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The genusCicadulaZetterstedtis reported for the first time from Pakistan with a newly recordedCicadulasimlaensisViraktamath & Yeshwanth. In addition,Platymetopius fidelis(Distant) is also rediscovered from Pakistan and studied here. A brief diagnosis along with the digital photographs of the habitus and male genitalia of bothspecies are provided. General distribution of these species in both Palaearctic and Oriental regions is also briefly discussed.

Stirellus pakistanicus Shah & Duan sp. n. (Fig. 1A–J) Measurement. Male: 3.9mm. Description. Coloration. General color chocolate brown (Fig. 1A–D). Crown anterior portion black with two conspicuous black spots normally present in middle adjacent to eyes (Fig. 1A, 1C). Eye reddish brown, surrounded with yellow band (Fig. 1A–D). Face black with yellowish patch on frontoclypeus (Fig. 1D). Pronotum with black band along the anterior margin, whitish across the middle while yellowish brown at the posterior margin (Fig. 1A, 1C). Mesonotum and scutellum yellowish brown (Fig. 1A, 1C). Thorax and abdomen black in ventral and lateral view (Fig. 1B). Abdominal segments separated by yellowish bands (Fig. 1B). Forewing dark brownish with prominent venation (Fig. 1A–B). Morphology. Crown narrower than pronotum, with anterior margin slightly angulate and smoothly curved to face (Fig. 1A–D). Crown almost equal to or a little longer than breadth between eyes (Fig. 1A, 1C). Ocellus next to eye slightly below anterior margin of crown (Fig. 1B, 1D). Antennae long and thin. Face slightly longer than broad (Fig. 1D). Pronotum median length a little smaller than median length of crown with posterior margin slightly concave (Fig. 1A, 1C). Mesonotum and scutellum generally shorter than pronotum. Scutellum with a transverse impression in middle (Fig. 1C). Forewing macropterous (Fig. 1A–B). Male genitalia. Pygofer lobe broad basally with apex truncate (Fig. 1E). Pygofer side with few dorsal macrosetae (Fig. 1E). Subgenital plate large, triangular, with a row of uniseriate macrosetae (Fig. 1F). Valve triangular. Style narrow with digitate apophysis, laterally curved (Fig. 1G). Connective H-shaped with stem slightly shorter than arms (Fig. 1H). Aedeagal shaft long, bent at near base, nearly parallel to aedeagal base, with a slight dorsal curve at mid-length in lateral view, apex pointed and slightly curved dorsad, gonopore apical (Fig. 1I–J). Material examined. Holotype: Pakistan: Punjab: ♂, Attock, 33°51′4.2192′′ N, 72°15′59.292′′ E, sweeping hand net, 16 June 2021, coll. Waqar Khan and Bismillah Shah. Distribution. Pakistan [Attock (Fig. 2)]. Remarks. The new species described in this paper appears to key out to S. indrus (couplet 8) in our previous key (See Shah et al., 2020b). However it can readily be distinguished from S. indrus by the crown with anterior portion black (Fig. 1A, 1C); two conspicuous black spots normally present in middle adjacent to eyes (Fig. 1A, 1C); and face black with yellowish patch on frontoclypeus (Fig. 1D). Also, the new species has coloration similar to some species described from the Americas (e.g., Stirellus bicolor, Stirellus picinus) and Australia (Stirellus petfordensis). Etymology. This species is named for the country Pakistan, where the type specimen has been collected., Published as part of Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad & Duan, Yani, 2022, A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan, pp. 344-350 in Zootaxa 5125 (3) on pages 346-347, DOI: 10.11646/zootaxa.5125.3.6, http://zenodo.org/record/6443920, {"references":["Shah, B., Naveed, H. & Duan, Y. N. (2020 b) A new species and four new records of Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species. Zootaxa, 4895 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 4895.3.5"]}

Stirellus rotundus (Singh-Pruthi, 1930) Distribution. Pakistan [Muzaffarabad (Fig. 2)], India., Published as part of Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad & Duan, Yani, 2022, A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan, pp. 344-350 in Zootaxa 5125 (3) on page 347, DOI: 10.11646/zootaxa.5125.3.6, http://zenodo.org/record/6443920, {"references":["Singh-Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68."]}

Shah, Bismillah, Khan, Waqar, Naveed, Hassan, Shakeel, Muhammad, Duan, Yani (2022): A new species in the grassland leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan. Zootaxa 5125 (3): 344-350, DOI: https://doi.org/10.11646/zootaxa.5125.3.6

Maiestas maculata (Singh-Pruthi) n. rec. (Fig. 2) Cicadula maculata Singh-Pruthi, 1930: 58���59, figs 80���81, plate V, fig. 2. Thamnotettix prabha Singh-Pruthi, 1930: 62, figs 85, 86, plate V, figs 6, 6a. Synonymised by Webb & Viraktamath, 2009: 41. Recilia prabha, Ghauri, 1980: 166���169, figs 1, 3���11. Deltocephalus (Recilia) maculata, Dash & Viraktamath, 1998: 32, figs 260���269. Maiestas maculata, Webb & Viraktamath, 2009: 22, comb. nov. Maiestas maculata, Zhang & Duan, 2011: 37���39, figs 33���35, plate IV: E, plate V: P, plate VI: P. Length. Male: 3.4mm Description. Coloration. Pale to yellowish brown (Fig. 2A���D). Crown with variable black spots (Fig. 2A, 2C). Face stramineous to yellowish brown, with black patches adjacent to antennal bases (Fig. 2D). Pronotum occasionally with black patches. Visible mesonotum with basal angles black (Fig. 2A, 2C). Legs with or without blackish spots (Fig. 2B). Morphology. Head as broad as pronotum (Fig. 2A, 2C). Forewing macropterous or submacropterous; inner anteapical cell open at base, sometimes outer anteapical cell absent (Fig. 2A���B). Male genitalia. Subgenital plate moderately long, subtriangular, lateral margin somewhat convex (Fig. 2F). Style with preapical lobe short, apophysis digitate, long and faintly sinuate (Fig. 2F). Connective almost as long as aedeagus (Fig. 2G). Aedeagus boat-shaped dorsally (Fig. 2H), straight in lateral aspect and narrow at base, broader in middle and thereafter gradually narrowed to acute upturned apex (Fig. 2I). Material examined. Pakistan: Azad Kashmir: 1♂, Banjosa, 33��48���36.9432 N, 73��49���14.304 E, sweep net, 26 July 2018, coll. Bismillah Shah (AAU). Distribution. Pakistan (new record), China, India. Remarks. This species, originally described from India (Calcutta and Sikkim) is recorded here for the first time from Pakistan. It can readily be distinguished from other species of the genus by its external appearance, e.g., with variable black patches on head and thorax, and shape of the aedeagus and style., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on pages 404-405, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and Description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Ghauri, M. S. K. (1980) Illustrated redescription of two of Pruthi's species of Cicadelloidea from India. Reichenbachia Staatliches Museum fur Tierkunde in Dresden, 18, 165 - 171.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1"]}

Maiestas setosa (Ahmed, Murtaza & Malik) Recilia setosa Ahmed, Murtaza & Malik, 1988: 412, fig. 2. Maiestas setosa, Webb & Viraktamath, 2009: 20, comb. nov.; Naveed et al., 2019: 287. Material examined. No material examined. Distribution. Pakistan. Remarks. The identity of this species remains uncertain due to the poor original description and figures and because the type series from Karachi could not be found in the repository indicated in the original description, Zoological Museum, University of Karachi (Khatri & Webb, 2010: 11)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 406, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Ahmed, M., Murtaza, B. & Malik, K. F. (1988) Some new aphrodine leafhoppers from grasslands of Karachi, Pakistan. Pakistan Journal of Zoology, 20, 409 - 421.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1"]}

Maiestas sp. 1 (Fig. 7) M. samuelsoni, Naveed et al., 2019, fig. 3L���M, misidentification. Material examined. Pakistan: 2♂♂, Khyber Pakhtunkhwa: Pirsabak, 34��0���37.1196 N, 72��3���3.6648 E, sweep net, 12 July 2018; 3♂♂, 4♀♀, same data, light trap; 2♂♂, Balakot, 34��34���18.1668 N, 73��21���46.6668 E, sweep net, 12 August 2019; Punjab: 1♂, Hazro, 33��54���44.208 N, 72��29���12.084 E, sweep net, 31 July 2018. All collected by Bismillah Shah (AAU). Remarks. The identity of this species is uncertain. It is similar to the following un-named species in having the forewings with additional cross-veins and similar male genitalia but its colour pattern is different. Its male genitalia are also similar to those of some other Maiestas species, e.g., M. samuelsoni (Knight). Although resembling the latter species (originally described from the Pacific region) in shape of aedeagus and style, it differs by its longer head with more distinct markings and forewing with extra cross-veins. It is likely that the material studied here is the same species recorded from Pakistan as M. samuelsoni by Naveed et al. (2019). These authors recorded three males (NWAFU) from the same region (Khyber Pakhtunkhwa) as the specimens examined here, but with the figures of Zhang & Duan (2011) from Chinese specimens, the latter also possibly misidentified. The Pakistan specimens deposited in NWAFU need to be re-examined to confirm their identity., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 412, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1"]}

Maiestas trispinosa (Dash & Viraktamath) n. rec. (Fig. 6) Deltocephalus (Recilia) trispinosus Dash & Viraktamath, 1998: 35, figs 296���304 Maiestas trispinosa, Webb & Viraktamath, 2009: 38, comb. nov. Length. Male: 4.1mm. Description. Coloration. Yellowish brown (Fig. 6A���D). Head and thorax pale yellow (Fig. 6A, 6C). Crown with six small, dark brownish marks at anterior margin and some yellowish brown patches on each side of mid-line (Fig. 6A, 6C). Pronotum with ochraceous shades and six yellow brown longitudinal stripes (Fig. 6A, 6C). Mesonotum and scutellum pale with yellowish brown spots (Fig. 6A, 6C). Eye tinged with yellowish brown (Fig. 6A���D). Frontoclypeus chocolate brown with pale transverse stripes (Fig. 6D). Leg marked with dark brown (Fig. 6B). Forewing light brown, with prominent pale venation (Fig. 6A���B). Morphology. Head slightly wider than pronotum, almost equal in length to width between eyes (Fig. 6A, 6C). Ocellus next to eye on anterior margin. Pronotum median length almost equal to the median length of crown (Fig. 6A, 6C). Forewing macropterous, almost same length as hind wings (Fig. 6A���B). Male genitalia. Pygofer quadrate (Fig. 6E). Subgenital plate moderately long, triangular, lateral margins almost straight, with a row of uniseriate macrosetae, apically rounded and curved dorsad (Fig. 6F). Valve triangular (Fig. 6F). Style narrow, with prominent preapical shoulder, apex digitate and sinuate (Fig. 6F). Connective almost equal in length to aedeagus (Fig. 6G���I). Aedeagal shaft extended far beyond pygofer, curved ventrally (Fig. 6E); shaft compressed dorsoventrally, in lateral view broader in basal two thirds then abruptly narrowed to acute apex, in dorsoventral view with needle-like apex (Fig. 6G���H); pair of subapical spine-like lateral processes adpressed to shaft (Fig. 6G���H). Material examined. Pakistan: Punjab: 2♂♂, Bansra Gali, 33��54���17.532 N, 73��22���13.9656 E, sweep net, 1 August 2019, coll. Bismillah Shah (AAU). Distribution. Pakistan (new record), India. Remarks. This species can be distinguished by the lateral spine-like processes of the aedeagus. The Pakistan specimens show very slight differences compared to the original Indian material with respect to the shape of the subgenital plates and style apical process, as seen in the original figures given by Dash & Viraktamath (1998) and images of type material sent by C. Viraktamath., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on pages 408-412, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Zhang, Y. L. & Duan, Y. N. (2011) Review of the Deltocephalus group of leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae). Zootaxa, 2870 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2870.1.1","Dash, P. C. & Viraktamath, C. A. (1995) Two new species of grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) from Orissa, India. Hexapoda, 7, 71 - 78."]}

