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Metaseiulus (Metaseiulus) neoflumenis Moraes & Kreiter Metaseiulus (Metaseiulus) neoflumenis Moraes & Kreiter in Moraes et al., 2000: 256. Metaseiulus (Metaseiulus) neoflumenis.��� Moraes et al., 2004: 285. Specimens examined. O���Higgins Region. Municipality of Machal��, July 2012, on Azara dentata R. et P., associated with Eriophyidae. Santiago Region, knoll Cantillana, October 2010. on A. dentata, associated with Eriophyidae. Measurements of a collected female: dorsal shield length 332 long, maximum width 225; j 1 18, j 3 27, j 4 20, j 5 19, j 6 25, J 2 32, J 5 10, z 2 25, z 3 26, z 4 30, z 5 20, Z 1 35, Z 4 41, Z 5 45, s 4 35, s 6 42, S 5 21, r 3 25, R 1 41, St IV 30. Remarks. These measurements are similar to those of the original description of the species., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 334, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Moraes, G. J. de, Kreiter, S. & Lofego, A. C. (2000) (1999) Plant mites (Acari) of the French Antilles. 3. Phytoseiidae (Gamasida). Acarologia, 40, 237 - 264.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1"]}

Neoseiulus alpinus (Schweizer) Amblyseius obtusus var. alpinus Schweizer, 1922: 41. Typhlodromips alpinus.��� Moraes et al., 1986: 136. Neoseiulus alpinus.��� Moraes et al., 2004: 100. Specimens examined. Santiago Region. El Melocot��n, Municipality of San Jos�� de Maipo, on Malus domestica Borkh, April 2001, associated with Panonychus ulmi Koch., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 328, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Schweizer, J. (1922) Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandlungen der Naturferschenden Gesellschaft in Basel, 33, 23 - 112, 4 tabs.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1"]}

Neoseiulus californicus (McGregor) Typhlodromus californicus McGregor, 1954: 89. Typhlodromus chilenensis Dosse, 1958: 55 (synonym according to Athias-Henriot 1977). Amblyseius chilenensis.��� Gonzalez & Schuster, 1962: 10. Neoseiulus californicus.��� Moraes et al., 1986: 73: 2004: 109; Xu et al., 2013: 332. Specimens examined. Santiago Region. Municipality of La Pintana, on Aloysia triphylla Palau, July 2002, associated with Tetranychus urticae Koch; on Baccharis salicina Torr. & A. Gray., July 2002, associated with Eriophyidae; on Bidens aurea (Aiton), April 2000, associated with T. urticae; on Convolvulus arvensis L., February 2001, associated with Tetranychus desertorum Banks; on Galega officinalis L., October 2000, associated with Tetranychus desertorum Banks; on Malus domestica Borkh, February 2002, associated with Tydeidae; on Malva nicaensis All., July 2002, associated with Brevipalpus sp.; on Medicago sativa L., March 2002, associated with Tetranychus desertorum Banks; on Poligonum sp., March 2002, associated with Tetranychus desertorum Banks; on Prunus persica L., February 2002, associated with Tydeidae; on Pyrus communis L., February 2002, associated with Panonychus ulmi (Koch) and Oligonychus yothersi (McGregor); on Trifolium sp., March 2002, associated with T. urticae. Municipality of Buin, on Prunus avium (L.), April 1990, associated with Bryobia rubrioculus (Scheuten) and Panonychus ulmi (Koch); on Prunus domestica L., March 1990, associated with Aculus fockeui (Nalepa & Trouessart). Municipality of Renca, on Solanum lycopersicum L., March 2000, associated with Aculops lycopersici (Massee). Measurements of a collected female: dorsal shield 362 long, 182 maximum width. Setae j 1 22, j 3 32, j 4 21, j 5 21, j 6 25, J 2 30, J 5 12, z 2 35, z 4 37, z 5 22, Z 1 32, Z 4 50, Z 5 70, s 4 40, S 2 37, S 4 37, S 5 32, r 3 25, R 1 22, St IV 52. Remarks. These measurements are similar to those given by Schuster & Pritchard (1963) for Neoseiulus californicus and to those given in the original description of Neoseiulus (Amblyseius) chilenensis by Gonzalez & Schuster (1962)., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on pages 328-329, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["McGregor, E. A. (1954) Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 53, 89 - 92.","Dosse, G. (1958) Ubereinege neue Raubmilbenarten. Pflanzenschutzber, 21, 125 - 133.","Athias-Henriot, C. (1977) Nouvelles notes sur les Amblyseiini. III. Sur le genre Cydnodromus: Redefinition, composition (Parasitiformes, Phytoseiidae). Entomophaga, 22, 61 - 73. https: // doi. org / 10.1007 / BF 02372991","Gonzalez, R. H. & Schuster, R. O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletin Tecnico. Estacion Experimental Agronomica. Universidad de Chile, Facultad de Agronomia, Chile, 16, 1 - 35.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Xu, X. - N., Wang, B. - M., Wang, E. - D. & Zhang, Z. - Q. (2013) Comments on the identity of Neoseiulus californicus sensu lato (Acari: Phytoseiidae) with a redescription of this species from southern China. Systematic & Applied Acarology, 18 (4), 329 - 344. https: // doi. org / 10.11158 / saa. 18.4.3"]}

Metaseiulus (Metaseiulus) relictus Trincado & Martin, n. sp. (Figs 18���24) Diagnosis. This species is most similar to Metaseiulus brevicollis Gonzalez & Schuster, 1962, also described from Chile. However, the latter differs by having fixed cheliceral digit with only two teeth in addition to the apical tooth, R 1 on dorsal shield and about as long as s 6, and dorsal shield lightly reticulate. FEMALE : (4 specimens measured) Dorsum (Fig. 18). Dorsal shield smooth; with seven distinguishable pairs of pores and 13 distinguishable pairs of lyrifissures; 360 (335��� 370) long, maximum width 185 (175���205); j 1 25 (23���27), j 3 30 (29���34), j 4 12 (12���20), j 5 15 (14���15), j 6 16 (15���20), J 2 20 (12���20), J 5 13 (12���14), z 2 17 (16���27), z 3 27 (25���30), z 4 30 (30���32), z 5 12 (10���13), Z 4 52 (51���60), Z 5 74 (65���77), s 4 47 (41���47), s 6 47 (43���50), S 5 18 (18���19), R 1 20 (17���22). All setae smooth. Peritreme (Fig. 18). Extending forward almost to level of j 1. Venter (Fig. 18). Sternal shield smooth; with three pairs of setae, and two pairs of lyrifissures; distances between St 1��� St 3 71 (68���73), St 2��� St 2 65 (60���73). Genital shield smooth; distance between St 5��� St 5 60 (55���74). Ventrianal shield smooth, shield-shaped, slightly constricted at level of JV 2; 120 (120���128) long, 95 (95���100) wide at ZV 2 level and 90 (84���90) at anus level, with four pairs of pre-anal setae and a pair of small, rounded pre-anal pores mesad and slightly posterior of JV 3. Seta JV 5 60 (60���65). Two pairs of slender metapodal plates, the posterior distinctly larger. Chelicera (Fig. 20). Antiaxial lyrifissure distinct. Fixed digit 43 (40���45) long, with seven teeth in addition to apical tooth and setiform pilus dentilis. Movable digit 36 (35���40) long, with two teeth in addition to apical tooth. Legs (Fig. 21). Macrosetae present only on legs III and IV, all with a very tiny distal knob: Sge III 22 (21���23), Sge IV 48 (47���48), Sti IV 36, St IV 46 (44���46). Chaetotaxy of genu II 2 2/1, 2/0 1; genu III 1 2/1, 2/0 1. Spermatheca (Fig. 22). Calyx bell-shaped, 9 (8���10) long. Atrium discrete. Male: (3 specimens measured) Dorsum. Dorsal shield similar to female; 310 (300��� 310) long, maximum width 165 (160���165); j 1 20 (16���22), j 3 29 (28���30), j 4 16 (15���17), j 5 15, j 6 15 (15���16), J 2 16 (16���17), J 5 11 (10���12), z 2 16 (14���20), z 4 27 (25���28), z 5 11 (10���12), Z 4 27 (25���28), Z 5 60 (59���65), s 4 38 (32���40), S 5 16 (16���18), R 1 18 (16���20). All setae smooth. Peritreme. Extending between j 1 and j 3. Venter (Fig. 23). Sternogenital shield smooth with five pairs of setae and two pairs of liryfissures; distances between St 1��� St 3 68 (66���71), St 2��� St 2 52 (52���55). Ventrianal shield transversely striate, subtriangular, 140 (137��� 150) long, 150 (150���155) wide at anterior corners, with five pairs of pre-anal setae and a pair of round pre-anal pores. Metapodal shields indistinct. Chelicera (Fig. 24). Fixed digit 35 (30���40) long, with five teeth in addition to apical tooth and a setiform pilus dentilis. Movable digit 25 (25���35) long, with one tooth in addition to apical tooth; spermadactyl with shaft 19 (18��� 21) and foot 12 (9���12) long. Legs. Macrosetae present only on legs III and IV, but differently from female, distal knob not distinct: Sge III 17 (16���18), Sge IV 33 (30���33), Sti IV 25, St IV 35 (35���37). Etymology: The specific name relictus refers to the type locality of this species, a humid relict forest located at the southern extreme of the coastal desert of north-central Chile. Type locality. Holotype female, collected on 24 March 2014, from Parque National Fray Jorge, Coquimbo, Chile, on Escallonia illinita C. Presl, associated with Brevipalpus sp. Deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura ���Luiz de Queiroz���, Universidade de S��o Paulo. One paratype female and three paratype males, same collection records as holotype; all deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura ���Luiz de Queiroz���, Universidade de S��o Paulo; 11 paratype females and 6 paratype males, on Senecio sp., other collection records same as holotype, deposited at Museo Nacional de Historia Natural, Santiago to Chile., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on pages 332-334, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Gonzalez, R. H. & Schuster, R. O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletin Tecnico. Estacion Experimental Agronomica. Universidad de Chile, Facultad de Agronomia, Chile, 16, 1 - 35."]}

Graminaseius graminis (Chant) Typhlodromus (Amblyseius) graminis Chant, 1956: 34. Neoseiulus graminis.��� Moraes et al., 1986: 81. Amblyseius graminis.��� Moraes et al., 2004: 26. Specimens examined. Region of Santiago. Municipality of San Pedro, on Rafanus sativus L., February 2002, associated with Tetranychus urticae Koch., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 328, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Chant, D. A. (1956) Some mites of the subfamily Phytoseiinae (Acarina: Laelaptidae) from southeastern England, with descriptions of new species. The Canadian Entomologist, 88, 26 - 37. https: // doi. org / 10.4039 / Ent 8826 - 1","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1"]}

Neoseiulus viticolus Trincado & Martin, n. sp. (Figs 11–17) Diagnosis: This species is most similar to Neoseiulus brigarinus Beard, 2001, Neoseiulus cucumeris (Oudemans, 1930) and Neoseiulus noosae (McMurtry & Schicha, 1987). They all differ from the new species here described by having distinct atrium, and ventrianal shield much wider at level at ZV 2 than at level of anus. The former two species further differ by having three teeth on the fixed digit and one tooth on movable digit, while the latter further differs by having shorter Z 5. FEMALE : (4 specimens measured) Dorsum (Fig. 11). Dorsal shield reticulated, except for a transverse band between setae S 4 and S 5; with seven distinguishable pairs of pores and 11 pairs of lyrifissures; 398 (375– 412) long, 210 (205–218) wide; j 1 24 (23–25), j 3 26 (25–28), j 4 18, j 5 16 (15–18), j 6 19 (18–20), J 2 20, J 5 13, z 2 21 (20–23), z 4 23 (23–25), z 5 16 (15–18), Z 1 20, Z 4 43 (40–48), Z 5 56 (55–58), s 4 32 (30–35), S 2 29 (28–32), S 4 23 (20–25), S 5 23, r 3 24 (23–25), R 1 23 (20–25). All setae smooth, except for Z 4 and Z 5, serrate. Peritreme (Fig. 11). Extending forward almost to level of j 1. Venter (Fig. 12). Sternal shield smooth over most of its extent, with scant anterolateral striae; with three pairs of setae, and two pairs of lyrifissures; distances between St 1– St 3 71 (68–73), St 2– St 2 73 (68–78). Genital shield smooth; distance between St 5– St 5 81 (75–85). Ventrianal shield transversely striate in the anterior half and irregularly striate elsewhere, vase-shaped 136 (133–140) long, 92 (88–100) wide at ZV 2 level and 88 (85–90) at anus level, with three pairs of pre-anal setae and a pair of small, rounded pre-anal pores posteromesad of JV 2. Seta JV 5 41 (38–45). All ventral setae smooth. Two pairs of slender metapodal shields, the posterior distinctly larger. Chelicera (Fig. 13). Antiaxial lyrifissure distinct. Fixed digit 28 (25–30) long, with five teeth in addition to apical tooth and setiform pilus dentilis. Movable digit 31 (30–33) long, with two teeth in addition to apical tooth. Legs (Fig. 14). Macrosetae present only on leg IV, with tiny distal knob: Sge 29 (28–30), Sti 23, St 41 (38–45). Chaetotaxy of genu II 2 2/0, 2/0 1; genu III 1 2/1, 2/0 1. Spermatheca (Fig. 15). Calyx bell-shaped, 18 long. Atrium discrete. Male: (3 specimens measured) Dorsum. Dorsal shield similar to female; 311 (305–315) long, 212 (200–222) wide; j 1 21 (21–22), j 3 25 (23–26), j 4 16 (13–16), j 5 17 (15–18), j 6 17 (16–17), J 2 16 (16–17), J 5 11 (10–11), z 2 18 (17–18), z 4 24 (21–26), z 5 15 (14– 15), Z 1 19 (17–23), Z 4 34 (31–37), Z 5 42 (39–46), s 4 32 (27–36), S 2 22 (19–25), S 4 20 (19–21), S 5 20, r 3 22 (20–24), R 1 18 (16–20). All setae smooth, except for Z 4 and Z 5, serrated. Peritreme. Extending forward almost to level of j 3. Venter (Fig. 16). Sternogenital shield mostly smooth, with scant lateral striae; with five pairs of setae, and two pairs of liryfissures; distances between St 1– St 3 66 (66–67), St 2–St2 62 (61–62). Ventrianal shield reticulate, subtriangular, 188 (185–192) long, 141 (137–145) wide at anterior corners, with five pairs of pre-anal setae and a pair of small, rounded pre-anal pores posteromesad of JV 2. Metapodal shields indistinct. Chelicera (Fig. 17). Fixed digit 22 (21–22) long, with five teeth in addition to apical tooth and setiform pilus dentilis. Movable digit 20 (19–21) long, with one tooth in addition to apical tooth; spermadactyl with shaft 19 (15–21) and foot 8 (7–9) long. Legs. Macrosetae present only on leg IV, with tiny distal knob: Sge 21 (19–25), Sti 18 (15–19), St 33 (30–35). Etymology. The specific name viticolus refers to Vitis vinifera L., the plant species from which the types of this new species were collected. Type locality. Holotype female collected on 29 September 2009, from Pirque, Santiago Region, Chile, on Vitis vinifera L., associated with Brevipalpus chilensis Baker. Deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura “Luiz de Queiroz”, Universidade de São Paulo. One paratype female and one paratype male, same collection records as holotype; deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura “Luiz de Queiroz”, Universidade de São Paulo; two paratype females and one paratype male, same collection data as holotype, deposited at Museo Nacional de Historia Natural, Santiago to Chile.

