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Three new species of the Clubiona corticalis group in China are described: Clubiona bidactylinasp. nov., C. camelasp. nov., and C. subhuimingsp. nov.

Clubiona flammaforma sp. nov. Figs 1, 2, 7 Type material. Holotype: ♁, CHINA: Hubei Province, Enshi City, Badong county, Yanduhe Town, Songziyuan Village, 31.35279°N, 110.39937°E, 1836 m, 27.IV.2016, W. Liu, T. Tian & C. Zeng leg. (HNU-HB-IV-1607). Paratypes: 1♁, same data as holotype (HNU-HB-IV-1607); 2♀, same place as holotype, 31.35067°N, 110.42625°E, 1340 m, 28.IV.2016, W. Liu et al. leg. (HNU-HB-IV-1608). Etymology. The specific epithet is the combination of the Latin flamma (fire) and formis (form), referring to the flame-shaped RTA, adjective. Diagnosis. Clubiona flammaforma sp. nov. resembles C. rostrata Paik, 1985 (see Zhu & Zhang 2011: fig. 266A–E) in having similar course of both embolus in male and copulatory ducts in female, but can be distinguished by: (1) the RTA trifurcate (Fig 1F) (vs. uniramous, with a ventral protuberance in C. rostrata); (2) the embolic base with a stronger dentiform process extended upwards distally in ventral view (Fig 1D) (vs. with a weak tooth extended horizontally in C. rostrata); (3) the posterior margin of epigyne recurve, without median excavation (Fig 2C) (vs. margine procurve, with oblong excavation in C. rostrata); (4) the proximal portion of copulatory ducts separated from each other and arc-shaped (Fig 2D) (vs. close to each other and straight in C. rostrata). Description. Male (holotype) (Fig. 1A, B). Total length 3.91; carapace 1.98 long, 1.35 wide; abdomen 2.15 long, 1.12 wide. Carapace dark yellow; fovea reddish and longitudinal, cervical groove inconspicuous, radial grooves visible. Eyes: AER slightly recurved, PER slightly precurved in dorsal view. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.12, PLE 0.10, AME–AME 0.04, AME–ALE 0.03, PME–PME 0.17, PME–PLE 0.10, MOQL 0.20, MOQA 0.26, MOQP 0.42. Chelicerae yellowish, with five promarginal teeth and seven retromarginal teeth. Endites and labium yellow. Sternum yellow, almost oval. Legs yellow; tibiae I and II with two pairs of ventral spines, metatarsus I and II with a pair of ventral spines. Leg measurements: I 4.04 (1.13, 1.76, 0.77, 0.45), II 4.22 (1.19, 1.77, 0.81, 0.45), III 3.59 (1.14, 1.25, 0.78, 0.42), IV 5.84 (1.91, 1.88, 1.59, 0.46). Abdomen elongate oval, dorsum yellowish white, with brown pinnate pattern posteriorly and two pairs of muscular depressions; venter yellowish white. Spinneret yellowish white. Palp (Fig. 1C–G). Femur and patella without apophysis. Tibia as long as patella and about 1/3 length of cymbium, RTA longer than tibia, sclerotized and trifurcated, flame-shaped, ventral branch tuberculate with blunt tip, lateral branch strongest with pointed tip, dorsal branch spiniform with pointed tip. Cymbium longer than wide. Tegulum relatively flattened, longer than wide. Conductor groovelike, membranous, located distally on retrolateral side of tegulum. Embolus slender, angled across the distal end of tegulum and extended to about 2/3 length of tegulum; embolic base broad with a stronger dentiform process extended upwards distally. Sperm duct slender and twisted in ventral view. Female (one of HNU-HB-IV-1608) (Fig. 2A–B). Total length 4.82. Carapace 2.09 long, 1.50 wide; abdomen 2.71 long, 1.58 wide. Eye sizes and interdistances: AME 0.13, ALE 0.14, PME 0.14, PLE 0.12, AME–AME 0.08, AME–ALE 0.06, PME–PME 0.22, PME–PLE 0.13, ALE–PLE 0.07, MOQL 0.27, MOQA 0.32, MOQP 0.47. Leg measurements: I 3.76 (1.14, 1.54, 0.70, 0.38), II 4.06 (1.26, 1.66, 0.72, 0.42), III 3.73 (1.14, 1.30, 0.90, 0.39), IV 5.77 (1.88, 1.77, 1.63, 0.49). Darker than male, patterns same as in male. Epigyne (Fig.2C, D).With trapezoid epigynal plate.Copulatory openings united at postmedian potion of epigyne. Copulatory ducts longitudinally arc-shaped and extended upwards, then twisted into oval loops. Spermathecae with two oval chambers respectively, interior chambers smaller than lateral chambers. Fertilization ducts located anteriorly inserted on spermathecal interior chambers. Distribution. Known only from the type locality (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on pages 521-523, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Zhu M. S. & Zhang B. S. (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp."]}

Two new species of the genus Clubiona Latreille, 1804 are described: C. tianpingshan sp. nov. (♂♀) and C. flammaforma sp. nov. (♂♀) from central and south China. The female of C. subcylindrata Wang et al., 2018 is described for the first time. Detailed descriptions, photographs of somatic features and copulatory organs, as well as a distribution map of these three species, are provided.

Clubiona subcylindrata Wang, Chen & Zhang, 2018 Figs 3, 4, 7 Clubiona subcylindrata Wang et al. 2018: 328, figs. 16A–C, 17A–D (♁). Holotype from Fanjing Mountain Reserve, Heiwanhe (27°50.778′N, 108°46.362′E; elev. 533 m), 26.IX.2013, L.Y. Wang, D. Wang & X.K. Jiang leg. (Museum of Hebei University, Baoding, China, examined by Luyu Wang). New Records. 4♁ 5♀, CHINA: Hunan Province, Zhangjiajie City, Tianpingshan Nature Reserve, 29.76101°N, 110.04879°E, 551 m, 22.X.2014, J.H. Gan et al. leg. (HNU-HN-X-1417). Diagnosis. The males of Clubiona s ubcylindrata resemble those of C. cylindrata Liu et al., 2007 (see Liu et al. 2007: figs 13–15) in having a bulged tegulum and a slender embolus, but can be distinguished by: (1) the RTA finger-shaped with abruptly sharpened tip in retrolateral view (Fig. 3E) (vs. spinous in C. cylindrata); (2) the sperm duct invisible in ventral view (Fig. 3D) (vs. visible in C. cylindrata). The females of this species resemble those of C. cylindrata Liu et al., 2007 (see Liu et al. 2007: figs 11, 12) and C. lyriformis Song & Zhu, 1991 (see Song et al. 1991: fig. 4A, B) in having twisted tubular spermathecae, but can be distinguished by: (1) the anterior portion of atrium slightly wider than posterior portion (Fig. 4C) (vs. the anterior portion of atrium about half width of posterior portion in C. cylindrata); (2) the posterior margin of atrium opened (Fig. 4C) (vs. enclosed in C. lyriformis). Description. Male (one of HNU-HN-X-1417) (Fig. 3A, B). Total length 7.34. Carapace 3.19 long, 2.33 wide; abdomen 4.24 long, 2.03 wide. Carapace yellow, fovea reddish and longitudinal, cervical and radial grooves conspicuous. Eyes: AER slightly recurved, PER almost straight in dorsal view. Eye sizes and interdistances: AME 0.16, ALE 0.18, PME 0.14, PLE 0.15, AME–AME 0.12, AME–ALE 0.06, PME–PME 0.28, PME–PLE 0.15, ALE–PLE 0.08. MOQL 0.46, MOQA 0.52, MOQP 0.72. Chelicerae yellow with five promarginal teeth and two retromarginal teeth. Endites and labium yellowish brown. Sternum yellowish, almost oval. Legs yellow; tibiae I and II with two pairs of ventral spines, metatarsus I and II with one pair of ventral spines. Leg measurements: I 8.74 (2.46, 3.54, 1.80, 0.94), II 10.29 (2.87, 4.18, 2.23, 1.01), III 7.55 (2.28, 2.61, 1.96, 0.70), IV 10.65 (2.93, 3.59, 3.26, 0.87). Abdomen elongate oval, dorsum yellow with conspicuous tufts of hairs anteriorly and two pairs of muscular depressions, venter light yellow with four rows of dots medianly. Spinneret yellow. Palp (Figs. 3C–G) Femur and patella without apophysis. Tibia and patella longer than wide, about 1/2 length of cymbium. Tibia with two apophyses; VTA shorter with blunt tip, RTA longer than VTA, finger-shaped with abruptly sharpened tip. Cymbium longer than wide. Tegulum strongly bulged, posterior margin extended to patella. Conductor membranous and leaf-shaped, arising from the top of tegulum. Embolus slender and flagelliform, arising from the middle and prolateral side of tegulum. Sperm duct invisible in ventral view. Female (one of HNU-HN-X-1417) (Figs. 4A, B). Total length 8.51. Carapace 3.67 long, 2.57 wide; abdomen 4.70 long, 2.83 wide. Eye sizes and interdistances: AME 0.17, ALE 0.19, PME 0.15, PLE 0.17, AME–AME 0.16, AME–ALE 0.08, PME–PME 0.40, PME–PLE 0.13, ALE–PLE 0.10. MOQL 0.46, MOQA 0.53, MOQP 0.73. Leg measurements: I 8.20 (2.46, 3.35, 1.52, 0.87), II 8.42 (2.46, 3.39, 1.68, 0.89), III 7.08 (2.13, 2.40, 1.90, 0.65), IV 10.06 (2.92, 3.54, 2.78, 0.82). Colors and patterns same as in male. Epigyne (Fig. 4C, D). Epigynal plate almost rounded. Atrium almost calabash-shaped, longer than wide, and located anteriorly; posterior margin opened. Copulatory openings located at the posterolateral margin of atrium. Copulatory ducts semicircular. Spermathecae slender, tubular. Bursae oval. Fertilization ducts short and curved. Distribution. Guizhou and Hunan Provinces of China (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on page 524, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Wang, L. Y., Chen, H. M., Wu, P. L., Zhang, F. & Zhang, Z. S. (2018) Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae). Zoological Systematics, 43 (4), 317 - 333. https: // doi. org / 10.11865 / zs. 201827","Liu, P., Yan, H. M., Griswold, C. & Ubick, D. (2007) Three new species of the genus Clubiona from China (Araneae: Clubionidae). Zootaxa, 1456, 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1456.1.3","Song, D. X., Zhu, M. S., Gao, S. S. & Guan, J. D. (1991 a) Six species of clubionid spiders (Araneae: Clubionidae) from China. Journal of Xinjiang University, 8, 66 - 72."]}