Maiestas indica (Singh-Pruthi) (Fig. 1E) Allophleps indica Singh-Pruthi, 1936: 120, fig. 132; plate 9, fig. 3. Allophleps delhiensis Rao & Ramakrishnan, 1990: 111, figs 1���9. Synonymised by Dash & Viraktamath, 1998: 35. Deltocephalus (Recilia) indicus, Dash & Viraktamath, 1998: 35���36, fig. 305. Maiestas indica, Webb & Viraktamath, 2009: 21, comb. nov.; Naveed et al. (2019: 287). Material examined. No material examined. Distribution. Pakistan, India. Remarks. This species was originally described based on both male and female specimens but a female from Pakistan ���Layallpur [Faisalabad], Punjab, at light; 10.X.29 (Coll. A. Rahman)��� (ZSI), was listed as the ��� holotype ��� together with ���four other specimens from the type locality and numerous specimens from several other localities in the Indian Museum collection���. The specimens other than the female holotype may be considered paratypes but it is not clear from which specimen the male genitalia were figured and the male genitalia slide is now missing (pers. com. C. Viraktamath). Therefore, Pruthi���s figure of the male could be from a specimen either from Pakistan or India. No further material of this species from Pakistan has been studied since the original description and although the original material was examined by Dash & Viraktamath (1998), including a male paratype, their figures are from their Indian specimens (pers. com. C. Viraktamath). Despite the above uncertainties, assuming that the female holotype and the male specimen illustrated by Pruthi are conspecific, the identity of the species is clear based on the aedeagus with a distinctive large subapical ventral process (Fig. 1E). The species can also be recognized by its accessary forewing cross-veins, similar to M. pruthii and Maiestas sp. 1 and Maiestas sp. 2, but from these it also differs by its shorter head as shown in Pruthi���s original figure., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 403, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11, 101 - 131.","Rao, V. R. S. & Ramakrishnan, U. (1990) New species of Aconurella and Paramesodes (Homoptera: Cicadellidae) from India. Journal of the Bombay Natural History Society, 87, 268 - 271.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Maiestas sinuata Shah & Duan sp. n. (Fig. 3) Length. Male: 3.3mm. Description. Coloration. Yellowish brown (Fig. 3A–B). Head and thorax creamy white with yellow markings (Fig. 3A, 3C). Crown with eight small, dark brownish marks, with ochraceous patches on each side of median longitudinal suture (Fig. 3A, 3C). Pronotum with four yellow longitudinal stripes (Fig. 3A, 3C). Mesonotum and scutellum pale with yellowish brown spots (Fig. 3A, 3C). Eye color tinged with yellowish grey. Ocellus white (Fig. 3A–C). Frontoclypeus dark brown with pale transverse stripes (Fig. 3D). Legs marked with dark brown (Fig. 3B). Forewing brown, with prominent pale venation (Fig. 3A–B). Morphology. Head slightly wider than pronotum and slightly longer than width between eyes (Fig. 3A, 3C). Ocellus next to eye on anterior margin (Fig. 3A–C). Pronotum median length almost equal to the median length of crown (Fig. 3A, 3C). Forewing macropterous, exceeding abdomen when at rest (Fig. 3A–B). Male genitalia. Pygofer side longer than height in lateral aspect with rounded apical margin (Fig. 3E). Subgenital plate moderately long, triangular, with row of marginal uniseriate macrosetae (Fig. 3F). Style narrow, with prominent preapical shoulder; apex digitate, slightly curved laterally (Fig. 3F). Connective almost equal in length to aedeagus (Fig. 3G–H). Aedeagal shaft in dorsal view broad at base (Fig. 3G), gradually tapered towards pointed apex; in lateral view, slightly sinuate with basal half concave ventrally and apical half concave dorsally, finely tapered from base to apex (Fig. 3H). Material examined. Holotype ♂: Pakistan: Punjab, Kahuta, 33°35′28.3776 N, 73°22′12.234 E, sweep net, 25 July 2018, coll. Bismillah Shah (AAU). Distribution. Pakistan. Etymology. The species named for the slightly sinuate aedeagal shaft in lateral view. Remarks. This species has a similar sinuate aedeagal shaft to M. chandrai Fletcher & Dai (2019) (replacement name for M. viraktamathi Fletcher & Dai, 2018), from Australia, but differs from the latter species in external appearance and other genitalia structures, i.e., the subgenital plate with apical fine setae, style with apical process thicker and aedeagus lacking a ventrobasal “heel”, the latter feature placing M. chandrai in the M. albomaculatus group of Dash & Viraktamath (1998).

Maiestas Distant, 1917 Maiestas Distant, 1917: 312 ( Type species: Maiestas illustris Distant, 1917). For full details of synonymy, see Webb & Viraktamath, 2009: 14���15. Diagnosis. Generally pale brown with c rown often with line of submarginal angular spots and pronotum often with longitudinal stramineous bands (Fig 1A), sometimes without these markings (Fig. 2A). Crown triangularly produced, ocellus usually close to eye on anterior margin. Forewing with three anteapical cells, inner cell open. Pygofer without appendage. Subgenital plate short and ovate with outer margin convex to slightly concave; with uniseriate row of macrosetae laterally, sometimes with apical tuft of fine setae. Connective linear with arms longer than stem. Aedeagus fused to connective; shaft elongate, weakly curved dorsally, usually without processes, gonopore apical on dorsal surface, indistinct. Distribution. Old World. Remarks. Maiestas was revised by Webb & Viraktamath (2009). The genus is similar externally to Deltocephalus Burmeister and Recilia but differs by the shape of the aedeagus; in Maiestas the shaft is boat-shaped, at most weekly curved dorsally, sometimes with an apical ventral spine-like process and gonopore apical on dorsal surface. To the previous treatment of the genus from Pakistan (Naveed et al., 2019), a new species and two new country records are added in the present work, together with two species of uncertain identity (see Maiestas spp)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 402, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Distant, W. L. (1917) The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr J. Stanley Gardiner, M. A. Vol. VI, No. VII-Rhynchota, Part II: Suborder Homoptera. Transactions of the Linnean Society of London, 17, 273 - 322. https: // doi. org / 10.1111 / j. 1096 - 3642.1917. tb 00469. x","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Maiestas pruthii (Metcalf) (Fig. 4F���L) Deltocephlaus notatus Singh-Pruthi, 1936: 128���129, fig. 139, plate IX, fig. 10 (Prim.hom.: Deltocephalus notatus Melichar, 1896). Deltocephalus pruthii Metcalf, 1967: 1173 (nom. nov. pro Deltocephalus notatus Singh-Pruthi, 1936). Deltocephalus (Recilia) pruthii, Dash & Viraktamath, 1998: 22���23, figs 150���158. Maiestas pruthii, Webb & Viraktamath, 2009: 20, comb. nov.; Khatri & Webb, 2010: 11, plate 2a, fig. 13, misidentification? (see Remarks); Naveed et al., 2019: 286, figs 2A���C (incorrectly cited as M. subviridis, pers. com. Naveed), Fig. 3i, misidentification? (see Remarks). Material examined. No material examined. Distribution. Pakistan, India. (see Remarks) Remarks. The identity of this species is somewhat uncertain. It was described from the holotype male and allotype female from Pakistan, Layallpur [Faisalabad], but the male genitalia slide of the holotype (ZSI) is now missing (pers. com. Chandra Viraktamath). Although the types were examined by Dash & Viraktamath (1998) their figures are from Indian (Dehli) specimens (pers. com. Chandra Viraktamath) and although showing a very slight difference in the style apex to that of Singh-Pruthi���s original they show a similar relatively long apical extension of the aedeagus and rounded shaft apex (Figs 4J���L). Therefore, Dash & Viraktamath���s material is tentatively considered correctly identified but, to be certain, topotypical material of M. pruthii, i.e., from Faisalabad, should be obtained and examined. The Pakistan material identified as this species by Khatri & Webb (2010) with their figures reproduced by Naveed et al. (2019) show a more tapered aedeagus apex and shorter apical process and vertex with a small dot either side of the coronal suture apex, are here considered to be a different species (see Maiestas sp. 2 below). Despite the shortcomings of Singh-Pruthi���s original genitalia figures, his habitus figure of this species is very good and shows a species with a relatively long acute head with a pair of inverted U-shaped markings basally and forewings with extra cross veins. The extra forewing cross veins are also present in M. indica (described from the same type locality as M. pruthii) and Maiestas sp. 1 and 2 (see Remarks under these species). In conclusion, until the type genitalia or topotypical specimens can be found, the identity of this species is based on the figures of Dash & Viraktamath (1998) together with the images given here from the same specimen (Figs 4F���L)., Published as part of Shah, Bismillah, Naveed, Hassan, Webb, Michael D. & Duan, Yani, 2021, Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records, pp. 401-416 in Zootaxa 5060 (3) on page 406, DOI: 10.11646/zootaxa.5060.3.6, http://zenodo.org/record/5635912, {"references":["Singh-Pruthi, H. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11, 101 - 131.","Melichar, L. (1896) Cicadinen (Hemiptera-Homoptera) von Mittel-Europa. Dames, Berlin, 364 pp. https: // doi. org / 10.5962 / bhl. title. 8568","Metcalf, Z. P. (1967) s. n. In: General Catalogue of the Homoptera. Fascicle VI. Cicadelloidea. Part 10. Euscelidae Section II. Agricultural Research Service, U. S. Department of Agriculture, Washington, D. C., 1078 - 2074.","Dash, P. C. & Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 & 2), 1 - 59.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1","Khatri I., & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H., Sohail, K., Islam, W., Zhang, Y. & Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16"]}

