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Two new species of the genus Amaurobius are described from China: A. guangwushanensis sp. nov. (♂♀) from Sichuan and A. wulongdongensis sp. nov. (♂♀) from Shaanxi. With the addition of two new species, the number of Amaurobius species endemic to China now reaches four. Morphological descriptions, photos, and illustrations of copulatory organs, as well as a map of distribution records, are provided.

Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang, Zhang, Zhisheng (2021): Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics 46 (1): 16-74, DOI: 10.11865/zs.2021102

Genus Allotrochosina Roewer, 1960 Allotrochosina Roewer, 1960: 927. Type species: Lycosa schauinslandi Simon, 1899, from New Zealand. Remarks. This is the first record of this genus from China. Prior to this study, three species were known, all from Australia and New Zealand (WSC, 2020). Diagnostic characteristics include: lack of bristles on tip of the male palp; subtegulum elongated, situated along the prolateral margin of the cymbium; embolus and terminal apophysis reduced and crowded together at tip of bulb (Vink, 2001; Framenau, 2008)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 17, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Roewer, C. F. 1960. Araneae Lycosaeformia II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba, Mission G. F. de Witte, 55: 519 - 1040.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2.","Vink, C. J. 2001. A revision of the genus Allotrochosina Roewer (Araneae: Lycosidae). Invertebrate Taxonomy, 15 (4): 461 - 466.","Framenau, V. W. 2008. A new species in the wolf spider genus Allotrochosina from New South Wales, Australia (Araneae, Lycosidae). Journal of Arachnology, 35: 463 - 469."]}

Genus Piratula Roewer, 1960 Piratula Roewer, 1960: 677. Type species: Pirata hygrophilus Thorell, 1872, from Europe. Remarks. Currently, 27 species are known worldwide, of which 11 species are reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 55, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Roewer, C. F. 1960. Araneae Lycosaeformia II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba, Mission G. F. de Witte, 55: 519 - 1040.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Genus Lysania Thorell, 1890 Lysania Thorell, 1890a: 312. Type species: Lysania pygmaea Thorell, 1890, from Malaysia. Remarks. Currently, 4 species are known from Asia, of which 2 species are reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 40, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Thorell, T. 1890 a. Arachnidi di Pinang raccolti nel 1889 dai Signori L. Loria e L. Fea. Annali del Museo Civico di Storia Naturale di Genova, 30: 269 - 383.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Trochosa honggiana Barrion, Barrion-Dupo & Heong, 2012 (Figs 61A–C, 62A–B) Trochosa honggiana Barrion, Barrion-Dupo & Heong, 2012: 20, figs 21A–B (♀). Material examined. 1♀ (holotype), Haiko, Honggi Village, 9.IV.2011, A. T. Barrion, J.L.A. Catindig & S.C. Villareal leg. (HANU). Diagnosis. The female of this species is similar to T. ruricoloides Schenkel, 1963 (Figs 63G–H, 65C–D) but differs by the robust septum and the large spermathecal heads of the epigyne (Figs 61B–C, 62A–B). Description. Female (Fig. 61A) total length 21.97. Carapace 8.23 long, 6.21 wide; opisthosoma 12.29 long, 7.44 wide. Carapace brown. Eye region black. Cervical groove and radial furrows indistinct. Eye sizes and interdistances: AME 0.30, ALE 0.26, PME 0.75, PLE 0.61; AME–AME 0.21, AME–ALE 0.15, PME–PLE 0.69. Clypeus height 0.71. Chelicerae elongate, black-brown. Labium black-brown. Endites black-brown, longer than wide. Sternum black-brown and scutellate, with brown setae. Legs brown, with black pigmentation. Leg measurements: I 22.44 (6.58, 8.06, 4.68, 3.12); II 20.52 (5.86, 7.08, 4.31, 3.27); III 19.17 (5.15, 6.32, 4.68, 3.02); IV 27.38 (7.17, 8.94, 7.56, 3.71). Opisthosoma oval. Dorsum of opisthosoma grey-brown, cardiac mark indistinct. Ventrum black. Epigyne (Figs 61B–C, 62A–B). Septum broad, inverse T-shaped. Spermathecal heads large and round, spermathecal stalks wide, only constricted near spermathecal heads. Fertilization ducts short. Male unknown. Distribution. China (Hainan)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 61-64, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Schenkel, E. 1963. Ostasiatische Spinnen aus dem Museum d'Histoire naturelle de Paris. Memoires du Museum National d'Histoire Naturelle de Paris (A, Zool.), 25: 1 - 481."]}

Allotrochosina huangi Wang, Li & Zhang, sp. nov. (Figs 1A–B, 3A–I, 4A–D, 5A–E) Type material. Holotype ♂, Ledong County, Jianfengling Nature Reserve, Mingfeng Valley, 18°44.485′N, 108°50.812′E, elev. 960 m, 28.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Paratypes. 9♂ 13♀, same data as holotype (SWUC); 2♂ 4♀, Jianfengling Nature Reserve, Wufengqu, 18°44.415′N, 108°51.802′E, elev. 900 m, 18 May 2011, Y.Y. Zhou leg. (IZCAS). Etymology. The specific name comes from the family name of the collector, Guiqiang Huang; noun (name) in genitive case. Diagnosis. The new species resembles A. schauinslandi (Simon, 1899) (Vink, 2002: 18, figs 7, 34, 41, 68, 95; Framenau, 2008: 467, fig. 7) but differs by the crescent-shaped embolus (Figs 3C–G, 4A–D, 5A–C); the embolus is needle-shaped, and the embolic division has a small beak-shaped projection in A. schauinslandi. Females can be differentiated by the shape and orientation of the spermathecae (Figs 3H–I, 5D–E). Description. Male. Total length 2.66–2.91. Holotype (Fig. 3A) total length 2.85. Carapace 1.60 long, 1.11 wide; opisthosoma 1.24 long, 0.78 wide. Live specimens with a long stripe of white setae from the base of the median eyes to the base of the spinnerets. Carapace yellow-brown. Eye region black. Cervical groove distinct, radial furrows indistinct. Eye sizes and interdistances: AME 0.06, ALE 0.05, PME 0.23, PLE 0.17; AME–AME 0.05, AME–ALE 0.07, PME–PME 0.14, PME–PLE 0.16. Clypeus 0.08 high. Chelicerae yellow-brown, with 3 promarginal and 2 retromarginal teeth. Labium yellowish brown. Endites yellow-brown. Sternum yellowish brown and scutellate, with sparse, yellow-brown setae. Legs yellowish brown. Leg measurements: I 4.11 (1.03, 1.36, 1.08, 0.64); II 3.70 (1.04, 1.28, 0.87, 0.51); III 3.69 (1.02, 1.15, 1.01, 0.51); IV 5.20 (1.30, 1.66, 1.50, 0.74). Dorsum yellow-brown, with black pigmentation. Ventrum yellowish brown. Palp (Figs 3C–G, 4A–D, 5A–C). Terminal apophysis small, hook-shaped, covered by conductor. Median apophysis liver-shaped, slightly sclerotized. Embolus crescent-shaped. Conductor wide, membranous, tapering distally. Female (paratype, Fig. 3B) total length 5.37. Carapace 2.52 long, 1.70 wide; opisthosoma 2.85 long, 1.71 wide. Eye sizes and interdistances: AME 0.05, ALE 0.06, PME 0.26, PLE 0.20; AME–AME 0.05, AME–ALE 0.10, PME–PME 0.15, PME–PLE 0.21. Clypeus height 0.08. Leg measurements: I 4.97 (1.43, 1.70, 1.12, 0.72); II 4.66 (1.32, 1.56, 1.11, 0.67); III 4.61 (1.30, 1.53, 1.19, 0.59); IV 6.55 (1.74, 2.07, 1.85, 0.89). Leg formula: 4123. Opisthosoma oval. Dorsum of opisthosoma yellow-brown, with black pigmentation. Ventrum yellow-brown. Epigyne (Figs 3H–I, 5D–E). Spermathecal heads round. Spermathecal stalks V-shaped, joined at the base, separated from each other by two times the length of a fertilization duct. Fertilization ducts relatively large, crescent-shaped. Distribution. China (Hainan)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 17-21, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Vink, C. J. 2002. Lycosidae (Arachnida: Araneae). Fauna of New Zealand, 44: 1 - 94.","Framenau, V. W. 2008. A new species in the wolf spider genus Allotrochosina from New South Wales, Australia (Araneae, Lycosidae). Journal of Arachnology, 35: 463 - 469.","Tanaka, H. 1985. Descriptions of new species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, Tokyo, 33: 51 - 87."]}

