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The mole crab Emerita portoricensis Schmitt, 1935 was originally described solely on the basis of few key characters that were not precisely defined, giving reason to question subsequent reports of its distribution. The present study, prompted by recent collections documenting coloration in life, undertakes a comprehensive redescription of the species based on specimens of varied sizes from Puerto Rico, Belize, Costa Rica, and Panama. Collections from the northern Caribbean that at first take appear to represent a northernmost record of E. brasiliensis Schmitt, 1935 or southernmost occurrence of E. talpoida (Say, 1817), may be assignable E. portoricensis as now recognized. Among western Atlantic species, E. portoricensis and E. benedicti have to date been considered to have the dactylus of the first pereopod terminally subacute or sharply pointed, which purportedly separates them from E. brasiliensis and E. talpoida, western Atlantic species in which this article is terminally rounded. However, in E. portoricensis this character varies with specimen size and the magnification at which the distal extreme of the dactylus is examined, being rounded to varying degrees in all but the largest specimens. Even in sexually mature specimens of less than maximum size, this rounded tip is armed by a minute corneous spine in E. portoricensis, although it is less prominent than the terminal spine on the consistently more acute dactylus of E. benedicti at all adult sizes. Also, the carapace color in live specimens of E. portoricensis, as documented for specimens collected in both Belize and Panama, differs from that of E. brasiliensis, E. talpoida, and E. benedicti by typically including longitudinal and diagonal dark bars of olive brown on the branchial regions and a light longitudinal bar marking the posterior quarter of the median line. Posterior to the cervical groove, fine rugae of the carapace that form broken transverse lines are at most little diminished across the mid-dorsal longitudinal line in E. portoricensis and E. benedicti, somewhat more broken in E. brasiliensis, and distinctly diminished to all but absent at the midline in E. talpoida. Previously reported BINs in the Barcode of Life database include sequenced specimens from Costa Rica herein accepted as E. portoricensis. We exclude populations from Brazil that have been mis-assigned to E. portoricensis.

A new amphipod species belonging to the genus Gammarus is described from the Gökgöl Cave, Zonguldak Province, Türkiye. The newly-identified species is relatively small (13 mm) and is a member of the Gammarus pulex-group by the presence of numerous long setae along the posterior margins of pereopods 3 and 4. The specimens were sampled from a shallow pond located in the dark zone (about 1 km inside the entrance) of the cave. Minute eyes, setose (both peduncle and flagellar segments) second antenna, slightly swollen flagellar segments of the second antenna, setose pereopods 3 and 4 and relatively short endopod/exopod ratio of the third uropod are the character combination of the newly-identified species in addition to lacking body pigmentation. The molecular phylogeny, based on the concatenated dataset (28S+COI, 1495 bp) indicated that the new species was resolved from the other Gammarus species by high bootstrap (NJ: 100, ML: 100). In addition to Gammarus tumafsp. nov., mtDNA COI and nuclear DNA 28S gene data of Gammarus baysaliÖzbek et al., 2013 were recorded for the first time. The newly-identified species was well-differentiated from the genetically closest species, G. baysali, with genetic distance of 12.22% and 0.55% for the COI and 28S genes, respectively. Detailed descriptions and drawings of the extremities of the holotype male were given and the morphology of the newly-identified species is compared with its relatives.

Austinixa bragantina Coelho, 2005, described from Ilha Canela, Bragança, state of Pará, Brazil, is established as a junior synonym of A. aidae (Righi, 1967) based on morphological (gross morphology and scanning electron microscopy) and molecular (16S rRNA mitochondrial marker) evidence. The phylogenetic tree was obtained through the Maximum Likelihood method aiming to contextualize A. bragantina among related species. We provide a redescription, an updated synonymy, and remarks on the intra-specific morphological variability of A. aidae to clarify some taxonomic inconsistencies previously reported. Austinixa hardyi Heard & Manning, 1997, a species of uncertain taxonomic status, was confirmed genetically to also be a synonym of A. aidae.

Karasawa, Hiroaki, Ohara, Masaaki, Kato, Hisayoshi (2023): Validation of the names of four species of Decapoda and one species of Isopoda from the Lower Cretaceous (Barremian) Arida Formation of central Japan. Zootaxa 5277 (1): 198-200, DOI: 10.11646/zootaxa.5277.1.12, URL: http://dx.doi.org/10.11646/zootaxa.5277.1.12

CHAETOGAMMARUS TRICHIATUS MARTYNOV, 1932 (FIGS 8–11) Brief redescription: Based on specimen ZMMU Mb-1227, ♂, Russia, Krasnodar Krai, Sochi Urban Okrug, Lazarevsky District, mouth of the Ashe River, in the stream under stones (43°57.376ʹN, 39°15.954ʹE), 13 May 2019, coll. D. Palatov & I. Marin. Relatively large species (≤ 15 mm); females smaller than males. Head with oblique anteroventral lobe with distally produced anterior margin (Fig. 6B), eyes large, reniform, well pigmented. Body unpigmented, moderately elongated, generally smooth, non-carinate. Urosomites 1–3 smooth, without elevations; urosomites 1 and 2 with solitary strong median and submedian spines; urosomite 3 with two submedian spines and a pair of median simple setae (Fig. 6A). Antenna 1 feebly setose, with small aesthetascs; accessory flagellum eight-segmented (Figs 7A, 8A). Antenna 2 is about one-third shorter than antenna 1, massive and deeply setose in males and less setose in females, without calceoli and aesthetascs (Figs 7B, 8C, D). Upper lip (labrum) with convex distal part (Fig. 9A). Lower lip (labium) usually with mostly reduced inner lobes (Fig. 9B). Mandible with well-developed palp and setose basal segment (Fig. 9C–F). Maxilla 1 with outer lobe bearing hairbrush-like distal setae, subequal to inner lobe; inner lobe distally expanding with a row of marginal setae (Fig. 9G). Maxilla 2 with outer lobe wider than inner lobe, bluntly expanding distally (Fig. 9I). Maxilliped with outer and inner plates wide (Fig. 9J). Gnathopod 1 is smaller than gnathopod 2, with weak sexual dimorphism, stronger in males; propodus (palm) teardrop shaped, with oblique palmar margin in males and females (Fig. 8E, F). Gnathopod 2 is significantly larger in males, rectangularly elongated, with straight palmar margin in both males and females (Fig. 8H, I). Basis (article 2) of pereopods 5–7 with feebly marked ventral lobes (Fig. 10D, F, G); pereopod 3 with highly developed coiled setation along posterior margin (Figs 7D, 10A); basis of pereopod 7 elongated, ~2.5 times as long as wide, without ventral lobe (Figs 7C, 10G). Pleopods with two elongated hooks and one or two thick bristles in retinacules (Fig. 11F, G). Uropod 3 exopod broad, furnished with clusters of long, distally coiled setae, about four to six times as long as wide, with reduced distal article furnished with numerous long, simple distal setae (Figs 6D, 11K, J). Epimeral plates 1–3 with sharply produced inferoposterior corners (Fig. 11A–C). Telson deeply cleft into suboval lobes, abruptly tapering distally, bearing clusters of strong, stout setae distally and submedially (Figs 6A, 11D, E). Remarks: Males of C. trichiatus can be distinguished from males of other species of Chaetogammarus by the highly developed coiled setation on pereopod 3 posterior margin and uropod 3 exopod, acutely produced anterolateral head lobes, and telson lobes abruptly tapering distally. Chaetogammarus trichiatus is most similar to C. ischnus, but differs from that species in the following traits: significantly more elongate appendages (Fig. 5C); epimeres 2 and 3 with more produced posterodistal corners; presence of coiled setae along the outer margin of uropod 3 exopod (Fig. 6D); longer basis of pereopod 7 (length = 3 × width vs. length = 2 × width; Fig. 7C). Distribution: Chaetogammarus trichiatus is endemic to lakes, lower (slow current) river stretches and river mouths along the north-eastern Black Sea coast, from Abrau to the Shahe River, and a separate population is known from New Athos (Fig. 4). Found only on substrates with pebbles or large stones and boulders, apparently avoiding sandy and silty substrates. Type locality: Estuarine part of Khosta River, Sochi area, Russia. Presently absent at its original type locality owing to anthropogenic influence and the urban reconstruction of the lower part of the river., Published as part of Copilaș-Ciocianu, Denis, Palatov, Dmitry, Rewicz, Tomasz, Sands, Arthur F., Arbačiauskas, Kęstutis, Haaren, Ton Van, Hebert, Paul D. N., Grabowski, Michał & Marin, Ivan, 2023, A widespread Ponto-Caspian invader with a mistaken identity: integrative taxonomy elucidates the confusing taxonomy of Trichogammarus trichiatus (= Echinogammarus) (Crustacea: Amphipoda), pp. 821-846 in Zoological Journal of the Linnean Society 198 on page 834, DOI: 10.1093/zoolinnean/zlad010, http://zenodo.org/record/8152446, {"references":["Martynov AB. 1932. Contribution to the knowledge of the freshwater fauna of the Black Sea coast of Caucasus. I. Amphipoda. TraVaux de l'Institut Zoologique de l'Academie des Sciences de l'URSS 1: 73 - 98."]}

Amphipods have diversified greatly in the Ponto-Caspian region. Although many of these species are prominent invaders their systematics remains unclear. Taking an integrative approach, we investigate the taxonomy of Trichogammarus trichiatus, a widespread invader in European inland waters. It was initially described from the north-eastern Black Sea coast as Chaetogammarus trichiatus by Martynov in 1932. A similar taxon, Chaetogammarus tenellus major, was described by Cărăușu from the western Black Sea in 1943 but later synonymized with C. trichiatus. Chaetogammarus trichiatus was itself shuffled between Chaetogammarus and the Atlanto-Mediterranean Echinogammarus, currently being assigned to Trichogammarus. Our analyses (six DNA markers, 60 measurements and scanning electron microscopic imaging) reveal that T. trichiatus and C. tenellus major are distinct species; the former is a Caucasian endemic, whereas the latter invaded Europe. Unexpectedly, T. trichiatus is an incipient species molecularly nested in Chaetogammarus ischnus, despite pronounced morphological and geographical differentiation. We also recover Chaetogammarus as polyphyletic, yet its member species are nested in the Ponto-Caspian radiation, thus distinct from Echinogammarus. Consequently, we reassign T. trichiatus to Chaetogammarus (Chaetogammarus trichiatus), synonymize Trichogammarus with Chaetogammarus and place C. tenellus major in the new genus Spirogammarus gen. nov. (Spirogammarus major comb. & stat. nov.). Chaetogammarus necessitates further systematic refinement.

GENUS SPIROGAMMARUS COPILAȘ-CIOCIANU, PALATOV, MARIN & GRABOWSKI GEN.NOV. PALATOV, MARIN & GRABOWSKI GEN.NOV. Z o o B a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. o r g: a c t: D 5 2 6 9 6 B 2 - A E B A - 4 0 D B - A 7 C C - BF712AAD3780 Type and only species: Chaetogammarus tenellus major CărăuȘu, 1943. Diagnosis: Large Echinogammarus -like gammarid amphipods (≤ 16 mm). Body smooth and non-carinate. Head anterolateral lobe straight. Antenna 1 poorly setose, accessory flagellum well developed (up to nine articles). Antenna 2 flagellum and peduncle segments densely set with tufts of setae twice as long as the width of underlying segment (distally coiled in males), calceoli absent. Gnathopod 1 propodus subequal to gnathopod 2 propodus. Posterior margin of carpal and meral (sometimes ischial) articles of pereopod 3 set with dense and distally coiled setae as long as/longer than the width of underlying segment; pereopods 4–7 with sparse and short setae, dominated by spines. Uropod 3 exopod with dense setae as long as/longer than the width of underlying segment (coiled in males); endopod short and rudimentary; pleonites unarmed dorsally; epimeral plates with spines only (second straight; third with produced inferoposterior corner); urosomites 1–2 without elevations, armed with four dorsolateral clusters of two to four spines. Telson fully cleft, as long as broad, armed with distal and lateral spines, with lobes abruptly tapering distally. Etymology: From the Latin spira, meaning coil/ spiral, with reference to the coiled setae found on the antennae, gnathopods, pereopods and uropod 3 in males. The gender is masculine. Remarks: Spirogammarus can be distinguished readily from any species of Chaetogammarus and Echinogammarus s.l. by the presence of dense, distally coiled setae on the carpus and merus of pereopod 3 that are as long as/longer than the width of the underlying segment in both sexes. To our knowledge, no Echinogammarus -like taxa have females with coiled setae on the carpus and the merus of pereopod 3 (Pinkster, 1993). Spirogammarus is similar to Echinogammarus tibaldii Pinkster & Stock, 1970 with respect to the highly developed setosity of antenna 2, pereopod 3 and weakly setose pereopods 5–7 (Pinkster, 1993). However, Spirogammarus is distinguished by the weak setation of antenna 1 (highly developed), higher number of articles in the accessory flagellum (nine vs. five), coiled setae on gnathopod propodi and uropod 3 (non-coiled), and absence of long setae along the distal margin of epimeres. For further comparisons, see Supporting Information, Table S4., Published as part of Copilaș-Ciocianu, Denis, Palatov, Dmitry, Rewicz, Tomasz, Sands, Arthur F., Arbačiauskas, Kęstutis, Haaren, Ton Van, Hebert, Paul D. N., Grabowski, Michał & Marin, Ivan, 2023, A widespread Ponto-Caspian invader with a mistaken identity: integrative taxonomy elucidates the confusing taxonomy of Trichogammarus trichiatus (= Echinogammarus) (Crustacea: Amphipoda), pp. 821-846 in Zoological Journal of the Linnean Society 198 on pages 831-832, DOI: 10.1093/zoolinnean/zlad010, http://zenodo.org/record/8152446, {"references":["Carausu S. 1943. Amphipodes de Roumanie I. Gammarides de type Caspian. Bucharest: Institutul de Cercetari Piscicole al Romaniei.","Pinkster S. 1993. A revision of the genus Echinogammarus (Stebbing, 1899) with some notes on related genera. Memorie del Museo CiVico di Storia Naturale, (lia serie), (Sezione scienze della Vita. A, Biologia) 10: 1 - 183."]}

GENUS CHAETOGAMMARUS MARTYNOV, 1924 Type species: Gammarus ischnus Stebbing, 1899. I n c l u d e d s p e c i e s: C h a e t o g a m m a r u s h y r c a n u s Pjatakova 1962, C. pauxillus (Sars, 1896), C. placidus (Sars, 1896), C. aearpachoaeskyi (Sars, 1894) and C. trichiatus Martynov, 1932. Non-included species: Echinogammarus karadagiensis Grintsov, 2009 and E. mazestiensis Marin & Palatov, 2020 will be transferred to a new genus (I. Marin, D. Palatov, and D. CopilaȘ- Ciocianu, unpub. obs.). Amended diagnosis: Small- to large-sized species (≤ 15 mm); females smaller than males. Head with oblique anteroventral lobe, sometimes distally produced. Eyes large, reniform, well pigmented. Body non-carinate. Antenna 1 feebly setose, accessory flagellum two- to eight-segmented; antenna 2 usually shorter that antenna 1, without calceoli, deeply setose in males and less setose in females. Lower lip (labium) with mostly reduced inner lobes. Gnathopod 1 smaller than gnathopod 2, sexually dimorphic, stronger in males, propodus generally teardrop shaped; gnathopod 2 sexually dimorphic, stronger in males, propodus trapezoidal with oblique palmar margin. Basis of pereopods 5–7 without or with weak posteroventral lobes, dominated by spines. Uropod 3 with exopod about four to six times as long as wide, generally dominated by spines. Urosomites 1 and 2 without elevations, armed with clusters of spines; urosomite 3 with a pair of submedian spines. Telson deeply cleft into suboval lobes, bearing clusters of spines distally and submedially. Synonymized genera: Trichogammarus Hou & Sket, 2016. Distribution: The native range of the genus is restricted to the Ponto-Caspian basin. Its species occur in the Caspian Sea at up to 500 m depth, and in lagoons, estuaries and lower stretches of rivers that drain into the Black, Azov and Caspian seas., Published as part of Copilaș-Ciocianu, Denis, Palatov, Dmitry, Rewicz, Tomasz, Sands, Arthur F., Arbačiauskas, Kęstutis, Haaren, Ton Van, Hebert, Paul D. N., Grabowski, Michał & Marin, Ivan, 2023, A widespread Ponto-Caspian invader with a mistaken identity: integrative taxonomy elucidates the confusing taxonomy of Trichogammarus trichiatus (= Echinogammarus) (Crustacea: Amphipoda), pp. 821-846 in Zoological Journal of the Linnean Society 198 on pages 833-834, DOI: 10.1093/zoolinnean/zlad010, http://zenodo.org/record/8152446, {"references":["Martynov AB. 1924. Etudes sur les Crustaces de Mer du Bassin du bas Don et leur distribution ethologique. Annuaire du Musee Zoologique de l'Academie des Sciences de Russie 25: 1 - 115.","Pjatakova GM. 1962. Novye formy gammarid Kaspiiskogo Morya. Doklady Akademii Nauk Azerbaidzhanskoi SSR 18: 47 - 51.","Sars GO. 1896. Les Crustacees caspiennes. Contributions pour server a la connaissance de la faune carcinologique de la mer Caspienne. Bulletin de I'Academie Imperiale des Sciences de St. - Petersbourg 4: 421 - 489.","Martynov AB. 1932. Contribution to the knowledge of the freshwater fauna of the Black Sea coast of Caucasus. I. Amphipoda. TraVaux de l'Institut Zoologique de l'Academie des Sciences de l'URSS 1: 73 - 98.","Grintsov V. 2009. A new amphipod species Echinogammarus karadagiensis sp. n. (Amphipoda, Gammaridae) from Crimean coasts (Black Sea). Vestnik Zoologii 43: e 23.","Hou Z, Sket B. 2016. A review of Gammaridae (Crustacea: Amphipoda): the family extent, its evolutionary history, and taxonomic redefinition of genera. Zoological Journal of the Linnean Society 176: 323 - 348."]}

SPIROGAMMARUS MAJOR (CĂRĂUȘU, 1943) COMB. & STAT. NOV. Chaetogammarus tenellus major CărăuȘu, 1943. Diagnosis: Same as for genus (see above). Remarks: Spirogammarus major can be distinguished from C. trichiatus (in parentheses) by: its larger size (16 vs. 12 mm; Fig. 5C), absence of setae on the basal segment of mandibular palp, subequal gnathopod propodi (propodus 2 longer than propodus 1), straight anterolateral head lobes (acutely produced; Fig. 6B), denser setation along the inner side of the gnathopod palm (Fig. 6C), urosomites armed with clusters of two to four spines (solitary spines; Fig. 6A), uropod 3 exopod with dense setation in both sexes (only males have dense setation; Fig. 6D), and uropod 3 endopod in males is armed with spines and long setae (setae absent in both sexes; Fig. 6D). Both taxa are similar with respect to the setosity of antennae (Fig. 7A, B) and pereopod 3 (Fig. 7D). Distribution: Lagoons, estuaries and lower stretches of rivers that drain along the north-western to western shore of the Black Sea. Our morphomolecular analyses confirm that S. major is a widespread invasive species in European inland waters, being widely distributed along the Danube, lower Rhine, Oder and Dnieper rivers (Fig. 4). Type locality: Lake Shabla (Shablensko Ezero), Bulgaria., Published as part of Copilaș-Ciocianu, Denis, Palatov, Dmitry, Rewicz, Tomasz, Sands, Arthur F., Arbačiauskas, Kęstutis, Haaren, Ton Van, Hebert, Paul D. N., Grabowski, Michał & Marin, Ivan, 2023, A widespread Ponto-Caspian invader with a mistaken identity: integrative taxonomy elucidates the confusing taxonomy of Trichogammarus trichiatus (= Echinogammarus) (Crustacea: Amphipoda), pp. 821-846 in Zoological Journal of the Linnean Society 198 on page 833, DOI: 10.1093/zoolinnean/zlad010, http://zenodo.org/record/8152446, {"references":["Carausu S. 1943. Amphipodes de Roumanie I. Gammarides de type Caspian. Bucharest: Institutul de Cercetari Piscicole al Romaniei."]}

Genus Politolana Bruce, 1981 Bruce (1981): 958–959; Kensley & Schotte (1989): 140–143; Wetzer et al. (1987): 1–11; Riseman & Brusca (2002): 64. Type species. Aega polita Stimpson (1853); by original designation. Remarks. The genus Politolana has 13 species, which are distributed in the North Atlantic (seven species), South Atlantic (three species), and Pacific (three species). Riseman & Brusca (2002) reviewed the genus based on a phylogenetic analysis and demonstrated that it is polyphyletic, given that only nine of the 12 species analyzed formed a monophyletic clade, while the three Pacific species were identified as non- Politolana sensu strictu. Riseman & Brusca (2002) restrict Politolana sensu strictu to the Atlantic Ocean, where the majority of its diversity is found in the North Atlantic, in the region of the Gulf of Mexico. However, this arrangement requires careful reconsideration, in particular, the inclusion of other cirolanid genera, a more ample character matrix, and more comprehensive discussion and analysis, including a molecular approach, to better understand the phylogenic relationships among the species and related genera.