Neoseiulus bicaudus (Wainstein) Amblyseius bicaudus Wainstein, 1962: 146. Neoseiulus bicaudus.��� Moraes et al., 1986: 72; 2004: 108; Wang et al., 2015: 456. Specimens examined. Santiago Region. Municipality of La Pintana, on Malva nicaensis All, April 2002, associated with Tetranychus desertorum Banks; on Vitis vin��fera L., June 2002, associated with Brevipalpus chilensis Baker. Measurements of a collected female: dorsal shield 400 long, 160 maximum width; j 1 25, j 3 28, j 4 10, j 5 15, j 6 17, J 2 18, J 5 12, z 2 17, z 4 15, z 5 7, Z 1 18, Z 4 35, Z 5 110, s 4 30, S 2 32, S 4 33, S 5 40, r 3 29, R 1 25, St IV 61. Remarks. Measurements of the specimens collected fit the redescription of the species provided by Ferragut et al. (2010)., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 328, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Wainstein, B. A. (1962) Some new predatory mites of the family Phytoseiidae (Parasitiformes) of the USSR fauna. Entomologicheskoe Obozrenie, 41, 230 - 240. [translation in Entomological Review, 41, 139 - 146]","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Wang, B. - M., Wang, Z. - H., Jiang, X. - H., Zhang, J. - P. & Xu, X. - N., (2015) Re-description of Neoseiulus bicaudus (Acari: Phytoseiidae) newly recorded from Xinjiang, China. Systematic and Applied Acarology, 20 (4), 455 - 461. https: // doi. org / 10.11158 / saa. 20.4.11","Ferragut, F., Perez Moreno, I., Iraola, V. & Escudero, A. (2010) Acaros depredadores em las plantas cultivadas. Familia Phytoseiidae. Ediciones Agrotecnicas, Madrid, 202 pp."]}

Galendromus (Galendromus) occidentalis (Nesbitt) Typhlodromus occidentalis Nesbitt, 1951: 29. Galendromus (Galendromus) occidentalis.��� Moraes et al. 1986: 190; 2004: 268. Specimens examined. Santiago Region. Municipality of La Pintana, on Convolvulus arvensis L., March 2000, associated with Tetranychus urticae Koch; on Ficus carica L., March 2000, associated with Eriophyes fici Essig; on Malus domestica Borkh, February 2002, associated with Tydeidae; on Pelargonium hortorum L. H. Bailey, June 2002, associated with Tydeidae; on Phaseolus vulgaris L., March 2002, associated with Tetranychus desertorum Banks; Prunus salicina, April 2002, associated with Oligonychus sp.; on Trifolium sp., March 2002, associated with T.. urticae; Buin, on Malus domestica Borkh, April 1990, associated with Panonychus ulmi (Koch); on Prunus avium (L.), April 1990, associated with Panonychus ulmi (Koch) and T. urticae. Measurements of two collected females: dorsal shield 315, 342 long, 170, 180 width; j 1 20; j 3 52; j 4 31; j 5 37, 46; j 6 56, 62; j 2 55, 68; J 5 9, 10; z 2 53, 59; z 3 47, 54; z 4 52, 58; z 5 37, 47; Z 4 58, 67; z 5 57, 62; s 4 54, 62; s 6 62, 71; S 2 62, 72; S 5 57, 65; r 3 42, 49. Remarks. The two females with the same collection data were found to have very different sizes, and for this reason, both were measured. Both of them are distinctly smaller than reported by Chant & Yoshida-Shaul (1984) for a type specimen (dorsal shield 380 long); concurrently, most setae of the larger specimens are about 10���20% shorter than reported by those authors for that type specimen., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on pages 331-332, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Nesbitt, H. H. J. (1951) A taxonomic study of the Phytoseiidae (Family Laelaptidae) predaceous upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 1 - 64, 32 pls.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Chant, D. A. & Yoshida-Shaul, E. (1984) A world review of four species groups in the genus Typhlodromus Scheuten (Acari: Phytoseiidae): brevicollis, luculentis, carinulatus, and pinnatus. Canadian Journal of Zoology, 62, 2631 - 2642. https: // doi. org / 10.1139 / z 84 - 383"]}

Arrenoseius robertogonzalezi Trincado & Martin, n. sp. Diagnosis. This species is most similar to Arrenoseius morgani (Chant, 1957), but the latter differs by having much longer j 1, j 3, s 4, Z 4 and Z 5 and the much shorter z 2 and z 4, and by having no teeth on the movable cheliceral digit. Female: (4 specimens measured) Dorsum (Fig. 1). With seven distinguishable pairs of pores and 11 distinguishable pairs of lyrifissures; 389 (360���407) long, 320 (300���330) maximum width; j 1 17 (16���18), j 3 25 (24���26), j 4 8 (7���9), j 5 8 (6���9), j 6 8 (6���9), J 2 8 (7���9), J 5 10 (8���11), z 2 16 (14���18), z 4 24 (23���25), z 5 6 (6���7), Z 1 11 (10���12), Z 4 71 (68���75), Z 5 89 (85���96), s 4 54 (50���57), S 2 17 (15���18), S 4 15 (14���16), S 5 17 (15���19), r 3 21 (20���22), R 1 16 (15���17). All setae smooth. Peritreme (Fig. 1). Extending forward to level of j 1. Venter (Fig. 2). Sternal shield reticulate; with two pairs of lyrifissures; distances between St 1��� St 3 61 (60���64), St 2��� St 2 64 (60���65). Genital shield with striae forming an inverted U-shaped pattern; distance between St 5��� St 5 92 (89���94). Ventrianal shield reticulate, shield-shaped 123 (120���130) long and 142 (100���165) wide at ZV 2 level, with three pairs of pre-anal setae and a very pair of small, rounded pre-anal pores posteromesad of JV 2. Seta JV 5 50 (50���51). All ventral setae smooth. Two pairs of metapodal plates, the inner rounded and much smaller than the elongate outer pair. Chelicera (Fig. 3). Antiaxial lyrifissure distinct. Fixed digit 34 (33���35) long, with eight teeth in addition to apical tooth and setiform pilus dentilis. Movable digit 35 (35���36) long, with two teeth in addition to apical tooth. Legs (Fig. 4). Macroseta present only on tarsus IV, 61 (60���64). Chaetotaxy of genu II 2 2/1, 2/0 1; genu III 1 2/ 1, 2/0 1. Spermatheca (Fig. 5). Calyx saccular, 18 (17���20) long. Atrium distinct. Etymology. This species is named after Prof. Roberto H. Gonz��lez, eminent Chilean acarologist and entomologist, who regretfully passed away recently. Type locality: Holotype female collected on 2 April 1991, from Municipality of Buin, Santiago Region, Chile, on Festuca sp. Deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura ���Luiz de Queiroz���, Universidade de S��o Paulo. Three paratype females collected on 9 July 2012, from Quinta de Tilcoco, O���Higgins Region, Chile, on Viburnum tinus L., associated with Tydeidae; all deposited at Departamento de Entomologia e Acarologia, Escola Superior de Agricultura ���Luiz de Queiroz���, Universidade de S��o Paulo. Two paratype females collected on 5 November 2002, from San Pedro, Santiago Region, on Passiflora incarnata L, associated with Tydeidae, deposited at Museo Nacional de Historia Natural, Santiago to Chile., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on pages 324-325, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Chant, D. A. (1957) Descriptions of some phytoseiid mites (Acarina, Phytoseiidae). Part I. Nine new species from British Columbia with keys to the species of British Columbia. Part II. Redescriptions of eight species described by Berlese. The Canadian Entomologist, 89, 289 - 308. https: // doi. org / 10.4039 / Ent 89289 - 7"]}

Chileseius camposi Gonzalez & Schuster Chileseius camposi Gonzalez & Schuster, 1962: 7. Chileseius camposi.— Moraes et al., 1986: 182, 2004: 58. Avioseius pennatus Karg 1976: 513 (synonymy according to Chant & McMurtry, 2003: 11). Chileseius pennatus.— Chant & McMurtry, 2003: 11. Specimens examined. Holotype female: Curicó, Chile, on Vitis vinífera L., associated with Brevipalpus chilensis Baker, 8 April 1961, collected by L. E. Campos; deposited at USNM United States National Insect & Mite Collection, US National Museum of Natural History (Smithsonian), held at Systematic Entomology Laboratory (SEL), Beltsville Agricultural Research Center West (BARC West), United States Department of Agriculture (USDA), Beltsville, Maryland, USA. Paratype female: Vilcún, Temuco, Chile, on apple, associated with Tetranychidae, 28 April 1961, collected by L. M. Smith; deposited at Museo Luis Peña Guzmán, Facultad de Agronomía, Universidad de Chile, Santiago. Other specimens: Santiago Region. Municipality of La Pintana, on Punica granatum L., February 2008, associated with Eriophyes granati (Canestrini & Massalongo); on Rubus ulmifolius Schott., September 2002, associated with Tydeidae. Municipality of Curacaví, on Schinus latifolius (Gill. ex Lindl.), February 2000, associated with Tydeidae. FEMALE : (holotype, paratype and 4 additional specimens measured) Dorsum (Fig. 6). Dorsal shield with seven distinguishable pairs of pores and 11 distinguishable pairs of lyrifissures; 377 369 (344–405) long, 208 215 (187–230) maximum width; j 1 22 27 (20–30), j 3 22 31 (25–37), j 4 17 20 (15–25), j 5 17 20 (15–27), j 6 18 23 (15–32), J 2 22 31 (20–39), J 4 22 32 (20–32), J 5 12 15 (12–18), z 2 22 32 (25–40), z 4 24 35 (24–44), z 5 19 25 (18–32), z 6 23 30 (22–38), Z 1 27 37 (30–43), Z 3 23 32 (25–40), Z 4 45 55 (45– 58), Z 5 58 62 (55–70), s 4 27 42 (34–50), S 2 32 43 (31–50), S 4 34 48 (40–63), S 5 34 46 (37–53), r 3 23 31 (28–37), R 1 22 34 (28–39). Holotype with J 4, J 5, Z 4, Z 5, S 2 S 4 and S 5 serrate; other setae, smooth. All setae relatively stout; setae on the opisthonotal region of the dorsal shield setae on small tubercles. Peritreme (Fig. 6). Extending forward to level of j 1. Venter (Fig. 7). Sternal shield smooth; with two pairs of lyrifissures; distances between St 1– St 3 66 66 (64–67), St 2– St 2 62 68 (63–75). Genital shield smooth; distance between St 5– St 5 67 71 (67–75). Ventrianal shield striate, shield-shaped 129 127 (117–135) long, 89 98 (89–100) wide at ZV 2 level and 56 75 (56–80) at anus level, with three pairs of pre-anal setae and a pair of small, rounded pre-anal pores posteromesad of JV 2. Seta JV 5 35 48 (35– 55). All ventral setae smooth. Two pairs of slender metapodal plates, the posterior distinctly larger. Chelicera (Fig. 8). Antiaxial lyrifissure distinct. Fixed digit 35 35 long, with five or six teeth in addition to apical tooth and setiform pilus dentilis. Movable digit 32 32 long, with two teeth in addition to apical tooth. Legs (Fig. 9). Macrosetae present only on leg IV, blunt (in some specimens collected in this study, slightly knobbed). Sti IV 18 24 and St IV 28 34. Chaetotaxy of genu II 2 2/0, 2/0 1; genu III 1 2/1, 2/0 1. Spermatheca (Fig. 10). Calyx dish-shaped, 5 4 (3–5) long. Atrium discrete. Remarks. In the original description of the species, the ventrianal shield is shown smooth, but our examination of the holotype showed it to be reticulate, as also observed in the additional specimens examined. The spermatheca was illustrated, but not described in the original description. The illustration could suggest the calyx to be trumpetshaped. However, our examination of the holotype and of the additional specimens examined suggests the calyx to be dish-shaped and the atrium being difficult to discern. In this publication, this species is considered to have most dorsal shield setae highly variable in length; measurements of the holotype are within the range determined for most setae of the specimens collected in this study, but j 3, J 5, Z 1, s 4, S 4, S 5, r 3 and R 1 of the holotype are shorter than observed in the specimens collected. In addition, differently from the holotype, in some of the specimens collected in this study Z 3 is distinctly serrate.

Neoseiulus fallacis (Garman) Iphidulus fallacis Garman, 1948: 13. Neoseiulus fallacis.��� Moraes et al., 1986: 79; 2004: 119. Specimens examined. Santiago Region. Municipality of San Jos�� de Maipo, on Prunus domestica L., June 2003, associated with Tydeidae. Measurements of a collected female: 380 long 212, maximum width; j 1 22, j 3 50, j 4 40, j 5 37, j 6 55, J 2 69, J 5 10, z 2 50, z 4 52, z 5 30, Z 1 62, Z 4 65, Z 5 82, s 4 67, S 2 65, S 4 60, S 5 45, r 3 32, R 1 42, St IV 62. Remarks. Although the dorsal shield of the specimen collected has about the same dimensions as reported by Schicha & Elshafie (1980) for Australian specimens, its dorsal setae are up to 20% longer., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 329, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Garman, P. (1948) Mite species from apple trees in Connecticut. Connecticut Agricultural Experiment Station, Bulletin, 520, 1 - 27.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Schicha, E. & Elshafie, M. (1980) Four new species of phytoseiid mites from Australia, and three species from America redescribed (Acari: Phytoseiidae). Journal of the Australian Entomological Society, Australia, 19, 27 - 36."]}

Metaseiulus (Metaseiulus) camelliae Chant & Yoshida-Shaul Typhlodromus camelliae Chant & Yoshida-Shaul, 1983: 1053. Typhlodromina camelliae.��� Moraes et al., 1986: 236. Metaseiulus (Metaseiulus) camelliae.��� Moraes et al., 2004: 278. Specimens examined. Santiago Region. Municipality of La Pintana, on Citrus aurantium L., July 2002, associated with Planococcus citri. Municipality of Buin, on Vitis vinifera L., February 1994, associated with Brevipalpus chilensis Barker and June 2002, associated with Tydeidae. Measurements of a collected female: dorsal shield 350 long, 200, maximum width; j 1 20, j 3 20, j 4 15, j 5 12, j 6 17, J2 22, J5 12, z 2 20, z 3 20, z 4 22, z 5 12, Z 4 35, z 5 45, s 4 25, s 6 26, S 5 20, r 3 22, R 1 29. These measurements are similar to those of the original description of the species., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 332, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Chant, D. A. & Yoshida-Shaul, E. (1983) A world review of five similar species groups in the genus Typhlodromus Scheuten. Part II. The conspicuus and cornus groups (Acarina: Phytoseiidae). Canadian Journal of Zoology, 61, 1041 - 1057. https: // doi. org / 10.1139 / z 83 - 138","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1"]}

Amblyseius tamatavensis Blommers Amblyseius (Amblyseius) tamatavensis Blommers, 1974: 144. Amblyseius tamatavensis.��� Moraes et al., 1986: 31; 2004: 52; Abo-Shnaf et al., 2016: 610. Specimens examined. Isla de Pascua, on Thespesia populnea (L.), April 2008, associated with Brevipalpus sp.; on Citrus Limon (L.), April 2008, associated with Brevipalpus sp.. Measurements of a collected female: dorsal shield 350 long, maximum width 210. Setae: j 1 34, j 3 54, j 4 5, j 5 5, j 6 5, J2 4, J5 5, z 2 5, z 4 5, z 5 6, Z 1 6, Z 4 123, Z 5 250, s 4 93, S 2 5, S 4 5, r 3 6, R 1 6, Sge IV 104, Sti IV 84, St IV 71. Remarks. These measurements are similar to those of the original description of the species., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 323, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Blommers, L. (1974) Species of the genus Amblyseius Berlese, 1914, from Tamatave, east Madagascar (Acarina: Phytoseiidae). Bulletin Zoologisch Museum Universiteit van Amsterdam, 3, 143 - 155.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Abo-Shnaf, R. I. A., Sanchez, L. & de Moraes, G. J. (2016) Plant inhabiting Gamasina mites (Acari: Mesostigmata) from the Dominican Republic, with descriptions of four new species of Lasioseius (Blattisociidae) and complementary descriptions of other species. Systematic & Applied Acarology, 21 (5), 607 - 646. https: // doi. org / 10.11158 / saa. 21.5.5"]}

Euseius fructicolus (Gonzalez & Schuster) Amblyseius fructicolus Gonzalez & Schuster, 1962: 12 Euseius fructicolus.��� Moraes et al., 1986: 45; 2004: 69; Lopes et al., 2015: 182; Ferragut & Navia, 2015: 542. Specimens examined. Santiago Region. Municipality of Buin, on Prunus avium (L.), April 1990, associated with Panonychus ulmi (Koch) and February 1991, associated with Bryobia rubrioculus (Scheuten); on Quillaja saponaria Mol., August 2002 and Rafanus sativus L., July 2002, associated with Brevipalpus sp.. Municipality of San Jos�� de Maipo: on Aristotelia chilensis (Mol.), February 2000, associated with Tetranychus urticae Koch; on Citrus limon L., September 2002, associated with Panonychus citri (McGregor). Municipality of Curacav��, on Cissus striata (Ruiz & Pav.), March 1994, associated with Tydeidae; on Persea lingue (Ruiz et Pav.), February 2002 associated with Oligonychus yothersi (McGregor). Municipality La Pintana: on Aloysia triphylla Palau, July 2002 and Corylus avellana L., March 2000, associated with T. urticae; on Aristotelia chilensis (Mol.), August 2002, associated with Tydeidae; on Azara serrata Ruiz & Pav., September 2002; on Melia azedarach Mill., March 2000, Pelargonum hortorum L. H. Bailey, April 2001, Malva nicaensis All., July 2002, Buddleja globosa Hope, July 2002 and Ligustrum sinense Lour., August 2002, associated with Brevipalpus sp.; on Corylus sativa Poit. & Turpin, April 2002 and Schinus poligamus (Cav.), July 2002, associated with Oligonychus sp.; on Crinodendron patagua Mol., July 2002 and Schinus molle L., August 2002, associated with Eriophyidae; on Juglans regia L., March 2000, associated with Oligonychus yothersi (McGregor); on Maytenus boaria Mol., July 2002, associated with Saissetia oleae Oliver; on Malus domestica Borkh., March 2000, associated with Aculus schlechtendali (Nalepa); on Prunus domestica L., March 2002, associated with Panonychus ulmi (Koch) and April 2001, associated with Aculus fockeui (Nalepa & Trouessart); on Prunus salicina Lindl., March 2002, associated with Bryobia rubrioculus (Scheuten); on Vitis vinifera L., April 2001, associated with Oligonychus vitis Zaher & Shehata and February 2002, associated with Colomerus vitis (Pagenstecher). Municipality of Renca, on Solanum lycopersicum L., March 2000, associated with Aculops lycopersici (Masse). Valparaiso Region���Municipality of Los Andes, on V. vinifera L., January1991, associated with Drepanothrips reuteri (Uzel). Municipality of Valparaiso: knoll Miraflores, on Ricinus communis L., August 1999, associated with Tetranychidae. Measurements of a collected female: dorsal shield 325 long, 210 maximum width; j 1 27, j 3 37, j 4 17, j 5 12, j 6 10, J 2 15, J 5 8, z 2 22, z 4 30, z 5 17, Z 1 17, Z 4 20, Z 5 47, s 4 37, S 2 22, S 4 17, S 5 22, r 3 17, R 1 11, Sge IV 32, Sti IV 31, St IV 50. Remarks. These measurents are similar to those of the original description., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 327, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Gonzalez, R. H. & Schuster, R. O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletin Tecnico. Estacion Experimental Agronomica. Universidad de Chile, Facultad de Agronomia, Chile, 16, 1 - 35.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Lopes, P. C., McMurtry, J. A. & de Moraes, G. J. (2015) Definition of the concordis species group of the genus Euseius (Acari: Phytoseiidae), with a morphological reassessment of the species included. Zootaxa, 4048 (2), 174 - 190. https: // doi. org / 10.11646 / zootaxa. 4048.2.2","Ferragut, F. & Navia, D. (2015) Phytoseiid mites (Acari: Phytoseiidae) from Patagonia and Tierra del Fuego. Zootaxa, 3990 (4), 525 - 550. https: // doi. org / 10.11646 / zootaxa. 3990.4.3"]}

