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Summary: Fossil grass silica short cell phytoliths (GSSCP) have been used to reconstruct the biogeography of Poaceae, untangle crop domestication history and detect past vegetation shifts. These inferences depend on accurately identifying the clade to which the fossils belong. Patterns of GSSCP shape and size variation across the family have not been established and current classification methods are subjective or based on a 2D view that ignores important 3D shape variation.Focusing on Poaceae subfamilies Anomochlooideae, Pharoideae, Pueliodieae, Bambusoideae and Oryzoideae, we observed in situ GSSCP to establish their orientation and imaged isolated GSSCP using confocal microscopy to produce 3D models. 3D geometric morphometrics was used to analyze GSSCP shape and size. Classification models were applied to GSSCP from Eocene sediments from Nebraska, USA, and Anatolia, Turkey.There were significant shape differences between nearly all recognized GSSCP morphotypes and between clades with shared morphotypes. Most of the Eocene GSSCP were classified as woody bamboos with some distinctive Nebraska GSSCP classified as herbaceous bamboos.3D morphometrics hold great promise for GSSCP classification. It accounts for the complete GSSCP shape, automates size measurements and accommodates the complete range of morphotypes within a single analytical framework. [ABSTRACT FROM AUTHOR]

This paper examines the contribution of microscopic multi-proxy approaches to the study of early husbandry practices and animal diet by integrated micromorphological, phytolith, and calcitic dung spherulite analyses of midden deposits at the three neighbouring Neolithic sites of Boncuklu (9th–8th millennium cal BC), Pınarbaşı (7th millennium cal BC) and Çatalhöyük (8th–6th millennium cal BC) in the Konya Plain, Central Turkey. The results reveal considerable chronological and contextual variation in human-animal inter-relations in open areas between different communities and sites. At Boncuklu, middens display well-defined areas where phytoliths and substantial accumulations of omnivore faecal matter low in spherulite content have been identified. By contrast, open spaces at the Late Neolithic campsite of Pınarbaşı comprise large concentrations of herbivore dung material associated with neonatal ovicaprine remains from spring birthing. Here, the deposits represent repeated dung-burning events, and include high concentrations of dung spherulites and phytoliths from wild grasses, and leaves and culms of reeds that, we suggest here, derive from fodder and fuel sources. Late middens at Çatalhöyük are characterised by thick sequences derived from multiple fuel burning events and rich in ashes, charred plants, articulated phytoliths – mainly from the husk of cereals, as well as the leaves and stems of reeds and sedges – and omnivore/ruminant coprolites, the abundance of the latter declining markedly in the latest levels of occupation. The application of this integrated microscopic approach to open areas has contributed to unravelling the complexity of formation processes at these sites, providing new insights into herding practices, diet, and the ecological diversity of Neolithic communities in Central Anatolia. [ABSTRACT FROM AUTHOR]

An investigation of wheat husk phytoliths from a midden deposit at Neolithic Çatalhöyük provided the opportunity to investigate the impact of taphonomic processes on conjoined phytolith size. Wheat husk phytoliths from a possible crop processing deposit were examined using three methods. The results indicate that the size of conjoined forms decreases significantly as a result of laboratory extraction and slide preparation. Observations of the phytolith layer in thin section show some of the depositional and post-depositional processes affecting phytolith size. These results indicate that these taphonomic factors need further investigation before conjoined phytolith size can be used to infer past water availability and agricultural practices. [ABSTRACT FROM AUTHOR]

Abstract: Samples taken from middens at the Neolithic site of Çatalhöyük in Turkey have been analysed using IR spectroscopy backed up by powder XRD and SEM–EDX. Microcomponents studied include fossil hackberries (providing evidence of ancient diet and seasonality), mineral nodules (providing evidence of post-depositional change) and phytoliths (mineralised plant cells, providing evidence of usage of plant species). Finely laminated ashy deposits have also been investigated allowing chemical and mineralogical variations to be explored. It is found that many layers which appear visually to be quite distinctive have, in fact, very similar mineralogy. [Copyright &y& Elsevier]

Abstract: The arrival of hipparionine horses in the eastern Mediterranean region around 11 Ma was traditionally thought to mark the simultaneous westward expansion of savanna vegetation across Eurasia. However, recent paleoecological reconstructions based on tooth wear, carbon isotopes, and functional morphology indicate that grasses played a minor role in Late Miocene ecosystems of the eastern Mediterranean, which were more likely dry woodlands or forests. The scarcity of grass macrofossils and pollen in Miocene floras of Europe and Asia Minor has been used to support this interpretation. Based on the combined evidence, it has therefore been suggested that Late Miocene ungulate faunal change in the eastern Mediterranean signals increased aridity and landscape openness, but not necessarily the development of grass-dominated habitats. To shed new light on the Miocene evolution of eastern Mediterranean ecosystems, we used phytolith assemblages preserved in direct association with faunas as a proxy for paleovegetation structure (grassland vs. forest). We extracted phytoliths and other biogenic silica from sediment samples from well-known Early to Late Miocene (∼20–7 Ma) faunal localities in Greece, Turkey, and Iran. In addition, a Middle Eocene sample from Turkey yielded phytoliths and served as a baseline comparison for vegetation inference. Phytolith analysis showed that the Middle Eocene assemblage consists of abundant grass phytoliths (grass silica short cells) interpreted as deriving from bambusoid grasses, as well as diverse forest indicator phytoliths from dicotyledonous angiosperms and palms, pointing to the presence of a woodland or forest with abundant bamboos. In contrast, the Miocene assemblages are dominated by diverse silica short cells typical of pooid open-habitat grasses. Forest indicator phytoliths are also present, but are rare in the Late Miocene (9–7 Ma) assemblages. Our analysis of the Miocene grass community composition is consistent with evidence from stable carbon isotopes from paleosols and ungulate tooth enamel, showing that C4 grasses were rare in the Mediterranean throughout the Miocene. These data indicate that relatively open habitats had become common in Turkey and surrounding areas by at least the Early Miocene (∼20 Ma), >7 million years before hipparionine horses reached Europe and arid conditions ensued, as judged by faunal data. [Copyright &y& Elsevier]

• Animal dung is a valuable archaeological resource for reconstructing human activity. • An integrated geoarchaeological approach identifies new penning areas at Çatalhöyük. • Results suggest proximity to and management of herds preceding site expansion. • Foddering/grazing practices display highly variable regimes among herd animals. • Results show patterns of co-habitation of humans with animals, their food and dung. Over the last few decades a variety of geoarchaeological methods and ethnoarchaeological and experimental approaches have demonstrated the fundamental importance of animal dung deposits for reconstructing past human life-ways. Through simultaneous examination in micromorphological thin-section and integrated phytolith and faecal spherulite analyses, this study provides direct evidence for animal management and organisation of space at Neolithic Çatalhöyük by examining livestock penning deposits across the settlement. The identification of new extensive areas of penning distributed within the boundaries of the early occupation of the site suggests greater proximity to and management of herds immediately prior to a phase of settlement expansion, access to wider networks and resources, and increased exploitation of the wider landscape. Phytolith assemblages from in situ dung accumulations also provide new insights into foddering/grazing practices showing highly variable herbivorous regimes, including both dicotyledonous and grass-based diets with an important proportion of grasses used as fodder and/or grazing during the early occupation of Çatalhöyük. This study provides direct evidence of the proximity of humans and herds, continuity and change in animal management strategies and farming practices, and concepts of space at the site. [ABSTRACT FROM AUTHOR]