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Apseudopsis rogi Esquete, sp. nov. Material examined. Holotype ���male II MNCN 20.04 / 10183, station LC 4 -C, Los Cristianos harbour, Tenerife, 28 �� 2 ��� 49 ������ N, 16 �� 42 ��� 53 ������W, 11 m depth, unvegetated sand, ���Cak Foster��� dredge, September 2013, coll. E. Ramos & R. Riera. Paratypes ���copulatory female allotype MNCN 20.04 / 10184; three male II MNCN 20.04 / 10185; male II MNCN 20.04 / 10186; male II MNCN 20.04 / 10187; preparatory female MNCN 20.04 / 10188, two copulatory females and eight male II, DBUA0001872.01-02 station LC 4 -C, 28 �� 2 ��� 49 ������ N, 16 �� 42 ��� 53 ������W, 11 m depth, unvegetated sand; ���Cak Foster��� dredge, September 2013, coll. E. Ramos & R. Riera. Other material ���six juveniles, six preparatory females, four males, DBUA0001873.01-03, station LE 4 -C, 27 �� 46 ��� 55 ������N, 17 �� 54 ��� 9 ������W, 6 m depth, unvegetated sand; All ���Cak Foster��� dredge, September 2013, coll. E. Ramos & R. Riera. Etymology. Named after Roger N. Bamber, also known as ���Rog���, in recognition of his great contribution to the knowledge of the Tanaidacea. Diagnosis. Apseudopsis with no marked posterolateral apophyses nor ventral hyposphenia on pereonites; rostrum pointed with rounded anterolateral margins; antennular inner flagellum with three segments, outer flagellum with nine segments; pereopod 1 merus without dorsodistal spine, propodus with four ventral spines; male II cheliped basis as long as broad, merus with dorsoproximal spur, propodus ventral margin perpendicular to cutting edge, cutting edge with proximal invagination followed by rounded apophysis; uropod outer ramus of three segments. Description. Male II paratype MNCN 20.04 / 10185. Body (Figure 2 A) length 3 mm, 5.3 times as long as broad. Cephalothorax about as long as broad, as long as pereonites 1���2, rostrum pointed with rounded shoulders (anterolateral margins); eyelobes with ommatidia present. Pereon with pereonites all without apophyses, without hyposphenians, margins with scattered setae. Pleon 0.14 times as long as body, pleonites (Figure 2 A, B) alike, lateral margins pointed with simple setae. Pleotelson (Figure 2 B) about as long as broad, lateral margins with simple setae, distal margin with two subdistal pairs of one simple and one penicillate setae. Antennule (Figure 2 C) peduncle article 1 3.1 times as long as broad, inner margin with simple setae, outer margin with proximal tuft of three penicillate setae, medial tuft of two simple and two penicillate setae, subdistal tuft of five simple setae and two penicillate setae; article 2 0.4 times as long as article 1, subdistally with nine simple setae and four penicillate setae; article 3 0.4 times as long as article 2, inner margin with five simple setae, outer margin with three distal setae. Flagellum common article naked; inner flagellum of three segments, segments 1 and 3 with setae, outer flagellum of nine segments, all segments with setae, segments 1���7 with one aesthetasc each. Antenna (Figure 2 D) peduncle article 1 1.2 times as long as broad, inner lobe with two small setae; article 2 1.3 times as long as broad, with inner small seta, bearing squama with 15 long marginal setae; article 3 0.3 times as long as article 2, 0.7 times as long as broad, with one long inner seta; article 4 1.6 times as long as article 3, with inner penicillate seta; article 5 1.5 times as long as article 4, twice as long as broad, with two inner proximal simple setae, two distal simple setae, two inner distal penicillate setae and two inner medial penicillate setae. Flagellum of six segments, all bearing inner short and outer long setae. Mouthparts. Labrum (not illustrated) bilobed, with setules. Right mandible (Figure 3 A) molar (Figure 3 B) triturative, with spiniform processes; pars incisiva with four denticles; setiferous lobe with five bifurcate and trifurcate setae. Palp article 1 1.3 times as long a broad, with ten setae on inner margin; article 2 1.7 times as long as article 1, 4.1 times as long as broad, with row of five pectinate spines, proximal spine longest; article 3 0.5 times as long as article 2, 3.7 as long as broad, with seven subterminal pectinate spines, one subterminal seta and three terminal setae. Left mandible (Figure 3 C) with tridentate lacinia mobilis and setiferous lobe bearing four multifurcate setae; pars incisiva with four denticles. Maxillule (Figure 3 D) inner endite with marginal apophysis, bearing five distal setulose setae; outer endite bearing eight distal spines and two subdistal pectinate setae, outer and inner margin with fine setae; palp (Figure 3 E) with four terminal setae. Maxilla (Figure 3 F) outer margin serrated, outer lobe of fixed endite with simple setae, and bifurcate, trifurcate and pectinate spines. Inner lobe of fixed endite with row of numerous simple setae in front of two pectinate setae. Outer lobe of movable endite with five pectinate setae, a row of six simple setae, and two strong, plumose setae with medial setules on outer margin. Inner lobe of movable endite with five pectinate setae. Labium (not illustrated) with outer margin serrated, palp with three distal setae and marginal setae and setules. Maxilliped basis simple; palp (Figure 3 G) first article with one longer inner seta and outer distal spine; second article inner margin with 21 setae, outer margin with one distal seta; third article with eight setae along inner margin; distal article with six distal setae; endite (Figure 3 H) inner margin with row of spines, distal margin with setules, row of bilobed setae, one subdistal long seta, and outer corner with row of five simple setae. Epignath (not illustrated) as in other species of the genus, with one circumplumose seta. Cheliped (Figure 4 A) exopod first article 3.1 times as long as broad, naked, distal article pentagonal, with five plumose setae; basis robust, as long as broad, with one ventral spine and seta, ventrodistal tuft of three setae and five dorsal setae. Merus posteriorly with a dorsal marked spur, ventrally with tuft of nine setae. Carpus 2.3 times as long as broad, with one dorsodistal and five ventral setae, and one inner distal simple seta. Chela robust and compact, propodus 1.2 times as long as broad, dorsal margin almost parallel to ventral margin, ventral margin perpendicular to cutting edge, fixed finger with four ventral and three dorsodistal setae, palm inner face with four setae near dactylus insertion and a row of seven setae and three more setae near cutting edge, cutting edge with proximal invagination followed by rounded apophyses, rest slightly convex with spinules. Dactylus with two dorsal setae. Pereopod 1 (Figure 5 A) exopod first article 3.1 times as long as broad, naked, distal article pentagonal, with five plumose setae; coxa with anterior apophysis and anterodorsal a tuft of five simple setae. Basis about twice as long as broad, with two dorsal short simple setae and tuft of simple setae on ventrodistal corner. Ischium with two ventral simple setae. Merus broader distally, with lateral row of four setae, dorsodistal tuft of setae, ventral margin with row of setae, and ventrodistal spine. Carpus 0.8 times as long as merus, bearing two distal and two ventral setae, a dorsodistal tuft of setae, and two ventral and one dorsodistal spines. Propodus about as long as carpus, with two distal and four ventral spines; dorsal and ventral margins with setae. Dactylus and unguis about 0.8 times as long as propodus, dactylus with two dorsal and one ventral spinules. Pereopod 2 (Figure 5 B) basis cylindrical, 2.7 times as long as broad, with five dorsoproximal setae, five ventrodistal setae and three ventral medial setae. Ischium 0.7 times as long as broad, with four ventral setae. Merus broader distally, with four dorsodistal setae, row of twelve ventral setae, and one ventrodistal spine. Carpus 0.9 times as long as merus, with dorsal and ventral setae, one dorsodistal and one ventrodistal spine. Propodus as long as carpus plus half of merus, three times as long as broad, dorsal margin with six setae, ventral margin with eight setae, subproximal and subdistal thin spines, distal margin with three two thin spines and three setae. Dactylus and unguis together 0.7 times as long as propodus, with dorsodistal setule. Pereopod 3 (Figure 5 C) basis with two subproximal setae, one medial seta, two subdistal setae, and four ventrodistal setae. Ischium wider than long, with one ventral setae. Merus with two ventral spines and eight ventral setae. Carpus as long as merus, with lateral row of three spines and six setae, ventral margin with row of setae. Propodus with lateral row of three spines, one ventral spine, three terminal serrate spines, one dorsal penicillate seta and simple setae on ventral, dorsal and terminal margins. Dactylus and unguis about as long as propodus, dactylus with dorsal setule. Pereopod 4 (Figure 5 D) basis about twice as long as broad, naked. Ischium with four ventral setae. Merus with three ventral spines and seven simple setae. Carpus 1.4 times as long as merus, with nine ventral spines and several simple setae. Propodus 0.8 times as long as carpus, with seven ventral setae, one dorsal penicillate seta, distally with six short and nine long pectinate spines. Dactylus and unguis together 0.9 times as long as propodus. Pereopod 5 (Figure 5 E) basis about twice as long as broad, with two dorsal penicillate setae and two ventrodistal simple setae. Ischium wider than long, with two simple setae. Merus with three ventral spines, five ventral setae and one dorsal seta. Carpus twice as long as merus, ventrally with eight spines and 13 simple setae. Propodus 0.8 times as long as carpus, with eight ventral spines, four ventral setae and one dorsal penicillate seta. Dactylus and unguis together 0.8 times as long as propodus, dactylus with one ventral setule. Pereopod 6 (Figure 5 F) basis fusiform, about twice as long as broad, dorsal and ventral margins with row of twelve plumose setae, ventral margin distally with row of five simple setae, ventrodistal corner with four simple setae. Ischium wider than long, with two ventral simple setae. Merus wider distally, dorsal margin with three plumose setae, ventral margin with eleven simple setae, distal margin with one medial simple seta, ventrodistal corner with spine. Carpus 1.5 times as long as merus, with two ventral rows of five spines, ventral margin with nine simple setae, dorsal margin with seven plumose setae. Propodus 0.7 times as long as carpus, with a row of four lateral spines, ventral margin with one proximal short spine followed by a row of pinnate spines that continues along distal margin, dorsodistal margin with one long spine. Dactylus and unguis about as long as propodus, dactylus bearing a dorsal setule. Pleopods 1 and 2 bases with five inner and six outer plumose setae; pleopods 3 and 4 with five inner and three outer plumose setae; pleopod 5 (Figure 5 G) with three inner and three outer plumose setae. Both rami with numerous distal and outer marginal plumose setae, endopod with one inner marginal plumose seta. Uropods (Figure 2 B) basis with 5 distal setae. Endopod with over 30 segments, many of them with simple setae. Exopod with three segments, proximal segment shortest, distal segment about as long as first two segments, bearing two distal setae. Preparatory female paratype MNCN 20.04 / 10188. Pereonite 6 with hyposphenium. Cheliped (Figure 4 B) exopodite present, as in male; basis 1.5 times as long as broad, dorsal margin with three proximal and two medial simple setae, ventral margin with simple seta and spine and pair of distal setae. Merus as long as basis, ventral margin with tuft of nine simple setae. Carpus cylindrical, 0.9 times as long as merus, ventral margin with 12 simple setae, dorsodistal corner with two setae, outer face with three setae proximally and two setae subdistally. Propodus dorsal margin with one dorsal setae and four setae near dactylus insertion, ventral margin four setae near claw; palm with two setae and tuft of six setae near dactylus insertion; cutting edge with row of setules. Dactylus with tuft of tree dorsal setae. Otherwise as male. Juveniles. Antennule inner flagellum of 7���9 segments. Otherwise as preparatory female. Remarks. Species of Apseudopsis are distinguished by a combination of characters that do not vary through the developmental stages, as follows (see Esquete et al., 2012 a): Shape of the rostrum, presence of apophysis on pereonites, presence or absence of dorsodistal spine on pereopod 1 merus, and number of ventral spines on pereopod 1 propodus. Other diagnostic characters include the number and position of hyposphenia on juveniles II and females, and features of the male II cheliped. The number of segments of the antennular flagella are useful for mature specimens (i.e. preparatory, copulatory and intermediate females and males II) of certain species. Apseudopsis rogi sp. nov. has a pointed rostrum with rounded shoulders, pereonites without apophyses, absence of dorsodistal spine on pereopod 1 merus, four ventral spines on pereopod 1 propodus, females with hyposphenium on pereonite 6 only. The most notable character of this new species is the male cheliped carpus with spur, which is unique for the genus. Apart from that, Apseudopsis arguinensis from Mauritania is different from Apseudopsis rogi in having hyposphenia on female pereonite 2 and 6 and six ventral spines on the pereopod 1 propodus. Apseudopsis isochelatus, also from Mauritania, differs from A. rogi in having lateral invaginations and hyposphenia in all pereonites, a more slender cheliped basis, carpus and chela in males, presence of a dorsodistal spine on the merus and five ventral spines on pereopod 1 propodus, and absence of anteroproximal protuberance with setae on pereopod 2. Other Atlantic and Mediterranean species of Apseudopsis are very similar to A. rogi in having a pointed rostrum with rounded shoulders and four ventral spines on pereopod 1 propodus, but differ in other relevant characters: A. adami, from the northwestern Iberian Peninsula has posterolateral apophysis on pereonites 2���5 that are absent on A. rogi and ten instead of nine segments on the antennular inner flagellum in mature specimens; A. mediterraneus (Băcescu, 1961) has posterolateral apophyses on pereonites 1���6, hyposphenia on pereonites 2 and 6 and a maximum of eight segments on the antennal main flagellum; A. bacescui (Guţu, 2002), from the Mediterranean Sea, has posterolateral apophyses on pereonites 1���6, hyposphenians on pereonites 2, 3 and 6 and a maximum of seven segments on the antennal main flagellum. Apseudopsis ostroumovi Băcescu & Cărăuşu, 1947, from the Black Sea, has posterolateral apophyses on pereonites 1���6, hyposphenia on pereonites 2���6 and a maximum of eight segments on the antennal main flagellum. A summary of these and other characters distinguishing species of Apseudopsis are given in the key, below. With regard to observed sex ratios and female fecundity, in sample LE 4, the male: female ratio was 4: 6. All males were male II stage, and no copulatory females were found. By contrast, in sample LC 4 the male: female ratio was 8: 4, and the four females were copulatory, with a fecundity of 6���8 embryos., Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on pages 250-256, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Family APSEUDIDAE Leach