Genus Hippasa Simon, 1885 Hippasa Simon, 1885: 31. Type species: Pirata agelenoides Simon, 1884, from India. Remarks. Currently 37 species are known worldwide, of which 3 species are reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 36, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Simon, E. 1885. Materiaux pour servir a la faune arachnologiques de l'Asie meridionale. I. Arachnides recueillis a Wagra-Karoor pres Gundacul, district de Bellary par M. M. Chaper. II. Arachnides recueillis a Ramnad, district de Madura par M. l'abbe Fabre. Bulletin de la Societe Zoologique de France, 10: 1 - 39.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Zoica hainan Wang, Li & Zhang, sp. nov. (Figs 72A–H, 73A–D, 74A–D) Type material. Holotype ♂, Haikou City, Dongzhaigang Mangrove Nature Reserve, 19°57.113′N, 110°33.983′E, elev. - 2 m, 20.VIII.2007, C.X. Wang leg. (IZCAS); Paratypes (7♂ 8♀). 4♂ 4♀, same data as holotype (IZCAS); 1♂ 3♀, Haikou City, Dongzhaigang Mangrove Nature Reserve, 19°57.113′N, 110°33.983′E, elev. - 2 m, 20.VIII.2007, S. Li leg. (IZCAS); 2♂ 1♀, Ding’an County, Tongren Village, 19°30.953′N, 110°18.904′E, elev. 58 m, 27. VI.2019, L.Y. Wang, T. Yuan, K. Yu & J.X. Zhao leg. (SWUC). Etymology. The specific epithet refers to the type locality; noun in apposition. Diagnosis. The new species resembles Z. unciformis Li, Wang & Zhang, 2013 (Li, Wang & Zhang, 2013: 30, figs 5A– E, 6A–H) but differs by the lateral apophysis with two small projections, the inner projection spinelike, the outer projection longer than wide with a sharp tip (Figs 72C–F, 73A–D, 74A–B); in Z. unciformis, the lateral apophysis is sinuous, slightly curved, with a small outgrowth. Terminal apophysis covers embolus in the new species (Figs 72C–F, 73A–D, 74A–B) but not in Z. unciformis. Spermathecae round in the new species (Figs 72G–H, 74C–D) but oval in Z. unciformis. Description. Male. Total length 0.99–1.39. Holotype (Fig. 72A) total length 1.05. Carapace 0.56 long, 0.40 wide; opisthosoma 0.50 long, 0.33 wide. Carapace uniformly yellow-brown. Eye region black. Eye sizes and interdistances: AME 0.08, ALE 0.08, PME 0.12, PLE 0.09; AME–AME 0.02, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.04. Clypeus height 0.07. Chelicerae elongate, yellowish brown, with three promarginal and two retromarginal teeth. Labium yellowish brown. Endites yellowish brown. Sternum yellowish brown, with sparse brown setae. Legs yellowish brown. Leg measurements: I 2.57 (0.79, 0.93, 0.39, 0.46); II 2.41 (0.72, 0.82, 0.47, 0.40); III 1.69 (0.50, 0.43, 0.31, 0.45); IV 3.20 (0.91, 1.06, 0.76, 0.47). Opisthosoma brown. Venter yellowish brown. Palp (Figs 72C–F, 73A–D, 74A–B). Lateral apophysis with two small projections, inner projection spinelike, outer projection longer than wide with sharp tip. Terminal apophysis leaf-shaped with pointed tip. Conductor wide, membranous, with a pointed tip. Embolus stout, covered by terminal apophysis. Female (paratype, Fig. 72B) total length 1.40. Carapace 0.68 long, 0.49 wide; opisthosoma 0.65 long, 0.49 wide. Eye sizes and interdistances: AME 0.07, ALE 0.08, PME 0.13, PLE 0.09; AME–AME 0.02, AME–ALE 0.02, PME–PME 0.05, PME–PLE 0.06. Clypeus height 0.05. Leg measurements: I 2.75 (0.84, 0.90, 0.55, 0.46); II 2.58 (0.78, 0.85, 0.53, 0.42); III 2.54 (0.79, 0.80, 0.54, 0.41); IV 3.16 (0.89, 1.13, 0.75, 0.39). Opisthosoma yellow-brown. Epigyne (Figs 72G–H, 74C–D). Copulatory openings located posteriorly, separated by the diameter of the opening. Spermathecal heads round. Spermathecal stalks wide. Fertilization ducts long and hooklike. Distribution. Known only from the type locality, Hainan Island, China. Funding This work was supported by the National Natural Science Foundation of China (31672278, 31702005), the Key Natural Science Foundation of Chongqing (cstc2019jcyj-zdxmX0006) and the Investigation Project of Basic Science and Technology (2018 FY100305) to Zhisheng Zhang and Luyu Wang. Acknowledgements The manuscript benefited greatly from comments by two anonymous referees. Sarah Crews checked the English. Theo Blick checked etymology. We thank Feng Zhang and Tianlang Lv for the loan of specimens and Guiqiang Huang for his assistance during field work and collection., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 70-72, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340

Wadicosa fidelis (O. Pickard-Cambridge, 1872) (Figs 66A–H, 67A–D, 68A–D) Lycosa fidelis O. Pickard-Cambridge, 1872: 319 (♂). For full list of publications and synonyms concerning this species, see WSC (2020). Description. See Kronestedt & Zyuzin (2009) for both sexes. Habitus, male palp and epigyne as in Figs 66A–H, 67A– D, 68A–D. Material examined. 4♂ 7♀, Dongfang City, Datian Town, Jubian Bridge, 19°08.451′N, 108°50.641′E, elev. 59 m, 28.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Distribution. South China, Mediterranean, West, Central, South, Southeast and East Asia., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 66-68, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Pardosa sumatrana (Thorell, 1890) (Figs 47A–I, 48A–F) Lycosa sumatrana Thorell, 1890b: 136 (♂ ♀). For full list of publications and synonyms concerning this species, see WSC (2020). Pardosa tieshinglii Barrion, Barrion-Dupo & Heong, 2012: 16, figs 17A–G (♂ ♀). syn. nov. Pardosa villarealae Barrion, Barrion-Dupo & Heong, 2012: 17, figs 18A–E (♂). syn. nov. Description. See Yin et al. (1997) for both sexes. Habitus, male palp and epigyne as in Figs 47A–I, 48A–F. Material examined. 1♂ (holotype of P. tieshinglii), Panja Town, Songtao Reservoir area, 9.IV.2011, A.T. Barrion, J.L. A Catindig & S.C. Villareal leg. (HANU); 1♂ (holotype of P. villarealae), Dapo Town, Dapo Village, 25.III.–9.IV.2011, A. T. Barrion, J.L. A Catindig & S.C. Villareal leg. (HANU); 1♂, Baisha County, Hongxin Village, 13. V.2011, C. Zhang leg. (SWUC); 1♂ 2♀, Baisha County, Yacha Farm, Ten Team, 19°16.481'N, 109°27.727'E, elev. 146 m, 24.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Distribution. South China, Bhutan, Bangladesh, India, Indonesia, Myanmar, Nepal, Philippines, Sri Lanka. Remarks. The palp structure and epigyne morphology leave no doubt that P. tieshinglii and P. villarealae are junior synonyms of P. sumatrana., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 50-52, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Thorell, T. 1890 b. Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2.","Thorell, T. 1891. Spindlar fran Nikobarerna och andra delar af sodra Asien. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 24 (2): 1 - 149."]}

Piratula piratoides (Bösenberg & Strand, 1906) (Figs 55A–H, 56A–D, 57A–D) Tarentula piratoides Bösenberg & Strand, 1906: 318, pl. 13, fig. 336 (♀). For full list of publications and synonyms concerning this species, see WSC (2020). Description. See Yin et al. (2012) for both sexes. Habitus, male palp and epigyne as in Figs 55A–H, 56A–D, 57A–D. Material examined. 1♂ 5♀, Dongfang City, Datian Town, Jubian Bridge, 19°08.451′N, 108°50.641′E, elev. 59 m, 28.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Distribution. China (north to Heilongjiang, south to Yunnan), Russia, Korea, Japan., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 57, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2.","Yin, C. M., Peng, X. J., Yan, H. M., Bao, Y. H., Xu, X., Tang, G., Zhou, Q. S., Liu, P. 2012. Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp."]}

Arctosa tangguoi Wang, Li & Zhang, sp. nov. (Figs 16A–E, 17A–D, 18A–B) Type material. Holotype ♂, Lingshui County, Diaoluo Mountain Nature Reserve, 18°41.445′N, 109°52.410′E, elev. 620 m, 16.IV.2009, G. Tang leg. (IZCAS). Etymology. The specific epithet is taken from the name of Mr. Guo Tang in honour of his great contribution to Chinese arachnology; noun (name) in genitive case. Diagnosis. The new species resembles A. hainan Wang, Li & Zhang, sp. nov. (Figs 10C–F, 11A–F, 12A–B) but differs by the sword-shaped median apophysis (Figs 16B–E, 17A–B, D, 18A–B), which is slightly curved with a hook-shaped tip in the latter (Figs 10C–F, 11A–D, F, 12A–B). Terminal apophysis axe-shaped in the new species (Figs 16B–E, 17A–C, 18A– B), knife-shaped in A. hainan Wang, Li & Zhang, sp. nov. (Figs 10C–F, 11A–E, 12A–B). Embolus crescent-shaped in the new species (Figs 16B–E, 17A–C, 18A–B), longer than wide with broad tip in the latter (Figs 10C–F, 11A–E, 12A–B). Description. Male holotype (Fig. 16A) total length 4.40. Carapace 2.55 long, 1.81 wide; opisthosoma 1.91 long, 1.39 wide. Carapace black-brown. Eye region black-brown. Fovea longitudinal. Cervical groove and radial furrows indistinct. Eye sizes and interdistances: AME 0.08, ALE 0.09, PME 0.21, PLE 0.16; AME–AME 0.06, AME–ALE 0.04, PME–PME 0.13, PME–PLE 0.16. Clypeus height 0.06. Chelicerae brown. Labium yellow-brown. Endites yellow-brown, longer than wide. Sternum yellow-brown, with sparse brown setae. Legs yellow-brown, with brown pigmentation. Leg measurements: I 5.23 (1.65, 1.92, 1.01, 0.65); II 5.14 (1.46, 1.76, 1.13, 0.79); III 4.97 (1.39, 1.55, 1.32, 0.71); IV 7.25 (1.87, 2.27, 2.06, 1.05). Dorsum of opisthosoma black-brown, with 5 pairs of white spots, cardiac mark distinct. Ventrum yellow-brown. Palp (Figs 16B–E, 17A–D, 18A–B). Median apophysis sword-shaped, with an anterior furrow. Terminal apophysis longer than wide, distally axe-shaped. Conductor small, membranous. Embolus crescent-shaped with an outgrowth medially. Female unknown. Distribution. Known only from the type locality, Hainan Island, China., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on pages 25-26, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340

