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Oil sands mining in northeastern Alberta, Canada, generates tailings containing sand, silt, clay, water, and residual bitumen with hydrocarbons. The impact of tailings on revegetation is a major environmental concern and poses a significant land reclamation challenge. Oil sands companies have recently developed technologies that use thickeners in combination with carbon dioxide to produce non‐segregating tailings (NST), to accelerate the consolidation of tailings while sequestering greenhouse gases. Effects of these tailings on plant re‐establishment have yet to be determined. We investigated the impact of NST on biomass and physiology of 20 boreal woody plant species grown in environmentally controlled growth rooms. The seedlings were grown for 8 weeks in NST capped with boreal forest topsoil or peat‐mineral soil mix or petroleum coke in 50‐cm‐long polyvinyl chloride pipes. We found that the biomass of balsam fir, beaked hazelnut, blueberry, green alder, Labrador tea, lowbush cranberry, paper birch, and raspberry was severely reduced by NST, while the effects on balsam poplar, Bebb's willow, chokecherry, dogwood, saskatoon, and white spruce were relatively weak. This was also the case for jack pine, which showed poor survival in NST. The negative impact of NST on plants can be largely explained by elevated sodium and decreased foliar nutrient concentrations. The impact of NST capping with petroleum coke on plant growth was non‐significant. Differences among species in their survival and biomass responses to NST appeared to be partly a reflection of their natural habitats. Findings from this study can help guide revegetation oil sands reclamation strategies. [ABSTRACT FROM AUTHOR]

Retention harvesting is advocated as an alternative to intensive timber harvesting, such as clear‐cutting, to better maintain or facilitate recovery of biodiversity and other ecological values in managed forests. However, it is not clear how long the benefits of retention harvests persist.We investigated responses of understory vascular plant cover, richness, diversity (inverse Simpson index) and composition to a gradient in dispersed retention (2% [clear‐ cut], 10%, 20%, 50% and 75% retention; unharvested reference [100% retention]) at 3, 6, 11, and 17 years after harvest, in four boreal mixedwood forest types (deciduous (broadleaf)‐dominated, deciduous‐dominated with conifer understory, mixed, and conifer‐dominated) in western Canada.Understory cover and richness tended to increase in the short‐term (3 years), peaked at 6–11 years with differences following the gradient of harvesting intensity, then plateaued or declined in the second decade (17 years), by which time there were minimal or no differences among harvesting levels, including the reference. Responses for diversity were minimal. In contrast, composition varied along the gradient of harvesting intensity and showed little recovery towards the unharvested condition over the 17‐year period. Generally, for plant community composition, clear‐cut and lower retention treatments (10%, 20%) were similar to one another but differed from the higher retention and unharvested reference treatments.Synthesis and applications: Retention harvests can moderate the negative impacts of harvesting and facilitate the recovery of biodiversity. Our results suggest that for the cover, richness and diversity of understory vascular plants, this moderating influence is weak and short‐lived. However, higher levels of retention can temper changes in understory composition relative to the unharvested forest, but full recovery is likely to be slow and will be complicated by post‐harvest regeneration dynamics. [ABSTRACT FROM AUTHOR]

Questions: Edge influence on forest biodiversity is an important environmental effect associated with habitat fragmentation, but extrapolating the influence of edges across the broader landscape has been difficult, especially for situations where multiple edges exist in close proximity. We asked whether there were differences in edge effects between two types (3 m vs 8 m width) of low‐severity linear disturbance (seismic lines) and whether there were interactions of edge effects when seismic lines occur in dense networks; that is, do multiple narrow seismic lines have a stronger or weaker edge influence than a single narrow seismic line. Location: Treed peatlands in northeastern Alberta, Canada. Methods: Seismic lines are created during oil and gas exploration and are responsible for dissection of boreal forests in western Canada. We sampled vascular plants along transects perpendicular to seismic lines in moderate‐rich and poor treed fens. We used the "Randomization Test of Edge Influence" (RTEI) to calculate the magnitude and distance of edge effects and then compared these between narrower (3 m) versus wider (8 m) lines and between single narrow lines versus multiple narrow lines (parallel and ~50 m apart). Results: In moderate‐rich fens, we found a positive edge influence on understorey diversity from both wide and narrow seismic lines. We also found a weakening edge interaction on diversity, that is, single narrow seismic lines had a stronger edge influence on diversity than did multiple narrow seismic lines. In treed poor fens, multiple narrow seismic lines had a negative edge effect on tree density, understorey abundance, richness, and composition. In addition, we found strengthening edge interactions in treed poor fens on tree density, graminoid cover, and understorey composition. Conclusions: Even narrow linear disturbances, such as seismic lines, can have significant edge effects and these are exacerbated when lines occur in dense networks. [ABSTRACT FROM AUTHOR]