Paiva, Ricardo J. C., Souza-Filho, Jesser F. (2023): Three new species of the family Cirolanidae Dana, 1852 (Isopoda, Cymothoida) from the shallow Brazilian bathyal. Zootaxa 5319 (1): 1-26, DOI: 10.11646/zootaxa.5319.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5319.1.1

Genus Natatolana Bruce, 1981 Natatolana Bruce, 1981: 957, Bruce 1986: 52, Brusca & Iverson 1985: 37, Botosaneanu et al. 1986: 412, Wetzer et al. 1987: 2, Brandt 1988: 102, Kensley & Schotte 1989: 139, Brusca et al. 1995: 74, Keable 2006: 139. Type species. Cirolana hirtipes Milne Edwards, 1840; by original designation. Genus remarks. Natatolana is similar to both Politolana and Dolicholana, although it can be readily distinguished from Politolana by the shape of pereopods 5–7. The basis in Politolana is narrow and has a few setae, while in Natatolana it is expanded and has many long plumose setae (Bruce, 1981). In Natatolana, in addition, the appendix masculina arises basally rather than sub-medially as observed in Politolana (Bruce, 1981). Natatolana also differs from Dolicholana in having the frontal lamina ventral surface entirely flat and not forming a 90° angle with the clypeus. Natatolana is the second most diverse cirolanid genus, with 74 species, including one fossil form from the Lower Cretaceous (Bruce & Olesen 1995, Brusca et al. 1995, Keable & Bruce 1997, Keable 2006, Vegan et al. 2019). Natatolana species have been recorded in all the world’s oceans, and the cosmopolitan distribution if these isopods can be explained by their pelagic or partially pelagic way of life (Bruce 1981, Brusca et al. 1995, Bruce 2003). Natatolana species have mostly been recorded from the intertidal zone to depths of 1000 m, and only eight species are known to occur at depths of more than 1000 m, with Natatolana rekohu Bruce, 2003 reaching a depth of 2769 m off New Zealand, and Natatolana natalis (Menzies & George, 1972). However, Keable (2006) reported Natatolana from depths of around 6000 m, based on a specimen from the Peru-Chile Trench (USNM 120961), although its poor condition impeded the identification of the species.

Dolicholana brucei Paiva & Souza-Filho, 2015 Material examined. 1 female (9.5 mm), Revizee Central, station # 6752, Box Corer, 16°49’58.8”S 38°35’27.6”W, 250 m, 21 June 2002, MNRJ 20209. Distribution. Rio Grande do Norte and Bahia (Brazil), 150–250 m depth. Remarks. Dolicholana brucei was described from material collected from the Potiguar Basin, off Rio Grande do Norte (Brazil), at a depth of 150 m. Its latitudinal distribution is now extended southward to Bahia and its depth range to 250 m. The female specimen examined here agrees with the original description, although the two distal robust setae of the pleotelson are broken.

Politolana cleoniceae spec. nov. (Figs 13–17) Material examined. Holotype: female, 45.5 mm, (dissected and drawn, dissected appendages in alcohol 75%), Pernambuco, Continental Shelf, baited bottom traps, 8°30’07.5”S, 34°41’37.7”W, 307 m, 06 October 2015, MOUFPE 15530. Type locality. Brazil, 8°30’07.5”S, 34°41’37.7”W. Etymology. Named in honour of Cleonice Peres de Carvalho, mother of first author. Diagnosis. Head interocular furrow present, submarginal sulcus present, incomplete dorsal groove. Antenna 1 not reaching pereonite 1. Antenna 2 reaching pereonite 1 posterior margin. Pereopods 1–4 basis without plumose marginal setae. Pereopod 7 merus with anterodistal angle rounded and very expanded. Pleonite 1 visible. Pleotelson as long as wide, smooth, apex truncated with a subacute medial process, distal margin with plumose setae, or with 2 RS. Description of female holotype. Body 2.8 times as long as greatest width, dorsal surfaces smooth, widest at pereonite 6, lateral margins slightly convex. Rostral point absent, frontal margin rounded. Eyes separated by about 80% head width, eye colour white. Pereonite 1 and coxae 2–3 each with posteroventral angle rounded; coxae 4–7 with entire oblique carina; pereonites 5–7 posterior margins smooth. Pleon with pleonite 1 not concealed by pereonite 7, pleonites 2–5 posterior margin smooth; pleonite 2 posterolateral angles forming subacute point, not posteriorly produced; pleonite 3 posterolateral margins not extending to pleonite 5 posterior margin, acute; pleonite 4 posterolateral margin acute extended beyond pleonite 5 posterior margin; pleonite 5 with posterolateral angles overlapped by pleonite 4 lateral margin. Pleotelson shorter than anterior width (0.74 times); without longitudinal carina; without dense patch of setae; lateral margins convex, smooth; posterior margin broadly rounded with a middle subacute process, with plumose setae, and a row of RS. Antenna 1 peduncle articles 1 and 2 distinct, articulated, article 2 0.3 times as long as article 1, article 3 and 4 0.8 times as long as combined lengths of articles 1 and 2, article 3 1.3 times as long as wide; flagellum moderately inflated, with 20 articles, extending beyond eye midway. Antenna 2 peduncle article 4 almost as long as wide, 1.45 times as long as article 3, with 6 short simple setae (distal); article 5 1.8 times as long as article 4, 2.8 times as long as wide, anterodistal angle with cluster of 3 short simple seta; flagellum with 31 articles, extending to pereonite 3. Frontal lamina 3.75 times longer than greatest width, lateral margins straight, slightly widening anteriorly, not visible in dorsal. Mandible molar process anterior margin with more than 20 flat teeth; without proximal cluster of long simple setae; right mandible spine row composed of 12 spines; palp article 2 with 17 distolateral setae, article 3 with 24 slender setae. Maxillula mesial lobe with 3 large CPS RS; lateral lobe with 10 RS. Maxilla lateral lobe with 2 long simple setae; middle lobe with 16 long simple setae; mesial lobe with 2 simple setae and 12 distal pappose setae. Maxilliped palp article 1 mesial margin without setae, lateral margin without setae; article 2 mesial margin with 6 slender setae, lateral margin with 9 simple setae; article 3 mesial margin with 10 slender setae, lateral margin with 19 slender setae; article 4 mesial margin with 9 slender setae, lateral margin with 12 simple setae; article 5 distal margin to mesial margin with 16 simple setae, lateral margin with 6 simple setae; endite with 5 long CPS and 2 coupling setae. Pereopod 1 basis 3 times as long as greatest width, superior margin setose, superior distal angle without cluster of setae, inferior margin without setae, inferior distal angle with a cluster of 4 long and slender setae; ischium 1.5 times as long as basis, inferior margin with 12 simple setae, inferior distal margin without setae, superior margin with a middle cluster of 6 simple setae, superior distal margin not produced over merus and with a line of long and slender setae; merus inferior margin with a row of 6 simple setae, superior distal angle produced and reaching 1/2 of carpus, superior margin with a row of both simple and RS; carpus inferior margin with 4 setae, inferior distal angle without setae; propodus 4.1 times as long as wide, inferior margin with 9 simple setae and 2 RS, inferior distal margin with 1 RS and 3 simple setae, superior distal margin with 4 short setae; dactylus 0.5 times as long as propodus. Pereopod 2 ischium inferior margin with 6 long simple setae, inferior distal margin with 6 simple setae, superior distal margin not produced over merus and with a cluster of long and slender setae, mesial margin with a cluster of long and slender setae; merus inferior margin with 1 stout RS, inferior distal margin with 2 RS, superior distal angle produced and reaching 1/4 of propodus, superior margin with a row of both long and simple; carpus inferior margin with 7 stout RS, inferior distal margin with 3 long acute RS; propodus 1.45 times as long as wide, inferior margin without setae, inferior distal margin with 1 short RS and 2 simple setae; dactylus 0.9 times as long as propodus. Pereopod 3 similar to pereopod 2. Pereopods 4–6 similar to each other. Pereopod 7 basis inferior distal angle with a 7 long and plumose setae; superior margin and facial middle line densely setose (plumose setae); basis slightly ovate, 2 times as long as greatest width; ischium 0.5 times as long as basis, 1.5 times as long as wide, widened distally; merus 0.6 times as long as ischium, 1.4 times as long as wide; carpus 0.8 times as long as ischium, 2.4 times as long as wide; propodus 1.2 times as long as ischium, 6 times as long as wide, inferior margin with 4 RS; dactylus 0.45 times as long as propodus. Pereon-pleon articulation tight, with pleonite 1 visible, not completely overlapped by pereonite 7. Pleopod 1 exopod 1.7 times as long as wide, lateral margin convex, distally broadly rounded with distolateral notch, with PMS from entire lateral to distal one-third mesial margin, with ~60 PMS; endopod 1.5 times as long as wide, distally narrowly rounded, with PMS from distal one-third, mesial margin with PMS from distal one-third, endopod with ~40 PMS; peduncle 0.4 times as wide as long, mesial margin with 5 coupling setae. Pleopod 2 exopod with ~90 PMS, endopod with ~30 PMS. Pleopod 3 exopod with incomplete suture, with ~100 PMS, endopod with ~20 PMS. Pleopod 4 exopod with incomplete suture, with ~80 PMS, endopod with ~20 PMS. Pleopod 5 exopod with incomplete suture, with ~35 PMS from distal two-third, endopod with 7 PMS distally. Pleopods 2–5 endopod distal margin with rounded. Uropod peduncle ventrolateral margin with 7 PMS, posterior lobe 0.9 times as long as endopod; rami slightly shorter than pleotelson, lateral, distal and mesial margin lined with long PMS. Endopod apically acute; lateral margin convex, without prominent excision; mesial margin almost straight. Exopod not extending to end of endopod, 4.4 times as long as greatest width, apically acute; lateral and mesial margin straight, lateral margin with 6 RS. Distribution. Known only from the type locality, off Pernambuco state (Brazil), at 307 m depth (Fig. 7). Remarks. Politolana cleoniceae spec. nov. is closest to Politolana haneyi Riseman & Brusca, 2002, especially considering features such as the close pereon–pleon articulation, in which pleonite 1 is visible, and is not overlapped completely by pereonite 7, with the pleon arched (in the dorsal–ventral view) in a uniform sequence with the pereon, and the endopod of the uropod being shorter than the exopod. However, the new species can be differentiated from P. haneyi by the following combination of characters (the characters of P. haneyi are shown within parentheses): antenna 2 does not reach the posterior margin of pereonite 1 (vs. reaches the anterior margin of pereonite 3), coxa 7 slender with a visible lateral oblique suture (vs. suture absent), basis of pereopods 1–4 with no stout CPS (vs. with 6-7 stout CPS), anterodistal angle of the menus of pereopod 7 rounded and very expanded (vs. straight, expanded discreetly), the endopod of the uropod with no cluster of long distal setae and distomedial margin excavated at an angle of approximately 90° (vs. clusters of long distal setae present and distomedial margin concave), internal basolateral angle of pleopod 5 with no lobe (vs. a long hook-shaped lobe near the peduncle), pleotelson with rounded distal margin bearing a medial distal spine with a robust seta on each side (vs. distal margin with no distal medial spine, but four robust distal setae).

Genus Dolicholana Bruce, 1986 Dolicholana Bruce, 1986: 122; Wetzer et al. 1987: 2 (key), Keable 1999: 396, Bruce et al. 2002: 145. Type species. Cirolana elongata H. Milne-Edwards, 1840; by original designation. Remarks on the genus. The genus Dolicholana Bruce, 1986 includes four species: Dolicholana brucei Paiva & Souza-Filho, 2015, Dolicholana elongata (Milne Edwards, 1840), Dolicholana enigma Keable, 1999, and Dolicholana porcellana (Barnard, 1936). Dolicholana resembles both Natatolana Bruce, 1981 and Politolana Bruce, 1981, and in order to distinguish these genera, Bruce (1986) emphasized the projection of the frontal lamina, the morphology of the peduncle of antenna 1, and the absence of plumose setae on the endopod of pleopods 3–5 in Dolicholana. Keable (1999) emended the diagnosis of the genus and added a further character, the formation of a 90° angle between the frontal lamina and the clypeus, which is unique in the cirolanid genera. Keable (1999) described some additional diagnostic characters of the genus Dolicholana and, in the case of the pleotelson, he stated that the dorsal surface is smooth with an anterodorsal depression, with no tubercles, pits or ridges, the margins are convex, meeting at an acute or rounded apex, with robust setae, and abundant plumose setae restricted to the posterolateral margins, but no marginal teeth-like serrations. In the adult males, the appendix masculina of pleopod 2 arises basally, although in juvenile D. enigma, it arises medially. In the new species described here, however, the distal margin of the pleotelson has no robust setae, and the adult appendix masculina arises medially (Fig. 6B). Despite these differences, we decided to allocate the new species to the genus Dolicholana until a more ample phylogenetic review of the Cirolanidae can clarify the relationships between this and other related genera, as suggested by Keable (1999).

Dolicholana ivanyae spec. nov. (Figs 1–6) Material examined. Holotype: male, 15 mm, (dissected and drawn, 8 slides), Revizee Sul, station # 6743, Box Corer, 23°55’40”S, 42°47’60”W, 508 m, 15 February 1998, MNRJ 20208. Paratypes: 1 manca (8 mm, drawn, pereopod 7 present not full developed), Santos Basin, S „o Paulo, Brazil, Station BS-N-07, 137G266, 26º03’26.15”S, 45º42’28.88”W, 24 May 2002, 440 m, MZUSP 25641. 1 manca (7 mm, lacking pereopod 7), Revizee Central, station # 6752, Box Corer, 15°49’58.8”S 38°35’27.6”W, 250 m, 16 February 1998, MNRJ 20207. Type locality. Brazil, 23°55’40”S, 42°47’60”W. Etymology. Named after Ivanya Vieira da Silva Paiva, wife of the first author. Diagnosis. Head interocular furrow present, submarginal furrow absent, dorsal furrow complete. Eyes separated by about 50% head width, strongly pigmented. Frontal lamina 1.5 times as long as wide, linguiform, anterior margin rounded, visible in dorsal view and reaching antenna 1 article 2. Pereopod 7 different from pereopod 6 in shape and ornamentation, densely setose, basis, ischium and merus anterior and posterior margins with very long plumose setae. Pleonite 1 not visible in dorsal view. Pleonite 3 posteroventral angle subacute and entire. Pleopods 3–5 endopod lacking marginal setae. Pleopod 5 exopod with complete suture. Pleotelson longer than wide, apex broadly rounded, distal margin with plumose setae, lacking RS. Description of male holotype. Body 3.1 times as long as greatest width, dorsal surface smooth, widest at pereonite 5, lateral margins slightly convex. Rostral point absent, frontal margin truncate. Eyes separated by about 50% head width, eye colour black. Pereonite 1 and coxae 2–3 each with posteroventral angle rounded; coxae 2–7 with entire oblique carina; pereonites 5–7 posterior margins smooth. Pleon with pleonite 1 almost concealed by pereonite 7; pleonites 2–5 posterior margin smooth; pleonite 2 posterolateral angles forming subacute point, not posteriorly produced; pleonite 3 posterolateral margins acute and not extending to pleonite 5 posterior margin; pleonite 4 posterolateral margin rounded extended beyond pleonite 5 posterior margin; pleonite 5 with posterolateral angles overlapped by pleonite 4 lateral margins. Pleotelson shorter than anterior width (0.87 times), without longitudinal carina, without dense patch of setae; lateral margins convex, smooth; posterior margin broadly rounded, with plumose setae. Antenna 1 peduncle articles 1 and 2 distinct and articulated; article 2 0.80 times as long as article 1; article 3 and 4 0.60 times as long as articles 1 and 2 length combined; article 3 almost as long as wide; flagellum moderately widened medially, with 12 articles, extending beyond to eye posterior margin. Antenna 2 peduncle article 4 1.5 times as long as wide, almost as long as article 3, with 3 short simple setae (distal); article 5 1.2 times as long as article 4, 2.4 times as long as wide, anterodistal angle with cluster of 3 short simple seta; flagellum with 22 articles, extending to pereonite 2. Frontal lamina long and linguiform, ventral surface entirely flat, 1.5 times longer than greatest width, lateral margins slightly convex, anterior margin rounded, visible in dorsal and reaching antenna 1 article 2. Mandible molar process anterior margin with 20 flat teeth; with proximal cluster of long simple setae; right mandible spine row composed of 8 spines; palp article 2 with 20 distolateral setae, article 3 with 13 slender setae. Maxillula mesial lobe with 3 large CPS RS and 1 short large CPS RS; lateral lobe with 11 RS. Maxilla lateral lobe with 5 long simple setae; middle lobe with 11 long simple setae; mesial lobe with 11 distal pappose setae. Maxilliped palp article 1 mesial margin without setae, lateral margin distally with 5 long CPS; article 2 mesial margin with 11 slender setae, lateral margin with 2 long and plumose setae; article 3 mesial margin with 14 slender setae, lateral margin with 6 long CPS and 2 slender setae; article 4 mesial margin with 14 slender setae and 4 pappose setae, lateral margin with 3 long CPS; article 5 distal margin to mesial margin with 15 slender setae and 6 pappose setae, lateral margin with 2 long and plumose setae; endite with 5 long CPS and 2 coupling setae. Pereopod 1 basis 2 times as long as greatest width, superior margin with 8 short plumose setae, superior distal angle with lacking cluster of setae, inferior margin with 6 long and slender setae, inferior distal angle with a cluster of 8 slender setae; ischium 0.8 times as long as basis, inferior margin with 9 simple setae and 2 RS, inferior distal margin without setae, superior margin with a middle and long CPS, superior distal margin produced over merus and with a cluster of long and slender setae, mesial margin with a cluster of long and slender setae; merus inferior margin with a row of 13 simple setae, superior distal angle produced and reaching 2/3 of carpus, superior margin with a row simple and RS; carpus inferior margin lacking setae, inferior distal angle with 2 RS; propodus 2.5 times as long as wide, inferior margin with 6 simple setae and 4 RS, inferior distal margin with 1 RS and 6 simple setae, superior distal margin with 4 short setae; dactylus 0.7 times as long as propodus. Pereopod 2 ischium inferior margin with 12 long simple setae, inferior distal margin without setae, superior distal margin produced over merus and with a cluster of long and slender setae, mesial margin with a cluster of long and slender setae; merus inferior margin with 2 stout RS and 6 acute RS, set as single row, inferior distal margin with 2 RS, superior distal angle produced and reaching 2/3 of carpus, superior margin with a row of both long and simple and 7 stout RS, mesial margin with a row of 4 stout RS and 13 simple and long setae; carpus inferior margin with 3 stout RS, inferior distal margin with 3 long acute RS and 4 simple seta; propodus 2.2 as long as wide, inferior margin with 2 long RS and 4 simple setae, inferior distal margin with 1 short RS and 3 simple setae; dactylus 0.8 times as long as propodus. Pereopod 3 similar to pereopod 2. Pereopods 4–6 similar to each other. Pereopod 7 both inferior and superior margins densely setose (plumose setae); basis slightly ovate, 2.2 times as long as greatest width; ischium 0.6 times as long as basis, 1.7 times as long as wide, widened distally; merus 0.5 times as long as ischium, 1.8 times as long as wide; carpus 0.5 times as long as ischium, 2.6 times as long as wide; propodus 0.5 times as long as ischium, 4.6 times as long as wide, inferior margin with 1 RS; dactylus 0.7 times as long as propodus. Penes distinct flat rounded lobes, separated by 15% of sternal width. Pleopod 1 exopod 1.3 times as long as wide, lateral margin slightly convex, distally broadly rounded with strongly oblique medial margin, mesial margin strongly convex, with PMS from distal two-third, with ~90 PMS; endopod 1.4 times as long as wide, distally narrowly rounded, with PMS from distal one-third, mesial margin with PMS from distal one-third, endopod with ~20 PMS; peduncle 0.45 times as wide as long, mesial margin with 7 coupling setae. Pleopod 2 exopod with ~90 PMS, endopod with ~21 PMS; appendix masculina inserted medially, with parallel margins, as long as endopod, distally subacute. Pleopod 3 exopod with ~100 PMS, endopod without PMS. Pleopod 4 exopod with ~75 PMS, endopod without PMS. Pleopod 5 exopod biarticulate, with ~55 PMS from distal two-third, endopod without PMS. Pleopods 2–5 endopod distal margin subacute. Uropod peduncle ventrolateral margin with 1 long RS, 1 short RS and 2 PMS, lateral margin with medial 4 short acute RS and 2 PMS, posterior lobe 0.80 times as long as endopod; rami extending beyond pleotelson, lateral, distal and mesial margin lined with long PMS. Endopod apically truncate; lateral margin straight, without prominent excision; mesial margin straight. Exopod not extending to end of endopod, 3.3 times as long as greatest width, apically rounded; lateral and mesial margin straight. Distribution. Bahia, Rio de Janeiro and S„o Paulo (Brazil), 250–508 m depth (Fig. 7). Variation. Two smaller individuals, mancae with 7 mm (MNRJ 20207) and juvenile with 8 mm (MZUSP 25641) have pleonite 1 not covered by pereonite 7, while in adult male holotype (15 mm, MNRJ 20208) the pereonite 7 hidden almost completely the pleonite 1. This character was not observed in other species of Dolicholana, which may be due to differing ontogenetic development of the species. Remarks. Dolicholana ivanyae spec. nov. is similar to D. brucei, recently described from Brazil (the Potiguar Basin off Rio Grande do Norte), and D. elongata, from the Pacific Ocean, in having a long frontal lamina that is visible in the dorsal view between the antennae, and the broadly rounded distal margin of the pleotelson. Dolicholana ivanyae spec. nov. can nevertheless be differentiated from D. brucei due to the frontal lamina surpassing article 2 of the peduncle of antenna 1 (vs. slightly surpassing peduncle 1) and visible dorsally between the antennae (vs. not visible dorsally), eyes separated by about 50% of the width of the head (vs. 18%), head with truncated anterior margin (vs. rounded), dorsal groove present (vs. absent), pereopod 7, densely setose, anterior and posterior margins of the basi-ischium and merus with very long plumose setae (vs. short plumose setae), uniarticulate endopods of pleopods 3 and 4 (vs. biarticulate), and distal margin pleotelson with no RS (vs. distal margin with two RS). Dolicholana ivanyae spec. nov. differs from D. elongata based on the following combination of characters: head with truncated (vs. slightly concave) anterior margin, dorsal groove present (vs. absent), frontal lamina very elongated surpassing article 2 of the peduncle of antenna 1 (vs. slightly surpassing article 1), pereopod 7 basis anterior and posterior margins densely setose with very long setae (vs. short setae), endopods of pleopods 3 and 4 uniarticulated (vs. biarticulate), pleotelson with distal margin lacking RS (vs. two RS), epimera 3 and 4 with a subacute (vs. rounded) posteroventral angle. A comparison of selected morphological parameters for all the species of the genus Dolicholana, including the new species, is provided in Table 1.