Stirellus capitatus (Distant) n. rec. (Fig. 1) Viridomarus capitatus Distant, 1918: 70. Viridomarus brevialatus Xing, Dai & Li, 2009: 590. Viridomarus laticellus Xing, Dai & Li, 2009: 592. Stirellus capitatus, Li, Dai & Xing, 2011: 271. Stirellus capitatus, Duan, Webb & Zhang, 2016: 108. Measurement. Male: 3.0���4.0mm; female: 4.6mm. Description. Coloration. Stramineous. The vertex lateral margins generally pale stramineous (Fig. 1 A���C); vertex tip with small blackish spot anteriorly (Fig. 1A, 1C). Morphology. Crown longer than twice breadth between the eyes, elongately subtriangular, tapering to apex and widely subacute (Fig. 1 A���C). Ocellus on anterior margin far from eye. Eye comparatively small (Fig. 1 A���C). Anteclypeus narrowing to apex, not exceeding beyond normal curve of genae. Pronotum almost as long as 0.5 times crown median length (Fig. 1 A���C). Mesonotum and scutellum shorter than pronotum (Fig. 1 A���C). Forewing macropterous to submacropterous, generally exposing abdomen apex in female and occasionally in male (Fig. 1 A���B). Male genitalia. Pygofer side with a small number of macrosetae present dorsally, apex slightly angulate (Fig. 1F). Subgenital plate highly convex laterally, macrosetae almost uniseriate centrally (Fig. 1G). Style apophysis digitate, curved laterally (Fig. 1J). Connective with stem shorter than arms (Fig. 1I). Aedeagal shaft long, strongly bent dorsally near base, tip sharp, somewhat curved dorsad, gonopore apical (Fig. 1K). Material examined. Pakistan: Khyber Pakhtunkhwa: 1♀, Jalbai, 34��0���43.8444 N, 72��16���0.0228 E, sweeping hand net, 19 July 2019, coll. Bismillah Shah. Distribution. Pakistan, China, India, Thailand. Remarks. This species can be recognized by its comparatively long head, and by the tip of vertex with small dark brown spot. This is the first record of this species from Pakistan., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 401, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Xing, J. C., Dai, R. H. & Li, Z. Z. (2009) A taxonomic study on the genus Viridomarus Distant (Hemiptera: Cicadellidae: Euscelinae). Acta Zootaxonomica Sinica, 34 (3), 590 - 595.","Li, Z. Z., Dai, R. & Xing, J. (2011) Deltocephalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, 336 pp.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus indrus (Distant) n. rec. (Fig. 2) Typhlocyba indra Distant, 1908: 415. Aconura bipunctella Matsumura, 1914: 232. Paivanana indra, Distant, 1918: 95. Paivanana centristriata Dai & Li, 2003 in Dai, Chen & Li, 2003: 31. Stirellus indrus, Li, Dai & Xing, 2011: 273; Duan, Webb & Zhang, 2016: 116. Measurement. Male: 2.9mm; female: 3.6mm. Description. Coloration. Normally pale greyish (Fig. 2 A���C). Crown with two conspicuous black spots normally between anterior margins of eyes, and to some extent adjacent to eyes than each other (Fig. 2A, 2C). Face having two blackish round marks on each side below the antennal sockets (Fig. 2D). Forewing with longitudinal brown bands, tinged with purple or green color (Fig. 2 A���B). Morphology. Head subconical, nearly subequal in width to pronotum (Fig. 2A, 2C), with anterior margin smoothly curved to face, acutely angled (Fig. 2D). Crown somewhat longer than breadth between eyes (Fig. 2A, 2C). Ocellus next to the eye at anterior margin. Eye comparatively large (Fig. 2 A���D). Anteclypeus tapering towards apex, exceeding beyond the normal curve of genae. Pronotum median length almost equal to the median length of crown (Fig. 2A, 2C). Mesonotum and scutellum smaller than pronotum. Forewing macropterous, normally exposing the apex of abdomen in females and occasionally in males (Fig. 2 A���B). Male genitalia. Pygofer side with few dorsal macrosetae, almost truncate apically (Fig. 2E). Subgenital plate a little concave laterally, macrosetae uniseriate laterally (Fig. 2F). Style narrow, with apophysis slender, digitate, to some extent curved laterally (Fig. 2F). Aedeagal shaft long, bent near base, parallel to base of aedeagus, with tip pointed, gonopore apical (Fig. 2I). Female hind margin of 7 th sternum almost straight. Material examined. Pakistan: Khyber Pakhtunkhwa: 1♀, Nowshera, 34��0���53.8056 N, 71��58���59.3184 E, sweeping hand net, 21 July 2019, coll. Bismillah Shah; 1♀, Faqir Baba, 33��54���53.4888 N, 72��9���56.2176 E, sweep-ing hand net, 20 July 2019, coll. Bismillah Shah; Punjab: 1��, Pir Sohawa, 33��45���37.674 N, 73��3���57.528 E, sweep-ing hand net, 6 August 2018, coll. Bismillah Shah. Distribution. Pakistan, China, India. Remarks. Though there is considerable variation in color pattern of this species, it can be recognized by the vertex with two conspicuous black spots between the anterior margins of the eyes, and by a brown longitudinal line on the forewing., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 401-402, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Matsumura, S. (1914) Die Jassinen und einige neue Acocephalinen Japans. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, 5, 165 - 240.","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Dai, R. H., Chen, X. X. & Li, Z. Z. (2003) A new species of the genus Paivanana (Homoptera: Cicadellidae: Euscelinae) from China. Entomotaxonomia, 25 (1), 31 - 33.","Li, Z. Z., Dai, R. & Xing, J. (2011) Deltocephalinae from China (Hemiptera: Cicadellidae). Popular Science Press, Beijing, 336 pp.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus rotundus (Pruthi) n. rec. (Fig. 4) Arya rotunda Pruthi, 1930: 40. Measurement. Male: 4.0mm. Description. Coloration. Pale to greyish (Fig. 4 A–D). Crown reddish-yellow (Fig. 4A, 4C). Eye and ocellus black (Fig. 4B, 4D). Pronotum with transverse darker greyish shade (Fig. 4A, 4C). Face light brown. Frons with reddish-brown transverse striae in lateral area (Fig. 4D). Forewing brown or stramineous, with brownish venation (Fig. 4 A–B). Legs brown. Abdomen black both dorsally and ventrally in males (Fig. 4 A–B). Morphology. Crown longer than width between eyes, rounded at anterior margin (Fig. 4A, 4C). Eye large, oblique, and projected over pronotal angles laterally (Fig. 4 A–D). Ocellus placed at anterior margin of vertex near eye, barely visible from above (Fig. 4D). Face as long as wide. Antennae long and thin. Pronotum a little longer than vertex with the posterior margin slightly concave (Fig. 4A, 4C). Scutellum with a transverse impression in middle. Male genitalia. Pygofer considerably produced in posterior direction (Fig. 4E). Subgenital plate large, triangular, with a row of uniseriate macrosetae (Fig. 4F). Valve triangular. Style narrow with digitate apophysis, laterally curved (Fig. 4F). Connective with arms well separated, stem slightly shorter than arms (Fig. 4F). Aedeagal shaft long, slender, parallel to aedeagal base, slightly sinuate, tip pointed, curved dorsad, gonopore apical (Fig. 4 G–H). Material examined. Pakistan: Azad Kashmir: 1³, Muzaffarabad, 34°21′44.1144 N, 73°27′10.62 E, sweeping hand net, 14 August 2019, coll. Bismillah Shah. Distribution. Pakistan, India. Remarks. Pruthi (1930) described this species for the first time from India. This is the first record of the species from Pakistan., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 404-406, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68."]}

Stirellus pakhtunensis Shah & Duan sp. n. (Fig. 3) Measurement. Male: 3.6mm. Description. Coloration. General color pale to greyish brown (Fig. 3 A–D). Crown with two spots near posterior margin (Fig. 3A, 3C). Pronotum and scutellum with two dark brown longitudinal stripes (Fig. 3A, 3C). Eye red, surrounded with yellow band (Fig. 3 A–D). Pronotum slightly darker (Fig. 3A, 3C). Scutellum yellowish brown. Forewing brownish with prominent white venation (Fig. 3 A–B). Morphology. Crown almost subequal to pronotum, almost as long as distance between eyes, anteriorly slightly angulate (Fig. 3A, 3C). Eye relatively large. Ocellus next to eye slightly below anterior margin of crown (Fig. 3B, 3D). Face as long as broad, with faded transverse markings near eyes (Fig. 3D). Pronotum median length nearly as long as median length of crown (Fig. 3A, 3C). Mesonotum and scutellum shorter than pronotum. Forewing macropterous; appendix distinctive, prolonged around wing apex; 4 apical and 3 anteapical cells, inner anteapical cell opened at base. (Fig. 3 A–B). Male genitalia. Pygofer lobe broad basally with apex truncate, without macrosetae (Fig. 3E). Subgenital plate slightly convex laterally, apically truncate, macrosetae not uniseriate laterally (Fig. 3F). Valve triangular. Styles narrow, apophysis digitate, slightly curved laterally (Fig. 3F). Connective with stem equal in length to arms (Fig. 3F). Aedeagal shaft long, geniculate basally, nearly parallel to aedeagal base, with preapical anterior expansion visible in lateral and dorsal views, apex pointed and slightly curved dorsad, gonopore apical (Fig. 3 G–H). Material examined. Holotype: Pakistan: Khyber Pakhtunkhwa: 1³, Manki, 34°0′34.5096 N, 73°19′44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. This species can be easily distinguished from other species in this genus by the distinctive external appearance. Etymology. This species is named for Pakhtun-khwa province in Pakistan where the type specimen has been collected.

Shah, Bismillah, Naveed, Hassan, Duan, Yani (2020): A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species. Zootaxa 4895 (3): 398-410, DOI: https://doi.org/10.11646/zootaxa.4895.3.5

Stirellus thattaensis Mahmood, Sultana & Waheed (See Shah et al., 2020: 195, Fig. 4) Stirellus thattaensis Mahmood, Sultana & Waheed, 1972: 82. Material examined. No material examined. Distribution. Pakistan. Remarks. Mahmood et al., (1972) has given detailed description of this species originally described from Pakistan. S. thattaensis is very similar to S. lahorensis in external appearance but differs in male genitalia., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 407, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84."]}

Stirellus Osborn & Ball Athysanus (Stirellus) Osborn & Ball, 1902: 250 (Type: Athysanus bicolor Van Duzee, 1892). Stirellus, Kirkaldy, 1907: 58. Doratulina Melichar, 1903: 198 (Type: Doratulina jocosa Melichar, 1903). Phrynophyes Kirkaldy, 1906: 237 (Type: Phrynophyes phrynophyes Kirkaldy, 1906). Anemochrea Kirkaldy, 1906: 329 (Type: Anemochrea mitis Kirkaldy, 1906). Anemolua Kirkaldy, 1906: 329 (Type: Anemolua hanuala Kirkaldy, 1906). Giffardia Kirkaldy, 1906: 336 (Type: Giffardia dolichocephala Kirkaldy, 1906). Arya Distant, 1908: 338 (Type: Arya rubrolineatus Distant, 1908). Trebellius Distant, 1918: 52 (Type: Trebellius albifrons Distant, 1918). Campbellinella Distant, 1918: 69 (Type: Campbellinella illustrata Distant, 1918). Viridomarus Distant, 1918: 69 (Type: Viridomarus capitatus Distant, 1918). Bituitus Distant, 1918: 70 (Type: Bituitus projectus Distant, 1918). Paternus Distant, 1918: 71 (Type: Paternus pusana Distant, 1918). Volusenus Distant, 1918: 72 (Type: Volusenus lahorensis Distant, 1918). Nandidrug Distant, 1918: 74 (Type: Nandidrug speciosus Distant, 1918). Allectus Distant, 1918: 75 (Type: Allectus notata Distant, 1918). Galerius Distant, 1918: 78 (Type: Galerius indicatrix Distant, 1918). Paivanana Distant, 1918: 95 (Type: Typhlocyba indrus Distant, 1908). Gillettiella Osborn, 1930: 689 (Type: Deltocephalus labiatus Gillette, 1898). Bella Pruthi, 1930: 44 (Type: Bella apicalis Pruthi, 1930). Penestirellus Beamer & Tuthill, 1934: 21 (Type: Penestirellus catalinus Beamer & Tuthill, 1934). Sunda Pruthi, 1936: 112 (Type: Sunda ribeiroi Pruthi, 1936). Pseudaconura Linnavuori, 1952: 182 (Type: Pseudaconura luxorensis Linnavuori, 1952). Malagasiella Evans, 1954: 110 (Type: Malagasiella minima Evans, 1954). Umesaona Ishihara, 1961: 246 (Type: Umesaona asiatica Ishihara, 1961). Doratulina (Cymbopogonella) Viraktamath, 1976: 79 (Type: Doratulina longivertex Viraktamath, 1976). Anaconura Emeljanov, 1999: 559 (Type: Stirellus aristidae Emeljanov, 1968). Synonymized by Duan et al., 2019. Coloration and morphology. Yellowish brown to greenish in color. Head variably shaped, slightly to strongly produced. Ocellus present slightly below or on anterior margin. Frontoclypeus straight to slightly convex in profile. Clypeal sulcus prominent. Forewing variably developed, brachypterous to macropterous, usually with three anteapical cells, inner anteapical cell opened at base. Male genitalia. Pygofer triangular in lateral view, dorsal margin sloping posteriorly and mostly straight or concave distally with short tooth present on the apical half near ventral margins. Valve long, triangular. Style with preapical angle distinct. Connective Y-shaped or H-shaped. Aedeagal base triangular, shaft elongate and slender with no basal hinge and strongly curved anterodorsally. Gonopore apical or subapical. Distribution. Worldwide. Remarks. Duan et al. (2019) provided a detailed description of the tribe. Stirellus is heterogeneous and the largest genus of the tribe, distributed worldwide. Variation in external morphological characters including body proportions and coloration explain why many diverse forms were considered previously to represent separate genera. Previously Khatri & Webb (2010) reported 3 known Stirellus species from Pakistan. Recently, Naveed et al. (2020) added 2 additional new species to make a total of 5 known species from Pakistan. Checklist of the species of Stirellus Osborn & Ball from Pakistan Stirellus kumratensis Naveed & Zhang, 2020. Stirellus lahorensis Distant, 1918. Stirellus mankiensis Shah & Duan sp. n. Stirellus neoconvexus Naveed & Zhang, 2020. Stirellus thattaensis Mahmood, Sultana & Waheed, 1972. Stirellus tolla Pruthi, 1930 n. rec. Stirellus viridulus Pruthi, 1930. Key to species of Stirellus Osborn & Ball from Pakistan 1. Crown 1.5x longer than breadth between eyes (Fig. 2A).............................................. S. lahorensis ��� Crown less than 1.5x or equal to breadth between eyes (Figs 1 A���B, 3A���C, 4A, 5A, 6A, 8A, 9A)........................2 2. Species with yellowish green in color (Figs 6 A���C, 8A���C).................................................... 3 ��� Species with ochraceous to brownish in color (Figs 1 A���D, 3A���E, 9A���C).........................................5 3. Crown anterior margin very slightly angulate (Figs 6A, 6C, 8A, 8D)................................... S. tolla n. rec. ��� Crown anterior margin acutely angled (Figs 4A, 5A)......................................................... 4 4. Male pygofer long, with rounded apex (Fig. 5C)................................................... S. viridulus ��� Male pygofer short with pointed apex (Fig. 4E)................................................... S. thattaensis 5. Subgenital plate with macrosetae uniseriate laterally (Fig. 1H)........................................ S. kumratensis ��� Subgenital plate with macrosetae not uniseriate laterally (Fig. 3K, 10B).......................................... 6 6. Connective stem shorter than anterior arms (Fig. 3L), aedeagal shaft with blunt apex (Fig. 3 M���N)........... S. neoconvexus ��� Connective stem longer than anterior arms (Fig. 10C), aedeagal shaft with pointed apex (Fig. 10 D���E).... S. mankiensis sp. n., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species, pp. 189-202 in Zootaxa 4763 (2) on pages 190-191, DOI: 10.11646/zootaxa.4763.2.3, http://zenodo.org/record/3758208, {"references":["Osborn, H. & Ball, E. D. (1902) A review of the North American species of Athysanus (Jassidae). The Ohio Naturalist. A Journal Devoted More Especially to the Natural History of Ohio, 2, 231 - 256.","Van Duzee, E. P. (1892) New North American Homoptera. IV. The Canadian Entomologist, 24, 113 - 117. https: // doi. org / 10.4039 / Ent 24113 - 5","Kirkaldy, G. W. (1907) Leafhoppers supplement (Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 3, 1 - 186.","Melichar, L. (1903) Homopteren-fauna von Ceylon. Verlag von Felix L. Damer, Berlin, 248 pp.","Kirkaldy, G. W. (1906) Leaf-hoppers and their natural enemies. (Pt. IX Leaf-hoppers. Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 1 (9), 271 - 479.","Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Osborn, H. (1930) North American leafhoppers of the Athysanella group (Homoptera: Cicadellidae). Annals of the Entomological Society of America, 23, 687 - 720. https: // doi. org / 10.1093 / aesa / 23.4.687","Gillette, C. P. (1898) A few new species of Deltocephalus and Athysanus from Colorado. Bulletin of the Colorado Agricultural Experiment Station, 43, 23 - 29.","Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Beamer, R. H. & Tuthill, L. D. (1934) Some new species and a new genus of deltocephaloid leafhoppers. (Homoptera: Cicadellidae). Journal of the Kansas Entomological Society, 7, 1 - 24.","Pruthi, H. S. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11 (3), 101 - 131.","Linnavuori, R. (1952) Studies on some Palearctic Hemiptera. Annales Entomologici Fennici, 18, 181 - 187.","Evans, J. W. (1954) Les Cicadellidae de Madagascar (Homopteres). Memoires de l'Institut Scientifique de Madagascar, 4, 87 - 137.","Ishihara, T. (1961) Homoptera of Southeast Asia collected by the Osaka city university biological expedition to Southeast Asia 1957 - 1958. Nature & Life in Southeast Asia, 1, 225 - 257.","Viraktamath, C. A. (1976) New species of Doratulina and Bumizana (Homoptera: Cicadellidae) from Karnataka. Oriental Insects, 10 (1), 79 - 86. https: // doi. org / 10.1080 / 00305316.1976.10432323","Emeljanov, A. F. (1999) A key to genera of the subfamily Deltocephalinae s. l. (Homoptera: Cicadellidae) from Kazakhstan, Middle Asia, and Mongolia with description of new genera and subgenera. Entomological Review, 79 (5), 547 - 562.","Emeljanov, A. F. (1968) Notes on the tribe Stirellini (Homoptera: Cicadellidae). Zoologicheskii Zhurnal, 47 (2), 249 - 253.","Duan, Y. N., Zhang, Y. T. & Dietrich, C. H. (2019) Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records. Zootaxa, 4603 (2), 201 - 241. https: // doi. org / 10.11646 / zootaxa. 4603.2.1","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6","Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84."]}