Genus TANAIS Latreille, Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 258, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Key to the Atlantic and Mediterranean species of Apseudopsis 1. Pereopod 1 merus with dorsodistal spine.................................................................... 2 - Pereopod 1 merus without dorsodistal spine................................................................. 4 2. Pereonites 2–6 with apophyses on anterior and posterior corners; antennula inner flagellum of four segments, outer flagellum of 4–8 segments in mature specimens; pereopod 1 propodus with five ventral spines; male with hyposphenia on pereonites 2– 5, female with hyposphenia on pereonites 2–6....................... A. elisae (Băcescu, 1961) [Monaco, Mediterranean] - Pereonites without anterolateral or posterolateral apophyses.................................................... 3 3. Antennula inner flagellum of 7–9 segments, outer flagellum of 15–16 segments in mature specimens; pereopod 1 propodus with four spines on ventral margin; male with hyposphenia on pereonites 2–5, female with hyposphenia on pereonites 1–6...................................................................... A. robustus (Sars 1882) [Mediterranean] - Antennula inner flagellum of four segments, outer flagellum of 8–10 segments; pereopod 1 propodus with three spines on ventral margin; only female with hyposphenia, on pereonites 2–6..... A. latreillii (Milne-Edwards, 1828 ) [European Atlantic] 4. Pereonites without apophyses............................................................................ 5 - Some pereonites with anterolateral or posterolateral apophyses................................................. 6 5. Rostrum pointed with rounded shoulders; pereopod 1 propodus with four ventral spines; male cheliped merus with spur, chela fixed finger cutting edge with proximal invagination; females with hyposphenium on pereonite 6 only...................................................................................... A. rogi Esquete, sp. nov. [Canary Islands] - Rostrum pointed, short, subtriangular; pereopod 1 propodus with three ventral spines; male cheliped merus without spur, chela fixed finger with semicircular apophysis; females with hyposphenia on pereonites 3 and 6....................................................................................... A. apocryphus (Guţu, 2002) [Israel, Mediterranean] 6. Pereopod 1 propodus with five or more ventral spines......................................................... 7 - Pereopod 1 propodus with four or fewer ventral spines....................................................... 10 7. Pereopod 1 propodus with five ventral spines.............................................................. 8 - Pereopod 1 propodus with six ventral spines............................................................... 9 8. Rostrum short; all pereonites with posterolateral acute apophyses; female with hyposphenium on pereonite 6 only; male chela fixed finger cutting edge with semicircular apophysis.............. A. annabensis (Guţu, 2002) [Algeria, Mediterranean] - Rostrum relatively long, acute, with rounded shoulders; all pereonites with posterolateral hook-like apophyses, female with hyposphenia on pereonites 2 and 6 (males unknown)............................ A. cuanzanus Bochert, 2012 [Angola] 9. Rostrum short, pointed; all pereonites with posterolateral apophyses; female with hyposphenia on pereonites 2 and 6; male chela fixed finger with semicircular apophysis.............................. A. arguinensis (Guţu, 2002) [Mauritania] - Rostrum long, acute; pereonites 2–5 with short posterolateral hooks; females with hyposphenia on pereonites 2–6; cheliped not sexually dimorphic, slender, merus about 3.5 times as long as broad, chela cutting edges not ornamented....................................................................................... A. isochelatus Guţu, 2006 [Mauritania] 10. Rostrum acute, with rounded lobes; pereonites with posterolateral apophyses; pereopod 1 propodus with three ventral spines (sole specimen known is a copulatory female)........................ A. minimus (Guţu, 2002) [Israel, Mediterranean] - Pereopod 1 propodus with four ventral spines.............................................................. 11 11. Pereonites 3–6 with anterolateral and posterolateral apophyses................................................. 12 - No pereonite with anterolateral apophyses................................................................. 13 12. Rostrum long, acute; anterolateral apophyses on pereonites 3–6 directed forward; females with hyposphenia on pereonites 2, 3, 5 and 6 (male unknown).................................. A. acutifrons (Sars 1882) [Gulf of Naples, Mediterranean] - Rostrum short, pointed; anterolateral apophyses on pereonites 3–6 not directed forward, but perpendicular to the main axis of the body; females with hyposphenia on pereonites 1–6; male chela fixed finger with triangular apophysis on cutting edge........................................................... A. ostroumovi (Băcescu & Cărăuşu, 1947) [Black Sea] 13. Pereonites 1–6 with posterolateral apophyses; mature specimens’ outer flagellum of antennule of 6–8 segments.......... 14 - Only pereonites 2–6 with posterolateral apophyses; mature specimens’ outer flagellum of antennule of ten segments; females with hyposphenium on pereonite 6 only; male chela fixed finger with semicircular and triangular apophyses on cutting edge.................................................. A. adami Esquete & Bamber, 2012 . [Atlantic Iberian Peninsula]. 14. Females with hyposphenia on pereonites 2, 3 and 6; male chela with apophysis on fixed finger cutting edge and proximal apophysis on dactylus cutting edge.............................. A. bacescui (Guţu, 2002) [Balearic Sea, Mediterranean]. - Females with hyposphenia on pereonites 2 and 6. Male cheliped fixed finger cutting edge with proximal semicircular apophyses and invagination......................... A. mediterraneus (Guţu, 2002) [Israel, Mediterranean coast of France].