Clubiona tianpingshan sp. nov. Figs 5–7 Type material. Holotype: ♁, CHINA: Hunan Province, Zhangjiajie City, Tianpingshan Nature Reserve, 29.79004°N, 110.09161°E, 1442 m, 23.X.2014, J.H. Gan, Y.H. Gong, C. Wang & B. Zhou leg. (HNU-HN-X-1418). Paratypes: 1♀, same data as holotype (HNU-HN-X-1418); 2 ♁ 2♀, Tianpingshan Nature Reserve, 29.77190°N, 110.06751°E, 1348 m, 24.X.2014, J.H. Gan et al. leg. (HNU-HN-X-1419). Etymology. The specific epithet comes from the type locality, noun. Diagnosis. The males of Clubiona tianpingshan sp. nov. resemble those of C. huiming Wang et al., 2018 (see Wang et al. 2018: figs 8, 9) in having helical embolus and simple RTA and VTA, but can be distinguished by: (1) the embolic base about 1/3 width of the tegulum in ventral view (Fig. 5D) (vs. about 1/6 width of the tegulum in C. huiming); (2) the conductor arising from the anterior 1/3 of the tegulum and the terminal portion above the top of tegulum in retrolateral view (Fig. 5E) (vs. arising from the median portion of the tegulum and its terminal portion below the top of tegulum in C. huiming). The females of the new species resemble those of C. applanata Liu et al., 2007 (see Liu et al. 2007: figs 1, 2) and C. subapplanata Wang, et al., 2018 (see Wang et al. 2018: figs 14C, D, 15E, F) in having an oval atrium located anteriorly, a pair of large bursae and a pair of spermathecae with two chambers, but can be distinguished from the two species by (1) the margin of atrium thinner and posterior margin W-shaped in ventral view (Fig. 6C) (vs. thicker and rounded posteriorly in C. applanate and C. subapplanata); (2) the copulatory duct longer and twisted into two circular loops in dorsal view (Fig. 6D) (vs. shorter in C. applanate and C. subapplanata); (3) the posterior tubular chambers of spermathecae extended vertically in dorsal view (Fig. 6D) (vs. extended transversely in C. applanate and C. subapplanata). Description. Male (holotype) (Fig. 5A, B). Total length 3.85; carapace 1.80 long, 1.24 wide; abdomen 2.07 long, 1.29 wide. Carapace yellow; fovea reddish and longitudinal, cervical and radial grooves inconspicuous. Eyes: AER and PER slightly recurved in dorsal view. Eye sizes and interdistances: AME 0.08, ALE 0.11, PME 0.09, PLE 0.10, AME–AME 0.04, AME–ALE 0.03, PME–PME 0.11, PME–PLE 0.06, ALE–PLE 0.04, MOQL 0.27, MOQA 0.22, MOQP 0.37. Chelicerae yellow, each margin with five teeth. Endites and Labium yellowish brown. Sternum yellowish, almost oval. Legs yellowish; tibiae I and II with three pairs of ventral spines, metatarsus I and II with two pairs of ventral spines. Leg measurements: I 4.20 (1.28, 1.65, 0.75, 0.52), II 4.30 (1.34, 1.60, 0.89, 0.47), III 3.84 (1.17, 1.32, 0.93, 0.41), IV 5.46 (1.68, 1.84, 1.44, 0.50). Abdomen oval, dorsum light yellow, with conspicuous tufts of hairs anteriorly, and two pairs of muscular depressions, venter light yellow. Spinneret yellow. Palp (Fig. 5C–F). Femur without apophysis. Patella wider than long in dorsal view, with a thumb-shaped retroapical apophysis. Tibia shorter and narrower than patella, about 1/3 length of cymbium, with three apophyses; RTA sclerotized and flat, almost triangular with pointed tip in retrolateral view; VTA smaller than RTA, triangular with blunt tip; LTA smallest and with blunt tip. Cymbium longer than wide. Tegulum enlarged and protruded, slightly longer than wide. Conductor beak-shaped with sharp apices, located retrolaterally and arising distally from the 1/3 portion of tegulum in retrolateral view. Embolus arising from the top of tegulum, helical and extended anticlockwise, basal portion about 1/3 wide of tegulum, distal portion filiform. Sperm duct U-shaped in ventral view. Female (HNU-HN-1418) (Fig. 6A, B). Total length 4.23; carapace 1.88 long, 1.37 wide; abdomen 2.37 long, 1.40 wide. Eye sizes and interdistances: AME 0.08, ALE 0.12, PME 0.11, PLE 0.11, AME–AME 0.06, AME–ALE 0.03, PME–PME 0.19, PME–PLE 0.10, ALE–PLE 0.08, MOQL 0.31, MOQA 0.23, MOQP 0.40. Leg measurements: I 3.96 (1.21, 1.58, 0.75, 0.42), II 4.07 (1.29, 1.67, 0.73, 0.38), III 3.65 (1.06, 1.33, 0.89, 0.37), IV 5.05 (1.52, 1.70, 1.36, 0.47). Colors and patterns same as in male. Epigyne (Fig. 6C, D). Epigynal plate almost rounded. Atrium almost oval, wider than long, about 3/5 width of epigyne, located anteriorly, posterior margin thickened and W-shaped. Copulatory openings located in the posterior margin of atrium. Copulatory ducts twisted into two circular loops. Spermathecae meso-laterally located with two chambers respectively; anterior chamber globular, posterior chamber tubular and extended vertically. Bursae almost globular, located posteriorly. Fertilization ducts short and curved. Distribution. Known only from the type locality (Fig. 7)., Published as part of Li, Li-Fen, Liu, Ping, Li, Bing & Peng, Xian-Jin, 2023, Two new species of the sac-spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) and the female of C. subcylindrata from China, pp. 520-530 in Zootaxa 5263 (4) on pages 524-528, DOI: 10.11646/zootaxa.5263.4.3, http://zenodo.org/record/7835794, {"references":["Wang, L. Y., Chen, H. M., Wu, P. L., Zhang, F. & Zhang, Z. S. (2018) Spider diversity in Fanjing Mountain Nature Reserve, Guizhou, China, II: Clubionidae (Araneae). Zoological Systematics, 43 (4), 317 - 333. https: // doi. org / 10.11865 / zs. 201827","Liu, P., Yan, H. M., Griswold, C. & Ubick, D. (2007) Three new species of the genus Clubiona from China (Araneae: Clubionidae). Zootaxa, 1456, 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1456.1.3"]}

Carteronius myene Bonaldo & Labarque sp. nov. Figs 13A, B; 14A, B; Map 1 Type. ♀ holotype from Parc National de Moukalaba-Doudou, Département de Ndougou, Province de Ogooué Maritime (02°35’S, 10°14’E), GABON, III–IV.2003, O.S.G. Olivier & M. Burger leg. (forêt marecaguese) (MRAC 220.827). Etymology. The specific name is a noun in apposition referring to the Myene people, which settled fishing communities along the coast of Gabon. Diagnosis. Females of Carteronius myene sp. nov. resemble those of C. gentilis comb. nov. by the slightly curved epigynal transversal ridge (Figs 14A, 19C), but can be recognized by the posterior sector relatively larger; spermathecae barely visible by transparency in anterior sector (Fig. 14A). Description. Male. unknown. Female. (MRAC 220.827). Measurements: Total length 11.29, CL 5.29, CW 4.85, AL 6.33, AW 4.59, SL 2.64, SW 2.30. Eye diameters and interdistances: AME 0.42, ALE 0.34, PLE 0.29, PME 0.32, AME-AME 0.32, AME-ALE 0.37, ALE-ALE 2.32, PME-PME 0.39, PME-PLE 0.62, PLE-PLE 2.71. Length of leg segments: I 4.68+2.28+4.48+3.46+1.34=16.24; II 4.96+2.29+4.27+3.58+1.41=16.58; III 3.59+1.71+2.83+2. 81+1.00=11.29; IV 4.03+1.69+3.43+3.44+1.15=13.74. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, proximal tooth largest; Leg spination: femora: II do 0-1-0, III do 0-1-0 ve 0-1-0, IV do 1-0-0; tibiae: I ve 2-2-2-2-2-2, II ve 2-2-2-2-2, III ve 2-2, IV ve 1-0-1; metatarsi: I ve 2-2-2-2, II ve 1 p -1 p -0-1 r -2, III pl 0-0-0-1 rl 0-0-0-1 ve 2-2-1, IV pl 2-0-1-0 ve 0-1-0 -0. Coloration: Carapace and chelicerae dark reddish-brown. Endites, labium and sternum dark reddish-brown. Legs reddish-brown, with femora I and II darker. Abdomen dark gray, with several small white spots, two pairs of small white spots in middle dorsally, and three faint chevrons posteriorly. Venter gray, with irregular white spots forming two transversal lines (Fig. 13A). Epigynum: CDv long, gently arched, ST2 tapering, anteriorly located, gland ducts present, approximately same size as ST1; CDd folded ventrally (Fig. 14B). Other material examined. None. Distribution. Only known from Gabon (Map 1).

Carteronius simoni Bonaldo & Shimano sp. nov. Fig. 17; Map. 1 Holotype. ♀ from GABON: leg. Mocquery (No further data) (MNHN-17.436). Etymology. The specific name is a patronym honoring French arachnologist Eugène Simon, who described Carteronius. Diagnosis. Females of C. simoni sp. nov. are similar to those of C. lumumba sp. nov. by the inconspicuous lateral plates of the posterior epigynal sector (Figs 16D, 17C) but can be recognized by the wider, divergent copulatory ducts (Fig. 17C). Description. Male. unknown. Female. (MNHN-17.436) Measurements: Total length 9.15, CL 4.14, CW 3.55, AL 4.99, AW 4.03, SL 1.68, SW 1.82. Eye diameters and interdistances:AME 0.35, ALE 0.24, PLE 0.25, PME 0.30, AME-AME 0.30, AME-ALE 0.40, ALE-ALE 2.15, PME-PME 0.56, PME-PLE 0.61, PLE-PLE 2.60. Length of leg segments: I 2.80+1.47+2.58+2.10+1.09=10.04; II 2.95+1.40+2.48+2.12+0.96=9.91; III 2.35+1.09+1.77+1.76+0.78 =7.75; IV 2.80+1.14+2.43+2.27+0.87=9.51. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two subequal teeth. Leg spination: tibiae: I ve 2-2-2-2-2, II ve 2-2-2-2, IV ve 0-0-1 p; metatarsi: I ve 2-2, II ve 2-2, III ve 2-1 p -2, IV pl 0-0-0-1 rl 0-1-0-1 ve 1 p -1 r - 1 p. Coloration: carapace, chelicerae, endites, labium and sternum reddish. All leg segments yellowish-red. Abdomen pale gray dorsally, with indistinct white spots (Fig. 17A). Epigynum: CDv long, straight, ST2 a small globe, anteriorly located, gland ducts inconspicuous, smaller than ST1; CDd large, S-shaped (Fig. 17D). Other material examined. None. Distribution. Only known from Gabon (Map 1)., Published as part of Eb. Bonaldo, Ramírez, Martín J., Om. Labarque, Shimano, Yulie, Silva-Junior, Cláudio J. & Haddad, Charles R., 2022, Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group, pp. 343-373 in Zootaxa 5205 (4) on page 363, DOI: 10.11646/zootaxa.5205.4.3, http://zenodo.org/record/7307035

Carteronius arboreus Bonaldo & Haddad sp. nov. Figs 7, 8; Map 1 Types. ♂ holotype from Bas Congo, Mayombe, Luki Forest Reserve (05°37’S, 13°05’E), 28.IX.2007, DEMOCRATIC REPUBLIC OF THE CONGO, D. de Bakker & J. P. Michiels leg. (sieving along trail in primary rainforest) (MRAC 223.475). ♀ paratype, same locality and collectors, 10.XI.2006 (Fog 3, primary rainforest) (MRAC 220.925). Etymology. The specific name is a Latin adjective meaning arboreal, in reference to the fact that most of the known specimens were collected by canopy fogging. Diagnosis. Males of Carteronius arboreus sp. nov. are similar to those of C. ashanti sp. nov. by the presence of a sub-apical embolar process (Figs 8A, 10A), but differ by the dorsal lobe of the RTA, which is broad and retrolaterally oriented, and the rounded medial lobe, which is small in relation to the dorsal lobe, and share the same base (Fig. 8B). In C. ashanti sp. nov., the dorsal lobe is spoon-shaped and the medial lobe longer and fang-shaped, with its own base. Females resemble those of C. sudanus comb. nov. in the strongly recurved epigynal transversal ridge (Figs 4C, 8C), but differ by the lateral plates of the epigynal posterior sector being strongly sclerotized (Fig. 8C). Description. Male. (MRAC 223475). Measurements: Total length 6.03, CL 3.16, CW 2.71, AL 2.95, AW 2.24, SL 1.39, SW 1.49. Eye diameters and interdistances: AME 0.28, ALE 0.17, PLE 0.18, PME 0.16, AME-AME 0.65, AME-ALE 0.16, ALE-ALE 1.34, PME-PME 0.33, PME-PLE 0.37, PLE-PLE 1.57. Length of leg segments: I 3.08+1.17+2.67+2.29+1.22=10.43; II 3.10+1.24+3.10+2.24+1.27=10.95; III 2.07+0.93+1.61+1.65+0.96=7.22; IV 2.55+0.94+2.11+2.25+1.05=8.90. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: I do 0-1-0 pl 0-0-1-0, II do 0-1-0, III do 0-1-0, IV do 0-1-0; tibiae: I ve 2-2-2-2-2, II ve 1 p -1 r -1 p -2-2-2, III ve 1 r -1 p -2, IV rl 0-1-0-1 ve 1 p -0-1 p; metatarsi: I ve 2-2, II ve 2-2, III pl 0-1-0-1 rl 0-1-0-1 ve 2-2, IV pl 0-1-0-1 rl 0-1-0-1 ve 1 r -1 p -1 r -0. Coloration: carapace and chelicerae reddish-brown. Endites, labium and sternum reddish-brown. Legs I and II: coxae and trochanters reddish-brown; femora reddish-brown, yellowish distally; tibiae, metatarsi and tarsi yellowish. Legs III and IV yellowish. Abdomen dark gray dorsally, with two well defined white bands; white ventrally (Fig. 7A). Palp: RTA with apical spur short, curved and pointed, dorsal lobe with apical edges bent ventrally, ventral lobe rounded and excavated. Tegulum with short retrolateral apical tegular process, spermatic duct with long loop (Fig. 8A, B). Female. (MRAC 220925). Measurements: Total length 10.58, CL 4.10, CW 3.86, AL 6.37, AW 4.91, SL 1.97, SW 1.93. Eye diameters and interdistances: AME 0.31, ALE 0.21, PLE 0.21, PME 0.18, AME-AME 0.25, AME-ALE 0.29, ALE-ALE 1.75, PME-PME 0.45, PME-PLE 0.49, PLE-PLE 2.13. Length of leg segments: I 3.87+1.81+3.70+2.88+1.28=13.54; II 3.70+1.73+3.66+2.99+1.31=13.39; III 2.75+1.32+2.17+2.08+0.91=9.23; IV 3.25+1.31+2.76+2.84+0.98=11.14. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: I do 0-1-0 pl 0-1-0, II do 0-1-0, III do 0-1-0-1 p, IV do 0-1-0-1 p; tibiae: I ve 2-2-2-2-2-2-2, II ve 2-2-2-1 r -1 p -1 r -1 p -1 r, III ve 1 p -1 r -2, IV rl 0-1-0-1 ve 1p-0-1 p; metatarsi: I ve 2-0-2-0, II ve 2-2-2, III pl 0-1-0-1 rl 0-0-0-1 ve 2-2-1, IV pl 0-1-0-1 rl 1-0-1 ve 1 p -1 r -1 p - 0-0. Coloration: Carapace and chelicerae dark reddish-brown. Endites, labium and sternum reddish-brown. Legs reddish-brown, with femora I and II darker. Abdomen gray, dorsum with scattered small white spots, denser in middle, forming irregular white longitudinal band; posteriorly with large triangular white spot. Ventrally gray with two irregular lateroventral white bands (Fig. 7C). Epigynum: CDv slight folded posteriorly, ST2 globose, anteriorly located, larger than ST1, CDd almost straight (Fig. 8D). Other material examined. DEMOCRATIC REPUBLIC OF THE CONGO: Bas Congo, Mayombe, Luki Forest Reserve (05°37’S, 13°05’E), 18.IX.2007, D. de Bakker & J.P. Michiels leg. (Fog 5, old secondary forest), 1♀ (MRAC); same data but 22.IX.2007, 1♀ (MRAC); same data but 30.IX.2007, 1♂ (MRAC). Distribution. Only known from the Democratic Republic of the Congo (Map 1).