• Depth-to-water index can be used to identify understory distribution. • Conifer stands had higher understory diversity and abundance on drier sites. • Mixedwood and broadleaf stands had higher understory abundance on drier sites. • Depth-to-water index can be useful for guiding conservation. For the purpose of informing biodiversity conservation efforts in managed landscapes, we explored whether and how understory plant communities (abundance, diversity, composition) were related to a topographic moisture index, called depth-to-water, in the boreal mixedwood forests of northwestern Alberta. Depth-to-water is an index of relative site moisture derived from the Wet Areas Mapping tool using a fine-scale digital elevation model based upon remotely-sensed lidar (light detection and ranging) data. Sample plots were placed along the depth-to-water moisture gradient in three forest types: conifer-dominated, mixedwood, and broadleaf -(deciduous) dominated. Understory vascular plant diversity, abundance, and composition were measured for each plot. We found understory attributes were related to the depth-to-water index with the relationships varying among forest types. In coniferous stands, diversity and abundance (cover) were higher on drier sites. In broadleaf and mixedwood stands, understory abundance was higher on drier sites, but diversity was not related to the depth-to-water index. Lastly, composition was significantly, but weakly, related to the depth-to-water index in all three forest types. Our study shows that this moisture index, based on remotely-sensed data, can be used to characterize patterns in understory vascular plant communities; hence it can be useful for identifying areas of particular interest for conservation or management. [ABSTRACT FROM AUTHOR]

Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

Site moisture is an important component of the forest landscape for maintaining biodiversity, including forest‐floor bryophytes. However, little is known about its role in shaping understorey responses to harvesting.We investigated the influence of site wetness, determined using a remotely sensed, topographic depth‐to‐water (DTW) index, on responses of bryophyte cover, richness, diversity and composition to variable retention harvesting (comparing: 2% [clear‐cut], 20% and 50% dispersed green tree retention and uncut controls [100% retention]) in three boreal forest cover‐types (broadleaf, mixed and conifer forests) in western Canada. The DTW index provides an approximation of DTW at or below the soil surface and was derived from wet‐areas mapping based on discrete Airborne Laser Scanning data acquired over an experimentally harvested landscape located in north‐western Alberta, Canada.The effectiveness of leaving retention (vs. clear‐cutting) for conserving bryophyte communities depended on site wetness, as indicated by DTW, with the specifics varying among forest types. In broadleaf forests, bryophyte cover and richness were generally low and not much affected by harvesting but drier sites had higher richness and a few more unique species. In mixed and conifer forests, leaving retention (vs. clear‐cutting) on wetter (vs. drier) sites was more effective for conserving bryophyte cover, wetter sites had higher total species richness and more species were exclusive to wetter sites.Synthesis and applications. Site wetness, as indicated using the remotely sensed topographic site wetness index "depth‐to‐water" mediates bryophyte responses to variable‐retention harvests. Specifically, our results suggested that in conifer and mixed forests it would be more beneficial to target wetter sites for retention patches or dispersed retention whereas in broadleaf forests there might be a slight advantage to targeting drier sites. Our study demonstrates that this tool could be used to inform management decisions around leaving dispersed or patch retention. Site wetness, as indicated using the remotely sensed topographic site wetness index "depth‐to‐water" mediates bryophyte responses to variable‐retention harvests. Specifically, our results suggested that in conifer and mixed forests it would be more beneficial to target wetter sites for retention patches or dispersed retention whereas in broadleaf forests there might be a slight advantage to targeting drier sites. Our study demonstrates that this tool could be used to inform management decisions around leaving dispersed or patch retention. [ABSTRACT FROM AUTHOR]