Stirellus mankiensis Shah & Duan (See Shah et al., 2020: 200���201, Figs 9���10) Stirellus mankiensis Shah & Duan, 2020 in Shah et al., 2020: 200. Measurement. Male: 3.8���4.0mm; female: 5.0mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 11����, 2♀♀, Manki, 34��0���34.5096 N, 73��19���44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. The species can be distinguished from other species of Stirellus by its distinctive external appearance and crown with anterior margin slightly bulbous. This species is somewhat similar to Stirellus neospeciosus Duan, Webb & Zhang in the form of the aedeagus but can easily be separated by the subgenital plate with macrosetae not uniseriate and the connective stem longer than arms; it is also very different from in external appearance., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 402-403, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3"]}

Stirellus kumratensis Naveed & Zhang (See Shah et al., 2020: 192, Fig. 1) Stirellus kumratensis Naveed & Zhang, 2020 in Naveed et al., 2020: 481. Measurement. Male: 3.5–3.6mm; female: 3.9mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 2³³, 1♀, Abbottabad, 34°10′7.5036 N, 73°13′17.3892 E, sweeping hand net, 23 July 2018, coll. Bismillah Shah; 1♀, Barkot, 34°35′58.3944 N, 73°22′41.8764 E, sweep-ing hand net, 11 August 2019, coll. Bismillah Shah; 14³³, 11♀♀, Balakot, 34°34′18.1668 N, 73°21′46.6668 E, sweeping hand net, 12 August 2019, coll. Bismillah Shah; Punjab: 6³³, 3♀♀, Pathriata, 33°52′29.6292 N, 73°27′23.5044 E, sweeping hand net, 29 July 2019, coll. Bismillah Shah; 1³, 7♀♀, Bansra Gali, 33°54′17.532 N, 73°22′13.9656 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; 4♀♀, Bhurban, 33°56′27.3228 N, 73°26′48.7428 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; 4³³, 3♀♀, Sunny Bank, 33°55′1.1244 N, 73°23′39.8256 E, sweeping hand net, 15 August 2019, coll. Bismillah Shah; 5♀♀, Lower Topa, 33°53′51.198 N, 73°25′53.8932 E, sweeping hand net, 16 August 2019, coll. Bismillah Shah; 3♀♀, New Murree, 33°52′36.8148 N, 73°27′2.4912 E, sweeping hand net, 16 August 2019, coll. Bismillah Shah; Azad Kashmir: 1♀, Muzaffarabad, 34°21′44.1144 N, 73°27′10.62 E, sweeping hand net, 14 August 2019, coll. Bismillah Shah. Distribution. Pakistan. Remarks. This species looks similar to Stirellus laetus (Melichar) but can easily be separated by the crown with a pair of orange longitudinal stripes along the mesal margins of eyes, the male pygofer with macrosetae, the style apophysis small and the aedeagus with a spine-like apex., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 402, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6"]}

Stirellus viridulus (Pruthi) (See Shah et al., 2020: 196, Fig. 5) Paternus viridula Pruthi, 1930: 42. Paternus viridulus, Metcalf, 1967: 2350. Stirellus viridulus, Khatri & Webb, 2010: 1���47. Measurement. Male: 3.3���3.7mm; female: 3.9mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 1��, 4♀♀, Pirsabak, 34��0���37.1196 N, 72��3���3.6648 E, light trap, 12 July 2018, coll. Bismillah Shah; 1��, Kund Park, 33��55���50.0304 N, 72��14���5.0064 E, sweeping hand net, 30 July 2018, coll. Bismillah Shah; 2����, 1♀, Khairabad, 33��54���15.1596 N, 72��13���37.8264 E, sweeping hand net, 20 August 2018, coll. Bismillah Shah; 1��, 2♀♀, Faqir Baba, 33��54���53.4888 N, 72��9���56.2176 E, sweeping hand net, 20 July 2019, coll. Bismillah Shah; 2����, Manki, 34��0���34.5096 N, 73��19���44.4432 E, sweeping hand net, 20 August 2019, coll. Bismillah Shah; Punjab: 28����, 12♀♀, Kuldana, 33��55���15.8232 N, 73��23���58.0812 E, sweep-ing hand net, 1 August 2019, coll. Bismillah Shah. Distribution. Pakistan, India. Remarks. This species closely resemble S. lahorensis (Distant) in external appearance and male genitalia but varies in color and in the pygofer lobe. The caudal region of pygofer lobe in S. viridulus (Pruthi) is straight but curved in S. lahorensis (Distant)., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 409, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Metcalf, Z. P. (1967) General catalogue of the Homoptera. Fascicle VI. Cicadelloidea. Part 10. Section III. Euscelidae. U. S. Department of Agriculture, Agriculture Research Service, Washington, D. C., pp. 2075 - 2695.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1"]}

Stirellus tolla (Pruthi) n. rec. (Figs 6���8) Aconura tolla Pruthi, 1930: 39. Measurement. Male: 3.00��� 3.50mm. Coloration and Morphology. General color yellowish green (Fig. 6 A���C). Crown with yellow hues. Eye dark greyish, surrounded with orange band (Figs 6 A���C, 8A, 8C). Pronotum posterior region blackish with minute dots. Mesonotum with basal triangles orange (Fig. 6A, 6C). Scutellum yellowish. Forewing pale with white venation (Fig. 6 A���B). Crown narrower than pronotum, almost as long as distance between eyes, anteriorly slightly angulate (Figs 6A, 6C, 8A, 8D). Eye relatively large. Ocellus next to eye slightly below anterior margin of crown. Face as long as broad, with transverse markings near eyes (Fig. 6D). Pronotum median length nearly as long as median length of crown (Figs 6A, 6C, 8A, 8D). Mesonotum and scutellum shorter than pronotum (Fig. 6A, 6C). Forewing macropterous (Figs 6 A���B, 8A, 8G). Male genitalia. Pygofer elongated, without macrosetae, apex as in Figs 7A, 8H. Subgenital plate slightly concave laterally, apically rounded, macrosetae uniseriate laterally (Fig. 7C). Style narrow, apophysis digitate, curved laterally (Figs 7C, 8K). Connective with stem slightly shorter than arms (Figs 7D, 8L). Aedeagal shaft long, strongly curved basally, almost parallel to aedeagal base, with preapical anterior expansion visible in lateral and posterior views, apex pointed and slightly curved anterad (Figs 7 E���F, 8M). Material examined. 1♂, Pakistan: Khyber Pakhtunkhwa: Khairabad Kund, 33��54���15.1596 N, 72��13���37.8264 E, Sweeping hand net, 20 August 2018, coll. Bismillah Shah. Remarks. This species was not placed in the previous checklist because of unconfirmed identity. Present studies have confirmed its identity and existence in Pakistan. This species differs from most other species of the genus in the shape and size of the crown, which is shorter and bluntly angulate at the anterior margin, by the eyes surrounded with an orange band and the pronotum posterior region blackish with minute dots. This species is somewhat similar to Stirellus daii Duan, Webb & Zhang in genital characters but differs in external appearance. Its male genitalia also show similarity with Stirellus rubrolineatus Distant but differ in subgenital plates., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species, pp. 189-202 in Zootaxa 4763 (2) on pages 196-197, DOI: 10.11646/zootaxa.4763.2.3, http://zenodo.org/record/3758208, {"references":["Pruthi, H. S. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68."]}

Stirellus lahorensis (Distant) (See Shah et al., 2020: 193, Fig. 2) Volusenus lahorensis Distant, 1918: 72. Stirellus peshawarensis Mahmood, Sultana & Waheed, 1972: 80. Paternus jhokensis Ahmed & Aziz, 1988: 805. Stirellus lahorensis, Khatri & Webb, 2010: 1���47; Naveed et al., 2020: 480. Measurement. Male: 3.4���3.6mm; female: 4.1mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 11����, 7♀♀, Buner, 34��23���39.5592 N, 72��36���54.4212 E, light trap, 20 July 2018, coll. Bismillah Shah. Distribution. Pakistan, India, Iran. Remarks. This species resembles S. viridulus (Pruthi) in male genitalia but S. lahorensis (Distant) has the crown more angulate and strongly produced than S. viridulus (Pruthi)., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on page 402, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84.","Ahmed, M. & Aziz, A. (1988) Two new species of grassland leafhoppers (Cicadellidae: Homoptera) in lower Sind, Pakistan. Sarhad Journal of Agriculturae, 4 (6), 805 - 811.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6"]}

Shah, Bismillah, Naveed, Hassan, Duan, Yani (2020): Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa 4763 (2): 189-202, DOI: 10.11646/zootaxa.4763.2.3