Natatolana hadassae spec. nov. (Figs 8–12) Material examined. Holotype: female, 30.1 mm (dissected and drawn, dissected appendages in alcohol 75%), Potiguar Basin, Rio Grande do Norte, MT 61 station, bottom fish net, 04°48’S, 036°9’W, 400 m, 08 May 2011, MOUFPE 15647. Type locality. Brazil, 04°48’S, 036°9’W. Etymology. Named after Hadassa Vieira de Carvalho Paiva, daughter of the first author. Diagnosis. Head interocular groove present, incomplete; submarginal furrow absent; dorsal groove absent. Eye unpigmented. Antenna 1 not reaching pereonite 1. Antenna 2 reaching pereonite 3. Pleopod 3–5 exopod with incomplete suture. Pleotelson broader than long, smooth, apex truncated with a subacute medial process, distal margin with 10 RS. Uropod rami lanceolate, apically acute. Description of female holotype. Body 2.8 times as long as greatest width, dorsal surfaces smooth, widest at pereonite 6, lateral margins slightly convex. Rostral point absent, frontal margin rounded. Eyes separated by about 80% head width, eye unpigmented. Pereonite 1 and coxae 2–3 each with posteroventral angle rounded; coxae 5–7 with entire oblique carina; posterior margins of pereonites 5–7 smooth. Pleon with pleonite 1 not concealed by pereonite 7, pleonites 2–5 posterior margin smooth; pleonite 2 posterolateral angle forming subacute point, not posteriorly produced; pleonite 3 posterolateral margins not extending to pleonite 5 posterior margin, acute; pleonite 4 posterolateral margin acute extended beyond pleonite 5 posterior margin; pleonite 5 with posterolateral angles overlapped by pleonite 4 lateral margins. Pleotelson shorter than anterior width (0.74 times); without longitudinal carina; without dense patch of setae; lateral margins convex, smooth; posterior margin broadly rounded with a middle subacute process, with plumose setae, and a row of 10 RS. Antenna 1 peduncle articles 1 and 2 distinct, articulated, article 2 0.3 times as long as article 1, article 3 and 4 0.8 times as long as articles 1 and 2 length combined, article 3 1.3 times as long as wide; flagellum moderately inflated, with 20 articles, extending beyond to eye halfway. Antenna 2 peduncle article 4 almost as long as wide, 1.45 times as long as article 3, with 6 short simple setae (distal); article 5 1.8 times as long as article 4, 2.8 times as long as wide, anterodistal angle with cluster of 3 short simple seta; flagellum with 31 articles, extending to pereonite 3. Frontal lamina 3.75 times longer than greatest width, lateral margins straight, slightly widening anteriorly, not visible in dorsal. Mandible molar process anterior margin with more than 20 flat teeth; without proximal cluster of long simple setae; right mandible spine row composed of 12 spines; palp article 2 with 17 distolateral setae, article 3 with 24 slender setae. Maxillula mesial lobe with 3 large CPS RS; lateral lobe with 10 RS. Maxilla lateral lobe with 2 long simple setae; middle lobe with 16 long simple setae; mesial lobe with 2 simple setae and 12 distal pappose setae. Maxilliped palp article 1 mesial margin without setae, lateral margin without setae; article 2 mesial margin with 6 slender setae, lateral margin with 9 simple setae; article 3 mesial margin with 10 slender setae, lateral margin with 19 slender setae; article 4 mesial margin with 9 slender setae, lateral margin with 12 simple setae; article 5 distal margin to mesial margin with 16 simple setae, lateral margin with 6 simple setae; endite with 5 long CPS and 2 coupling setae. Pereopod 1 basis 3 times as long as greatest width, superior margin setose, superior distal angle without cluster of setae, inferior margin without setae, inferior distal angle with a cluster of 4 long and slender setae; ischium 1.5 times as long as basis, inferior margin with 12 simple setae, inferior distal margin without setae, superior margin with a middle cluster of 6 simple setae, superior distal margin not produced over merus and with a line of long and slender setae; merus inferior margin with a row of 6 simple setae, superior distal angle produced and reaching 1/2 of carpus, superior margin with a row of both simple and RS; carpus inferior margin with 4 setae, inferior distal angle without setae; propodus 4.1 times as long as wide, inferior margin with 9 simple setae and 2 RS, inferior distal margin with 1 RS and 3 simple setae, superior distal margin with 4 short setae; dactylus 0.5 times as long as propodus. Pereopod 2 ischium inferior margin with 6 long simple setae, inferior distal margin with 6 simple setae, superior distal margin not produced over merus and with a cluster of long and slender setae, mesial margin with a cluster of long and slender setae; merus inferior margin with 1 stout RS, inferior distal margin with 2 RS, superior distal angle produced and reaching 1/4 of propodus, superior margin with a row of both long and simple; carpus inferior margin with 7 stout RS, inferior distal margin with 3 long acute RS; propodus 1.45 times as long as wide, inferior margin without setae, inferior distal margin with 1 short RS and 2 simple setae; dactylus 0.90 times as long as propodus. Pereopod 3 similar to pereopod 2. Pereopods 4–6 similar to each other. Pereopod 7 basis inferior distal angle with a 7 long and plumose setae; superior margin and facial middle line densely setose (plumose setae); basis slightly ovate, 2 times as long as greatest width; ischium 0.5 times as long as basis, 1.5 times as long as wide, widened distally; merus 0.60 times as long as ischium, 1.4 times as long as wide; carpus 0.80 times as long as ischium, 2.4 times as long as wide; propodus 1.2 times as long as ischium, 6 times as long as wide, inferior margin with 4 RS; dactylus 0.45 times as long as propodus. Pleopod 1 exopod 1.7 times as long as wide, lateral margin convex, distally broadly rounded with distolateral notch, with PMS from entire lateral to distal one-third mesial margin, with ~60 PMS; endopod 1.5 times as long as wide, distally narrowly rounded, with PMS from distal one-third, mesial margin with PMS from distal one-third, endopod with ~40 PMS; peduncle 0.4 times as wide as long, mesial margin with 5 coupling setae. Pleopod 2 exopod with ~90 PMS, endopod with ~30 PMS. Pleopod 3 exopod with incomplete suture, with ~100 PMS, endopod with ~20 PMS. Pleopod 4 exopod with incomplete suture, with ~80 PMS, endopod with ~20 PMS. Pleopod 5 exopod with incomplete suture, with ~35 PMS from distal two-third, endopod with 7 PMS distally. Pleopods 2–5 endopod distal margin with rounded. Uropod peduncle ventrolateral margin with 7 PMS, posterior lobe 0.90 times as long as endopod; rami slightly shorter than pleotelson, lateral, distal and mesial margin lined with long PMS. Endopod apically acute; lateral margin convex, without prominent excision; mesial margin almost straight. Exopod not extending beyond endopod, 4.4 times as long as greatest width, apically acute; lateral and mesial margin straight, lateral margin with 6 RS. Distribution. Known only from the type locality, Potiguar Basin, Rio Grande do Norte (Brazil), at 400 m depth (Fig. 7). Remarks. Considering the Natatolana species known from the Atlantic Ocean and Mediterranean Sea, Natatolana hadassae spec. nov. is most similar to Natatolana caeca (Dollfus, 1903), Natatolana gracilis Hansen, 1890, and Natatolana imicola (Dollfus, 1903) due to its unpigmented eyes. However, the new species differs from N. caeca by the incomplete (vs. complete) interocular furrow, antenna 2 reaching the middle of pereonite 3 (vs. reaching the posterior margin of pereonite 2), oblique groove present (vs. absent) in all the coxae, posterolateral margin of pleonite 4 subacute (vs. rounded), ischium of pereopod 3 as produced (vs. less produced) as those of pereopods 1–2, and distal margin of the pleotelson with 10 distal RS (vs. 8 RS). Natatolana hadassae spec. nov. can be distinguished from N. gracilis, which has also been also recorded from Brazilian waters, by the following combination of characters: ventrolateral margin of the head with a distinct depression (vs. no depression), posterolateral margin of pleonite 3 subacute (vs. rounded), frontal lamina 3.5 times longer than wide, with subparallel lateral margins widening to a rounded apex (vs. frontal lamina 3.1 times longer than wide, with concave lateral margins narrowing to a subacute apex), pereopods 1–7 more setose than in N. gracilis, especially the basis and ischium, pereopod 7 with well-marked medial carina (vs. poorly-defined medial carina), and uropod of the exopod with subacute (vs. rounded) apex. Natatolana hadassae spec. nov. can be distinguished readily from N. imicola by the fact that pleonite 1 is not concealed by pereonite 7 (vs. almost concealed by pereonite 7), posteroventral angle of pleonite 4 and coxa 7 acute (vs. slightly rounded), and exopods of pleopods 3–5 with poorly developed (vs. strongly developed) sutures.

Genus Paromola Wood-Mason in Wood-Mason & Alcock, 1891 Type species. Dorippe cuvieri Risso, 1815, by original designation., Published as part of Nyborg, Torrey, Garassino, Alessandro & Vega, Francisco J., 2023, Validation of Paromola roseburgensis Nyborg & Garassino (Brachyura: Homolidae) from the early Eocene of Oregon, USA, pp. 441-442 in Zootaxa 5318 (3) on page 441, DOI: 10.11646/zootaxa.5318.3.11, http://zenodo.org/record/8166786, {"references":["Wood-Mason, J. & Alcock, A. (1891) A note on the result of the last season's deep-sea dredging: natural history notes from H. M. Indian Marine Survey Steamer \" Investigator \", Commander, R. F. Hoskyn, R. N. Commanding, no. 21. Annals and Magazine of Natural History, Series 6, 7 (39), 258 - 272. https: // doi. org / 10.1080 / 00222939109460605"]}

Nyborg, Torrey, Garassino, Alessandro, Vega, Francisco J. (2023): Validation of Paromola roseburgensis Nyborg & Garassino (Brachyura: Homolidae) from the early Eocene of Oregon, USA. Zootaxa 5318 (3): 441-442, DOI: 10.11646/zootaxa.5318.3.11, URL: http://dx.doi.org/10.11646/zootaxa.5318.3.11

Tanidromites maerteni sp. nov. urn:lsid:zoobank.org:act: 6AF2E747-B3FE-4417-A4D9-3B94B2A5693A Tanidromites maerteni Fraaije, Van Bakel, Guinot & Jagt, 2013: 250, fig. 1 [unavailable]. Type material. The holotype, MNHN.F.A47612 (leg. L. Maerten), is deposited in the collections of the Muséum National d’Histoire Naturelle, Département Histoire de la Terre (Paris). The type specimen originates from the town quarry of the Commune de Maizet, Calvados (northwest France) from a level which is locally referred to as the “Oolithes ferrugineuses de Bayeux”, the age of which is early late Bajocian (Niortense ammonite Zone) (Gauthier et al. 1996). Etymology. In honour of Lionel Maerten (Ver-sur-Mer, France), who collected the holotype and kindly donated it to the Muséum National d’Histoire Naturelle collections. Diagnosis. A tanidromitid with lateral epibranchial spine; slightly concave portion of cervical groove at central posterior margin of mesogastric region; two faint, yet clearly visible, V-shaped grooves on central part of mesogastric region; relatively deep and long postcervical grooves; pustulate ornament on cardiac, epibranchial and epigastric regions (in accordance with Fraaije et al. 2013: 252). Remarks. A full description and figures of Tanidromites maerteni sp. nov. are supplied by Fraaije et al. (2013).

Paromola roseburgensis sp. nov. (Fig. 1) Paromola roseburgensis Nyborg & Garassino, 2017: 140–142, figs. 3, 4 [unavailable]. Type material. HOLOTYPE: UWBM 104435, carapace preserving dorsal surface, front and one left pereiopod, carapace length 24.2 mm, width 23.0 mm, Early Eocene, Roseburg Formation, Coos County, Oregon, USA. Diagnosis. Carapace widest at level of mesobranchial region, narrowing in anterior one-third; dorsal regions welldefined, marked by shallow incised grooves; rostrum long, pointed, moderately upturned; pseudorostral spines, slightly longer than rostrum, ornamented with small median external spinule; anterolateral margin with laterally-projecting spine; posterolateral margin strongly convex, rimmed by three laterally-projecting spines; posterior margin consisting of three arcs, one axially convex and two laterally concave; strongly spinose dorsal carapace. Etymology. The species name alludes to the Roseburg Formation where the studied specimen was collected. Remarks. A full description and figures of Paromola roseburgensis and stratigraphic account are given by Nyborg & Garassino (2017)., Published as part of Nyborg, Torrey, Garassino, Alessandro & Vega, Francisco J., 2023, Validation of Paromola roseburgensis Nyborg & Garassino (Brachyura: Homolidae) from the early Eocene of Oregon, USA, pp. 441-442 in Zootaxa 5318 (3) on page 441, DOI: 10.11646/zootaxa.5318.3.11, http://zenodo.org/record/8166786, {"references":["Nyborg, T. & Garassino, A. (2017) New Occurrences of Fossil Homolidae from the Eastern Pacific. Boletin de la Sociedad Geologica Mexicana, 69 (1), 135 - 148. https: // doi. org / 10.18268 / bsgm 2017 v 69 n 1 a 6"]}

Albaidaplax ispalensis sp. nov. Goneplax gulderi. — Mayoral et al. 1998: 508, 509, fig. 2(5). Goneplax cf. saccoi [sic].— Vía 1948: 146, fig. 6.— Solé & Vía 1989: 34. Albaidaplax ispalensis Garassino, Pasini & Castro, 2013: 362–363, fig. 4 [unavailable]. Type material. HOLOTYPE: MGUS AA5, Albaida del Aljarafe (Guadalquivir Basin, Seville, Andalusia, Spain, Early Pliocene). PARATYPES: MGUS AA2, AA3, AA4, AA6, AA45, same locality as holotype. Diagnosis. Carapace transversely rectangular, slightly wider than long; straight front slightly turned downwards; front as wide as the orbits; inner edge of supraorbital margin distinct; wide orbits slightly expanded distally with one short tooth on outer orbital angle; supraorbital margins gently sinuous, without fissures; sinuous, spineless suborbital margins; anterolateral margins slightly convex, with one short tooth; long, gently rounded posterolateral margins, slightly converging posteriorly; long, straight posterior margin; deep gastric pits; smooth dorsal surface of carapace; slight horizontal ridges, moderately convex, without clear indication of regions. Chelipeds (pereopod 1) stout, heavy, surfaces smooth; merus moderately long; carpus subtriangular, spineless; palm stout, globular; dactylus and pollex slender, as long as the propodus, with toothless occlusal margin. Ambulatory legs (pereopods 2–5) conspicuously long, slender, smooth. Etymology. From Ispal, name of an ancient village nearby Sevilla, city located close to the quarry where the specimens were discovered. Remarks. A full description and figures of Albaidaplax ispalensis and the stratigraphic account are given in Garassino et al. (2013)., Published as part of Garassino, Alessandro, Pasini, Giovanni & Castro, Peter, 2023, Validation of Albaidaplax ispalensis Garassino, Pasini & Castro, a fossil goneplacid crab from Spain and Italy (Crustacea: Decapoda: Goneplacidae), pp. 297-298 in Zootaxa 5318 (2) on pages 297-298, DOI: 10.11646/zootaxa.5318.2.12, http://zenodo.org/record/8162352, {"references":["Mayoral, E., Muller, P. & Muniz, F. (1998) Lower Pliocene decapod crustaceans from the southwestern Iberian Peninsula (Guadalquivir Basin, Sevilla, Spain). Geobios, 31 (4), 505 - 510. https: // doi. org / 10.1016 / S 0016 - 6995 (98) 80122 - 8","Via, L. (1948) Braquiuros fosiles de Barcelona y sus alrededores. Anales de la Escuela de Peritos Agricolas y de Servicios Tecnicos de Agricultura, 7, 143 - 152.","Sole, J. & Via, L. (1989) Crustacis Decapodes fossils dels Paisos Catalans. Batalleria, 2, 23 - 42.","Garassino, A., Pasini, G. & Castro, P. (2013) Revision of the fossil species of Goneplax Leach, 1814 (Crustacea, Decapoda, Brachyura, Goneplacidae). Boletin de la Sociedad Geologica Mexicana, 65 (2), 355 - 368. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 16"]}