Given the habitat moisture (air humidity or soil moisture) preferences of many forest bryophytes, we explored whether the depth-to-water (DTW) index, derived from remotely sensed Light Detection and Ranging (LiDAR) data, was related to fine-scale patterns of spatial variation in bryophyte abundance, diversity, and composition. The goal was to assess the utility of the topographic DTW index as a tool to decipher trends in bryophyte assemblages along a site wetness gradient in the boreal mixedwood forest. Discrete Airborne Laser Scanning (ALS) data were acquired over the entire Ecosystem Management Emulating Natural Disturbance (EMEND) experimental site located in northwestern Alberta, Canada (56° 46′ 13″ N, 118° 22′ 28″ W), based on which we calculated a mathematical index of approximate depth to water at or below the soil surface at 1 m resolution using the Wet-Areas Mapping model. Bryophytes (mosses and liverworts) were sampled in permanent sample plots in unmanaged forest stands of varying dominant canopy tree composition. The relationships between DTW and bryophyte cover, richness, diversity, and composition in broadleaf (deciduous)-, mixed, and conifer-dominated boreal forest stands were analyzed using linear mixed-effect models and multivariate analyses. Bryophyte cover was highest in conifer-dominated forest, which occupied the wetter end of the DTW gradient, followed by mixed forest, whereas broadleaf forest, which occupied the drier end of the DTW gradient, had the lowest cover but highest bryophyte diversity. Bryophyte cover in conifer-dominated forests was positively related to site moisture (negatively related to the DTW index). In contrast, bryophyte species richness and diversity were negatively related to site moisture (increased at higher DTW values) in all forest types. DTW explained significant variation in bryophyte species composition in mixed forests, while indicator species analysis identified species with preferences for wet, moist, and dry site conditions in each forest type. Our results corroborate the importance of site moisture as a driver of bryophyte assemblages but, interestingly, there were important differences among forest types, which themselves are distributed across a gradient of site moisture. Our study demonstrates the utility of the topographic DTW index for understanding fine-scale (plot-level) variation in bryophyte assemblages in forested landscapes. [ABSTRACT FROM AUTHOR]

Given a seed source, the quality of available substrates is a key factor in determining the success of white spruce ( Picea glauca (Moench) Voss) natural regeneration. We examined the influence of substrate and competing vegetation on survival and growth of natural regeneration of white spruce up to 4 years following harvesting in deciduous-dominated upland boreal mixedwood sites. Feather moss, thick soil surface organic layers, litter, and solid wood were poor substrates for establishment. Early successional mosses establishing on mineral soil, thin organics, and rotten wood were generally favourable microsites but were not highly available on postharvest sites. Mineral soil substrates were not as suitable as expected, likely because on a postlogged site, they are associated with unfavourable environmental characteristics (e.g., low nutrient availability, exposure). There was some evidence that survival and growth of seedlings were improved by surrounding vegetation in the first years, but heavy competing vegetation had a negative impact on older seedlings. Burial by aspen litter greatly increased seedling mortality, especially when combined with a brief period of submergence due to heavy spring snowmelt. The results provide insight into conditions under which natural regeneration could be an option for establishing white spruce following harvesting of deciduous-dominated boreal mixedwood forests. [ABSTRACT FROM AUTHOR]

Abstract: Variable‐retention harvest is widely recognized as an alternative to more intensive methods such as clear‐cutting. However, present information is inadequate to judge the impact of variable retention on biodiversity of indigenous forest organisms intolerant of canopy removal, such as forest‐inhabiting bryophytes. We examined how bryophyte species cover, richness, diversity and composition change with time in response to a broad range of dispersed retention harvest treatments (2% [clear‐cut], 10%, 20%, 50%, 75% retention of original basal area) contrasted with uncut controls [100% retention]) in broadleaf deciduous, mixedwood and conifer‐dominated boreal forests in North West Alberta, Canada. Bryophytes were studied in 432 permanent sample plots within 72 compartments before harvest and at 3, 6 and 11 years after harvest. Clear‐cut and lower (10% and 20%) retention levels resulted in lower cover and richness of bryophytes than in unharvested control compartments in mixed and conifer‐dominated forests, but less so in deciduous‐dominated forests, which generally supported low cover and richness. Species composition in each forest type varied along the gradient of harvesting intensity; clear‐cuts and lower levels of retention supported similar composition, as did control plots and those representing higher retention levels. Over time, the retention harvest treatments became more similar to uncut controls. Synthesis and applications. Variable‐retention harvests can better maintain bryophyte biodiversity in managed boreal mixedwood forests, as compared to clear‐cuts. We found the efficacy of retention harvest scaled with harvest intensity. Higher levels of retention better moderated the negative impacts of harvesting on bryophyte assemblages across all forest types. Our results suggest, however, that even 10% retention will facilitate faster post‐harvest recovery of bryophytes, as compared to clear‐cutting. [ABSTRACT FROM AUTHOR]