Stirellus neoconvexus Naveed & Zhang (See Shah et al., 2020: 194, Fig. 3) Stirellus neoconvexus Naveed & Zhang, 2020 in Naveed et al. 2020: 481. Measurement. Male: 3.5–3.7mm; female: 3.9mm. Material examined. Pakistan: Khyber Pakhtunkhwa: 8³³, 6♀♀, Maidan, 34°57′6.678 N, 71°48′29.5632 E, sweeping hand net, 13 July 2018, coll. Bismillah Shah; 1♀, Abbottabad, 34°10′7.5036 N, 73°13′17.3892 E, sweep-ing hand net, 23 July 2018, coll. Bismillah Shah; 1♀, Ayubia, 34°1′48.5544 N, 73°24′24.4332 E, sweeping hand net, 28 July 2019, coll. Bismillah Shah; 4³³, 1♀, Lora, 33°53′31.8804 N, 73°16′50.7288 E, sweeping hand net, 30 July 2019, coll. Bismillah Shah; Punjab: 1³, 2♀♀, Bansra Gali, 33°54′17.532 N, 73°22′13.9656 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; 1³, Bhurban, 33°56′27.3228 N, 73°26′48.7428 E, sweeping hand net, 1 August 2019, coll. Bismillah Shah; Azad Kashmir: 1♀, Rawalakot, 33°50′34.098 N, 73°47′18.366 E, sweeping hand net, 26 July 2018, coll. Bismillah Shah. Distribution. Pakistan. Remarks. This species is similar to Stirellus convexus Thomson but differs in having the male pygofer side with macrosetae present dorsally, the shape of the style and in having the aedeagal shaft with a slight expansion dorsally. This species is also similar to Stirellus multipunctatus Duan, Webb & Zhang but can readily be distinguished by the medially interrupted orange-red or brownish red stripes on the head, and by the shape of the pygofer and aedeagus., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, A new species and four new records of Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with a revised key to species, pp. 398-410 in Zootaxa 4895 (3) on pages 403-404, DOI: 10.11646/zootaxa.4895.3.5, http://zenodo.org/record/4326829, {"references":["Shah B., Naveed H., Duan Y. N. (2020) Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera: Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species. Zootaxa, 4763 (2), 189 - 202. https: // doi. org / 10.11646 / zootaxa. 4763.2.3","Naveed, H., Shah, B., Sohail, K. & Zhang, Y. L. (2020) Review of the leafhopper genus Stirellus Osborn & Ball, 1902 (Hemiptera: Cicadellidae: Deltocephalinae) with description of two new species from Pakistan. Zootaxa, 4722 (5), 479 - 485. https: // doi. org / 10.11646 / zootaxa. 4722.5.6"]}

Stirellus thattaensis Mahmood, Sultana & Waheed (Fig. 4) Stirellus thattaensis Mahmood, Sultana & Waheed, 1972: 82. Material examined. No specimen examined. Remarks. Mahmood et al., (1972) provided a detailed description of this species. S. thattaensis is quite close to S. lahorensis in external appearance but differs in male genital characters., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2020, Taxonomic review of the leafhopper genus Stirellus Osborn & Ball (Hemiptera Cicadellidae: Deltocephalinae: Stenometopiini) from Pakistan with description of a new species, pp. 189-202 in Zootaxa 4763 (2) on page 195, DOI: 10.11646/zootaxa.4763.2.3, http://zenodo.org/record/3758208, {"references":["Mahmood, S. H., Sultana, S. & Waheed, A. (1972) Two new species of Stirellus Osborn & Ball (Homoptera: Cicadellidae: Deltocephalinae) from West Pakistan. Pakistan Journal of Zoology, 4 (1), 79 - 84."]}

Grammacephalus punjabensis Shah & Duan sp. n. (Figs 11–12) Measurement. Male: 5.5 mm. Coloration and morphology. Yellowish brown (Fig. 11 A–C). Crown dark brown with median longitudinal white stripe reaching tip of anterior margin, also with pair of lateral ochraceous stripes, pair of posterior yellowish spots present on each side of median white stripe. Eyes color tinged with red (Fig. 11A, C). Face whitish yellow with brownish narrow band along anterior margin (Fig. 11D). Pronotum with pale irregular spots, as well as with bright median stripe. Mesonotum and scutellum yellowish brown with pale spots (Fig. 11A, C). Femora and tibiae with dark brown marks (Fig. 11B). Crown narrower than pronotum, subconical, longer than 2x width between eyes (Fig. 11A, C). Anteclypeus with submedian constriction, apex extending beyond normal curve of genae (Fig. 11D). Pronotum shorter than crown. Mesonotum and scutellum nearly as long as pronotum (Fig. 11A, C). Male genitalia. Pygofer lobe broad basally while narrowed towards posterior margin, with posterior short setae covering membranous area, tuft of long microsetae present near posterior margin. Pygofer process long, directed postero-dorsally, surpassing pygofer lobe, with subapical posteroventrally directed spine near apex (Fig. 12A, 12B). Subgenital plate triangular, broad basally, tapered towards apex with with lateral uniseriate macrosetae. Style elongated, with small apophysis (Fig. 12D). Connective Y-shaped with arms well separated, stem short (Fig. 12E, F). Aedeagus broad at base with well-developed dorsal apodeme, gonopore apical, aedeagal shaft slightly narrowed at mid-length, widened near apex in dorsal view, sinuate and evenly tapered distally in lateral view (Fig. 12 E–G). Material examined. Holotype: male, Pakistan: Punjab: Murree Hills, 33°54′23.508 N, 73°23′ 37.284 E, 2000m, Sweeping hand net, 23 July 2018, coll. Bismillah Shah. Etymology. This species is named for Punjab province in Pakistan where the type specimen has been collected. Remarks. This species resembles G. furcatus in external characters but can readily be distinguished by the subgenital plate with lateral margins not incurved at mid-length and the aedeagus without dorsal apical processes.

Grammacephalus harpago Linnavuori (Fig. 4D, E) Grammacephalus harpago Linnavuori, 1978: 476. Material examined. No specimen examined. Remarks. This species also resembles G. pugio in external features and in having a retrorse dorsal spine on the pygofer process, but is much smaller in size and can readily be differentiated by the subapical position of the spine on the male pygofer process., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 74, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Linnavuori, R. (1978) Revision of the Ethiopian Cicadellidae (Homoptera). Paraboloponinae and Deltocephalinae. Scaphytopiini and Goniagnathini. Revue de Zoologie et de Botanique Africaines, 92 (2), 457 - 500."]}

Grammacephalus furcatus Dai & Zhang (Fig. 2) Grammacephalus furcatus Dai & Zhang, 2006 in Dai et al., 2006: 39. Material examined. No specimen examined. Remarks. This species is similar to G. rahmani externally but can be differentiated by the bifurcate pygofer process and aedeagus with a pair of dorsally curved appendages. This species is also similar to G. pallidus in aedagal shape but differs in having the pygofer process branched and the aedeagus with short distal processes., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 74, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Dai, W., Zhang, Y. L., Viraktamath, C. A. & Webb, M. D. (2006) Two new Asian Scaphoideini leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae) with description of a new genus. Zootaxa, 1309 (1), 37 - 44. https: // doi. org / 10.11646 / zootaxa. 1309.1.3"]}

Grammacephalus niveimarginatus (Melichar) (Fig. 4F, G) Platymetopius niveimarginatus Melichar, 1904: 43. Grammacephalus niveimarginatus, Linnavuori, 1978: 477. Material examined. No specimen examined. Remarks. The male of this species is unknown., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 80, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Melichar, L. (1904) Neue Homopteren aus Sud-Schoa, Galla und den Somal-Landern. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- botanischen Gesellschaft in Wien, 54, 25 - 48. https: // doi. org / 10.5962 / bhl. part. 27685","Linnavuori, R. (1978) Revision of the Ethiopian Cicadellidae (Homoptera). Paraboloponinae and Deltocephalinae. Scaphytopiini and Goniagnathini. Revue de Zoologie et de Botanique Africaines, 92 (2), 457 - 500."]}

Grammacephalus indicus Viraktamath & Anantha Murthy (Fig. 5) Grammacephalus indicus Viraktamath & Anantha Murthy, 1999: 42; Khatri & Webb, 2016: 16; Naveed & Zhang, 2018: 1817. Material examined. No specimen examined. Remarks. This species varies in the color pattern of the crown and is close to G. pallidus in external characters but can be easily differentiated by the male genitalia., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 74, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Viraktamath, C. A. & Anantha Murthy, H. V. (1999) A revision of the leafhopper tribe Scaphoideini from India and Nepal (Insecta: Hemiptera: Cicadellidae: Deltocephalinae). Senckenbergiana Biologica, 79, 39 - 55.","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Grammacephalus rahmani (Singh Pruthi) (Fig. 9) Platymetopius rahmani Singh Pruthi, 1930: 33. Grammacephalus rahmani, Mahmood, 1979; Viraktamath, 1981: 7; Khatri & Webb, 2010: 16; Naveed & Zhang, 2018: 1817. Material examined. No specimen examined. Remarks. This species was described based on syntypes from Punjab, Lyallpur, Oct., 1939 (A. Rahman). Later Mahmood (1979) also recorded this species from Pakistan. This species resembles G. pugio but differs in lacking a retrorse spine on the pygofer process., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 82, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Singh Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11, 1 - 68.","Mahmood, S. H. (1979) A Revision of the leafhoppers (Cicadellidae: Homoptera) of Pakistan and adjoining countries of the Oriental region. Final Technical Report PK-ARS- 15, 10, 103.","Viraktamath, C. A. (1981) Indian species of Grammacephalus (Homoptera: Cicadellidae). Colemania, 1 (1), 7 - 12.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Grammacephalus Haupt Grammacephalu s Haupt, 1929: 258. Type species: Platymetopius pugio Noualhier, 1895. Caffretus Evans, 1947: 258. Type species: Caffretus turneri Evans, 1947. Coloration and morphology. Forewing with whitish costal area. Crown acutely triangular in dorsal view. Expanded genae visible dorsally behind eyes. Ocelli present near eyes. Face elongated. Pronotum broader than crown. Forewing without extra cross veins, appendix rounded reaching second apical cell. Male genitalia. Pygofer short and broad, usually with attenuate apical lobe or long ventral process. Anal tube usually extended beyond pygofer posterior margin. Valve semicircular. Subgenital plate triangular. Style apophysis small, digitate, preapical lobe rounded, moderately developed. Connective Y-shaped. Aedeagal shaft gracile, long, gonopore apical or subapical. Female. 7 th sternite comparatively large. Remarks. Linnavuori (1978) reviewed the African species of the genus, including 2 new combinations and 3 new species. Viraktamath (1981) proposed 2 new combinations and described 1 new species from the Indian subcontinent. Later, 1 new species was added to the genus by Viraktamath & Anantha Murthy (1999) from India. Subsequently, Dai et al. (2006) described a new species from China. Khatri & Webb (2010) reported 3 known species whereas Naveed & Zhang (2018) documented a total of 5 known species from Pakistan., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 72, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Haupt, H. (1929) Neueinteilung der Homoptera-Cicadina nach phylogenetisch zu wertenden Merkmalen. Zoologische Jahrbcher, 58, 173 - 286, 86 illustr.","Noualhier, M. (1895) [in Puton, A. & Noualhier, M. 1895: 176] Supplment a la liste des Hemiptres d'Akebs. Review Entomologique, 6, 298 - 311.","Evans, J. W. (1947) A natural classification of leaf-hoppers (Jassoidea: Homoptera). Part 3. Jassidae. Transactions of the Royal Entomological Society of London, 98, 105 - 271. https: // doi. org / 10.1111 / j. 1365 - 2311.1947. tb 01054. x","Linnavuori, R. (1978) Revision of the Ethiopian Cicadellidae (Homoptera). Paraboloponinae and Deltocephalinae. Scaphytopiini and Goniagnathini. Revue de Zoologie et de Botanique Africaines, 92 (2), 457 - 500.","Viraktamath, C. A. (1981) Indian species of Grammacephalus (Homoptera: Cicadellidae). Colemania, 1 (1), 7 - 12.","Viraktamath, C. A. & Anantha Murthy, H. V. (1999) A revision of the leafhopper tribe Scaphoideini from India and Nepal (Insecta: Hemiptera: Cicadellidae: Deltocephalinae). Senckenbergiana Biologica, 79, 39 - 55.","Dai, W., Zhang, Y. L., Viraktamath, C. A. & Webb, M. D. (2006) Two new Asian Scaphoideini leafhoppers (Hemiptera: Cicadellidae: Deltocephalinae) with description of a new genus. Zootaxa, 1309 (1), 37 - 44. https: // doi. org / 10.11646 / zootaxa. 1309.1.3","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Grammacephalus kempi (Singh Pruthi) (Fig. 6) Platymetopius kempi Singh Pruthi, 1934: 28. Grammacephalus kempi, Viraktamath, 1981: 9. Material examined. No specimen examined. Remarks. G. kempi can be readily distinguished from other members of the genus by the longer crown with some partial longitudinal stripes and a larger pale irregular area preapically, the serrate male pygofer processs, the apically upturned subgenital plate, the denticulate style apex, the footlike apex of the aedeagus and the female 7 th sternum with lateral lobes surpassing the 8 th sternite. This species has no close relative in the genus., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 74, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Singh Pruthi, H. (1934) Entomological investigations on the spike disease of sandal (14). Jassidae (Homopt.). Indian Forest Records Entomology Series, 19 (4), 1 - 30.","Viraktamath, C. A. (1981) Indian species of Grammacephalus (Homoptera: Cicadellidae). Colemania, 1 (1), 7 - 12."]}

