as well as the validity of the two species and the validity of other Eurasian Palaeotragus. This study reveals that the Chinese Palaeotragus has a continuous sequence of size variability and premolar features, which is insufficient to separate the mandible of the two species. In addition, the type species P. rouenii can be clearly distinguished from the Chinese Palaeotragus by the relatively long lower premolar row, possibly due to the difference in feeding habits. The holotype of P. coelophrys is closely related to Chinese Palaeotragus, while the P. coelophrys mandible from Maragheh (MMTT37 2517) should belong to different species. [ABSTRACT FROM AUTHOR]

A rhinocerotid mandibular remain from the Upper Pliocene Tsubusagawa Formation (around 3.5 Ma) was described and discussed its taxonomic status and paleobiogeographical implication. Compared with Late Miocene to Early Pleistocene and living rhinocerotid species, the studied specimen resembles the mandible of the Pliocene Eurasian taxa of the subtribe Rhinocerotina, Dihoplus and Pliorhinus in having a straight ventral margin, constant mandibular height and probably reduced lower second incisor. This specimen, however, lacks detailed diagnostic characters of these genus. In addition, the taxonomic validity of these taxa has been argued so far. Therefore, this specimen was identified as Rhinocerotina gen. et sp. indet. Japanese Pliocene rhinocerotid records and the present study suggest that the Rhinocerotina was distributed in the central to southwestern part of proto-Japan during the mid-Pliocene. If the studied specimen is truly a species of Dihoplus or Pliorhinus, its close relative might have migrated from northern China as in the case of the ancestor of Japanese Pliocene proboscidean, Stegodon miensis. Further discovery of Japanese Pliocene rhinocerotid specimens could resolve this issue. [ABSTRACT FROM AUTHOR]

The late Miocene Monte Capanne and Porto Azzurro plutons are investigated by means of coupled U–Pb zircon and 40Ar/39Ar white mica dating to test the occurrence of long‐lived magmatic systems in the upper crust. Zircon crystallized for >1 Ma in both plutonic systems, with supersolidus conditions overlapping for ~220 ka indicating previously unrecognized co‐existence of the two reservoirs. The development of the Porto Azzurro high T‐aureole is post‐dated by continuous igneous zircon crystallization until ~6.0 Ma. By linking crystallization to post‐emplacement cooling of late‐stage pulses in both western and eastern Elba we constrain long‐lived sizeable reservoirs (possibly the same reservoir) in the Tyrrhenian upper crust between ~8 and 6 Ma. [ABSTRACT FROM AUTHOR]

A new species Arachnocoris furtivinabis Ma, Du et Yao sp. nov. is described and illustrated, which represents the first fossil record of Arachnocoris, also the first record in the family Nabidae from Cenozoic Dominican amber. Based on the distinctive morphological characters in this new species, including antennomere II much shorter than twice of antennomere I, labial segment III slightly longer than segment II, and parameres round, it is differentiated from other species within Arachnocoris. Compared with the spider-webdwelling taxa of the extant and extinct species in Heteroptera, the evolution of predation strategies for this special group during the Miocene was discussed. [ABSTRACT FROM AUTHOR]

A new, moderately large shell of a fossil species from the family Fasciolariidae Gray, 1853 is described from the Middle Miocene at Bengbreng Hill, West Java, Indonesia. The shell of this new species is related to species of the genus Clavilithes Swainson, 1840. The characteristics of a globular body whorl, low spire, and moderately long siphonal canal, distinguish it from species of the genus Clavilithes that have a fusiform shape, high spire, and usually long siphonal canal. This leads to its placement in a new genus Indolithes new genus. The new fossil species is compared with two other fossil species of Fasciolariidae from Java, Clavilithes fennemai (K. Martin, 1906) and C. verbeeki (K. Martin, 1895). [ABSTRACT FROM AUTHOR]

The origin and evolution in the Southern Hemisphere of the Austral teals, consisting of the grey-teal and brown-teal species complexes, remains poorly understood owing to limited molecular data. With the group containing multiple independent examples of flight loss, understanding the evolutionary history of the group is of significant interest for functional genomic studies into the evolution of flightlessness. Here, we present the first whole-mitogenome-based phylogeny of the Austral teals. We show that the group diverged from a common ancestor with mallards in the late Miocene and soon after radiated into the brown-teal and grey-teal lineages, in addition to the widely distributed pintails and green-winged teals. The brown-teal species complex, which includes the volant brown teal, the flight-impaired, sub-Antarctic Auckland Island teal, and the flightless, Chatham Island and sub-Antarctic Campbell Island teals, radiated within the past 0.9–1.8 Myr. The divergence of the extinct Chatham Island teal and the colonization of the Auckland and Campbell Islands occurred from mainland New Zealand. Morphological changes towards flightlessness are also present in the volant brown teal on mainland New Zealand, suggesting that this group was on the pathway to flightlessness, a trend that accelerated in some insular island lineages. [ABSTRACT FROM AUTHOR]

Elongated biconvex Carex fossil fruits are relatively common in fossil assemblages, but they are of much uncertain taxonomic affinity. Three fossil-species have been erected to accommodate the materials from the respective type localities (C. klarae, C. paucifloriformis and C. paucifloroides) and a number of superficially similar-looking fossils were reported from other sites. The formerly proposed affinities mostly pointed to the extant C. pauciflora (subgen. Euthyceras) or Carex sect. Cyperoideae (subgen. Vignea), depending on the authors. But despite the similar superficial resemblance of the fossil remains to these not very closely related sedge taxa, other groups are also possible matches. We used a comparative approach employing stereomicroscope and SEM to analyze the affinities of several Old World "mummified" fossils (ca. 23 to 0.1 Ma) of elongated biconvex fruits. Several samples showed a fair preservation of previously unattested microscopic characters. In particular, one of them allowed us to detect an indirect record of fine silica body morphologies (impressions) on the periclinal walls, which permitted the sound taxonomic placement of the remains from the Miocene of Siberia as a Carex sect. Cyperoideae. In all the other fossils analyzed, the epidermal cell outline and the central position of the main silica body were sufficient to discard assignment to subgen. Euthyceras, despite a purported affinity to C. pauciflora reported in previous literature. In the absence of detailed information on silica bodies, epidermal cell (exocarp) details in fossil specimens do not seem to be determinant for the systematic placement. However, the addition of diagnostic style characters pointed to sect. Cyperoideae as the best match with the available information. Our data support a continued presence of Carex sect. Cyperoideae in the Old World since the Miocene, which candidates Eurasia as a possible place of origin for it, despite DNA-based reconstructions alone proposed North America. This study underlines the importance of detailed taxonomic studies to properly address the identity of fossils and unravel the biogeographic history of plants. [ABSTRACT FROM AUTHOR]

The article focuses on Oligocene and Miocene blowtorch volcanism within the California Borderland and its influence on the plate-boundary transition. It details the methods for sample collection, including ROV deployments and geological analyses, and presents results from 40Ar/39Ar age determinations and whole rock geochemistry. It also includes a regional plate motion model constructed using GPlates software.