Genus LEPTOCHELIA Dana, Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 258, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Key to the Atlantic and Mediterranean species of Apseudopsis 1. Pereopod 1 merus with dorsodistal spine.................................................................... 2 - Pereopod 1 merus without dorsodistal spine................................................................. 4 2. Pereonites 2���6 with apophyses on anterior and posterior corners; antennula inner flagellum of four segments, outer flagellum of 4���8 segments in mature specimens; pereopod 1 propodus with five ventral spines; male with hyposphenia on pereonites 2��� 5, female with hyposphenia on pereonites 2���6....................... A. elisae (Băcescu, 1961) [Monaco, Mediterranean] - Pereonites without anterolateral or posterolateral apophyses.................................................... 3 3. Antennula inner flagellum of 7���9 segments, outer flagellum of 15���16 segments in mature specimens; pereopod 1 propodus with four spines on ventral margin; male with hyposphenia on pereonites 2���5, female with hyposphenia on pereonites 1���6...................................................................... A. robustus (Sars 1882) [Mediterranean] - Antennula inner flagellum of four segments, outer flagellum of 8���10 segments; pereopod 1 propodus with three spines on ventral margin; only female with hyposphenia, on pereonites 2���6..... A. latreillii (Milne-Edwards, 1828 ) [European Atlantic] 4. Pereonites without apophyses............................................................................ 5 - Some pereonites with anterolateral or posterolateral apophyses................................................. 6 5. Rostrum pointed with rounded shoulders; pereopod 1 propodus with four ventral spines; male cheliped merus with spur, chela fixed finger cutting edge with proximal invagination; females with hyposphenium on pereonite 6 only...................................................................................... A. rogi Esquete, sp. nov. [Canary Islands] - Rostrum pointed, short, subtriangular; pereopod 1 propodus with three ventral spines; male cheliped merus without spur, chela fixed finger with semicircular apophysis; females with hyposphenia on pereonites 3 and 6....................................................................................... A. apocryphus (Guţu, 2002) [Israel, Mediterranean] 6. Pereopod 1 propodus with five or more ventral spines......................................................... 7 - Pereopod 1 propodus with four or fewer ventral spines....................................................... 10 7. Pereopod 1 propodus with five ventral spines.............................................................. 8 - Pereopod 1 propodus with six ventral spines............................................................... 9 8. Rostrum short; all pereonites with posterolateral acute apophyses; female with hyposphenium on pereonite 6 only; male chela fixed finger cutting edge with semicircular apophysis.............. A. annabensis (Guţu, 2002) [Algeria, Mediterranean] - Rostrum relatively long, acute, with rounded shoulders; all pereonites with posterolateral hook-like apophyses, female with hyposphenia on pereonites 2 and 6 (males unknown)............................ A. cuanzanus Bochert, 2012 [Angola] 9. Rostrum short, pointed; all pereonites with posterolateral apophyses; female with hyposphenia on pereonites 2 and 6; male chela fixed finger with semicircular apophysis.............................. A. arguinensis (Guţu, 2002) [Mauritania] - Rostrum long, acute; pereonites 2���5 with short posterolateral hooks; females with hyposphenia on pereonites 2���6; cheliped not sexually dimorphic, slender, merus about 3.5 times as long as broad, chela cutting edges not ornamented....................................................................................... A. isochelatus Guţu, 2006 [Mauritania] 10. Rostrum acute, with rounded lobes; pereonites with posterolateral apophyses; pereopod 1 propodus with three ventral spines (sole specimen known is a copulatory female)........................ A. minimus (Guţu, 2002) [Israel, Mediterranean] - Pereopod 1 propodus with four ventral spines.............................................................. 11 11. Pereonites 3���6 with anterolateral and posterolateral apophyses................................................. 12 - No pereonite with anterolateral apophyses................................................................. 13 12. Rostrum long, acute; anterolateral apophyses on pereonites 3���6 directed forward; females with hyposphenia on pereonites 2, 3, 5 and 6 (male unknown).................................. A. acutifrons (Sars 1882) [Gulf of Naples, Mediterranean] - Rostrum short, pointed; anterolateral apophyses on pereonites 3���6 not directed forward, but perpendicular to the main axis of the body; females with hyposphenia on pereonites 1���6; male chela fixed finger with triangular apophysis on cutting edge........................................................... A. ostroumovi (Băcescu & Cărăuşu, 1947) [Black Sea] 13. Pereonites 1���6 with posterolateral apophyses; mature specimens��� outer flagellum of antennule of 6���8 segments.......... 14 - Only pereonites 2���6 with posterolateral apophyses; mature specimens��� outer flagellum of antennule of ten segments; females with hyposphenium on pereonite 6 only; male chela fixed finger with semicircular and triangular apophyses on cutting edge.................................................. A. adami Esquete & Bamber, 2012 . [Atlantic Iberian Peninsula]. 14. Females with hyposphenia on pereonites 2, 3 and 6; male chela with apophysis on fixed finger cutting edge and proximal apophysis on dactylus cutting edge.............................. A. bacescui (Guţu, 2002) [Balearic Sea, Mediterranean]. - Females with hyposphenia on pereonites 2 and 6. Male cheliped fixed finger cutting edge with proximal semicircular apophyses and invagination......................... A. mediterraneus (Guţu, 2002) [Israel, Mediterranean coast of France]., Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 257, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Genus APSEUDOPSIS Norman, 1899, Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 250, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Genus TANAIS Latreille