Metaseiulus (Metaseiulus) brevicollis Gonzalez & Schuster Metaseiulus brevicollis Gonzalez & Schuster, 1962: 19. Galendromus (Cursoriseius) brevicollis.��� Moraes et al., 1986: 186. Metaseius (Metaseius) brevicollis.��� Moraes et al., 2004: 278. Specimens examined. Santiago Region. Municipality of La Pintana, on Buddleja globosa Hope, July 2002, associated with Brevipalpus sp. Measurements of a collected female: dorsal shield 377 long, 200 maximum width; j 1 25, j 3 35, j 4 20, j 5 20, j 6 27, J 2 30, J 5 13, z 2 27, z 3 27, z 4 30, z 5 20, Z 4 52, Z 5 72, s 4 42, s 6 37, S 5 50, r 3 32, R 1 35. Remarks. These measurements are similar to those reported for the holotype in the original description and in the redescription by Chant &Yoshida-Shaul (1984)., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 332, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Gonzalez, R. H. & Schuster, R. O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletin Tecnico. Estacion Experimental Agronomica. Universidad de Chile, Facultad de Agronomia, Chile, 16, 1 - 35.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1","Chant, D. A. & Yoshida-Shaul, E. (1984) A world review of four species groups in the genus Typhlodromus Scheuten (Acari: Phytoseiidae): brevicollis, luculentis, carinulatus, and pinnatus. Canadian Journal of Zoology, 62, 2631 - 2642. https: // doi. org / 10.1139 / z 84 - 383"]}

Amblyseius herbicolus (Chant) Typhlodromus (Amblyseius) herbicolus Chant, 1959: 84. Amblyseius herbicolus.��� Daneshvar & Denmark, 1982: 5; Moraes et al., 1986: 14; 2004: 27. Specimens examined. Isla de Pascua, on Thespesia populnea (L.), April 2008, associated with Brevipalpus sp.; on Mangifera indica L., April 2008, associated with Tydeidae; on Citrus limon (L.), August 2008, associated with Brevipalpus sp.. Measurements of a collected female: dorsal shield 370 long, maximum width 239. Setae j 1 39, j 3 41, j 4 9, j 5 6, j 6 6, J 2 8, J5 11, z 2 14, z 4 10, z 5 6, Z 1 9, Z 4 94, Z 5 47, s 4 98, S 2 11, S 4 10, S 5 9, r 3 14, R 1 10, Sge IV 110, S ti IV 82, St IV 74. Remarks. These measurements are similar to those of Guanilo et. al. (2008) for specimens from Peru., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 323, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Chant, D. A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species. The Canadian Entomologist, 12 (Supplement), 1 - 166.","Daneshvar, H. & Denmark, H. A. (1982) Phytoseiids of Iran (Acarina: Phytoseiidae). International Journal of Acarology, 8, 3 - 14. https: // doi. org / 10.1080 / 01647958208683272","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp."]}

Phytoseiulus longipes Evans Phytoseiulus longipes Evans, 1958: 307. Mesoseiulus longipes.��� Gonzalez & Schuster, 1962: 18, Moraes et al., 1986: 66. Phytoseiulus longipes.��� Moraes et al., 2004: 167. Specimens examined. Santiago Region. Municipality of La Pintana, on Convolvulus arvensis L., February 2001, associated with Tetranychus desertorum Banks and Tetranychus urticae Koch; and on Marrubium vulgare L., February 2001, associated with T. urticae. Measurements of a collected female: 376 long, 223, maximum width; j 1 20, j 3 88, j 4 15, j 6 87, J 5 4, z 2 32, z 4 93, z 5 9, Z 1 101, Z 4 9, Z 5 94, s 4 123, r 3 50, R 1 73, St IV 97. Remarks. Setal measurements were not given in the original description of this species. The measurements of the specimens collected in this study are close to those given by Gonzalez & Schuster (1962) for specimens also collected in Chile and by Van der Merwe (1968) for specimens from South Africa, neighbour of Zimbabwe, country from where the type specimens were collected. Despite slightly larger than the specimens from South Africa, the specimens collected in this study have setae j 6, Z 1 and Z 5 about 10% shorter (as also reported by Gonzalez & Schuster, 1962) and St IV about 20% longer than South African specimens, differently from what was reported by Gonzalez & Schuster (1962)., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 331, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Evans, G. O. (1958) A new mite of the genus Phytoseiulus Evans (Acarina: Phytoseiidae) from southern Rhodesia. Journal of the Entomological Society of South Africa, 21, 306 - 308.","Gonzalez, R. H. & Schuster, R. O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletin Tecnico. Estacion Experimental Agronomica. Universidad de Chile, Facultad de Agronomia, Chile, 16, 1 - 35.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1","Van der Merwe, G. G. (1968) A taxonomic study of the family Phytoseiidae (Acari) in South Africa with contributions to the biology of two species. Entomology Memoirs, South Africa Department of Agricultural Technical Services, South Africa, 18, 1 - 198."]}

Typhlodromus (Typhlodromus) pyri Scheuten Typhlodromus pyri Scheuten, 1857: 111. Typhlodromus pyri.��� Moraes et al., 1986: 246; 2004: 367. Specimens examined. Maule Region. Municipality of Talca, August 2008, on Corylus avellana L., associated with Tydeidae. Ays��n Region, January 2012, on Juglans regia L., associated with Tydeidae. Chile Chico, Municipality of Coyhaique, February 2013, on Malus domestica L., associated with Bryobia sp., Tydeidae and Oligonychus sp.. Measurements of a collected female: dorsal shield 362 long, 162, maximum width; j 1 25, j 3 30, j 4 17, j 5 17, j 6 17, J 2 20, J 5 7, z 2 17, z 3 27, z 4 27, z 5 17, Z 4 40, Z 5 65, s 4 32, s 6 30, S 2 40, S 4 41, j3 30, R 1 22, St IV 37. Remarks. These measurements are similar to those reported by Chant & Yoshida-Shaul (1987) for the neotype., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 334, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Scheuten, A. (1857) Einiges uber Milben. Archivfur Naturgeschichte, 23, 104 - 112.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp."]}

Neoseiulus anonymus (Chant & Baker) Amblyseius anonymus Chant & Baker, 1965: 21. Neoseiulus anonymus.��� Moraes et al., 1986: 68; 2004: 102. Specimens examined. Santiago Region. Municipality of Pudahuel, on Convulvulus arvensis L., March 2011 and Marrubium vulgare L., May 2011, associated with Tetranychus urticae Koch. Measurements of a studied female: dorsal shield 341 long, 154 maximum width; j 1 25, j 3 56, j 4 50, j 5 50, j 6 62, J 2 56, J 5 12, z 2 53, z 4 50, z 5 37, Z 1 56, Z 4 75, Z 5 76, s 4 55, S 2 68, S 4 35, S 5 30, r 3 37, R 1 43, St IV 50. Remarks. Most of these measurements of the specimen studied in this work are slightly longer than reported by Schicha & Elshafie (1980) for the holotype of N. anonymus, which is compatible with the larger size of the former; the exceptions are the slightly shorter S 4 and macroseta of tarsus IV (respectively 43 and 54���67 in the holotype)., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 328, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Chant, D. A. & Baker, E. W. (1965) The Phytoseiidae (Acarina) of Central America. Memoirs of the Entomological Society of Canada, 41, 1 - 56.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A Catalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, 353 pp.","Schicha, E. & Elshafie, M. (1980) Four new species of phytoseiid mites from Australia, and three species from America redescribed (Acari: Phytoseiidae). Journal of the Australian Entomological Society, Australia, 19, 27 - 36."]}

Chileseius paracamposi Yoshida-Shaul & Chant Chileseius paracamposi Yoshida-Shaul & Chant, 1991: 95. Chileseius paracamposi.��� Moraes et al., 2004: 58; Ferragut & Navia, 2015: 527. Specimens examined. Magallanes Region. Municipality of Punta Arenas, on Nothophagus pumilio (P. et. E.), November 2012., Published as part of Trincado, R. D., Martin, J. P. I., M��ndez Rosa, D. D., Lopes, P. C. & De Moraes, G. J., 2018, Phytoseiid mites (Acari: Phytoseiidae) from Chile, with descriptions of three new species and a redescription of Chileseius camposi, pp. 322-340 in Zootaxa 4482 (2) on page 327, DOI: 10.11646/zootaxa.4482.2.5, http://zenodo.org/record/1440610, {"references":["Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434 (1), 1 - 494. https: // doi. org / 10.11646 / zootaxa. 434.1.1","Ferragut, F. & Navia, D. (2015) Phytoseiid mites (Acari: Phytoseiidae) from Patagonia and Tierra del Fuego. Zootaxa, 3990 (4), 525 - 550. https: // doi. org / 10.11646 / zootaxa. 3990.4.3"]}

Amblyseius tamatavensis Blommers, 1974 Amblyseius tamatavensis Blommers 1974: 144; Denmark and Muma 1989: 13; Moraes et al. 1991: 119. Amblyseius maai Tseng 1976: 123 (synonymy according to Denmark and Muma, 1989). Amblyseius aegyptiacus Matthysse and Denmark 1981: 343. Specimens examined — Pernambuco, Bonito, 14 August 2005 (1 ♀), 4 September 2005 (1 ♀; 2 ♂) and 23 October 2005 (6 ♀; 4 ♂);`gua Preta, 10 September 2005 (1 ♀); CearAE, Paraipaba, 5 September 2005 (1 ♀) on Annona muricata. Pernambuco, Bonito, 14 August 2005 (13 ♀; 9 ♂) on Annona squamosa., Published as part of Sousa, J. M. de, Gondim Jr., M. G. C., Lofego, A. C. & Moraes, G. J. de, 2015, Mites on Annonaceae species in northeast Brazil and in the state of Para, pp. 5-18 in Acarologia 55 (1) on page 7, DOI: 10.1051/acarologia/20152147, http://zenodo.org/record/5402210, {"references":["Blommers L. 1974 - Species of the genus Amblyseius Berlese, 1914, from Tamatave, east Madagascar (Acarina: Phytoseiidae) - Bull. Zool. Mus. Univ. Amst., 3: 143 - 155.","Denmark H. A., Muma M. H. 1989 - A revision of the genus Amblyseius Berlese, 1914 (Acari: Phytoseiidae) - Occas. Pap. Fla. State Collect. Arthropods, 4: 1 - 149.","Moraes G. J., Mesa N. C., Braun A. 1991 - Some Phytoseiid mites of Latina America (Acari: Phytoseiidae) - Int. J. Acarol., 17: 117 - 139.","Tseng Y. H. 1976 - Systematics of the mite family Phytoseiidae from Taiwan, with a revised key to genera of the world (II) - J. Agr. Assoc. China, 94: 85 - 128."]}

Neoseiulus californicus (McGregor) Typhlodromus californicus McGregor, 1954: 89 Amblyseius californicus, Schuster & Pritchard, 1963: 271 Neoseiulus californicus, Moraes et al., 2004: 109; Chant & McMurtry, 2007: 25; Guanilo et al., 2008: 27 Measurements of 5 adult females collected are: dorsal shield 359 (340 – 375) long and 174 (168 – 187) wide; j1 22 (20 – 23), j3 30 (25 – 33), j4 25 (23 – 28), j5 24 (20 – 27), j6 28 (25 – 33), J2 31 (28 – 33), J5 13 (13 – 15), z2 25 (23 – 28), z4 28 (25 – 33), z5 24 (23 – 28), Z1 36 (33 – 40), Z4 50 (48 – 53), Z5 65 (55 – 73), s4 32 (30 – 38), S2 40 (38 – 45), S4 37 (35 – 38), S5 31 (30 – 33), r3 25 (24 – 25), R1 23 (22 – 23); distances between st1 – st3 63 (53 – 68), st2 – st2 63 (53 – 68) and st5 – st5 67 (63 – 73); ventrianal shield 119 (113 – 127) long, 86 (95 – 105) wide at level of ZV2; calyx of spermatheca 10 (9 – 10) long; St IV 52 (48 – 60). Fixed and movable cheliceral digits with 3 and 2 teeth, respectively. Chaetotaxy — genu II 2, 2/0-2/0, 1; genu III 1, 2/1-2/0, 1. Remarks — Measurements of the specimens collected are similar to what was reported in redescriptions of the species based on specimens from California (Schuster & Pritchard, 1963) and Peru (Guanilo et al., 2008), except for the anterolateral dorsal shield setae and Z5, about 10 % shorter in the specimens collected. Specimens collected — More than 50 specimens on Solanum aethiopicum L. and S. melongena L., infested by T. evansi and T. urticae, at Malika, between March and July 2010. World distribution — This species has been reported from subtropical to temperate areas in the Americas, Europe, Asia and northern Africa. A few years ago (Pringle & Heunis, 2006) it was introduced to South Africa for the control of mite pests in apple orchards; thus, our finding is the second record of this species in sub-Saharan Africa (Zannou et al., 2006)., Published as part of Kade, N., Gueye-Ndiaye, A., Duverney, C. & de Moraes, G. J., 2011, PHYTOSEIID MITES (ACARI: PHYTOSEIIDAE) FROM SENEGAL Niokhor K, Absa G - N, Claude D and Gilberto J. de M, pp. 133-138 in Acarologia 51 (1) on pages 135-136, DOI: 10.1051/acarologia/20112001, http://zenodo.org/record/5392488, {"references":["McGregor E. A. 1954 - Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae) - South. Calif. Acad. Sci. Bull., 53: 89 - 92.","Schuster R. O., Pritchard A. E. 1963 - Phytoseiid mites of California - Hilgardia, 34: 191 - 285.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: pp. 494.","Chant D. A., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - West Bloomfield: Indira Publishing House, pp. 219.","Guanilo A. D., Moraes G. J. de, Knapp M. 2008 - Phytoseiid mites (Acari: Phytoseiidae) from Peru, with descriptions of four new species - Zootaxa, 1880: 1 - 47.","Pringle K. L., Heunis J. M. 2006 - Biological control of phytophagous mites in apple orchards in the Elgin area of South Africa using the predatory mite, Neoseiulus californicus (McGregor) (Mesostigmata: Phytoseiidae): a benefit-cost analysis - Afr. Entomol., 14: 113 - 121.","Zannou I. D., Moraes G. J. de, Ueckermann E. A., Oliveira A. R., Yaninek J. S., Hanna R. 2006 - Phytoseiid mites of the genus Neoseiulus Hugues (Acari: Phytoseiidae) from sub-Saharan Africa - Int. J. Acarol., 32: 241 - 276. doi: 10.1080 / 01647950608684467"]}

Paraphytoseius horrifer (Pritchard & Baker) Amblyseius (Ptenoseius) horrifer Pritchard & Baker, 1962: 295 Paraphytoseius horrifer, Moraes et al., 2004: 161; Chant & McMurtry, 2007: 49; Moraes et al., 2007: 10 Measurements of 2 adult females collected are: dorsal shield 300 ��� 304 long and 163 ��� 170 wide; j1 35 ��� 38, j3 83 ��� 85, j4 and j5 3 ��� 5, j6 5 ��� 6, J5 3 ��� 5, z2 8 ��� 10, z4 8 ��� 10, z5 3 ��� 5, Z1 7 ��� 9, Z4 73 ��� 75, Z5 95 ��� 100, s4 118 ��� 125, r3 43 ��� 45, R1 30 ��� 32; distances between st1 ��� st3 68 ��� 70, st2 ��� st2 68 ��� 69 and st5 ��� st5 85 ��� 88; ventrianal shield 113 ��� 115 long, 68 ��� 70 wide at level of ZV2, 65 wide at anus level; Sge II 13 ��� 15, Sge IV 28 ��� 30, Sti IV 32 ��� 38, St (basitarsus) IV 43 ��� 45, St (telotarsus) 33 ��� 35. Chaetotaxy ��� genu II 2, 2/0-2/0, 1; genu III 1, 2/1-2/0, 1. Remarks ��� Measurements of the specimens collected are similar to those given by Moraes et al. (2007) for specimens from sub-Saharan Africa, including the holotype. The presence of a macroseta on genu II and not on genu III, observed in the specimens collected and in the specimens examined by Moraes et al. (2007) is not common for other phytoseiid mites. Usually, the presence of a macroseta on a given leg implies its presence on subsequent legs. Specimens collected ��� 28 females and 9 males on Centaurea perrotettii DC., Chrosophora senegalensis Vis. and Urena lobata L, at KBD, in November 2009. World distribution ��� This species has been reported almost only from sub-Saharan Africa; the exception refers to a single report from India (Gupta, 1981)., Published as part of Kade, N., Gueye-Ndiaye, A., Duverney, C. & de Moraes, G. J., 2011, PHYTOSEIID MITES (ACARI: PHYTOSEIIDAE) FROM SENEGAL Niokhor K, Absa G - N, Claude D and Gilberto J. de M, pp. 133-138 in Acarologia 51 (1) on page 136, DOI: 10.1051/acarologia/20112001, http://zenodo.org/record/5392488, {"references":["Pritchard A. E., Baker W. E. 1962 - Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world - Hilgardia, 33: 205 - 309.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: pp. 494.","Chant D. A., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - West Bloomfield: Indira Publishing House, pp. 219.","Moraes G. J. de, Zannou I. D., Ueckermann E. A., Oliveira A. R., Yaninek J. S., Hanna R. 2007 - Phytoseiid mites of the tribes Afroseiulini, Kampimodromini and Phytoseiulini, and complementary notes on mites of the tribes Euseiini and Neoseiulini (Acari: Phytoseiidae) from sub-Saharan Africa - Zootaxa, 1628: 1 - 22.","Gupta S. K. 1981 - Phytoseiidae (Acari) of Kumaon and Garhwal Himalaya. Contribution to acarology in India - Proceedings of the I All India Symposium of Acarology. April 23 - 25, 1979. Bangalore, India, p. 21."]}