This study investigated the calcareous nannofossil assemblages in detail from the early Miocene aged Lice Formation outcropping in the Kahramanmaraş basin. The biostratigraphy of calcareous nannofossils was outlined and paleoenvironmental features determined. In 81 samples taken from three measured sections in the region, 17 calcareous nannofossil genus and 48 nannofossil species were identified. These calcareous nannofossil genus and species identified the Lice Formation as being in the CNM4 nannofossil biozone. The abundance and diversity of early Miocene calcareous nannofossil species varied in the measured sections, with the samples generally moderate‐poor, apart from a few samples. The relative abundance of individuals of Cyclicargolithus floridanus, Coccolithus pelagicus, Reticulofenestra hagii and Sphenolithus moriformis species, with paleoecological importance identified in the study region, indicate that in early Miocene times, the basin in which the Lice Formation deposited was meso‐eutrophic with excess nutrient input, temperate and generally stable shallow marine conditions. [ABSTRACT FROM AUTHOR]

The folded Zagros is a part of the active Zagros belt in the southwest of Iran. This area has main faults and active rock formations, the activity of these faults has somehow affected the strata and sediments of different geological periods. In this study First, by studying satellite images and aerial photographs, five locations were selected in different parts of Fars and the hinterland of Bandar Abbas, and finally, the tectonostratigraphy of these locations was compared with each other. The sinology column and the paleogeographical map of these regions in the Miocene period were reconstructed Using the data obtained from five places of folded Zagros (Bastak, Bahar anticline, Arjan plain, Saadi mountain and Yasouj). The obtained results and the resulting diagrams were obtained using the RockWorks geological program. the minimum and maximum stratified thicknesses of the Miocene period formation in these five studied points are related to Arjan Plain and Saadi mountain sections By comparing the obtained results and the Hesar diagrams and the isopach map. It was done between the studied locations, which shows the highest thickness of this formation in the Arjan plain section and the lowest thickness is in Saadi Mountain. The facies changes of sedimentary rocks of the Miocene period seem to be due to the operation and activity of bedrock faults such as Kare Bes, Sabzpushan, Sarvestan and Kazeron faults in this region. [ABSTRACT FROM AUTHOR]

The High‐Pressure Phyllites‐Quartzites (PQ) unit of the External Hellenides is exposed in tectonic windows in the central and northern Peloponnese (Greece). Understanding the deformation history of this unit is essential to interpreting the Oligo‐Miocene evolution of the External Hellenides belt and its associated exhumation events. This study integrates new field observations and microtectonic analyses with previous studies to offer a comprehensive deformation model of the PQ unit since the Late Oligocene. The first deformation phase (D1), captures the progressive incorporation of the PQ into an orogenic wedge. This phase is largely overprinted and only preserved as relict features. The second phase (D2) displays coeval top‐to‐the‐ENE and top‐to‐the‐WSW localized ductile shear. A transition is observed from top‐to‐the‐ENE non‐coaxial deformation at the upper parts of the nappe to intense isoclinal folding (refolding S1) at the lower structural levels. We associate D2 with the ductile syn‐orogenic exhumation of the PQ within an extrusion wedge, accompanied by greenschist‐facies retrogression. In the third phase (D3), semi‐brittle to brittle extensional fault planes cut through the previous ductile structures. D3 faults exhibit extensional kinematics in all directions on the flanks of exhumation domes. This phase correlates with a late‐orogenic doming event, marking the final exhumation stage of the PQ unit in the upper crust. The exhumation of high‐pressure units results from the interplay between ductile syn‐orogenic extrusion and continuous underplating within the subduction zone. This underplating maintains vertical movements and uplift of the units, initiating a 3D upper‐crustal extensional collapse along low‐angle normal faults. Plain Language Summary: The formation of mountain ranges is a subject of great interest to geologists. This process unfolds in cycles, typically beginning with the plunging, or subduction, of the oceanic portion of a tectonic plate into the mantle. Occasionally, a subducting fragment is too buoyant to sink, thus obstructing the process and leading to the erection of a mountain range. Traces of these early stages are not frequently observed, as they are either carried away in the mantle or deformed during the ensuing collision. In this study, we examine the Phyllites‐Quartzites unit, a sedimentary remnant from a plate subducted 25 million years ago (Ma) at the Aegean Arc, which managed to resurface due to its buoyancy, the upward thrust from other tectonic fragments, and the eventual collapse of the overlying mountain range. By analyzing the deformation markers retained by this unit, we discern that they fall into three phases: (D1) evidence of burial under high‐pressure conditions (35–50 km depth and 500°C) around 25 Ma; (D2) subsequent rise around 15 Ma under warm and ductile conditions due to buoyancy and underplating; and (D3) dismantling of the overlying mountain chain at the surface around 5 Ma, characterized by cooler, brittle faulting. Key Points: The Phyllites‐Quartzites (PQ) unit underwent three distinct deformation phases (D1–D3) in the central and northern Peloponnese (External Hellenides)The exhumation of the high‐pressure PQ (D2) resulted from ductile syn‐orogenic extrusion and underplating within the subduction zoneA brittle doming event (D3) records vertical uplift and exhumation in the upper portions of the orogenic prism, facilitated by multi‐directional Low‐Angle Normal Faults [ABSTRACT FROM AUTHOR]