Bahiacaris n. gen. http://zoobank.org/ urn:lsid:zoobank.org:act: 0B9F091B-F98E-49A7-B794-E250DB722900 Bahiacaris Schweitzer, Santana, Pinheiro & Feldmann, 2019: 72 [unavailable]. Diagnosis. Carapace high and short; rostrum about 30% total length of carapace, minutely serrate on upper margin. Scaphocerite not extending beyond rostrum. Pleonal somite 2 overlapping somites 1 and 3, variable in size. Telson sharp in lateral view, with apparent square tip in dorsal view; tip with setae; with 2 pairs of movable spines dorsally. Uropodal endopod and exopod narrow, much longer than wide; exopod inner margin weakly serrate, outer margin with setal pits. Third maxillipeds shorter than pereiopods 3–5. Pereiopods 1 and 2 shorter and with more bulbous articles than pereiopods 3–4. Type species. Atyoida roxoi Beurlen, 1950, by present designation. Type age. Lower Cretaceous (Aptian). Etymology. The genus name is derived from the state of Bahia in Brazil, where the specimen was collected, and ‘caris’, from the Greek for crab, a common stem in the group. Gender: feminine. Remarks. A full description and stratigraphic account of Bahiacaris n. gen. is given by Schweitzer et al. (2019)., Published as part of Schweitzer, Carrie E., Santana, William, Pinheiro, Allysson & Feldmann, Rodney M., 2023, Validation of Bahiacaris Schweitzer, Santana, Pinheiro & Feldmann (Crustacea, Decapoda, Caridea) from the Cretaceous (Aptian) of Brazil, pp. 299-300 in Zootaxa 5318 (2) on page 299, DOI: 10.11646/zootaxa.5318.2.13, http://zenodo.org/record/8162451, {"references":["Schweitzer, C. E., Santana, W., Pinheiro, A. & Feldmann, R. M. (2019) Redescription and illustration of caridean shrimp from the Cretaceous (Aptian) of Brazil. Journal of South American Earth Sciences, 90, 70 - 75. https: // doi. org / 10.1016 / j. jsames. 2018.12.001","Beurlen, K. (1950) Alguns restos de Crustaceos Decapodes d'agua doce fosseis no Brasil. Anais da Academia Brasiliera de Ciencias, 22 (4), 453 - 459."]}

Praedatrypaea Poore, Dworschak, Robles, Mantelatto & Felder, 2019 Remarks. Hyžný et al. (2013) considered Callianassa ocozocoautlaensis as morphologically close to extant Callianassa modesta De Man, 1905. Recently, the latter species was together with three more species placed into a distinct genus, Praedatrypaea Poore, Dworschak, Robles, Mantelatto & Felder, 2019. Thus, the fossil species in question is placed within the genus Praedatrypaea, being the only known representative of the genus known from the fossil record., Published as part of Hyžný, Matúš, Vega, Francisco J. & Coutiño, Marco A., 2023, Validation of Callianassa ocozocoautlaensis Hyžný, Vega & Coutiño, a fossil ghost shrimp (Malacostraca: Decapoda: Axiidea) from the Upper Cretaceous of Chiapas, Mexico, pp. 295-296 in Zootaxa 5318 (2) on page 295, DOI: 10.11646/zootaxa.5318.2.11, http://zenodo.org/record/8162338, {"references":["Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. L. & Felder, D. L. (2019) A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05","Hyzny, M., Vega, F. J. & Coutino, M. A. (2013) Ghost shrimps (Decapoda: Axiidea: Callianassidae) of the Maastrichtian (Late Cretaceous) Ocozocoautla Formation, Chiapas (Mexico). Boletin de la Sociedad Geologica Mexicana, 65, 255 - 264. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 7","De Man, J. G. (1905) Diagnoses of new species of macrurous decapod Crustacea from the \" Siboga-Expedition \". Tijdschrift der Nederlandsche Dierkundige Vereeniging, 9, 587 - 614."]}

Hyžný, Matúš, Vega, Francisco J., Coutiño, Marco A. (2023): Validation of Callianassa ocozocoautlaensis Hyžný, Vega & Coutiño, a fossil ghost shrimp (Malacostraca: Decapoda: Axiidea) from the Upper Cretaceous of Chiapas, Mexico. Zootaxa 5318 (2): 295-296, DOI: 10.11646/zootaxa.5318.2.11, URL: http://dx.doi.org/10.11646/zootaxa.5318.2.11

Praedatrypaea ocozocoautlaensis sp. nov. (Fig. 1A–1D) urn:lsid:zoobank.org:act: D461B591-EAB8-4FFE-91C0-5E59E73E4F4D Callianassa (s.l.) ocozocoautlaensis Hyžný, Vega & Coutiño, 2013: 259, figs. 3.1–3.3. [Unavailable] Holotype. The holotype is deposited in the Museo de Paleontología "Eliseo Palacios Aguilera", Secretaría de Medio Ambiente e Historia Natural/ Dirección de Paleontología, Calzada de los Hombres Ilustres s/n, Colonia Antiguo Parque Madero, Tuxtla Gutiérrez, 29000, Chiapas, México, under the repository number IHNFG-4664. Diagnosis. Callianassid with chelipeds of unequal size and dissimilar shape. Major cheliped manus upper margin arcuate, lower margin straight, fixed finger shorter than manus, lateral surface with longitudinal depression, fingers of approximately the same length, occlusal margins armed. Minor cheliped carpus subtriangular, propodus long and slender, both fingers slender, unarmed and pointed. Etymology. After the Ocozocoautla Formation. Occurrence. Lower Maastrichtian of the Ocozocoautla Formation, Chiapas (Mexico). Remarks. A full description and figures of the species are given by Hyžný et al. (2013)., Published as part of Hyžný, Matúš, Vega, Francisco J. & Coutiño, Marco A., 2023, Validation of Callianassa ocozocoautlaensis Hyžný, Vega & Coutiño, a fossil ghost shrimp (Malacostraca: Decapoda: Axiidea) from the Upper Cretaceous of Chiapas, Mexico, pp. 295-296 in Zootaxa 5318 (2) on pages 295-296, DOI: 10.11646/zootaxa.5318.2.11, http://zenodo.org/record/8162338, {"references":["Hyzny, M., Vega, F. J. & Coutino, M. A. (2013) Ghost shrimps (Decapoda: Axiidea: Callianassidae) of the Maastrichtian (Late Cretaceous) Ocozocoautla Formation, Chiapas (Mexico). Boletin de la Sociedad Geologica Mexicana, 65, 255 - 264. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 7"]}

Garassino, Alessandro, Pasini, Giovanni, Castro, Peter (2023): Validation of Albaidaplax ispalensis Garassino, Pasini & Castro, a fossil goneplacid crab from Spain and Italy (Crustacea: Decapoda: Goneplacidae). Zootaxa 5318 (2): 297-298, DOI: 10.11646/zootaxa.5318.2.12, URL: http://dx.doi.org/10.11646/zootaxa.5318.2.12

Clistocoeloma villosum (A. Milne-Edwards, 1869) (Figs. 1A–C, 5A, B, 6D, 9A–D) Sesarma villosum A. Milne-Edwards, 1869: 31 [type locality: Upolu, Samoa]. Sesarma villosa — De Man 1887: 644, 645. Sesarma (Sesarma) villosa — De Man 1897: 153–161 (in part).— De Man 1898: 702 (list) (in part), pl. 29 fig. 30b–d, 30dd. Sesarma (Holometopus) villosa — Tesch 1917: 208–210 (in part). Sesarma (Holometopus) villosum — Rathbun 1907: 35.— Crosnier 1965: 56 (in part).—Serène 1968: 107 (list)— Rahayu & Takeda 2000: 40. Clistocoeloma suvaense — Edmondson 1951: 238, 242 (in part) [not Clistocoeloma suvaense Edmondson, 1951]. Sesarma villosum — Ng & Liu 1999: 229. Clistocoeloma villosum — Davie 2002: 221 (in part).— Ng et al. 2008: 220 (list).— Lee et al. 2013: 420.— Li et al. 2019: 1211 (in part), 1214 (in part). Type material. Lectotype: male (18.6 × 17.0 mm) (ZMH K4188), Upolu, Samoa, no other data. Diagnosis. Carapace subquadrate, nearly as wide as long; postfrontal lobes distinct, divided by longitudinal grooves, median pair broad, prominent, distal pair less distinct, not divided (Fig. 1A). Inner orbital angle distinct, rounded, with convex outer margin, gap absent between tip of inner orbital angle and frontal margin (Fig. 1C). Anterolateral margin of carapace straight, smooth, external orbital angle less pronounced; posterolateral margin slightly convex. Chelipeds dorsal surface of dactylus of chela with 16 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Fig. 5A), each tubercle on dorsal surface of movable dactylus with small median chitinous granule (Fig. 5A). Single longitudinal pectinated ridge with 27–30 chitinous comb-like tubercles on dorsal surface of chela, distal end of ridge with granules; row of 8 granules behind pectinated ridge on ventral surface of palm (Fig. 5A); inner surface of palm granulated, with single vertical row of 8 granules in adult males (Fig. 5B). P5 merus relatively short, wide, merus length approximately 0.44 times width (Fig. 6D). G1 slightly curved, distal region slightly wide; chitinous tip relatively wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (Fig. 9A–D). Description. Carapace subquadrate, nearly as wide as long; covered with coat of short setae, among which stands small tufts of setae resembling tubercles; carapace smooth when denuded. Postfrontal lobes distinct, divided by longitudinal grooves, median pair broad, prominent, distal pair less distinct, not divided (Fig. 1A). Inner orbital angle distinct, rounded, with convex outer margin, no gap present between tip of inner orbital angle and frontal margin (Fig. 1C). Anterolateral margin of carapace straight, smooth, external orbital angle less pronounced; posterolateral margin slightly convex. Entire margin of carapace covered with dense, short setae. Antenna short. Antennular fossa transverse. Third maxilliped with merus rounded. Epistomal margin narrow (Fig. 1C). Chelipeds equal or subequal in size; large, robust in adult male; outer surface of palm smooth when denuded; dorsal surface of carpus covered with small tufts of setae resembling tubercles, smooth when denuded; dorsal surface of dactylus of chela with 16 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Fig. 5A), each tubercle on dorsal surface of movable dactylus with small median chitinous granule (Fig. 5A); single longitudinal pectinated ridge with 27–30 chitinous comb-like tubercles on dorsal surface of chela, distal end of ridge with granules; row of 8 granules behind pectinated ridge on ventral surface of palm (Fig. 5A); inner surface of palm granulated, with single vertical row of 8 granules in adult males (Fig. 5B). P2–P5 covered with dense coat of setae, with small tufts of setae resembling tubercles, longer setae on dorsal and ventral margins of entire leg except dactylus; dactylus shortest, with sharp tip (Fig. 1A); P5 merus relatively short, wide, merus length approximately 0.44 times of width (Fig. 6D). Male thoracic sternum covered with short setae, smooth when denuded; sternites 1 and 2 fused, with suture between fused sternites 3 and 4; sternopleonal cavity extends to edge of sternites 1 and 2. Male pleon short, wide, long setae fringed margins of pleon; somite 3 widest; telson slightly broad, triangular with convex margins, rounded tip (Fig. 1B); male pleonal locking mechanism absent, without tubercle on sternite 5. G1 slightly curved, distal region slightly wide; chitinous tip relatively wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (Fig. 9A–D). G2 shorter than G1, slightly curved, tip rounded. Female characters not known. Remarks. The original description by A. Milne-Edwards (1869) was inadequate by modern standards and no figure was provided. He did not indicate how many specimens he had but the measurements for one specimen from Upolu was listed; no type was designated. Subsequently, De Man (1879) mentioned that the male specimen from the “Museum zu Hamburg ” was send to him by Dr. Pfeffer to examine, and that it is the “Original-Exemplar” of A. Milne-Edwards from Upolu. De Man (1898: pl. 29, fig. 30b–d) subsequently figured the chela of this male specimen. Crosnier (1965: 56) incorrectly assumed that the type was in Paris when he commented that “le type S. villosum, qui devrait se trouver au Muséum national d’Histoire naturelle de Paris, paraît avoir disparu”. The present type specimen matches the descriptions, measurements and figures provided by A. Milne-Edwards (1869) and De Man (1879, 1898) very well and we are confident that it is the type of S. villosum A. Milne-Edwards, 1869. As we are not sure if A. Milne-Edwards (1869) had only one specimen, the specimen we have on hand is considered syntypic and is here designated the lectotype. One of the old labels with the specimen states that the lectotype specimen was from the Godeffroy Museum. Based on the eight sales catalogues of Godeffroy Museum that were published between 1864 and 1881 by curator J.D.E. Schmeltz, the species was listed once in catalogue V in 1874 (see Bieler & Petit 2012). The Godeffroy Museum catalogue number of the specimen listed, Sesarma villosa, (see Schmeltz 1874: 76) matches the catalogue number on the label for the lectotype of C. villosum. Tesch (1917) noted that Sesarma (Holometopus) villosum might be a species of Clistocoeloma because of the pattern of setae described for the species, although he did not formally transfer it there. Serène (1968) listed the species under Sesarma (Holometopus) while indicating that there is a photographed specimen in the “Institute of Oceanography of Nhatrang, Vietnam ”. The actual locality and collection information for the photographed specimen from the collection in the Vietnamese institute is not known and its identity cannot be confirmed. Ng & Liu (1999) noted that S. villosum should be in the genus Clistocoeloma but did not provide any explanation. Sesarma villosum was then briefly discussed by Rahayu & Takeda (2000: 40) before Davie (2002: 221) formally transferred the species to Clistocoeloma, citing McNeill’s suggestion (1968: 79). Komai et al. (2004: 41, 42) briefly discussed the generic assignment of this species in Clistocoeloma, following Davie (2002: 220). This species has been widely recorded from the IWP, i.e., Sumatra (De Man 1897, 1898), Caroline Islands (Rathbun 1907), New Guinea (Tesch 1917), Philippines (McNeill 1968), Taiwan (Li et al. 2019), Japan (Komai et al. 2004; Yuhara et al. 2017; Takakura & Komai 2019), and South Korea (Lee et al. 2010), but comparison of the morphology of the specimens from these localities shows that these specimens are of a different species, and is here described as new (see Clistocoeloma nobile n. sp.) Records of C. villosum from Nosy Be, off Madagascar (Crosnier 1965; Lenz 1910), Tuléar, Madagascar (Crosnier 1965), and Kenya (Schubart et al. 2006) are shown to be a different species. Various specimens from that region have been examined and the species clearly differs from C. villosum, and is more similar in morphology to C. suvaense and C. nobile n. sp. The morphological differences are also supported by DNA sequence differences (unpublished data) and the new species will be described in a separate study by the first author together with S. Cannicci (pers. comm.). Ecological notes. Little is known about this species except that the specimens were collected from mangroves. Distribution. Known for certain only from Upolu, Samoa (type locality)., Published as part of Ng, Bee Yan Lee Ngan Kee & Ng, Peter K. L., 2023, On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata), pp. 217-236 in Zootaxa 5318 (2) on pages 218-220, DOI: 10.11646/zootaxa.5318.2.3, http://zenodo.org/record/8162486, {"references":["Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]","De Man, J. G. (1887) Uebersicht der indo-pacifischen Arten der Gattung Sesarma Say, nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschriebenen Decapoden. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 2 (3 - 4), 639 - 722, figs. 1 - 4, pl. 17.","De Man, J. G. (1897) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 9 (1), 75 - 218. [in German]","De Man, J. G. (1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Sechster (Schluss-) Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (4), 677 - 708, pls. 28 - 38.","Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640","Rathbun, M. J. (1907) Reports on the scientific results of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer \" Albatross, \" from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. IX. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer \" Albatross, \" from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. X: The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2), 25 - 74.","Crosnier, A. (1965) Crustaces decapodes. Grapsidae et Ocypodidae. Faune de Madagascar, 18, 1 - 143, figs. 1 - 260, pls. 1 - 11. [in French]","Rahayu, D. L. & Takeda, M. (2000) A new species of the genus Clistocoeloma (Crustacea: Decapoda: Grapsidae) from Irian Jaya, Indonesia. Bulletin of the National Science Museum, Tokyo, Series A, 26 (2), 35 - 41.","Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38.","Ng, P. K. L. & Liu, H. - C. (1999) The taxonomy of Sesarma tangi Rathbun, 1931, and S. stormi De Man, 1895 (Crustacea: Decapoda: Brachyura: Grapsidae: Sesarminae), with establishment of a new genus for S. stormi. Zoological Studies, 38 (2), 228 - 237.","Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda - Anomura, Brachyura. In: Wells, A. & Houston, W. W. K. (Eds.), Zoological Catalogues of Australia. Vol. 19.3 B. CSIRO Publishing, Melbourne, pp. i - xiv + 1 - 641.","Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286.","Lee, B. Y., Ng, N. K. & Ng, P. K. L. (2013) On the identity of Clistocoeloma balansae A. Milne-Edwards, 1873, and C. tectum (Rathbun, 1914), with description of a new species from the West Pacific (Crustacea: Decapoda: Sesarmidae). Zootaxa, 3641 (4), 420 - 432. https: // doi. org / 10.11646 / zootaxa. 3641.4.8","Li, J. - J., Hsu, J. - W., Ng, N. K. & Shih, H. - T. (2019) Eight new records of crabs (Decapoda, Brachyura: Sesarmidae, Varunidae) from the coasts of Taiwan. Crustaceana, 92 (10), 1207 - 1230. https: // doi. org / 10.1163 / 15685403 - 00003935","Bieler, R. & Petit, R. E. (2012) Molluscan taxa in the publications of the Museum Godeffroy of Hamburg, with a discussion of the Godeffroy Sales Catalogs (1864 - 1884), the Journal des Museum Godeffroy (1873 - 1910), and a history of the museum. Zootax a, 3511 (1), 1 - 80. https: // doi. org / 10.11646 / zootaxa. 3511.1.1","Schmeltz, J. D. E. (1874) Museum Godeffroy Catalog V. Nebst einer Beilage Enthaltend topographische and zoologische Notizen. L. Friederichsen & Co., Hamburg, xxxviii + 215 pp. [February, in German]","Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. & Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63.","McNeill, F. A. (1968) Crustacea, Decapoda and Stomatopoda. Great Barrier Expedition 1928 - 29, Scientific Reports, 7 (1), 1 - 98, pls. 1 + 2.","Yuhara, T., Yokooka, H. & Taru, M. (2017) Range extension of the sesarmid crab Clistocoeloma villosum along the eastern Pacific coast of the Izu Peninsula, Japan. Marine Biodiversity Records, 10, 19. https: // doi. org / 10.1186 / s 41200 - 017 - 0122 - 1","Takakura, H. & Komai, T. (2019) New records of a semiterrestrial sesarmid crab Clistocoeloma villosum (A. Milne-Edwards, 1869) (Crustacea: Decapoda: Brachyura: Grapsoidea) from Izu and Boso Peninsulas, Japan. Cancer, 28, 25 - 31. [in Japanese with English abstract]","Lee, S. - K., Jung, J. & Kim, W. (2010) A new report on sesarmid crab Clistocoeloma villosum (Crustacea: Decapoda: Brachyura) from Korea. Korean Journal of Systematic Zoology, 26 (2), 179 - 181. https: // doi. org / 10.5635 / KJSZ. 2010.26.2.179","Lenz, H. (1910) Crustaceen von Madagascar, Ostafrika und Ceylon. In: Voeltzkow, A. (Ed.), Reise in Ostafrika in den Jahren 1903 - 1905 mit Mitteln der Hermann und Elise geb. Heckmann Wentzel-Stiftung ausgefuhrt. Wissenschaftliche Ergebnisse. Systematische Arbeiten. Vol. 2. Schweizerbart, Nagele und Sproesser, Stuttgart, pp. 539 - 576, figs. 1 - 4 [in German]","Schubart, S. D., Cannicci, S., Vannini, M. & Fratini, S. (2006) Molecular phylogeny of grapsoid crabs (Decapoda, Brachyura) and allies based on two mitochondrial genes and a proposal for refraining from current superfamily classification. Journal of Zoological Systematics and Evolutionary Research, 44 (3), 193 - 199. https: // doi. org / 10.1111 / j. 1439 - 0469.2006.00354. x"]}

Albaidaplax gen. nov. Albaidaplax Garassino, Pasini & Castro, 2013: 362 [unavailable]. Type species. Albaidaplax ispalensis sp. nov. Diagnosis. As for the type species Etymology. From Albaida del Aljarafe, village nearby the quarry where the specimens were discovered. Gender feminine., Published as part of Garassino, Alessandro, Pasini, Giovanni & Castro, Peter, 2023, Validation of Albaidaplax ispalensis Garassino, Pasini & Castro, a fossil goneplacid crab from Spain and Italy (Crustacea: Decapoda: Goneplacidae), pp. 297-298 in Zootaxa 5318 (2) on page 297, DOI: 10.11646/zootaxa.5318.2.12, http://zenodo.org/record/8162352, {"references":["Garassino, A., Pasini, G. & Castro, P. (2013) Revision of the fossil species of Goneplax Leach, 1814 (Crustacea, Decapoda, Brachyura, Goneplacidae). Boletin de la Sociedad Geologica Mexicana, 65 (2), 355 - 368. https: // doi. org / 10.18268 / BSGM 2013 v 65 n 2 a 16"]}