Eb.Bonaldo, Ramírez, Martín J., Om.Labarque, Shimano, Yulie, Silva-Junior, Cláudio J., Haddad, Charles R. (2022): Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group. Zootaxa 5205 (4): 343-373, DOI: https://doi.org/10.11646/zootaxa.5205.4.3

Carteronius teke Bonaldo & Bosselaers sp. nov. Figs 13C, D, 14C, D; Map 1 Type. ♀ holotype from Kivu, Rwankwi [01°19’S, 29°22’E], DEMOCRATIC REPUBLIC OF THE CONGO, VI.1946, J. Leroy leg. (MRAC 168.643). Etymology. The specific name is a noun in apposition referring to the Teke people, one of the three kingdoms that ruled Congo before the arrival of Europeans. Diagnosis. Females of Carteronius teke sp. nov. differ from all other species of Carteronius by the presence of completely straight epigynal transversal ridge (Fig. 14C). Description. Male. Unknown. Female. (MRAC 168.643) Measurements:Total length 13.45, CL 4.90, CW 4.39, AL 7.99, AW 5.24, SL 2.44, SW 2.15. Eye diameters and interdistances:AME 0.38, ALE 0.25, PLE 0.28, PME 0.25, AME-AME 0.36, AME-ALE 0.40, ALE-ALE 2.30, PME-PME 0.57, PME-PLE 0.67, PLE-PLE 2.81. Length of leg segments (sequence from femur to tarsus, and total): I 4.21+2.04+3.57+2.92+1.25=13.99; II 4.15+1.86+3.89+2.79 +1.19=13.88; III 2.83+1.43+2.52+2.20+1.04=10.02; IV 3.42+1.42+3.03+3.14+1.28=12.29. Chelicerae: promargin with three spaced teeth, median tooth largest; retromargin with two spaced teeth, subequal in size. Leg spination: femora: IV do 1-0-0; tibiae: I ve 1 p -2-2-2-2-2, II ve 0-0-2-2-2-2, III ve 0-0-1 p -0, IV ve 1 p -0-1 p -0; metatarsi: I ve 2-2-2-2, II ve 1 r -1 p -2-2, III ve 0-2-2-0, IV ve 0-2-2-0. Coloration: carapace and chelicerae dark reddish-brown. Endites, labium and sternum reddish-brown. Legs I and II reddish-brown, III and IV dark yellow. Abdomen dorsally pale gray, ventrally white with darker band converging at spinnerets (Fig. 13C). Epigynum: CDv long, arched, ST2 tapering, medially located, gland ducts inconspicuous, much smaller than ST1; CDd folded ventrally (Fig. 14D). Other material examined. None. Distribution. Only known from the Democratic Republic of the Congo (Map 1)., Published as part of Eb. Bonaldo, Ramírez, Martín J., Om. Labarque, Shimano, Yulie, Silva-Junior, Cláudio J. & Haddad, Charles R., 2022, Switching identities: a revision of the Afrotropical spider genus Carteronius Simon 1897 (Araneae, Corinnidae), senior synonym of Mandaneta Strand, 1932, with a new genus of the Pronophaea group, pp. 343-373 in Zootaxa 5205 (4) on pages 359-361, DOI: 10.11646/zootaxa.5205.4.3, http://zenodo.org/record/7307035

A new genus of Clubionidae Wagner, 1887, Femorbiona Yu & Li, gen. nov., is described, with Clubiona brachyptera Zhu & Chen, 2012 (♂♀; Hainan, China) as the type species. Three species are included in Femorbionagen. nov.: F. brachypteracomb. nov., F. phami Yu & Li, sp. nov. (♂♀; Hai Phong, Vietnam), and F. shenzhen Yu & Li, sp. nov. (♂♀; Guangdong, China).

Spiders of the genusClubionaLatreille, 1804 from Xishuangbanna, Yunnan Province, China are studied. A total of 47 species is reported and illustrated, including 14 new species and two new synonyms. Twelve of the new species belong to four species groups:C. dengpaoYu & Li,sp. nov.,C. subdidentataYu & Li,sp. nov.,C. tixingYu & Li,sp. nov.,C. xiaociYu & Li,sp. nov.,C. xiaokongYu & Li,sp. nov.,C. yejieiYu & Li,sp. nov.,C. zhaoiYu & Li,sp. nov.andC. zhigangiYu & Li,sp. nov.from theC. corticalisgroup;C. miiYu & Li,sp. nov.andC. subtongiYu & Li,sp. nov.from theC. ternatensisgroup;C. bannaYu & Li,sp. nov.from theC. filicatagroup; andC. menglunYu & Li,sp. nov.from theC. trivialisgroup. The remaining two new species,C. shuangsiYu & Li,sp. nov.andC. wangchengiYu & Li,sp. nov., are not readily assignable to any of the existing species groups. The female ofC. cochlearisYu & Li, 2019, the female ofC. tianeYu & Li, 2019, the female ofC. bicornisYu & Li, 2019, the male ofC. lalaJäger & Dankittipakul, 2010 and the true female ofC. suthepicaDankittipakul, 2008 are described for the first time. Two new synonyms are:C. vukomiJäger & Dankittipakul, 2010syn. nov. =C. circulataZhang & Yin, 1998;C. melanotheleThorell, 1895syn. nov. =Clubiona melanostictaThorell, 1890. A checklist ofClubionaspecies from Xishuangbanna is provided. The DNA barcodes of almost all of the species were obtained for species delimitation, matching of sexes and future use.

Clubiona qianlei J. Zhang, F. Zhang & H. Yu, sp. nov. Figs 1–4, 6 Type material. Holotype: ♂, CHINA: Hubei Province, Xianning City, Tongshan County, Jiugongshan Nature Reserve, Shilonggou (N29.41044598º, E114.64803038º, 480 m), 29.IV.2020, Q.L. Lu and Y. Zhong leg. Paratype: 1♀ (YHCLU0276), Jiugongshan Nature Reserve, Yunzhonghu (N29.40443038º, E114.67254442º, 1240 m), 1.VII.2020, Q.L. Lu et al. leg.; 1♂ (YHCLU0318), same data as holotype. Etymology. This species is a patronymic named after Mr. Qianle Lu (Shenzhen City, China), collector of the types, who has greatly helped us in our research. Diagnosis. The male of C. qianlei sp. nov. differs from those of all other group members by the absence of subapical barb on the RTA’s ventral process, and by the tegular apophysis width ca. 2/3 of bulb width (Figs 2B, C, 3B). In contrast, all other group members have subapical barb on the RTA’s ventral process, tegular apophysis narrower than half of bulb width, such as C. reclusa (Almquist 2006: 374, figs 323a, b) and C. interjecta (Figs 5A, C; Tang et al. 2005: 79, figs 3D, E). The female is similar to those of C. interjecta in the general appearance of the epigyne, but differs by: (1) copulatory openings slit-like, indistinct (Figs 4A–C) (vs. nearly circular, relatively conspicuous; Fig. 5D; Tang et al. 2005: 79, fig. 3B; Zhu & Zhang 2011: 358, fig. 257B); (2) dorsal part of spermatheca (SPd) with hyaline outermost surface (Figs 4D, E) (vs. surface sclerotized; Fig. 5E; Tang et al., 2005: 79, fig. 3C; Zhu & Zhang, 2011: 358, fig. 257B); (3) the two SPds separated by about one diameter (Figs 4D, E) (vs. closely spaced or separated by less than 0.5 diameter; Fig. 5E; Tang et al. 2005: 79, fig. 3C; Zhu & Zhang 2011: 358, fig. 257B) Description. Male (holotype): Total length 5.90. carapace 2.95 long, 2.05 wide; abdomen 2.95 long, 1.62 wide. Living holotype dark brown (Fig. 1A). Carapace yellowish-brown in ethanol (Figs 3D, F), slightly darker in front, without a distinct pattern; cephalic region distinctly narrowed, thoracic groove and radial grooves indistinct; tegument smooth, clothed with fine hairs. Eyes: AER slightly recurved, PER slightly wider than AER, almost straight in dorsal view. Eye sizes and interdistances: AME 0.15, ALE 0.14, PME 0.18, PLE 0.16, AME–AME 0.12, AME–ALE 0.07, PME–PME 0.33, PME–PLE 0.21, MOQL 0.34, MOQA 0.37, MOQP 0.60. Chelicerae protruding, robust, uniformly brownish red, with four teeth on promargin and three on retromargin. Sternum (Fig. 3E) yellowish white, 1.57 long, 1.02 wide. Labium coloured as chelicerae, endites brown. Legs light yellow, without distinct markings. Leg measurements: I 8.41 (2.37, 3.41, 1.66, 0.97), II 9.99 (2.55, 3.39, 3.11, 0.94), III 7.31 (2.21, 2.57, 1.90, 0.63), IV 10.36 (2.95, 3.44, 3.09, 0.88). Abdomen (Figs 3D–F) elongate, oval, dorsum centrally with a lengthwise heart mark, reaching half of abdomen length, with a pair of muscular depressions located on middle level of heart mark; laterally with lengthwise reticular pattern, posteriorly furnished with a fuzzy pattern; venter medially with two longitudinal dotted lines. Palp (Figs 2 A−D, 3A−C): Tibia short, ca. 1/3 length of cymbium. Retrolateral tibial apophysis (RTA) heavily sclerotized and bifurcated, slightly longer than tibia, broad at base; ventral process (VP) with sharp tip, shaped like a leaf in retrolateral view; dorsal process (DP) triangular, ca. 1/2 ventral process length. Genital bulb elongated and with a relatively flat tegulum, ca. 1.8 × longer than wide, with distinctive, sinuate sperm duct. Tegular apophysis (TA) represented by a large and flat sclerite, inverted triangular, anterior margin slightly concaved and rough, situated prolaterally on the tegulum, about 2/3 width of bulb. Embolar base (EB) inserted at approximately ten o’clock on tegulum, arising on the dorsal, hidden side of the tegular apophysis; free part of embolus (E) filamentous, as long as tegulum width, tip directed antero-mesally. Conductor (C) thick, finger-like, ca. 1/5 of tegulum length, originating from retrolateral, apical area of tegulum. Female. As in male, but slightly larger in size and lighter in colour. Living paratype light brown (Fig. 1B). Habitus in ethanol as Figs 4F, G. Total length 6.80; carapace 2.92 long, 2.09 wide; abdomen 3.88 long, 2.86 wide. Eye sizes and interdistances: AME 0.16, ALE 0.18, PME 0.19, PLE 0.11, AME–AME 0.05, AME–ALE 0.06, PME– PME 0.25, PME–PLE 0.20. MOQL 0.45, MOQA 0.38, MOQP 0.58. Sternum 1.56 long, 0.94 wide. Measurements of legs: I 6.49 (1.89, 2.68, 1.16, 0.76), II 6.68 (1.93, 2.72, 1.31, 0.72), III 5.97 (1.76, 2.13, 1.57, 0.51), IV 9.17 (2.63, 3.09, 2.76, 0.69). Epigyne (Figs 4 A−E): Epigyne plate a darkened disc, nearly triangular, anterior and lateral margins indistinct, posterior margin heavily sclerotized, middle part of posterior margin protruding; arrangement of the various parts of the vulva indistinctly visible through the tegument. Copulatory openings (CO) small, indistinct, represented by a short slit, situated at concave bends of epigynal plate posterior margin. Copulatory ducts (CD) firstly running along the posterior margin of epigyne, leading to large glands (GL), then ascending obliquely, finally connecting to anteriorly located spermathecae. Dorsal part of spermathecae (SPd) globular, innermost surface sclerotized, outermost surface hyaline, the two SPd separated by about one diameter. The ventral part of spermathecae (SPv) tubular or finger-like, curved medially, the two SPvs closely spaced. Fertilization ducts (FD) acicular, membranous, slightly shorter than diameter of SPv, located on dorsal surface of SPd. Natural History. Leaf-dwelling spiders. Photos of live specimens were taken after the spiders dropped on the ground due to beating. Distribution. Know only from the type locality, Mt. Jiugong, Hubei, China (Fig. 6). Clubiona interjecta L. Koch, 1879 Figs 5−6 Clubiona interjecta L. Koch 1879: 89, pl. 3, fig. 7 (♂ ♀); Zhang & Hu 1989: 57, figs 3, 18 (♂ ♀); Song et al. 1999: 416, figs 246F−G, 249E−F (♂ ♀); Tang et al. 2005: 79, figs 3A−E (♂ ♀); Zhu & Zhang 2011: 358, f. 257A-E (♂ ♀), World Spider Catalog 2022 (full list of taxonomic references). Material examined. CHINA: Hebei Province: Chengde City, Weichang County, 1♂ 5♀, Saihanba National Forest Park (N42.584537º, E117.837489º, 1500 m), 3.VIII.2018, X.B. Guo, H. Wang, J.X. Lai and Y.F. Li leg; Shijiazhuang City, Pingshan County, 1♂, Tuoliang Nature Reserve (N38.731813º, E113.824548º, 2000 m), 12. V.2018, X.B. Guo and Z.Y. Li leg; Zhangjiakou City, Zhuolu County, 27♀, Xiao Wutai Mountain Nature Reserve, Shanjiankou Village (N40.029531º, E115.063527º, 1300 m), 6.VII.2018, X.B. Guo, S. Qiao, J.X. Lai and Y.F. Li leg; Shanxi Province: Xinzhou City, Ningwu County, 1♂, Luya mountain (N38.837989º, E112.083789º, 1900 m), 6.VII.2011, C. Jin leg; Sichuan Province: Ganzi County, 1♂, Renguo village (N31.667885º, E99.803766º, 3420 m), 20. VI.2018, X.B. Guo Leg. Emended diagnosis. As the only two members of the reclusa -group in China, Clubiona interjecta and C. qianlei sp. nov. share almost all of the group characters, but they can be separated by the shape of the copulatory organs, and by some somatic characters. C. interjecta can be distinguished from C. qianlei sp. nov. by the following characters: for the males, RTA’s ventral process with a subapical barb, dorsal process finger-like in C. interjecta (Fig. 5C; Tang et al. 2005: 79, fig. 3E) (vs. subapical barb absent, dorsal process triangular; Fig. 2B); tegular apophysis about 1/3 width of bulb, distally narrowed and bearing a dentiform process (Fig. 5A; Tang et al. 2005: 79, fig. 3D) (vs. tegular apophysis about 2/3 width of bulb, distal margin wide and rough; Figs 2C, 3B); for the females, copulatory openings distinct, nearly circular in C. interjecta (Fig. 5D; Tang et al. 2005: 79, fig. 3B) (vs. copulatory openings indistinct, slit-like; Figs 4 A−C); SPd’ surface sclerotized, the two SPds nearly closely spaced (Fig.5E; Tang et al. 2005: 79 fig. 3C) (vs. SPd’ outermost surface hyaline, the two SPds separated by about one diameter; Figs 4D, E). In addition, the two species can be separated by the abdominal pattern: dorsum of abdomen anteriorly with distinct, longitudinal heart mark, reaching 1/2 of abdomen length, posteriorly with 4~6 transverse chevrons in C. interjecta (Figs 5F, G; Tang et al. 2005: 79, fig. 3A), but only with indistinct heart mark and muscular depressions in C. qianlei sp. nov. (Figs 3D, 4F). Description. See Tang et al. (2005). Male palp as in Figs 5 A−C, epigyne as in Figs 5D, E, habitus as in Figs 5F, G. Distribution. Russia (West Siberia to Far East), Mongolia, China (Inner Mongolia, Hebei, Shanxi, Henan, Sichuan, Jilin, Heiongjiang, as in Fig. 6).