The Sulu Sea is a small marginal sea in the western Pacific, but it is a very complex and tectonically active region, situated amidst the convergence of the Eurasian, Pacific, and India-Australian plates. Deciphering its geodynamic evolution is crucial, but our understanding of its opening, closure, and tectonic history remains inadequate. The main aim of this study was to systematically study the opening and subsequent closure of the Sulu Sea though discerning tectonic unconformities, structural features, and subduction-collision tectonic zones around margins of the sea. The interpreted sections and gravity anomaly data indicate that the NE Sulu Sea has undergone Neogene extension and contraction due to subduction and collision along the northern margins of the Sulu Sea, whereas the SE Sulu Sea gradually extended from northwest to southeast during the Middle Miocene and has subsequently subducted since the Middle Miocene along the southeastern margins of the Sulu Sea. Several subduction and collision boundaries with different characteristics were developed including continent-continent collision, arc-continent collision, and ocean-arc subduction. The different margins of the Sulu Sea showed distinct asynchronous subduction and collision processes. The northern margins of the Sulu Sea can be divided into three subduction-collision tectonic zones from west to east: the Sabah-Nansha block collision has occurred in NE Borneo since the Early Miocene, followed by the SW Palawan-Cagayan arc collision in SW Palawan Island since the Middle Miocene, and the NW Palawan-Mindoro arc collision since the Late Miocene with further oblique subduction of the Philippine Sea Plate. The southeastern margins can also be divided into two subduction tectonic zones from south to east: the SE Sulu Sea has subducted southward beneath the Celebes Sea since the Middle Miocene, followed by the southeastward subduction beneath the Philippine Sea Plate since the Pliocene. Since the Miocene, the interactions among the Australia-India, the Philippine Sea, and the Eurasian plates have formed the circum-Sulu Sea subduction-collisional margins characterized by microplate collisions, deep-sea trough development, and thick sediments filling in the orogenic foreland. This study is significant for gaining insights into the opening and closure of marginal seas and the dynamics of multiple microplates in Southeast Asia. [ABSTRACT FROM AUTHOR]

The South Asian summer monsoon (SASM) significantly intensified during the Middle Miocene (17–12 Ma), but the driver of this change remains an open question. The uplift of the Himalaya (HM) and the Iranian Plateau (IP) and global CO 2 variation are prominent factors among suggested drivers. Particularly, the impact of high CO 2 levels on the Miocene SASM has been little studied, despite the wide range of reconstructed CO 2 values around this period. Here we investigate their effects on the SASM using the fully coupled Ocean–Atmosphere Global Climate Model, CESM1.2, through a series of 12 sensitivity experiments. Our simulations show that the IP uplift plays a dominant role in the intensification of the SASM, mainly in the region around northwestern India. The effect of the HM uplift is confined to the range of the HM and its vicinity, producing orographic precipitation change. The topography forcing overall out-competes CO 2 variation in driving the intensification of the SASM. In the case of extremely strong CO 2 variation, the effects of these two factors are comparable in the core SASM region, while in the western region, the topographic forcing is still the dominant driver. We propose a thermodynamical process linking the uplift of the IP and the enhanced SASM through the release of latent heat. When compared with reconstructions, the simulated response of SASM to the IP uplift is in good agreement with observed precipitation and wind field, while the effects of the HM uplift and CO 2 variation are inadequate to interpret the proxies. [ABSTRACT FROM AUTHOR]

The Late Miocene vertebrate fauna from the Entre Ríos Province, Argentina, comes from two units: the marine Paraná Formation and the Lower Member of Ituzaingó Formation (LMIF). This latter includes a lower conglomerate set, with fine quartz gravel, and clay and chalcedony clasts, and fossil remains, traditionally known as ‘Mesopotamiense’ or ‘Conglomerado osífero’. We present new fossil vertebrate remains from the LMIF at the Puerto Víboras locality (Hernandarias, Entre Ríos Province) including Leptodactylus sp. (Amphibia, Anura), Crocodylia indet. Caimaninae indet. and Paranasuchus gasparinae (Alligatoridae), Chelidae indet. (Testudines, Pleurodira), Chelonoidis sp. (Testudines, Cryptodira), Plohophorus paranensis (Cingulata, Glyptodontidae), and Macraucheniinae indet. (Macraucheniidae, Litopterna). The record of Leptodactylus sp. represents the oldest record of the genus and the first amphibian record for the Neogene of the Entre Ríos Province. The material described herein and assigned to Chelonoidis sp. allows to confirm for the first time the presence of a testudinid in the Late Miocene of the Entre Ríos Province. [ABSTRACT FROM AUTHOR]

The South American Dry Diagonal, also called the Diagonal of Open Formations, is a large region of seasonally dry vegetation extending from northeastern Brazil to northern Argentina, comprising the Caatinga, Cerrado, and Chaco subregions. A growing body of phylogeography literature has determined that a complex history of climatic changes coupled with more ancient geological events has produced a diverse and endemic‐rich Dry Diagonal biota. However, the exact drivers are still under investigation, and their relative strengths and effects are controversial. Pleistocene climatic fluctuations structured lineages via vegetation shifts, refugium formation, and corridors between the Amazon and Atlantic forests. In some taxa, older geological events, such as the reconfiguration of the São Francisco River, uplift of the Central Brazilian Plateau, or the Miocene inundation of the Chaco by marine incursions, were more important. Here, we review the Dry Diagonal phylogeography literature, discussing each hypothesized driver of diversification and assessing degree of support. Few studies statistically test these hypotheses, with most support drawn from associating encountered phylogeographic patterns such as population structure with the timing of ancient geoclimatic events. Across statistical studies, most hypotheses are well supported, with the exception of the Pleistocene Arc Hypothesis. However, taxonomic and regional biases persist, such as a proportional overabundance of herpetofauna studies, and the under‐representation of Chaco studies. Overall, both Pleistocene climate change and Neogene geological events shaped the evolution of the Dry Diagonal biota, though the precise effects are regionally and taxonomically varied. We encourage further use of model‐based analyses to test evolutionary scenarios, as well as interdisciplinary collaborations to progress the field beyond its current focus on the traditional set of geoclimatic hypotheses. [ABSTRACT FROM AUTHOR]

