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Key to the members of the Brachyopa quadrimaculosa species group in the Eastern Mediterranean (following Doczkal & Dziock 2004) 1. Scutellum entirely covered in microtrichia................................. Brachyopa bimaculosa Doczkal & Dziock - Scutellum largely bare of microtrichia, at least on posterior 1 / 2 ��� 1 / 3.............................................. 2 2. Proepimeron with a few pile anteriorly. Scutum with rounded bare spots at the transverse suture (Figs 3 C & D)........... 3 - Proepimeron bare. Scutum with microtrichose transverse suture................................................. 5 3. Lateral sides of scutum uniformly covered with light grey microtrichia....... Brachyopa vernalis van Steenis & van Steenis - Lateral sides of scutum with stripes of dark grey microtrichia... (as in Fig. 3 C)..................................... 4 4. Postalar callus with a very small bare spot (Fig. 3 C); male terminalia illustrated in Kaplan & Thompson (1981: 209, Figs 16��� 18)............................................... Brachyopa quadrimaculosa Thompson in Kaplan & Thompson - Postalar callus with large bare spot (see van Steenis & van Steenis 2004: Fig. 4).................................................................................................. Brachyopa cruriscutum van Steenis & van Steenis 5. Sensory pit present; scutum with bare spots and stripes (Figs 3 A & B); tergites with pale pile; male terminalia in Figs 4���5.................................................................................. Brachyopa minima sp. nov. - Sensory pit absent; scutum completely microtrichose except for the postalar callus; second tergite with black pile posteromedi- ally in female (can be absent in male); (male terminalia illustrated in Vujić (1991: 147, Fig. 4 B)....................................................................................................... Brachyopa insensilis Collin, Published as part of P��rez-Ba����n, Celeste, Radenkovi��, Snezana, Vuji��, Ante, St��hls, Gunilla, Rojo, Santos, Grkovi��, Ana & Petanidou, Theodora, 2016, Brachyopa minima (Diptera: Syrphidae), a new species from Greece with notes on the biodiversity and conservation of the genus Brachyopa Meigen in the Northern Aegean Islands, pp. 217-234 in Zootaxa 4072 (2) on page 223, DOI: 10.11646/zootaxa.4072.2.5, http://zenodo.org/record/270665, {"references":["Doczkal, D. & Dziock, F. (2004) Two new species of Brachyopa Meigen from Germany, with notes on B. grunewaldensis Kassebeer (Diptera, Syrphidae). Volucella, 7, 35 - 59.","Kaplan, M. & Thompson, F. C. (1981) New Syrphidae from Israel (Diptera). Proceedings of the Entomological Society of Washington, 83, 198 - 212."]}

Brachyopa minima Vujić & P��rez-Ba����n sp. nov. Diagnosis. Small species (6���7 mm). Scutum (Figs 3 A, B) covered with grey microtrichia except: pair of small bare spots posteromedian to the postpronotum; a pair of large bare spots anterior to transverse suture and median to notopleural sulcus; spots at the median ends of transverse sulcus covered by brown microtrichia; postalar callus with a bare spot; two stripes bare of microtrichia on posterior part of scutum located laterally. Male terminalia on Figs 4���5; surstylus with well-developed ventral lobe (Figs 4 A, 5 A); dorsal and ventral margins of surstylus straight (Figs 4 A, 5 A); hypandrium wide with complex apical appendages (Figs 5 C, D). Type material. HOLOTYPE: Greece, Lesvos, Karini, 13.04.2011, 1 male (G0322), leg. Vujić (in FSUNS). PARATYPES: Greece, Lesvos, Karini, 13.04.2011, 4 males (G018���G020, G023), 1 female (G0324), 13.04.2013, 1 female (G 2805), leg. Vujić (in FSUNS); 30.04.2008, 1 male (G 3069), leg. Vujić (in FSUNS); Greece, Lesvos, Karini, 26.04.2007, 1 male (G 3068) 1 female (G 3067), leg. P��rez-Ba����n and Vujić (in FSUNS). Greece, Lesvos, Karini, 3.05. 2007, leg. P��rez-Ba����n, 1 female, DNA voucher specimen MZH _Y 544 (in MZH). Etymology. The specific epithet minima means "smallest" and refers to the size of the species, which is smaller than most other members of the genus. Description. MALE. Body length 6���7 mm, wing length 7 mm. Head (Figs 1 A, C; 2 A���C). Length to height ratio of head 0.97. Face moderately protruding (Fig. 1 C). Length to width ratio of clypeus 1.4. Length to width ratio of subcranial cavity 1.6; 1.7 times as wide as the shortest distance from the eye to the subcranial cavity. Orbital strip (Figs 2 A, B) shallow, in narrower part as wide as the maximum width of arista. Pile on all parts of the head capsule pale, except black pile on upper part of occiput near eye margin. Frons and vertex (Fig. 1 A) microtrichose, only narrow area above lunule and around ocelli bare. Length of eye contiguity about 0.4 times the length of frons, shorter than ocellar triangle. Distance between posterior ocellus and eye margin about equal to two ommatidium diameters. Hypostomal bridge grey-reddish, paler anteriorly, entirely microtrichose, except bare hypostomal sulcus. Face (Figs 2 A, B) microtrichose except for a fascia from the tentorial sulcus to the