Grammacephalus raunoi Viraktamath (Fig. 10) Grammacephalus raunoi Viraktamath, 1981: 9; Naveed & Zhang, 2018: 1817. Material examined. No specimen examined. Remarks. Naveed & Zhang (2018) recorded this species from Pakistan. It was previously known only in India. It resembles G. rahmani in coloration but can be differentiated easily by the male genitalia characters., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 82, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Viraktamath, C. A. (1981) Indian species of Grammacephalus (Homoptera: Cicadellidae). Colemania, 1 (1), 7 - 12.","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Grammacephalus pugio (Noualhier) (Fig. 4 A���C) Platymetopius pugio Noualhier, in Puton & Noualhier 1895: 176. Enantiocephalus pugio, Haupt, 1927: 25. Grammacephalus pugio, Haupt, 1929: 258. Caffretus turneri Evans, 1947: 258. Grammacephalus pugio djirofticus Dlabola, 1960: 17. Grammacephalus pugio, Linnavuori, 1978: 476. Material examined. No specimen examined. Remarks. This species can be distinguished from other members of the genus by male genitalia characters. It is possible that G. pugio is a synonym of G. niveimarginatus but the male of the latter species is unknown so far., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 81, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Noualhier, M. (1895) [in Puton, A. & Noualhier, M. 1895: 176] Supplment a la liste des Hemiptres d'Akebs. Review Entomologique, 6, 298 - 311.","Haupt, H. (1927) Homoptera Palestinae I. Bulletin. The Zionist Organization. Institute of Agriculture and Natural History. Agricultural Experiment Station, 8, 5 - 43.","Haupt, H. (1929) Neueinteilung der Homoptera-Cicadina nach phylogenetisch zu wertenden Merkmalen. Zoologische Jahrbcher, 58, 173 - 286, 86 illustr.","Evans, J. W. (1947) A natural classification of leaf-hoppers (Jassoidea: Homoptera). Part 3. Jassidae. Transactions of the Royal Entomological Society of London, 98, 105 - 271. https: // doi. org / 10.1111 / j. 1365 - 2311.1947. tb 01054. x","Dlabola, J. (1960) Iranische Zikaden (Homoptera: Auchenorrhyncha). (Ergebnisse der Entomologischen Reisen Willi Richter, Stuttgart, in Iran 1954 und 1956 - Nr. 31). Stuttgarter Beitraege zur Naturkunde aus dem staatliche Museum fur Naturkunde in Stuttgart, 41, 1 - 24.","Linnavuori, R. (1978) Revision of the Ethiopian Cicadellidae (Homoptera). Paraboloponinae and Deltocephalinae. Scaphytopiini and Goniagnathini. Revue de Zoologie et de Botanique Africaines, 92 (2), 457 - 500."]}

Grammacephalus pallidus Linnavuori (Fig. 8) Grammacephalus pallidus Linnavuori, 1978: 479; Viraktamath, 1981: 8; Khatri & Webb, 2010: 16; Naveed & Zhang, 2018: 1817. Material examined. No specimen examined. Remarks. This species was previously known from Sudan and India. Khatri & Webb (2010) reported it from Pakistan. This species resembles G. minabicus Dlabola and G. genoicus but can be distinguished from them by the male genitalia., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 81, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Linnavuori, R. (1978) Revision of the Ethiopian Cicadellidae (Homoptera). Paraboloponinae and Deltocephalinae. Scaphytopiini and Goniagnathini. Revue de Zoologie et de Botanique Africaines, 92 (2), 457 - 500.","Viraktamath, C. A. (1981) Indian species of Grammacephalus (Homoptera: Cicadellidae). Colemania, 1 (1), 7 - 12.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Grammacephalus genoicus Dlabola (Fig. 3) Grammacephalus genoicus Dlabola, 1984: 52; Khatri & Webb, 2010: 16; Naveed & Zhang, 2018: 1817. Material examined. No specimen examined. Remarks. Khatri & Webb (2010) reported this species, originally described from Iran, from Pakistan. Males differ from others of the genus in lacking a pygofer process and having the anal tube relatively short., Published as part of Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, pp. 71-85 in Zootaxa 4688 (1) on page 74, DOI: 10.11646/zootaxa.4688.1.3, http://zenodo.org/record/3514828, {"references":["Dlabola, J. (1984) Typen revision einiger mediterraner bzw. Nordafrikanischer Hysteropterum (s. l.) (Auchenorrhyncha-Issidae). Acta Faunistica Entomologica Musei Nationalis Pragae, 17 (195), 27 - 67.","Khatri, I. & Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1","Naveed, H. & Zhang, Y. L. (2018) Taxonomy of the leafhopper genus Grammacephalus (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan with description of a newly recorded species. Journal of Entomology and Zoology Studies, 6 (2), 1816 - 1818."]}

Stirellus laetus (Melichar, 1903) n. rec. (Figs 15–16) Deltocephalus laetus Melichar, 1903: 202. Stirellus laetus, Webb & Viraktamath, 2009: 30. Description. Length. Male: 2.8mm. Coloration. General color usually stramineous with orange-red and dark brown markings (Fig. 15A, B). Crown with transverse preapical orange band, pronotum with anterior margin orange, mesonotum with basal triangle orange (Fig. 15A). Face with two black round markings on each side just below antennal sockets; frontoclypeus ochraceous with fuscous arcs (Fig. 15C). Forewing translucent without fuscous tint (Fig. 15A, B). Femora and tibiae with dark brown marks. Abdominal segments variably infused with dark brown (Fig. 15B). Morphology. Head nearly as long as pronotum, subconical, anterior margin rounded to face, angled in dorsal and lateral view (Fig. 15A, B). Crown nearly as long as width between eyes (Fig. 15A). Anteclypeus parallel-sided, extending beyond normal curve of genae (Fig. 15C). Pronotum median length nearly as long as median length of crown. Mesonotum and scutellum together longer than pronotum (Fig. 15A). Forewing submacropterous, usually exposing apex of abdomen (Fig. 15A, B). Male genitalia. Pygofer side with few macrosetae dorsally, steeply declivous posteriorly, apex narrow, posterior margin bluntly rounded (Fig. 16A). Subgenital plate slightly convex laterally, macrosetae uniseriate, well separated from lateral margin, not extended to apex (Fig. 16C). Style apophysis relatively long, digitate, nearlynearly straight, preapical lobe forming right angle (Fig. 16D). ConnectiveConnective H-shaped, stemstem broadbroad andand nearlynearly asas longlong asas armsarms (Fig. 16E). Aedeagal shaft strongly bent dorsad near base, long, sinuate, parallel to aedeagal base in lateral view, apex hooked and blunt; in posterior view abruptly constricted near apex, apex pointed (Fig. 16F, G). Female. Hind margin of seventh sternum slightly concave. Material examined. 1Ƌ, Thailand Loei Phu Ruea NP Pah Lo Noy, 17° 33′ 502" N 101° 20′ 868" E, 1343m, Pan trap, 5–6 ix 2006, Nukoonchai Jaroenchai, leg T817 (INHS). Distribution. India; Sri Lanka; Thailand. Remarks. This species is here recorded from Thailand for the first time., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 220, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Melichar, L. (1903) Homopteren-fauna von Ceylon. VerlagVerlag vonvon FelixFelix L. Damer, Berlin, 248 pp.","Webb, M. D. & Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163, 1 - 64."]}