A toxodontid (Notoungulata) specimen (CHN-UNLC-P-V-4) is the first fossil from the Paso de las Carretas Formation that crops out in the Río Quinto Basin (Neogene), to the east of San Luis city (Argentina). The geological study provides a new stratigraphic section and the position of sediments in the valley depocentre permits inferring an older age than that of the Río Quinto Formation, which implies an early Late Miocene age for the Paso de las Carretas Formation. This age agrees with the identified cf. Ocnerotherium intermedium (taxonomic determination supported by a phylogenetic analysis), as this species was recognised in the Chasicoan Stage/Age of San Luis Province. CHN-UNLC-P-V-4 includes a mandibular fragment with ventral extension, preserving m1–3, a symphysis fragment with incisors, and the partial upper tooth series. A body mass of ~ 1700 kg is estimated. The mandible is similar to that of Dinotoxodon and Gyrinodon, but teeth (mainly upper ones) approach CHN-UNLC-P-V-4 to O. intermedium. As the mandibular extension of this species was not established, but likely present and lower teeth have some differences, we maintain an open nomenclature for CHN-UNLC-P-V-4. The similar incisors to the holotype of ‘Neoadinotherium triangulatum’ supports the latter might belong to O. intermedium, as previously suggested. [ABSTRACT FROM AUTHOR]

The Bounty Channel is a large-scale submarine channel system located in the eastern continental margin of New Zealand. Extending along the axis of the Bounty Trough, the channel system comprises three main tributaries (C1–C3) at its head, which merge downstream into a trunk channel leading to a terminal submarine fan. In this study, we use high-quality two-dimensional multichannel seismic data to investigate the formation and evolution of tributary channels C1 and C2. Four types of seismic facies are identified in the tributary channels: fill-type, mounded divergent, wavy, and subparallel facies. These seismic facies are correspondingly interpreted as topographic depression or channel fills, levees, sediment waves, and hemipelagic deposits. The Late Miocene tributary channels were developed above a pre-existing NE–SW-oriented depression. The Pliocene to Quaternary tributary channels are characterized by preferential development of higher levees on their left hand, and the presence of sediment waves on the lower levees of their right-hand, signaling an effect of the Coriolis force. The formation and evolution of the tributaries are primarily linked to regional tectonics, including increased convergence rate between the Pacific and Australian plates along the Alpine Fault in the Late Miocene and enhanced uplift and erosion at the Southern Alps during the Pliocene. [ABSTRACT FROM AUTHOR]

Salinas Grandes de Hidalgo (La Pampa Province) is a relevant locality of the Cerro Azul Formation in central Argentina because of its abundant and diverse vertebrate remains, including representatives of Reptilia, Aves and mostly Mammalia. Concerning previous records, the present faunal list adds one Argyrolagidae marsupial, one Ctenomyidae and two Caviidae rodents, as well as the first detailed analysis and description of glyptodonts from the Cerro Azul Formation in La Pampa Province. New geological information is added about the two fossiliferous outcrops included in Salinas Grandes de Hidalgo area. The classically studied outcrop located in the saline lake and the nearby one in the so-called ex-Autódromo, both herein correlated based on lithostratigraphy, particularly by the presence of paleosols in the intermediate section of the sedimentary succession. These sandy silts deposits in the base of the outcrops indicate the development of poorly confined flows and intervals of surface stabilisation by paedogenesis. We offer a biochronological interpretation based on a recently published U-Pb detrital zircons dating (5.9 ± 0.4 Ma) and the presence of significant taxa such as octodontoid rodents recovered from the dated levels and below it. From a paleoenvironmental viewpoint, we infer variations from woodlands to open environments. [ABSTRACT FROM AUTHOR]

High latitude metatherians from the Middle Miocene of Patagonia (Argentina and Chile) are only known from medium-sized Sparassodonta, and undescribed taxa collected in several fossil localities of Collon Cura (Argentina) and Río Frías (Chile) Formations, although the latter has several species reported and mostly recognised for the slightly older Santa Cruz Formation. In this contribution we describe five new genera and six new species of metatherians from Collon Cura Formation at Cerro Zeballos, Chubut province (Argentina), based on 11 specimens that notably increased the diversity of Collon Cura Formation. The specimens were assigned to the orders Paucituberculata (Palaeothentoidea), Sparassodonta (Hathliacynidae), and Polydolopimorphia (Argyrolagidae), and are taxa of small size, including the smallest palaeothentoids known to date. Also, the Palaeothentoidea of Cerro Zeballos are amongst the youngest of the superfamily in Patagonia. The new specimens described herein show morphological differences with other metatherian associations already known, probably related to adaptive changes coincident with the Middle Miocene Climatic Transition and the beginning of a faunal turnover. [ABSTRACT FROM AUTHOR]

Background: Calcareous nannofossils are minute microfossils widely present in marine strata. Their identification holds significant value in studies related to stratigraphic dating, paleo-environmental evolution, and paleoclimate reconstruction. However, the process of identifying these fossils is time consuming, and the discrepancies between the results obtained from different manual identification methods are substantial, hindering quantification efforts. Therefore, it is necessary to explore automated assisted identification of fossil species. This study mainly focused on 18 key fossil species from the Miocene era. Five convolutional neural network (CNN) models and 10 data augmentation techniques were compared. These models and techniques were employed to analyze and collectively train two- and three-dimensional fossil morphologies and structures obtained from three different fossils observed under single-polarized light microscopy, orthogonal polarized light microscopy, and scanning electron microscopy. Finally, the model performance was evaluated based on the predictive outcomes on the test set, using metrics such as confusion matrix and top-k accuracy. Result: The results indicate that, for the calcareous nannofossil images, the most effective data augmentation approach is a combination of four methods: random rotation, random mirroring, random brightness, and gamma correction. Among the CNN models, DenseNet121 exhibits the optimal performance, achieving an identification accuracy of 94.56%. Moreover, this model can distinguish other fossils beyond the 18 key fossil species and non-fossil debris. Based on the confusion matrix, the evaluation results reveal that the model has strong generalization capability and outputs highly credible identification results. Conclusion: Drawing on the identification results from CNN, this study asserts a robust correlation among extinction photographs, planar images, and stereoscopic morphological images of fossil species. Collective training facilitates the joint extraction and analysis of fossil features under different imaging methods. CNN demonstrates many advantages in the identification of calcareous nannofossils, offering convenience to researchers in various fields, such as stratigraphy, paleo-ecology, paleoclimate, and paleo-environments of ancient oceans. It has great potential for advancing the development of marine surveys and stratigraphic recognition processes in the future. [ABSTRACT FROM AUTHOR]

