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The thirty-eight-year record (1984–2021) of glacier mass balance measurement indicates a significant glacier response to climate change in the North Cascades, Washington that has led to declining glacier runoff in the Nooksack Basin. Glacier runoff in the Nooksack Basin is a major source of streamflow during the summer low-flow season and mitigates both low flow and warm water temperatures; this is particularly true during summer heat waves. Synchronous observations of glacier ablation and stream discharge immediately below Sholes Glacier from 2013–2017, independently identify daily discharge during the ablation season. The identified ablation rate is applied to glaciers across the North Fork Nooksack watershed, providing daily glacier runoff discharge to the North Fork Nooksack River. This is compared to observed daily discharge and temperature data of the North Fork Nooksack River and the unglaciated South Fork Nooksack River from the USGS. The ameliorating role of glacier runoff on discharge and water temperature is examined during 24 late summer heat wave events from 2010–2021. The primary response to these events is increased discharge in the heavily glaciated North Fork, and increased stream temperature in the unglaciated South Fork. During the 24 heat events, the discharge increased an average of +24% (±17%) in the North Fork and decreased an average of 20% (±8%) in the South Fork. For water temperature the mean increase was 0.7 °C (±0.4 °C) in the North Fork and 2.1 °C (±1.2 °C) in the South Fork. For the North Fork glacier runoff production was equivalent to 34% of the total discharge during the 24 events. Ongoing climate change will likely cause further decreases in summer baseflow and summer baseflow, along with an increase in water temperature potentially exceeding tolerance levels of several Pacific salmonid species that would further stress this population. [ABSTRACT FROM AUTHOR]