Clubiona reclusa species-group Diagnosis. See Mikhailov (1995)., Published as part of Zhang, Jianshuang, Chen, Lulu, Ding, Yanmei, Zhang, Feng & Yu, Hao, 2022, On the Clubiona reclusa species-group in China, with the description of Clubiona qianlei sp. nov. (Araneae, Clubionidae), pp. 412-421 in Zootaxa 5129 (3) on page 413, DOI: 10.11646/zootaxa.5129.3.5, http://zenodo.org/record/6501213, {"references":["Mikhailov, K. G. (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta, 4 (2), 33 - 48."]}

The representation of the Clubiona reclusa species-group in China is here established in two species: Clubiona qianlei sp. nov. from Central China and C. interjecta L. Koch, 1879, which is mainly distributed in northern China. Detailed description, diagnosis, photographs of the new species are given. DNA barcodes (a partial fragment of the mitochondrial cytochrome oxidase subunit I gene, COI) of the new species were obtained to confirm matching of the sexes and for future use in molecular studies. Supplementary micrographs of C. interjecta are provided for the first time, alongside an emended diagnosis, to demonstrate the validity of C. qianlei sp. nov.. A distribution map of the reclusa group species in China is given.

The monotypic genus Bucliona Benoit, 1977 (type Clubiona dubia O. Pickard-Cambridge, 1870 from Saint Helena Island), previously considered a junior synonym of Clubiona Latreille, 1804, is resurrected. Three species are assigned to the genus Bucliona: B. dubia comb. reval., B. kirilli sp. n. (♂, Kenya), and B. jucunda (Karsch, 1879) comb. n., ex Clubiona (♂♀, Far East Asia).

Bucliona kirilli Yu & Li sp. n. Figs 2E, F, 5, 6, 7A, C Type material. Holotype ♂ (NMK), KENYA: Nyeri County, Naro Moru Town, Mount Kenya National Park, Metrological Station (S0.1702��, E37.2140��, 3000 m), 16.VIII.2018, Kioko G. leg. Other material examined. KENYA: Laikipia County, Nanyuki Town, Mount Kenya National Park, Sirimon Gate, Bamboo Forest (S0.2675��, E37.28276��, 3176 m), 1♂ (MGEU), 17.VIII.2018, Kioko G. leg. Etymology. The species is named in honour of Kirill G. Mikhailov from Moscow State University for his contributions to the systematics of clubionid spiders and on the occasion of his 60 th birthday; noun (name) in genitive case. Diagnosis. Males of B. kirilli sp. n. resemble those of B. dubia but differ by the parallel sides and a triangular tip of the RTA (vs. wide base and claw-like tip), as well as by the tip of the embolus which is directed antero-mesally (vs. anteriorly). Description. Male. Total length 7.66; carapace 3.65 long, 2.54 wide; abdomen 4.01 long, 2.14 wide. Carapace (Figs 6A���C) uniformly yellow-brown. Labium and endites dark reddish brown. Sternum coloured as carapace. Eyes: AER slightly recurved, PER wider than AER and straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.17, ALE 0.20, PME 0.17, PLE 0.18, AME���AME 0.14, AME���ALE 0.15, PME���PME 0.39, PME���PLE 0.28, MOQL 0.47, MOQA 0.44, MOQP 0.71. Legs yellowish, without distinct pattern. Leg measurements: I and II missing, III (2.50, ���, ���, ���), IV 11.45 (3.20, 3.95, 3.33, 0.97). Abdomen (Figs 6A���C) with wide dorsal scutum covering almost entire abdomen; dorsum anteriorly with triangular, dark, median stripe, reaching 1/3 of abdomen length, centrally without distinct patterns (faded in the preserved holotype but with 2 transverse chevrons in the second female), posteriorly with 3 transverse chevrons; venter anteriorly without any markings, posteriorly with 2 broken longitudinal stripes. Palp (Figs 5A���E, 7A, C): Tibia relatively long, over 1/2 cymbium length, ca. 1.9 times longer than wide, retrolateral tibial apophysis (RTA) about 1/2 of tibia length, about 5 times longer than wide, slightly wider at base, gradually tapering towards apex, tip triangular, sharply pointed. Embolus ��-shaped in ventral view, bent at almost a right angle, tip directed antero-mesally, base (EB) slightly longer than free part (E), embolus 6.6 times longer than wide. Tegular apophysis (TA) heavily sclerotized, arising retrolatero-distally from tegulum, finger shaped, directed proximally, distally curved retrolaterally. Female. Unknown. Distribution. Known only from the type locality., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on pages 203-206, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237

Bucliona dubia (O. Pickard-Cambridge, 1870) comb. reval. Fig. 1 Clubiona dubia O. Pickard-Cambridge, 1870: 532, pl. 42, fig. 1 (♀); O. Pickard-Cambridge, 1873: 213, pl. 24, fig. 3 (♂). Bucliona dubia: Benoit 1977: 68, figs 26a���c, 27a���d (♂ ♀). Clubiona (Bucliona) dubia: Mikhailov 1997: 96, figs 1���4 (♂ ♀). Type examined. Photograph of the holotype female from OUMNH, Mr. Melliss���s Saint Helena Island collection, May 1869. Diagnosis and description. See Benoit (1977) and Mikhailov (1997). Epigyne and habitus of the holotype female are as in Fig. 1A���D. Comments. The holotype female was thought to be lost (Benoit 1977), however Zo�� Simmons, the curator of OUMNH has managed to find the vial with the holotype female. Distribution. The species is known from Saint Helena Island only., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on page 198, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237, {"references":["Pickard-Cambridge, O. (1870) Notes on some spiders and scorpions from St Helena, with descriptions of new species. Proceedings of the Zoological Society of London, 37 (3, 1869), 531 - 544, pl. 42.","Benoit, P. L. G. (1977) Fam. Clubionidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences Zoologiques (Zool. - Ser. 8 °), 220, 64 - 81.","Mikhailov, K. G. (1997) Redescription of Clubiona dubia O. Pickard-Cambridge, 1869 (Aranei Clubionidae) from Saint Helena Island. Arthropoda Selecta, 6 (1 / 2), 95 - 97."]}

Genus Bucliona Benoit, 1977, status revalidated Bucliona Benoit, 1977: 68. Bicluona Mikhailov, 1994: 52 (described as a subgenus of Clubiona, type Liocranum jucundum Karsch, 1879). Bucliona: Mikhailov 1997: 95 (downgraded to subgenus of Clubiona and synonymized with Bicluona). Type species. Clubiona dubia O. Pickard-Cambridge, 1870 from Saint Helena Island, by monotypy. Diagnosis. Bucliona differs from Clubiona sensu stricto (type species Araneus pallidulus Clerck, 1757) by: having a distinct mesal ridge on the male chelicerae (Fig. 2A, C) (vs. lacking in Clubiona); the presence of a dorsal abdominal scutum in males (Figs 4E, F, 6A, C) (vs. absent); the undivided, short tibial apophysis of the male palp (Figs 3B, 5B, 7B���D) (vs. divided in many species); the presence of a tegular (= median) apophysis and the lack of a meandering sperm duct (Figs 3B���E, 5A���E) (vs. lack of a tegular apophysis and presence of a meandering sperm duct); the subtegulum with a prolateral extension (Fig. 8A) (vs. without, Figs 8B���F); the tip of the embolus directed anteriorly or antero-mesally (Figs 3D, 5D, 7A, B) (vs. retrolaterally or posteriorly). Females of Bucliona differ from those of Clubiona sensu stricto in having an epigyne with a distinct, undivided epigynal fovea with an anterior hood and a pair of posterior hoods (Figs 1C, D, 4A, C) (vs. fovea and hoods lacking) and longer copulatory ducts (Fig. 4D). Both the males and females of Bucliona can be recognized by the anterior tibiae with a prolateral ���scopula���, which is reduced to a single row (Fig. 2D���F) (vs. dense scopula ventrally on the anterior tibiae). Description. Medium to large size, with the body length of males 4.90���7.75 and of females 6.50���9.50. Male. Body yellow-brown, legs uniformly coloured as carapace (Figs 4E, F, 6A���C). Carapace (Figs 4E, F, 6A, C): elongate-oval, widest at midpoint, pars cephalica slightly elevated above thorax, pars thoracica distinctly wider and slightly higher than pars cephalica; integument smooth; distinctly darker in ocular region, without distinct pattern; cervical groove and radial grooves distinct; fovea a short slit on posterior half of carapace, longitudinal and reddish. Clypeus height distinctly narrower than AME (Figs 2A, C). Chelicerae (Figs 2A, C, 4E, F, 6A���C): robust and red brownish, consisting of a coniform paturon and claw-shaped fang; cheliceral paturon with distinct mesal ridge, distally with several setae; fang furrow with 3 promarginal and 2 retromarginal teeth. Both endites and labium longer than wide; endites depressed posteriorly, slightly convergent anteriorly, with dense scopulae on inner margin; labium nearly trapezoidal and depressed laterally (Fig. 6B). Sternum (Fig. 6B): shield shaped and brown, longer than wide, anteriorly straight; posterior region protruding strongly between coxae IV. Eyes: in dorsal view, AER slightly recurved, PER almost straight (Figs 4E, 6A); in anterior view, AER almost straight, PER slightly recurved (Figs 2A, C); AME very slightly smaller than ALE, or equal in diameter, AME closer to ALE than to each other; in dorsal view, PME and PLE nearly equal in diameter, well-separated, PME located about 2���3 diameters apart, PME���PLE distance ca. 2/3 PME���PME distance. Legs: leg formula 4213; anterior legs with conspicuous scopulae disto-prolaterally on tibiae, and entire metatarsi and tarsi (Figs 2D, F); anterior femora with 3���5 dorsal spines, posterior femora with 5���7 dorsal spines; all patellae with 0���1 dorsal or retrolateral spine; tibiae I���II with 2 or 3 pairs of ventral spines, 1 retrolateral, and 1 prolateral spine; metatarsi I���II with 1 or 2 pair of spines, 1 retrolateral, and 1 prolateral spine; tibiae and metatarsi of posterior legs with more spines than anterior legs but spination varies among different individuals. Abdomen (Figs 4E, F 6A���C): elongate-oval, tapering posteriorly, dorsal scutum over 3/4 of length of abdomen; without distinct colour patterns in the type species and B. jucunda (with median band and chevrons in B. kirilli sp. n., see description below). Palp: femur and patella unmodified, tibia cylindrical with single, unbranched retrolateral apophysis (RTA), apophysis shorter than diameter of tibia; cymbium unmodified, lacking spines; bulb elongate-oval; subtegulum (ST) with characteristic prolateral extension (Figs 3A, C, 5A, C, 8A); sperm duct not meandering, U-shaped in ventral view of tegulum; tegulum with well-developed hook-shaped distal apophysis (TA); embolus helical, originating prolatero-distally, either filamentous and wrapping around dorsal side of tegulum (B. kirilli sp. n.) or shorter, tip directed either anteriorly (B. dubia) or antero-mesally (B. jucunda). Female. General characters as in males, from which the females differ by (1) the cheliceral base without a mesal ridge (Fig. 2B); (2) the abdomen without a scutum (Figs 1A, 4G); and (3) a denser ���scopula��� on anterior legs (Fig. 2E). Epigyne: epigynal plate with large fovea, located posteriorly, with 1 anterior (AH) and 2 posterior hoods (PH); copulatory openings (CO) small and indistinct, located in central portion of fovea; copulatory ducts (CD) hyaline, strongly convoluted; receptacles (R) small and located posteriorly; bursae (BS) situated anteriorly, balloon shaped, larger than receptacles. Composition. Bucliona dubia (Saint Helena Island), B. kirilli sp. n. (Kenya), and B. jucunda (Far East Russia, China, Korea, Japan)., Published as part of Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao & Li, Shuqiang, 2021, Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae), pp. 195-207 in Zootaxa 5006 (1) on pages 196-197, DOI: 10.11646/zootaxa.5006.1.21, http://zenodo.org/record/5157237, {"references":["Benoit, P. L. G. (1977) Fam. Clubionidae. In: La faune terrestre de l'ile de Sainte-Helene IV. Annales, Musee Royal de l'Afrique Centrale, Sciences Zoologiques (Zool. - Ser. 8 °), 220, 64 - 81.","Mikhailov, K. G. (1994) Bicluona Mikhailov, subgen. n., a new subgenus of spiders of the genus Clubiona (Aranei, Clubionidae) from eastern Asia. Zoologicheskii Zhurnal, 73 (11), 52 - 57.","Karsch, F. (1879) Baustoffe zu einer Spinnenfauna von Japan. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 36, 57 - 105.","Mikhailov, K. G. (1997) Redescription of Clubiona dubia O. Pickard-Cambridge, 1869 (Aranei Clubionidae) from Saint Helena Island. Arthropoda Selecta, 6 (1 / 2), 95 - 97.","Pickard-Cambridge, O. (1870) Notes on some spiders and scorpions from St Helena, with descriptions of new species. Proceedings of the Zoological Society of London, 37 (3, 1869), 531 - 544, pl. 42."]}