Arctosa depectinata (Bösenberg & Strand, 1906) (Figs 1C–D, 8A–H, 9A–D) Tarentula depectinata Bösenberg & Strand, 1906: 314, pl. 7, fig. 332 (♂ ♀). For full list of publications and synonyms concerning this species, see WSC (2020). Description. See Wang, Marusik & Zhang (2012) for both sexes. Habitus, male palp and epigyne as in Figs 1C–D, 8A– H, 9A–D. Material examined. 3♂ 4♀, Dongfang City, Datian Town, Jubian Bridge, 19°08.451′N, 108°50.641′E, elev. 59 m, 28.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC- 2♂ 3♀, HANU- 1♂ 1♀). Distribution. China (Fujian, Guangdong, Hainan, Jiangxi, Shandong, Yunnan), Japan., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 24, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Genus Ocyale Audouin, 1826 Ocyale Audouin, 1826: 149. Type species: Ocyale atalanta Audouin, 1826, from North Africa. Remarks. Currently, 3 species are known in Asia, and 1 species is reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 42, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Audouin, V. 1826. Explication sommaire des planches d'arachnides de l'Egypte et de la Syrie. In: Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee fran ҫaise, Vol. 1 (4). C. L. F. Panckoucke, Paris. pp. 99 - 186.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Allotrochosina limu Wang, Li & Zhang, sp. nov. (Figs 6A–C, 7A–B) Type material. Holotype ♂, Qiongzhong County, Limu Mountain Nature Reserve, Zhufeng, 19°10.874′N, 109°45.322′E, elev. 940 m, 4.V.2011, Y.Y. Zhou leg. (IZCAS). Etymology. The specific epithet refers to the type locality; noun in apposition. Diagnosis. The new species resembles A. huangi Wang, Li & Zhang, sp. nov. (Figs 3C–G, 4A–D, 5A–C) but differs by the heart-shaped median apophysis (Figs 6B–C, 7A–B,); the median apophysis is liver-shaped in A. huangi (Figs 3C–F, 4A–B, D, 5A–C,). Embolus rodlike, pointed at the end in the new species (Figs 6B–C, 7A–B), crescent-shaped in A. huangi (Figs 3C–G, 4A–C, 5A–B). In retrolateral view, the terminal apophysis is horn-shaped in the new species (Figs 6B–C, 7A– B) but hook-shaped in A. huangi (Figs 3C–G, 4A–C, 5A–C). Description. Male holotype (Fig. 6A) total length 3.25. Carapace 1.78 long, 1.34 wide; opisthosoma 1.40 long, 1.00 wide. Carapace yellow-brown. Eye region black. Cervical groove and radial furrows indistinct. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.18, PLE 0.15; AME–AME 0.05, AME–ALE 0.03, PME–PME 0.11, PME–PLE 0.12. Clypeus height 0.06. Chelicerae yellow-brown. Labium yellowish brown. Endites yellowish brown. Sternum yellowish brown with sparse brown setae. Legs yellow-brown. Leg measurements: I 4.88 (1.33, 1.66, 1.17, 0.72); II 4.81 (1.39, 1.57, 1.14, 0.71); III 4.55 (1.32, 1.39, 1.18, 0.66); IV 6.07 (1.66, 1.84, 1.70, 0.87). Dorsum of opisthosoma yellow-brown, cardiac mark indistinct. Ventrum yellowish brown. Palp (Figs 6B–C, 7A–B). Terminal apophysis sclerotized, horn-shaped, with pointed end. Embolus long, rodlike, with pointed end. Conductor membranous, wider than long, tongue-shaped, with blunt end. Median apophysis sclerotized, heartshaped. Female unknown. Distribution. Known only from the type locality, Hainan Island, China.

Lycosa grahami Fox, 1935 (Figs 26A–F, 27A–D) Lycosa grahami Fox, 1935: 455, fig. 3 (♀). For full list of publications and synonyms concerning this species, see WSC (2020). Description. See Yin et al. (1997) for both sexes. Habitus, male palp and epigyne as in Figs 26A–F, 27A–D. Material examined. 1♂ 1♀, Haikou City, Xiuying District, near Overseas Chinese Middle School, 25.XI.2015, T. L. Lv leg. (SWUC). Distribution. China (Hainan, Sichuan, Yunnan)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 37, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Fox, I. 1935. Chinese spiders of the family Lycosidae. Journal of the Washington Academy of Sciences, 25: 451 - 456.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Wadicosa okinawensis (Tanaka, 1985) (Figs 2D–F, 69A–H, 70A–E, 71A–D) Pardosa okinawensis Tanaka, 1985: 78, figs 41–44 (♂ ♀). For full list of publications and synonyms concerning this species, see WSC (2020). Description. See Tanaka (1985) for both sexes. Habitus, male palp, and epigyne as in Figs 69A–H, 70A–E, 71A–D. Material examined. 4♂ 10♀, Haikou City, Qiongshan District, Shangdao Village, Nandu River, 19°58.208′N, 110°25.116′E, elev. - 6 m, 20.III.2015, L.Y. Wang & G.Q. Huang leg. (SWUC). Distribution. China (Hainan), Japan. Remarks.ThisisthefirstrecordofthisspeciesinChina., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 68, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Tanaka, H. 1985. Descriptions of new species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, Tokyo, 33: 51 - 87.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Genus Trochosa C.L. Koch, 1847 Trochosa C.L. Koch, 1847: 95. Type species: Aranea lupus ruricola De Geer, 1778, from Europe. Remarks. Currently, 92 species are known worldwide, of which 15 species are reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 57, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Koch, C. L. 1847. Die Arachniden. Nurnberg, Vierzehnter Band, pp. 89 - 210, Funfzehnter Band, pp. 1 - 136, Sechszehnter Band, pp. 1 - 80.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Genus Pirata Sundevall, 1833 Pirata Sundevall, 1833: 24. Type species: Araneus piraticus Clerck, 1757, from Sweden. Remarks. Currently, 52 species are known worldwide, of which 7 species are reported from China (WSC, 2020)., Published as part of Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang & Zhang, Zhisheng, 2021, Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae), pp. 16-74 in Zoological Systematics 46 (1) on page 53, DOI: 10.11865/zs.2021102, http://zenodo.org/record/5366340, {"references":["Sundevall, C. J. 1833. Conspectus Arachnidum. Londini Gothorum. pp. 1 - 39.","WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2."]}

Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng, Zhang, Zhisheng (2018): Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae). Zoological Systematics 43 (3): 317-333, DOI: 10.11865/zs.201827

Clubiona globosa Wang, Chen & Zhang ZS, sp. nov. (Figs 6A–E, 7A–I) Material examined. Holotype ♂, China, Guizhou, Fanjing Mountain Reserve, Macaohe (27°53.86′N, 108°44.51′E; elev. 694 m), 2.X.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. Paratypes. 1♂, Yamugou (27°47.353′N, 108°43.765′E; elev. 547 m), 1.X.2013, L.Y. Wang, D. Wang & X.K. Jiang leg.; 4♀, Heiwanhe (27°50.800′N, 108°46.294′E; elev. 588 m), 20– 21.VII.2014, Z.S. Zhang & L.Y. Wang leg.; 1♀, Heiwanhe, 11.X.2014, C. Jin & Z.Z. Gao leg.; 2♂ 1♀, Heiwanhe, 14.X.2014, C. Jin & Z.Z. Gao leg.; 1♀, Heiwanhe, 20.X.2014, C. Jin & Z.Z. Gao leg. Etymology. The specific name comes from the Latin word “ globosus ”, referring to the shape of the male pedipalpal bulb; adjective Diagnosis. The new species is similar to C. qiyunensis Xu, Yang & Song, 2003 (Wu & Zhang, 2014: 211, figs 13–23). Males differ from the latter by the strong femoral apophysis, longer than femoral width, and the tiny embolus (Figs 6A–C, 7C–E) (femoral apophysis shorter than femoral width, spine-like embolus of C. qiyunensis). Females differ by the oval copulatory opening (roundish in C. qiyunensis) (Figs 6D, 7F, 7H). Males of this new species are also similar to that of C. pollicaris Wu, Zheng & Zhang, 2015 (Wu, Zheng & Zhang, 2015: 20, figs 15–16, 19, 25–27), but differ from the latter by the femoral apophysis shorter than two times of femoral width (longer than two times of femoral width in C. pollicaris). Description. Male (Fig. 7A). Total length 4.02–5.14. Holotype total length 5.14. Carapace 2.17 long, 1.63 wide; abdomen 2.87 long, 1.48 wide. Carapace yellow brown. Eye sizes and interdistances: AME 0.09, ALE 0.14, PME 0.12, PLE 0.13; AME–AME 0.10, AME–ALE 0.08, PME–PME 0.25, PME–PLE 0.15, ALE–PLE 0.07. MOA 0.32 long, front width 0.30, back width 0.50. Clypeus height 0.05. Chelicerae yellow brown, with 6 promarginal and 4 retromarginal teeth. Gnathocoxae yellow brown, longer than wide. Labium yellow brown, longer than wide. Sternum yellowish brown, heartshaped, with sparse brown hairs. Pairs of ventral spines: Tibia I 2; Metatarsus I 1; Tibia II 2; Metatarsus II 1. Leg measurements: I 4.38 (1.28, 1.72, 0.86, 0.52); II 4.97 (1.48, 2.01, 0.93, 0.55); III 4.12 (1.19, 1.43, 1.03, 0.47); IV 5.92 (1.69, 2.03, 1.56, 0.64). Leg formula 4213. Abdomen oval. Both dorsum and venter yellow brown. Male pedipalp (Figs 6A–C, 7C–E). Femoral apophysis strong, 5 times longer than basal width. Patella with blunt and wide prolateral apophysis, its width about 1/2 of patellar diameter. RTA small and blunt, located distally. Bulb nearly spherical, widest proximally. Embolus arising from tegulum distally, very small. Sperm duct narrow. Female paratypes total length 5.23–7.09. One of them (Fig. 7B) total length 6.71. Carapace 2.38 long, 1.84 wide; abdomen 4.21 long, 2.42 wide. Eye sizes and interdistances: AME 0.09, ALE 0.14, PME 0.11, PLE 0.14; AME–AME 0.09, AME–ALE 0.06, PME–PME 0.28, PME–PLE 0.15, ALE–PLE 0.09. MOA 0.38 long, front width 0.30, back width 0.52. Clypeus height 0.07. Chelicerae with 4 promarginal and 6 retromarginal teeth. Leg measurements: I 5.11 (1.46, 2.04, 1.00, 0.61); II 5.86 (1.75, 2.33, 1.15, 0.63); III 4.93 (1.46, 1.69, 1.24, 0.54); IV 7.18 (2.02, 2.49, 2.00, 0.67). Leg formula 4213. Other characters as in male. Epigyne (Figs 6D–E, 7F–I). Copulatory opening small, located centrally. Copulatory ducts strongly twisted, connected with a pair of large, transparent, oval bursae and tube-like spermathecae. Fertilization ducts small. Distribution. Known only from the type locality, Guizhou, China. Comment. The new species belongs to the C. corticalis- group., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on pages 322-324, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468