Clistocoeloma suvaense Edmondson, 1951 (Figs. 2A, 3A–F, 4A–C, 5C, D, 6A–C, 8A, B, 9E–H, 10A–H) Clistocoeloma suvaense Edmondson, 1951: 238–241, figs. 37a–i, 38 [type locality: reef in Suva Habor, Fiji]. Clistocoeloma suvaense — Rahayu & Takeda 2000: 38 (list).— Komai et al. 2004: 41 (list).— Ng et al. 2008: 220 (list). Type material. Holotype: male (21.0 × 18.8 mm) (BPBM 3976), Suva, Fiji, no collection data. Other material examined: Fiji: 1 male (18.8 × 16.9 mm) (ZRC 2022.0985), river mouth, Navaca mangrove, Taveuni Island, Fiji, 16°05’19”S 179°05’48”E, coll. B.Y. Lee, S. Choy & B. Rashni, 26 July 2019. — 1 male (17.8 × 16.3 mm) (ZRC 2022.0986), mangrove at Suva Point Beach, near My Suva Picnic Park and Suva City Council Park, Fiji, 18°00’28”S 178°02’42”E, coll. B.Y. Lee & B. Rashni, 22 July 2019.— 1 male (12.1 × 10.9 mm) (ZRC 2022.0987), Bure Bure mangrove, Naqai Creek, near Naqai Bridge, Taveuni Island, Fiji, 16°04’28”S 179°05’16”W, coll. B.Y. Lee, S. Choy & B. Rashni, 25 July 2019.— 2 males (13.7 × 12.2 mm, 12.5 × 11.4 mm), 1 female (13.3 × 11.9 mm) (ZRC 2022.0988), river mouth, Navaca mangrove, Taveuni Island, Fiji, 16°05’19”S 179°05’48”E, coll. B.Y. Lee, S. Choy & B. Rashni, 26 July 2019.— 2 males (17.1 × 15.5 mm, 15.2 × 14.2 mm), 2 ovigerous females (16.6 × 15.0 mm, 16.2 × 14.5 mm), 1 female (18.7 × 17.0 mm) (ZRC 2022.0989), mangrove at Suva Point Beach, near My Suva Picnic Park and Suva City Council Park, Fiji, 18°00’28”S 178°02’42”E, coll. B.Y. Lee & B. Rashni, 22 July 2019. Samoa: 1 male (16.2 × 14.3 mm) (ZRC 1973.11.2.490) Upolu, coll. Godeffroy Museum. Diagnosis. Carapace slightly wider than long, almost square; postfrontal lobes distinct, divided by longitudinal grooves; median pair broad, prominent, nearly equal in proportion; distant pair broad, prominent, nearly equal in proportion (Figs. 2A, 3A, 4A). Anterolateral margin of carapace with distinct external orbital angle, blunt second tooth blunt separated by distinct shallow groove from external orbital tooth; posterolateral margin convex (Figs. 2A, 3A, 4A). Inner orbital angle present, truncate, slight gap between inner orbital angle and frontal margin (Figs. 3C, 4C). Chelipeds dorsal surface of dactylus of chela with 22–27 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Figs. 3E, 5C); absent or weak granules in female and juveniles; presence of single longitudinal pectinated ridge with 33–36 chitinous comb-like tubercles on dorsal surface of chela; row of 9–11 granules behind pectinated ridge on chela (Figs. 3E, 5C); inner surface of palm granulated, with single vertical row of 6–10 granules in adult males (Figs. 3F, 5D); absent in females and juveniles. P5 merus relatively short, wide, length approximately 0.39–0.46 times width (Fig. 6A–C). G1 straight, distal region relatively wide; chitinous tip wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (Figs. 9E–H, 10A–H). Female vulvae with central operculum, ovate, slightly protruded; sternal vulvar cover slightly raised above operculum (Fig. 8B). Description. Carapace slightly wider than long, almost square; covered with dense coat of setae, with small tufts of setae resembling tubercles; carapace margin fringed with dense, short setae; carapace smooth when denuded. Postfrontal lobes distinct, divided by longitudinal grooves; median pair broad, prominent, nearly equal in proportion; distant pair broad, prominent, nearly equal in proportion (Figs. 2A, 3A, 4A). Anterolateral margin of carapace with distinct external orbital angle, blunt second tooth blunt separated by distinct shallow groove from external orbital tooth; posterolateral margin convex (Figs. 2A, 3A, 4A). Inner orbital angle present, truncate, slight gap between inner orbital angle and frontal margin (Figs. 3C, 4C). Antenna short. Antennules folding transversely. Third maxilliped with merus relatively ovate. Epistomial margin relatively narrow (Figs. 3C, 4C). Chelipeds subequal in size; large, robust in male; outer surface of palm smooth when denuded; dorsal surface of carpus with small tufts of setae resembling tubercles, smooth when denuded; dorsal surface of dactylus of chela with 22–27 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Figs. 3E, 5C); absent or weak granules in female and juveniles. Presence of single longitudinal pectinated ridge with 33–36 chitinous comb-like tubercles on dorsal surface of chela; row of 9–11 granules behind pectinated ridge on chela (Figs. 3E, 5C). Inner surface of palm granulated, with single vertical row of 6–10 granules in adult males (Figs. 3F, 5D); absent in females and juveniles. P2–P5 with dense coat of setae, with small tufts of setae resembling tubercles, longer setae on dorsal and ventral margins except distal portion of dactylus (Figs. 2A, 4A); P5 merus relatively short, wide, length approximately 0.39–0.46 times width (Fig. 6A–C). Male thoracic sternum covered with short setae, smooth when denuded; sternites 1 and 2 fused, with suture between fused sternites 3 and 4; sternopleonal cavity extends to edge sternites 1 and 2. Male pleon wide, stout, long setae fringed pleon margin; somite 3 widest; somite 6 with curved edge; telson wide, dome-shaped (Figs. 3B, D, 4B); male pleonal locking mechanism absent, without tubercle on sternite 5. G1 straight, distal region relatively wide; chitinous tip wide, short; tip with dense tufts of setae, single row of plumose setae on exterior margin; chitinous tip visible when denuded (Figs. 9E–H, 10A–H). G2 shorter than G1, slightly curved, tip rounded. Female pleon rounded, relatively wider than long; telson dome-shaped (Fig. 8A). Vulvae with central operculum, ovate, slightly protruded; sternal vulvar cover slightly raised above operculum (Fig. 8B). Colouration. Chelipeds pale yellow fingers with pale orange to reddish palm (Fig. 2A). In life, entire crab covered in thick layer of mud. Remarks. Clistocoeloma suvaense was described by Edmondson (1951) based on one male and one female specimen collected from a reef in Suva Harbour. He compared it with other species that were recognised in Clistocoeloma at the time, i.e., C. balansae. C. merguiense and C. tectum (Rathbun, 1914). Clistoceoloma villosum was not treated or discussed as it was not in the genus at the time. Edmondson (1951) actually had three specimens from the Bishop Museum, two of which were collected from Suva Habour, Fiji, while one male specimen was collected from a Samoan reef. While Edmondson (1951) was certain that the two Fijian specimens belonged to his new species, he was more doubtful about the Samoan one; noting that the latter had no distinct lobes on the anterolateral margins (lobes present on the anterolateral margins of C. suvaense s. str.) and the dorsal surface of the dactylus of the male chela with 16 granules (with 25 granules in C. suvaense s. str.). We have not examined Edmondson’s (1951) Samoan specimens, but his observations match exactly what is known for C. villosum, we believe they are conspecific. Clistocoeloma suvaense is known with certainty only from its type locality, although there is one specimen also from Samoa (see below) and published records from Singapore (Wee & Ng 1994; Tan & Ng 1994). In addition to the lectotype specimen of C. villosum, another Wroblewsky specimen originally from the Godeffroy Museum was found in the ZRC, although how it got there is not known, possibly from the time R. Serène was based in Singapore in the 1960s. It is noteworthy that during that time, Serène was revising the IWP Sesarmidae with Soh Cheng Lan (e.g., see Serène & Soh 1970) and he had gathered material in Singapore for this work. The morphology of this Wroblewsky specimen, however, clearly matches what is here defined as C. suvaense and not C. villosum. Based on the specimen label, it has the same locality information as the lectotype of C. villosum. In addition to the very different morphology, the measurements of this specimen also do not match those listed in A. Milne-Edwards (1869) in his original description. It is unclear whether A. Milne-Edwards actually examined this specimen or if there is a mistake in the labels. There is also no mention of additional specimens of “ C. villosum ” in the eight sales catalogues of Godeffroy Museum that were published between 1864 and 1881 by curator J.D.E. Schmeltz other than that for the original specimen in A. Milne-Edwards (1896) or later papers by the author on the Godeffroy Museum collections (e.g. A. Mine-Edwards 1873a). For the moment, we accept the record of C. suvaense as being also present in Samoa. Even though both C suvaense and C. villosum are found together in Samoa, it is important to note that currently, no specimens of C. villosum has been found in Fiji. Recent collections there only found C. suvaense. Records of “ C. suvaense ” from Singapore by Wee & Ng (1994) and Tan & Ng (1994) belong to C. nobile n. sp. instead. Although C. suvaense is superficially similar to C. villosum and C. nobile n. sp., they are relatively easy to distinguish, especially adult males. Clistocoeloma suvaense clearly differs from C. villosum s. str. in the presence of a distinct external orbital angle on the anterolateral margin of the carapace (Figs. 2A, 3A, 4A) (the external orbital angle on the anterolateral carapace margin is not distinct and low in C. villosum; Fig. 1A); the presence of a broad, rounded inner orbital angle (Figs. 3C, 4C) (the inner orbital angle is narrow and ovate in C. villosum; Fig. 1C); possessing 25–26 granules on the dorsal surface of the dactylus of the adult male chela (Figs. 3E, 5D) (16 granules in C. villosum; Fig. 5A); and the G1 is straighter with a proportionately broader distal portion (Figs. 9E–H, 10A–H) (G1 slightly curved with a narrower distal portion for C. villosum; Fig. 9A–D). Fresh collections of C. suvaense from the type locality show that the species is not uncommon in Fiji, and all the recent specimens agree very well with the type material. Ecological notes. It was mentioned that the specimens were collected from “a reef in Suva Harbor in 1933” (Edmondson 1951: 240). Fresh material was collected from the back mangrove at a higher elevation and found on muddy substrate under large loose rocks in Suva and Taveuni Island, Fiji. Specimens were found with layer of mud covering the entire crab. Distribution. Currently only known for certain from the type locality, Suva, as well as Taveuni Island, Fiji; the record from Samoa is uncertain., Published as part of Ng, Bee Yan Lee Ngan Kee & Ng, Peter K. L., 2023, On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata), pp. 217-236 in Zootaxa 5318 (2) on pages 221-224, DOI: 10.11646/zootaxa.5318.2.3, http://zenodo.org/record/8162486, {"references":["Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38.","Rahayu, D. L. & Takeda, M. (2000) A new species of the genus Clistocoeloma (Crustacea: Decapoda: Grapsidae) from Irian Jaya, Indonesia. Bulletin of the National Science Museum, Tokyo, Series A, 26 (2), 35 - 41.","Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. & Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63.","Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286.","Rathbun, M. J. (1914) New species of crabs of the families Grapsidae and Ocypodida. Proceedings of the United Stated National Museum, 47 (2044), 69 - 85. https: // doi. org / 10.5479 / si. 00963801.2044.69","Wee, Y. C. & Ng, P. K. L. (Eds.) (1994) A first look at biodiversity in Singapore. National council on the Environment, Singapore, 163 pp.","Tan, C. G. S. & Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655","Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]"]}