Family LEPTOCHELIIDAE Lang

Family LEPTOCHELIIDAE Lang, Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 258, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Esquete, Patricia, Ramos, Eva, Riera, Rodrigo (2016): New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis. Zootaxa 4093 (2): 248-260, DOI: http://dx.doi.org/10.11646/zootaxa.4093.2.6

Tanais dulongii (Audouin, 1826) Material examined. One female, length 2.5 mm, station LC 4 -C, 28 �� 2 ��� 49 ������ N, 16 �� 42 ��� 53 ������W, 11 m depth, unvegetated sand, ���Cak Foster��� dredge, September 2013, coll. E. Ramos & R. Riera. Remarks. Tanais dulongii was described from the coast of Egypt and has a broad distribution on Atlantic and Mediterranean coasts (Hamers et al. 2000; Bamber 2012) where it dwells littoral algae. In the Macaronesian region, early records of the species in the Azores were partially clarified by Bamber & Costa (2009), and later Bamber (2012) confirmed its presence in Cabo Verde and Madeira. This is the first record of T. dulongii for Tenerife, and the second for the Canary Islands after Sanz et al. (2003) found it in subtidal seabeds in SW Lanzarote., Published as part of Esquete, Patricia, Ramos, Eva & Riera, Rodrigo, 2016, New data on the Tanaidacea (Crustacea: Peracarida) from the Canary Islands, with a description of a new species of Apseudopsis, pp. 248-260 in Zootaxa 4093 (2) on page 258, DOI: 10.11646/zootaxa.4093.2.6, http://zenodo.org/record/261881

Genus LEPTOCHELIA Dana

Setosabatieria triangularis sp. nov. (Figure 3, Table 2) Type locality. Los Cristianos beach (Tenerife, Canary Islands). Type material. Holotype. Adult male 1.7 mm long, mounted on a slide (coordinates 28 º02´58 ´´N / 16 º 42´54 ´´W; 3 m depth) [TFMCBMNA/00011]. Collected by R. Riera, November 2000. Allotype, adult female, 2.1 mm long, mounted on a slide (coordinates 28 º02´58 ´´N / 16 º 42´54 ´´W; 3 m depth) [DBAULL NE/TC 1 S 11]. Collected by R. Riera, May 2000. Paratypes. Adult male, 2.4 mm long, mounted on a slide (coordinates 28 º02´58 ´´N / 16 º 42´54 ´´W; 3 m depth) [DBAULL NE/TC 7 S 11]. Collected by R. Riera, November 2000. Collected by R. Riera, May 2000. Adult female, 1.6 mm long, mounted on a slide (coordinates 28 º02´58 ´´N / 16 º 42´54 ´´W; 3 m depth) [DBAULL NE/TC 7 S 12]. Collected by R. Riera, November 2000. Meristic data. Cristianos subtidal: May, 1 female (Allotype), November, 1 female (f 2), 2 males (Holotype and m 2). Etymology. The specific name refers to the triangular gubernaculum of this species. Description. Male: Body slender, attenuating towards both ends. Head round and slightly set off. Cuticle with very fine transverse striations, sometimes inconspicuous in the posterior half. Amphids are 68 % of the corresponding body diameter in width, consisting of 2.5 turns, located at 6 m from the anterior end. Buccal cavity conical and small, without noticeable teeth. Inner labial setae lacking. Six outer labial setae 2 m long and 4 cephalic setae 1.6 cephalic diameters long, situated on the anterior part of the head. Subcephalic setae 13 m long, numerous, arranged in 6–9 rows and located at 21–60 m from the anterior end. Pharynx narrow and cylindrical. Ventral gland and nerve ring not seen. The reproductive system is diorchic, with two opposed testes, almost inconspicuous. Spicules 1.7 anal diameters long, paired, arcuated and without marked capitulum. Gubernaculum 0.75 anal diameters long, with slender apophysis and distally compressed. 15 precloacal supplements 3 m wide, the posteriormost located at 21 m from the cloaca. Precloacal seta 7 m long, situated at 18 m from the cloaca. Tail 5.3 anal diameters long, cylindrical and with slightly swollen posterior tip. Subterminal setae 17 m long, located at 5 m from the posterior end. Caudal setae 25 m long. Spinneret conspicuous. Female: They are similar to males. Tail length slightly shorter than in males (3.2–4.5 anal diameters). The reproductive system is didelphic, with two opposed and reflexed ovaries. Vulva located in the anterior half of the body, at the level of 45.7 % of the total body length. Discussion. Setosabatieria triangularis new species most closely resembles S. fibulata (Wieser, 1954) in the shape of copulatory apparatus, but S. triangularis new species differs in having amphids 2.5 turns, occupying 68 % of the corresponding body diameter, while S. fibulata possess more developed amphids (90 % cbd), with 4.25 turns. S. triangularis new species can be distinguished from the remaining species of the genus: S. hilarula (De Man, 1922) and S. jingjingae (Guo & Warwick, 2001) in the number of amphid turns (3.25 in S. hilarula and 3.5 in S. jingjingae) and the size of the gubernaculum, very slender with a triangular apophysis in S. triangularis new species but narrower and slender in S. hilarula and S. jingjingae. Ecology. This species was recorded in Los Cristianos subtidal from fine sands (150– 180 µm, median particle size). The organic matter content ranged from 0.006 % to 1.01 % and 19.32–27.35 % carbonates.