Background and Aims The geographical origin and evolutionary mechanisms underpinning the rich and distinctive New Caledonian flora remain poorly understood. This is attributable to the complex geological past of the island and to the scarcity of well-resolved species-level phylogenies. Here, we infer phylogenetic relationships and divergence times of New Caledonian palms, which comprise 40 species. We use this framework to elucidate the biogeography of New Caledonian palm lineages and to explore how extant species might have formed. Methods A phylogenetic tree including 37 New Caledonian palm species and 77 relatives from tribe Areceae was inferred from 151 nuclear genes obtained by targeted sequencing. Fossil-calibrated divergence times were estimated and ancestral ranges inferred. Ancestral and extant ecological preferences in terms of elevation, precipitation and substrate were compared between New Caledonian sister species to explore their possible roles as drivers of speciation. Key Results New Caledonian palms form four well-supported clades, inside which relationships are well resolved. Our results support the current classification but suggest that Veillonia and Campecarpus should be resurrected and fail to clarify whether Rhopalostylidinae is sister to or nested in Basseliniinae. New Caledonian palm lineages are derived from New Guinean and Australian ancestors, which reached the island through at least three independent dispersal events between the Eocene and Miocene. Palms then dispersed out of New Caledonia at least five times, mainly towards Pacific islands. Geographical and ecological transitions associated with speciation events differed across time and genera. Substrate transitions were more frequently associated with older events than with younger ones. Conclusions Neighbouring areas and a mosaic of local habitats shaped the palm flora of New Caledonia, and the island played a significant role in generating palm diversity across the Pacific region. This new spatio-temporal framework will enable population-level ecological and genetic studies to unpick the mechanisms underpinning New Caledonian palm endemism. [ABSTRACT FROM AUTHOR]

Phylogenetic analyses of mitochondrial and nuclear data of 31 specimens of Cyphocharax from trans‐Andean rivers support the presence of one lineage of Cyphocharax aspilos in Lago Maracaibo and three cryptic lineages of Cyphocharax magdalenae: (1) Cauca‐Magdalena and Ranchería, (2) León and Atrato, and (3) Chucunaque‐Tuira, Santa María, and Chiriquí basins of Central America. Results suggest that the Serranía del Perijá facilitated Late Miocene cladogenetic events, whereas post‐Isthmian C. magdalenae expansion was enabled by gene flow across the lower Magdalena valley and Central American lowlands. Time‐calibrated phylogenetics indicate that the C. magdalenae colonized lower Central America in the Pliocene (3.7 MYA; Ma), the divergence Atrato‐Magdalena occurred in Late Pliocene (3.0 Ma) and the split Ranchería‐Magdalena during the Middle Pleistocene (1.3 Ma). Updated geographic distribution data support the hypothesis that the Cordillera de Talamanca functions as a barrier to northward expansion of C. magdalenae in Central America. [ABSTRACT FROM AUTHOR]

The Peruvian Province, from 6° S in Peru to 42° S in Chile, is a highly productive coastal marine region whose biology and fossil record have long been studied separately but never integrated. To understand how past events and conditions affected today's species composition and interactions, we examined the role of extinction, colonization, geologic changes to explain previously unrecognized peculiar features of the biota and to compare the Peruvian Province's history to that of other climatically similar temperate coasts. We synthesized all available data on the benthic (or benthically feeding) biota, with emphasis on fossilizable taxa, for the interval from the Miocene (23–5.4 Ma) and Pliocene (5.4–2.5 Ma) to the present. We outline the history of ecological guilds including primary producers, herbivores, predators, and suspension‐feeders and document patterns of extinction, colonization, and geographic restriction. We identify twelve unusual attributes of the biota, most of which are the result of repeated episodes of extinction. Several guilds present during the Miocene and Pliocene are not represented in the province today, while groups such as kelps and perhaps intertidal predatory sea stars are relative newcomers. Guilds on soft bottoms and in sheltered habitats were severely affected by extinction, whereas those on hard bottoms were most affected by colonists and held their own in diversity. The Peruvian Province has not served as a biogeographic refuge, in contrast to the coasts of Australasia and Argentina, where lineages no longer present in the Peruvian Province survive. The loss of sheltered habitats since the Pliocene explains many of the present‐day peculiarities of the biota. The history of the province's biota explains its unique attributes. High productivity, a rich Southern Hemisphere heritage, and colonization from the north account for the present‐day composition and unusual characteristics of the biota. [ABSTRACT FROM AUTHOR]

The Miocene sequence in the Roer Valley Rift System consists of alternating open‐to‐shallow marine, coastal and fluvio‐deltaic deposits. In this study, well logs, bio‐chronostratigraphy and seismostratigraphy are used to characterize major units and their bounding unconformities and to infer sediment dispersal patterns. Three major unconformities occur in the sequence: the early, middle and late Miocene unconformities (EMU, MMU and LMU). The EMU formed due to tectonic motions related to the Savian phase. After formation of the EMU, a broad depocentre developed in the south‐eastern part of the Roer Valley Graben (RVG). Sediment accumulation increased during this period and peaked in the middle Langhian, after which it diminished again to a low level during the late Serravallian. The decrease in sediment accumulation coincided with a period of tectonic subsidence along the major bounding fault zones (i.e. the Peel Boundary Fault System, the Feldbiss Fault System and the Veldhoven Fault System). The resulting transgression caused sediment starvation in the central RVG. Subsequently, global sea‐level fall during the early Tortonian caused large‐scale erosion, and formation of incised valleys on the highs adjacent to the RVG (Peel Block and Campine Block), as well as the south‐eastern RVG, forming the MMU. However, sedimentation continued during this period in the central part of the RVG where no erosional hiatus developed. From the Tortonian onwards, accumulation rates increased again. The depocentre shifted towards the north‐west and clinoforms developed in the RVG. During the latest Miocene, the depocentre was concentrated along the south‐western margin of the RVG. Meanwhile, the depositional environment of the entire RVRS gradually shallowed as the LMU was formed. [ABSTRACT FROM AUTHOR]