Clistocoeloma nobile n. sp. (Figs, 2B, C, 5E, F, 6E–K, 7A–C, 8C, D, 11A–D) Sesarma (Sesarma) villosa — De Man 1897: 153–161 (in part).— De Man 1898: 702 (list) (in part), pl. 29 fig. 30a, 30e [not Sesarma villosum A. Milne-Edwards, 1869]. Sesarma (Holometopus) villosum — McNeill 1968: 79 (in part).— Rathbun 1907: 35. Sesarma (Holometopus) villosa — Tesch 1917: 208–210 (in part), 231 (list), 237 (key), 239 (footnote), 260, pl. 17 fig. 2 [not Sesarma villosum A. Milne-Edwards, 1869]. Chiromantes villosum — Nomoto et al. 1999: 9, pl. 1 fig. 6.— Kishino et al. 2001a: 17–18, pl. 2 fig. 2.— Kishino et al. 2001b: 127 (list), table 2.— Shokita et al. 2002: 78 (list), 85, photo 4A–1 fig. A [not Sesarma villosum A. Milne-Edwards, 1869]. Cleistocoeloma (sic) suvaense — Tan & Ng 1994: 82 (list) [not Clistocoeloma suvaense Edmondson, 1951]. Clistocoeloma suvaensis —Wee & Ng 1996: 82 (list) [not Clistocoeloma suvaense Edmondson, 1951]. Clistocoeloma villosum — Davie 2002: 221 (in part).— Komai et al. 2004: 38, 40–42, fig. 1B, 3.— Karasawa et al. 2006: 60, 61, fig. 1. — Schubart et al. 2009: 3 (list), fig. 18 (in part).— Lee et al. 2010: 179–181, fig. 1 A–H.—Maenosono & Sakai 2016: 6, fig. 2D.— Yuhara et al. 2017: [1–4], fig. 2.— Yuhara & Yokooka 2019: 8–11, figs. 2C, 3.— Li et al. 2019: 1208 (list), 1211–1214, figs. 1B, 3. – Takakura & Komai 2019: 25–30, figs. 1–5 [not Sesarma villosum A. Milne-Edwards, 1869]. Clistocoeloma cf. suvaense — Li 2015: fig. 15 [not Clistocoeloma suvaense Edmondson, 1951]. Type material. Holotype: male (15.3 × 12.7 mm) (ZRC 2022.0970), stn SW7 (SS-00221), pontoon at south lagoon, St. John’s Island, Singapore, 1°12.92’N, 103°51.09’E, coll. CMBS II Expedition, 20 May 2013. Paratypes: 1 female (18.7 × 15.2 mm) (ZRC 2022.0971), same locality and collection information as Holotype. — 3 males (16.2 × 13.8 mm, 15.9 × 14.2 mm, 12.6 × 11.0 mm), 1 female (13.2 × 11.2 mm) (ZRC 2002.0154), Kusu Island, Singapore, coll. C.D. Schubart, 15 April 2000. — 1 male (15.0 × 13.0 mm) (ZRC 2002.0155), Sungei Buloh, Singapore, coll. C.D. Schubart & N. Sivasothi, 16 August 1999. — 6 males (16.9 × 14.3 mm, 15.8 × 13.5 mm, 15.2 × 13.1 mm, 15.1 × 13.0 mm), 2 females (15.1 × 12.8 mm, 12.7 × 10.3 mm) (ZRC 2022.0972), Lazarus Island, Singapore, coll. B.Y. Lee et al., 30 September 2018. Other material examined. South Korea: 1 female (12.8 × 10.8 mm) (WS900), Gosan, Jeju Island, coll. J. Jung, May 2003.— 1 female (14.5 × 12.1 mm) (WS899), Gosan, Jeju Island, coll. J. Jung, May 2003.— 1 male (16.2 × 14.2 mm) (WS898), Gosan, Jeju Island, coll. J. Jung, May 2003. Japan: 3 males (18.2 × 15.0 mm, 12.7 × 11.0 mm, 9.4 × 8.1 mm), 1 female (19.0 × 15.8 mm) (ZRC 2022.0990), Yagaji-jima, Okinawa, coll. J.J. Li, 18 July 2016.— 1 damaged female (11.9 × 10.1 mm) (ZRC 1999.0254), Nago City, Yagaji Island, coll. 23 May 1978.— 1 male (9.4 × 8.0 mm) (ZRC 2011.1037), Kumeijima, Okinawa, coll. T. Komai, 16 November 2009.— 1 female (10.3 × 8.8 mm) (ZRC 2011.1036), Ohara, Kumejima, 26°20’57.8”N 126°43’33.9”E, coll. Kumeijima 2009 Expedition, 18 November 2009. Taiwan: 1 ovigerous female (15.8 × 13.4 mm) (ZRC 2022.0991), Xiangjiaowan, coll. J.J. Li, 25 August 2014.— 1 ovigerous female (16.0 × 13.2 mm) (ZRC 2022.0992), Dintanzi, Hengchun Township, Pintung County, coll. Z.C. Ji, 3 September 2017. Hong Kong: 1 male (12.4 × 10.7 mm), 1 female (12.5 × 10.5 mm) (ZRC 2022.0993), To Kwa Peng, Sai Kung, coll. K. Wong, 9 May 2016.— 1 male (17.3 × 15.0 mm) (ZRC 2022.0994), Kak Po, Northeast Hong Kong, coll. C. Cheng & A. Quadros, 18 September 2017. Philippines: 1 male (17.4 × 14.9 mm), 1 female (14.9 × 12.7 mm) [photographed] (ZRC 2022.0973), stn M52, Danao embayment, Panglao Island, coll. Panglao Marine Biodiversity Project, 30 June 2004.— 1 female (15.9 × 13.4 mm) (ZRC 2022.0974), Kawasan, Cebu, coll. H.-C. Liu, 25 November 2001. Vietnam: 1 male [photographed] (17.4 × 15.2 mm) (ZRC 2022.0975), stn CD-I-08 (22), Bay Canh Island, 08°40.125’N 106°40.933’E, coll. 14 April 2010.— 1 female [photographed] (14.1 × 12.0 mm) (ZRC 2022.0976), stn CD-I-08 (24), Bay Canh Island, 08°40.125’N 106°40.933’E, coll. 14 April 2010.— 1 male [photographed] (16.0 × 13.5 mm) (ZRC 2022.0977), stn CD-I-08 (23), Bay Canh Island, 08°40.125’N 106°40.933’E, coll. 14 April 2010.— 8 males (16.5 × 14.2 mm, 14.0 × 12.0 mm, 7.6 × 6.3 mm, 6.1 × 5.1 mm), 17 females (16.2 × 13.1 mm, 14.9 × 12.6 mm, 7.6 × 6.5 mm, 5.0 × 4.3 mm) (ZRC 2022.0978), stn CD-I-08, Bay Canh Island, 08°40.125’N 106°40.933’E, coll. 14 April 2010. Malaysia: 1 male (14.0 × 11.8 mm) (ZRC 2022.0979), Sungei Baharu mangrove, Pulau Tioman, coll. LSM4263 students, 18 July 2014.— 1 male (11.6 × 9.8 mm) (ZRC 2016.0140), Telok Bakau, Pulau Tulai, Pulau Tioman, Pahang, coll. H.H. Tan & M.A.H. Chua, 16 July 2016.— 1 male (12.8 × 11.0 mm) (ZRC 2022.0980), Sungei Baharu mangrove, Pulau Tioman, coll. B.Y. Lee, 15 July 2014.— 2 males (12.5 × 11.0 mm, 8.2 × 7.8 m) (ZRC 2022.0981), Sungei Baharu mangrove, Pulau Tioman, coll. B.Y. Lee, 15 July 2014.— 1 female (17.6 × 14.6 mm) (ZRC 2000.1655), Pulau Manukan, Kota Kinabalu, Sabah, coll. C.D. Schubart, 22 June 2000.— 2 females (12.8 × 10.5 mm, 9.0 × 7.7 mm), 1 ovigerous female (12.5 × 10.1 mm), 1 juvenile female (5.5 × 4.9 mm) (ZRC 1969.10.13.3), Johore Straits, coll. C.L. Soh, 5 October 1969.— 3 males (12.8 × 11.0 mm, 9.8 × 8.8 mm, 9.7 × 8.6 mm), 4 females (15.8 × 13.8 mm, 13.4 × 11.3 mm, 9.3 × 8.0 mm, 7.7 × 6.6 mm) (ZRC 2022.0982), near Sungei Lengara, Langkawi Island, coll. A.D. Tran et al., 20 November 2006.— 2 males (12.5 × 11.0 mm, 9.6 × 8.3 mm) (ZRC 2022.0983), river mouth of Sungei Temurun, Langkawi, coll. T. Naruse, 20 November 2006.— 10 males (13.3 × 11.4 mm, 13.2 × 11.0 mm, 8.7 × 7.3 mm, 7.4 × 6.6 mm), 3 females (11.5 × 9.7 mm, 6.7 × 5.7 mm, 6.7 × 5.5 mm) (ZRC 1969.10.14.1–10), Johore Straits, coll. C.L. Soh, 12 October 1969. Indonesia: 1 male (11.2 × 9.7 mm), near NAD resort, Mawali Bay, Pulau Lembeh, Manado, Sulawesi, coll. H.H. Tan, December 2018. Diagnosis. Carapace slightly wider than long,almost square; postfrontal lobes weak, divided by weak longitudinal grooves; median pair slightly prominent, nearly equal in proportions. Inner orbital angle present, rounded, distinct, slight gap between inner orbital angle and frontal margin (Fig. 7C). Anterolateral margin of carapace with distinct external orbital angle, second tooth blunt, weak or absent; posterolateral margin slightly convex (Figs. 2B, C, 7A). Chelipeds dorsal surface of dactylus of chela with between 20–31 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Fig. 5E); presence of single longitudinal pectinated ridge with between 28–45 chitinous comb-like tubercles on dorsal surface of chela; row of 10–20 granules clustered behind pectinated ridge on chela; inner surface of palm granulated, vertical rows of 6–10 granules in adult male (Fig. 5F). P5 merus relatively short, wide, length approximately 0.36–0.43 times width (Fig. 6E–K). G1 slightly curved, distal region slightly wide; chitinous tip narrow, short; tip with dense tufts of setae, single row of plumose setae on exterior lateral margin; chitinous tip visible when denuded (Fig. 11A–D). Female vulvae with central operculum, round, slightly protruded (Fig. 8D). Description. Carapace slightly wider than long, almost square; covered with dense coat of setae, including small tufts of setae resembling tubercles; carapace margin fringed with dense, short setae; carapace smooth when denuded. Postfrontal lobes weak, divided by weak longitudinal grooves; median pair slightly prominent, nearly equal in proportions. Inner orbital angle present, rounded, distinct, slight gap between inner orbital angle and frontal margin (Fig. 7C). Anterolateral margin of carapace with distinct external orbital angle, second tooth blunt, weak or absent; posterolateral margin slightly convex (Figs. 2B, C, 7A). Antenna short. Antennules folding transversely. Third maxilliped with merus relatively rounded. Epistomial margin relatively narrow (Fig. 7C). Chelipeds subequal in size; large, robust in male; outer surface of palm smooth when denuded; dorsal surface of carpus covered with small tufts of setae resembling tubercles, smooth when denuded.; dorsal surface of dactylus of chela with between 20–31 evenly spaced tubercles, proximal tubercles small, increasing in size medially, decreasing in size distally (Fig. 5E); absent or weak granules in female and juveniles; presence of single longitudinal pectinated ridge with between 28–45 chitinous comb-like tubercles on dorsal surface of chela; row of 10–20 granules clustered behind pectinated ridge on chela; inner surface of palm granulated, vertical rows of 6–10 granules in adult male (Fig. 5F). P2–P5 covered with dense coat of setae, with small tufts of setae resembling tubercles, longer setae on dorsal and ventral margins except distal portion of dactylus (Figs. 2B, C, 7A); P5 merus relatively short, wide, length approximately 0.36–0.43 times width (Fig. 6E–K). Male thoracic sternum sparsely covered with short setae, smooth when denuded; sternites 1 and 2 fused, with suture between fused sternites 3 and 4; sternopleonal cavity extends nearly to edge of sternites 1 and 2. Male pleon wide and stout, long setae fringed pleon margin; somite 3 widest; somite 6 with curved edge; telson wide, dome-shaped (Fig. 7B); male pleonal locking mechanism absent, without tubercle on sternite 5. G1 slightly curved, distal region slightly wide; chitinous tip narrow, short; tip with dense tufts of setae, single row of plumose setae on exterior lateral margin; chitinous tip visible when denuded (Fig. 11A–D). G2 shorter than G1, slightly curved, tip rounded. Female pleon rounded, relatively wider than long; telson dome-shaped (Fig. 8C). Vulvae with central operculum, round, slightly protruded; sternal vulvar cover slightly raised above operculum (Fig. 8D). Colouration. Chelipeds with fingers yellowish white and purple to purplish red palm (Fig. 2B, C). In life, entire crab covered in a thin layer of mud. Etymology. The term, “ nobile ” alludes to the distinctive colour of the chela of the species; purple being a colour normally associated with royalty. Remarks. Clistocoeloma nobile n. sp. has a mix of diagnostic characters of C. villosum and C. suvaense. In the form of the carapace, C. nobile n. sp. has a weak external orbital angle on the anterolateral margin of carapace (Figs. 2B, C, 7A), and this is intermediate in form between C. villosum (which has no external orbital angle visible; Fig. 1A) and C. suvaense (which has a distinct external orbital angle; Figs. 2A, 3A, 4A). This character is very consistent in the large series of specimens of C. nobile n. sp. examined. With regards to the armature on the dorsal margin of the dactylus of the male chela, C. nobile n. sp. has 20–31 granules (Fig. 5E), in contrast to only16 granules present in C. villosum (Fig. 5A). The number of granules in C. suvaense (22–27; Figs. 3E, 5C), however, overlaps with that of C. nobile n. sp. The G1of C. suvaense is also very similar to that of C. nobile n. sp., with the lateral view appearing stout with a distinct angle before chitinous tip in C. suvaense (Figs. 9F, 10B, F) while the lateral view appearing narrower with a weak angle visibile before the chitinous tip in C. nobile n. sp. (Fig. 11B). The lateral view of the G1 of C. villosum, on the other hand, appears narrow with no clear angle before chitinous tip (Fig 9B). In addition, C. nobile n. sp. can also be separated from C. suvaense in having a rounded and slightly protruded vulvar cover (Fig. 8D); which is ovate and less protruded vulvar cover in C. suvaense (Fig. 8B). Females of C. villosum s. str. are not known. On the basis of these characters, it is clear that most of the records of “ C. villosum ” from the IWP, including Singapore, belong to C. nobile n. sp. instead. For the record by Tesch (1917: 208–210, pl. 17 fig. 2), his description and figures of the female specimen match C. nobile n. sp.. Tesch (1917) had noted that the New Guinea specimens has a distinct external orbital angle, in contrast to the type of C. villosum described by De Man (1987: 644) which has an indistinct one. The older records of “ Cleistocoeloma suvaense ” from Singapore by Tan & Ng (1994) and Wee & Ng (1994), are based on specimen records in the ZRC, are here referred to C. nobile n. sp. instead. A number of records of “ C. villosum ” from the IWP, however, are in doubt as we have neither specimens from these areas nor good descriptions and/or figures on which to base our decisions (see later). Ecological notes. Clistocoeloma nobile n. sp. is not a mangal species, although it has been found in habitats near mangroves. Most of the Singapore specimens were found in muddy substrates amongst the rocky intertidal areas in the supralittoral zone, typically found under large rocks with muddy substrate. For the Japanese specimens (as C. villosum), it was observed by Komai et al. (2004) that it can be found underneath rocks or fallen leaves within the landward side of the mangrove, while Yuhara et al. (2017) noted that this species was found underneath gravels and cobble stones around the edges of mangrove forests. For the Korean material, Lee et al. (2010) noted that the species occurred in “slightly moist soil under fallen leaves”. Distribution. Singapore (type locality), Sumatra (De Man 1897, 1898), Caroline Islands (Rathbun 1907), New Guinea (Tesch 1917), the Philippines (McNeill 1968), Taiwan (Li 2015; Li et al. 2019), Japan (Komai et al. 2004; Yuhara et al. 2017; Takakura & Komai 2019), and South Korea (Lee et al. 2010)., Published as part of Ng, Bee Yan Lee Ngan Kee & Ng, Peter K. L., 2023, On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata), pp. 217-236 in Zootaxa 5318 (2) on pages 226-234, DOI: 10.11646/zootaxa.5318.2.3, http://zenodo.org/record/8162486, {"references":["De Man, J. G. (1897) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 9 (1), 75 - 218. [in German]","De Man, J. G. (1898) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Sechster (Schluss-) Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (4), 677 - 708, pls. 28 - 38.","Milne-Edwards, A. (1869) Notes sur quelques nouvelles especes de genre Sesarma. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 5, 25 - 31 [in French]","McNeill, F. A. (1968) Crustacea, Decapoda and Stomatopoda. Great Barrier Expedition 1928 - 29, Scientific Reports, 7 (1), 1 - 98, pls. 1 + 2.","Rathbun, M. J. (1907) Reports on the scientific results of the expedition to the tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer \" Albatross, \" from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding. IX. Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer \" Albatross, \" from October, 1904, to March, 1905, Lieut. - Commander L. M. Garrett, U. S. N., commanding. X: The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2), 25 - 74.","Tesch, J. J. (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the Indo-Pacific species. Zoologische Mededelingen, Leiden, 3 (2 - 3), 127 - 260, pls. 15 - 17. https: // doi. org / 10.5962 / bhl. title. 10640","Nomoto, A., Yodo, S., Kimura, S., Kishino, T., Sakano, M. & Wada, K. (1999) Six rare brachyuran species of the family Grapsidae, recorded from the Kinokawa River Estuary, Wakayama Prefecture. Nanki Seibutu, 41 (1), 5 - 9 [in Japanese]","Kishino, T., Yonegawa, T., Nomoto, A., Kimura, S. & Wada, K. (2001 a) Twelve rare species of brachyuran crabs recorded in the brackish waters of Amami-Oshima, Kagoshima Prefecture, Japan. Nanki Seibutsu, 43 (1), 15 - 22. [in Japanese]","Kishino, T., Yonegawa, T., Nomoto, A., Kimura, S. & Wada, K. (2001 b) Brachyuran crab species recorded in the brackish waters of Amami-Oshima, Kagoshima Prefecture, Japan. Nanki Seibutsu, 43 (2), 125 - 131. [in Japanese]","Shokita, S., Nagai, T., Fujita, Y., Naruse, T., lto, A., Nagamatsu, T., Yamazaki, T., Shinjo, M. & Nagata, Y. (2002) [= Distribution and abundance of crustaceans in the Ohura River mangrove swamp of Okinawa Island, Japan]. In: A Comprehensive Study for Mangrove Ecosystem in Okinawa. Research Institute for Subtropics, Naha, Okinawa, pp. 73 - 86. [in Japanese with an English summary]","Tan, C. G. S. & Ng, P. K. L. (1994) An annotated checklist of mangrove brachyuran crabs from Malaysia and Singapore. Hydrobiologia, 285, 75 - 84. https: // doi. org / 10.1007 / BF 00005655","Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38.","Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda - Anomura, Brachyura. In: Wells, A. & Houston, W. W. K. (Eds.), Zoological Catalogues of Australia. Vol. 19.3 B. CSIRO Publishing, Melbourne, pp. i - xiv + 1 - 641.","Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y. & Shokita, S. (2004) New Records of Four Grapsoid Crabs (Crustacea: Decapoda: Brachyura) from Japan, with Notes on Four Rare Species. Natural History Research, 8 (1), 33 - 63.","Karasawa, T., Kimura, S., Kuroda, M. & Nomoto, A. (2006) Record of Clistocoeloma villosum (Crustacea, Brachyura, Sesarmidae) from the Waka River Estuary in Wakayama Prefecture, Japan. Nanki-Seibutu, 48 (1), 60 - 62. [in Japanese]","Schubart, C. D., Liu, H. - C. & Ng, P. K. L. (2009) Revision of Selatium Serene & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), with description of a new genus and two new species. Zootaxa, 2154 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2154.1.1","Lee, S. - K., Jung, J. & Kim, W. (2010) A new report on sesarmid crab Clistocoeloma villosum (Crustacea: Decapoda: Brachyura) from Korea. Korean Journal of Systematic Zoology, 26 (2), 179 - 181. https: // doi. org / 10.5635 / KJSZ. 2010.26.2.179","Yuhara, T., Yokooka, H. & Taru, M. (2017) Range extension of the sesarmid crab Clistocoeloma villosum along the eastern Pacific coast of the Izu Peninsula, Japan. Marine Biodiversity Records, 10, 19. https: // doi. org / 10.1186 / s 41200 - 017 - 0122 - 1","Yuhara, T. & Yokooka, H. (2019) Life history of the sesarmid crab, Clistocoeloma villosum, on the southern coast of the Izu Peninsula, central Japan. Plankton & Benthos Research, 14 (1), 8 - 13. https: // doi. org / 10.3800 / pbr. 14.8","Li, J. - J., Hsu, J. - W., Ng, N. K. & Shih, H. - T. (2019) Eight new records of crabs (Decapoda, Brachyura: Sesarmidae, Varunidae) from the coasts of Taiwan. Crustaceana, 92 (10), 1207 - 1230. https: // doi. org / 10.1163 / 15685403 - 00003935","Takakura, H. & Komai, T. (2019) New records of a semiterrestrial sesarmid crab Clistocoeloma villosum (A. Milne-Edwards, 1869) (Crustacea: Decapoda: Brachyura: Grapsoidea) from Izu and Boso Peninsulas, Japan. Cancer, 28, 25 - 31. [in Japanese with English abstract]","Li, J. - J. (2015) Land crabs of Christmas Island and Taiwan. Taiwan Natural Science, 126, 84 - 91. [in Chinese]","Wee, Y. C. & Ng, P. K. L. (Eds.) (1994) A first look at biodiversity in Singapore. National council on the Environment, Singapore, 163 pp."]}

Genus Clistocoeloma A. Milne-Edwards, 1873 Type species. Clistocoeloma balansae A. Milne-Edwards, 1868, by monotypy., Published as part of Ng, Bee Yan Lee Ngan Kee & Ng, Peter K. L., 2023, On the identities of the sesarmid crabs, Sesarma villosum A. Milne-Edwards 1869, and Clistocoeloma suvaense Edmondson, 1951, with the description of a new species from the West Pacific (Decapoda, Brachyura, Thoracotremata), pp. 217-236 in Zootaxa 5318 (2) on page 218, DOI: 10.11646/zootaxa.5318.2.3, http://zenodo.org/record/8162486

Schweitzer, Carrie E., Santana, William, Pinheiro, Allysson, Feldmann, Rodney M. (2023): Validation of Bahiacaris Schweitzer, Santana, Pinheiro & Feldmann (Crustacea, Decapoda, Caridea) from the Cretaceous (Aptian) of Brazil. Zootaxa 5318 (2): 299-300, DOI: 10.11646/zootaxa.5318.2.13, URL: http://dx.doi.org/10.11646/zootaxa.5318.2.13

Parapleisticantha aie n. sp. (Figs. 1D, 14–18) Type material. Holotype: male (cl 18.7 mm, pcl 16.5 mm, cw 12.4 mm, pcw 11.5 mm) (MNHN-IU-2014-18677a), stn CP 4502, New Ireland, 02°32’S 150°44’E, 193–206 m, coll. N.O. Alis, KAVIENG 2014 expedition, 7 September 2014. Paratypes: 1 female (cl 14.9 mm, pcl 12.6 mm, cw 10.0 mm, pcw 9.2 mm) (MNHN-IU-2014-18677b), same data as holotype; 1 ovigerous female (cl 17.7 mm, pcl 15.4 mm, cw 12.9 mm, pcw 12.2 mm) (ZRC 2022.73 ex MNHN-IU-2014-10019), stn DW 4415, New Ireland, 02°31’S 150°38’E, 212–214 m, coll. N.O. Alis, KAVIENG 2014 expedition, 27 August 2014. All specimens from Papua New Guinea. Diagnosis. Small-sized species (ovigerous female less than 17 mm long); regions well defined; base of epigastric region with 2 short spines, meso- and metagastric regions with 1 median spine each, protogastric spines with 1 short oblique spine, cardiac region with 2 laterally positioned short spines, intestinal region with 2 short spines, branchial region with 1 prominent anterior spine and 1 short posterior spine, lateral branchial margin with 7–9 larger spines and many smaller ones, hepatic region with 2 strong spines; rest of carapace covered with short spinules and sharp granules (Figs. 14, 15A–C, 16A, B); pseudorostral spines clearly diverging, forming V-shape, each spine relatively short, with 4 lateral accessory spines; ventral surface with 2 spines; distal border of antennular fossa with strong distal anteriorly directed spine; supraorbital eave relatively low, narrow, preorbital tooth long, margin of eave with 5 spines, proximal one small, no obvious large antorbital spine; intercalated spine distinct, smaller than prominent postorbital spine (Fig. 15A–D); interantennular spine bifid distally (Fig. 15D); basal antennal article long with 2 distal spines, one larger, outer margin with 3 spines, inner margin with 1 short spine, first article with prominent lateral spine (Fig. 16C, D). Third maxilliped pediform: ischium with 2 longitudinal rows of sharp granules on outer surface, lateral row with 8 tubercles, mesial row with 5 tubercles; merus subtriangular, longer than broad, with 4 sharp tubercles on outer surface, exopod with row of 9 tubercles or granules (Fig. 16E, F). Cheliped palm not inflated; propodal margin below articulation of dactylus and pollex with low subtruncate granulated process; cutting edge fingers with evenly sized teeth, those on proximal part of dactylus slightly larger (Fig. 17B). Distal margin of pleonal somite 6 deeply concave (Fig. 16E). G1 relatively slender, slightly curved along basal two-thirds, distal part bent outwards, forming ca. 60° angle, distal part elongate, with small subdistal process (Figs. 17C–E, 18A–D). G2 about a third length of G1, tip spatuliform (Figs. 17F, 18E). Female. The females agree well with the male in all non-sexual aspects. The female pleon has somites 1–5 free (each with a median tubercle), with somites 1–3 relatively narrow and somites 4–6 increasingly wider; somite 6 is functionally fused to the telson although the suture is still visible; the structure appearing almost round in ventral view, prominently dome-shape with the telson semicircular (Fig. 17G). The vulva is large and open laterally on a raised part of sternite 6 (Fig. 17H). Colour. The carapace is pale yellow to orange with the chelipeds and ambulatory legs orangish-red with patches of white (Fig. 1D). Etymology. The name is derived from the French word “aie” for an exclamation of pain, alluding to the prickly features of the species. The name is used as a Latin noun in apposition. Remarks. The present new species is closest to P. japonica, known from only a few specimens, all from Japan (Yokoya 1933: 140, text-fig. 50; Takeda & Miyake 1969: 494, pl. 18, fig. A, text-fig. 9c, d; Sakai 1976: 174; Sakai 1986: 238; 1940: 55; Richer de Forges et al. 2013: 17, figs. 1–3, 6A–E). Parapleisticantha aie n. sp., however, differs from P. japonica in having the carapace proportionately longer (Figs. 14, 15A, B; cf. Richer de Forges et al. 2013: figs. 1A, B, 3A, B); the basal antennal article is distinctly more elongate (Fig. 16C, D; cf. Richer de Forges et al. 2013: fig. 1C); the third maxilliped ischium is proportionately longer (Fig. 16F; cf. Richer de Forges et al. 2013: fig. 2C); and most significantly, the G1 has the distal part less distinctly curved with the tapering tip more elongate and there is a small subdistal fold (Fig. 17C–E, 18C, D) (versus G1 distal part curving at almost right angles with the tip shorter and there is no trace of a subdistal flap in P. japonica; cf. Richer de Forges et al. 2013: fig. 6A–D). The holotype male is not fully mature as its chelae are still relatively slender (Fig. 17B); in adult males of Parapleisticantha, the chelae are inflated and stout (Richer de Forges et al. 2013: figs. 2E, 5E)., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on pages 19-20, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["Yokoya, Y. (1933) On the distribution of Decapod Crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by the S. S. Soyo-Maru, during the year 1923 - 1930. Journal of the College of Agriculture, Imperial University of Tokyo, 12 (1), 1 - 226.","Takeda, M. & Miyake, S. (1969) Crabs from the East China Sea. III Brachygnatha Oxyrhyncha. Journal of the Faculty of Agriculture, Kyushu University, 15 (4), 469 - 525. https: // doi. org / 10.5109 / 22797","Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vols. 1 - 3. Kodansha Ltd., Tokyo, pp. xxix + 773 pp. (English Txt), 461 pp. (Japanese Txt), 16 pp., 251 pls. (Pls. Vol.)","Sakai, T. (1986) Rare crabs and their genus in Japan. Research on Crustacea, 15, 1 - 4. https: // doi. org / 10.18353 / rcustacea. 15.0 _ 1","Richer de Forges, B., Ng, P. K. L. & Ahyong, S. T. (2013) Parapleisticantha Yokoya, 1933, a valid genus of deep-sea inachid spider crab from Japan and the Philippines (Crustacea: Decapoda: Brachyura: Majoidea), with the description of a new species. Zootaxa, 3635 (1), 15 - 26. https: // doi. org / 10.11646 / zootaxa. 3635.1.2"]}