Key to Parapionosyllis species from the Canary Islands 1. Unidentate ventral simple chaetae................................................................................. 2 �� Bidentate ventral simple chaetae ........................... Parapionosyllis abriguensis sp. nov. 2. Dorso��ventral gradation in length of blades of compound chaetae............................... 3 �� Short��bladed compound chaetae, without a dorso��ventral gradation in length of fal�� cigerous blades ............................................................................................. P. labronica 3. Dorsal simple chaeta with 1 thick subdistal spine......................................................... 4 �� Dorsal simple chaetae with 2���3 thick subdistal spines ..................... P.macaronesiensis 4. Blades of the dorsal��most compound chaetae twice as long as the ventralmost blades ........................................................................................................................... P.elegans �� Blades of the dorsal��most compound chaetae more than three times longer than the ventralmost blades ............................................................................................ P. minuta, Published as part of Riera, Rodrigo & Brito, Jorge N����ez And Mar��a Del Carmen, 2006, Parapionosyllis (Polychaeta: Syllidae: Exogoninae) from Tenerife (Canary Islands, Spain) with description of a new species and new records, pp. 17-26 in Zootaxa 1110 on pages 23-24, DOI: 10.5281/zenodo.171520

Paracomesoma sigmoidalis sp. nov. (Figures 1, 2, Table 1) Type locality. Los Abrigos beach (Tenerife, Canary Islands). Type material. Holotype, adult male 2.6 mm long, mounted on a slide (coordinates 28 ��08��34 ����N / 16 �� 26��21 ����W; 3 m depth) [TFMCBMNA/00010]. Collected by R. Riera, May 2000. Allotype, adult female, 2.2 mm long, mounted on a slide (coordinates 28 ��02��59 ����N / 16 �� 42��55 ����W; 3 m depth) [DBAULL NE/TC 7 S 11]. Paratypes. Adult male, 2.6 mm long, mounted on a slide (coordinates 28 ��08��34 ����N / 16 �� 26��21 ����W; 3 m depth) [DBAULL NE/TA 11 S 11]. Collected by R. Riera, March 2000. Adult male, 2.3 mm long, mounted on a slide (coordinates 28 ��08��34 ����N / 16 �� 26��21 ����W; 3 m depth) [DBAULL NE/TAS 12]. Additional specimens (two males and four females) in R. Riera collection. Meristic data. Abrigos subtidal: May, 2 males (Holotype and m 4), March, 1 male (m 2), December, 1 male (m 3) and 3 females (f 3, f 4 and f 5); Cristianos subtidal: November, 1 female (Allotype), April, 1 female (f 2). Etymology. The specific name refers to the S��shaped spicules of this species. Description. Male: Body slender, attenuating towards both ends. Head slightly round and not set off. Cuticle smooth, without marked punctations nor lateral differentiation. Amphids are 50 % of the corresponding body diameter in width, multispiral of 3.5 turns, located at 5 m from the anterior end. Buccal cavity conical and small, with three small teeth, difficult to discern. Inner labial setae inconspicuous. 6 outer labial setae 0.3 cephalic diameters long and 4 cephalic setae 1.6 cephalic diameters long, situated at the median part of the head. Subcephalic setae 14 m long, located at 28 m from the anterior end. Pharynx slender and cylindrical. Ventral gland and nerve ring not seen. The reproductive system is diorchic, with two opposed testes, difficult to discern. Spicules 3.6���5.2 anal diameters long, slender, S��shaped, without marked capitulum. Gubernaculum 0.8���0.9 anal diameters long, slender, proximally compressed. An elliptical plaque present at the distal end of the gubernaculum. 35 precloacal supplements, each penetrated by a duct; posteriormost precloacal supplement situated at 45 m from the cloaca. Tail 2.4���4.4 anal diameters long, cylindrical and filiform in most of its length, with a swollen posterior tip. 2 subterminal setae 15 m long, located at 5 ���m from the posterior tip. Spinneret well developed. Female: They are similar to males. Total body length slightly shorter than in males (1.7���2.3 mm). Tail longer than in males (5���6 anal diameters). The reproductive system is didelphic, with two outstretched ovaries, almost inconspicuous. Vulva located in the posterior half of the body, at the level of 53.9���55.2 % of the total body length. Discussion. Paracomesoma sigmoidalis new species is closely related to P. inaequale Jensen & Gerlach, 1977, in general body shape, but the latter species differs in having a longer tail, 6.3���7.5 anal diameters in males and 8���9 anal diameters in females, and the number of precloacal supplements (25���32 in P. inaquale and 35 in P. sigmoidalis). P. sipho (Gerlach, 1956) and P. hexasetosum (Chitwood, 1937) resemble P. sigmoidalis new species although they differ, like not as P. inaequale, in the presence of a larger tail and the number of precloacal supplements (30 in P. sipho and 40 or more in P. hexasetosum). Moreover, P. sipho has more developed amphids (80 % cbd) and P. sigmoidalis new species can be distinguished from the latter two species by having subcephalic setae more anteriorly located. Holotype m 2 m 3 m 4 Allotype f 2 f 3 f 4 f5 m 1 f 1 Maximum body diameter 70 50 46 65 37.3 40 39.3 39.3 39.5 Ecology. This species was recorded in Los Abrigos subtidal from medium sands (280 ��m, median particle size). The organic matter content was 0.78 % and 5.47 % carbonates. In Los Cristianos it was collected from fine sands (150���160 ��m, median particle size). The organic matter content ranged from 0.006 % to 0.53 % and 19.32���22.56 % carbonates., Published as part of Riera, Rodrigo, N����ez, Jorge & Brito, Mar��a Del Carmen, 2006, Two new species of Comesomatidae Filipjev, 1922 (Nematoda: Chromadorida) from sandy bottoms of Tenerife, Canary Islands, pp. 53-61 in Zootaxa 1126 on pages 55-57, DOI: 10.5281/zenodo.171831