Stirellus Osborn & Ball, 1902 Athysanus (Stirellus) Osborn & Ball, 1902: 250 (Type: Athysanus bicolor Van Duzee, 1892). Stirellus, Kirkaldy, 1907: 58. Doratulina Melichar, 1903: 198 (Type: Doratulina jocosa Melichar, 1903). Phrynophyes Kirkaldy, 1906: 237 (Type: Phrynophyes phrynophyes Kirkaldy, 1906). Anemochrea Kirkaldy, 1906: 329 (Type: Anemochrea mitis Kirkaldy, 1906). Anemolua Kirkaldy, 1906: 329 (Type: Anemolua hanuala Kirkaldy, 1906). Giffardia Kirkaldy, 1906: 336 (Type: Giffardia dolichocephala Kirkaldy, 1906). syn. nov. Arya Distant, 1908: 338 (Type: Arya rubrolineatus Distant, 1908). Trebellius Distant, 1918: 52 (Type: Trebellius albifrons Distant, 1918). Campbellinella Distant, 1918: 69 (Type: Campbellinella illustrata Distant, 1918). Viridomarus Distant, 1918: 69 (Type: Viridomarus capitatus Distant, 1918). Bituitus Distant, 1918: 70 (Type: Bituitus projectus Distant, 1918). Paternus Distant, 1918: 71 (Type: Paternus pusana Distant, 1918). Volusenus Distant, 1918: 72 (Type: Volusenus lahorensis Distant, 1918). Nandidrug Distant, 1918: 74 (Type: Nandidrug speciosus Distant, 1918). Allectus Distant, 1918: 75 (Type: Allectus notata Distant, 1918). Galerius Distant, 1918: 78 (Type: Galerius indicatrix Distant, 1918). Paivanana Distant, 1918: 95 (Type: Typhlocyba indrus Distant, 1908). Gillettiella Osborn, 1930: 689 (Type: Deltocephalus labiatus Gillette, 1898). Bella Singh-Pruthi, 1930: 44 (Type: Bella apicalis Singh-Pruthi, 1930). Penestirellus Beamer & Tuthill, 1934: 21 (Type: Penestirellus catalinus Beamer & Tuthill, 1934). Sunda Singh-Pruthi, 1936: 112 (Type: Sunda ribeiroi Singh-Pruthi, 1936). Pseudaconura Linnavuori, 1952: 182 (Type: Pseudaconura luxorensis Linnavuori, 1952). Malagasiella Evans, 1954: 110 (Type: Malagasiella minima Evans, 1954). Umesaona Ishihara, 1961: 246 (Type: Umesaona asiatica Ishihara, 1961). Doratulina (Cymbopogonella) Viraktamath, 1976: 79 (Type: Doratulina longivertex Viraktamath, 1976). Anaconura Emeljanov, 1999: 559 (Type: Stirellus aristidae Emeljanov, 1968). syn. nov. Diagnosis. Head variably produced. Ocelli on or slightly below anterior margin. Frontoclypeus straight to moderately convex in profile, clypeal sulcus prominent. Forewing variably developed, usually with three anteapical cells, inner anteapical cell open basally. Pygofer side with dorsal margin usually straight or concave distally, short tooth usually present on apical half near ventral margin. Valve long, triangular. Style preapical angle distinct. Connective Y-shaped or H-shaped. Aedeagus base triangular, shaft without basal hinge, usually elongate and slender, strongly curved anterodorsally. Distribution. Worldwide. Remarks. The composition and morphological definition of this genus remain controversial. It is presently defined based on the absence of the distinctive characters used to define other genera of the tribe. Our current concept of Stirellus includes 25 other named genus-group taxa, here considered junior synonyms. As noted by Emeljanov (1962), Stirellus is a large and heterogeneous assemblage of species distributed worldwide. Although there is considerable interspecific (and sometimes intraspecific) variation in body proportions, wing development and coloration, the male genitalia vary far less obviously among species of Stirellus than in other leafhopper genera. The differences among species in body proportions and coloration, partly explain why many such diverse forms were previously considered to represent separate genera. Most of these taxa have been recognized by previous authors as synonyms of either Stirellus or Doratulina (Linnavuori 1959, Emeljanov 1962, Vilbaste 1965). We follow Emeljanov (1966) and Duan et al. (2016) in treating Doratulina as a junior synonym of Stirellus. Comprehensive phylo- genetic analyses including molecular data will be needed to elucidate the status of Stirellus, the taxa here treated as synonyms, and relationships among the many included species., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 205, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Osborn, H. & Ball, E. D. (1902) A review of the North American species of Athysanus (Jassidae). The Ohio Naturalist. A Journal Devoted More Especially to the Natural History of Ohio, 2, 231 - 256.","Van Duzee, E. P. (1892) New North American Homoptera. IV. The Canadian Entomologist, 24, 113 - 117. https: // doi. org / 10.4039 / Ent 24113 - 5","Kirkaldy, G. W. (1907) Leafhoppers supplement (Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 3, 1 - 186.","Melichar, L. (1903) Homopteren-fauna von Ceylon. VerlagVerlag vonvon FelixFelix L. Damer, Berlin, 248 pp.","Kirkaldy, G. W. (1906) Leaf-hoppers and their natural enemies. (Pt. IX Leaf-hoppers. Hemiptera). Report of Work of the Experiment Station of the Hawaiian Sugar Planters' Association. Division of Entomology Bulletin, 1 (9), 271 - 479.","Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Distant, W. L. (1918) Rhynchota-Homoptera. In: Shipley, A. E. & Marshall, A. K. (Eds.), The Fauna of British India, including Ceylon and Burma, 7, pp. 1 - 210.","Osborn, H. (1930) North American leafhoppers of the Athysanella group (Homoptera: Cicadellidae). Annals of the Entomological Society of America, 23, 687 - 720. https: // doi. org / 10.1093 / aesa / 23.4.687","Gillette, C. P. (1898) A few new species of Deltocephalus and Athysanus from Colorado. Bulletin of the Colorado Agricultural Experiment Station, 43, 23 - 29.","Singh-Pruthi, H. (1930) Studies on Indian Jassidae (Homoptera). Part I. Introductory and description of some new genera and species. Memoirs of the Indian Museum, 11 (1), 1 - 68.","Beamer, R. H. & Tuthill, L. D. (1934) Some new species and a new genus of deltocephaloid leafhoppers. (Homoptera: Cicadellidae). Journal of the Kansas Entomological Society, 7, 1 - 24.","Singh-Pruthi, H. (1936) Studies on Indian Jassidae (Homoptera). Part III. Descriptions of some new genera and species, with first records of some known species from India. Memoirs of the Indian Museum, 11 (3), 101 - 131.","Linnavuori, R. (1952) Studies on some Palearctic Hemiptera. Annales Entomologici Fennici, 18, 181 - 187.","Evans, J. W. (1954) Les Cicadellidae de Madagascar (Homopteres). Memoires de l'Institut Scientifique de Madagascar, 4, 87 - 137.","Ishihara, T. (1961) Homoptera of Southeast Asia collected by the Osaka city university biological expedition to Southeast Asia 1957 - 1958. Nature & Life in Southeast Asia, 1, 225 - 257.","Viraktamath, C. A. (1976) New species of Doratulina and Bumizana (Homoptera: Cicadellidae) from Karnataka. Oriental Insects, 10 (1), 79 - 86. https: // doi. org / 10.1080 / 00305316.1976.10432323","Emeljanov, A. F. (1999) A key to genera of the subfamily Deltocephalinae s. l. (Homoptera: Cicadellidae) from Kazakhstan, Middle Asia, and Mongolia with description of new genera and subgenera. Entomological Review, 79 (5), 547 - 562.","Emeljanov, A. F. (1968) Notes on the tribe Stirellini (Homoptera: Cicadellidae). Zoologicheskii Zhurnal, 47 (2), 249 - 253.","Emeljanov, A. F. (1962) New tribes of leafhoppers of the subfamily Euscelinae (Auchenorrhyncha: Cicadellidae). Entomologicheskoe Obozrenie, 41, 388 - 397.","Linnavuori, R. (1959) Revision of the Neotropical Deltocephalinae and some related subfamilies (Homoptera). Annales Zoologici Societatis Zoologicae Botanicae Fennicae ' Vanamo', 20 (1), 1 - 370.","Emeljanov, A. F. (1966) On the tribe Stirellini trib. n. and its taxonomic position (Homoptera: Cicadellidae). Zoologicheskii Zhurnal, 45 (4), 609 - 610.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus rubrolineatus (Distant, 1908) n. rec. (Fig. 20) Length. Male: 4.8–5.1mm; female: 5.9mm. Material examined. 1♀, Thailand Chaiyaphum Pa Hin Ngam NP Savana (Thepana waterfall), 15° 38′ 948" N 101° 25′ 625" E, 604m, Malaise, 1–7 x 2006, K Sa-nog & B. Adnafai, leg T657 (INHS). Distribution. China; India; Thailand. Remarks. This species is newly recorded from Thailand. Duan et al. (2016) described this species adequately. We supplement their description by including color illustrations. The hind margin of female sternite VII is slightly concave (Fig. 20D)., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 224, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Distant, W. L. (1908) Rhynchota-Homoptera. In: Bingham, C. T. (Ed.), The Fauna of British India, including Ceylon and Burma, 4, pp. 1 - 501.","Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus kitwensis sp. nov. (Figs 27���28) Description. Length. Male: 3.4mm; female: 3.6mm. Coloration. General color stramineous with ochraceous and brown or black markings (Fig. 27A, B, D). Crown with transverse apical and preapical band sometimes joined by black longitudinal stripe along midline, pronotum with white median area partly or completely enclosed by brown or black pigment, mesonotum with basal triangles brown or black and more extensive dark pigment sometimes present (Fig. 27A, D). Face with dorsal, median and ventral transverse black bands variably developed (Fig. 27E). Forewing translucent with symmetrical pattern consisting of large irregular orange areas and numerous small orange spots (Fig. 27A, B, D). Femora and tibiae with dark brown marks (Fig. 27B). Abdominal sternites dark variably infused with white (Fig. 27F). Morphology. Head slightly narrower than pronotum, subconical, anterior margin rounded to face, acutely angled (Fig. 27A, B, D). Vertex nearly as long as 2x width between eyes (Fig. 27A, D). Anteclypeus nearly parallelsided, extending slightly beyond normal curve of genae (Fig. 27E). Pronotum median length slightly shorter than median length of crown. Mesonotum and scutellum together slightly shorter than pronotum (Fig. 27A, D). Forewing macropterous, usually exposing apex of abdomen in female, with supernumerary crossveins (Fig. 27A, B, D). Male genitalia. Pygofer side long, with few macrosetae dorsally, dorsal margin nearly straight posteriorly, posterior margin rounded (Fig. 28A). Subgenital plate slightly convex laterally, macrosetae uniseriate laterally (Fig. 28C). Style apophysis attenuated, laterallylaterally curved, preapicalpreapical lobelobe roundedrounded (Fig. 28D). ConnectiveConnectiveConnective Y-shaped, stemstemstem broadened distally and nearly as long as arms (Fig. 28E). Aedeagal shaft bent strongly dorsad near base, relatively short and sinuate in lateral view with apex attenuated, parallel to aedeagal base, slightly widened preapically then tapered to sharp point in dorsal view, with subapical membranous area (Fig. 28 F���H). Female. Hind margin of seventh sternum shallowly concave (Fig. 27F). Material examined. Holotype: ��, Zambia: Copperbelt Prov. ~ 12km SSE of Kitwe Kumasamba Lodge, 12�� 54��� 325" S 28�� 14��� 373" E, 1151m, 31 x���10 xi 2007, sweep, ZA-01, coll. J.N. Zahniser (INHS). Paratype: 1♀, Zambia: Northwest Prov. ~ 28.5 km N Mwinilunga, 11�� 30��� 16.1" S 24�� 22��� 44.8" E, Chitunta plain grassland, 1384m, 7 xi 2007, sweep, ZA-03, coll. J.N. Zahniser (INHS). Etymology. The species epithet is based on the holotype locality. Distribution. Zambia. Remarks. This species differs from other species of Stirellus in the unique color pattern (Fig. 27A, D); forewing with supernumerary crossveins (Fig. 27A, B, D), evenly rounded male pygofer apex, and aedeagal shaft with a subapical membranous area (Fig. 28 F���H)., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 231, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252

Kinonia Ball, 1933 n. rec. Kinonia Ball, 1933: 224 (Type: Kinonia elongata Ball, 1933, by original designation). Diagnosis. Head acutely produced in dorsal and lateral view. Ocelli on anterior margin of crown. Frontoclypeus straight in profile, clypeal sulcus prominent. Forewing rounded apically, usually not extended to apex of abdomen, branches of R not strongly reflexed, usually with three anteapical cells, inner anteapical cell open basally, appendix absent. Pygofer side with short tooth on apical half near ventral margin. Valve long, triangular. Style preapical angle distinct. Connective H-shaped. Aedeagus with shaft relatively short and broad in lateral and posterior view, hinged at the base. Distribution. Mexico; USA. Remarks. This genus only includes the type species, Kinonia elongata. Emeljanov (1962) treated it as a junior synonym of Stirellus but Oman et al. (1990) retained Kinonia as a valid genus. Although it is similar to Stirellus, as delimited here, in most respects, it differs in having the aedeagus relatively broad and hinged at the base of the shaft, a trait characteristic of most Chiasmini Distant. Thus, further analysis may show that this genus may be more appropriately placed in the latter tribe. Nevertheless, Kinonia agrees with the current definition of Stenometopiini in other respects and phylogenetic analysis of combined morphological and molecular data placed this genus as sister to Stirellus with strong branch support (Zahniser & Dietrich, 2013). Kinonia was originally described from the southwestern USA. It is here recorded from Mexico for the first time., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 202, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Ball, E. D. (1933) Some new genera and species of western leafhoppers. Bulletin of the Brooklyn Entomological Society, 28, 223 - 228.","Emeljanov, A. F. (1962) New tribes of leafhoppers of the subfamily Euscelinae (Auchenorrhyncha: Cicadellidae). Entomologicheskoe Obozrenie, 41, 388 - 397.","Zahniser, J. N. & Dietrich, C. H. (2013) A review of the tribes of Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). European Journal of Taxonomy, 45, 1 - 211. https: // doi. org / 10.5852 / ejt. 2013.45"]}

Stirellus neospeciosus Duan, Webb & Zhang, 2016 n. rec. (Fig. 19) Stirellus neospeciosus Duan, Webb & Zhang, 2016: 121. Length. Male: 3.8���4.0mm; female: 4.1���4.4mm. Material examined. 1♀, Thailand Nan Doi Phu Kha NP Office 14, 19�� 12��� 488" N 101�� 4��� 907" E, 1375m, Malaise, 8���15 xii 2007, Charoen & Nikomi, leg T3278 (INHS). Distribution. China; Thailand. Remarks. This species is newly recorded from Thailand. Duan et al. (2016) have described this species adequately. We supplement their description by including color illustrations. The hind margin of female sternite VII is slightly concave or straight (Fig. 19C)., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on pages 222-224, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Duan, Y. N., Webb, M. D. & Zhang, Y. L. (2016) Review of the leafhopper tribe Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) from China with description of four new species. Zootaxa, 4171 (1), 101 - 138. https: // doi. org / 10.11646 / zootaxa. 4171.1.4"]}

