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The evolution of the Apennines thrust-and-fold belt is related to heterogeneous process of subduction and continental delamination that generates extension within the mountain range and compression on the outer front of the Adria lithosphere. While normal faulting earthquakes diffusely occur along the mountain chain, the sparse and poor seismicity in the compressional front does not permit to resolve the ambiguity that still exists about which structure accommodates the few mm/yr of convergence observed by geodetic data. In this study, we illustrate the 2012 Emilia seismic sequence that is the most significant series of moderate-to-large earthquakes developed during the past decades on the compressional front of the Apennines. Accurately located aftershocks, along with P-wave and Vp/Vs tomographic models, clearly reveal the geometry of the thrust system, buried beneath the Quaternary sediments of the Po Valley. The seismic sequence ruptured two distinct adjacent thrust faults, whose different dip, steep or flat, accounts for the development of the arc-like shape of the compressional front. The first shock of May 20 (Mw 6.0) developed on the middle Ferrara thrust that has a southward dip of about 30°. The second shock of May 29 (Mw 5.8) ruptured the Mirandola thrust that we define as a steep dipping (50–60°) pre-existing (Permo-Triassic) basement normal fault inverted during compression. The overall geometry of the fault system is controlled by heterogeneity of the basement inherited from the older extension. We also observe that the rupture directivity during the two main-shocks and the aftershocks concentration correlate with low Poisson ratio volumes, probably indicating that portions of the fault have experienced intense micro-damage. [ABSTRACT FROM AUTHOR]