Genus Paracomesoma Hope & Murphy 1972 Type species. Paracomesoma dubium (Filipjev, 1918). Cuticle with transverse striations or punctations, sometimes difficult to discern. Lateral differentiation lacking. Buccal cavity with three small triangular teeth. Spicules narrow and slender, not jointed. Gubernaculum small, sometimes inconspicuous, and without apophysis. Males with two opposed testes and females with two outstretched ovaries. Up to now, the genus Paracomesoma comprises 6 species: P. dubium (Filipjev, 1918), P. hexasetosum (Chitwood, 1937), P. inaequale Jensen & Gerlach, 1977, P. longispiculum (Timm, 1961), P. quadrisetosum (Chitwood, 1937) and P. s i p h o (Gerlach, 1956)., Published as part of Riera, Rodrigo, N����ez, Jorge & Brito, Mar��a Del Carmen, 2006, Two new species of Comesomatidae Filipjev, 1922 (Nematoda: Chromadorida) from sandy bottoms of Tenerife, Canary Islands, pp. 53-61 in Zootaxa 1126 on page 54, DOI: 10.5281/zenodo.171831

Key to Parapionosyllis species from the Canary Islands 1. Unidentate ventral simple chaetae................................................................................. 2 ­ Bidentate ventral simple chaetae ........................... Parapionosyllis abriguensis sp. nov. 2. Dorso­ventral gradation in length of blades of compound chaetae............................... 3 ­ Short­bladed compound chaetae, without a dorso­ventral gradation in length of fal­ cigerous blades ............................................................................................. P. labronica 3. Dorsal simple chaeta with 1 thick subdistal spine......................................................... 4 ­ Dorsal simple chaetae with 2–3 thick subdistal spines ..................... P.macaronesiensis 4. Blades of the dorsal­most compound chaetae twice as long as the ventralmost blades ........................................................................................................................... P.elegans ­ Blades of the dorsal­most compound chaetae more than three times longer than the ventralmost blades ............................................................................................ P. minuta

Riera, Rodrigo, Brito, Jorge Núñez And María Del Carmen (2006): Parapionosyllis (Polychaeta: Syllidae: Exogoninae) from Tenerife (Canary Islands, Spain) with description of a new species and new records. Zootaxa 1110: 17-26, DOI: 10.5281/zenodo.171520

Parapionosyllis Fauvel, 1923: 289. Parapionosyllis minuta (Pierantoni, 1903) Pionosyllis minuta Pierantoni, 1903: 239, fig. 5. Parapionosyllis minuta. ��� Fauvel, 1923: 292, fig. 111 f; San Mart��n, Vi��itez & Campoy, 1981: 68, fig. 8; Campoy, 1982: 263, fig. 17; San Mart��n, 1984: 198, fig. 44; San Mart��n, 2003: 288, figs. 158���159. Material examined.���Los Abrigos beach (Tenerife), Lat. N. 28 ��08��58 ���� Long. W. 16 �� 42��54 ����, 3 m depth, 1 specimen, June 2000, coll. R. Riera. Ecology.���This species was recorded in medium grained sands with an organic matter of 1.54 % and 6.84 % of carbonates. The species has been recorded from a variety of habitats and reaches its highest abundances in sandy substrates, green algae (Caulerpa spp.) and seagrass meadows (Zostera and Posidonia) (Vi��itez, 1976). Distribution.���East Atlantic and Mediterranean (San Mart��n, 2003). This is the first record of this species from the Canary Islands. Parapionosyllis macaronesiensis Brito, N����ez & San Mart��n, 2000 Parapionosyllis macaronesiensis Brito, N����ez & San Mart��n 2000: 1147, fig. 1. Parapionosyllis sp.��� N����ez, Pascual, Delgado & San Mart��n 1995: 6. Material examined.���Los Abrigos beach (Tenerife), 3 m depth, 1 specimen, March 2001, coll. R. Riera. Ecology. ���This species was recorded in medium grained sands with an organic content. of 0.91 % and 6.32 % of carbonates. It is a strictly interstitial species of sandy subtidal bottoms (Brito, 2002). In Madeira it was collected in coarse grained sands with shell remains, rich in carbonates (N����ez et al., 1995). Distribution. ���East Atlantic: Central Macaronesian Region (Madeira, Selvagens Islands and Canary Islands). Parapionosyllis abriguensis sp. nov. (Figure 1, Table 1) Type locality. ���Los Abrigos beach (Tenerife, Canary Islands) Type material. ��� Holotype (microscopic slide) (TFMCBM AN/000222) Los Abrigos beach (Tenerife), Lat. 28 ��08��30 N Long. 16 �� 26��10 ����W, 3 m depth, July 2000, coll. R. Riera. Paratypes, three specimens (microscopic slide): (DBAULL PO/TA 12 S 11; /TA 4 S 55; / TA 3 S 55), Los Abrigos beach (Tenerife), same coordinates; 3 m depth, August 2000, coll. R. Riera. Description. ���Body long, tapering towards both ends, without any pigmentation. Body 0.8���1.6 mm long, and 61���100 m wide at proventricle level. Prostomium semicircular to ovate, slightly wider than long, apparently without eyes. Antennae skittleshaped, median antenna slightly longer (77 m) than lateral ones (72���75 m). Lateral antennae inserted near of half margin of the prostomium, median antenna inserted behind on the posterior margin (Fig. 1 A). Palps broad, similar in length to prostomium, fused along basal half, leaving a mid��dorsal seam and frontal scar. Single pair of tentacular cirri, originating laterally, more or less similar in length (71 ��m) and width to lateral antennae (Fig. 1 A). Peristomium dorsally reduced. Dorsal cirri on all chaetigers, slightly shorter (43���68 m) than lateral antennae and with granular material distally. Parapodial lobe triangular��shaped, with a ventral cirri digitiform (23���32 m) similar in length to parapodial lobe (Fig. 1 C). Apparently lacking subdermal dorsal glands. Anterior parapodia each with one long��bladed compound chaeta and up to five falcigers with shorter blades; progressively, number of falcigers decreasing in median and posterior parapodia (Fig. 1 E). Long blades unidentate, with tips rounded and uniform fine serration along one margin. Dorsalmost falcigers blades about 31 m on most anterior parapodia, 39 m in midbody and 40 m posteriorly. Remaining shorter falcigers with dorso��ventral gradation in length of blades, in midbody 14 m above, 9 m below. Solitary dorsal simple chaeta from anterior parapodia, unidentate, provided with spinulated serrations distally, without thick spines (Fig. 1 D). Posterior parapodia also with a solitary ventral simple chaeta, sigmoid and bidentate (Fig. 1 F). Solitary thick acicula in each parapodium, distally rounded hollow at the tip (Fig. 1 G). Pharynx wide, extending through about three segments, provided with a conspicuous, conical middorsal tooth on the anterior rim. Proventricle shorter than pharynx, extending through about three segments, and with about 16���18 rows of muscle cells. Pygidium with one pair of long, slender anal cirri (125 m) and one caudal appendix (18 m) (Fig. 1 B). Remarks.��� Parapionosyllis abriguensis sp. nov. is characterized by having a unidentate dorsal simple chaeta with spinulated serrations without any thick spines, bidentate ventral simple chaeta, marked dorso��ventral gradation and by the length of the falcigerous blades and smooth dorsal cirri. Parapionosyllis floridana San Mart��n, 1991 is the species most similar to P. abriguensis, both are the two only species of this genus having bidentate simple ventral chaetae and lacking eyes, but it differs in having pseudoarticulated dorsal cirri and a less marked dorso��ventral gradation in the length of the falcigerous blades (10���33 m in P. floridana and 9���39 m in P. abriguensis). Ecology. ���This species was recorded in fine and medium sands. The organic matter content ranged from 0.77 % to 0.85 % and carbonates from 5.13 % to 9.57 %. Etymology. ���The derivatio nominis refers to the type locality, Los Abrigos beach. Distribution. ���Tenerife, Canary Islands., Published as part of Riera, Rodrigo & Brito, Jorge N����ez And Mar��a Del Carmen, 2006, Parapionosyllis (Polychaeta: Syllidae: Exogoninae) from Tenerife (Canary Islands, Spain) with description of a new species and new records, pp. 17-26 in Zootaxa 1110 on pages 19-21, DOI: 10.5281/zenodo.171520, {"references":["Fauvel, P. (1923) Polychetes Errantes. In: Faune de France, vol 5, Le Chevalier (ed.), Paris, 488 pp.","Pierantoni, U. (1903) La gestazione esterna (Contributo alla biologia de alla embriologia dei Sillidi). Archivo Zoologico Italiano de Torino, 1, 231 - 252.","San Martin, G., Vieitez, J. M. & Campoy, A. (1981) Contribucion al estudio de la fauna de Anelidos Poliquetos de las costas espanolas: Poliquetos Errantes recoletados en la bahia de Palma de Mallorca. Boletin del Instituto Espanol de Oceanografia, 6, 63 - 87.","Campoy, A. (1982) Fauna de Espana. Fauna de Anelidos Poliquetos de la Peninsula Iberica. EUNSA, (Ediciones de la Universidad de Navarra), Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica, vol. 7, 781 pp.","San Martin, G. (1984) Estudio biogeografico, faunistico y sistematico de los Poliquetos de la familia Silidos (Syllidae: Polychaeta) en Baleares. PhD Thesis, 581 pp.","San Martin, G. (2003) Annelida, Polychaeta II: Syllidae. In Fauna Iberica, vol. 21. Ramos, M. A et al. (eds.), Museo Nacional de Ciencias Naturales, CSIC, Madrid, 554 pp.","Vieitez, J. M. (1976) Ecologia de Poliquetos y Moluscos de la playa de Meira (Ria de Vigo). Investigaciones Pesqueras, 40 (1), 223 - 238.","Brito, M. C., Nunez, J. & San Martin, G. (2000) Parapionosyllis macaronesiensis, a new species of Exogoninae (Polychaeta: Syllidae) from the Macaronesian Region. Proceedings of the Biological Society of Washington, 113 (4), 1147 - 1150.","Nunez, J., Pascual, M., Delgado, J. D. & San Martin, G. (1995) Interstitial polychaetes from Madeira with a description of Syllides bansei Perkins, 1981. Bocagiana, 179, 1 - 7.","Brito, M. (2002) Estudio de las comunidades intersticiales del sebadal (Cymodocea nodosa) en Canarias, con especial referencia a los anelidos poliquetos. PhD Thesis, 618 pp.","San Martin, G. (1991) Sphaerosyllis and Parapionosyllis (Polychaeta: Syllidae) from Cuba and Florida. Ophelia suppl., 5, 321 - 328."]}