The extent to which past states influence present and future ecosystem characteristics (ecosystem memory (EM)) is challenging to assess because signals of past ecological conditions fade with time. Using data about seven different taxa, we show that ecological gradients initiated by wildfires up to three centuries earlier affect biotic recovery after variable retention harvest in the boreal mixedwood forest. First, we show that fire history over the last 300 years is reflected in pre-harvest species-specific stand basal area (BA), with longer times since high severity fire associated with proportionally higher BA of shade-tolerant softwood species than shade-intolerant hardwoods. Second, using patterns in the BA of pre-harvest tree species we link fire history to species composition of pre-harvest assemblages of bryophytes, herbs, shrubs, regenerated trees, songbirds, spiders and carabid beetles. Finally, we use variance partitioning to compare the importance of species-specific pre- versus post-harvest BA for explaining the structure of these seven biotic assemblages two, five and ten years after harvest. We detected persistent significant effects of pre-harvest BA in all post-harvest biotic assemblages up to ten years after harvest. Pre-harvest BA was more strongly associated with early post-harvest understory plant and carabid beetle assemblages than was post-harvest BA, but the opposite was true for spiders, songbirds and regenerated trees. EM effects were detected two, five and ten years after harvest but temporal patterns varied according to taxa. Thus, EM of fire history can persist at least ten years after variable retention harvest and such effects appear to be stronger for understory plants than for animals. We conclude that management of biological legacies to increase post-disturbance EM will increase overall resilience and sustainability of these mixedwood forests. [ABSTRACT FROM AUTHOR]

As boreal forests rapidly warm due to anthropogenic climate change, long‐term baseline community data are needed to effectively characterize the corresponding ecological changes that are occurring in these forests. The combined seasonal dynamics (SEADYN) and annual dynamics (ANNDYN) data set, which documents the vegetative changes in boreal forests during the snow‐free period, is one such source of baseline community data. These data were collected by George H. La Roi and colleagues in Alberta, Canada from 1980 to 2015 within permanent sampling plots established in the Hondo‐Slave Lake area (eight stands; 1980–2015) in central Alberta and the Athabasca Oil Sands (AOS) region (17 stands; 1981–1984) near Fort McMurray in northeastern Alberta. Various data were collected, with temporal and spatial coverage differing by data set. These data sets include, but are not limited to, cover of each identified vascular plant and bryoid (moss, liverwort, and lichen) species; forest mensuration; forest litter production; and soil temperature and moisture. Notably, permanent sampling plots were set up as a grid, which will facilitate analyses of spatial relations. These data can be used to analyze long‐term changes in seasonal dynamics and succession within boreal forest communities and serve as a baseline for comparison with future forest conditions in unmanaged, managed, and reclaimed forests. Data are released under a CC‐BY license; please cite this data paper when using the data for analyses. [ABSTRACT FROM AUTHOR]

Aims: Reclamation following oil sands mining in northeastern Alberta (Canada) creates adverse reforestation soil conditions, including extreme pH values. We elucidated pH tolerance limits of boreal plant species and how pH affects nutrient uptake in these plants. Methods: We measured growth, gas exchange, and foliar nutrient concentration of 15 common northern boreal forest plants after eight weeks exposure to root zone pH ranging from 5.0 to 9.0. Cluster analyses were used to group these species based on their pH responses. Results: Based on their growth and gas exchange responses to pH, the 15 plant species could be divided into five groups, each of which contained species that commonly co-occur in particular boreal forest site types. For the foliar nutrient responses to pH, the 15 species could be grouped into only two categories; both showed decreases in foliar N, P, Fe and Zn concentration with increasing pH, with a more pronounced effect on the group that included trembling aspen, paper birch and chokecherry. Conclusions: The evidence of differential adaptation to pH by habitat type suggests the importance of soil pH as a factor affecting boreal plant species distribution and could be helpful for selection of species suitable for reclamation of sites with altered soil pH. [ABSTRACT FROM AUTHOR]