Stirellus petfordensis sp. nov. (Figs 31–32) Description. Length. Male: 4.4mm. Coloration. General color stramineous with dark brown, brown and orange-red marks (Fig. 31A, B). Crown with large apical pair of dark brown spots and smaller pair of median orange-red marks. Pronotum with irregular anterior row of brown spots, mesonotum with basal triangles brown (Fig. 31A). Frontoclypeus brown with sordid white marks, dark brown spot below each eye, gena unmarked (Fig. 31C). Forewing translucent, apex brown (Fig. 31A, B). Femora and tibiae with dark brown marks. Morphology. Head narrower than pronotum, weakly subconical, anterior margin rounded to face, angled (Fig. 31 A–B). Crown ~ 1.3x width between eyes (Fig. 31A). Apex of anteclypeus widened, extending slightly beyond normal curve of genae (Fig. 31C). Pronotum median length nearly as long as median length of crown. Mesonotum and scutellum together longer than pronotum (Fig. 31A). Forewing macropterous (Fig. 31A, B). Male genitalia. Pygofer side without macrosetae, dorsal margin slightly convex, posterior margin rounded (Fig. 32A). Subgenital plate convex laterally, macrosetae uniseriate laterally (Fig. 32C). Style preapical angle indistinct, apophysis robust, digitate, slightlyslightly sinuatesinuate (Fig. 32D).. ConnectiveConnectiveConnective H-shaped H-shaped, stemstemstem slightlyslightlyslightly shortershortershorter thanthanthan armsarmsarms (Fig. 32E). Aedeagal shaft bent strongly dorsad near base, nearly straight through most of length in lateral view, parallel to aedeagal base, apex hooked anterad and attenuate, apex gradually tapered to point in dorsal view, with subapical membranous area (Fig. 32F, G). Material examined. Holotype: Ƌ, Australia, Q[ueensland], 7 km. W. Petford, 24 ii 1997, L.B. O’Brien (ANIC). Etymology. The species epithet is based on the type locality. Distribution. Australia. Remarks. This species is similar to Stirellus indrus (Distant), but it can be distinguished by the following combination of characters: marks on vertex, the proportions of the vertex and pygofer side without macrosetae., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on pages 236-238, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252

Stirellus lesioensis sp. nov. (Figs 23���24) Description. Length. Male: 4.1���4.2mm.. Coloration. General color stramineous with brown or orange-red marks (Fig. 23 A���C). Crown with pair of small apical submedial spots, four spots between anterior eye angles and two spots on posterior margin; pronotum, mesonotum and scutellum symmetrically marked with brown or orange-red (Fig. 23A, B). Frontoclypeus brown with sordid white arcs reaching only part way across (Fig. 23D, E). Forewing translucent with fuscous tint and white veins (Fig. 23 A���C). Femora and tibiae with dark brown marks (Fig. 23C). Abdominal sternites variably infused with dark brown. Morphology. Head nearly subequal in width to pronotum, subconical, anterior margin rounded to face, angled in dorsal and lateral view (Fig. 23 A���C). Crown with anterior margin forming right angle in dorsal view, slightly longer than width between eyes (Fig. 23A, B). Anteclypeus parallel-sided, extending slightly beyond normal curve of genae (Fig. 23D, E). Pronotum median length slightly longer than median length of crown. Mesonotum and scutellum together nearly as long as pronotum (Fig. 23A, B). Forewing macropterous, with few to many supernumerary crossveins (Fig. 23 A���C). Male genitalia. Pygofer side long, with few macrosetae dorsally, dorsal margin slightly concave distally, posterior margin nearly truncate (Fig. 24A). Subgenital plate slightly concave laterally, macrosetae uniseriate laterally (Fig. 24C). Style apophysis digitate, laterallylaterally curved, preapicalpreapical lobelobe poorlypoorly developed, roundedrounded (Fig. 24D). Con-Con-Connective Y-shaped, stem broad and shorter than arms (Fig. 24E). Aedeagal shaft bent dorsad near base, long, very slender, sinuate in lateral view, apex abruptly tapered to point in dorsal view (Fig. 24 F���H). Material examined. Holotype: ��, REP OF CONGO Dept Pool, Abio Lesio-Louna Pk, 03�� 06��� 020" S 15�� 31��� 440" E, 330m, MT, 29 viii 2008, coll. Braet & Sharkey (INHS). Paratype: 1��, same data as holotype (INHS). Etymology. The species epithet is based on the type locality. Distribution. Republic of Congo. Remarks. This species can be distinguished from other species of Stirellus by the distinctive color pattern of the crown (Fig. 23A, B), presence of supernumerary crossveins on the forewing, and the relatively long, apically truncate male pygofer (Fig. 24A, B). The Australian species S. hanuala also has supernumerary crossveins on the forewing but its forewing is truncate apically rather than rounded., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on pages 225-227, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252

Kinonia elongata Ball, 1933 n. rec. (Figs 1���2) Kinonia elongata Ball, 1933: 224. Description. Length. Male: 3.3���3.6mm; female: 4.0��� 4.9mm. Coloration. Generally yellow green throughout without obvious markings other than variable brown infuscation on abdomen (Fig. 1 A���F). Face mostly ochraceous (Fig. 1G). Forewing translucent (Fig. 1 A���F). Femora and tibiae with dark brown marks (Fig. 1F). Abdominal sternites variably infused with dark brown (Fig. 1H). Morphology. Head slightly wider than pronotum, subconical, anterior margin rounded to face (Fig. 1 A���F). Crown longer than 2x width between eyes (Fig. 1 A���E). Anteclypeus tapering to apex, slightly tumid medially, extending beyond normal curve of genae (Fig. 1F). Pronotum median length shorter than median length of crown. Mesonotum and scutellum nearly as long as pronotum (Fig. 1 A���E). Forewing exposing apex of abdomen (Fig. 1 A���F). Male genitalia. Pygofer side with few macrosetae dorsally, dorsal margin convex, posterior margin nearly truncate (Fig. 2A). Subgenital plate ovoid, strongly convex laterally, apex obliquely truncate with weak median apical tooth, macrosetae uniseriate (Fig. 2C). Style apophysis digitate, slim, bent laterad adjacent to obtuse preapical lobe, then straight and tapered distally (Fig. 2D). Connective H-shaped, stem nearly as long as weakly divergent anterior arms (Fig. 2E). Aedeagal shaft robust, evenly curved anterodorsad and slightly tapered in lateral view with attenuate posteroapical projection, broadened from base to distal third, then tapered to apex in posterior view; gonopore large, apical (Fig. 2 F���H). Female. Hind margin of seventh sternum broadly concave medially, angulate laterally (Fig. 1H). Material examined. 2����, Sabino Can[yon] Ar[izona]. 14 vii 1932, coll. R.H. Beamer (INHS); 1��, 6♀♀, Mexico: Durango. Hacienda El Potrero 4.5mi N. Borbollones, 2280m, 25 x 1995, coll. C.H. Dietrich vacuum 95- 0 55 (INHS); 9����, 19♀♀, Mexico: Chihuahua. Rt. 432 29km NE Parral. 0.5km E Rio Chico. 1800m, 30 x 1995, coll. C.H. Dietrich, sweeping, 95-075 (INHS). Distribution. Mexico; USA. Remarks. This species is here recorded from Mexico for the first time., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on pages 202-205, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Ball, E. D. (1933) Some new genera and species of western leafhoppers. Bulletin of the Brooklyn Entomological Society, 28, 223 - 228."]}

Stirellus sagittarius (Naudé, 1926) (Figs 21–22) Aconura sagittaria Naudé, 1926: 54. Stirellus sagittarius, Theron, 1975: 189. Length. Male: 3.0– 3.6mm; female: 4.1–5.1mm. Material examined. 4ƋƋ, 4♀♀, South Africa: KZN Prov. Kwa Dlamini-White Mt. Resort, 29° 08′ 186" S 29° 36′ 408" E, 1532m, 26 xii 2004, #04-66, coll. J.N. Zahniser, sweep (INHS). Distribution. Namibia; South Africa. Remarks. Theron (1975) described this species adequately. We supplement his description by including color illustrations., Published as part of Duan, Yani, Zhang, Yating & Dietrich, Christopher H., 2019, Six new species of Stenometopiini (Hemiptera: Cicadellidae: Deltocephalinae) with redescription of additional species and new distributional records, pp. 201-240 in Zootaxa 4603 (2) on page 225, DOI: 10.11646/zootaxa.4603.2.1, http://zenodo.org/record/2682252, {"references":["Naude, T. J. (1926) Cicadellidae of South Africa. A taxonomic and faunistic study. Entomology Memoirs, Union of South Africa, Department of Agriculture, 1 (4), 1 - 106, pls. I-X.","Theron, J. G. (1975) The Naude species of South African Cicadellidae (Hemiptera). IV. Species assigned to the genera Aconura Lethierry and Deltocephalus Burmeister. Journal of the Entomological Society of Southern Africa, 38 (2), 189 - 206."]}

Stirellus madagascarensis sp. nov. (Figs 29–30) Description. Length. Male: 4.7–4.8mm; female: 4.9–5.1mm. Coloration. General color light brown with darker brown and orange-red marks (Fig. 29 A–E). Crown, with pair of orange-red medial spots variably developed (Fig. 29 A–F). Frontoclypeus uniformly light brown with dorsal pair of orange-red marks (Fig. 29E, F). Forewing translucent infused with dark brown (Fig. 29 A–E). Femora and tibiae usually without dark brown markings (Fig. 29E). Abdominal sternites dark, variably infused with white (Fig. 29G). Morphology. Head narrower than pronotum, subconical, anterior margin rounded to face, angled (Fig. 29 A–E). Vertex nearly as long as 1.5x width between eyes (Fig. 29 A–D). Apex of anteclypeus widened, extending slightly beyond normal curve of genae (Fig. 29F). Pronotum median length longer than median length of crown. Mesonotum and scutellum together nearly as long as pronotum (Fig. 29 A–D). Forewing macropterous, usually exposing apex of abdomen in female (Fig. 29 A–D). Male genitalia. Pygofer side without macrosetae, long, dorsal margin deeply concave, posterior margin rounded (Fig. 30A). Subgenital plate subquadrate, macrosetae arranged in irregular band and well separated from lateral margin (Fig. 30C, D). Style preapical angle indistinct, apophysis robust, digitate, laterallylaterally curvedcurved (Fig. 30E). Con-Con-Connective H-shaped, stem broadened distally and nearly as long as arms (Fig. 30F). Aedeagal shaft broadly curved dorsad in lateral view divergent from aedeagal base, slightly sinuate, apex attenuate, dorsal view slender with apex abruptly tapered to acute point, with subapical membranous area (Fig. 30G, H). Female. Hind margin of seventh sternum shallowly concave (Fig. 29G). Material examined. Holotype: Ƌ, Madagascar: Province Fianarantsoa, Parc National Ranomafana Belle Vue at Talatakely, elev 1020m, 14–23 iv 2002, 21° 15′ 99" S 47° 25′ 21" E, collector R. Harin‘Hala California Acad of Sciences malaise, secondary tropical forest, MA-02-09C-25 (CAS). Paratypes: 3ƋƋ, 3♀♀, the same with holotype; 1Ƌ, 2♀♀, Madagascar: Province d’Antanananarivo, 3km 41°NE Andranomay, 11.5km 147° SS, elev 1300m, 5–13 xii 2000, 18° 28′ 24" S 47° 57′ 36" E, BLF2375, collector Fisher, Griswold et al. Calif. Acad. Sci. Collection (CAS); 1Ƌ, 1♀, Madagascar: Province d’Antsiranana:, parc National Montagne d’Ambre, 12.2km 211° SSW Joffreville, elev 1300m, 2–7 ii 2001, 12° 35′ 47" S 49° 9′ 34" E, BLF2853, collector Fisher, Griswold et al. (CAS, INHS); 2ƋƋ, 1♀, Madagascar: Province Fianarantsoa, Parc National Ranomafana Vohiparara at broken bridge, elev 1110m, 22 xi–8 xii 2001, 21° 13′ 57" S 47° 22′ 19" E, collector R. Harin‘Hala California Acad of Sciences malaise trap in high altitude rainforest, MA-02-09A-05(INHS); 3ƋƋ, 8♀♀, Madagascar: Province Fianarantsoa, Parc National Ranomafana radio tower at forest edge, elev 1130m, 21 xii 2001 – 4 vi 2002, 21° 15′ 05" S 47° 24′ 43" E, collector R. Harin‘Hala California Acad. of Sciences malaise mixed tropical forest (CAS, INHS). Etymology. The species epithet is based on the type locality. Distribution. Madagascar. Remarks. This species can be distinguished from other species of Stirellus by the following combination of characters: general color brown without extensive markings (Fig. 29 A–E); subgenital plate distinctly convex laterally, macrosetae not uniseriate (Fig. 30 C–D); aedeagal shaft with subapical membranous area (Fig. 30G, H). Only one other species of Stenometopiini has been recorded from Madagascar so far: Stirellus minimus (Evans). It differs from S. madagascarensis sp. nov. in its less elongated body, less reflexed costal crossveins, rounded subgenital plate, and strongly acute angle between the aedeagal atrium and shaft. Malagasiella was treated as a junior synonym of Doratulina by Zahniser & Nielson (2012). The latter genus was considered a junior synonym of Stirellus by Duan et al. (2016).