The Malay Basin is a mature hydrocarbon province currently being re‐assessed for CO2 storage. Selecting an appropriate storage site requires a comprehensive understanding of the structural and stratigraphic history of the basin. However, previous studies have been limited to observations from either regional 2D seismic lines or individual 3D seismic volumes. In this study, we access and utilise a basin‐wide (ca. 36,000 km2) 3D seismic and well database to describe the structural and stratigraphic features of the basin, particularly those within the uppermost ca. 4 km (Oligocene to Recent) and gain new insights into the basin's evolution. E–W transtensional rift basins first developed due to sinistral shear across an NW‐SE strike‐slip zone. The NW‐SE basin morphology seen today was generated during the late Oligocene–early Miocene, during which time dextral motion across marginal hinge zones created en‐echelon antithetic, extensional faults and pull‐apart basins, especially well preserved along the western margin of the basin. Collisional forces to the southeast during the early to middle Miocene resulted in the shallowing of the basin, intermittent connection to the South China Sea and a cyclic depositional pattern. Around 8 Ma (late Miocene), a significant uplift of the basin resulted in a major unconformity with up to 4.2 km of erosion and exhumation in the southeast. In the centre and northwest of the basin, the inversion of deeper E–W rifts resulted in the folding of Miocene sequences and the formation of large anticlines parallel to the rift‐bounding faults. The Pliocene to Pleistocene history is more tectonically quiescent, but some extensional faulting continued to affect the northwest part of the basin. Larger glacio‐eustatic sea‐level fluctuations during this time resulted in major changes in sedimentation and erosion on the Sunda Shelf, including the formation of a middle‐Pliocene unconformity. These structural events have created a variety of hydrocarbon traps across the basin of different ages, including transpressional anticlines, rollover anticlines and tilted fault blocks. Each of these has discrete and distinct trap elements with important implications for their CO2 storage potential. [ABSTRACT FROM AUTHOR]

The present work deals with the carnivorans found in five Neogene localities in eastern Romania: the Vallesian locality of Păun, the Turolian localities of Pogana 1 and Creţeşti 1, and the Ruscinian localities of Bereşti and Măluşteni. The presented material includes cranial and postcranial specimens, some of which are herein described for the first time. This study includes a re-description and re-depiction of the type material of Lutra rumana Simionescu, 1922 and Promephitis malustenensis Simionescu, 1930, in order to clarify the taxonomic position of these two enigmatic taxa. The carnivoran faunal lists of all five localities are reviewed offering interesting insights into the temporospatial range of the discovered forms. Most localities have only yielded a limited number of species, except for Măluşteni, which shows evidence of at least nine different coexisting forms. Notable occurrences in terms of biostratigraphy include Paludolutra sp. in Pogana 1, Protictitherium crassum (Depéret, 1892) in Creţeşti 1, and a phocid from Măluşteni. This review aims to aid the understanding of the Neogene ecosystems in the north of the Paratethys in terms of faunal contents, biostratigraphy and palaeoecology. [ABSTRACT FROM AUTHOR]

Here, we studied several Pachyrukhinae (Mammalia: Notoungulata: Typotheria: Hegetotheriidae) fossils recovered from the late Early Miocene (Santacrucian SALMA) beds of the Cura-Mallín Formation at the Laguna del Laja (Biobío Region, Chile). The specimens are referred to a new species of the genus Pachyrukhos Ameghino, 1885, P. ngenwinkul sp. nov. The holotype of this taxon is a relatively well-preserved articulated skull and mandibles, which was investigated using 3D X-ray Microscopy. The new taxon is distinguished from other Pachyrukhos species by having a p2 smaller than p3, M3 mesiodistally shorter (around 30%) than M1, and upper premolars of a rather subtriangular contour. Pachyrukhos ngenwinkul is the most common mammal and the smallest-sized (with ~2 kg of body mass) notoungulate found in the Santacrucian assemblage at the Laguna del Laja. The discovery of this new taxon improves our understanding of the Neogene pachyrukhine diversity in southern South America and highlights the potential of the Chilean Andean Cenozoic deposits in revealing a hidden diversity of extinct continental mammals. [ABSTRACT FROM AUTHOR]

In this study, we describe the dental remains of a very minute beaver from the early Late Miocene locality Hammerschmiede (MN 7/8) located at the Northern Alpine Foreland Basin (Southern Germany, Bavaria). The finds represent the so far most comprehensive (>1.000 dental specimens) collection of the trogontheriine castorid Euroxenomys minutus. Metrically and morphologically, the large data set demonstrates an extensive intraspecific variability of this beaver from Hammerschmiede that covers most finds of other European localities and confirms previous species assignments. The subspecies Euroxenomys minutus rhenanus from Dorn-Dürkheim (Germany), clearly isolates from this range of variation and can thus be validated. The performed age-frequency distributions for Euroxenomys minutus from the local stratigraphic levels HAM 5 (rivulet) and HAM 4 (river) correspond roughly and also resemble Euroxenomys from the swamp deposit Rudabánya (Hungary). Since previous mortality studies on the also occurring larger beaver Steneofiber depereti from Hammerschmiede demonstrated dominance of behavioural or internal factors, this cross-ecosystem similarity in mortality for the small beaver species is likely explained by external factors, such as predation. Supported by a taphonomic bone analysis, a multitude of predators are finally considered to prey on Euroxenomys minutus and indicate a predatory bone assemblage. [ABSTRACT FROM AUTHOR]

Prospecting fieldwork in a new fossiliferous locality of the marine Early Miocene Gaiman Formation (Patagonia, Argentina) results in the discovery of abundant fossil cetaceans (n = 29). We analyse stratigraphic, sedimentologic, palaeontologic, and taphonomic data with the aim of understanding the factors that control the preservation and distribution of these specimens. The cetacean sample is dominated by small odontocetes exhibiting five different preservation categories. Based on sedimentological features, three distinct facies were recognised in the stratigraphic section. Stratigraphically condensed deposits of facies C, related to a ravinement surface and subsequent transgressive deposits, enclose 52% of the specimens, grading from isolated postcranial elements to articulated and associated postcranial and cranial elements. In contrast, facies A and B exhibit predominantly isolated postcranial elements. Facies C shows higher degrees of corrosion, encrustation and fragmentation than those specimens from facies A and B, reflecting differences in preservation quality among the facies. The higher abundance and evidence of long exposure times recorded for facies C seems to be resulted mostly from physico-chemical factors after stratigraphic condensation. Finally, intrinsic biological features and ecological preferences of the specimens could explain the dominance of small-sized odontocetes in a restricted inner shelf to lower shoreface environmental setting. [ABSTRACT FROM AUTHOR]