Neoseiulus barkeri Hughes Neoseiulus barkeri Hughes, 1948: 141; Moraes et al., 2004: 104; Chant & McMurtry, 2007: 22; Zannou et al., 2006: 249. Measurements of the single adult female collected are: dorsal shield 355 long and 195 wide; j1 15, j3 18, j4 and j5 15, j6 18, J2 19, J5 10, z2 19, z4 20, z5 16, Z1 21, Z4 33, Z5 45, s4 23, S2 23, S4 20, S5 18, r3 15, R1 13; distances between st1 – st3 63, st2 – st2 65 and st5 – st5 55; ventrianal shield 113 long, 90 wide at level of ZV2; calyx of spermatheca 25 long; St IV 55. Chaetotaxy —genu II 2, 2/0-2/0, 1; genu III 1, 2/1-2/0, 1. Remarks — Measurements of the only specimen collected fit well those given by Zannou et al. (2006) for specimens from sub-Saharan Africa. Specimens collected — 1 female, on Solanum aethiopicum L., at Malika, in March 2007., Published as part of Kade, N., Gueye-Ndiaye, A., Duverney, C. & de Moraes, G. J., 2011, PHYTOSEIID MITES (ACARI: PHYTOSEIIDAE) FROM SENEGAL Niokhor K, Absa G - N, Claude D and Gilberto J. de M, pp. 133-138 in Acarologia 51 (1) on page 135, DOI: 10.1051/acarologia/20112001, http://zenodo.org/record/5392488, {"references":["Hughes A. M. 1948 - The mites associated with stored food products. London: Ministry of Agriculture and Fisheries, H. M. Stationary Office, pp. 168.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: pp. 494.","Chant D. A., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - West Bloomfield: Indira Publishing House, pp. 219.","Zannou I. D., Moraes G. J. de, Ueckermann E. A., Oliveira A. R., Yaninek J. S., Hanna R. 2006 - Phytoseiid mites of the genus Neoseiulus Hugues (Acari: Phytoseiidae) from sub-Saharan Africa - Int. J. Acarol., 32: 241 - 276. doi: 10.1080 / 01647950608684467"]}

Ueckermannseius saltus (Denmark & Matthysse) (Fig. 20) Euseius saltus Denmark & Matthysse, in Matthysse & Denmark, 1981: 348 Amblyseius (Amblyseius) saltus, Ueckermann & Loots, 1988: 87 Amblyseius saltus, Moraes et al., 1989: 82 Typhlodromalus saltus, Moraes et al., 2004: 203 Ueckermannia saltus, Chant & McMurtry, 2005 a: 203 FEMALE ��� (Specimens measured ��� Benin: 3; Burundi: 4; Cameroon: 3; Ghana: 4; Kenya: 5; Malawi: 3; Mozambique: 2; Nigeria: 3; Sierra��Leone: 3; Democratic Republic of Congo: 2). Dorsum ��� Dorsal shield mostly smooth, with few striae along anterolateral margins, 344 (320���388) long and 228 (188���263) wide. Setae j 1 25 (22���28), j 3 25 (19���28), j 4 8 (5���10), j 5 7 (5���10), j 6 11 (8���13), J 2 13 (10���16), J 5 9 (6���13), z 2 17 (14���20), z 4 16 (13���19), z 5 10 (6���13), Z 1 15 (11���18), Z 4 17 (15���21), Z 5 23 (20��� 26), s 4 19 (16���25), S 2 18 (15���21), S 4 17 (14���20), S 5 17 (14���20), r 3 18 (14���25), R 1 17 (14���21). All setae smooth. Peritreme ��� Extending almost to j 1. Venter ��� Distances between St 1 ���St 3 62 (56���68), St 2 ���St 2 66 (62���70), St 5 ���St 5 69 (62��� 78). Ventrianal shield 100 (88���108) long, 53 (48���58) wide at level of Zv 2 and 70 (61���75) wide at level of anus, with a pair of elliptical pores posterior to and slightly mesad of Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 31 (30���33) long, with 4 teeth; fixed digit 30 (28���30) long, with 10 teeth and a discernibe pilus dentilis. Spermatheca ��� Calyx bulged near atrium, followed by a slender region and flared distally, 34 (19���50) long; atrium distinct. Legs ��� Macrosetae sharp��tipped; Sge II 16 (13���19), Sge III (24���32), Sti III 32 (26��� 38), Sge IV 35 (27���40), Sti IV 51 (42���58), St IV 53 (48���61). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. SPECIMENS EXAMINED: Benin: 2 km S Cotiakou, on Butyrospermum parkii, 20 �� XI�� 1989, A. Onzo; 14 km S Tamarou, on Manihot esculenta, 22 ��XI�� 1989, A. Onzo; Gamia, on unknown plant, 22 ��XI�� 1989, G.J. Moraes. Burundi: 16 km NW Bujumbura, on Solanum capsicoides, 11 ��XII�� 1989, J.S. Yaninek; ISABU, Bujumbura, on Bauhinia sp., 11 ��XII�� 1989, J.S. Yaninek; 16 km NW Bujumbura, on Ricinus communis, 11 ��XII�� 1989, J.S. Yaninek; 17 km SW Makamba, on unknown plant, 14 ��XII�� 1989, J.S. Yaninek. Cameroon: 11 km W Diang, on M. esculenta, 04��II�� 1991, L. Louis; Manga, on M. esculenta, 19 ��XII�� 1995, L. Louis. Democratic Republic of Congo: 3 km E Lukula, Bas�� Congo, on M. esculenta, 12 ��VII�� 1991, A. Onzo; Nkolo, Bas��Congo, on M. esculenta, 12 �� VII�� 1991, A. Onzo. Ghana: 3 km NE Suhum, on Colocasia sp., 08��XI�� 1989, G.J. Moraes; 18 km N Kumasi, on M. esculenta, 09��XI�� 1989, J.S. Yaninek; 3 km NW Aboi, on M. esculenta, 11 ��XI�� 1989, J.S. Yaninek. Kenya: 17 km N Lunga Lunga, on M. esculenta, 28 �� XI�� 1989, J.S. Yaninek; 17 km N Lunga Lunga, on Physalis angulata, 28 ��XI�� 1989, J.S. Yaninek; 3km N Lunga Lunga, on M. esculenta, 28 ��XI�� 1989, J.S. Yaninek; 5 km N Mutunga, on M. esculenta, 30 ��XI�� 1989, J.S. Yaninek. Malawi: 12.9 km SE Mzuzu, on M. esculenta, 20 ��V�� 2002, I.D. Zannou; 10 km NW Nkhata��Bay��Mzuzu junction, on Ageratum conyzoides, 20 ��V�� 2002, M.J. Chitete. Mozambique: 5.9 km N Unguana, on M. esculenta, 17 ��IV�� 2001, I.D. Zannou; 8 km S Movia, Nacala��Velha, on Mucuna sp., 29 ��IV�� 2001, I.D. Zannou. Nigeria: IITA Station, Ibadan, on M. esculentus, 14 ��XII�� 1983, J.S. Yaninek, IITA Station, Ibadan, on M. esculenta, 05��IX�� 1996, J. Adisa; Alabata, on M. esculenta, 30 ��XI�� 1996, Joel. Sierra��Leone: 36 km N Makeni, on M. esculenta, 16 ��XII�� 1991, A. F. Kanu; Lunsar hospital, on carica papaya, 16 ��XII�� 1991, B. Kristensen; Lunsar Mabesoh, on unknown plant, 16 ��XII�� 1991, M.M. Bocharie. WORLD DISTRIBUTION ��� Benin, Burundi, Cameroon, Democratic Republic of Congo, Ghana, Kenya, Malawi, Mozambique, Nigeria, Sierra��Leone., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 38-40, DOI: 10.5281/zenodo.171564, {"references":["Matthysse, J. G. & Denmark, H. A. (1981) Some phytoseiids of Nigeria (Acarina: Mesostigmata). The Florida Entomologist, USA, 64, 340 - 357.","Ueckermann, E. A. & Loots, G. C. (1988) The African species of the subgenera Anthoseius De Leon and Amblyseius Berlese (Acari: Phytoseiidae). Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa, South Africa, 73, 168 pp.","Moraes, G. J. de, McMurtry, J. A., van den Berg, H. & Yaninek, J. S. (1989) Phytoseiid mites (Acari: Phytoseiidae) of Kenya, with descriptions of five new species and complementary descriptions of eight species. International Journal of Acarology, USA, 15 (2), 79 - 93.","Moraes, G. J. de, McMurtry, J. A., Denmark, H. A., Campos, C. B. (2004) A revised catatalog of the mite family Phytoseiidae. Zootaxa, New Zealand, 434, 1 - 494.","Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224."]}

Ueckermannseius neohavu Moraes, Zannou & Oliveira, sp. nov. (Fig. 18) DIAGNOSIS ��� This species is distinct by having Z 5 ca. 3.7 times as long as S 5, Jv 4 present and Jv 5 ca. 70 % as long as ventrianal shield and macrosetae of legs III and IV sharp��tipped. FEMALE ��� (5 specimens measured). Dorsum ��� Dorsal shield mostly smooth, with reticulations along anterolateral margins and few striae and reticulations on posterior half, 416 (403���432) long and 291 (278���301) wide. Setae j 1 40 (38���42), j 3 26 (24���28), j 4 19 (18���20), j 5 18 (16���18), j 6 15 (15���16), J 2 15 (14���15), J 5 7 (6���8), z 2 24 (22���25), z 4 25 (24���25), z 5 15 (14���16), Z 1 17 (16���18), Z 4 16 (14���17), Z 5 52 (48���58), s 4 27 (25���28), S 2 15 (14���17), S 4 14 (13���15), S 5 14 (13���15), r 3 19 (18���20), R 1 17 (16���18). All setae smooth. Peritreme ��� Extending to level between j 1 and j 3. Venter ��� Distances between St 1 ���St 3 69 (66���70), St 2 ���St 2 74 (72���76), St 5 ���St 5 78 (72��� 86). Ventrianal shield 128 (122���134) long, 74 (71���75) wide at level of Zv 2 and 73 (71���74) wide at level of anus, with a pair of elliptical pores posterior to and slightly mesad of Jv 2. One pair of metapodal shields; Caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 46 (45���47) long, with at least 4 teeth; fixed digit 36 (35���36) long, with at least 11 teeth and a discernible pilus dentilis. Spermatheca ��� Calyx with distal 3 / 5 slender and basal 2 / 5 bulged, 49 (46���53) long; atrium distinct. Legs ��� Macrosetae sharp��tipped; Sge II 30 (29���32), Sge III 46 (45���47), Sti III 38 (35��� 39), Sge IV 74 (72���76), Sti IV 58 (55���61) and St IV 73 (70���77). Chaetotaxy of genu III: 1�� 2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype and 4 paratypes females from Cassia sp., 11 km W Diang, Cameroon, 04��II�� 1991, L. Louis deposited at ESALQ��USP. One paratype female, same data as holotype, deposited at IITAIM. ETYMOLOGY ��� The epithet neohavu refers to the similarity between this species and U. havu. REMARKS ��� This new species resembles U. havu, but the latter differs by having longer setae z 2, z 4 and s 4, knobbed Jv 5, and knobbed macrosetae on legs III and IV and by lacking Jv 4. Ueckermannseius eastafricae Moraes, Zannou & Oliveira, sp. nov. and U. macrosetosus are also similar to the new species, but the former has Z 5 ca. 2.5 times as long as S 5 and knobbed macrosetae on legs III and IV, and the latter has shorter peritreme (extending between j 3 and z 2) and knobbed macrosetae on legs III and IV., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 34-36, DOI: 10.5281/zenodo.171564

Ueckermannseius parahavu Moraes, Zannou & Oliveira, sp. nov. (Fig. 19) DIAGNOSIS ��� This species is characterized by s 4 ca. 75 % as long as distance between its base and the base of z 5; j 4, j 6 and J 2 shorter than a third the distance between bases of their respective homologues of both sides, peritreme extending beyond base of j 3 and Jv 4 present. FEMALE ��� (11 specimens measured). Dorsum ��� Dorsal shield mostly smooth, with few striae along anterolateral margins, 356 (328���371) long and 237 (221���253) wide. Setae j 1 33 (30���37), j 3 35 (32���42), j 4 12 (11���13), j 5 12 (10���14), j 6 12 (10���14), J 2 12 (11���14), J 5 8 (5���10), z 2 21 (19���24), z 4 23 (16���27), z 5 13 (10���16), Z 1 13 (11���16), Z 4 13 (10���14), Z 5 21 (18��� 26), s 4 30 (26���35), S 2 15 (11���18), S 4 11 (10���14), S 5 12 (8���14), r 3 18 (16���22), R 1 14 (11���16). All setae smooth. Peritreme ��� Extending to level of j 1. Venter ��� Distances between St 1 ���St 3 70 (67���74), St 2 ���St 2 62 (58���66), St 5 ���St 5 70 (64��� 78). Ventrianal shield 114 (96���128) long, 56 (51���64) wide at level of Zv 2 and 60 (58���67) wide at level of anus, with a pair of elliptical pores posterior to and slightly mesad of Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 35 (35���36) long, with 1 large tooth and 3 small teeth; fixed digit 30 (30���31) long, with 11 teeth and a discernible pilus dentilis. Spermatheca ��� Calyx tubular, with distal half slender and lighly sclerotized and proximal half bulged, 32 (27���35) long; atrium distinct. Legs ��� Macrosetae sharp��tipped; Sge II 25 (22���27), Sge III 33 (30���37), Sti III 27 (26��� 29), Sge IV 50 (45���53), Sti IV 42 (38���46), St IV 64 (58���70). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype female and 1 paratype female from unknown plant, 15 km W Nsuta, Brong Ahafo, Ghana, 09��XI�� 1989, J.S. Yaninek, deposited at ESALQ��USP; 3 paratype female from unknown plant, 12 km E Obuasi, Ashanti, Ghana, 10 ��XI�� 1989, J.S. Yaninek, deposited at ESALQ��USP. One paratype female from Securinega virosa, Agbotagon, Benin, 17 ��VI�� 1991, B. Eklou, deposited at IITAIM; 1 paratype female from Byrsocarpus coccineus, Agbotagon, Benin, 12 ��VIII�� 1991, B. Eklou, deposited at ESALQ��USP; 1 paratype female from Albizia adianthifolia, Ikpinle, Benin, 11 ��XI�� 1991, A. Onzo, deposited at ESALQ��USP. One paratype female from Mangifera indica, 11 km W Diang, Cameroon, 04��I�� 1991, L. Louis, deposited at IITAIM. One paratype female from unknown plant, 2 km N Namalonge, Uganda, 13 ��X�� 1990, J.S. Yaninek, deposited at IITAIM. ETYMOLOGY ��� The epithet parahavu refers to similarity between this new species and T. havu. REMARKS ��� Ueckermannseius bundibugyoensis Moraes, Zannou & Oliveira, sp. nov. is similar to this new species, but it differs by having j 4, j 6 and J 2 longer, each at least half as long as distance between bases of their respective homologues. Ueckermannseius havu, U. macrosetosus and U. neohavu Moraes, Zannou & Oliveira, sp. nov. also resemble this new species, but they differ by having shorter peritreme (not reaching j 3) and longer Z 5. The first 2 species further differ by having knobbed macrosetae on leg IV, whereas the last species further differs by having some striation on posterior half of dorsal shield and anterior third of ventrianal shield, in addition U. havu differs by lacking Jv 4. Ueckermannseius saltus (Denmark & Matthysse) is also similar to this new species, but differs by having most setae near the margin of the dorsal shield similar in length., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 36-38, DOI: 10.5281/zenodo.171564

Tenuipalpus isabelae n. sp. (Figure 1 A���D) Diagnosis: This species belongs to the keiensis subgroup of the proteae group, established by Meyer (1993), whose adult females are characterized by having 6 pairs of dorsolateral opisthosomal setae, one pair of Ic 3 and 2 pairs of Ic 4. Females of this new species resemble T. barticanus De Leon (1965), but differ by having 2 setae each on genua I and trochanter III (one seta each in T. barticanus), and prodorsal seta sc 2 about 4 times longer than in T. barticanus. Par�� Veiga & Flechtmann (1980) Adult female: (Specimens measured: 5). Subcapitulum (rostrum) extending to base of femur I. Palpus (Fig. 1 C) with three segments; proximal segment without setae, intermediate segment with a dorsodistal pectinate seta; distal segment with one eupathidium 8 (7���9). Idiosoma (Fig. 1 A) 273 (260���282) long, 196 (187���200) wide. Rostral shield deeply cleft, with pointed lobes. Propodosoma with anterior margin irregular; dorsocentral region delimitated by two longitudinal grooves, with transversal striation; lateral regions with diagonal striation. Setae v 2 7, sc 1 8, smooth; sc 2 43 (40���50), thick and serrate. Propodosomal pores absent. Dorsal surface of hysterosoma with 2 curved, roughly longitudinal grooves; region between grooves, anterior to opisthosomal pores, with transverse striation; remaining surface primarily smooth. Setae c 1 5 (4���8), c 3 16 (14���20), d 3 8 (6���9), f 2 27 (25���30), f 3 25 (23���28), h 1 19 (18���22), all serrate; h 2 100 (90���110) flagelliform, serrate proximally. Setae d 1, e 1, e 3 absent. Ventral surface (Fig. 1 D) with fine longitudinal striae between Ic 1 and Ic 2 and between Ic 3 and Ic 4. Genitoventral plate lightly sclerotized, with fine transversal striae. Setae Ic 3 a 8 (7���9), Ic 4 a 62 (50���80) and Ic 4 b 55 (50���60), ag 19 (15���23); g 1 and g 2 18 (15���23), on a transversal line. Chaetotaxy of legs I to IV: coxa 2 / 2 / 1 / 1, trochanter 1 / 1 / 2 / 1, femur 4 / 4 / 2 / 1, genu 2 / 2 /0/ 0, tibia 5 / 5 / 3 / 3; tarsi I and II each with a serrate seta overlying distal solenidium. Adult male: unknown. Deutonymph: Idiosoma (Fig. B) 210 long, 185 wide. Setae v 2 5, sc 1 7, c 1 2, d 3 6 smooth, sc 2 40, c 3 22, f 2 33, f 3 27, h 1 19, all serrate; h 2 50, flagelliform, serrate proximally. Dorsal seta e 3 absent. Type material: Holotype female, 3 paratype females and 1 paratype deutonymph, ex Actinostemon sp. (Euphorbiaceae), VIII�� 2002, N.C. Mesa, Reserva Ecol��gica de Ibicatu, Piracicaba���SP, Brazil (22 o 46 ��� 43 ���S and 47 o 49 ��� 32 ���W). Deposited in the reference collection of Departamento de Entomologia, Fitopatologia e Zoologia Agr��cola, Setor Zoologia, Escola Superior de Agricultura Luiz de Queiroz, Universidade de S��o Paulo, Piracicaba���SP, Brazil. One paratype female, same collection data, deposited in the National Museum of Natural History, Smithsonian Institution, Washington, DC, USA. Etymology: This species is named in honor Isabel Cristina Zuluaga Mesa, daughter of the senior author., Published as part of Mesa, N. C., De Moraes, G. J. & Ochoa, R., 2006, Two new species of Tenuipalpus (Acari: Tenuipalpidae) from southeastern Brazil, pp. 45-51 in Zootaxa 1138 on pages 45-48, DOI: 10.5281/zenodo.172028, {"references":["Meyer, M. (1993) A revision of the genus Tenuipalpus Donnadieu (Acari: Tenuipalpidae) in the Afrotropical region. Entomology Memoir Department of Agriculture Repub. South Africa, 88, 1 - 84.","De Leon, D. (1965) New Tenuipalpidae (false spider mites) from British Guiana with notes on four described species. Florida Entomologist, 48, 65 - 75.","Veiga, A. F. de S. L. & Flechtmann, C. H. W. (1980) O acaro plano Tenuipalpus pacificus Baker 1945 (Acari, Prostigmata: Tenuipalpidae) em avencas e samambaias (Pteridophyta) em Pernambuco e Para, Brasil. Anais da Sociedade Entomologica do Brasil, 9, 155 - 158."]}

Genus Amblydromalus Chant & McMurtry Amblydromalus Chant & McMurtry, 2005 a: 203 Amblyseius (Amblyseius) section Amblydromus Wainstein, 1962: 15 (preoccupied by Amblydromus Muma, 1961: 297) Amblyseius limonicus species group Moraes et al., 1994: 209 Amblyseius (Typhlodromalus) Muma, 1961: 288 (part) Typhlodromalus, De Leon, 1966: 87 (part) Type species — Amblyseius limonicus Garman and McGregor, 1956: 9 The species of this genus that are reported in the present study fit the description of the limonicus species group of Chant & McMurtry (2005 a), except for slight differences which are mentioned under the new species here described.