Genus Oxypleurodon Miers, 1885 Oxypleurodon Miers, 1885: 588 [type species: Oxypleurodon stimpsoni Miers, 1885, by monotypy]. Oxypleurodon��� Miers 1886: 38.��� Tavares 1991: 167, 169.��� Ng et al. 2008: 104.��� Richer de Forges & Ng 2009a: 2.��� Richer de Forges & Ng 2009b: 250.��� Richer de Forges 2010: 646. Nasutocarcinus Tavares 1991: 160 (list), 169.��� Ng et al. 2008: 104. (For the complete synonymy of the genus, see Richer de Forges 2010: 646). Remarks. There are currently 31 recognised species in this genus (Richer de Forges & Ng 2009a, b; Richer de Forges 2010; Richer de Forges & Corbari 2012; Lee et al. 2015, 2017). Garth (1958) synonymised the Indo-West Pacific genus Oxypleurodon Miers, 1885, under Sphenocarcinus A. Milne-Edwards, 1878, following Alcock (1895, 1899). Sphenocarcinus was then synonymised under Rochinia A. Milne-Edwards, 1875, by Griffin & Tranter (1986). Subsequently, Tavares (1991) resurrected the genera, Oxypleurodon and Sphenocarcinus (see Lee et al. 2015; Tavares & Santana 2018), and established a new genus for species with a very elongated pseudorostral spine, Nasutocarcinus Tavares, 1991. Richer de Forges & Ng (2009b) reviewed the taxonomy of both genera, noting that Oxypleurodon is quite distinct from Rochinia, and synonymised Nasutocarcinus under Oxypleurodon (see also Lee et al. 2015). With the description of two new species from PNG, there are now 33 species in Oxypleurodon., Published as part of Lee, Bee Yan, Forges, Bertrand Richer De & Ng, Peter K. L., 2019, Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai & Sato, 2017, and descriptions of two new genera (Crustacea: Decapoda: Brachyura: Majoidea), pp. 1-44 in Zootaxa 4619 (1) on page 5, DOI: 10.11646/zootaxa.4619.1.1, http://zenodo.org/record/3248363, {"references":["Miers, E. J. (1885) The Brachyura. In: Tizard, T. H., Moseley, H. N., Buchanan, J. Y. & Murray, J. (Eds.), Narrative of the cruise of H. M. S. Challenger with a general account of the scientific results of the expedition. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, R. N., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. & c. Regius Professor of Natural History in the University of Edinburgh Director of the civilian scientific staff on board and now of John Murray, one of the naturalists of the Expedition. Narrative 1 (2). Published by Order of Her Majesty's Government, London, Edinburgh and Her Majesty Stationery Office, Dublin, pp. 585 - 592.","Miers, E. J. (1886) Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 1876. In: Wyville Thomson, C. & Murray, J. (Eds.), Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, R. N., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. & c. Regius Professor of Natural History in the University of Edinburgh Director of the civilian scientific staff on board and now of John Murray, LL. D., Ph. D., & c. one of the naturalists of the Expedition, Zoology. 17 (2). Published by Order of Her Majesty's Government, London, Edinburgh and Her Majesty Stationary Office, Dublin, pp. l [= 50] + 1 - 362, pls. 1 - 29.","Tavares, M. S. (1991) Redefinition des genres Rochinia A. Milne Edwards, Sphenocarcinus A. Milne Edwards et Oxypleurodon Miers, et etablissement du genre Nasutocarcinus gen. nov. (Crustacea, Brachyura, Majidae). Bulletin du Museum national d'Histoire naturelle, Paris, 4 e serie, Section A, 13 (1 - 2), 159 - 179.","Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286, figs. 1 - 198, 2 figs.","Richer de Forges, B. & Ng, P. K. L. (2009 a) New genera, new species and new records of Indo-West Pacific spider crabs (Crustacea: Brachyura: Epialtidae: Majoidea). Zootaxa, 2025, 1 - 20.","Richer de Forges, B. & Ng, P. K. L. (2009 b) On the Majoid genera Oxypleurodon Miers, 1886, and Sphenocarcinus A. Milne-Edwards, 1875 (Crustacea: Brachyura: Epialtidae), with descriptions of two new genera and five new species. Raffles Bulletin of Zoology, Supplement, 20, 247 - 266.","Richer de Forges, B. (2010) Majoid crabs from the Mozambique Channel with the description of a new species of Oxypleurodon Miers, 1886 (Decapoda, Brachyura). In: Fransen, C., De Grave, S. & Ng, P. K. L. (Eds.), Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, pp. 645 - 653. https: // doi. org / 10.1163 / 9789047427759 _ 046","Richer de Forges, B. & Corbari, L. (2012) A new species of Oxypleurodon Miers, 1886 (Crustacea, Brachyura, Majoidea) from the Bismarck Sea, Papua New Guinea. Zootaxa, 3320 (1), 56 - 60. https: // doi. org / 10.11646 / zootaxa. 3320.1.4","Lee, B. Y., Corbari, L. & Richer de Forges, B. (2015) Deep-sea spider crabs of the genus Oxypleurodon Miers, 1885 (Decapoda, Brachyura, Majoidea, Epialtidae), from the NANHAI 2014 cruise in the South China Sea, with a description of a new species. Crustaceana, 88 (12 - 14), 1255 - 1263. https: // doi. org / 10.1163 / 15685403 - 00003488","Lee, B. Y., Richer de Forges, B. & Ng, P. K. L. (2017) Deep-sea spider crabs of the families Epialtidae Macleay, 1838 and Inachidae MacLeay, 1838, from the South China Sea, with descriptions of two new species (Decapoda, Brachyura, Majoidea). European Journal of Taxonomy, 358, 1 - 37. https: // doi. org / 10.5852 / ejt. 2017.358","Alcock, A. (1895) Materials for a carcinological fauna of India. No. 1. The Brachyura Oxyrhyncha. Journal of the Asiatic Society of Bengal, 44 (Part 2, No. 2), 157 - 291, pls. 3 - 5. https: // doi. org / 10.5962 / bhl. title. 16033","Alcock, A. (1899) An Account of the Deep-sea Brachyura collected by the Royal Indian Marine Survey Ship Investigator. Trustees of the Indian Museum, Calcutta, iii + 85 + [2] pp., IV pls. https: // doi. org / 10.5962 / bhl. title. 10330","Tavares, M. S. & Santana, W. (2018) Refining the genus Rochinia A. Milne-Edwards, 1875: reinstatement of Scyramathia A. Milne-Edwards, 1880 and Anamathia Smith, 1885, and a new genus for Amathia crassa A. Milne-Edwards, 1879, with notes on its ontogeny (Crustacea: Brachyura: Epialtidae). Zootaxa, 4418 (3), 201 - 227. https: // doi. org / 10.11646 / zootaxa. 4418.3.1"]}

Nyborg, Torrey, Garassino, Alessandro, Vega, Francisco J. (2023): Validation of Cretalamoha boweni Nyborg & Garassino (Brachyura: Homolidae) from the early Campanian, Upper Cretaceous of British Columbia, Canada. Zootaxa 5318 (1): 148-150, DOI: 10.11646/zootaxa.5318.1.8, URL: http://dx.doi.org/10.11646/zootaxa.5318.1.8

Genus Dorhynchus Wyville Thomson, 1873 Type species. Dorhynchus thomsoni Wyville Thomson, 1873, by monotypy. Remarks. Manning & Holthuis (1981: 280–281) commented that the species in Achaeopsis Stimpson, 1857, can be split into two groups: one with shorter pseudorostral spines, relatively shorter and stouter P2–P5, the P2 merus is shorter than the carapace length, the P2 and P3 dactyli are gently curved and the P4 and P5 dactyli falciform, and occurring in waters less than 200 m depth (A. spinulosa Stimpson, 1857, type species); and the other with longer pseudorostral spines, relatively longer and more slender P2–P5, the P2 merus is longer than the carapace length, the P2–P5 dactyli all gently curved and occurring in waters greater than 200 m depth (Dorhynchus thomsoni Wyville Thomson, 1873; Lispognathus furcatus A. Milne-Edwards, 1880; Stenorhynchus ramusculus Baker, 1906, Achaeopsis rostrata Sakai, 1932). They suggested that the second group of species should be referred to Dorhynchus Wyville Thomson, 1873, instead. This was followed by Davie (2002) and has been adopted ever since. We are not fully convinced that this is a natural dichotomy, and some of the characters of the present new species challenge this separation. The type species of Dorhynchus (Dorhynchus thomsoni) is unusual in that the pseudorostral spines, frontal carapace regions, epistome and antennules are transversely much narrower (cf. Barnard 1950: fig. 4d) compared to most of the other species placed by Manning & Holthuis (1981) in the genus. Only the more recently described D. basi MacPherson, 1983, from Namibia shares similar characters (MacPherson 1983: figs. 21A, B, 22). The carapace of D. furcatus, however, is actually very similar to that of A. spinulosa, with both possessing relatively short pseudorostral spines, and notably, the lateral branchial spine is directed dorsally and not laterally (A. Milne-Edwards, 1880b: pl. 31A fig. 4). In other species of Dorhynchus (including D. thomsoni and the present new species, D. profundus), the lateral branchial spine is directed laterally. In addition, the male anterior thoracic sternum (sternites 1–4) of D. thomsoni is relatively wide (cf. Barnard 1950: fig. 4c) while that of D. profundus n. sp. is transversely narrower, with sternites 3 and 4 laterally constricted (Fig. 4E). The condition of the male anterior thoracic sternums of the other species is not known. The ambulatory dactyli of D. furcatus, nevertheless, are like the other described species of Dorhynchus (A. Milne-Edwards, 1880b: pl. 31A fig. 4), with the exception of the present new species (see below). A reappraisal of Achaeopsis and Dorhynchus is clearly necessary, with all the constituent species re-examined to clarify generic delineations. As for the lengths of the P2 and P3 merus, we find that they are sometimes shorter than the pcl; one male (ZRC 2022.0067) has the P2 merus 1.15 times the pcl, while its P3 merus is 0.94 times the pcl; while a female (MNHN-IU-2015-888) has the P2 and P3 meri 0.89 and 0.86 times the pcl, respectively. In the specimen of D. rostratus examined, the P2 and P3 meri are 1.18 and 1.04 times the pcl. These proportions therefore need to be used very carefully. A large series of specimens of a few species will be needed to ascertain its reliability and degree of variation in members of Dorhynchus. A note on the nomenclature of D. furcatus is necessary. The two names Lispognathus furcatus A. Milne-Edwards, 1880a, and Lispognathus furcillatus A. Milne-Edwards, 1880b, certainly refer to the same species as their accompanying data is the same. The genus Lispognathus was first established in A. Milne-Edwards (1880a) and the only described species was “ Lispognathus furcatus A. Milne-Edwards, 1880a ”. The genus was again described in a follow-up paper by A. Milne-Edwards (1880b) but this time, the only species described was “ Lispognathus furcillatus A. Milne-Edwards, 1880b ” (same spelling used in the plates). Why A. Milne-Edwards changed the spelling is not known but both are clearly objective synonyms as they share the same type series., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on pages 2-3, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["Wyville Thomson, C. (1873) The Depths of the Sea. An account of the general results of the dredging cruises of H. M. SS. ' Porcupine' and ' Lightning' during the summers of 1868, 1869, and 1870, under the scientific direction of Dr. Carpenter, F. R. S., J. Gwyn Jeffreys, F. R. S., and Dr Wyville Thomson, F. R. S. 1 st Edition. Macmillan and Co., London, xxi + 527 pp., 84 figs., 8 pls. https: // doi. org / 10.5962 / bhl. title. 16330","Manning, R. B. & Holthuis, L. B. (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, i - xii, 1 - 379, figs. 1 - 88, table 1. https: // doi. org / 10.5479 / si. 00810282.306","Stimpson, W. (1857) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars III. Crustacea Maioidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 216 - 221 (23 - 28). https: // doi. org / 10.5962 / bhl. title. 51447","Baker, W. H. (1906) Notes on South Australian decapod Crustacea. Part IV. Transactions of the Royal Society of South Australia, 30, 104 - 117, pls. 1 - 3.","Sakai, T. (1932) Notes on Some Rare Materials of Japanese Oxyrhyncha. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (4), 41 - 59, figs. 1 - 8, pls. 2 & 3.","Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum, 38, 1 - 837, figs. 1 - 154.","Macpherson, E. (1983) Crustaceos Decapodos capturados en las costas de Namibia. Resultados Expediciones Cientificas Buque oceanogr. Cornide Saavedra, 11 (Supplement Investigacion Pesquera), 3 - 79.","Milne-Edwards, A. (1880 b) Etudes sur les xiphosures et les crustaces podophthalmaires. In: Mission scientifique au Mexique et dans l'Amerique centrale. Recherches Zoologique pour servir a l'histoire de la faune de l'Amerique centrale et du Mexique, publiees sous la direction de M. H. Milne Edwards, membre de l'Institut. Cinquieme partie. Livraison 7, 265 - 312, pls. 31 A + 44 + 49 - 54.","Milne-Edwards, A. (1880 a) Etudes preliminaires sur les Crustaces, 1 ere partie. In: Reports on the results of dredging under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the U. S. Coast Survey Steamer \" Blake, \" Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding, VIII. Bulletin of the Museum of Comparative Zoology at Harvard College, 8, pp. 1 - 68, pls. 1, 2."]}

Grypachaeus aff. tenuicollis Takeda, 1978 (Figs. 1E, F, 23, 24) Material examined. 1 male (rostrum broken, pcl 10.1 mm, cw 6.0 mm, pcw 5.0 mm) (ZRC 2022.0075, ex MNHN- IU-2014-18677), stn 4502, New Ireland, Papua New Guinea, 02°32’S 150°44’E, 193–206 m, coll. N.O. Alis, KAVIENG 2014 expedition, 7 September 2014; 1 female (cl 7.6 mm, pcl 7.1 mm, cw 4.3 mm, pcw 4.0 mm) (ZRC 2002.505), stn V4698-107, East China Sea, 162 m, coll. cruise by Institute of Oceanography, Academia Sinica, China, 6 June 1975; 1 ovigerous female (cl 8.1 mm, pcl 7.2 mm, cw 4.9 mm, pcw 4.3 mm) (ZRC 2022.0077), stn T1, on mud with sponges, Bolod, Panglao Island, Bohol, Philippines, 9°32.4'N 123°47.3'E, 83–102 m, coll. PANGLAO 2004 Expedition, 30 May 2004; 1 ovigerous female (cl 11.6 mm, pcl 10.4 mm, cw 6.5 mm, pcw 5.8 mm) (ZRC 2012.1160), stn AT44, Urèlapa island, Vanuatu, coll. SANTO 2006 expedition, 29 September 2006. Remarks. The present specimen from Papua New Guinea is problematic because it is clearly neither G. hyalinus (Alcock & Anderson, 1894) nor G. tenuicollis Takeda, 1978. Grypachaeus hyalinus has a prominently shorter carapace with the “neck” (frontal and epigastric regions) shorter, the meso-, metagastric branchial regions are distinctly wider and the ambulatory legs are relatively short (cf. Alcock & Anderson 1896: pl. 17 fig. 3). Griffin (1974) reported on specimens from Mombasa and the Gulf of Aden and figured a G1 that is relatively straighter with the small dorsal fold closer to the tip (Griffin 1974: fig. 1e, f) (versus gently curved with the dorsal fold further from the tip in the present specimen; Fig. 23C). Griffin (1976: 190) also reported G. hyalinus from the Philippines but without any figures. Grypachaeus tenuicollis is a very distinctive species, with a very slender carapace, the “neck” is long and slender, the pseudorostral spines are relatively long and slender, and there is only one laterally directed spine on the hepatic region, the ambulatory legs are proportionately shorter, the G1 more slender and C-shaped with the distal part turned sharply at almost 90º (cf. Takeda 1978: figs. 1, 2, 7, 9–11) (versus carapace less elongate, the branchial regions are relatively wider, the ambulatory legs are proportionately longer, and the G1 is proportionately stouter less curved with the distal part tapering; Fig. 23A–C). The differences here suggest that the species on hand is new, but the present specimen is too incomplete to formally describe for now: its pseudorostrum is broken off, and both P5 are missing. There is a second specimen, a female (MNHN-IU-2015-85; stn DW4313, north of Normanby island, southeast of Entrecasteaux Archipelago, Papua New Guinea, 09°49’S 151°34’E, 105–175 m, coll. N.O. Alis, MADEEP cruise, 3 May 2014) (Fig. 1E) that was photographed but we have not been able to examine the specimen. It appears to agree very well with the male examined here and it still has the left P5 intact, but its pseudorostrum is also broken off. We have on hand one adult female specimen from Vanuatu (ZRC 2012.1160) that may be conspecific with the Papua New Guinea specimens, having the same carapace and “neck” proportions, the two hepatic spines directed obliquely, and the meso-, metagastric and branchial regions more quadrate in shape (Fig. 24F). This Vanuatu specimen has an intact pseudorostral spine and this is long (Fig. 24F). Its ambulatory legs, however, are proportionately shorter (Fig. 24C); but this may be due to sexual dimorphism. We also have two other specimens from the East China Sea (ZRC 2002.505) and Philippines (ZRC 2022.0077), and neither are G. hyalinus. Compared to G. tenuicollis, they are still distinctly wider at the gastric and branchial regions, the “neck” is distinctly shorter, the pseudorostral spines are relatively short, and there are two lateral spines on the hepatic region with the anterior one obliquely directed (Fig. 24D, E). As the two specimens are females, it is difficult to ascertain whether these are the result of sexual dimorphism, although it seems unlikely; the male and female specimens from Papua New Guinea do not show such prominent differences. Their short pseudorostral armature is different from those of G. tenuicollis s. str. and the Vanuatu specimen (pseudorostral spines long) and indicates this character is not associated with sexual dimorphism as all are females., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on pages 30-32, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["Takeda, M. (1978) A new spider crab from off Iriomote Island, the Ryukyu Islands. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 4 (2), 117 - 121, figs. 1 - 11.","Alcock, A. & Anderson, A. R. S. (1894) Natural History Notes from H. M. Indian Marine Survey Steamer ' Investigator, ' Commander C. F. Oldham, R. N., commanding. Series II, No. 17. List of the Shore and Shallow-water Brachyura collected during the Season 1893 - 1894. Journal of the Asiatic Society of Bengal, 63 (2 - Natural Science), 197 - 209.","Alcock, A. & Anderson, A. R. S. (1896) Illustrations of the Zoology of the Royal Indian Marine Surveying Steamer Investigator, under the command of Commander C. F. Oldham, R. N. Crustacea. Part IV. Published under the Authority of Captain J. Hert, R. N., C. I. E., Director of the Royal Indian Marine. Printed and sold by the Superintendent of Government Printing, Calcutta, pls. XVI - XXVII.","Griffin, D. J. G. (1974) Spider crabs (Crustacea: Brachyura: Majidae) from the International Indian Ocean Expedition 1963 - 64. Smithsonian Contributions to Zoology, 182, i - iv + 1 - 35. https: // doi. org / 10.5479 / si. 00810282.182","Griffin, D. J. G. (1976) Spider crabs of the family Majidae (Crustacea: Brachyura) from the Philippine Islands. Journal of Natural History, 10 (2), 179 - 222, figs. 1 - 11, tabs. 1 + 2. https: // doi. org / 10.1080 / 00222937600770131"]}