Clubiona reclusa O. Pickard-Cambridge, 1863 Material. 1 ♂, 1 ♀ (NMNHS), Vitosha Mts, Bistritsa village, N 42.5927 ˚, E 23.3401 ˚, 1070 m, 03.07.1982, 1 ♀ (NMNHS), 17.10.1982, L. Penev leg.; 1 ♂ (NMNHS), Vitosha Mts, Tintyava hut, N 42.6178 ˚, E 23.2584 ˚, 1620 m, 27.06.1984, 1 ♀ (NMNHS), 19.07.1984, C. Deltshev leg. Remarks. Widespread in Europe and Turkey to South Siberia and Kazakhstan (Nentwig et al. 2021; World Spider Catalog 2021)., Published as part of Naumova, Maria, Blagoev, Gergin & Deltshev, Christo, 2021, Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae), pp. 228-257 in Zootaxa 4984 (1) on page 231, DOI: 10.11646/zootaxa.4984.1.18, http://zenodo.org/record/4927030, {"references":["World Spider Catalog (2021) World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 28 February 2021) https: // doi. org / 10.24436 / 2"]}

This study is a part of an ongoing comprehensive inventory of Bulgarian spiders. A total of fifty spider species belonging to thirteen families are reported for the first time from Bulgaria. Four species are rejected from the Bulgarian spider checklist due to misidentification: Callilepis concolor Simon, 1914, Centromerus capucinus (Simon, 1884), Hoplopholcus labyrinthi (Kulczyński, 1903) and Mansuphantes prope fragilis (Thorell, 1875). Another four species (Drassodes villosus (Thorell, 1856), Entelecara flavipes (Blackwall, 1834), Lepthyphantes notabilis Kulczyński, 1887 and Singa semiatra L. Koch, 1867) are rejected after a new interpretation of the locations. Six species were omitted from the list of Bulgarian spiders as obviously doubtful records (Dysdera nicaeensis Thorell, 1873, Haplodrassus rufipes (Lucas, 1846), Harpactea hispana (Simon, 1882), Macaroeris cata (Blackwall, 1867), Nomisia celerrima (Simon, 1914) and Tenuiphantes monachus (Simon, 1884)). A new synonymy is proposed for Cyclosa strandjae Drensky, 1915 (= Cyclosa sierrae Simon, 1870 syn. nov.). In addition, new images with essential taxonomic value are provided for twenty-five species to facilitate their identification or to illustrate their intraspecific variability. To ensure correct identification, DNA barcoding was additionally used in some species.

The Clubiona zilla-group is a relatively small species group, distributed exclusively in East Asia, with only three species clearly documented so far. Clubiona hooda Dong & Zhang, 2016, which was previously placed in the C. trivialis-group, is assigned to the C. zilla-group in the present paper. A new spider of the C. zilla-group from Jiugong Mountain in China is described under the name of C. jiugong sp. nov. Detailed descriptions and photographs of the new species are provided.

CLUBIONIDAE Unidentified sp.: Auplopus militaris., Published as part of Kurczewski, Frank E., West, Rick C., Waichert, Cecilia, Kissane, Kelly C., Ubick, Darrell & Pitts, James P., 2020, New and unusual host records for North American and South American spider wasps (Hymenoptera: Pompilidae), pp. 1-112 in Zootaxa 4891 (1) on page 111, DOI: 10.11646/zootaxa.4891.1.1, http://zenodo.org/record/4309249

Kurczewski, Frank E., West, Rick C., Waichert, Cecilia, Kissane, Kelly C., Ubick, Darrell, Pitts, James P. (2020): New and unusual host records for North American and South American spider wasps (Hymenoptera: Pompilidae). Zootaxa 4891 (1): 1-112, DOI: https://doi.org/10.11646/zootaxa.4891.1.1

Clubiona milingae Barrion-Dupo, Barrion & Heong, 2013 was described from a single male and no additional specimens have been recorded. The original description was brief and the illustrations were inadequate. Clubiona milingae is redescribed and illustrated based on new material from the type locality and the new distribution region (Jianfeng Mountains and Limu Mountains of Hainan Island, China). The female is reported for the first time.

In the present paper, a worldwide checklist of Clubiona japonica-group spiders is provided based on published literature and authors’ collections. A new japonica-group species, Clubiona grucollaris sp. n. (♀♂) from Guizhou Province and Hainan Island of China is diagnosed, described, and illustrated. A distribution map of this species is given.

Clubiona zhanggureni sp. nov. Figs 13���14 Type material. Holotype: ♂ (IZCAS Ar 34714), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Secondary tropical montane evergreen broad-leaved forest (N21��57.809���, E101��12.173���, 888 m), 4 August 2007, leg. G. Zheng (Fogging-CBIII). Paratype: 1♂ (IZCAS Ar 34715), same data as holotype. Etymology. The specific name is after Prof. Dr. Zhang, Guren from Sun Yat-Sen University for his contribution on the systematics of Chinese clubionid species; noun (name) in genitive case. Diagnosis. Males of Clubiona zhanggureni sp. nov. resemble those of C. digitata Dankittipakul, 2012 (Dankittipakul et al. 2012: figs 1���3) in having similarly shaped embolus, and almost membranous conductor, but differ by: (1) retrolateral tibial apophysis with sharp apex (Fig. 13B) (apically blunt in C. digitata); (2) beak-shaped conductor with slightly sclerotized apex (Figs 13B, E) (triangular conductor completely membranous in C. digitata); (3) sperm duct represented by a reversed S-shaped course in ventral view (Fig. 13D) (3-shaped course in C. digitata). Description. Male (holotype): Total length 6.16; prosoma 2.77 long, 1.99 wide; opisthosoma 3.12 long, 1.51 wide. Prosoma (Figs 14 A���C), long-oval, pars cephalica distinctly narrowed, in profile strongly raised, highest highest between PME and fovea; integument smooth, clothed with short fine hairs. Carapace reddish-brown, darker in front, without distinct colour pattern; fovea longitudinal and dark. Chelicerae protruding and wine-colored, with four promarginal and two retromarginal teeth. Labium and endites reddish-brown, longer than wide. Sternum yellowish-brown. Eyes: AER slightly recurved, PER wider than AER and almost straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.15, ALE 0.10, PME 0.16, PLE 0.14, AME��� AME 0.11, AME���ALE 0.10, PME���PME 0.27, PME���PLE 0.17, MOQL 0.23, MOQA 0.29, MOQP 0.46. Legs light brown, without distinct color markings. Leg formula: IV, I, II, III; leg measurements: I 8.95 (2.39, 3.00, 3.00, 0.55), II 7.39 (1.98, 3.07, 1.64, 0.70), III 6.27 (1.76, 2.10, 1.84, 0.57), IV 9.18 (2.34, 3.13, 3.02, 0.69). Opisthosoma (Figs 14 A���C) lanceolate, grayish, with conspicuous anterior tufts of hairs, dorsum with dense grey hairs and a broken dark median band, reaching half of opisthosoma length, posteriorly with six or seven chevrons; venter brown. Spinnerets purplish. Palp (Figs 13 A���E): RTA dark, small but strong, triangular, sharply pointed; bulb nearly oval, proapically and apically membranous, slightly excavated on prolatero-apical side to accommodate embolus; sperm duct distinct and sinuate, reversed S-shaped; embolus spiniform, originated at 9 o���clock position in prolateral view, tip slightly overpasses the genital bulb; conductor membranous except beak-shaped and slightly sclerotized apex; tegular apophysis absent. Female: Unknown. Distribution. Presently known only from the type locality., Published as part of Yu, Hao & Li, Shuqiang, 2019, On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China, pp. 201-230 in Zootaxa 4679 (2) on pages 216-220, DOI: 10.11646/zootaxa.4679.2.1, http://zenodo.org/record/3772397, {"references":["Dankittipakul, P., Tavano, M. L., Chotwong, W. & Singtripop, T. (2012) New synonym and descriptions of two new species of the spider genus Clubiona Latreille, 1804 from Thailand (Araneae, Clubionidae). Zootaxa, 3532, 51 - 63. https: // doi. org / 10.11646 / zootaxa. 3532.1.4"]}

Clubiona bicornis sp. nov. Figs 15–16 Type material. Holotype: ♂ (IZCAS Ar 34716), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Primary tropical seasonal rain forest (N21º55.035′, E101º16.500′, 558 m), 22 July 2007, leg. G. Zheng (Fogging-GZI). Etymology. The species epithet is a Latin adjective and refers to the two-branched retrolateral tibial apophysis. Diagnosis. Males of Clubiona bicorn is sp. nov. can be easily distinguished from other members of the C. trivialis -group, with the exception of C. cheni Yu & Li, 2019 (Yu & Li 2019a: 171, figs 13A–E, 14E–F) by the flat body and similar palp. The palpal organ of C. bicorn is sp. nov. resemble those of C. cheni in having similarly shaped prolateral tibial apophysis, and retrolateral tibial apophysis with a long tip extending to mid length of cymbium, but differ by: (1) retrolateral tibial apophysis branched, both ventral and dorsal branch with sharp tip (Fig. 15B) (retrolateral tibial apophysis not branched and with blunt tip in C. cheni); (2) embolic base bearing only dentiform processes (Figs 15 A–E) (embolic base bearing two processes in C. cheni); (3) the wide but indistinct distal tegular hump (Fig. 15D) (developed more prominent in C. cheni). Description. Male (holotype): Total length 2.49; prosoma 1.09 long, 0.85 wide; opisthosoma 1.23 long, 0.70 wide. Prosoma (Figs 16 A–C), oval in dorsal view, ocular region slightly narrowed, widest between coxae II and III, in profile almost flat; integument smooth, clothed with numerous short fine hairs. Carapace orange, slightly darker in front, with a pair of unobvious short lines running longitudinally from behind AME. Chelicerae protruding and coloured as carapace, with three promarginal and two retromarginal teeth. Labium reddish-brown, endites orange, both longer than wide. Sternum yellowish-orange. Eyes: in dorsal view, both anterior and posterior eye rows recurved, PER wider than AER.AME dark, other eyes light; with black rings. Eye sizes and interdistances:AME 0.05, ALE 0.06, PME 0.06, PLE 0.05, AME–AME 0.02, AME–ALE 0.16, PME–PME 0.29, PME–PLE 0.08, MOQL 0.04, MOQA 0.09, MOQP 0.35. Legs yellowish-white, without distinct color markings. Leg formula: II, IV, I, III; leg measurements: I 2.24 (0.76, 0.95, 0.33, 0.19), II 2.53 (0.82, 0.95, 0.55, 0.21), III 1.69 (0.54, 0.61, 0.36, 0.18), IV 2.26 (0.66, 0.96, 0.41, 0.23). Opisthosoma (Figs 16 A–C) elongate-oval, dorsally white with two pairs of indistinct muscular depressions; venter and spinnerets light yellow. Palp (Figs 15 A–E): tibia short, with two apophyses, PTA digitiform, with a bent and blunt tip; RTA well-developed and forked, ventral branch small and with a row of teeth in retrolateral view, dorsal branch horn-shaped, sharp tip extending to mid length of cymbium; genital bulb elongated, with distinctive, sinuate sperm duct; tegular hump indistinct, fusing with conductor; embolic base bearing a dentiform processes; embolus originating from retrolateral side of tegulum, slender and flagelliform, angled across tegular hump, tip terminating at approximately two o’clock position relative to tegulum; conductor area relatively small, with approximately one-fifth the length of tegulum. Female: Unknown. Distribution. Presently known only from the type locality.