Although anthropogenic edges are an important consequence of timber harvesting, edges due to natural disturbances or landscape heterogeneity are also common. Forest edges have been well studied in temperate and tropical forests, but less so in less productive, disturbance-adapted boreal forests., We synthesized data on forest vegetation at edges of boreal forests and compared edge influence among edge types (fire, cut, lake/wetland; old vs. young), forest types (broadleaf vs. coniferous) and geographic regions. Our objectives were to quantify vegetation responses at edges of all types and to compare the strength and extent of edge influence among different types of edges and forests., Research was conducted using the same general sampling design in Alberta, Ontario and Quebec in Canada, and in Sweden and Finland. We conducted a meta-analysis for a variety of response variables including forest structure, deadwood abundance, regeneration, understorey abundance and diversity, and non-vascular plant cover. We also determined the magnitude and distance of edge influence (DEI) using randomization tests., Some edge responses (lower tree basal area, tree canopy and bryophyte cover; more logs; higher regeneration) were significant overall across studies. Edge influence on ground vegetation in boreal forests was generally weak, not very extensive (DEI usually < 20 m) and decreased with time. We found more extensive edge influence at natural edges, at younger edges and in broadleaf forests. The comparison among regions revealed weaker edge influence in Fennoscandian forests., Synthesis. Edges created by forest harvesting do not appear to have as strong, extensive or persistent influence on vegetation in boreal as in tropical or temperate forested ecosystems. We attribute this apparent resistance to shorter canopy heights, inherent heterogeneity in boreal forests and their adaptation to frequent natural disturbance. Nevertheless, notable differences between forest structure responses to natural (fire) and anthropogenic (cut) edges raise concerns about biodiversity implications of extensive creation of anthropogenic edges. By highlighting universal responses to edge influence in boreal forests that are significant irrespective of edge or forest type, and those which vary by edge type, we provide a context for the conservation of boreal forests. [ABSTRACT FROM AUTHOR]

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When species are introduced into new regions, there is great uncertainty whether the trait differences of the introduced species or regional factors, such as climate or edaphic properties, will serve as the dominant control of ecosystem properties or processes. In this study, we examined whether the introduction of Pinus contorta Douglas ex Loudon into Sweden has altered forest floor properties and processes or whether these properties are more strongly controlled by regional factors. We compared forest floor pH, potential N mineralization rates, bulk density, litter and forest floor depths, C and N concentrations and pool sizes, C:N ratios, and soil microbial communities using substrate-induced respiration and phospholipid fatty acid analysis among stands of introduced P. contorta (SwPc), native Swedish Pinus sylvestris L. (SwPs), and native Canadian P. contorta (CaPc). For most forest floor properties (pH, net NH4+ mineralization, bulk density, N mass, and the microbial phospholipid fatty acid community structure), SwPc sites were more similar to SwPs than to CaPc, whereas litter and forest floor depth were significantly higher in SwPc than the two other forest types. Our findings suggest that regional factors exerted a stronger control on most forest floor properties and processes than did species differences between the two Pinus species for the regions we studied. [ABSTRACT FROM AUTHOR]

Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

We explored interactions among plant growth forms in the understory of mature boreal mixedwood forests in western Canada by investigating the competitive influence of erect shrubs on herbs (forbs and grasses). We established 10 pairs of plots; all erect shrubs were removed in one plot of each pair (removals) and left intact in the other plot (controls). Two years later, we harvested all aboveground biomass of the herbaceous layer (herb biomass: this included graminoids, forbs, trailing shrubs, and species with a woody base but not woody stems) from the 20 plots. We tested for significant differences in understory species biomass and composition between control and removal plots and examined the influence of 25 environmental factors on species composition of the herbaceous layer. Competition intensity was measured by the natural logarithm of response ratio (ln RR) index based on herb biomass. After erect shrub removal, there was a significant increase in herb biomass, mostly due to an increase of the most common species (e.g., Cornus canadensis Linnaeus, Linnaea borealis Linnaeus). The values of competition intensity (ln RR) varied among herb species but were, overall, positive, indicating a release from competition following shrub removal. Composition of the herbaceous layer was significantly different between removal and control plots and was also significantly related to seven environmental factors, which explained 40% of the variation in composition. Our study suggests that there is asymmetric competition for light between erect shrub and herb species in boreal ecosystems. [ABSTRACT FROM AUTHOR]