Genus Orbicantha n. gen. Diagnosis of female. Carapace subcircular; gastrohepatic and frontal regions not elongated, lateral constriction with branchial regions weak (Fig. 19A, B); dorsal surface not swollen, gently convex; mesogastric region with 1 median spine with rounded tip; protogastric region with 2 small spines arranged obliquely; metagastric region with 1 median spine with rounded tip cardiac region with 2 short spines positioned side by side; intestinal region with 1 large median tubercle and 1 short spine lateral to it on each side; hepatic region not swollen with 2 or 3 spines with round tips; branchial region with oblique median row of 3 spines; lateral branchial margin with 6–8 large and many small spines (larger ones round-tipped); dorsal carapace surface with numerous distinct rounded granules (Figs. 19A–E, 22A–C); pterygostomial region protruding anteriorly, forming angular shelf-like structure in lateral view (Fig. 19D, E); suborbital region narrow; pseudorostrum short, as long as supraorbital eave, base wide with 2 distal spines; interantennular spine distinct, tip bifurcated, clearly visible in dorsal view; postorbital spine gently curved with rounded tip; intercalated spine distinct with rounded tip; margin of supraorbital eave with 4 short spines, antorbital spine relatively short, no obvious preorbital spine (Figs. 19B, C, F, 20B); basal antennal article elongate, outer margin lined with short spines, third article short, subquadrate, ca. one-third length of basal article (Fig. 19F); eyes with short, stout peduncle (Fig. 19C, F); epistome is distinctly quadrate. Third maxilliped covered with sharp and rounded granules, those on inner margins sharper, ischium subrectangular, merus subtriangular, proximal margin with strong sharp tubercles, anteroexternal angle expanded, auriculiform (Fig. 20A). Chelipeds slender, relatively short (Fig. 19A, 22A). Ambulatory legs relatively long, slender, P2 and P3 propodus and dactylus with long stiff setae that resemble spines and setae on ventral margins, appearing comb-like (Figs. 19A, 20E, F, 22A), P4 and P5 prominently reduced in size, ventral margins of propodus and dactylus without long spines or long stiff setae (Figs. 19A, 20G, H, 22A). Thoracic sternite 4 with prominent broad V-shaped ridge medially that demarcates sternopleonal cavity (Fig. 14A). Pleon subcircular in shape, domed; covering most of thoracic sternum; somites 1–5 free but barely mobile; somite 6 and telson functionally fused (Fig. 21A, C, D); vulvae on anterior part of sternite 6, subovate, with narrow rim (Fig. 21B). Etymology. The name is derived from the Latin “orbis” for round, alluding to the shape of the carapace; in arbitrary combination with the genus name Pleistacantha. Gender feminine. Remarks. Orbicantha n. gen. is clearly related to the following oregoniid pleistacanthine genera: Pleistacantha Miers, 1879, Parapleisticantha Yokoya 1933, and Pleisticanthoides Yokoya, 1933. It is markedly different from all of these genera in that its carapace shape is semicircular, with the gastrohepatic and frontal regions not elongated (Figs. 19A, B, 22A, B); the pseudorostral spines are very short with the distal spines rounded at the tip (Figs. 19B, F, 22B); the gastric, cardiac and branchial regions possessing distinct spines which have a rounded tip (Figs. 19A–D, 22A–C); the ocular peduncle is very short (Fig. 19F); the third antennal article is very short, being almost quadrate and much shorter than the basal antennal article (Fig. 19F); the epistome is distinctly quadrate (Fig. 19F); the buccal cavity is proportionately much wider (Fig. 19E); and the P4 and P5 are prominently reduced in size and more slender in form compared to P2 and P3, with the propodus and dactylus lacking long spines or setae (Figs. 19A, 20D, E, 22A). In species of Pleistacantha, Parapleisticantha and Pleisticanthoides, the carapace shape is distinctly pyriform, with the gastrohepatic regions separated from the branchial regions by a lateral constriction; the pseudorostral spines usually relatively longer with the distal spines sharp or distinctly tapering; the gastric, cardiac and branchial regions possessing normal sharp spines; the ocular peduncle is elongate; the third antennal article is at least half the length of the basal antennal article, often being subequal in length, being almost quadrate; the epistome is transversely narrower; the buccal cavity is proportionately narrower; and the P5, while reduced in size compared to the other pereopods, is not otherwise different in form, with P3 and P4 similar in form and structure (cf. Guinot & Richer de Forges 1986, 1992; Ahyong & Lee 2006; Ahyong & Ng 2007; Ng & Richer de Forges 2012; Richer de Forges et al. 2013; Ahyong et al. 2015, 2019; Ng et al. 2017). In addition, the ornamentation on most of the dorsal carapace surface of Orbicantha n. gen. consists of small, rounded granules, not spines (Figs. 19B–D, 22B); those on Pleistacantha and Parapleisticantha are invariably covered with small spines. While Pleisticanthoides simplex (Rathbun, 1932) has a spine-covered carapace, the other two species (P. cameroni Ng & Richer de Forges, 2012, and P. piccardorum Ng & Richer de Forges, 2012) have much smoother carapaces with no spines on most of the surface (cf. Ng & Richer de Forges 2012). The presence of four short spines on the supraocular eave (Fig. 19B, C) and two distal spines on the ventral edge of the basal antennal article (Fig. 19F) are characters shared with Parapleistacantha; although they completely differ in the above discussed features (cf. Figs. 15C, 16C, D; Richer de Forges et al. 2013). The differences with the Pleistacantha Miers, 1879 s. lato are obvious, with the latter possessing long pseudorostral spines usually diverging in V bearing long accessory spines versus very short triangular spines; pyriform carapace covered with long sharp spines in Pleistacantha versus rounded carapace with short granule-like spines in Orbicantha n. gen.

Family Epialtidae Macleay, 1838 Remarks. Lee et al. (2019) reported on a large collection of epialtid crabs from Papua New Guinea and adjacent areas. The present study found further specimens of the following species previously reported from Papua New Guinea: Crocydocinus ewok Lee, Richer de Forges & Ng, 2019; Samadinia ahyongi (McLay, 2009); S. boucheti (Richer de Forges & Ng, 2013) and Tunepugettia corbariae Lee, Richer de Forges & Ng, 2019. In this paper, we only treat a few species. The taxonomy of Griffinia lappacea (Rathbun, 1918), which had earlier been misidentified as G. takedai Richer de Forges & Ng 2012, is reappraised and new characters used to separate these two allied species. Variation in G. lappacea is also discussed. Griffinia gilloloensis (Rathbun, 1916) and Oxypleurodon luzonicum (Rathbun, 1916) are also recorded from Papua New Guinea. Specimens of Hyastenus ? borradailei (Rathbun, 1907) are now reported from Papua New Guinea for the first time and aspects of its taxonomy discussed., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on page 32, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In: Smith, A., Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa: ' together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe. Smith, Elder and Co., London, pp. 53 - 71, pls. 2 + 3.","Lee, B. Y., Richer de Forges, B. & Ng, P. K. L. (2019) Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai & Sato, 2017, and description of two new genera (Crustacea: Decapoda: Brachyura: Majoidea). Zootaxa, 4619 (1), 1 - 44. https: // doi. org / 10.11646 / zootaxa. 4619.1.1","Richer de Forges, B., Ng, P. K. L. & Ahyong, S. T. (2013) Parapleisticantha Yokoya, 1933, a valid genus of deep-sea inachid spider crab from Japan and the Philippines (Crustacea: Decapoda: Brachyura: Majoidea), with the description of a new species. Zootaxa, 3635 (1), 15 - 26. https: // doi. org / 10.11646 / zootaxa. 3635.1.2","Rathbun, M. J. (1918) Report on the spider crabs obtained by the F. I. S. \" Endeavour \" on the coasts of Queensland, New South Wales, Victoria, South Australia and Tasmania. Biological Results of the Fishing Experiments Carried On by the F. I. S. \" Endeavour \" 1909 - 14, 5 (1), 3 - 29, figs. 1 - 3, pls. 1 - 15. https: // doi. org / 10.5479 / si. 00963801.2135.527","Richer de Forges, B. & Ng, P. K. L. (2012) Griffinia takedai, a new species of deep sea majoid crab (Decapoda, Brachyura, Epialtidae) from the Philippines. In: Komatsu, H., Okuno, J. & Fukuoka, K. (Eds.), Studies on Eumalacostraca: a homage to Takeda Masatsune. Crustaceana Monographs, 17, 275 - 284. https: // doi. org / 10.1163 / 9789004202894 _ 023","Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. [Scientific results of the Philippine Cruise of the Fisheries Steamer \" Albatross, \" 1907 - 1910. - No. 34]. Proceedings of the United States National Museum, 50 (2135), 527 - 559."]}

Griffinia lappacea (Rathbun, 1918) (Figs. 25A–E, 26A, B, D, E, G–I) Antilibinia lappacea Rathbun, 1918: 12, fig. 3, pl. 7 fig. 3.—Hale 1927: 133, fig. 133; Barnard 1950: 37.—Griffin 1966: 267.— Sakai 1976: 201.— Griffin & Tranter 1986: 70. Pisidarum sp. Serène & Vadon, 1981:128, pl. 4F. Griffinia lappacea. — Richer de Forges 1994: 67, figs. 3A–D, 4B, C.— Ng et al. 2008: 100. Material examined. Solomon Sea: 1 ovigerous female (pcl 11.0 mm, pcw 7.4 mm) (MNHN-IU-2014-9637), stn CP 4330, 06°08’S 149°12’E, 315–625 m, coll. N.O. Alis, MADEEP cruise, 6 May 2014; 2 males (pcl 13.9 mm, pcw 9.8 mm [broken carapace]; pcl 12.0 mm, pcw 8.3 mm) (MNHN-IU-2014-9648), stn CP 4331, 06°07’S 149°12’E, 260 m, coll. N.O. Alis, MADEEP cruise, 6 May 2014; 1 ovigerous female (pcl 11.0 mm, pcw 8.1 mm) (MHNN-IU-2015-814), stn CP 4331, 06°07’S 149°12’E, 260 m, coll. N.O. Alis, MADEEP cruise, 6 May 2014; 1 female (pcl 7.5 mm, pcw 5.4 mm) (MNHN-IU-2015-63), stn CP 4337, 06°07’S 149°17’E, 287–447 m, coll. N.O. Alis, MADEEP cruise, 7 May 2014; 1 male (pcl 13.0 mm, pcw 9.4 mm), 1 ovigerous female (pcl 10.0 mm, pcw 7.5 mm) (MNHN- IU-2011-3529), stn CP 3708, 04°58’S 145°50’E, 502–529 m, coll. N.O. Alis, BIOPAPUA cruise, 2 October 2010. Papua New Guinea: 2 ovigerous female (pcl 12.1 mm, pcw 9.0 mm; pcl 12.7 mm, pcw 8.8 mm) (MNHN-IU-2011- 2341), 1 female (MNHN-IU-2011-2299), stn CP 3721, Vitiaz Strait, 06°03’S 147°37’ E, 542–554 m, coll. N.O. Alis, BIOPAPUA cruise, 7 October 2010; 2 ovigerous females (pcl 11.1 mm, pcw 8.1 mm; pcl 11.1 mm, pcw 8.2 mm) (MNHN-IU-2011-946), stn DW 3668, north of Rabaul, 04°08’S 151°58’E, 383–411 m, coll. N.O. Alis, BIOPAPUA cruise, 24 September 2010; 1 ovigerous female (pcl 11.1 mm, pcw 7.6 mm) (MHNH-IU-2014-9949), stn CP 4437, 02°23’S 150°37’E, 416–535 m, coll. N.O.Alis, KAVIENG 2014 expedition, 31August 2014; 1 ovigerous female (pcl 11.6 mm, pcw 9.4 mm) (MNHN-IU-2014-19049), stn CP 4423, New Ireland, 02°20’S 150°38’E, 550–649 m, coll. N.O. Alis, KAVIENG 2014 expedition, 28 August 2014; 1 ovigerous female (pcl 12.0 mm, pcw 8.5 mm) (MNHN- IU-2011-2889), stn CP 3709, off coast of Madang, 04°58’S 145°52’E, 640–675 m, coll. N.O. Alis, BIOPAPUA cruise, 2 October 2010; 1 ovigerous female (pcl 9.9 mm, pcw 7.9 mm) (MNHN-IU-2011-967), stn CP 3670, north of Rabaul, 04°06’S 151°56’E, 497–500 m, coll. N.O. Alis, BIOPAPUA cruise, 24 September 2010; 1 male (pcl 10.0 mm, pcw 7.3 mm) (MNHN-IU-2011-2203), stn CP 3672, north of Rabaul, 04°04’S 151°50’E, 702–724 m, coll. N.O. Alis, BIOPAPUA cruise, 24 September 2010; 1 male (pcl 10.3 mm, pcw 7.4 mm) (MNHN-IU-2011-2704), stn CP 3645, Tami Island, north of Huon, 06°44’S 147°50’E, 403–418 m, coll. N.O. Alis, BIOPAPUA cruise, 24 August 2010; 1 male (pcl 13.2 mm, pcw 9.5 mm) (MNHN-IU-2011-3194), coll. N.O. Alis, BIOPAPUA cruise, leg. 4, no other data; 1 male (pcl 13.0 mm, pcw 9.6 mm) (MNHN-IU-2011-1473), stn CP 3670, north of Rabaul, 04°06’S 151°56’E, 497–500 m, coll. N.O. Alis, BIOPAPUA cruise, 24 September 2010; 1 male (pcl 9.5 mm, pcw 7.6 mm) (MNHN-IU-2011-889), stn CP 3654, west of New Hanover, 02°14’S 150°16’E, 490–505 m, coll. N.O. Alis, BIOPAPUA cruise, 28 August 2010; 1 male (pcl 12.4 mm, pcw 9.0 mm) (ZRC 2018.1473), stn CP 4339, 06°10’S 149°18’E, 510–743 m, coll. MADEEP Expedition, 7 May 2014; 1 male (pcl 9.8 mm, pcw 6.7 mm), 2 females (pcl 11.5, pcw 8.4 mm; pcl 10.3 mm, pcw 7.3 mm) (ZRC 018.1474), stn CP 4444, 02°15’S 150°14’E, 417–421 m, coll. KAVIENG 2014, 1 September 2014. Remarks. Lee et al. (2013: 3, fig. 1A) identified a series of specimens from Papua New Guinea as G. takedai Richer de Forges & Ng, 2012, extending the range of the species to off Luzon in the Philippines. Together with the present material, we have re-examined all this material and now believe that they should all be referred to G. lappacea instead. The two species are close, but highly diagnostic of the latter taxon are the very long and inward curving hepatic spines, which reach well beyond the orbits (Rathbun 1918: 12, pl. 7 fig. 3; Richer de Forges 1994: fig. 3A–C). Griffinia lappacea has been reported from southern Australia (type locality) as well as the Northwest Shelf, Kai Islands (Indonesia) and Philippines (Serène & Vadon 1984; Richer de Forges 1994). Serène & Vadon (1984: 128, pl. 4F) had reported their specimen from northwest Mindoro in western Philippines as “ Pisidarum sp.” but Richer de Forges (1994: 67) noted that this should be G. lappacea instead. The specimens on hand show that the strength and length of the hepatic spine is very variable, being short in some specimens (Fig. 25C, E; Lee et al. 2013: fig. 1A) to distinctly longer (Fig. 25A, B, D). Comparisons with the type material of G. takedai shows several clear differences between the two species not observed before. Griffinia lappacea differs in almost always having a row of four strong tubercles on the inner part of the pterygostomial ridge, which are either distinct or basally fused to various degrees (Fig. 26A, B). In one small subadult female (pcl 7.5 mm, pcw 5.4 mm, MNHN-IU-2015-63), the left pterygostomial ridge had three tubercles but the right side had four tubercles, the first being small. This character is very consistent in all the specimens examined. In G. takedai, this pterygostomial ridge is defined by a row of four more elongate dentiform tubercles, with an additional median tubercle posterior to the row (Fig. 26C). In addition, we note that the hepatic spine of G. lappacea is always more anteriorly directed, even if short (Fig. 25A–E) whereas in G. takedai, the hepatic spine is always directed more obliquely laterally (Fig. 25F; Richer de Forges & Ng 2012: figs. 1, 2A). In G. lappacea, the posterior margin of the lobiform outer margin of the carpus of the male cheliped is almost always expanded and distinctly auriculiform (Fig. 25A–E); but in G. takedai, the structure is more symmetrical in shape and the posterior margin is rounded (Fig. 25F). When specimens of similar sizes are compared, the ambulatory meri of G. lappacea are also proportionately longer and more slender (Fig. 26D, E) compared to those of G. takedai (Fig. 26F; Richer de Forges & Ng 2012: fig. 3C). Significantly, the G1 of G. lappacea is also more sinuous in general shape, with the tapering distal part relatively longer (Fig. 26H–I); in G. takedai, the G1 is straighter with the distal part shorter (Fig. 26J–L; Richer de Forges & Ng 2012: fig. 4A, B)., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on page 34, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["Rathbun, M. J. (1918) Report on the spider crabs obtained by the F. I. S. \" Endeavour \" on the coasts of Queensland, New South Wales, Victoria, South Australia and Tasmania. Biological Results of the Fishing Experiments Carried On by the F. I. S. \" Endeavour \" 1909 - 14, 5 (1), 3 - 29, figs. 1 - 3, pls. 1 - 15. https: // doi. org / 10.5479 / si. 00963801.2135.527","Barnard, K. H. (1950) Descriptive Catalogue of South African Decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum, 38, 1 - 837, figs. 1 - 154.","Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vols. 1 - 3. Kodansha Ltd., Tokyo, pp. xxix + 773 pp. (English Txt), 461 pp. (Japanese Txt), 16 pp., 251 pls. (Pls. Vol.)","Griffin, D. J. G. & Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335.","Richer de Forges, B. (1994) A new genus of deep-sea majid crab: Griffinia gen. nov. (Crustacea, Decapoda, Brachyura). The Beagle, Records of the Museum and Art Gallery of the Northern Territory, 11, 65 - 72.","Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286.","Richer de Forges, B. & Ng, P. K. L. (2012) Griffinia takedai, a new species of deep sea majoid crab (Decapoda, Brachyura, Epialtidae) from the Philippines. In: Komatsu, H., Okuno, J. & Fukuoka, K. (Eds.), Studies on Eumalacostraca: a homage to Takeda Masatsune. Crustaceana Monographs, 17, 275 - 284. https: // doi. org / 10.1163 / 9789004202894 _ 023"]}

Genus Grypachaeus Alcock, 1895 Type species. Achaeus hyalinus Alcock & Anderson, 1894, by monotypy., Published as part of Ng, Peter K. L., Forges, Bertrand Richer De & Lee, Bee Yan, 2023, New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea, pp. 1-44 in Zootaxa 5318 (1) on page 30, DOI: 10.11646/zootaxa.5318.1.1, http://zenodo.org/record/8158079, {"references":["Alcock, A. (1895) Materials for a carcinological fauna of India. No. 1. The Brachyura Oxyrhyncha. Journal of the Asiatic Society of Bengal, 64 (2), 157 - 291, pls. 3 - 5. https: // doi. org / 10.5962 / bhl. title. 16033","Alcock, A. & Anderson, A. R. S. (1894) Natural History Notes from H. M. Indian Marine Survey Steamer ' Investigator, ' Commander C. F. Oldham, R. N., commanding. Series II, No. 17. List of the Shore and Shallow-water Brachyura collected during the Season 1893 - 1894. Journal of the Asiatic Society of Bengal, 63 (2 - Natural Science), 197 - 209."]}

Genus Cretalamoha gen. nov. Cretalamoha Nyborg & Garassino, 2017: 137 [unavailable]. Type species. Cretalamoha boweni sp. nov. Gender: feminine. Diagnosis. As for the type species. Etymology. After the Cretaceous, the geological age of the studied specimen, and Lamoha Ng, 1998, which shares some morphological characters with the new genus., Published as part of Nyborg, Torrey, Garassino, Alessandro & Vega, Francisco J., 2023, Validation of Cretalamoha boweni Nyborg & Garassino (Brachyura: Homolidae) from the early Campanian, Upper Cretaceous of British Columbia, Canada, pp. 148-150 in Zootaxa 5318 (1) on page 148, DOI: 10.11646/zootaxa.5318.1.8, http://zenodo.org/record/8158418, {"references":["Nyborg, T. & Garassino, A. (2017) New occurrences of fossil Homolidae from the Eastern Pacific. Boletin de la Sociedad Geologica Mexicana, 69 (1), 135 - 148. https: // doi. org / 10.18268 / bsgm 2017 v 69 n 1 a 6","Ng, P. K. L. (1998) Lamoha, a Replacement Name for Hypsophrys Wood Mason & Alcock, 1891 (Brachyura, Homolidae), a Junior Homonym of Hypsophrys Agassiz, 1859 (Pisces, Teleostei, Cichlidae). Crustaceana, 71, 121 - 125. https: // doi. org / 10.1163 / 156854098 x 00833"]}