Clubiona cochlearis sp. nov. Figs 1���2 Type material. Holotype: ♂ (IZCAS Ar 34701), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Paramichelia baillonii plantation (about 20 yr.) (N21��54.772', E101��16.043', 556 m), 18 July 2007, leg. G. Zheng (Fogging-RZI). Paratype: CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, 1♂ (IZCAS Ar 34702), Rubber plantation (about 20 yr.) (N21��54.498', E101��16.326', 586 m), 17 July 2007, leg. G. Zheng (Fogging-XZII). Etymology. The species epithet is a Latin adjective and refers to the spoon-shaped embolus. Diagnosis. This new species can be easily distinguished from those of all other species of the C. corticalis group by the strong, spoon-shaped embolus (Figs 1A, C���D). Description. Male (holotype): Total length 7.34; prosoma 3.29 long, 2.31 wide; opisthosoma 3.59 long, 1.91 wide. Prosoma (Figs 2 A���C) pyriform, ocular region distinctly narrowed, widest between coxae II and III; in profile highest just behind longitudinal fovea; integument smooth, clothed with short fine hairs. Carapace red brown, without distinct color pattern; fovea dark. Chelicerae protruding and coloured as carapace, with three promarginal and two retromarginal teeth. Labium and endites dark brown, longer than wide. Sternum brown. Eyes: AER slightly recurved, PER wider than AER and slightly procurved in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.14, ALE 0.17, PME 0.14, PLE 0.16, AME���AME 0.11, AME���ALE 0.08, PME��� PME 0.33, PME���PLE 0.18, MOQL 0.28, MOQA 0.41, MOQP 0.63. Legs brown, without distinct color markings. Leg formula: IV, II, I, III; leg measurements: I 7.78 (2.30, 3.38, 1.54, 0.96), II 9.29 (2.61, 3.85, 1.87, 1.05), III 7.43 (2.16, 2.47, 1.90, 0.69), IV 9.99 (2.81, 3.42, 2.82, 0.95). Opisthosoma (Figs 2 A���C) elongate-oval, dorsal scutum ovoid, lightly sclerotized, furnished with a thick tuft of hairs anteriorly; dorsum brown, without distinct color pattern; venter gray. Spinnerets brown. Palp (Figs 1 A���E): Tibia short, with two apophyses, VTA large and blunt, with partly membranous tip, thumb-like; RTA dark and strong, triangular, sharply pointed; bulb elongate-oval, strongly bulged and prolapsed, sperm duct indistinct in ventral view; embolus distinctly strong, with a expand and torsional tip, spoon-shaped, the embolic base situated prolateral on the tegulum; conductor membranous and blade-shaped, long, with blunt tip. Female: Unknown. Distribution. Presently known only from the type locality., Published as part of Yu, Hao & Li, Shuqiang, 2019, On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China, pp. 201-230 in Zootaxa 4679 (2) on pages 202-204, DOI: 10.11646/zootaxa.4679.2.1, http://zenodo.org/record/3772397

Clubiona kai Jäger & Dankittipakul, 2010 Figs 7–8 Clubiona kai Jäger & Dankittipakul, 2010: 25, figs 4–12 Material examined. CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, 1♂, Lvshilin Forest Park, Limestone tropical seasonal rain forest (N21º54.714′, E101º16.935′, 660 m), 16 November 2009, leg. G. Tang and Z.Y. Yao (Tang-Yao_No.11); 3♂, 7♀, Lvshilin Forest Park, Limestone tropical seasonal rain forest (N21º54.555′, E101º16.860′, 610 m), 29 November 2009, leg. G. Tang and Z.Y. Yao (Tang-Yao_No.33). Diagnosis. This species can be easily distinguished from those of the other species belonging to the corticalis - group, with the exception of C. didentata (Figs 5–6), by their bulged bulb with tegular apophysis and the epigynal plate with atrial membrane (tegular apophysis and atrial membrane are usually absent in other corticalis -group species). It can be separated from C. didentata by the larger, twisted embolus, smaller and partly membranous conductor, smaller and teeth-shaped tegular apophysis (Figs 7 A–E vs. Figs 5 A–E), and by the disc-shaped atrial membrane and the smooth bursae (Figs 8 A–C vs. Figs 6 A–C). Description. Male: Total length 3.60; prosoma 1.69 long, 1.32 wide; opisthosoma 1.30 long, 1.94 wide. Prosoma (Figs 8 D–E), ovoid in dorsal view, ocular region slightly narrowed, widest between coxae II and III; in profile, slightly higher between ocular area and longitudinal fovea, gradually sloping towards pars cephalica; integument smooth, clothed with short fine hairs. Carapace brown, darker anteriorly, without distinct color pattern; fovea reddish. Chelicerae protruding and wine-coloured, with five promarginal and two retromarginal teeth. Labium and endites light brown, longer than wide. Sternum yellowish white. Eyes: AER slightly recurved, PER wider than AER and slightly procurved in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.12, ALE 0.13, PME 0.14, PLE 0.13, AME–AME 0.05, AME–ALE 0.07, PME–PME 0.21, PME–PLE 0.12, MOQL 0.16, MOQA 0.15, MOQP 0.33. Legs yellowish-white, without distinct color markings. Leg formula: IV, II, III, I; leg measurements: I 3.60 (0.96, 1.68, 0.80, 0.38), II 4.26 (1.30, 1.80, 0.79, 0.48), III 3.68 (1.11, 1.16, 1.02, 0.37), IV 4.73 (1.37, 1.73, 1.45, 0.52). Opisthosoma (Figs 8 D–E) elongate-oval, marked with numerous brown spots. Palp (Figs 7 A–E): tibia short, only with retrolateral apophysis; RTA broad, flat and with a semicircular flange; bulb oval and strongly bulged, extending prolaterally and retrolaterally beyond cymbium, with several fanshaped markings; sperm duct inconspicuous; embolus with basal torsion and distal sickle-shaped bend; conductor small and papilliform, with membranous tip; tegular apophysis small and tooth-shaped, pointing prolatero-distally. Female: Total length 4.24; prosoma 1.60 long, 1.26 wide; opisthosoma 2.61 long, 1.63 wide. General color lighter than in male (Figs 8 F–G). Eye sizes and interdistances: AME 0.10, ALE 0.12, PME 0.13, PLE 0.11, AME– AME 0.02, AME–ALE 0.03, PME–PME 0.22, PME–PLE 0.10, MOQL 0.16, MOQA 0.15, MOQP 0.32. Legs yellowish-white, without distinct color markings. Leg formula: IV, I, III, II; leg measurements: I 3.56 (0.98, 1.46, 0.74, 0.38), II 3.03 (1.09, 0.98, 0.59, 0.38), III 3.34 (1.05, 1.07, 0.90, 0.32), IV 4.86 (1.29, 1.62, 1.45, 0.52). Epigyne (Figs 8 A–C): Epigynal plate distinctly longer than wide, margin not rebordered; atrium small, covered by atrial membrane; atrial membrane disc-shaped, located on the anterior margin of atrium; bursae prominently visible through epigynal plate in ventral view; copulatory openings small, located at basolateral atrial borders, leading to short copulatory ducts which descend obliquely to connect with spermathecae; spermathecae with bean-shaped proximal part and tubular distal part; fertilization ducts short and curved, acicular; bursae oblong, translucent, surface smooth except several transversal depressions. Distribution. Laos, China (Xishuangbanna, Yunnan Province). The new record presented here extends the known range of this species to the northwest.

Clubiona tongi sp. nov. Figs 9���10 Type material. Holotype: ♂ (IZCAS Ar 34705), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Garbage dump, Secondary tropical forest (N21��54.380', E101��16.815', 627 m), 23 November 2009, leg. G. Tang and Z.Y. Yao (Tang-Yao_No.21). Paratypes: CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, 1 ♂, 2♀ (IZCAS Ar 34706���34708), G213 roadside, Bamboo plantation (N21��53.622', E101��16.955', 581 m), 26 November 2009, leg. G. Tang and Z.Y. Yao (Tang- Yao_No.27). Etymology. The specific name is a patronymic in honor of Dr. Tong, Yanfeng for his contribution on the taxonomy of Chinese spiders; noun (name) in genitive case. Diagnosis. Clubiona tongi sp. nov. can be easily distinguished from other members of the C. hystrix-group, with the exception of C. theoblicki Yu & Li, 2019 (Yu & Li 2019a: 161, figs 7A���E; 8A���H; Yu & Li 2019b: 40) by the distinctly long embolus, the retrolateral tibial apophysis with a spine-like tip, and the distinctly long and convoluted copulatory ducts. It differs from C. theoblicki by: (1) the tegular hump with a blunt and semicircular tip, resembling a wave crest in ventral view (Fig. 9D) (tegular hump nearly quadrate in C. theoblicki); (2) the more or less lengthwise epigynal ridges (Figs 10 A���B) (vs. the diagonal ridges in C. theoblicki); (3) the anterior part of the copulatory ducts represented by two transversal loops (Figs 10 C���D) (forming two longitudinal loops in C. theoblicki). Description. Male (holotype): Total length 3.18; prosoma 1.60 long, 1.02 wide; opisthosoma 1.60 long, 0.98 wide. Prosoma (Figs 10 E���F), oval, pars cephalica slightly narrowed, widest between coxae II and III, in profile slightly higher just behind ocular region, gradually sloping posteriorly; integument smooth, clothed with short fine hairs. Carapace light orange, without distinct color pattern; longitudinal fovea reddish. Chelicerae coloured as carapace, with six promarginal and three retromarginal teeth. Labium and endites light orange, longer than wide. Sternum yellowish. Eyes: both anterior and posterior eye rows slightly recurved in dorsal view; AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.12, PLE 0.11, AME���AME 0.03, AME���ALE 0.02, PME���PME 0.14, PME���PLE 0.07, MOQL 0.13, MOQA 0.13, MOQP 0.25. Legs yellowish, without distinct color marking. Leg formula: IV, II, I, III; leg measurements: I 3.15 (0.91, 1.30, 0.56, 0.38), II 3.35 (0.98, 1.32, 0.63, 0.42), III 3.03 (0.92, 0.97, 0.79, 0.35), IV 4.68 (1.40, 1.54, 1.25, 0.50). Opisthosoma (Figs 10 E���F) elongate-oval, dorsally yellowish-white with a pair of muscular depressions; with conspicuous anterior hair tufts; venter and spinnerets light yellow. Palp (Figs 9 A���E): tibia short, only with retrolateral apophysis, RTA small, with a rhomboid base and a spine-like tip; genital bulb elongated, with relatively flat tegulum, sperm duct distinctive, Vshaped; tegulum distinctly longer than wide, apex with a semicircular tegular hump, base with a papilliform flange; embolus distinctly long, slender, basal portion wide, inserted prolaterally (approximately ten o���clock relative of tegulum), aligned clockwise along the tegular distal margin, apex filiform, terminated at approximately five o���clock position relative to tegulum. Female (paratype IZCAS Ar 34707): Total length 3.43; prosoma 1.51 long, 1.04 wide; opisthosoma 1.80 long, 0.96 wide. Not strikingly different from males but darker in color (Figs 10 G���H). Eye sizes and interdistances: AME 0.11, ALE 0.10, PME 0.10, PLE 0.07, AME���AME 0.04, AME���ALE 0.04, PME���PME 0.19, PME���PLE 0.08, MOQL 0.15, MOQA 0.14, MOQP 0.27. Legs yellowish, without distinct color marking. Leg formula: IV, II, I, III; leg measurements: I 2.57 (0.78, 1.03, 0.52, 0.25), II 2.64 (0.78, 1.05, 0.47, 0.35), III 2.41 (0.77, 0.80, 0.55, 0.30), IV 4.18 (1.41, 1.33, 1.05, 0.39). Epigyne (Figs 10 A���D): Epigynal plate slightly longer than wide, through which spermathecae and copulatory ducts well visible, posterior margin not rebordered; atrium absent; two copulatory openings small, contiguous, situated at medial portion of epigynal plate posterior margin, hidden in two lengthwise ridges in ventral view; copulatory ducts strongly entwined, ascending dorsally and expanding laterally two times, respectively, forming two horizontal loops and then connected to lateral spermathecae; both spermathecae and bursae globular, the former anteriad and smaller than the latter; bursae translucent with smooth surface; fertilization ducts acicular, membranous, located on dorsal-basal surface of spermathecae. Distribution. Presently known only from the type locality., Published as part of Yu, Hao & Li, Shuqiang, 2019, On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China, pp. 201-230 in Zootaxa 4679 (2) on pages 212-215, DOI: 10.11646/zootaxa.4679.2.1, http://zenodo.org/record/3772397, {"references":["Yu, H. & Li, S. (2019 a) Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae). Zootaxa, 4545 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 4545.2.1","Yu, H. & Li, S. (2019 b) Clubiona theoblicki, a name to replace Clubiona quadrata (Araneae, Clubionidae). Acta Arachnologica Sinica, 28 (1), 40. https: // doi. org / 10.3969 / j. issn. 1005 - 9628.2019.01.003"]}

Clubiona hystrix species-group Diagnosis. The Clubiona hystrix -group resembles the apiculata group in having the similar, simple tegulum with U-shaped sperm duct in male, and in having sclerotized spermathecae and the hyaline bursae in female, but can be distinguished by (see also Deeleman-Reinhold 2001; Dankittipakul & Singtripop 2014): (1) conductor absent; (2) embolus oriented clockwise along the margin of the tegulum instead of pointing apically; (3) retrolateral tibial apophysis relatively smaller and weakly developed; (4) copulatory opennings usually hidden in ridges, folds or under a hood; (5) copulatory ducts usually convoluted., Published as part of Yu, Hao & Li, Shuqiang, 2019, Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae), pp. 151-178 in Zootaxa 4545 (2) on page 161, DOI: 10.11646/zootaxa.4545.2.1, http://zenodo.org/record/2618741, {"references":["Deeleman-Reinhold, C. L. (2001) Forest Spiders of South East Asia: with a Revision of the Sac and Ground Spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp.","Dankittipakul, P. & Singtripop, T. (2014) A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae). Journal of Natural History, 48 (31 - 32), 1923 - 1936. https: // doi. org / 10.1080 / 00222933.2014.902140"]}