Clubiona subapplanata Wang, Chen & Zhang ZS, sp. nov. (Figs 14A–D, 15A–F) Material examined. Holotype ♂, China, Guizhou, Fanjing Mountain Reserve, Jinding (27°54.49′N, 108°41.86′E; elev. 2214 m), 29.IX.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. Paratypes. 1♀, same data as holotype; 5♀, Jinding (27°54.54′N, 108°41.88′E; elev. 2289m), 23.VII.2013, L.Y. Wang, T. Lu & X.K. Jiang leg.; 2♂ 2♀, Mianxuling (27°54.54′N, 108°39.83′E; elev. 1974 m), 13.X.2014, C. Jin & Z.Z. Gao leg. Etymology. The specific name refers to its similarity of C. applanata Liu, Yan, Griswold & Ubick, 2007; adjective. Diagnosis. The new species is similar to C. applanata Liu, Yan, Griswold & Ubick, 2007 (Liu et al., 2007: 64, figs 1– 5), but differs from the latter by the blunt RTA, the wider than long VTA, the presence of LTA and the wider conductor (in ventral view) (Figs 14A–B, 15C–D) (RTA relatively peaked, VTA as long as wide, LTA absent and conductor narrow in C. applanata) and a tiny copulatory opening of epigyne (14C–D, 15E–F) (copulatory opening small in C. applanata). Description. Male (Fig. 15A). Holotype total length 5.15. Carapace 2.42 long, 1.75 wide; abdomen 2.79 long, 1.37 wide. Carapace yellowish brown. Eye sizes and interdistances: AME 0.12, ALE 0.15, PME 0.13, PLE 0.15; AME–AME 0.06, AME–ALE 0.04, PME–PME 0.21, PME–PLE 0.11, ALE–PLE 0.07. MOA 0.36 long, front width 0.28, back width 0.45. Clypeus height 0.07. Chelicerae yellow brown, with 4 promarginal and 4 retromarginal teeth. Pairs of ventral spines: Tibia I 3; Metatarsus I 1; Tibia II 3; Metatarsus II 1. Leg measurements: I 5.31 (1.50, 2.23, 1.00, 0.58); II 5.80 (1.84, 2.33, 1.06, 0.57); III 5.22 (1.55, 1.83, 1.26, 0.58); IV 7.38 (2.13, 2.54, 2.04, 0.67). Leg formula 4213. Abdomen yellowish brown. Male pedipalp (Figs 14A–B, 15C–D) without femoral and patellar apophysis. Tibia with 3 apophyses: RTA blunt, as long as wide, VTA wider than long and LTA short, much wider than long. Embolus located anteriorly, anterior part curved ventrally. Conductor relatively long, wide medially and thin anteriorly. Sperm duct distinct, U-shaped in ventral view. Female paratypes total length 5.06–6.17. One paratype (Fig. 15B) total length 5.60. Carapace 2.61 long, 1.81 wide; abdomen 3.07 long, 1.73 wide. Eye sizes and interdistances: AME 0.12, ALE 0.16, PME 0.13, PLE 0.15; AME–AME 0.08, AME–ALE 0.04, PME–PME 0.25, PME–PLE 0.14, ALE–PLE 0.08. MOA 0.39 long, front width 0.31, back width 0.50. Clypeus height 0.07. Leg measurements: I 5.01 (1.55, 2.05, 0.88, 0.53); II 5.34 (1.60, 2.23, 0.93, 0.58); III 4.74 (1.45, 1.54, 1.20, 0.55); IV 6.89 (1.98, 2.31, 1.88, 0.72). Leg formula 4213. Other characters as for male. Epigyne (Figs 14C–D, 15E–F) with a small copulatory opening anteriorly. Copulatory ducts short and curved, with a pair of large, transparent bursae posteriorly. Spermathecae spherical, meso-laterally located and separated from each other by at least two times of their diameter. Fertilization ducts short. Distribution. Known only from the type locality, Guizhou, China. Comment. The new species belongs to the C. corticalis- group.

Pristidia ramosa Yu, Sun & Zhang, 2012 (Figs 19A–C) Pristidia ramosa Yu et al., 2012: 45, figs 1–16 (♂ ♀). Material examined. 1♀, Heiwanhe (27°50.80′N, 108°46.29′E; elev. 588 m), 18.VII.2013, L.Y. Wang, T. Lu & X.K. Jiang leg.; 1♀, Heiwanhe, 24.VII.2013, L.Y. Wang, T. Lu & X.K. Jiang leg.; 1♀, Zhiwuyuan (27°50′2.42″N, 108°45′30.08″E; elev. 532 m), 26.VII.2013, L.Y. Wang, T. Lu & X.K. Jiang leg. Description. See Yu et al. (2012). Habitus as shown in Fig. 19A, epigyne as in Figs 19B–C. Distribution. China (Fujiang, Guizhou, Guangxi, Jiangxi). Funding This study was supported by the National Natural Science Foundation of China (31471974, 31672278) and Fundamental Research Funds for the Central Universities (XDJK 2017B003) to Zhisheng Zhang, and the Foundation of Biologic Resource and Environment Big Data ([2015] 4013) to Huiming Chen. Acknowledgements Thanks are greatly given to the editor and two anonymous reviewers. Early version of this manuscript benefited from the comments of Yuri Marusik (Institute for Biological Problems of the North, Magadan, Russia), Kiril Mikhailov (Moscow Lomonosov State University, Russia) and Charles Haddad (University of the Free State, South Africa) for their helpful suggestions. We thank Dong Wang, Xuankong Jiang, Tian Lu (SWUC) and Junxia Zhang, Chi Jin, Zhizhong Gao (MHBU) for their assistance during the field work., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on page 332, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468, {"references":["Yu, H., Sun, Z. X., Zhang, G. R. 2012. New taxonomic data on the sac spiders (Arachnida: Araneae: Clubionidae) from China, with description of a new species. Zootaxa, 3299: 44 - 60."]}

Clubiona pterogona Yang, Song & Zhu, 2003 (Figs 13A–C) Clubiona pterogona Yang et al., 2003: 8, figs 3A–C (♀). Material examined. 1♀, Mianxuling (27°54.626′N, 108°40.288′E; elev. 1997m), 25.VII.2013, L.Y. Wang, T. Lu & X.K. Jiang leg. Description. See Yang et al. (2003). Female habitus as shown in Fig. 13A. Epigyne as in Figs 13B–C. Male unknown. Distribution. Known only from Mt. Fanjing, Guizhou, China. Comments. Judging from the shape of epigyne this species may belong to the C. marmorata- group as defined by Mikhailov (1995). Three species described here on the basis of males belong to the C. corticalis -group and therefore cannot be conspecific with this species., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on page 327, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468, {"references":["Yang, J. Y., Song, D. X., Zhu, M. S. 2003. Three new species and a new discovery of male spider of the genus Clubiona from China (Araneae: Clubionidae). Acta Arachnologica Sinica, 12: 6 - 13.","Mikhailov, K. G. 1995. Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta, 4 (2): 33 - 48."]}

Clubiona huiming Wang, Zhang & Zhang, sp. nov. (Figs 8A–B, 9A–C) Material examined. Holotype ♂, China, Guizhou, Fanjing Mountain Reserve, Mianxuling (27°54.54′N, 108°39.83′E; elev. 1974 m), 30.IX.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. Paratype. 1♂, between Jinding and Jiandaoxia, 13.X.2014, C. Jin & Z.Z. Gao leg. Etymology. The specific name honours Dr. Huiming Chen, a researcher on biospeleology and local animal resource of Guizhou, China; noun. Diagnosis. Males of the new species are similar to C. subapplanata Wang, Chen & Zhang ZS, sp. nov. (Figs 14A–B, 15A, C–F), but differs from the latter by the VTA as long as wide, the long and slender embolus and the tapering and coiled conductor (Figs 8A–B, 9B–C) (VTA wider than long, embolus short and conductor reduced in C. subapplanata Wang, Chen & Zhang ZS, sp. nov.). Description. Male (Fig. 9A). Holotype total length 4.16. Carapace 2.10 long, 1.58 wide; abdomen 2.07 long, 1.27 wide. Carapace yellowish brown. Eye sizes and interdistances: AME 0.08, ALE 0.14, PME 0.12, PLE 0.14; AME–AME 0.07, AME–ALE 0.04, PME–PME 0.17, PME–PLE 0.10, ALE–PLE 0.05. MOA 0.31 long, front width 0.25, back width 0.40. Clypeus height 0.07. Chelicerae yellow brown, with 5 promarginal and 5 retromarginal teeth. Pairs of ventral spines: Tibia I 3; Metatarsus I 1; Tibia II 3; Metatarsus II 1. Leg measurements: I 4.92 (1.42, 1.98, 0.96, 0.56); II 5.35 (1.61, 2.13, 1.01, 0.60); III 4.57 (1.34, 1.62, 1.11, 0.50); IV 6.58 (1.96, 2.09, 1.94, 0.59). Leg formula 4213. Abdomen yellowish brown. Male pedipalp (Figs 8A–B, 9B–C). Femur without apophysis. Patella longer than wide. Tibia shorter than patella, with 3 apophyses. RTA almost conical, located distally. VTA almost as long as wide. LTA wide and short, located proximally. Embolus long and coiled, arising distally from tegulum. Conductor wide in mid-part and tapering toward its apex. Sperm duct U-shaped. Female. Unknown. Distribution. Known only from the type locality, Guizhou, China. Comment. The new species belongs to the C. corticalis -group., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on pages 324-325, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468