Ueckermannseius danhomeensis Moraes, Zannou & Oliveira, sp. nov. (Fig. 15) DIAGNOSIS ��� This species is different from the other Ueckermannseius species by having funnel��shaped calyx of spermatheca. FEMALE ��� (3 specimens measured). Dorsum ��� Dorsal shield smooth, with few striae along lateral margins of proscutum, 359 (353���368) long and 238 (228���243) wide. Setae j 1 30, j 3 33 (33���35), j 4 9 (8���10), j 5 9 (8��� 10), j 6 9 (8���10), J 2 12 (11���13) J 5 8, z 2 13, z 4 10 (10���11), z 5 9 (8���10), Z 1 12 (11���13), Z 4 13, Z 5 51 (50���53), s 4 15 (15���16), S 2 13 S 4 13, S 5 12 (11���13), r 3 15 (14���15), R 1 10 (10���11). All setae smooth, except Z 5 which is lightly serrate. Peritreme ��� Extending almost to j 1. Venter ��� Distances between St 1 ���St 3 58 (56���60), St 2 ���St 2 78 (75���80), St 5 ���St 5 80. Ventrianal shield 117 (117���118) long, 56 (55���56) wide at level of Zv 2 and 67 (67���68) wide at level of anus, with a pair of elliptical pores slightly posteromesad of Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 35 long, with 4 teeth; fixed digit 32 (30���33) long, with 10 teeth and a distinguishable pilus dentilis. Spermatheca ��� Calyx funnel��shaped, 24 (23���24) long; atrium distinctly bulbous. Legs ��� Macrosetae sharp��tipped; Sge I 27 (25���29), Sge II 35, Sge III 45 (43���48), Sti III 33 (32���35), Sge IV 68 (65���73), Sti IV 49 (46���50), St IV 98 (96���100). Chaetotaxy of genu III: 1��2 / 0,2 / 1 �� 1. MALE ��� (1 specimen measured). Dorsum ��� Dorsal shield pattern as in female, 255 long and 170 wide. Setae j 1 23, j 3 48, j 4 8, j 5 5, J 2 8, J 5 5, z 2 10, z 4 8, z 5 8, Z 1 8, Z 4 8, s 4 13, S 2 10, S 4 10, S 5 8, r 3 10 (setae j 6, Z 5 and R 1 broken in available specimen). All setae smooth, except Z 5 which is lightly serrate. Peritreme ��� Extending to level between j 1 and j 3. Venter ��� Ventrianal shield subtriangular, with a small contriction at anus level and transversal striae anteriorly; 105 long, 148 wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral) and a pair of elliptical pores slightly posteromesad of Jv 2. Spermatodactyl ��� Shaft 28 long. Legs ��� Macrosetae sharp��tipped; Sge I 23, Sge II 25, Sge III 35, Sti III 28, Sge IV 50, Sti IV 40 and St IV 75. Chaetotaxy of genu III same as in female. LOCALITY AND TYPE MATERIAL ��� Holotype female, allotype male and 2 paratypes females from Albizia adianthifolia, Agbotagon, Benin; 18 ��XI�� 1991, A. Onzo, deposited at ESALQ��USP. ETYMOLOGY ��� The epithet danhomeensis refers to the name of the former most powerful kingdom of the region presently known as Benin, where the types of this new species were collected. REMARKS ��� The spermatheca of this new species does not fit the types of spermathecae mentioned by Chant & McMurtry (2005 a) as typical for the genus. Ueckermannseius aequidens, Ueckermannseius munsteriensis (Van der Merwe, 1965) and Ueckermannseius ultimus (Chant & Baker 1965) resemble this new species, but differ by having calyx of spermatheca longer and tubular. Ueckermannseius saltus (Matthysse & Denamrk) is also close to this new species, but it differs by having shorter Z 5 and calyx bulged near atrium, slender medially and flared distally., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 28-29, DOI: 10.5281/zenodo.171564, {"references":["Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224.","Van der Merwe, G. G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, South Africa, 28, 57 - 76.","Chant, D. A. & Baker, E. W. (1965) The Phytoseiidae (Acarina) of Central America. Memoirs of the Entomological Society of Canada, Canada, 41, 56 pp."]}

Euseius nyanzaensis Moraes, Zannou & Oliveira, sp. nov. (Fig. 22) DIAGNOSIS — This species is characterized by a combination of the following characteristics: short peritreme (not reaching z 2), Jv 1 inserted near anterior margin of ventrianal shield, Jv 4 present and macrosetae blunt on tibiae and with a tiny knob on genua. FEMALE — (1 specimen measured). Dorsum — Dorsal shield mostly smooth, striate along lateral margins anterior to S 2, 336 long and 216 wide. Setae j 1 26, j 3 18, j 4 13, j 5 13, j 6 11, J 2 11 J 5 5, z 2 16, z 4 16, z 5 13, Z 1 16, Z 4 16, Z 5 55, s 4 19, S 2 16, S 4 13, S 5 13, r 3 16, R 1 13. All setae smooth. Peritreme — Extending almost to z 2. Venter — Distances between St 1 –St 3 62, St 2 –St 2 62, St 5 –St 5 72. Ventrianal shield 112 long, 70 wide at level of Zv 2 and 67 wide at level of anus, with a pair of elliptical pores slightly posterior to Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp­tipped. Chelicera — Movable digit 27 long, apparently with 1 small tooth; fixed digit 25 long, apparently with 3 small teeth and a discernible pilus dentilis. Spermatheca — Calyx mostly slender, slightly bulged near atrium, flaring towards vesicle, 43 long; atrium distinct. Legs — Macrosetae blunt on tibiae and with a tiny knob on genua; Sge II 18, Sge III 26, Sti III 22, Sge IV 30, Sti IV 30, St IV 43. Chaetotaxy of genu III: 1­2 / 0,2 / 1 ­ 1. MALE — (1 specimen measured). Dorsum — Dorsal shield pattern as in female, 270 long and 185 wide. Setar j 1 20, j 3 23, j 4 10, j 5 10, j 6 10, J 2 10, J 5 5, z 2 15, z 4 15, z 5 10, Z 1 13, Z 4 15, Z 5 38, s 4 18, S 2 13, S 4 15, S 5 15, r 3 13, R 1 15. All setae smooth. Peritreme — Extending to level between z 2 and z 4. Venter — Ventrianal shield subtriangular, with transverse striae anteriorly; 115 long, 155 wide at the anterior corners; with 3 pairs of preanal setae, 2 pairs of lyrifissures, and a pair of small pores posterior to Jv 2. Spermatodactyl — Shaft 26 long. Legs — Macrosetae blunt on tibiae and with a tiny knob on genua; Sge II 13, Sge III 23, Sti III 18, Sge IV 28, Sti IV 23 and St IV 38. Chaetotaxy of genu III same as in female. LOCALITY AND TYPE MATERIAL — Holotype female and allotype male from unknown plant, 30 km W Yala, Nyanza Province, Kenya; 02­XII­ 1989; J.S. Yaninek, deposited at ESALQ­USP. ETYMOLOGY — The epithet nyanzaensis refers to the location where the types of this new species were collected. REMARKS — Euseius rhusi resembles this new species, but differs by having much longer peritreme (extending to level of j 1). Euseius distinctus is also similar to this species, but it lacks seta Jv 4 and has macrosetae of legs III and IV distinctly knobed.

Typhlodromalus aripo De Leon (Fig. 6) Typhlodromalus aripo De Leon, 1967: 21 Amblyseius aripo, Moraes & McMurtry, 1983: 132 Typhlodromalus aripo, Moraes et al., 2004: 195 Typhlodromalus aripo, Chant & McMurtry, 2005 a: 199 FEMALE ��� (Specimens measured ��� Benin: 2; Cameroon: 2; Congo Brazzaville: 1; Malawi: 1; Mozambique: 2; Tanzania: 2; Uganda: 2). Dorsum ��� Dorsal shield sculptured throughout, with mostly roundish to elongate elements between j 1 and Z 4, with an indentation on each side, near S 5 /Z 5; 355 (325���365) long and 191 (180���208) wide. Setae j 1 33 (30���35), j 3 41 (39���43), j 4 12 (11���13), j 5 12 (10��� 13), j 6 15 (14���16), J 2 14 (13���15), J 5 9 (8���10), z 2 18 (17���24), z 4 35 (32���38), z 5 10 (10���11), Z 1 20 (18���23), Z 4 56 (45���64), Z 5 75 (65���83), s 4 49 (44���51), S 2 30 (25���32), S 4 22 (18���24), S 5 11 (10���12), r 3 22 (21���24), R 1 18 (17���20). All setae smooth, except Z 4 and Z 5 which are serrate. Peritreme ��� Extending to base of j 1. Venter ��� Distances between St 1 ���St 3 67 (65���70), St 2 ���St 2 70 (68���73), St 5 ���St 5 82 (74��� 88). Ventrianal shield 110 (102���116) long, 68 (60���74) wide at level of Zv 2 and 67 (64���73) wide at level of anus, with a pair of elliptical pores posterior to Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 32 (31���32) long, with 3 teeth; fixed digit 29 (28���29) long, with 9 teeth and a clearly discernable pilus dentilis. Spermatheca ��� Calyx tubular 19 (18���20) long, frequently with distal 2 / 3 sclerotized; atrium distinct. Legs ��� Macrosetae blunt on tibiae and with tiny knob on genua and tarsus IV; Sge I 20 (19���21), Sge II 20 (19���21), Sge III 25 (21���28), Sti III 19 (18���20), Sge IV 53 (47���59), Sti IV 26 (23���28), St IV 73 (70���77). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� (Specimens measured ��� Cameroon: 1; Congo Brazzaville: 1; Malawi: 1; Mozambique: 1; Tanzania: 1). Dorsum ��� Dorsal shield pattern as in female, 275 (268���283) long and 159 (150���170) wide. Setae j 1 25 (24���26), j 3 32, j 4 10 (10���11), j 5 10 (9���11), j 6 12 (11���13), J 2 10 (10���12), J 5 7 (7���8), z 2 15 (14���15), z 4 24 (23���26), z 5 8 (7���10), Z 1 14 (13���15), Z 4 35 (33���38), Z 5 44 (41��� 48), s 4 35 (33���39), S 2 18 (16���21), S 4 13 (12���13), S 5 9 (8���10) r 3 18 (16���20), R 1 14 (12���15). All setae smooth, except Z 4 and Z 5 which are serrate. Peritreme ��� Extending to base of j 1. Venter ��� Ventrianal shield subtriangular, with transverse striae anteriorly; 108 (103��� 115) long, 142 (138���150) wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral), and a pair of elliptical pores approximately in level with Jv 2. Spermatodactyl ��� Shaft 22 (21���23) long. Legs ��� Macrosetae blunt on tibiae and with tiny knob on genua and tarsus IV; Sge I 17 (16���17); Sge II 18 (17���18), Sge III 16 (14���17), Sti III 17 (16���19), Sge IV 28 (25���31), Sti IV 22 (20���23) and St IV 51 (48���53). Chaetotaxy of genu III same as in female. SPECIMENS EXAMINED: Benin: Se, Mono Province, on Manihot esculenta, 21 �� XII�� 1999, D. Hounhagni. Cameroon: 2 km S Ombessa, on M. esculenta, 24 ��XI�� 1998, A. Onzo; Bagante release field, on M. esculenta, 24 ��XI�� 1998, A. Onzo. Republic of Congo: Lekana, on M. esculenta, 17 ��VIII�� 1999, P. Neuenschwander. Malawi: 1.5 km N Nkhotakota, Central Province, on M. esculenta, 11 ��V�� 2001, I.D. Zannou; 29.9 km S Matyewo, Central Province, on M. esculenta, 17 ��V�� 2001, I.D. Zannou. Mozambique: 3.5 km NE Muecate, Nampula Province, on M. esculenta, 01��V�� 2001, I.D. Zannou; 4 km N Muecate, Nampula Province, on M. esculenta, 01��V�� 2001, I.D. Zannou. Tanzania: 2 km N Mnazimmoja (Mingoyo), on M. esculenta, 22 ��XII�� 1999, B. Pallangyo; Rwani��Shenye, on M. esculenta, 22 ��XII�� 1999, E. Nsami. Uganda: 49 km E Iganga, on M. esculenta, 04��III�� 1998, A. Onzo; 39 km E Iganga, on M. esculenta, 05��III�� 1998, A. Onzo. REMARKS ��� This species was imported from Brazil (South America) and introduced into Africa in 1993 in classical biological control program of M. tanajoa. WORLD DISTRIBUTION (based on Hanna et al. 2005, Moraes et al., 2004 and Yaninek & Hanna, 2003) �� Benin, Burundi, Brazil, Cameroon, Centrafrique, Colombia, Costa Rica, Democratic Republic of Congo, El Salvador, Equatorial Guinea, Gabon, Ghana, Guadeloupe, Guinea��Conakry, Guyana, Ivory��Coast, Kenya, Jamaica, Liberia, Malawi, Mozambique, Nigeria, Paraguay, Republic of Congo, Sierra��Leone, Tanzania, Togo, Uganda, Zambia., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 11-13, DOI: 10.5281/zenodo.171564, {"references":["De Leon, D. (1967) Some mites of the Caribbean Area. Part I. Acarina on plants in Trinidad, West Indies. Allen Press Inc., Lawrence, Kansas, USA, pp. 1 - 66.","Moraes, G. J. de & McMurtry, J. A. (1983) Phytoseiid mites (Acarina) of northeastern Brazil with descriptions of four new species. International Journal of Acarology, USA, 9, 131 - 148.","Moraes, G. J. de, McMurtry, J. A., Denmark, H. A., Campos, C. B. (2004) A revised catatalog of the mite family Phytoseiidae. Zootaxa, New Zealand, 434, 1 - 494.","Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224.","Hanna, R., Onzo, A., Lingeman, R., Yaninek, J. S. & Sabelis, M. W. (2005) Seasonal cycles and persistence in an acarine predator-prey system on cassava in Africa. Population Ecology, Tokyo, 47, 107 - 117.","Yaninek, J. S. & Hanna, R. (2003) Cassava green mite in Africa: a unique example of successful classical biological control of a mite on a continental scale. In: Borgemeister, C., Langewald, J. (Eds), Biological Control in IPM System in Africa, CABI, UK, pp. 61 - 75."]}