The late Neogene megatooth shark, Otodus megalodon (Lamniformes: Otodontidae), is mostly known for its gigantic teeth and vertebrae. Re-examination of the rock matrix surrounding a previously described associated tooth set of O. megalodon from the upper Miocene of Japan resulted in the observation of numerous fragments of tessellated calcified cartilage and placoid scales. The morphology of each tessera and the arrangement of overall tessellated calcified cartilage are practically identical to those of extant chondrichthyans. Many placoid scales possess pronounced, rather broadly-spaced keels. A quantitative relationship between interkeel distances of keeled scales and reported cruising speeds across extant pelagic lamniforms and carcharhiniforms suggests that O. megalodon with a representative interkeel distance of ca. 100 µm was not a fast swimmer. We propose that O. megalodon was generally a slow cruising shark with occasional burst swimming for prey capture, where much of its metabolic heat through regional endothermy was possibly used to facilitate the digestion of large pieces of ingested meat as well as absorbing and processing nutrients. If so, the relative importance of the functional roles of regional endothermy possibly shifted from maintaining high cruising speeds to visceral food processing through the evolution towards gigantism in otodontids. [ABSTRACT FROM AUTHOR]

Since discovery and description of the species, Sahelanthropus tchadensis has been at the centre of a great discussion around its classification as a hominin - the first of our lineage - or an ape. Many studies have been conducted in order to clarify this question, especially studies based on the morphology of the cranium and the post-cranial remains. In this study, we analysed the posterior dentition of S. tchadensis in relation to those of other hominins and chimpanzees, using a multivariate comparative metric analysis. Our results suggest that the posterior dentition of the Chad material lies in the range of well-established early Plio-Pleistocene hominins, supporting its classification as part of the hominin lineage. [ABSTRACT FROM AUTHOR]

Rift basins are commonly characterised by multiple sediment sources (e.g. transverse and longitudinal) that change over time during different rifting evolution stages. Therefore, tracing sediment provenance to decipher the sedimentary source-to-sink analyses processes in rift basins is important to predict sandy reservoirs. The Xihu Sag in the East China Sea Shelf Basin, particularly its western slope, features large-scale sand bodies of unclear origin. We used petrography, heavy minerals and geochemistry to study sediment provenance in the Eocene Baoshi to Miocene Longjing formations. Our work suggests there are three distinct areas (namely zones A, B and C) affected by different source areas. Zone A, influenced by the Hupijiao uplift, primarily contains metamorphic rock minerals; Zone B, affected by the Haijiao and Hupijiao uplifts, contains igneous and metamorphic rock minerals; and Zone C, dominated by the Haijiao uplift, features igneous with some metamorphic sources, which increase over time, indicating stronger sediment input from the northern Hupijiao uplift. Rare earth element analysis suggests a continental margin and island arc setting, primarily with intermediate–acid felsic rock sources. However, a Eu negative anomaly in Eocene upper Pinghu Formation to the Miocene Longjing Formation indicates some basic source contributions, possibly from a volcanic belt east of the South Yellow Sea. This intermediate–basic volcanic rock belt is located to the northwest of the Hupijiao uplift and may provide clasts for zones A and B through axial channels. These findings indicate that varied sediment sources in the different zones and basin evolution stages with a growing extra-basinal contribution from the Eocene to Miocene, which align with the tectonic shift from the rifting to post-rift period, are essential for understanding the sedimentary filling in this area. Three distinct regions are identified, each influenced by different source areas. Rare earth elements indicate that the tectonic setting of the source terranes was active continental margin and continental island arc. From Eocene to Miocene, there was a growing influence of distal source areas entering the basin through the axial fluvial channels. Sediment provenance changes respond to the basin-scale tectonic evolution. [ABSTRACT FROM AUTHOR]

The Ponto-Caspian radiation of gammaroid amphipods is one of the world's most diverse lacustrine crustacean radiations as well as the only lacustrine amphipod radiation with a fossil record (dating back 9 Ma). However, the phyletic relationship between fossils and extant species was never formally investigated, hampering our evolutionary understanding of this diverse clade. Here, we present the first morphology-based phylogeny (114 characters) of Ponto-Caspian gammaroids that includes 99% of extant taxa, all previously known fossils, as well as newly discovered fossil specimens from the Late Miocene (10.5 Ma) of eastern Romania. Our phylogenetic and phenetic results strongly support the monophyly of all fossil taxa. However, despite its widespread Late Miocene distribution throughout the Paratethys Sea, this fossil clade left no modern descendants. Its exact phylogenetic position is ambiguous, although it is nested in the radiation's crown group. Morphometric analyses further indicate that fossil species occupy a narrower and peripheral morphospace relative to extant taxa. Our results support the assignment of the newly discovered Romanian fossils to a new genus, Eogmelina gen. nov. with two new species Eogmelina moldavica gen. et sp. nov. and Eogmelina prisca gen. et sp. nov. Finally, our findings call for a fundamental systematic restructuring of Ponto-Caspian gammaroids at genus and family level. [ABSTRACT FROM AUTHOR]