Genus Setosabatieria Platt, 1985 Type species. Setosabatieria hilarula (De Man, 1922). Cuticle smooth. Buccal cavity small, cup��shaped, without marked teeth. Rows of developed subcephalic setae. Gubernaculum with developed apophysis, dorsocaudally directed. Males with two opposed testes and females with two opposed and reflexed ovaries. The genus Setosabatieria includes 3 species (Guo & Warwick, 2001): S. fibulata (Wieser, 1954), S. hilarula (De Man, 1922) and S. jingjingae Guo & Warwick, 2001., Published as part of Riera, Rodrigo, N����ez, Jorge & Brito, Mar��a Del Carmen, 2006, Two new species of Comesomatidae Filipjev, 1922 (Nematoda: Chromadorida) from sandy bottoms of Tenerife, Canary Islands, pp. 53-61 in Zootaxa 1126 on page 58, DOI: 10.5281/zenodo.171831, {"references":["Platt, H. M. (1985) The freeliving marine nematode genus Sabatieria (Nematoda: Comesomatidae). Taxonomic revision and pictorial keys. Zoological Journal of the Linnean Society, 83, 27 - 78.","Guo, Y. Q. & Warwick, R. M. (2001) Three new species of free-living nematodes from the Bohai Sea, China. Journal of Natural History, 35, 1575 - 1586"]}

Paracomesoma sigmoidalis sp. nov. (Figures 1, 2, Table 1) Type locality. Los Abrigos beach (Tenerife, Canary Islands). Type material. Holotype, adult male 2.6 mm long, mounted on a slide (coordinates 28 º08´34 ´´N / 16 º 26´21 ´´W; 3 m depth) [TFMCBMNA/00010]. Collected by R. Riera, May 2000. Allotype, adult female, 2.2 mm long, mounted on a slide (coordinates 28 º02´59 ´´N / 16 º 42´55 ´´W; 3 m depth) [DBAULL NE/TC 7 S 11]. Paratypes. Adult male, 2.6 mm long, mounted on a slide (coordinates 28 º08´34 ´´N / 16 º 26´21 ´´W; 3 m depth) [DBAULL NE/TA 11 S 11]. Collected by R. Riera, March 2000. Adult male, 2.3 mm long, mounted on a slide (coordinates 28 º08´34 ´´N / 16 º 26´21 ´´W; 3 m depth) [DBAULL NE/TAS 12]. Additional specimens (two males and four females) in R. Riera collection. Meristic data. Abrigos subtidal: May, 2 males (Holotype and m 4), March, 1 male (m 2), December, 1 male (m 3) and 3 females (f 3, f 4 and f 5); Cristianos subtidal: November, 1 female (Allotype), April, 1 female (f 2). Etymology. The specific name refers to the S­shaped spicules of this species. Description. Male: Body slender, attenuating towards both ends. Head slightly round and not set off. Cuticle smooth, without marked punctations nor lateral differentiation. Amphids are 50 % of the corresponding body diameter in width, multispiral of 3.5 turns, located at 5 m from the anterior end. Buccal cavity conical and small, with three small teeth, difficult to discern. Inner labial setae inconspicuous. 6 outer labial setae 0.3 cephalic diameters long and 4 cephalic setae 1.6 cephalic diameters long, situated at the median part of the head. Subcephalic setae 14 m long, located at 28 m from the anterior end. Pharynx slender and cylindrical. Ventral gland and nerve ring not seen. The reproductive system is diorchic, with two opposed testes, difficult to discern. Spicules 3.6–5.2 anal diameters long, slender, S­shaped, without marked capitulum. Gubernaculum 0.8–0.9 anal diameters long, slender, proximally compressed. An elliptical plaque present at the distal end of the gubernaculum. 35 precloacal supplements, each penetrated by a duct; posteriormost precloacal supplement situated at 45 m from the cloaca. Tail 2.4–4.4 anal diameters long, cylindrical and filiform in most of its length, with a swollen posterior tip. 2 subterminal setae 15 m long, located at 5 “m from the posterior tip. Spinneret well developed. Female: They are similar to males. Total body length slightly shorter than in males (1.7–2.3 mm). Tail longer than in males (5–6 anal diameters). The reproductive system is didelphic, with two outstretched ovaries, almost inconspicuous. Vulva located in the posterior half of the body, at the level of 53.9–55.2 % of the total body length. Discussion. Paracomesoma sigmoidalis new species is closely related to P. inaequale Jensen & Gerlach, 1977, in general body shape, but the latter species differs in having a longer tail, 6.3–7.5 anal diameters in males and 8–9 anal diameters in females, and the number of precloacal supplements (25–32 in P. inaquale and 35 in P. sigmoidalis). P. sipho (Gerlach, 1956) and P. hexasetosum (Chitwood, 1937) resemble P. sigmoidalis new species although they differ, like not as P. inaequale, in the presence of a larger tail and the number of precloacal supplements (30 in P. sipho and 40 or more in P. hexasetosum). Moreover, P. sipho has more developed amphids (80 % cbd) and P. sigmoidalis new species can be distinguished from the latter two species by having subcephalic setae more anteriorly located. Holotype m 2 m 3 m 4 Allotype f 2 f 3 f 4 f5 m 1 f 1 Maximum body diameter 70 50 46 65 37.3 40 39.3 39.3 39.5 Ecology. This species was recorded in Los Abrigos subtidal from medium sands (280 µm, median particle size). The organic matter content was 0.78 % and 5.47 % carbonates. In Los Cristianos it was collected from fine sands (150–160 µm, median particle size). The organic matter content ranged from 0.006 % to 0.53 % and 19.32–22.56 % carbonates.