Clubiona trivialis species-group Diagnosis. Members of the Clubiona trivialis -group can be recognized by the following combination of characters (see also Dondale & Redner 1982 and Mikhailov 1990): male retrolateral tibial apophysis simple, erect, not conspicuously toothed; tegulum convex; embolic base usually with one to three dentiform processes; embolus arched around or angled across the distal end of tegulum; conductor groove-like, membranous, fused to tegulum, located distally on retrolateral side of tegulum; sperm duct sinuous, distinct. Female epigyne with large ventral plate and a semitranslucent dorsal plate; ventral epigynal plate disc-shaped, posterior margin heavily sclerotized and extending over epigastric furrow; copulatory openings located posteriorly on ventral epigynal plate, united at midline or separated; copulatory ducts slender, straight or arched laterad, parallel and close together at midline; spermathecae located anteriorly and connected posteriorly to bursae, both sandwiched between ventral and dorsal epigynal plates., Published as part of Yu, Hao & Li, Shuqiang, 2019, Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae), pp. 151-178 in Zootaxa 4545 (2) on page 171, DOI: 10.11646/zootaxa.4545.2.1, http://zenodo.org/record/2618741, {"references":["Dondale, C. D. & Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194.","Mikhailov, K. G. (1990) The spider genus Clubiona Latreille 1804 in the Soviet Far East, 1 (Arachnida, Aranei, Clubionidae). Korean Arachnology, 5, 139 - 175."]}

Clubiona jiandan sp. nov. Figs 19–20 Type material. Holotype: ♂ (IZCAS Ar 34720), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Secondary tropical montane evergreen broad-leaved forest (N21º57.784', E101º11.947', 895 m), 6 August 2007, leg. G. Zheng (Fogging-CBI). Paratypes: CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve: 1♂, 1♀ (IZCAS Ar 34721–34722), Secondary tropical montane ever- green broad-leaved forest (N21º57.809', E101º12.173', 888 m), 4 August 2007, leg. G. Zheng (Fogging-CBIII); 1♀ (IZCAS Ar 34723), Secondary tropical montane evergreen broad-leaved forest (N21º57.767', E101º12.431', 880 m), 6 August 2007, leg. G. Zheng (Fogging-CBII). Etymology. The specific name is derived from the Chinese pinyin 'jian dan', which means 'simple', referring to the simple genital bulb and the weakly developed tibial apophysis; adjective. Diagnosis. Males of Clubiona jiandan sp. nov. resembles those of C. yaoi Yu & Li, 2019 (Yu & Li 2019a: 152, figs 1A–E; 2A–H) in having a simple palpal bulb with U-shaped sperm duct, transparent, lamelliform conductor, and short embolus (Figs 19 C–E), but differ by: (1) tibia only with retrolateral apophysis (Fig. 19B) (with three apophyses in C. yaoi); (2) retrolateral tibial apophysis unbranched, small and weakly developed (Fig. 19B) (retrolateral tibial apophysis distally forked and well developed in C. yaoi); (3) dorsal cymbial apophysis absent (Figs 19 A–B); (4) embolus cone-shaped and partly membranous (Figs 19 C–E) (embolus spiniform in C. yaoi). Females also resemble those of C. yaoi by the general shape of vulva but can be recognized by the oval spermathecae (Figs 20 B–D) (globular in C. yaoi), and by the globular bursae (Figs 20 B–D) (oblong in C. yaoi). Taxonomic remarks. Clubiona jiandan sp. nov. possesses several characters associated with the apiculata - group and resembles C. yaoi (the only apiculata -group species recorded from China) for their characteristic genital organs (for a detailed diagnosis, see above). The males of the apiculata -group (known for C. yaoi and four apiculata group species from Borneo reported by Dankittipakul & Singtripop 2014) share the following distinctive suite of characters, here contrasted with the corresponding condition in this new species: (1) the presence of the tubercle at dorsal cymbium (C. jiandan sp. nov.: dorsal cymbial apophysis absent, Figs 19 A–B); (2) males with at least two palpal tibial apophyses, the retrolateral one usually well-developed (C. jiandan sp. nov.: tibia only with retrolateral apophysis, RTA small, simple and weakly developed, Fig. 19B). Female closely resembles those of the apiculata - group, in possessing a vulva with ascending copulatory ducts connected to bursae at mid length between copulatory openings and spermathecae, and a pair copulatory ducts lacking convolution. In addition, this new species can be separated from existing members of the apiculata -group by number of ventral spines in tibia I and II, namely two pairs in C. jiandan sp. nov., but three pairs in known apiculata -group species. Nevertheless, considerable uncertainty about placing this new species in the apiculata -group remains. Despite the fact that the general shape of palpal bulb and vulva of Clubiona jiandan sp. nov. is fairly typical of a member of the apiculata -group, it is currently impossible to discern any obvious derived features (i.e., well-developed RTA) that could indicate a close relationship to the apiculata -group. Description. Male (holotype): Total length 3.55; prosoma 1.74 long, 1.17 wide; opisthosoma 1.76 long, 0.96 wide. Prosoma (Figs 20 E–F), oval, pars cephalica slightly narrowed, widest between coxae II and III, in profile slightly higher just behind longitudinal fovea, gradually sloping towards pars cephalica; integument smooth, clothed with numerous short fine hairs. Carapace yellowish-orange, without distinct color pattern; fovea reddish. Chelicerae protruding and orange, with five promarginal and three retromarginal teeth. Labium and endites yellowish-orange, longer than wide. Sternum yellowish. Eyes: both AER and PER straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.11, ALE 0.11, PME 0.14, PLE 0.11, AME–AME 0.04, AME–ALE 0.02, PME–PME 0.02, PME–PLE 0.08, MOQL 0.14, MOQA 0.16, MOQP 0.32. Legs yellowish-white, without distinct color markings. Leg formula: IV, I, II, III; leg measurements: I 4.23 (1.27, 1.75, 0.76, 0.44), II 3.51 (0.99, 1.49, 0.66, 0.37), III 2.94 (0.88, 1.03, 0.75, 0.28), IV 4.52 (1.36, 1.55, 1.22, 0.38). Opisthosoma (Figs 20 E–F) elongate-oval, yellowish-white, dorsum without color marking; venter white; all spinnerets without distinct color marking. Palp (Figs 19 A–E): Tibia short, with single retrolateral apophysis; RTA small and flat, triangular; genital bulb elongated, with a relatively flat tegulum, sperm duct distinct, U-shaped; embolus, a small cone, originating from apical, prolateral side of tegulum, apex points ventrally; conductor represented by a transparent lamina, originating on apical, retrolateral area of tegulum. Female (paratype IZCAS Ar 34723): Total length 4.08; prosoma 1.45 long, 1.16 wide; opisthosoma 2.55 long, 1.38 wide. Not strikingly different from males but slightly darker in color (Figs 20 G–H). Eye sizes and interdistances: AME 0.13, ALE 0.09, PME 0.11, PLE 0.09, AME–AME 0.03, AME–ALE 0.03, PME–PME 0.19, PME–PLE 0.05, MOQL 0.13, MOQA 0.13, MOQP 0.31. Legs yellowish-white, without distinct color markings. Leg formula: IV, II, I, III; leg measurements: I 2.84 (0.72, 1.28, 0.48, 0.37), II 2.97 (0.93, 1.13, 0.57, 0.33), III 2.74 (0.82, 0.92, 0.68, 0.32), IV 3.88 (1.17, 1.32, 1.08, 0.32). Epigyne (Figs 20 A–D): Epigynal plate longer than wide, posterior margin concaved in the middle; atrium absent; spermathecae, bursae and copulatory ducts prominently visible through epigynal plate in ventral view; two copulatory openings distinctive and large, located at postero-lateral portion of epigynal plate; copulatory ducts ascending anteriorly, connect with spermathecae; spermathecae oval, basolaterally with digitiform head; bursae globular, connected to copulatory ducts at mid length between copulatory openings and spermathecae; fertilization ducts acicular, membranous, located on dorsal-lateral surface of spermathecae. Distribution. Presently known only from the type locality.

Clubiona yueya sp. nov. Figs 11���12 Type material. Holotype: ♂ (IZCAS Ar 34709), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanic Garden, 300 Acre-feet Bamboo plantation (N21��53.901', E101��16.884', 515 m), 7 August 2018, leg. H. Yu and Z.G. Chen (Yu_20180807_Fogging-9). Paratypes: 1 ♂, 3♀ (IZCAS Ar 34710���34713), same data as holotype. Etymology. The specific name is derived from the Chinese pinyin 'yu�� y��', which means 'crescent moon', referring to the crescent-shaped tegular apophysis; noun in apposition. Diagnosis. Males of Clubiona yueya sp. nov. can be easily distinguished from those of all other species of the C. japonica group by the crescent-shaped tegular apophysis (Fig. 11D) and the inconspicuous sperm duct (Figs 11 A���E) (vs. tegular apophysis absent and sperm duct distinct in almost all the C. japonica group species, such as Clubiona zhanggureni sp. nov.; Figs 13 C���E).). Females of C. yueya sp. nov. are similar to C. digitata Dankittipakul, 2012 (Dankittipakul et al. 2012: 52, figs 4���5, 17), C. filifera Dankittipakul, 2008 (Dankittipakul et al. 2012: 57, figs 18���19, 23���24), and C. grucollaris Yu, Zhang & Chen, 2017 (Yu et al. 2017a: 4, figs 13���14, 18���19) by the broad atrium with M-shaped anterior margin, and the globular bursae, but can be recognized by the small and bean-shaped spermathecae (Fig. 12C) (relatively long and tubular spermathecae in the latter three species). In addition, the new species can be consistently separable by its unique habitus: dorsal opisthosoma with three longitudinal bands (Figs 12D, F). Description. Male (holotype): Total length 5.42; prosoma 1.29 long, 1.63 wide; opisthosoma 3.07 long, 1.45 wide. Prosoma (Figs 12 D���E), long-oval, narrowed behind ocular area, in profile slightly raised, highest just in front fovea; integument smooth, clothed with short fine hairs. Carapace yellowish-white, darker in ocular area, with irregular reticular pattern behind posterior eyes; fovea longitudinal and red. Chelicerae dark brown, with three promarginal and two retromarginal teeth. Labium and endites ash brown, longer than wide. Sternum yellowish. Eyes: AER slightly recurved, PER wider than AER and almost straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.14, ALE 0.14, PME 0.14, PLE 0.12, AME���AME 0.09, AME���ALE 0.06, PME���PME 0.24, PME���PLE 0.17, MOQL 0.24, MOQA 0.33, MOQP 0.46. Legs light yellow, without distinct color markings. Leg formula: IV, II, I, III; leg measurements: I 5.93 (1.54, 2.49, 1.19, 0.72), II 6.82 (1.86, 2.79, 1.44, 0.74), III 5.43 (1.61, 1.90, 1.41, 0.51), IV 7.73 (2.14, 2.67, 2.33, 0.63). Opisthosoma (Figs 12 D���E) lanceolate, dorsal scutum indistinct, furnished with a thick tuft of hairs anteriorly; dorsum yellowish-white, posteriorly with dark purplish coloring which is formed into three longitudinal and linear markings; venter light yellow. Spinnerets dark. Palp (Figs 11 A���E): RTA dark, small but strong, digitiform, sharply pointed; bulb nearly pyriform, slightly excavated on prolatero-apical side to accommodate embolus; sperm duct indistinct; embolus spiniform, originated at 8���9 o��� clock position in prolateral view, its tip slightly overpasses the genital bulb; conductor with a slightly sclerotized and beak-shaped apex, its base part membranous; tegular apophysis large and heavily sclerotized, crescent-shaped in ventral view. Female (paratype IZCAS Ar 34711): Total length 5.13; prosoma 2.30 long, 1.74 wide; opisthosoma 2.77 long, 1.45 wide. General color darker than in male (Figs 12 F���G). Eye sizes and interdistances: AME 0.10, ALE 0.13, PME 0.12, PLE 0.11, AME���AME 0.04, AME���ALE 0.07, PME���PME 0.23, PME���PLE 0.15, MOQL 0.34, MOQA 0.32, MOQP 0.47. Legs yellowish-white, without distinct color markings. Leg formula: IV, II, I, III; leg measurements: I 4.77 (1.18, 2.08, 0.96, 0.56), II 5.01 (1.20, 2.10, 1.04, 0.66), III 3.93 (1.05, 1.45, 1.02, 0.41), IV 6.22 (1.27, 2.36, 1.85, 0.74). Epigyne (Figs 12 A���C): Epigynal plate slightly wider than long, margin not rebordered; atrium large, anterior atrial margin more or less ���M��� shaped; atrium about three times wider than long, blocked by mating plug before cleaning; two copulatory openings located at basolateral atrial borders; spermathecae consisting of bean-shaped proximal part and digitiform distal part; bursae globular and translucent, surface wrinkled and ribbed. Distribution. Presently known only from Xishuangbanna., Published as part of Yu, Hao & Li, Shuqiang, 2019, On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China, pp. 201-230 in Zootaxa 4679 (2) on pages 215-216, DOI: 10.11646/zootaxa.4679.2.1, http://zenodo.org/record/3772397, {"references":["Dankittipakul, P., Tavano, M. L., Chotwong, W. & Singtripop, T. (2012) New synonym and descriptions of two new species of the spider genus Clubiona Latreille, 1804 from Thailand (Araneae, Clubionidae). Zootaxa, 3532, 51 - 63. https: // doi. org / 10.11646 / zootaxa. 3532.1.4","Dankittipakul, P. & Singtripop, T. (2008) Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand. Zootaxa, 1747 (1), 34 - 60. https: // doi. org / 10.11646 / zootaxa. 1747.1.2","Yu, H., Zhang, J. S. & Chen, J. (2017 a) Checklist of the Clubiona japonica - group spiders, with the description of a new species from China (Araneae, Clubionidae). ZooKeys, 715, 1 - 16. https: // doi. org / 10.3897 / zookeys. 715.14645"]}