Clubiona fanjingshan Wang, Chen & Zhang ZS, sp. nov. (Figs 4A–B, 5A–C) Material examined. Holotype ♂, China, Guizhou, Fanjing Mountain Reserve, between Jinding and Huixiangping, 1.VIII.2001, J.X. Zhang & Z.S. Zhang leg. Paratype. 1♂, Mianxuling 27°54.54′N, 108°39.83′E; elev. 1974 m), 13.X.2014, C. Jin & Z.Z. Gao leg. Etymology. The specific name refers to the type locality (Mt. Fanjing is read as Fanjingshan in Chinese); noun. Diagnosis. Male of the new species is similar to that of C. subapplanata Wang, Chen & Zhang ZS, sp. nov. (Figs 14A– B, 15A, C–D), but can be distinguished by the wider than long RTA, the absence of VTA, the wide and arc-shaped conductor, and the slightly curved embolus (Figs 4A–B, 5B–C) (as long as wide RTA, narrow apex of conductor and strongly curved embolus in C. subapplanata Wang, Chen & Zhang ZS, sp. nov.). Description. Male (Fig. 5A). Holotype total length 6.13. Carapace 2.67 long, 1.93 wide; abdomen 3.45 long, 1.68 wide. Carapace yellowish brown. Eye sizes and interdistances: AME 0.11, ALE 0.14, PME 0.12, PLE 0.13; AME–AME 0.12, AME–ALE 0.05, PME–PME 0.24, PME–PLE 0.16, ALE–PLE 0.08. MOA 0.38 long, front width 0.35, back width 0.49. Clypeus height 0.09. Chelicerae yellow brown, with 4 promarginal and 5 retromarginal teeth. Pairs of ventral spines: Tibia I 3; Metatarsus I 1; Tibia II 3; Metatarsus II 1. Leg measurements: I 7.43 (2.13, 2.90, 1.50, 0.90); II 7.86 (2.20, 3.03, 1.77, 0.86); III 6.71 (1.93, 2.20, 1.73, 0.85); IV 8.99 (2.50, 3.01, 2.63, 0.85). Leg formula 4213. Abdomen yellowish brown. Male pedipalp (Figs 4A–B, 5B–C). Femur without apophysis. Tibia with 2 apophyses, RTA wide and short, LTA small, close to the base of tibia, proximad. Conductor wide, transversal, arc or band-shaped, prolaterad, tip distad. Embolus located distally, with sharply pointed, ventrad tip. Sperm duct long, forming a circular loop in proximal half, gradually tapering toward the base of embolus. Female. Unknown. Distribution. Known only from the type locality, Guizhou, China. Comment. The new species belongs to the C. corticalis -group., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on pages 320-322, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468

Clubiona dichotoma Wang, Chen & Zhang ZS, sp. nov. (Figs 2A–D, 3A–G) Material examined. Holotype ♂, China, Guizhou, Fanjing Mountain Reserve, Maxi’ao (28°01.16′N, 108°45.41′E; elev. 1239 m), 11.X.2014, C. Jin & Z.Z. Gao leg. Paratypes. 3♀, same data as holotype; 1♂, Mianxuling (27°54.54′N, 108°39.83′E; elev. 1974 m), 13.X.2014, C. Jin & Z.Z. Gao leg.; 1♀, between Jinding and Huixiangping, 1.VIII.2001, J.X. Zhang & Z.S. Zhang leg.; 1♀, Maxi’ao (28°01.16′N, 108°45.41′E; elev. 1239 m), 5.X.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. Etymology. The specific name comes from the Greek word “ dichotomus ” and refers to its bifid embolus; adjective. Diagnosis. The new species is similar to C. subapplanata Wang, Chen & Zhang ZS, sp. nov. (Figs 14A–D, 15A–F), but differs by the bifid embolus, the stronger conductor and the presence of a finger-like apophysis on male pedipalpal femur (Figs 2A–B, 3C–E) (thinner and not bifid embolus, thinner conductor and absent femoral apophysis in C. subapplanata Wang, Chen & Zhang ZS, sp. nov.), and the slit-like copulatory opening of epigyne (Figs 2C–D, 3F–G) (hole-like copulatory opening in C. subapplanata Wang, Chen & Zhang ZS, sp. nov.). Description. Male total length 4.96–5.06. Holotype (Fig. 3A) total length 5.06. Carapace 2.07 long, 1.50 wide; abdomen 2.83 long, 1.33 wide. Carapace yellowish brown. Eye sizes and interdistances: AME 0.09, ALE 0.14, PME 0.13, PLE 0.13; AME–AME 0.04, AME–ALE 0.02, PME–PME 0.15, PME–PLE 0.07, ALE–PLE 0.06. MOA 0.33 long, front width 0.22, back width 0.38. Clypeus height 0.06. Chelicerae yellow brown, with 5 promarginal and 4 retromarginal teeth. Pairs of ventral spines: Tibia I 3; Metatarsus I 1; Tibia II 3; Metatarsus II 1. Leg measurements: I 4.69 (1.33, 1.91, 0.90, 0.55); II 4.99 (1.43, 2.05, 0.97, 0.54); III 4.26 (1.26, 1.47, 1.03, 0.50); IV 6.07 (1.71, 2.04, 1.73, 0.59). Leg formula 4213. Abdomen yellowish brown. Male pedipalp (Figs 2A–B, 3C–E). Femur retrolaterally with a slanting finger-like apophysis bearing a spine, apophysis as long as wide, half-length of femur diameter. Tibia with 3 apophyses: retrolateral apophysis short, almost triangular; ventral apophysis short and blunt; lateral apophysis small, slightly peaked. Conductor strong, arising from the tegulum mesoretrolaterally and extending disto-prolaterally, widest in the mid part. Embolus bifurcated, arising from distal tegulum, tip ventrad. Sperm duct obvious, curved like a fishhook in ventral view. Female paratypes total length 4.40–5.73. One of them (between Jinding and Huixiangping, Fig. 3B) total length 5.73. Carapace 2.48 long, 1.73 wide; abdomen 3.46 long, 1.74 wide. Eye sizes and interdistances: AME 0.10, ALE 0.12, PME 0.11, PLE 0.12; AME–AME 0.07, AME–ALE 0.06, PME–PME 0.28, PME–PLE 0.15, ALE–PLE 0.10. MOA 0.33 long, front width 0.27, back width 0.49. Clypeus height 0.06. Chelicerae with 6 promarginal and 4 retromarginal teeth. Leg measurements: I 4.85 (1.45, 1.95, 0.95, 0.50); II 5.22 (1.56, 2.13, 0.99, 0.54); III 4.70 (1.35, 1.69, 1.15, 0.51); IV 6.86 (1.95, 2.42, 1.87, 0.62). Leg formula 4213. Other characters as for male. Epigyne (Figs 2C–D, 3F–G). Copulatory openings small, located anteriorly. Copulatory ducts short and curved, connecting with a pair of large, transparent bursae posteriorly. Spermathecae small, spherical, about half of the diameter of bursae, located meso-laterally, separated by more than one of their own diameter. Fertilization ducts thin and short. Distribution. Known only from the type locality, Mt. Fanjing, Guizhou, China. Comment. The species belongs to the C. corticalis -group., Published as part of Wang, Luyu, Chen, Huiming, Wu, Panlong, Zhang, Feng & Zhang, Zhisheng, 2018, Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae), pp. 317-333 in Zoological Systematics 43 (3) on pages 319-320, DOI: 10.11865/zs.201827, http://zenodo.org/record/5366468

Fu, Lina, Zhang, Zhisheng, Zhang, Feng (2016): Description Of Two New Otacilia Species From Anhui, China (Araneae, Phrurolithidae). Acta Zoologica Academiae Scientiarum Hungaricae 62 (2): 133-142, DOI: 10.17109/AZH.62.2.133.2016, URL: http://dx.doi.org/10.17109/azh.62.2.133.2016

Fu, Lina, Zhang, Zhisheng, Zhang, Feng (2016): New Otacilia species from Southwest China (Araneae: Phrurolithidae). Zootaxa 4107 (2): 197-221, DOI: http://doi.org/10.11646/zootaxa.4107.2.4