Setosabatieria triangularis sp. nov. (Figure 3, Table 2) Type locality. Los Cristianos beach (Tenerife, Canary Islands). Type material. Holotype. Adult male 1.7 mm long, mounted on a slide (coordinates 28 ��02��58 ����N / 16 �� 42��54 ����W; 3 m depth) [TFMCBMNA/00011]. Collected by R. Riera, November 2000. Allotype, adult female, 2.1 mm long, mounted on a slide (coordinates 28 ��02��58 ����N / 16 �� 42��54 ����W; 3 m depth) [DBAULL NE/TC 1 S 11]. Collected by R. Riera, May 2000. Paratypes. Adult male, 2.4 mm long, mounted on a slide (coordinates 28 ��02��58 ����N / 16 �� 42��54 ����W; 3 m depth) [DBAULL NE/TC 7 S 11]. Collected by R. Riera, November 2000. Collected by R. Riera, May 2000. Adult female, 1.6 mm long, mounted on a slide (coordinates 28 ��02��58 ����N / 16 �� 42��54 ����W; 3 m depth) [DBAULL NE/TC 7 S 12]. Collected by R. Riera, November 2000. Meristic data. Cristianos subtidal: May, 1 female (Allotype), November, 1 female (f 2), 2 males (Holotype and m 2). Etymology. The specific name refers to the triangular gubernaculum of this species. Description. Male: Body slender, attenuating towards both ends. Head round and slightly set off. Cuticle with very fine transverse striations, sometimes inconspicuous in the posterior half. Amphids are 68 % of the corresponding body diameter in width, consisting of 2.5 turns, located at 6 m from the anterior end. Buccal cavity conical and small, without noticeable teeth. Inner labial setae lacking. Six outer labial setae 2 m long and 4 cephalic setae 1.6 cephalic diameters long, situated on the anterior part of the head. Subcephalic setae 13 m long, numerous, arranged in 6���9 rows and located at 21���60 m from the anterior end. Pharynx narrow and cylindrical. Ventral gland and nerve ring not seen. The reproductive system is diorchic, with two opposed testes, almost inconspicuous. Spicules 1.7 anal diameters long, paired, arcuated and without marked capitulum. Gubernaculum 0.75 anal diameters long, with slender apophysis and distally compressed. 15 precloacal supplements 3 m wide, the posteriormost located at 21 m from the cloaca. Precloacal seta 7 m long, situated at 18 m from the cloaca. Tail 5.3 anal diameters long, cylindrical and with slightly swollen posterior tip. Subterminal setae 17 m long, located at 5 m from the posterior end. Caudal setae 25 m long. Spinneret conspicuous. Female: They are similar to males. Tail length slightly shorter than in males (3.2���4.5 anal diameters). The reproductive system is didelphic, with two opposed and reflexed ovaries. Vulva located in the anterior half of the body, at the level of 45.7 % of the total body length. Discussion. Setosabatieria triangularis new species most closely resembles S. fibulata (Wieser, 1954) in the shape of copulatory apparatus, but S. triangularis new species differs in having amphids 2.5 turns, occupying 68 % of the corresponding body diameter, while S. fibulata possess more developed amphids (90 % cbd), with 4.25 turns. S. triangularis new species can be distinguished from the remaining species of the genus: S. hilarula (De Man, 1922) and S. jingjingae (Guo & Warwick, 2001) in the number of amphid turns (3.25 in S. hilarula and 3.5 in S. jingjingae) and the size of the gubernaculum, very slender with a triangular apophysis in S. triangularis new species but narrower and slender in S. hilarula and S. jingjingae. Ecology. This species was recorded in Los Cristianos subtidal from fine sands (150��� 180 ��m, median particle size). The organic matter content ranged from 0.006 % to 1.01 % and 19.32���27.35 % carbonates., Published as part of Riera, Rodrigo, N����ez, Jorge & Brito, Mar��a Del Carmen, 2006, Two new species of Comesomatidae Filipjev, 1922 (Nematoda: Chromadorida) from sandy bottoms of Tenerife, Canary Islands, pp. 53-61 in Zootaxa 1126 on pages 58-60, DOI: 10.5281/zenodo.171831, {"references":["Guo, Y. Q. & Warwick, R. M. (2001) Three new species of free-living nematodes from the Bohai Sea, China. Journal of Natural History, 35, 1575 - 1586"]}

23 páginas, 13 figuras, 2 tablas., The present paper includes a morphological, ecological and biological updating of the three gorgonian associated species of Haplosyllis (Polychaeta, Syllidae) known to date: H. chamaeleon (symbiont with Paramuricea clavata in the Mediterranean), H. anthogorgicola (symbiont with Anthogorgia bocki in the Japanese seas) and H. villogorgicola , a new species living symbiotically with Villogorgia bebrycoides which is only known from Tenerife (Canary Islands, Eastern Central Atlantic). The new species is described on the basis of ecological, morphological, morphometric and statistical analysis of relevant characteristics. Each host colony harboured about 15 pale-yellowish worms, whose cryptic colouration mimicked that of the host. They occurred either on the host branches or partly hidden inside cavities formed by the fusion of two branches. The new species is characterized by the presence of simple chaetae with clearly bidentate tips all along the body, the presence of gland pore aggregates distributed in two lateral rows and two ventral patches on each palp and the absence of ciliary tufts on the pharyngeal papillae. H. villogorgicola sp. nov. is closely related to H. chamaeleon . Thus, it is compared with two populations of this species collected in the north-west and south-west Mediterranean. Stolons of H. chamaeleon are re-described as tetracerous and a peculiar posterior end regeneration process occurring in adult worms during the stolon formation is described. H. anthogorgicola is also re-described, with particular emphasis on its appendage and chaetal arrangements. The main features of the three associations are discussed in light of the current knowledge on symbiotic polychaetes, particularly cnidarian-associated syllids. © 2002 The Linnean Society of London, Biological Journal of the Linnean Society , 2002, 77 , 455–477., This paper is a contribution to the research project INTAS-97–0916.