Clubiona subasrevida sp. nov. Figs 17���18 Type material. Holotype: ♂ (IZCAS Ar 34717), CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, Secondary tropical montane evergreen broad-leaved forest (N21��57.809���, E101��12.173���, 888 m), 4 August 2007, leg. G. Zheng (Fogging-CBIII). Paratypes: CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve: 1♀ (IZCAS Ar 34718), same data as holotype; 1 ♂ (IZCAS Ar 34719), Secondary tropical montane evergreen broad-leaved forest (N21��54.813���, E101��12.634���, 876 m), 5 August 2007, leg. G. Zheng (Fogging-CBIV). Etymology. The specific name is a Latin substantive referring to its similarity to Clubiona asrevida Ono, 1992, being a combination of the preposition sub (near) and the epithet of that species. Diagnosis. Males of Clubiona subasrevida sp. nov. resemble those of C. asrevida (Ono 1992: 124, figs 5���7) in having similarly shaped bulb with sinuate sperm duct, and triangular retrolateral tibial apophysis, but differ by: (1) embolus with a sub-basal torsion, placed distally in relation to tegular hump (Figs 17 A���B, D) (embolus aligned clockwise along the tegular distal margin in C. asrevida); (2) tip of embolus extending approximately two-thirds length of tegulum (Figs 17B, D���E) (relatively shorter tip in C. asrevida). Females are similar to those of C. asrevida in the general shape of vulva (Ono 1994: 81, figs 19���22), but can be recognized by the contiguous copulatory openings (Figs 18 A���B) (copulatory ducts separated in C. asrevida). Description. Male (holotype): Total length 2.52; prosoma 1.14 long, 1.01 wide; opisthosoma 1.42 long, 0.83 wide. Prosoma (Figs 18 E���F), oval, with relatively broad ocular area, widest between coxae II and III, in profile slightly raised, highest just in front fovea; integument smooth, covered with numerous fine hairs. Carapace orange, pars cephalica darker in ocular area, without distinct colour pattern. Chelicerae protruding and coloured as ocular area, with seven promarginal and five retromarginal teeth. Labium and endites colored as carapace, longer than wide. Sternum light yellow. Eyes: in dorsal view, anterior eye row slightly recurved, posterior eye row slightly procurved, PER wider than AER. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.11, ALE 0.09, PME 0.09, PLE 0.10, AME���AME 0.11, AME���ALE 0.04, PME���PME 0.27, PME���PLE 0.04, MOQL 0.11, MOQA 0.20, MOQP 0.36. Legs yellowish, without distinct color markings. Leg formula: IV, I, II, III; leg measurements: I 2.73 (0.77, 1.18, 0.58, 0.21), II 2.24 (0.70, 0.77, 0.50, 0.27), III 1.87 (0.45, 0.76, 0.45, 0.21), IV 3.03 (0.90, 1.06, 0.77, 0.30). Opisthosoma (Figs 18 E���F) oval, dorsally yellowish-white with two pairs of muscular depressions; all spinnerets without distinct color marking. Palp (Figs 17 A���E): tibia short, with single retrolateral apophysis, RTA broad, flat and triangular, with a sharp tip; genital bulb elongated, sperm duct sinuous; tegular hump represented by a flange; embolic base smooth, represented by a large sclerite, inserted at approximately 11 o���clock of tegulum; embolus distinctly long, flagelliform, slender, bent 90 degrees ventrally, forming a loop-shaped torsion; embolic tip stretched proximally on groovelike conductor, extending basad more than two-thirds length of tegulum; conductor area relatively large, for approximately two-thirds the length of tegulum. Female (paratype IZCAS Ar 34718): Total length 2.81; prosoma 1.27 long, 0.96 wide; opisthosoma 1.42 long, 0.92 wide. Not strikingly different from males but slightly lighter in color (Figs 18 G���H). Eye sizes and interdistances: AME 0.10, ALE 0.07, PME 0.07, PLE 0.06, AME���AME 0.10, AME���ALE 0.03, PME���PME 0.22, PME���PLE 0.04, MOQL 0.08, MOQA 0.18, MOQP 0.29. Leg formula: IV, I, II, III; leg measurements: I 2.70 (0.77, 0.98, 0.65, 0.31), II 2.23 (0.60, 0.97, 0.40, 0.25), III 1.82 (0.49, 0.83, 0.30, 0.20), IV 2.25 (0.36, 0.52, 0.80, 0.57). Epigyne (Figs 18 A���D): Epigynal plate disc-shaped, distinctly wider than long, posterior margin heavily sclerotized; spermathecae and copulatory ducts are prominently through epigynal plate in ventral view; two copulatory openings large and conjoined, located at posterior portion of epigynal plate; hyaline copulatory ducts thick and straight, close together, extending above anterior level of spermathecae, and then oblique descending ventrally, connected laterally to bursae; spermathecae sub-globular, connected to digitiform spermathecal heads; bursae globular, situated anteriorly, much smaller than spermathecae; fertilization ducts acicular, membranous, located on dorsal surface of spermathecae. Distribution. Presently known only from the type locality., Published as part of Yu, Hao & Li, Shuqiang, 2019, On further species of the spider genus Clubiona Latreille, 1804 (Araneae, Clubionidae) from Xishuangbanna Rainforest, southwestern China, pp. 201-230 in Zootaxa 4679 (2) on pages 221-226, DOI: 10.11646/zootaxa.4679.2.1, http://zenodo.org/record/3772397, {"references":["Ono, H. (1992) Two new species of the families Araneidae and Clubionidae (Arachnida, Araneae) from Taiwan. Bulletin of the National Museum of Nature and Science Tokyo (A), 18, 121 - 126.","Ono, H. (1994). Spiders of the genus Clubiona from Taiwan (Araneae: Clubionidae). Acta Arachnologica, 43, 71 - 85. https: // doi. org / 10.2476 / asjaa. 43.71"]}

A further study of the spider genus Clubiona Latreille, 1804 from Xishuangbanna, Yunnan, China is presented. A total of ten species, eight of which new to science, are here addressed, raising from 18 to 27 the number of species of the genus known to the area. Clubiona didentata Zhang & Yin, 1998 is redescribed based on new specimens from type locality, with the first description of the female. C. kai Jäger & Dankittipakul, 2010 is newly recorded from China, and the female is reported for the first time. Seven of the new species here described belongs to four species groups: C. cochlearis sp. nov. (♂) and C. tiane sp. nov. (♂) from the C. corticalis species-group; C. tongi sp. nov. (♂♀) from the C. hystrix species-group; C. yueya sp. nov. (♂♀) and C. zhanggureni sp. nov. (♂) from the C. japonica species-group; and C. bicornis sp. nov. (♂) and C. subasrevida sp. nov. (♂♀) from the C. trivialis species-group. The eighth new species, C. jiandan sp. nov. (♂♀) is not readily assignable to any of the existing species groups.

Clubiona didentata Zhang & Yin, 1998 Figs 5–6 Clubiona didentata Zhang & Yin, 1998: 11, figs 6–8 Material examined. CHINA: Yunnan: Xishuangbanna, Mengla County, Menglun Town, Menglun Nature Reserve, leg. G. Tang and Z.Y. Yao: 1♂, G213 roadside, lowevergreen forest (N21º53.794′, E101º17.152′, 594 m), 27 Novem- ber 2009 (Tang-Yao _ No. 29); 2♀, G213 roadside, Bamboo plantation (N21º53.640′, E101º16.940′, 580 m), 3 De- cember 2009 (Tang-Yao _ No. 40); leg. H. Yu and Z.G. Chen: 1♂, 3♀, XTBG, Rubber-Tea Plantation (N21º55.239′, E101º15.854′, 572 m), 28 July 2018 (Yu _20180728_Searching-26). Diagnosis. Males of this species resembles those of C. kai Jäger & Dankittipakul, 2010 (Jäger & Dankittipakul 2010: 25, figs 4–6; Figs 7 A–E) in having a tegular apophysis, usually absent in other species of the corticalis group, but differ by: (1) the basal branch of embolus bulky and hidden in ventral view (Fig. 5D) (relatively thinner and clearly visible in C. kai; Fig. 7D); (2) embolus slightly curved (Figs 5 C–D) (embolus twisted in C. kai; Figs 7 C–D); (3) conductor large and heavily sclerotized (Figs 5 D–E) (conductor small and partly membranous in C. kai; Figs 7 D–E); (4) tegular apophysis large and triangular, pointing distally (Fig. 5D) (tegular apophysis small and teethshaped, pointing prolatero-distally; Fig. 7D). Females also resemble those of C. kai in having an atrial membrane and similar vulvae (Figs 6 A–C; Figs 8 A–C), but can be recognized by the atrial membrane nearly tongue-shaped (Figs 6 A–B) (disc-shaped in C. kai; Figs 8 A–B), bursae surface wrinkled (Fig. 6C) (bursae surface smooth in C. kai; Fig. 8C). Description. Male: Total length 3.48; prosoma 1.42 long, 0.95 wide; opisthosoma 1.89 long, 0.93 wide. Prosoma (Figs 6 D–E), ovoid in dorsal view, pars cephalica slightly narrowed, widest between coxae II and III; in profile highest just behind longitudinal fovea, gradually sloping towards pars cephalica; integument smooth, clothed with short fine hairs. Carapace yellowish, without distinct color pattern; fovea dark. Chelicerae light orange, with five promarginal and three retromarginal teeth. Labium and endites light orange, longer than wide. Sternum yellowish white. Eyes: AER slightly recurved, PER wider than AER and straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.06, ALE 0.09, PME 0.10, PLE 0.11, AME–AME 0.02, AME–ALE 0.03, PME–PME 0.15, PME–PLE 0.07, MOQL 0.13, MOQA 0.10, MOQP 0.25. Legs light yellow, without distinct color markings. Leg formula: IV, II, III, I; leg measurements: I 2.76 (0.60, 1.29, 0.51, 0.42), II 3.72 (1.25, 1.30, 0.82, 0.44), III 3.25 (0.98, 1.13, 0.73, 0.35), IV 4.68 (1.34, 1.50, 1.38, 0.46). Opisthosoma (Figs 6 D–E) oval, dorsally yellowish-white with a lengthwise light orange heart mark, reaching posterior half; muscular depression inconspicuous; with conspicuous anterior hair tuft; venter white. Spinnerets light yellow. Palp (Figs 5 A–E): tibia short, only with retrolateral apophysis; RTA wide and thumb-shaped, with blunt tip; bulb oval and distinct expanded, sperm duct indistinct; embolus dagger-like, with a bulky base and a sharp tip; conductor large and heavily sclerotized, linguiform; tegular apophysis large, shape like a equilateral triangle, it’s apex sharp and pointing distally. Female: Total length 3.72; prosoma 1.18 long, 0.92 wide; opisthosoma 2.04 long, 1.10 wide. General color lighter than in male (Figs 6 F–G). Eye sizes and interdistances: AME 0.09, ALE 0.10, PME 0.13, PLE 0.11, AME– AME 0.05, AME–ALE 0.06, PME–PME 0.21, PME–PLE 0.05, MOQL 0.14, MOQA 0.13, MOQP 0.27. Legs yellowish-white, without distinct color markings. Leg formula: IV, II, I, III; leg measurements: I 3.26 (0.94, 1.35, 0.61, 0.34), II 3.59 (1.05, 1.41, 0.73, 0.40), III 3.11 (0.89, 0.93, 0.80, 0.35), IV 4.41 (1.21, 1.52, 1.22, 0.45). Epigyne (Figs 6 A–C): Epigynal plate slightly wider than long, margin not rebordered; atrium small, with tongue-shaped membrane on anterior margin; spermathecae and bursae prominently visible through epigynal plate in ventral view; copulatory openings small, located at basolateral atrial borders; spermathecae situated anteriorly, with bean-shaped proximal part and acicular distal part; fertilization ducts short and curved; reniform bursae situated posteriorly, translucent, surface wrinkled. Distribution. Presently known only from Xishuangbanna.

Pristidia cervicornuta Yu, Zhang & Chen, 2017 is redescribed based on new material from the type locality, Diaoluo Mountains of Hainan Island, China. The female is described and illustrated for the first time. In addition, this paper further illustrates the male, and provides a supplementary description.

Yang, Daxing, Zhou, Guchun, Yang, Maofa, Peng, Xianjin (2019): Redescription of Clubiona gongi Zhang, Yin, Bao & Kim, 1997 (Araneae: Clubionidae) with the first description of the male. Zootaxa 4658 (1): 183-188, DOI: https://doi.org/10.11646/zootaxa.4658.1.10

Haddad, Charles R., Marusik, Yuri M. (2019): Clarifying the taxonomic status and distributions of the spider species collected during the Leonhard Schultze expeditions in western and central southern Africa (Arachnida: Araneae). Zootaxa 4608 (3): 451-483, DOI: https://doi.org/10.11646/zootaxa.4608.3.3

Clubiona corticalis (Walckenaer, 1802) MATERIAL. ��� La Salce, mixed forest of Populus and Pinus, 1406 m, 01.VII.2017, Isaia, Mammola & Milano leg., 1♀, coll. MI. CHOROTYPE. ��� TUE. MACROHABITAT. ��� Mixed forests. NOTE. ��� According to Nentwig et al. (2018), this species occurs preferably below 1200 m a.s.l. Our finding at 1400 m a.s.l. extends the current known altimetric distribution range of the species., Published as part of Milano, Filippo, Mammola, Stefano, Rollard, Christine, Leccia, Marie-France & Isaia, Marco, 2019, An inventory of the spider species of Barcelonnette (France), with taxonomic notes on Piniphantes agnellus n. comb. (Araneae, Linyphiidae), pp. 29-58 in Zoosystema 41 (4) on page 34, DOI: 10.5252/zoosystema2019v41a4, http://zenodo.org/record/3718482, {"references":["NENTWIG W., BLICK T., GLOOR D., HANGGI A. & KROPF C. 2018. - Spiders of Europe. Version 05.2018. https: // doi. org / 10.24436 / 1"]}