Otacilia aurita sp. nov. Figs 7 A���G, 8 A���E Type material. Holotype ♂, China, Sichuan Province, Miyi County, Baipo Town, 27 �� 7.135 ���N, 101 �� 54.896 ���E, 13 May 2015, Zhizhong Gao leg. Paratypes: 4 ♂, 1 ♀, same data as holotype. Etymology. The species name is derived from the Latin word ��� auritus ���, and refers to the ear-shaped bursae; adjective. Diagnosis. The new species is similar to O. pseudostella (Fu et al. 2014: 488, figs 3 A���G, 4 A���E), but can be distinguished from the latter species by: 1) a relatively short E (longer in O. pseudostella); 2) PEA sharp (relatively blunt in O. pseudostella); 3) DTA long, spine-like (shorter, with a wide base and a sharp tip in O. pseudostella); 4) BU ear-shaped (pear-like in O. pseudostella); 5) CO located anteriorly, concealing the margin of the concavity (CO more posterior, located in the center of the concavity in O. pseudostella); 6) short CD (longer in O. pseudostella). The new species is also similar to O. digitata sp. nov. (Fig. 9), but can be distinguished from the latter species by: 1) palp with two tibial apophyses (only one in O. digitata sp. nov.); 2) PEA long, spine-like (relatively shorter, triangular in O. digitata sp. nov.); 3) TA antero-retrolaterally located, small (larger in O. digitata sp. nov.). Description. Male (Fig. 7 A). Total length 2.25���3.06 (n= 7). Holotype: body 2.87 long; carapace 1.31 long, 1.12 wide; abdomen 1.53 long, 0.92 wide. Carapace dark brown. Eyes sizes and interdistances: AME 0.06, ALE 0.09, PME 0.06, PLE 0.07; AME���AME 0.05, AME���ALE 0.01, PME���PME 0.06, PME���PLE 0.05. MOA 0.20 long, front 0.17 wide, back 0.18 wide. Clypeus 0.08 high. Cheliceral promargin with three well-separated teeth and retromargin with two teeth close to each other. Labium and gnathocoxae brown. Legs brown. Leg measurements: leg I 4.70 (1.27, 0.36, 1.29, 1.07, 0.71), II 3.89 (1.12, 0.29, 0.95, 0.92, 0.61), III 3.39 (0.91, 0.32, 0.72, 0.87, 0.57), IV 4.95 (1.33, 0.42, 1.13, 1.29, 0.78). Abdomen dark grey, anterior half with large dorsal scutum, nearly occupying nearly 4 / 5 of abdomen length, posterior half without distinct chevron-shaped markings dorsally. Venter yellow brown. Palp (Figs 7 E���G, 8 A���C): DTA long, sharp distally, spine-like. RTA short. E sharp apically, with membranous base ventrally and spine-like PEA prolaterally. SD sclerotized. TA antero-retrolaterally located, small. Female (Fig. 7 B). Total length 3.12 (n= 1). One paratype: body 3.12 long; carapace 1.26 long, 1.14 wide; abdomen 1.82 long, 1.19 wide. Carapace grey, abruptly narrow anteriorly; fovea longitudinal. Eyes sizes and interdistances: AME 0.07, ALE 0.09, PME 0.06, PLE 0.07; AME���AME 0.05, AME���ALE 0.01, PME���PME 0.07, PME���PLE 0.06. MOA 0.20 long, front 0.18 wide, back 0.19 wide. Clypeus 0.07 high. Leg measurements: I 3.83 (1.10, 0.31, 0.98, 0.88, 0.56); II 3.61 (0.96, 0.33, 0.88, 0.83, 0.61); III 3.16 (0.83, 0.29, 0.68, 0.79, 0.57); IV 4.84 (1.27, 0.42, 1.16, 1.33, 0.66). Abdomen dark grey, without dorsal scutum or chevron-shaped markings dorsally. Epigyne (Figs 7 C���D, 8 D���E): Two small CO located separately in pair of shallow concavities posteriorly (Figs 7 C, 8 D). CD thin and short, connected to two transparent, ear-shaped BU. SP oval, located posteriorly. FD arising anteriorly from SP. Distribution. Known only from the type locality, Sichuan, China (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on pages 207-209, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Fu, L. N., Jin, C. & Zhang, F. (2014) Three new species of the genus Otacilia Thorell (Araneae: Phrurolithidae) from China. Zootaxa, 3869 (4), 483 - 492. http: // dx. doi. org / 10.11646 / zootaxa. 3869.4.10"]}

Otacilia papilion sp. nov. Figs 5 A–I, 6 A–E Type material. Holotype ♂, China, Guizhou City, Leishan County, Leigongshan Natural Reserve, Xiannv Lake, 26 ° 22.951 ′N, 108 ° 12.437 ′E, 22 October 2012, Luyu Wang & Xuankong Jiang leg. Paratypes: 4 ♂, 6 ♀, same data as holotype; 5 ♂, 3 ♀, Guizhou City, Leishan County, Leigongshan Natural Reserve, Xiannv Lake, 26 ° 22.951 ′N, 108 ° 12.437 ′E, 21 October 2012, Luyu Wang & Xiangwei Meng leg. Etymology. The species name is derived from the Latin word ‘ papilion ’, and refers to the butterfly-shaped epigyne; noun. Diagnosis. The new species is similar to O. komurai (Kamura 2005: 90, figs 7–11), but can be distinguished by: 1) sickle-shaped E (hook-shaped E in O. komurai); 2) sclerotized “bubbles” interpreted to be remnants of fused furrows near the FD (absent in O. komurai); 3) epigastric furrow with a bulge in the new species (absent in O. komurai); 4) MP absent (present in O. komurai). Description. Male (Fig. 5 A). Total length 2.44–2.82 (n= 11). Holotype: body 2.69 long; carapace 1.34 long, 1.17 wide; abdomen 1.37 long, 0.90 wide. Carapace dark brown. Eyes sizes and interdistances: AME 0.07, ALE 0.09, PME 0.07, PLE 0.08; AME–AME 0.04, AME–ALE 0.02, PME–PME 0.12, PME–PLE 0.05. MOA 0.22 long, front 0.18 wide, back 0.27 wide. Clypeus 0.10 high. Cheliceral promargin with three well-separated teeth, retromargin with six teeth close to each other. Labium and gnathocoxae yellow. Legs light yellow. Leg measurements: leg I 4.57 (1.16, 0.32, 1.42, 1.13, 0.54), II 3.76 (1.04, 0.33, 1.05, 0.86, 0.48), III 3.21 (0.91, 0.29, 0.64, 0.88, 0.49), IV 4.85 (1.29, 0.38, 1.07, 1.37, 0.74). Abdomen dark grey, anterior half with a grey, dorsal scutum, occupying nearly 1 / 5 of abdomen length; posterior half with five bright chevron-shaped markings dorsally. Venter yellowish brown. Palp (Figs 5 E–G, 6 A–C): RTA with wide base extending dorsally, tip finger-shaped. E short, sickle-shaped, tip pointed, with a membranous part retrolaterally. SD relatively thick, sclerotized. TA small, with sharp tip, originating antero-retrolaterally. Tegulum widest in middle part, its posterior margin almost reaching ventral apex of tibia retrolaterally. Female (Fig. 5 B). Total length 2.67–3.26 (n= 8). One paratype: body 2.98 long; carapace 1.39 long, 1.29 wide; abdomen 1.63 long, 1.08 wide. Carapace dark brown anteriorly. Eyes sizes and interdistances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.08; AME–AME 0.04, AME–ALE 0.01, PME–PME 0.11, PME–PLE 0.05. MOA 0.24 long, front 0.21 wide, back 0.28 wide. Clypeus 0.10 high. Leg measurements: I 5.00 (1.28, 0.36, 1.58, 1.23, 0.55); II 4.33 (1.14, 0.38, 1.17, 1.08, 0.56); III 3.49 (0.89, 0.32, 0.76, 0.98, 0.54); IV 5.24 (1.36, 0.38, 1.22, 1.49, 0.79). Abdomen dark grey, without dorsal scutum dorsally, markings as in male. Epigyne (Figs 5 C–D, 6 D–E): CO small, located in oval concavity. CD thick, anteriorly connected to two large transparent BU, with sclerotized “bubbles” interpreted as remnants of fused furrows. SP oval, located posteriorly, close to each other. FD originating anteriorly from SP. Epigyne butterfly-shaped ventrally. Distribution. Known only from the type locality, Guizhou, China (Fig. 18).

Otacilia Thorell, 1897 Diagnosis. Otacilia can be recognized by the following combination of characters: carapace oval, abruptly narrow anteriorly; labium and gnathocoxae wider than long; chelicerae each with two bristles (rarely with one bristle) on anterior side; leg formula: 4123; spination (see Table 1 for new species in this paper): femora I���IV d 1, I pl 0���4, II pl 0���3; tibiae I���II and metatarsi I���II each with several pairs of long ventral spines (few with one more rv than pv); male abdomen with a dorsal scutum on antero-dorsal part, female abdomen without scutum. Male palpal femur with ventral apophysis; tibia usually with single strong RTA, seldom with BTA or DTA; embolus originating antero-prolaterally, with membranous or weakly sclerotized tegular apophysis (or conductor); epigyne with or without median plate, vulva usually with pair of transparent bursae anteriorly and pair of strongly sclerotized spermathecae. Species groups of Chinese Otacilia. After reviewing 30 Chinese Otacilia species (including the nine new species described in this paper) and the type species, but not including O. paracymbium J��ger & Wunderlich, 2012, whose female is unknown, we confirm that Wang et al. ���s (2015) assignment of species to the two groups is accurate. Here we add a third species group for the genus Otacilia and outline and formally name the three species groups, also list the male and female diagnostic characters for each species group, followed by a list of all of the included Chinese species (Table 2)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on page 198, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Thorell, T. (1897) Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni birmani. I. Parallelodontes. Tubitelariae. Annali del Museo Civico di Storia Naturali di Genova, Series 2, 17, 161 - 267.","Jager, P. & Wunderlich, J. (2012) Seven new species of the spider genus Otacilia (Araneae: Corinnidae) from China, Laos and Thailand. Beitrage zur Araneologie, 7, 251 - 271.","Wang, L. Y., Chen, H. M., Zhou, K. X., Zhang, F. & Zhang, Z. S. (2015) Diversity of spiders in Fanjing Mountain Nature Reserve, Guizhou, China, I: Six new species of Phrurolithidae (Araneae). Zootaxa, 4012 (3), 447 - 464. http: // dx. doi. org / 10.11646 / zootaxa. 4012.3.2"]}

Otacilia digitata sp. nov. Figs 9 A���G Type material. Holotype ♂, China, Sichuan Province, Guangyuan City, Qingchuan County, Sanguo Town, Xiyanggou Natural Reserve, 32 �� 31.754 ���N, 105 �� 1.078 ���E, 4 May 2013, Xiangwei Meng leg. Etymology. The specific name is from a Latin word ��� digitatus ���, and refers to the finger-shaped RTA; adjective. Diagnosis. The new species is similar to O. pseudostella (Fu et al. 2014: 488, figs 3 A���G, 4 A���E), but can be distinguished by: 1) PEA short and sharp (longer and blunt in O. pseudostella); 2) an E that is relatively short (longer in O. pseudostella); 3) BTA absent (present in O. pseudostella); 4) a finger-shaped RTA (with a wide base and sharp tip in O. pseudostella). The new species is also similar to O. zhangi (Fu et al. 2014: 490, fig 5 A���F), but can be distinguished from it by: 1) PEA short and sharp (longer and blunt in O. zhangi); 2) a relatively short E (longer in O. zhangi); 3) a finger-shaped RTA (with a wide base and sharp tip in O. zhangi). Female unknown. Description. Male (Fig. 9 A). Total length 3.02 (n= 1). Holotype: body 3.02 long; carapace 1.44 long, 1.19 wide; abdomen 1.55 long, 1.05 wide. Carapace brown. Eyes sizes and interdistances: AME 0.07, ALE 0.09, PME 0.06, PLE 0.07; AME���AME 0.05, AME���ALE 0.02, PME���PME 0.08, PME���PLE 0.07. MOA 0.21 long, front 0.18 wide, back 0.24 wide. Clypeus 0.07 high. Cheliceral promargin with three well-separated teeth and retromargin with two teeth close to each other. Labium and gnathocoxae brown. Legs light yellow. Leg measurements: leg I 4.81 (1.34, 0.36, 1.37, 1.10, 0.64), II 4.08 (1.15, 0.31, 1.04, 0.96, 0.62), III 3.62 (1.02, 0.34, 0.73, 0.94, 0.59), IV 5.20 (1.37, 0.42, 1.28, 1.35, 0.78). Abdomen yellow brown, dorsal scutum covering whole dorsum, posterior half with three chevron-shaped markings dorsally. Venter yellow brown. Palp (Figs 9 B���G): With only one RTA, finger-shaped. E thin apically, prolaterally with thick, sharp PEA. SD sclerotized. TA originating antero-retrolaterally. Tegulum widest in middle part. Distribution. Known only from the type locality, Sichuan, China (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on page 209, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Fu, L. N., Jin, C. & Zhang, F. (2014) Three new species of the genus Otacilia Thorell (Araneae: Phrurolithidae) from China. Zootaxa, 3869 (4), 483 - 492. http: // dx. doi. org / 10.11646 / zootaxa. 3869.4.10"]}