mouth. Length to height ratio of postpedicellus 1���1.15, with small sensory pit (Fig. 2 C). Arista (Fig. 2 C) with very short pile (shorter than 1 / 3 the maximum width of arista), with orange base and darkened toward apex. Thorax. Black except for the postalar callus, which is partly reddish, and the metasternum and probasisternum which are more or less reddish. Scutum (Figs 3 A, B) entirely densely covered in grey microtrichia except for a pair of bare maculae posteromedian to the postpronotum; a pair of bare maculae anterior to transverse suture and median to notopleural sulcus (maculae at the median ends of transverse sulcus covered by brown microtrichia); postalar callus with a bare spot; two stripes bare of microtrichia on posterior part of scutum located laterally. Notopleural sulcus impressed. Thoracic bristles not well differentiated from the pile. Postpronotum pale haired, with few black pile mixed in. Scutum predominantly black haired except for its anterior margin and the notopleuron, which are pale haired, a few pale pile that are present at the supra-alar area, and the postalar callus that is pale haired. Scutellum rather trapezoidal, without a distinct depression, mainly orange, only slightly darkened anteriorly and anterolaterally, with two pairs of strong black marginal bristles; predominantly pale haired mixed with black pile medially; only anterior half covered with microtrichia (Fig. 2 D). Mediotergite microtrichose below subscutellum, except at posterior margin. Proepimeron and anterior flat part of mesanepisternum bare. Posterior surface of mid coxa without pile. Legs orange, 5 th tarsomere of all legs black, tarsomeres 1���4 of metatarsus darkened dorsally. Legs pale pilose except for the usual black bristles on hind femur ventrally, apex of mesotibia ventrally with strong black bristles, the ventral surface of mesotarsus with black bristles. Haltere yellow with light yellow capitulum. Wing without dark spots. Abdomen. Tergites and sternites orange, except first tergite and first sternite with dark grey area laterally, pile all pale. First tergite entirely microtrichose, second tergite extensively covered with microtrichia except for the posterior margin narrowly and large area at the posterolateral corners, which are bare. Sternites entirely densely microtrichose. Male terminalia on Figs 4���5. FEMALE. Differs from the male in the following characters: length to width ratio of frons 1.46���1.54. Frons (Fig. 1 B) entirely microtrichose except shiny anterior area above antennae, with small central microtrichose spot (this spot lacking in B. quadrimaculosa). A narrow pollinose stripe running from the dusted area of the frons downward along the eye margin, widening beside the lunule and merging with the wide pollinose cross band that covers the dorsal part of face (face less covered by microtrichia than in male, without bare spot below each antenna, contrary to B. quadrimaculosa) (Fig. 1 B). Length to height ratio of postpedicellus 1.14���1.15, a little larger than in male, with a minute sensory pit. Tibiae and tarsi darker than in male., Published as part of P��rez-Ba����n, Celeste, Radenkovi��, Snezana, Vuji��, Ante, St��hls, Gunilla, Rojo, Santos, Grkovi��, Ana & Petanidou, Theodora, 2016, Brachyopa minima (Diptera: Syrphidae), a new species from Greece with notes on the biodiversity and conservation of the genus Brachyopa Meigen in the Northern Aegean Islands, pp. 217-234 in Zootaxa 4072 (2) on pages 220-222, DOI: 10.11646/zootaxa.4072.2.5, http://zenodo.org/record/270665

An on-going study of the hoverfly fauna of the Northern Aegean Islands (Greece) has revealed the presence of four species of the genus Brachyopa Meigen. During the survey the following species were found: B. bicolor (Fallén), B. quadrimaculosa Thompson in Kaplan & Thompson, B. minima Vujić & Pérez-Bañón sp. nov. and an unidentified species very close to B. pilosa (Collin). Morphological characters and mitochondrial COI barcodes were used to link different life stages of B. minima, and to identify a larval specimen of B. bicolor. In this study adult and larval morphology and habitat preferences for B. minima are described. The description of larval morphology of B. bicolor and Brachyopa sp. aff. pilosa is amended too. An identification key to the adults of the B. quadrimaculosa group sensu Kassebeer (2002) in the Eastern Mediterranean (Greece, Israel and Turkey) is provided. The importance of specific microhabitats for the continued existence of these taxa is discussed. The study was funded by the European Union (European Social Fund – ESF) and Greek national funds through the Operational Program "Education and Lifelong Learning" of the National Strategic Reference Framework (NSRF)—Research Funding Program: THALES—Investing in knowledge society through the European Social Fund (Petanidou et al. 2013). Partial support came from the Ministry of Education, Science and Technological Development of the Republic of Serbia projects No. 173002 and the Provincial Secretariat for Science and Technological Development (project: Genetic diversity of agricultural ecosystems).