Ueckermannseius macrosetosus (Van der Merwe) (Fig. 17) Amblyseius (Amblyseius) macrosetosus Van der Merwe, 1965: 61 Amblyseius macrosetosus, Meyer & Rodrigues, 1966: 30 Typhlodromalus macrosetosus, Moraes et al. 2004: Ueckermannia macrosetosus, Chant & McMurtry, 2005 a: 203 FEMALE ��� (Specimens measured; Burundi: 2; Cameroon: 4; Democratic Republic of Congo: 3; Ghana: 2; Kenya: 2; Nigeria: 1; Rwanda: 4; Sierra��Leone: 1; Unganda: 6; Zimbabwe: 1). Dorsum ��� Dorsal shield smooth in the central area, with some striation along lateral margins, 376 (328���446) long and 251 (214���294) wide. Setae j 1 37 (32���42), j 3 30 (24���38), j 4 19 (16���24), j 5 18 (14���24), j 6 18 (14���24), J 2 17 (14���21), J 5 5 (3���8), z 2 28 (22���35), z 4 34 (24��� 48), z 5 18 (13���24), Z 1 22 (16���34), Z 4 21 (16���32), Z 5 57 (46���67), s 4 41 (32���56), S 2 24 (16��� 35), S 4 21 (16���35), S 5 21 (16���37), r 3 22 (16���27), R 1 19 (13���24). All setae smooth. Peritreme ��� Extending to level between j 3 and z 2. Venter ��� Distances between St 1 ���St 3 68 (61���75), St 2 ���St 2 73 (66���82), St 5 ���St 5 75 (67��� 85). Ventrianal shield 120 (104���130) long, 73 (62���90) wide at level of Zv 2 and 66 (61���78) wide at level of anus, with a pair of large pores posterior to Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 37 (36���38) long, with 4 teeth; fixed digit 30 (28���33) long, with 12 teeth and a clearly discernible pilus dentilis. Spermatheca ��� Calyx tubular, with distal 2 / 3 slender, 45 (38���58) long; atrium distinct. Legs ��� Macrosetae sharp��tipped on legs I, blunt on leg II, and knobbed on legs III and IV, Sge I 28 (27���29), Sge II 31 (27���35), Sge III 42 (38���48), Sti III 34 (27���38), Sge IV 63 (50���70), Sti IV 49 (39���58), St IV 64 (53���80). Chaetotaxy of genu III: 1��2 / 0,2 / 1 �� 1. MALE ��� (Specimen measured ��� Cameroon: 1). Dorsum ��� Dorsal shield pattern as in female, 250 long and 185 wide. Setae j 1 28, j 3 38, j 4 15, j 5 13, j 6 18, J 2 15, J 5 4, z 2 28, z 4 38, z 5 15, Z 1 23, Z 4 33, Z 5 53, s 4 45, S 2 28, S 4 25, S 5 28, r 3 15, R 1 13. All setae smooth. Peritreme ��� Extending to level between z 2 and z 4. Venter ��� Ventrianal shield subtriangular, with transversal striae anteriorly; 98 long, 155 wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral) and a pair of medium pores slightly posterior to Jv 2. Spermatodactyl ��� Shaft 23 long. Legs ��� With sharp��tipped macroseta on leg I, blunt macroseta on leg II and knobbed macrosetae on legs III and IV; Sge I 28, Sge II 29, Sge III 30, Sti III 25, Sge IV 45, Sti IV 38 and St IV 48. Chaetotaxy of genu III same as in female. SPECIMENS EXAMINED: Burundi: 5 km S Gitega, Rutana Province, on unknown plant, 13 ��XII�� 1989, J.S. Yaninek. Cameroon: 21 km E Ngoura, Eastern Province, on unknown plant, 03��II�� 1991, A. Onzo; 11 km W Diang, Eastern Province, on Elaeis guineensis, 04��II�� 1991, A. Onzo; 2 km W Minta, Central Province, on Elaeis guineensis, 04��II�� 1991, A. Onzo. Democratic Republic of Congo: 3 km N Masi��Manimba, on Hymenocardia acida, 26 ��VI�� 1991, A. Onzo; 3 km N Masi��Manimba, on Hypanhenia sp., 26 ��06�� 1991, A. Onzo; 4 km S Kikwit, on Morinda lucida, 26 ��06�� 1991, A. Onzo. Ghana: 3 km E Mankesim, Central Region, on Mallotus oppositifolius, 12 ��XI�� 1989, J.S. Yaninek; Cape Coast University, Central Region, on M. oppositifolius, 12 ��XI�� 1989, J.S. Yaninek. Kenya: 27.9 km S Mombasa, Coastal Province, on Zea mays, 28 ��XI�� 1989, J.S. Yaninek; Sondu, Nyanza Province, on unknown plant, 03��XII�� 1989, J.S. Yaninek. Nigeria: IITA Station, Ibadan, on unknown plant, 06��VIII�� 1986, F. Schulthess. Rwanda: ISAR Station, Rubosa, on unknown plant, 06��XII�� 1989, J.S. Yaninek; on Vernonia amygdalina, 08��XII�� 1989, J.S. Yaninek; ISAR Station, Kigali, on Citrus sp., 09��XII�� 1989, S.T. Kamara. Sierra��Leone: Luzar Hospital, on Mangifera indica, 16 ��XII�� 1991, B. Kristensen. Uganda: Luwero, on unknown plant, 05��X�� 1990, J.S. Yaninek; Namansa, on Artocarpus heterophyllus, 05��X�� 1990, J.S. Yaninek; 13 km W Mbarara, on Mangifera indica, 07��X�� 1990, J.S. Yaninek; 1 km N Muzizi River, Hoima, on Acanthus pubescens, 10 ��X�� 1990, J.S. Yaninek; Jinja District, on Ricinus communis, 05��XII�� 1990, Mugarura; 20 km W Tororo, on Acacia sp., 05��XII�� 1990, Mugarura. Zimbabwe: Tubalse, Research Institute, Harare, on Pensettis sp., 12 ��II�� 1985, unknown collector. REMARKS ��� Specimens from Kenya and Cameroon have knobbed or sharp��tipped Jv 5, while all the specimens from Burundi, Democratic Republic of Congo, Rwanda, Uganda and Sierra��Leone have knobbed Jv 5 and all the specimens from Ghana, Nigeria and Zimbabwe have sharp��tipped Jv 5. Only the peritreme of one specimen from Rwanda reaches the level of j 3 as in the original description, whereas in most of the specimens examined the peritremes extend to a level between j 3 and z 2. In one specimen from Kenya the peritreme extends to level of z 2 and in one specimen from Cameroon it extends to level between z 2 and z 4. WORLD DISTRIBUTION ��� Burundi, Cameroon, Democratic Republic of Congo, Ghana, Kenya, Mozambique, Nigeria, Rwanda, Sierra��Leone, South Africa, Uganda, Zimbabwe., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 32-34, DOI: 10.5281/zenodo.171564, {"references":["Van der Merwe, G. G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, South Africa, 28, 57 - 76.","Meyer, M. K. P. & Rodrigues, M. C. (1966) Acari associated with cotton in southern Africa (with reference to other plants). Garcia de Orta, Revista Junta Investigacoes de Ultramar, Portugal, 13 (2), 195 - 226.","Moraes, G. J. de, McMurtry, J. A., Denmark, H. A., Campos, C. B. (2004) A revised catatalog of the mite family Phytoseiidae. Zootaxa, New Zealand, 434, 1 - 494.","Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224."]}

Ueckermannseius eastafricae Moraes, Zannou & Oliveira, sp. nov. (Fig. 16) DIAGNOSIS ��� This new species is characterized by having s 4 ca. 40 % as long as distance between its base and the base of z 5, Z 5 ca. 2.5 times as long as S 5, Jv 5 blunt and ca. 60 % as long as ventrianal shield, and macrosetae of legs III and IV knobbed. FEMALE ��� (7 specimens measured). Dorsum ��� Dorsal shield mostly smooth, with some striae along lateral margins of proscutum, 390 (336���405) long and 274 (258���291) wide. Setae j 1 34 (30���38), j 3 23 (19��� 27), j 4 15 (11���18), j 5 14 (10���19), j 6 12 (10���16), J 2 13 (10���16), J 5 5 (3���6), z 2 20 (16���24), z 4 21 (13���29), z 5 14 (13���16), Z 1 16 (13���21), Z 4 16 (11���19), Z 5 39 (32���53), s 4 26 (19���35), S 2 18 (16���22), S 4 17 (14���21), S 5 16 (14���21), r 3 18 (16���21), R 1 18 (14���21). All setae smooth. Peritreme ��� Extending to level between j 1 and j 3. Venter ��� Distances between St 1 ���St 3 71 (62���77), St 2 ���St 2 72 (69���80), St 5 ���St 5 80 (67��� 90). Ventrianal shield 122 (96���130) long, 78 (67���85) wide at level of Zv 2 and 71 (62���75) wide at level of anus, with a pair of elliptical pores posterior to and slightly mesad of Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped, except Jv 5 which is blunt. Chelicera ��� Movable digit 38 long, with 4 teeth; fixed digit 34 (33���35) long, with 10 teeth and a distinguishable pilus dentilis. Spermatheca ��� Calyx tubular, with distal half slender and proximal half bulged, 35 long; atrium distinct. Legs ��� Macrosetae sharp��tipped on legs II and knobbed on legs III and IV; Sge II 28 (24���34), Sge III 39 (37���45), Sti III 31 (29���37), Sge IV 57 (51���62), Sti IV 44 (34���56), St IV 58 (51���69). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype female from Capsicum sp., 26 km NE Bundibugyo, Uganda, 09��X�� 1990, J.S. Yaninek, deposited at ESALQ��USP; 1 paratype female from Vernonia amygdalina, 5 km N Masaka, Uganda, 06��X�� 1990, J.S. Yaninek, deposited at IITAIM. Five paratypes females from unknown plant, 17 km S Sonda, Kenya, 03��XII�� 1989, J.S. Yaninek, deposited at ESALQ��USP. ETYMOLOGY ��� The epithet eastafricae refers to the region of Africa (east) where the types of this new species were collected. REMARKS ��� Ueckermannseius macrosetosus (Van der Merwe 1965) is similar to this new species, but it has much longer dorsal shield setae, shorter peritreme (extending between j 3 and z 2) and sharp��tipped Jv 5. Ueckermannseius havu (Pritchard & Baker 1962) also resembles this new species, but it has much longer j 3, z 2, z 4, s 4 and Z 5, shorter peritreme (extending between j 3 and z 2), a knobbed Jv 5 and lacks Jv 4., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 30-31, DOI: 10.5281/zenodo.171564, {"references":["Van der Merwe, G. G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, South Africa, 28, 57 - 76.","Pritchard, A. E. & Baker, E. W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world. Hilgardia, USA, 33, 205 - 309."]}

Typhlodromalus spinosus (Meyer & Rodrigues) (Fig. 11) Amblyseius spinosus Meyer & Rodrigues, 1966: 30 Kampimodromus spinosus, Quilici et al., 2000: 100 Typhlodromalus spinosus, Moraes et al., 2004: 204 Typhlodromalus spinosus, Chant & McMurtry, 2005 a: 199 FEMALE — (Species measured — Benin: 1; Burundi: 1; Kenya: 3; Democratic Republic of Congo: 1). Dorsum — Dorsal shield sculptured anteriorly to Z 4, with roundish elements centrally, irregular elements sublaterally and few striae along anterolateral margins, with an indentation on each side, near S 5 /Z 5; 332 (322–344) long and 202 (192–210) wide. Idiosomal setal pattern 10 A: 8 B/JV– 3:ZV, a characteristic which distinguishes the athiasae species group of Chant & McMurtry (2005 a). Setae j 1 29 (26–33), j 3 33 (29–38), j 4 15 (14– 16), j 5 16 (14–18), j 6 22 (19–25), J 2 24 (21–25), J 5 9 (8–10), z 2 22 (21–24), z 4 34 (30–40), z 5 16 (14–18), Z 1 23 (19–26), Z 4 48 (43–50), Z 5 65 (61–68), s 4 47 (43–53), S 2 42 (37–48), S 4 30 (22–38), r 3 18 (16–23), R 1 18 (14–21). Setae j 1, j 4, j 5, J 5, z 5, Z 1, r 3 and R 1 smooth; other setae serrate. S 5 absent. Peritreme — Extending anterior to base of j 1. Venter — Distances between St 1 –St 3 64 (61–68), St 2 –St 2 65 (62–69), St 5 –St 5 71 (67– 78). Ventrianal shield 103 (99–109) long, 51 (43–55) wide at level of Zv 2 and 62 (58–70) wide at level of anus, with a pair of elliptical pores roughly aligned with Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp­tipped, except Jv 5 which is serrate. Chelicera — Movable digit 36 (35–37) long, with 3 to 4 teeth; fixed digit 30 (29–30) long, with 9 to 10 teeth and a discernible pilus dentilis. Spermatheca — Calyx saccular, with a distinctly constricted region followed by a flared region distally, 16 (15–18) long; atrium distinct. Legs — Macrosetae blunt on genu I and tibiae III and IV and knobbed on genua II to IV and tarsus IV; Sge I 16 (13–20), Sge II 19 (14–23), Sge III 27 (22–33), Sti III 20 (19–23), Sge IV 43 (40–48), Sti IV 26 (24–30), St IV 55 (53–58). Chaetotaxy of genu III: 1 ­(2,2)/ 1 ­ 1. MALE — (Specimen measured — Kenya: 1) Dorsum — Dorsal shield pattern as in female, 246 long and 152 wide. Setae j 1 19, j 3 22, j 4 11, j 5 13, j 6 14, J 2 14, J 5 5, z 2 16, z 4 26, z 5 13, Z 1 16, Z 4 30, Z 5 40, s 4 34, S 2 24, S 4 16, r 3 13, R 1 13; r 3 and R 1 on dorsal shield. Setae j 1, j 4, j 5, J 5, z 5, Z 1, r 3 and R 1 smooth; other setae serrate. S 5 absent. Peritreme — Extending to level of j 1. Venter — Ventrianal shield subtriangular, with anterior transversal striae; 91 long, 136 wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral) and a pair of elliptical pores posterior to Jv 2. Spermatodactyl — Shaft 24 long. Legs — Macrosetae blunt on genu I and tibiae III and IV and knobbed on genua II to IV and tarsus IV; Sge I 14, Sge II 14, Sge III 19, Sti III 13, Sge IV 29, Sti IV 18 and St IV 40. Chaetotaxy of genu III same as in female. SPECIMENS EXAMINED: Benin: Gangnigon, on Securinega virosa, 22 ­VIII­ 1991, B. Eklou. Burundi: 16 km NW Bujumbura, on unknown plant, 11 ­XII­ 1989, J.S. Yaninek. Democratic Republic of Congo: 34 km E Matadi, Kenge, Bas­Congo, on Cajanus cajan, 11 ­VII­ 1991, A. Onzo. Kenya: 5 km N Kinango, Coastal Province, on Acacia sp., 29 ­XI­ 1989, J.S. Yaninek; 5 km N Kinango, Coastal Province, on unknown plant, 29 ­XI­ 1989, J.S. Yaninek; Katumani Research Station, on unknown plant, 26 ­XII­ 1989, J.S. Yaninek. REMARKS — Contrary to the original description and to the specimen used to illustrate the dorsal shield, some of the specimens examined in the present study have serrate j 4, j 5 and z 5; some specimens have blunt Sge II and Sge III. Typhlodromalus (= Amblyseius) serengati (El­Banhawy & Abou­Awad, 1990) was described based on specimens collected on an unspecified substrate in Dar Es Salaam, Tanzania. This species was not included in this paper because, based on its original description, we did not find significant differences between it and T. spinosus. Although this species is more similar to T. spinosus than to T. athiasae, the authors only compared it with the latter, in the original description. According to them, the main difference between those species is the shape of the calyx of spermatheca. The illustration provided in the original description shows the calyx to be very similar to that of T. spinosus. The authors also mentioned that S 4 is absent in T. serengati, but their illustration indicates S 5 rather than S 4 to be absent (as also occurs in T. athiasae and T. spinosus). The synonymy between T. serengati and T. spinosus is not officially proposed in this paper because the type of the former was not examined in this study. WORLD DISTRIBUTION — Benin, Burundi, Democratic Republic of Congo, Kenya, Mozambique, Reunion Island.

Euseius oligodous Moraes, Zannou & Oliveira, sp. nov. (Fig. 23) DIAGNOSIS — This species is distinct by a combination of the following characteristics: long peritreme (reaching base of j 1), reticulate dorsal shield, Jv 1 inserted near anterior margin of ventrianal shield and distinctly knobbed macrosetae on legs. FEMALE — (2 specimens measured). Dorsum — Dorsal shield reticulate, 335 (334–337) long and 225 wide. Setae j 1 27 (26– 28), j 3 14, j 4 11, j 5 12 (11–12), j 6 12 (11–12), J 2 14, J 5 9 (8–9), z 2 16 (15–17), z 4 14, z 5 11, Z 1 14 (13–14), Z 4 15 (14–15), Z 5 57 (55–58), s 4 18 (17–18), S 2 16, S 4 20 (19–20), S 5 18 (17–19), r 3 16 (15–16), R 1 14 (13–14). All setae smooth, except Z 5 which is lightly serrate. Peritreme — Extending to level of j 1. Venter — Distances between St 1 –St 3 55 (54–55), St 2 –St 2 68 (67–68), St 5 –St 5 75. Ventrianal shield 105 long, 58 wide at level of Zv 2 and 67 (65–69) wide at level of anus, with a pair of elliptical pores slightly posteromesad of Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp­tipped. Chelicera — Movable digit 27 (26–27) long, with 1 small tooth; fixed digit 25 long, with 4 small teeth and a discernible pilus dentilis. Spermatheca — Calyx funnel­shaped, 15 (14–15) long; atrium elongate. Legs — Macrosetae knobbed; Sge II 19, Sge III 26 (25–26), Sti III 23 (22–23), Sge IV 38, Sti IV 31 (30–31), St IV 44 (43–44). Chaetotaxy of genu III: 1­2 / 1,2 / 0­1. MALE — Unknown. LOCALITY AND TYPE MATERIAL — Holotype female and 1 paratype female from Anisophyllea boehmii, 30 km S Kasama, Zambia, 16 ­VI­ 1992, A. Onzo, deposited at ESALQ­USP. ETYMOLOGY — The epithet oligodous refers to the presence of few teeth on the fixed cheliceral digit of this new species. REMARKS — Euseius parcidentatus is similar to this new species, but it differs by having shorter dorsal shield setae, S 4 inserted very close to dorsal shield margin, longer and strongly knobbed macrosetae on legs III and IV and shorter atrium of spermatheca.