Otacilia leibo sp. nov. Figs 10 A���G, 11 A���E Type material. Holotype ♂, China, Sichuan Province, Leibo County, Ma Lake, 28 �� 25.536 ���N, 103 �� 47.115 ���E, 19 May 2012, Luyu Wang leg. Paratypes: 5 ♂, 12 ♀, same data as for holotype, Luyu Wang & Mingxin Liu leg. Etymology. The specific name is a noun in apposition of the type locality. Diagnosis. The new species is similar to O. pseudostella (Fu et al. 2014: 488, figs 3 A���G, 4 A���E), but can be distinguished from it by: 1) PEA sharp, pointed in ten o���clock direction (blunt, pointed to twelve o���clock in O. pseudostella); 2) a narrow tegulum (broad in O. pseudostella); 3) CO large (smaller and concealed by the margin of a pair of shallow concavities in O. pseudostella). The male of the new species is also similar to O. zhangi (Fu et al. 2014: 490, fig 5 A���F), but can be distinguished from it by: 1) PEA pointed to ten o���clock (pointed to twelve o���clock in O. zhangi); 2) the palp with a RTA and a BTA (with only an RTA in O. zhangi). The new species is similar to O. aurita sp. nov. (Figs 10���11), but can be distinguished from it by: 1) PEA pointed to ten o���clock (pointed to twelve o���clock in O. aurita sp. nov.); 2) E long (shorter, pointed to twelve o���clock in O. aurita sp. nov.); 3) CO anterior (relatively posterior in O. aurita sp. nov.); 4) CD long (shorter in O. aurita sp. nov.). Description. Male (Fig. 10 A). Total length 2.87���3.46 (n= 6). Holotype: body 3.40 long; carapace 1.59 long, 1.34 wide; abdomen 1.69 long, 1.12 wide. Carapace yellowish brown. Eyes sizes and interdistances: AME 0.08, ALE 0.10, PME 0.08, PLE 0.07; AME���AME 0.05, AME���ALE 0.02, PME���PME 0.10, PME���PLE 0.07. MOA 0.23 long, front 0.21 wide, back 0.25 wide. Clypeus 0.11 high. Cheliceral promargin with three well-separated teeth, retromargin with two teeth close to each other. Labium and gnathocoxae brown. Legs light yellowish. Leg measurements: leg I 5.39 (1.46, 0.37, 1.54, 1.27, 0.75), II 4.27 (1.15, 0.33, 1.10, 0.98, 0.71), III 3.68 (1.06, 0.34, 0.68, 0.96, 0.64), IV 5.64 (1.54, 0.47, 1.28, 1.49, 0.86). Abdomen yellow brown, anterior half with large dorsal scutum, nearly occupying nearly 4 / 5 of abdomen length; posterior half dark grey, without distinct chevron-shaped markings dorsally. Venter yellow brown. Palp (Figs 10 E���G, 11 A���C): BTA short, RTA with wide base and sharp tip. E long, originating at about 9 o'clock, thin apically, prolaterally with a sharp PEA, retrolaterally with membranous TA near E base. Tegulum widest in middle part. Female (Fig. 10 B). Total length 3.87���4.79 (n= 10). One paratype: body 4.47 long; carapace 1.77 long, 1.56 wide; abdomen 2.57 long, 1.68 wide. Carapace yellow brown. Eyes sizes and interdistances: AME 0.07, ALE 0.10, PME 0.09, PLE 0.08; AME���AME 0.05, AME���ALE 0.03, PME���PME 0.09, PME���PLE 0.06. MOA 0.24 long, front 0.21 wide, back 0.26 wide. Clypeus 0.12 high. Leg measurements: I 6.24 (1.74, 0.46, 1.82, 1.51, 0.71); II 5.22 (1.41, 0.42, 1.31, 1.25, 0.83); III 4.53 (1.32, 0.38, 1.02, 1.09, 0.72); IV 6.69 (1.77, 0.51, 1.58, 1.85, 0.98). Abdomen dark grey dorsally, without dorsal scutum and chevron-shaped markings dorsally. Epigyne (Figs 10 C���D, 11 D���E): CO large, widely separated, by distance slightly larger than their diameter, situated anteriorly. CD thin and long, connected to two transparent BU. SP small, separated by distance equal to their diameter, located posteriorly. FD arising anteriorly from SP. Distribution. Known only from the type locality, Sichuan, China (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on pages 209-212, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Fu, L. N., Jin, C. & Zhang, F. (2014) Three new species of the genus Otacilia Thorell (Araneae: Phrurolithidae) from China. Zootaxa, 3869 (4), 483 - 492. http: // dx. doi. org / 10.11646 / zootaxa. 3869.4.10"]}

Otacilia ovata sp. nov. Figs 12 A���H, 13 A���E Type material. Holotype ♂, China, Sichuan Province, Baoxing County, Fengtongzhai Town, Yanjing Village, 30 �� 32.321 ���N, 102 �� 55.273 ���E, 23 May 2015, Zhizhong Gao leg. Paratypes: 2 ♂, 1 ♀, same data as for holotype. Etymology. The species name is derived from the Latin word ��� ovatus ���, and refers to the oval, egg-shaped bursae; adjective. Diagnosis. The new species is similar to O. pseudostella (Fu et al. 2014: 488, figs 3 A���G, 4 A���E), but can be distinguished from the latter by: 1) a short E (longer in O. pseudostella); 2) PEA sharp (relatively blunt in O. pseudostella); 3) oval BU located posteriorly (pear-like and located anteriorly in O. pseudostella); 4) CO located centrally in the concavity (anteriorly, concealed by the margin of concavity in O. pseudostella). The new species is also similar to O. acuta sp. nov. (Figs 14���15), but can be distinguished from the latter by: 1) bulb with a sharp PEA prolaterally, directed to 10 o���clock (pointed ventrally in O. acuta sp. nov.); 2) palpal tibia with one broad RTA (with two distinct, thin apophyses with the same length in O. acuta sp. nov.); 3) BU oval, located posteriorly (relatively round, located anteriorly in O. acuta sp. nov.). Description. Male (Fig. 12 A). Total length 2.55���3.12 (n= 3). Holotype: body 3.12 long; carapace 1.56 long, 1.37 wide; abdomen 1.52 long, 1.14 wide. Carapace yellowish brown. Eyes sizes and interdistances: AME 0.09, ALE 0.11, PME 0.07, PLE 0.09; AME���AME 0.05, AME���ALE 0.01, PME���PME 0.09, PME���PLE 0.07. MOA 0.21 long, front 0.21 wide, back 0.23 wide. Clypeus 0.08 high. Cheliceral promargin with three well-separated teeth and retromargin with two teeth close to each other. Labium and gnathocoxae brown. Legs light brown. Leg measurements: leg I 5.91 (1.78, 0.41, 1.67, 1.42, 0.63), II 4.64 (1.28, 0.42, 1.21, 1.00, 0.73), III 3.97 (1.05, 0.36, 0.89, 1.01, 0.66), IV 6.10 (1.69, 0.48, 1.47, 1.54, 0.92). Abdomen brown, anterior half with a large dorsal scutum, occupying nearly 9 / 10 of abdomen length; posterior half without distinct chevron-shaped markings dorsally. Venter grey. Palp (Figs 12 A���H, 13 A���E): RTA with broad base bifurcated into thumb-like tip and triangular apophysis. Cymbium ventrally with several hairs around E tip. E beak-like, prolaterally with sharp PEA. SD sclerotized. Conductor membranous, sharp, originating from E base. Female (Fig. 12 B). Total length 3.74 (n= 1). One paratype: body 3.74 long; carapace 1.68 long, 1.54 wide; abdomen 2.02 long, 1.28 wide. Carapace brown. Eyes sizes and interdistances: AME 0.10, ALE 0.11, PME 0.07, PLE 0.08; AME���AME 0.05, AME���ALE 0.02, PME���PME 0.13, PME���PLE 0.07. MOA 0.26 long, front 0.23 wide, back 0.27 wide. Clypeus 0.09 high. Leg measurements: I 6.01 (1.65, 0.49, 1.72, 1.37, 0.78); II 5.10 (1.49, 0.38, 1.29, 1.18, 0.76); III 4.45 (1.26, 0.42, 0.91, 1.19, 0.67); IV 6.59 (1.79, 0.53, 1.57, 1.81, 0.89). Abdomen dark grey dorsally, anterior half without dorsal scutum; posterior half with three chevron-shaped markings dorsally. Epigyne (Figs 12 C���D, 13 D���E): CO small, located in large, shallow concavity anteriorly (Figs 12 C, 13 D), widely separated. Concavity with distinct anterior margin. CD thick and short, separated by two transparent BU, oval, located centrally; CD as long as BU. SP small, widely separated, with similar diameter to CD, located posteriorly. FD originating anteriorly from SP. Distribution. Known only from the type locality, Sichuan, China (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on pages 212-215, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Fu, L. N., Jin, C. & Zhang, F. (2014) Three new species of the genus Otacilia Thorell (Araneae: Phrurolithidae) from China. Zootaxa, 3869 (4), 483 - 492. http: // dx. doi. org / 10.11646 / zootaxa. 3869.4.10"]}