We provide new insights into the Badenian transgression (Langhian to early Serravallian) in western Serbia, focusing on the marginal part of the Central Paratethys. Nano- and microfossils (calcareous nannofossils, foraminifers, and ostracods) are used along with U–Pb radioisotope dating of a tuff layer near the Badenian lower boundary to reconstruct the timing of the middle Badenian marine flooding. Our findings reveal the occurrence of a second (main) Badenian transgression (TB2.4 third-order cycle) around 14.2–13.8 million years ago. The Badenian marine waters flooded the continental formation of the poorly defined early-middle Miocene sequences. The sedimentary succession (KC-4 core, 490–205 m) is grouped into five units (I–V). The lower two units (I–II) are non-marine alluvials that are characterized by coarse-grained siliciclastics. Unit III represents the basal coarse-grained clastites and sandy limestones of the initial shallow-marine environment and unit IV consists of marls and high-calcitic fine-grained clastites of neritic facies. Unit V contains carbonaceous marlstone and limestone deposited in a restricted marine shallow-water environment. Biostratigraphically, over 154 m of dominantly fine-grained clastites (units III–IV) confirm the presence of the marine middle and upper Badenian. Its lower part (366–325 m) corresponds to the nannofossil NN5 Zone and the youngest Upper Lagenidae Zone (local Zone Ammonia viennensis–Trilobatus trilobus), as well as a part of the Spirorutilus carinatus Zone. The upper part (325–212 m) corresponds to the nannofossil NN6 Zone as well as the foraminifer Bulimina-Bolivina Zone. The U–Pb age for zircon grains is 14.58 ± 0.08 Ma. Petrographic data from the tuff sample show a strong correlation with volcanic records along the S-SW margin of Central Paratethys (N Croatia, NE Bosnia, and central Serbia). The U–Pb age as well as tuff glass composition and mineralogy suggest the possible source of the tuff as from the Subotica area (N Serbia) or the Bükkalja Volcanic Field in NE Hungary. [ABSTRACT FROM AUTHOR]

Phenotypic plasticity can be adaptive in fluctuating environments by providing rapid environment–phenotype matching and this applies particularly in seasonal environments. African Bicyclus butterflies have repeatedly colonized seasonal savannahs from ancestral forests around the late Miocene, and many species now exhibit seasonal polyphenism. On a macroevolutionary scale, it can be expected that savannah species will exhibit higher plasticity because of experiencing stronger environmental seasonality than forest species. We quantified seasonality using environmental niche modeling and surveyed the degree of plasticity in a key wing pattern element (eyespot size) using museum specimens. We showed that species occurring in highly seasonal environments display strong plasticity, while species in less seasonal or aseasonal environments exhibit surprisingly variable degrees of plasticity, including strong to no plasticity. Furthermore, eyespot size plasticity has a moderate phylogenetic signal and the ancestral Bicyclus likely exhibited some degree of plasticity. We propose hypotheses to explain the range of plasticity patterns seen in less seasonal environments and generate testable predictions for the evolution of plasticity in Bicyclus. Our study provides one of the most compelling cases showing links between seasonality and phenotypic plasticity on a macroevolutionary scale and the potential role of plasticity in facilitating the colonization of novel environments. [ABSTRACT FROM AUTHOR]

Cynanchum belongs to the Apocynaceae family and is a morphologically diverse genus that includes around 200 shrub or perennial herb species. Despite the utilization of CPGs, few molecular phylogenetic studies have endeavored to elucidate infrafamilial relationships within Cynanchum through extensive taxon sampling. In this research, we constructed a phylogeny and estimated divergence time based on the chloroplast genomes (CPGs) of nine Cynanchum species. We sequenced and annotated nine chloroplast (CP) genomes in this study. The comparative analysis of these genomes from these Cynanchum species revealed a typical quadripartite structure, with a total sequence length ranging from 158,283 to 161,241 base pairs (bp). The CP genome (CPG) was highly conserved and moderately differentiated. Through annotation, we identified a total of 129–132 genes. Analysis of the boundaries of inverted repeat (IR) regions showed consistent positioning: the rps19 gene was located in the IRb region, varying from 46 to 50 bp. IRb/SSC junctions were located between the trnN and ndhF genes. We did not detect major expansions or contractions in the IR region or rearrangements or insertions in the CPGs of the nine Cynanchum species. The results of SSR analysis revealed a variation in the number of SSRs, ranging from 112 to 150. In five types of SSRs, the largest number was mononucleotide repeats, and the smallest number was hexanucleotide repeats. The number of long repeats in the cp genomes of nine Cynanchum species was from 35 to 80. In nine species of Cynanchum, the GC3s values ranged from 26.80% to 27.00%, indicating a strong bias towards A/U-ending codons. Comparative analyses revealed four hotspot regions in the CPG, ndhA-ndhH, trnI-GAU-rrn16, psbI-trnS-GCU, and rps7-ndhB, which could potentially serve as molecular markers. In addition, phylogenetic tree construction based on the CPG indicated that the nine Cynanchum species formed a monophyletic group. Molecular dating suggested that Cynanchum diverged from its sister genus approximately 18.87 million years ago (Mya) and species diversification within the Cynanchum species primarily occurred during the recent Miocene epoch. The divergence time estimation presented in this study will facilitate future research on Cynanchum, aid in species differentiation, and facilitate diverse investigations into this economically and ecologically important genus. [ABSTRACT FROM AUTHOR]

Subtropical evergreen broad‐leaved forest (EBLF) is the predominant vegetation type in eastern China. However, the majority of the region it covers in eastern China was an arid area during the Paleogene. The temporal history and essential factors involved in the evolution of subtropical EBLFs in eastern China remain enigmatic. Here we report on the niche evolution of Quercus section Cyclobalanopsis, which appeared in south China and Japan during the Eocene and became a dominant component of subtropical EBLFs since the Miocene in eastern Asia, using integrative analysis of occurrences, climate data and a dated phylogeny of 35 species in Cyclobalanopsis. Species within clades Cyclobalanoides, Lamellosa, and Helferiana mainly exist in the Himalaya–Hengduan region, adapting to a plateau climate, while species within the other clades mainly live in eastern China under the control of the East Asian monsoon. Reconstructed history showed that significant divergence of climatic tolerance in Cyclobalanopsis began around 19 million years ago (Ma) in the early Miocene. Simultaneously, disparities in precipitation of wettest/warmest quarter and annual precipitation were markedly enhanced in Cyclobalanopsis, especially in the recent eastern clades. During the Miocene, the marked radiation of Cyclobalanopsis and many other dominant taxa of subtropical EBLFs strongly suggest the rapid formation and expansion of subtropical EBLFs in eastern China. Our research highlights that the intensification of the East Asian monsoon and subsequent occupation of new niches by the ancient clades already present in the south may have jointly promoted the formation of subtropical EBLFs in eastern China since the early Miocene. [ABSTRACT FROM AUTHOR]