Amblydromalus nakuruensis Moraes, Zannou & Oliveira, sp. nov. (Fig. 3) DIAGNOSIS ��� This species is characterized by the relatively short Z 4 and relatively long funnel��shaped calyx of spermatheca. FEMALE ��� (1 specimen measured). Dorsum ��� Dorsal shield smooth, 349 long and 208 wide. Setae j 1 34, j 3 43, j 4 13, j 5 11, j 6 16, J 2 16, J 5 8, z 2 24, z 4 38, z 5 13, Z 1 18, Z 4 27, Z 5 62, s 4 59, S 2 35, S 4 16, S 5 21, r 3 24, R 1 13. All setae smooth. Peritreme ��� Extending to level of j 1. Venter ��� Distances between St 1 ���St 3 67, St 2 ���St 2 66, St 5 ���St 5 85. Ventrianal shield 114 long, 62 wide at level of Zv 2 and 70 wide at level of anus, with a pair of small pores posterior to and slightly mesad of Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 35 long, apparently with 4 teeth; fixed digit 38 long, with at least 9 teeth and a clearly discernible pilus dentilis. Spermatheca ��� Calyx funnel��shaped, 22 long; atrium not clearly discernable. Legs ���Macrosetae sharp��tipped; Sge II 27, Sge III 26, Sti III 24, Sge IV 43, Sti IV 26 (St IV broken). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype female from Triumphetta sp., 39 km W Nakuru, Kenya; 04��XII�� 1989, J.S.Yaninek; deposited at ESALQ��USP. ETYMOLOGY ��� The epithet nakuruensis refers to Lake Nakuru, in the proximity of the location where the type of this new species was collected. REMARKS ��� Differently from what is reported by Chant & McMurtry (2005 a) for limonicus species group, in this species Z 4 is ca. 50 % as long as the distance between its base and the base of Z 5, there is no distinguishable macroseta on genu I, and Jv 2 and Zv 2 are not distinctly anterior when compared with species in other groups. Females of this new species resemble Amblydromalus higuilloae (Denmark & Muma 1975), but the latter differs by having longer Z 4 and much shorter calyx of spermatheca. Amblydromalus laetus (Chant & Baker 1965) is also similar to this new species, but it has shorter z 2, longer Z 4, lightly serrate Z 5 and tubular calyx of spermatheca., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 6-7, DOI: 10.5281/zenodo.171564, {"references":["Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224.","Denmark, H. A. & Muma, M. H. (1975) The Phytoseiidae (Acarina: Mesostigmata) of Puerto Rico. The Journal of Agriculture of the University of Puerto Rico, Puerto Rico, 59, 279 - 304.","Chant, D. A. & Baker, E. W. (1965) The Phytoseiidae (Acarina) of Central America. Memoirs of the Entomological Society of Canada, Canada, 41, 56 pp."]}

Typhlodromalus longisetatus Moraes, Zannou & Oliveira, sp. nov. (Fig. 8) DIAGNOSIS ��� This new species resembles none of the Typhlodromalus species reported in Chant & McMurtry (2005 a), because of the subequal lengths of its dorsal shield setae (except J 5), most of which are longer than distances between their bases and the bases of the setae immediately behind. FEMALE ��� (1 specimen measured). Dorsum ��� Dorsal shield sculptured, with roundish elements between j 5 and J 2, behind s 4 and behind Z 1 and with some striae mostly along anterolateral margins, without indentation near S 5 /Z 5; 304 long and 208 wide. Idiosomal setal pattern 10 A: 8 B/JV�� 3: ZV, a characteristic which distinguishes the athiasae species group of Chant & McMurtry (2005 a). Setae j 1 38, j 3 48, j 4 29, j 5 32, j 6 51, J 2 50, J 5 8, z 2 32, z 4 48, z 5 26, Z 1 43, Z 4 64, Z 5 78, s 4 62, S 2 66, S 4 53, r 3 27, R 1 29. Setae serrate, except J 5, r 3 and R 1 which are smooth. S 5 absent. Peritreme ��� Extending anterior to base of j 1. Venter ��� Distances between St 1 ���St 3 61, St 2 ���St 2 66, St 5 ���St 5 77. Ventrianal shield 96 long, 61 wide at level of Zv 2 and 62 wide at level of anus, with a pair of elliptical pores slightly posterior to Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp��tipped, except Jv 4 and Jv 5 which are serrate. Chelicera ��� Movable digit 32 long, with 4 teeth; fixed digit 29 long, with 8 teeth and a distinguishable pilus dentilis. Spermatheca ��� Calyx slender, flaring slightly distally, 21 long; atrium distinct and slender. Legs ��� Macrosetae blunt on tibiae III and IV and knobbed on genua I to IV and tarsus IV; Sge I 22 Sge II 27, Sge III 34, Sti III 24, Sge IV 53, Sti IV 32, St IV 66. Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype female from unknown plant, 400 km N Adomi bridge, Atimpoku, Ghana, 13 ��XI�� 1989, J.S. Yaninek, deposited at ESALQ�� USP. ETYMOLOGY ��� The epithet longisetatus refers to the relatively long dorsocentral setae of the idiosoma of this new species. REMARKS ��� Differently from most of the species in this genus, as reported by Chant & McMurtry (2005 a), the margin of the dorsal shield of T. longisetatus sp. nov. does not have an indentation near S 5 /Z 5; and the dorsocentral setae of this species are relatively long. Similarly to this new species, T. athiasae, Typhlodromalus sinespinosus Moraes, Zannou & Oliveira, sp. nov., Typhlodromalus spinosus (Meyer & Rodrigues) and Typhlodromalus tenuicalyx Moraes, Zannou & Oliveira, sp. nov. do not have S 5; however, all of those species have dorsal shield setae quite variable in length. In addition, T. sinespinosus and T. spinosus have calyx of spermatheca sacculiform., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 15-17, DOI: 10.5281/zenodo.171564, {"references":["Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224."]}

Ueckermannseius havu (Pritchard & Baker) Amblyseius (Amblyseius) havu Pritchard & Baker, 1962: 257 Amblyseius havu, Ghai & Menon, 1967: 70 Amblyseius (Typhlodromalus) havu, Gupta, 1985: 361 Typhlodromalus havu, Moraes et al., 2004: 197 Ueckermannia havu, Chant & McMurtry, 2005 a: 203 FEMALE ��� (Specimens measured ��� Democratic Republic of Congo: holotype) Dorsum ��� Dorsal shield smooth in the central area and with some striation along lateral margins, [374] long and [263] wide. Setae j 1 [39], j 3 [41], j 4 [27], j 5 [22], j 6 [22], J 2 [24], J 5 [5], z 2 [34], z 4 [39], z 5 [22], Z 1 [31], Z 4 [28], Z 5 [63], s 4 [44], S 2 [36], S 4 [30], S 5 [30], r 3 [22], R 1 [24]. All setae smooth. Venter ��� Distances between St 1 ���St 3 [71], St 2 ���St 2 [77], St 5 ���St 5 [78]. Ventrianal shield [75] wide at level of Zv 2 (length and width at level of anus, of ventrianal shield not possible to measure); with a pair of elliptical pores posterior to Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped, except Jv 5 which is knobbed; Jv 4 absent. Spermatheca ��� Calyx with distal 3 / 5 slender and basal 2 / 5 bulged [56] long; atrium distinct. Legs ��� Macrosetae sharp��tipped on leg II and knobbed on legs III and IV; Sge II [38], Sge III [47], Sti III [39], Sge IV [68], Sti [55], St [75]. SPECIMENS EXAMINED: Democratic Republic of Congo (Belgian Congo): Holotype female from Lwiro, on Maesa rufescens, 16 ��V�� 1955, E.W. Baker. REMARKS ��� This species was described from specimens collected in Lwiro, Democratic Republic of Congo (Belgian Congo) on Maesa rufescens. No specimens were collected in this study. WORLD DISTRIBUTION ��� Democratic Republic of Congo., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 31-32, DOI: 10.5281/zenodo.171564, {"references":["Pritchard, A. E. & Baker, E. W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world. Hilgardia, USA, 33, 205 - 309.","Ghai, S. & Menon, M. G. R. (1967) Taxonomic studies on Indian mites of the family Phytoseiidae (Acarina). I. New species and new records of the genus Amblyseus Berlese from India (Acarina: Phytoseiidae) with a key to the Indian species. Oriental Insects, India, 1, 65 - 79.","Gupta, S. K. (1985) Plant mites of India. Zoogical Survey of India Handbook Series, Calcutta, India, 5, 1 - 520.","Moraes, G. J. de, McMurtry, J. A., Denmark, H. A., Campos, C. B. (2004) A revised catatalog of the mite family Phytoseiidae. Zootaxa, New Zealand, 434, 1 - 494.","Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224."]}

Typhlodromalus athiasae (Pritchard & Baker) (Fig. 7) Amblyseius (Amblyseiella) athiasae Pritchard & Baker, 1962: 291 Typhlodromalus athiasae, Moraes et al., 2004: 196 Typhlodromalus athiasae, Chant & McMurtry, 2005 a: 199 FEMALE — (Specimens measured — Benin: 2; Cameroon: 6; Kenya: 3; Nigeria: 1; Uganda: 1; Democratic Republic of Congo: 1 in addition to holotype). Dorsum — Dorsal shield sculptured throughout, with mostly roundish to elongate elements, with an indentation on each side, near S 5 /Z 5; 340 (320–371) [298] long and 201 (176–237) [196] wide. Idiosomal setal pattern 10 A: 8 B/JV­ 3:ZV, a characteristic which distinguishes the athiasae species group of Chant & McMurtry (2005 a). Setae j 1 30 (23– 37) [31], j 3 36 (32–43) [34], j 4 19 (16–27) [17], j 5 20 (16–27) [19], j 6 28 (22–38) [26], J 2 29 (22–38) [28], J 5 8 (6–10), z 2 26 (21–34) [19], z 4 37 (32–43) [34], z 5 20 (18–27) [18], Z 1 28 (22–38) [27], Z 4 50 (42–58) [44], Z 5 68 (61–83) [56], s 4 49 (42–58) [41], S 2 48 (40–53) [41], S 4 37 (30–40) [33], r 3 20 (14–25) [19], R 1 19 (14–28) [16]. Setae serrate, except J 5, r 3 and R 1 which are smooth. S 5 absent. Peritreme — Extending anterior to base of j 1. Venter — Distances between St 1 –St 3 63 (61–67) [58], St 2 –St 2 64 (58–69) [63], St 5 – St 5 74 (67–82) [75]. Ventrianal shield 108 (98–120) [94] long, 60 (53–69) [58] wide at level of Zv 2 and 68 (61–80) [63] wide at level of anus, with a pair of elliptical pores slightly posterior to Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharptipped, except Jv 5 which is serrate. Chelicera — Movable digit 33 (32–35) long, with 3 to 4 teeth; fixed digit 29 (28–29) long, with 6 to 10 [at least 7] teeth and a clearly discernable pilus dentilis. Spermatheca — Calyx slender, 33 (26–43); atrium indistinct. Legs — Macrosetae blunt on genua I and II and tibiae III and IV, and knobbed on genua III and IV and tarsus IV; Sge I 17 (11–22) [17], Sge II 18 (14–21) [17], Sge III 27 (24–30) [24], Sti III 20 (18–22) [17], Sge IV 41 (37–48) [39], Sti IV 26 (22–30) [24], St IV 56 (48–64) [48]. Chaetotaxy of genu III: 1­2 / 1,2 / 0­1. MALE — (Specimens measured — Kenya: 2) Dorsum — Dorsal shield pattern as in female, 221 (220–223) long and 141 (138–145) wide. Setae j 1 18 (18–19), j 3 25, j 4 13, j 5 14 (13–15), j 6 14 (14–15), J 2 15, J 5 6, z 2 19 (18– 20), z 4 26 (25–28), z 5 13, Z 1 16 (15–16), Z 4 33 (33–34), Z 5 44 (44–45), s 4 36 (35–36), S 2 31 (30–31), S 4 19 (18–20), r 3 13, R 1 15. Setae serrate, except J 5, r 3 and R 1 which are smooth. Peritreme — Extending to level of j 1. Venter — Ventrianal shield subtriangular, with transversal striae anteriorly; 86 (85–88) long, 119 (118–120) wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral), and a pair of elliptical pores approximately in level with Jv 2. Spermatodactyl — Shaft 15 long. Legs — Macrosetae blunt on genua I and II and tibiae III and IV, and knobbed on genua III and IV and tarsus IV; Sge I 14; Sge II 18, Sge III 19 (19–20), Sti III 17 (16–18), Sge IV 25, Sti IV 20 and St IV 37 (35–39). Chaetotaxy of genu III same as in female. SPECIMENS EXAMINED: Benin: Agbotagon, on Securinega virosa, 12 ­VIII­ 1991, B. Eklou; Agbotagon, on Mallotus oppositifolius, 18 ­XI­ 1991, A. Onzo. Cameroon: 12 km S Obala, Central Province, on Sida acuta, 05­II­ 1991, A. Onzo; 14 km SW Mvieng, Southern Province, on Urena lobata, 07­II­ 1991, A. Onzo; 105 km N Kribi, Littoral Province, on Solanum torvum, 08­II­ 1991, A. Onzo; 18 km SE Douala, on Chromolaena odorata, 08­II­ 1991, A. Onzo; 5 km N Muyuka, Littoral Province, on Stachytarpheta sp., 09­II­ 1991, A. Onzo; 5 km N Muyuka, Littoral Province, on Albizia zygia, 09­II­ 1991, A. Onzo. Democratic Republic of Congo (Belgian Congo): Holotype female from Stanleyville, on unknown tree, 20 ­IV­ 1955, E.W. Baker; 19 km W Kinzoavuete, Bas­ Congo, on C. odorata, 10 ­VII­ 1991, A. Onzo. Kenya: 6.5 km S Mombasa, Coastal Region, on Cocos nucifera, 28 ­XI­ 1989, J.S. Yaninek; 13 km W Kwale, on Lycopersicum esculentum, 29 ­XI­ 1989, J.S. Yaninek; 19 km N Kilifi Ferry, on unknown plant, 30 ­XI­ 1989, J.S. Yaninek. Nigeria: IITA Station, Ibadan, Oyo state, on unknown plant, 06­VIII­ 1986, J.S. Yaninek. Uganda: 20 km W Tororo, on Acacia sp., 1990, Mugarura. WORLD DISTRIBUTION — Benin, Burundi, Cameroon, Democratic Republic of Congo, Ivory Coast, Ghana, Kenya, Malawi, Mozambique, Nigeria, Rwanda, Tanzania, Togo, Uganda.

Typhlodromalus sinespinosus Moraes, Zannou & Oliveira, sp. nov. (Fig. 10) DIAGNOSIS — This species is different from the other Typhlodromalus species by a combination of the following characters: absence of S 5, dorsal shield with lateral setae (except Z 5) smooth and calyx of spermatheca basally bulged. FEMALE — (11 specimens measured). Dorsum — Dorsal shield sculptured, with roundish elements centrally, sparse striae throughout, with an indentation on each side, near S 5 /Z 5; 340 (320–384) long and 210 (192–229) wide. Idiosomal setal pattern 10 A: 8 B/JV– 3:ZV, a characteristic which distinguishes the athiasae species group of Chant & McMurtry (2005 a). Setae j 1 34 (30– 37), j 3 44 (38–46), j 4 16 (14–19), j 5 15 (13–19), j 6 20 (18–26), J 2 21 (16–24), J 5 8 (8–10), z 2 24 (21–26), z 4 39 (35–43), z 5 14 (11–16), Z 1 19 (16–24), Z 4 54 (48–58), Z 5 70 (62–75), s 4 58 (53–61), S 2 47 (38–53), S 4 28 (19–40), r 3 20 (16–24), R 1 19 (14–24). All setae smooth, except Z 4 and Z 5 which are serrate. S 5 absent. Peritreme — Extending anterior to base of j 1. Venter — Distances between St 1 –St 3 67 (64–70), St 2 –St 2 68 (64–72), St 5 –St 5 78 (67– 86). Ventrianal shield 115 (107–125) long, 59 (53–62) wide at level of Zv 2 and 68 (62–74) wide at level of anus, with a pair of elliptical pores slightly posterior to Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp­tipped, except Jv 5 which is serrate. Chelicera — Movable digit 38 (37–40) long, with 3 to 4 teeth; fixed digit 32 (31–33) long, with 9 to 10 teeth and a discernable pilus dentilis. Spermatheca — Calyx saccular, with a distinctly constricted region followed by a flared region distally, 18 (16–20) long; atrium distinct. Legs — Macrosetae blunt on genua I and II and tibiae III and IV, and knobbed on genua III and IV and tarsus IV; SgeI 21 (19–22), Sge II 25 (22–27), Sge III 33 (29–35), Sti III 23 (21–24), Sge IV 52 (48–56), Sti IV 30 (27–34), St IV 66 (59–70). Chaetotaxy of genu III: 1­2 / 1,2 / 0­1. MALE — Unknown. LOCALITY AND TYPE MATERIAL — Holotype female and 4 paratype females from Elephantopus mollis, 27 km S Yaounde, Central Province, Cameroon, 06­II­ 1991, A. Onzo, deposited at ESALQ­USP; 1 paratype female from Sida corymbosa, 1 km SW Nkoteng, Central Province, Cameroon, 05­II­ 1991, A. Onzo, deposited at IITAIM; 1 paratype female from Securinega virosa, 1 km SW Nkoteng, Central Province, Cameroon, 05­II­ 1991, A. Onzo, deposited at ESALQ­USP. One paratype female from Solanum melongena, Mile 71, Bo Road, Sierra­Leone, 14 ­I­ 1992, B.I. Kamara, deposited at IITAIM; 1 paratype female, from Synedrella nodiflora, North of Mayamba, Sierra­Leone, 17 ­I­ 1992, B.I. Kamara, deposited at ESALQ­USP. One paratype female from Gossypium sp., 7 miles W Busembatia, Uganda, 05­XII­ 1991, Mugarura, deposited at IITAIM. One paratype female from Erigeron floribundus, 23 km E Kawambosa, Zambia, 14 ­VI­ 1992, J.S. Yaninek, deposited at ESALQ­USP. ETYMOLOGY — The epithet sinespinosus refers to fact that most of the dorsal idiossomal setae are smooth, differing from the closely related T. spinosus. REMARKS — This new species is very similar to Typhlodromalus spinosus (Meyer & Rodrigues 1966), but the latter differs by having serrate dorsal shield setae, except j 1, j 4, j 5, J 5, z 5, Z 1, r 3 and R 1, which are smooth. Typhlodromalus olombo also resembles this new species, but it has S 5 present and Jv 5 smooth. Typhlodromalus longisetatus sp. nov. and T. tenuicalyx resemble this new species by the absence of S 5, but both have calyx of spermatheca slender; in addition, T. longisetatus has dorsal shield setae subequal in length (except J 5).