Otacilia flexa sp. nov. Figs 16 A���J, 17 A���E Type material. Holotype ♂, China, Yunnan Province, Funing County, Muyang National Public Ecological Forest, 23 �� 27.107 ���N, 105 �� 25.373 ���E, 7 May 2011, Zongxu Li leg. Paratypes: 5 ♂, 8 ♀, same data as for holotype, Zongxu Li & Guchun Zhou leg. Etymology. The species name is derived from the Latin word ��� flexus ��� and refers to the folded copulatory ducts; adjective in genitive case. Diagnosis. The new species is similar to O. longituba (Wang et al. 2012: 41, figs 4 A���E, 5 A���I), but can be distinguished from it by: 1) CO large, anteriorly, widely separated (Figs 16 C, 17 D) (smaller, located relatively posteriorly and close to each other in O. longituba); 2) boomerang-shaped BU (Figs 16 D, 17 E) (racket shaped in O. longituba); 3) DTA trifurcated apically (Figs 16 E, 17 C) (broad, with a relatively narrow base in O. longituba); 4) a longer spoon-shaped conductor (Figs 16 G, 17 B) (short in O. longituba). The new species is also similar to O. mira sp. nov. (Figs 1���2), but can be distinguished from the latter by: 1) DTA trifurcated apically (Figs 16 E, 17 C) (Sshaped, with relatively sharp apex in O. mira sp. nov.); 2) conductor present (absent in O. mira sp. nov.); 3) boomerang-shaped BU (Figs 16 D, 17 E) (hilt-like in O. mira sp. nov.); 4) SP larger (small in O. mira sp. nov.). Description. Male (Fig. 16 A). Total length 1.93���2.42 (n= 6). Holotype: body 2.33 long; carapace 1.03 long, 0.92 wide; abdomen 1.19 long, 0.83 wide. Carapace light yellowish. Eyes sizes and interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.06; AME���AME 0.03, AME���ALE 0.01, PME���PME 0.06, PME���PLE 0.05. MOA 0.13 long, front 0.10 wide, back 0.17 wide. Clypeus 0.07 high. Chelicerae with one strong anterior bristle; promargin with three well-separated teeth and retromargin with two teeth close to each other. Labium and gnathocoxae yellow. Legs light yellow. Leg measurements: leg I 2.84 (0.88, 0.26, 0.79, 0.58, 0.33), II 2.28 (0.69, 0.22, 0.52, 0.49, 0.36), III 2.19 (0.58, 0.25, 0.43, 0.52, 0.41), IV 3.13 (0.86, 0.28, 0.71, 0.76, 0.52). Abdomen dark grey, with anterior large dorsal scutum occupying nearly 4 / 5 of abdomen length; posterior half with six bright chevron-shaped markings dorsally. Venter yellow grey. Palp (Figs 16 E���H, 17 A���C): DTA strong, relatively thick, with wide base, trifurcated apically. Tibia wider than femur and patella. E long, tapering, needle-like, without PEA. SD strongly sclerotized. Conductor long, spoonshaped, its tip close to tip of E. Female (Fig. 16 B). Total length 2.46���2.76 (n= 4). One paratype: body 2.51 long; carapace 1.04 long, 0.90 wide; abdomen 1.42 long, 0.92 wide. Carapace light yellowish. Eyes sizes and interdistances: AME 0.04, ALE 0.07, PME 0.05, PLE 0.06; AME���AME 0.02, AME���ALE 0.01, PME���PME 0.05, PME���PLE 0.05. MOA 0.12 long, front 0.09 wide, back 0.16 wide. Clypeus 0.08 high. Leg measurements: I 2.80 (0.78, 0.28, 0.77, 0.63, 0.34); II 2.50 (0.76, 0.26, 0.48, 0.54, 0.44); III 2.37 (0.68, 0.23, 0.48, 0.54, 0.44); IV 3.39 (0.93, 0.29, 0.78, 0.84, 0.55). Abdomen light grey dorsally, without dorsal scutum; posterior half with seven light chevron-shaped markings dorsally. Epigyne (Figs 16 C���D, 17 D���E): CO large, widely separated, by distance equal to double their diameter. CD with two folds, connected to two large, transparent, boomerang-shaped BU anteriorly. SP round, large, located posteriorly, close to each other. FD originating anteriorly from SP. Distribution. Known only from the type locality, Yunnan, China (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on pages 217-220, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Wang, L. Y., Zhang, F. & Zhang, Z. S. (2012) Ant-like sac spiders from Jinyun Mountain Natural Reserve of Chongqing, China (Araneae: Corinnidae). Zootaxa, 3431, 37 - 53."]}

Otacilia mira sp. nov. Figs 1 A���H, 2 A���E Type material. Holotype ♂, China, Chongqing Municipality, Wanzhou Region, Wangerbao Natural Reserve, 30 �� 39.363 ���N, 108 �� 43.999 ���E, 19 September 2012, Luyu Wang & Xuankong Jiang leg. Paratypes: 8 ♂, 5 ♀, same data as holotype; 9 ♂, 10 ♀, same data as holotype, but collected on 15 March 2013; 1 ♀, Guizhou Province, Chishui City, Wuzhufeng scenic area, 28 �� 23.448 ���N, 105 �� 58.690 ���E, 2 October 2012, Luyu Wang leg. Etymology. The species name is derived from the Latin word ��� mirus ���, meaning strange, and refers to the unique structure of the epigyne; adjective. Diagnosis. This new species is similar to O. longituba (Wang et al. 2012: 41, figs 4 A���E, 5 A���I), but can be distinguished by: 1) large CO, widely separated, located centrally (relative close and located posteriorly in O. longituba); 2) epigastric furrow with a medial notch (lacking a notch in O. longituba); 3) DTA long, S-shaped, with relatively sharp apex (shorter, with a blunt apex in O. longituba); 4) tegulum inflated in the new species (uninflated in O. longituba). The new species is also similar to O. simianshan (Zhou et al. 2013: 1, figs 2 A���E, 3 A���G), but can be distinguished by: 1) large CO, located centrally and widely separated (smaller, relatively posterior, and close to each other in O. simianshan); 2) BU hilt-like in the new species (oval in O. simianshan); 3) DTA longer in the new species (relatively short in O. simianshan). Description. Male (Fig. 1 A). Total length 3.08���3.86 (n= 8). Holotype: body 3.18 long; carapace 1.54 long, 1.31 wide; abdomen 1.52 long, 0.93 wide. Carapace light yellowish. Eyes sizes and interdistances: AME 0.07, ALE 0.09, PME 0.07, PLE 0.08; AME���AME 0.05, AME���ALE 0.02, PME���PME 0.12, PME���PLE 0.08. MOA 0.25 long, front 0.19 wide, back 0.28 wide. Clypeus 0.11 high. Cheliceral promargin with three well-separated teeth and retromargin with five teeth near each other. Labium and gnathocoxae yellow. Legs light yellow. Leg measurements: leg I 6.02 (1.64, 0.38, 1.87, 1.45, 0.68), II 4.91 (1.34, 0.35, 0.37, 1.13, 0.72), III 4.14 (1.15, 0.32, 0.89, 1.12, 0.66), IV 6.47 (1.84, 0.38, 1.64, 1.75, 0.86). Abdomen light grey, anterior half with a small, grey dorsal scutum, occupying nearly 1 / 10 of abdomen length; posterior half light grey with five bright chevron-shaped markings dorsally. Venter light yellow. Palp (Figs 1 E���H, 2 A���C): DTA long, relatively curved, S-shaped, extending along the cymbium retrolaterally, its tip tapering. E gradually tapering, needle-like, tegulum membranous near base of E, without PEA. SD very short, strongly sclerotized. Female (Fig. 1 B). Total length 3.44���4.12 (n= 6). One paratype: body 3.54 long; carapace 1.65 long, 1.46 wide; abdomen 1.88 long, 1.11 wide. Carapace light yellowish anteriorly. Eyes sizes and interdistances: AME 0.08, ALE 0.09, PME 0.07, PLE 0.08; AME���AME 0.05, AME���ALE 0.02, PME���PME 0.12, PME���PLE 0.07. MOA 0.26 long, front 0.21 wide, back 0.29 wide. Clypeus 0.12 high. Leg measurements: I 6.10 (1.69, 0.37, 1.94, 1.43, 0.67); II 5.13 (1.35, 0.39, 1.41, 1.22, 0.76); III 4.62 (1.21, 0.33, 0.97, 1.33, 0.78); IV 6.61 (1.53, 0.58, 1.67, 1.92, 0.91). Abdomen light grey, without dorsal scutum dorsally, markings as in male. Epigyne (Figs 1 C���D, 2 D���E): CO large, widely separated, centrally located. CD long, folded, connecting with two large transparent BU anteriorly. SP egg-shaped, with similar width to CO, located posteriorly. FD originating anteriorly from SP. Epigynal plate with posterior medial notch. Distribution. China (Chongqing, Guizhou) (Fig. 18)., Published as part of Fu, Lina, Zhang, Zhisheng & Zhang, Feng, 2016, New Otacilia species from Southwest China (Araneae: Phrurolithidae), pp. 197-221 in Zootaxa 4107 (2) on pages 199-202, DOI: 10.11646/zootaxa.4107.2.4, http://zenodo.org/record/256257, {"references":["Wang, L. Y., Zhang, F. & Zhang, Z. S. (2012) Ant-like sac spiders from Jinyun Mountain Natural Reserve of Chongqing, China (Araneae: Corinnidae). Zootaxa, 3431, 37 - 53.","Zhou, G. C., Wang, L. Y. & Zhang, Z. S. (2013) Two new corinnid spiders from Simian Mountain of Chongqing, China (Araneae: Corinnidae). Acta Arachnologica Sinica, 22, 1 - 8."]}