Typhlodromalus tenuicalyx Moraes, Zannou & Oliveira, sp. nov. (Fig. 12) DIAGNOSIS ��� This species is characterized by a combination of the following characters: dorsal shield without indentation near S 5 /Z 5; S 5 absent, dorso��central setae short and calyx of spermatheca slender. FEMALE ��� (1 specimen measured). Dorsum ��� Dorsal shield sculptured anteriorly to J 2, with round elements centrally, irregular elements sublaterally and few striae along anterolateral margins, without indentation near S 5 /Z 5; 278 long and 168 wide. Idiosomal setal pattern 10 A: 8 B/JV�� 3:ZV, a characteristic which distinguishes the athiasae species group of Chant & McMurtry (2005 a). Setae j 1 26, j 3 34, j 4 11, j 5 11, j 6 14, J 2 16, J 5 6, z 2 14, z 4 32, z 5 11, Z 1 18, Z 4 43, Z 5 62, s 4 43, S 2 42, S 4 24, r 3 18, R 1 14. All setae smooth, except Z 4 and Z 5 which are serrate. S 5 absent. Peritreme ��� Extending beyond base of j 1. Venter ��� Sternal shield with bi��dentate median lobe on posterior margin. Distances between St 1 ���St 3 62, St 2 ���St 2 59, St 5 ���St 5 66. Ventrianal shield 98 long, 58 wide at level of Zv 2 and 56 wide at level of anus, with a pair of elliptical pores approximately aligned with Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 27 long, with 4 teeth; fixed digit 23 long, with 9 teeth and a distinguishable pilus dentilis. Spermatheca ��� Calyx slender, 9 long; atrium distinct, small. Legs ��� Macrosetae blunt or with very tiny knobs; Sge I 22 Sge II 21, Sge III 29, Sti III 20, Sge IV 43, Sti IV 29, St IV 58. Chaetotaxy of genu III: 1��2 / 1,2 / 0��1. MALE ��� Unknown. LOCALITY AND TYPE MATERIAL ��� Holotype female from Alchornea cordifolia, Umudike, Nigeria, 13 ��IV�� 1992; A. Onzo, deposited at ESALQ��USP. ETYMOLOGY ��� The epithet tenuicalyx refers to the slender calyx of the spermatheca of this new species. REMARKS ��� Differently from most of the species in this genus, as reported by Chant & McMurtry (2005 a), the margin of the dorsal shield of T. tenuicalyx sp. nov. does not have an indentation near S 5 /Z 5. Typhlodromalus sinespinosus Moraes, Zannou & Oliveira, sp. nov. and T. olombo are similar to this new species, but both have an indentation near S 5 /Z 5 and calyx saccular with a distinctly constricted region followed by a flared region distally. Moreover, T. sinespinosus has most dorsal shield setae longer and T. olombo has seta S 5 present. Typhlodromalus aripo, Amblydromalus congeae (De Leon 1965) and Typhlodromalus evansi (Chant 1959) also resemble this new species, but they have seta S 5 present, tubular calyx and much more evident atrium. Similarly to this new species, T. longisetatus does not have S 5, but its dorsal shield setae (except J 5) are of subequal lengths., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 23-24, DOI: 10.5281/zenodo.171564, {"references":["Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224.","De Leon, D. (1965) Phytoseiid mites from Puerto Rico with descriptions of new species (Acarina: Mesostigmata). The Florida Entomologist, USA, 48 (2), 121 - 131.","Chant, D. A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species. The Canadian Entomologist, Canada, Supplement 12, 166 pp."]}

Euseius rhusi (Van der Merwe) (Fig. 24) Amblyseius (Amblyseius) rhusi Van der Merwe, 1965: 63 Euseius rhusi, Moraes et al. 1986: 52 Typhlodromalus rhusi, Moraes et al., 2004: 203 Euseius rhusi, Chant & McMurtry, 2005 a: 215 FEMALE ��� (Specimens measured ��� Burundi: 1; Kenya: 4; Rwanda: 3). Dorsum ��� Dorsal shield mostly smooth, with few striae along anterolateral margins, 367 (336���384) long and 264 (238���285) wide. Setae j 1 25 (21���27), j 3 10 (10���13), j 4 8 (5���10), j 5 7 (5���10), j 6 9 (8���11), J 2 10 (8���11), J 5 8 (6���10), z 2 11 (8���14), z 4 11 (8���14), z 5 9 (6���10), Z 1 11 (8���14), Z 4 26, Z 5 36 (34���38), s 4 18, S 2 18, S 4 16, S 5 15 (14���15), r 3 15 (14���15), R 1 13 (13���14). All setae smooth. Peritreme ��� Extending almost to j 1. Venter ��� Sternal shield with tridentate median lobe on posterior margin. Distances between St 1 ���St 3 62 (58���66), St 2 ���St 2 66 (62���72), St 5 ���St 5 78 (70���86). Ventrianal shield 104 (96���112) long, 58 (53���62) wide at level of Zv 2 and 65 (58���72) wide at level of anus, with a pair of large pores posteromesad of Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharp��tipped. Chelicera ��� Movable digit 30 long, with 1 small tooth; fixed digit 27 (25���28) long, with 3 small teeth and a clearly distinguishable pilus dentilis. Spermatheca ��� Calyx mostly slender, bulged near atrium and flared distally, 42 (38��� 45) long; atrium distinct. Legs ��� Macrosetae sharp��tipped; Sge II 22 (19���24), Sge III 37 (35���40), Sti III 29 (26��� 32), Sge IV 66 (61���70), Sti IV 56 (51���62), St IV 63 (53���72). Chaetotaxy of genu III: 1��2 / 1,2 / 0��1 MALE ��� (Specimens measured ��� Kenya 3; Rwanda: 1). Dorsum ��� Dorsal shield pattern as in female, 268 (263���273) long and 208 (195���218) wide. Setae j 1 23 (23���25), j 3 8, j 4 5 (5���6), j 5 5, j 6 6 (5���8), J 2 8, J 5 5, z 2 8 (8���10), z 4 9 (8��� 10), z 5 5 (5���6), Z 1 8 (8���10), Z 4 9 (8���10), Z 5 26 (24���28), s 4 12 (10���13), S 2 10 (9���10), S 4 8 (8���10), S 5 8, r 3 11 (10���13), R 1 10 (8���13). All setae smooth. Peritreme ��� Extending to level of z 2. Venter ��� Ventrianal shield subtriangular, with transversal striae anteriorly and with a constriction at level of anus; 108 (100���115) long, 164 (160���168) wide at the anterior corners; with 3 pairs of preanal setae, 4 pairs of lyrifissures (1 anterior to Jv 1 and 3 anterolateral) and a pair of large pores posterior to Jv 2. Spermatodactyl ��� Shaft 25 (23���26) long. Legs ��� Macrosetae sharp��tipped; Sge II 25, Sge III 34 (30���38), Sti III 30 (28���33), Sge IV 53, Sti IV 43 (38���49) and St IV 54 (50���58). Chaetotaxy of genu III same as in female. SPECIMENS EXAMINED �� Burundi: 16 km S Rutana, ISABU��MOSO, on unknown plant, 13 ��XII�� 1989, J.S. Yaninek. Kenya: 16 km W Maseno, Nyanza Province, on Psidium guajava, 02��XII�� 1989, J.S. Yaninek; 30 km W Yala, Siaya District, on unknown plant, 02��XII�� 1989, J.S. Yaninek; 10 km E Kisii, on unknown plant, 03��XII�� 1989, J.S. Yaninek; Kisumu District, Ricinus communis, 03��XII�� 1989, J.S. Yaninek. Rwanda: 35 km SE Ruhonzri, Gakonki village, on Persea americana, 08��XII�� 1989 J.S. Yaninek; unknown locality, on Vernonia amygdalina, 08��XII�� 1989, S.J. Yaninek; 13 km SE Kigali, Prefecture of Kigali, on R. communis, 09��XII�� 1989, J.S. Yaninek. WORLD DISTRIBUTION ��� Burundi, Kenya, Mozambique, Reunion Island, Rwanda, South Africa., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on pages 48-49, DOI: 10.5281/zenodo.171564, {"references":["Van der Merwe, G. G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, South Africa, 28, 57 - 76.","Moraes, G. J. de, McMurtry, J. A. & Denmark, H. A. (1986) A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat. EMBRAPA - DDT, Brasilia, Brazil, 353 pp.","Moraes, G. J. de, McMurtry, J. A., Denmark, H. A., Campos, C. B. (2004) A revised catatalog of the mite family Phytoseiidae. Zootaxa, New Zealand, 434, 1 - 494.","Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224."]}

Genus Prasadromalus Chant & McMurtry Prasadromalus Chant & McMurtry, 2005 a: 203 Amblyseius (Typhlodromalus) Muma, 1961: 288 (part) Typhlodromalus, De Leon, 1966: 87 (part) Type species ­ Typhlodromalus breviscutus Moraes, Oliveira and Zannou, 2001: 2

Genus Ueckermannseius Chant & McMurtry Ueckermannia Chant & McMurtry, 2005 a: 201 Ueckermannseius Chant & McMurtry, 2005 b: 337 (replacement name for Ueckermannia) Amblyseius (Typhlodromalus) Muma, 1961: 288 (part) Typhlodromalus, De Leon, 1966: 87 (part) Type species��� Amblyseius (Amblyseius) munsteriensis Merwe, 1965: 67 The species reported subsequently under this genus fit its original description generally well, except for the relative lengths of s 4 and Z 5 in some of the species. While in the original description of this genus it was mentioned that s 4 and Z 5 are not greatly longer than other dorsolateral setae, most of the new species described here have Z 5 2 to 3 times longer than most other dorsolateral setae. Discussions with the authors of Ueckermannseius led to the conclusion that the latter species should be included in that genus. The original diagnosis of the genus is hereby modified to account for the fact that some species of Ueckermannseius have setae s 4 and Z 5 longer than other dorsolateral setae., Published as part of De Moraes, G. J., Zannou, I. D., Oliveira, A. R., Yaninek, J. S. & Hanna, R., 2006, Phytoseiid mites of the subtribes Typhlodromalina and Euseiina (Acari: Phytoseiidae: Euseiini) from sub��Saharan Africa, pp. 1-52 in Zootaxa 1114 on page 25, DOI: 10.5281/zenodo.171564, {"references":["Chant, D. A. & McMurtry A. J. (2005 a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, USA, 31: 187 - 224.","Chant, D. A. & McMurtry A. J. (2005 b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiide): Part VII. Typhlodromipsini n. tribe. International Journal of Acarology, USA, 31: 315 - 340","Muma, M. H. (1961) Subfamilies, genera, and species of Phytoseiidae (Acarina: Mesostigmata). Florida State Museum Bulletin, USA, 5 (7), 267 - 302.","De Leon, D. (1966) Phytoseiidae of British Guyana with keys to species (Acarina: Mesostigmata). Studies on the Fauna of Suriname and other Guyanas, 8, 81 - 102.","Van der Merwe, G. G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, South Africa, 28, 57 - 76."]}

Amblydromalus hum (Pritchard & Baker) (Fig. 1) Amblyseius (Amblyseius) hum Pritchard & Baker, 1962: 271 Typhlodromalus hum, Moraes et al., 2004: 198 Amblydromalus hum, Chant & McMurtry, 2005 a: 207 FEMALE — (Specimens measured — Democratic Republic of Congo: 1 specimen of slide labeled as “ holotype ”; Ghana: 2; Sierra­Leone: 3). Dorsum — Dorsal shield smooth in the central area, with some striation along lateral margins, 326 (322–331) [330] long and 218 (216–221) [220] wide. Setae j 1 29 (26–32) [33], j 3 34 (32–38) [36], j 4 8 (6–11) [8], j 5 7 (6–8) [8], j 6 9 (8–11) [8], J 2 10 (10–11) [10], J 5 6 (5– 8) [6], z 2 14 (11–18) [13], z 4 10 [11], z 5 8 (6–8) [9], Z 1 10 (10–11) [11], Z 4 10 (8–13) [8], Z 5 78 (69–83) [87], s 4 48 (43–56) [57], S 2 12 (10–16) [12], S 4 12 (10–16) [13], S 5 10 (8–14) [11], r 3 14 (11–16) [12], R 1 10 (6–13) [9]. All setae smooth, except Z 5 which is lightly serrate. Peritreme — Extending almost to j 1. Venter — Distances between St 1 –St 3 55 (53–58) [58], St 2 –St 2 71 (67–74) [75], St 5 – St 5 74 (72–77) [75]. Ventrianal shield 100 (96–102) [102] long, 47 (43–51) [49] wide at level of Zv 2 and 68 (62–70) [74] wide at level of anus, with a pair of large pores posteromesad of Jv 2. Two pairs of metapodal shields; caudoventral setae smooth and acute. Chelicera — Movable digit 28 long, with 4 teeth; fixed digit 25 long, with 8 teeth and a distinguishable pilus dentilus. Spermatheca — Calyx tubular, considerably narrower near atrium 29 (26–32) long; atrium bulbous. Legs — Macrosetae sharp­tipped; Sge I 34 (34–35) [36], Sge II 33 (30–34) [34], Sge III 39 (35–42) [41], Sti III 28 (26–30) [31], Sge IV 59 (56–64) [61], Sti IV 36 (35–40) [34], St IV 69 (67–72) [78]. Chaetotaxy of genu III: 1­2 / 1,2 / 0­1. SPECIMENS EXAMINED: Ghana: 2 km W Jakobu, Ashanti Region, on Pteridium sp., 09­XI­ 1989, J.S. Yaninek; 2 km W Jakobu, Ashanti Region, on Raphia farinifera, 10 ­ XI­ 1989, J.S. Yaninek. Democratic Republic of Congo (Belgian Congo): Holotype female from Kinshasa (= Leopoldville), on unknown tree, 20 ­IV­ 1955, E.W. Baker. Sierra­Leone: 15 km N Magbusaka, on Mangifera indica, 15 ­XII­ 1991, A.F. Kanu; Waterloo, on Anacardium occidentale, 18 ­XII­ 1991, J.S. Yaninek; Mile 71, Bo road, on Citrus sp., 14 ­I­ 1992, J.S. Yaninek. WORLD DISTRIBUTION — Democratic Republic of Congo, Ghana, Sierra­Leone.

Typhlodromalus olombo (Pritchard & Baker) (Fig. 9) Amblyseius (Amblyseius) olombo Pritchard & Baker, 1962: 252 Typhlodromalus olombo, Moraes et al., 2004: 201 Typhlodromalus olombo, Chant & McMurtry 2005 a: 199 FEMALE — (Specimens measured — Cameroon: 1; Democratic Republic of Congo: 1 in addition to 1 paratype; Kenya: 2; Uganda: 1). Dorsum — Dorsal shield sculptured anteriorly to Z 4, with roundish elements centrally, irregular elements sublaterally and few striae along anterolateral margins, with an indentation on each side, near S 5 /Z 5; 368 (317–387) [283] long and 226 (202–238) [180] wide. Setae j 1 35 (32–39) [33], j 3 42 (41–44) [42], j 4 15 (14–16) [16], j 5 16 (14–18) [16], j 6 21 (18–25) [20], J 2 24 (22–25) [21], J 5 8 (7–10), z 2 20 (18–21) [18], z 4 36 (32–39) [38], z 5 15 (13–16) [14], Z 1 23 (22–24) [17], Z 4 52 (48–60) [45], Z 5 77 (74–81) [71], s 4 53 (51–56) [52], S 2 46 (41–51) [47], S 4 26 (19–32) [29], S 5 13 (10–18) [8], r 3 23 (18–28) [22], R 1 16 (14–18) [13]. All setae smooth, except Z 4 and Z 5 which are serrate; Peritreme — Extending anterior to base of j 1. Venter — Distances between St 1 –St 3 67 (62–70) [63], St 2 –St 2 71 (64–74) [69], St 5 – St 5 83 (76–86). Ventrianal shield 118 (108–128) [123] long, 64 (60–67) [66] wide at level of Zv 2 and 70 (60–75) [63] wide at level of anus, with a pair of elliptical pores slightly posterior to Jv 2. One pair of metapodal shields; caudoventral setae smooth and sharptipped. Chelicera — Movable digit 41 (40–43) long, with 4 teeth; fixed digit 36 (35–38) long, with 9 teeth and a distinguishable pilus dentilis. Spermatheca — Calyx saccular with a distinctly constricted region followed by a flared region distally, 22 (21–23) long; atrium distinct. Legs — Macrosetae blunt on genua I and II and tibiae III and IV, and with a tiny knob on genua III and IV and tarsus IV; Sge I 24 (23–25) [23], Sge II 28 (25–30) [27], Sge III 33 (32–37) [34], Sti III 25 (23–28) [27], Sge IV 52 (44–56) [53], Sti IV 35 (32–37) [31], St IV 78 (62–86) [65]. Chaetotaxy of genu III: 1­2 / 1,2 / 0­1. SPECIMENS EXAMINED: Cameroon: Western Province, on Crassocephalum biafrae, 26 ­I­ 1991, J.S. Yaninek. Democratic Republic of Congo (Belgian Congo): paratype female, from Mt. Hoyo, on Hibiscus sp., 18 ­IV­ 1955, E.W. Baker; 19 km W Kinzoavuete, Bas­Congo, on Sida corymbosa, 10 ­VII­ 1991, A. Onzo. Kenya: 39 km W Nakuru, Valley Province, on Triumphetta sp., 04­XII­ 1989, J.S. Yaninek; 39 km W Nakuru, Valley Province, on unknown plant, 04­XII­ 1989, J.S. Yaninek. Uganda: Kabarole, Mountain Moon Hotel, on Manihot esculenta, 08­X­ 1990, Karamura. WORLD DISTRIBUTION — Cameroon, Democratic Republic of Congo, Kenya, Uganda.