Epiphytic mosses and liverworts contribute substantially to the bryophyte diversity of circumpolar boreal forests but are susceptible to altered growing conditions after forest harvesting. Management practices that retain some trees after harvest may enhance epiphyte survival; however, the effectiveness of this emerging method needs to be assessed. We examined the survival, composition, and nearest neighbour relationships of epiphytic bryophytes on trembling aspen (Populus tremuloides Michx.) across a range (10%–100%) of dispersed green-tree retention 5 years after harvest in boreal mixed-wood forest. Growth of the forest floor moss Hylocomium splendens (Hedw.) Schimp. in B.S.G. was used as an indicator of changes in moisture availability for epiphytes following harvesting. Epiphyte richness and abundance increased with canopy retention and were positively correlated with local abundance of coniferous trees. Positive associations among neighbouring species in intact forest demonstrated that interspecies relationships form naturally. However, there was a shift in species composition after harvesting and fewer interspecific associations with declining retention. These trends were accompanied by reduced Hylocomium splendens growth, which implies that moisture may be an important driver of epiphyte response. Although different levels of canopy retention were similarly capable of maintaining some epiphytes, the loss of species associated with intact forest will require consideration of alternative management practices for their conservation. Les mousses et les hépatiques épiphytes contribuent substantiellement à la diversité des bryophytes des forêts boréales circumpolaires, mais sont sensibles aux modifications des conditions de croissance, suite aux opérations forestières. Les méthodes d’aménagement qui conservent certains arbres après la récolte peuvent améliorer la survie des épiphytes; cependant, on doit évaluer l’efficacité de ces méthodes en émergence. Les auteurs ont examiné la survie, la composition et les relations avec les voisins les plus rapprochés des bryophytes épiphytes sur le peuplier faux-tremble (Populus tremuloides Michx.) pour l’ensemble d’une aire (10 % à 100 %) comportant des arbres vivants conservés et dispersés, 5 ans après la récolte dans une forêt boréale mixte. Ils ont utilisé la croissance d’une mousse du parterre forestier, l’Hylocomium splendens (Hedw.) Schimp. in B.S.G., comme indicatrice des modifications de la disponibilité de l’humidité pour les épiphytes, suite à la récolte des arbres. La richesse et l’abondance des épiphytes augmentent avec la rétention de la canopée et montrent des corrélations positives avec l’abondance localisée de conifères. Les associations positives avec les espèces avoisinantes dans la forêt intacte démontrent que des relations interspécifiques se forment naturellement. Cependant, on observe un déplacement de la composition en espèces après la récolte et moins d’associations interspécifiques avec une diminution de la rétention. Ces tendances s’accompagnent d’une réduction de la croissance de l’ Hylocomium splendens, ce qui implique que l’humidité pourrait être un agent causal important de la réaction des épiphytes. Bien que différents degrés de conservation de la canopée peuvent être également aptes à maintenir certaines épiphytes, la perte d’espèces associées aux forêts intactes nécessitera la définition de méthodes alternatives d’aménagement pour leur conservation. [ABSTRACT FROM AUTHOR]

Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

Copyright of Canadian Journal of Forest Research is the property of Canadian Science Publishing and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

Bryophyte diaspore banks are a potential source of reproductive propagules that may be able to colonize newly available habitat after forest harvesting. However, their role and the factors influencing the successful establishment of species remain poorly understood. To investigate these aspects of diaspore banks, we obtained mineral soil samples from mixed and coniferous stands of boreal mixed-wood forest in northern Alberta, Canada, which had recently experienced a range of harvesting intensities. Samples were germinated in growth cabinets under two light regimes. The composition of germinated bryophyte species was compared among forest types, harvesting intensities, and light levels, and also related to edaphic conditions and geographic location of the sample site. Germinated species composition was not related to forest type or harvesting intensity, but was associated with measured edaphic variables and geographic space, indicating that similarity in species composition reflected similarity in edaphic conditions and spatial proximity. This was partly because of spatial dependence of edaphic variables. Light intensity had a significant influence on the development of species assemblages and individual species responses. Richness and cover of acrocarpous mosses (fugitive, colonist, and shuttle life-history strategies) were significantly reduced under low light conditions, but pleurocarpous mosses (perennial strategy) were not affected. Shannon diversity and the frequency of reproduction were significantly greater with high light. Pleurocarpous mosses that are characteristic of intact forests germinated frequently, suggesting that diaspore banks may influence their recovery after disturbance. Diaspore banks are a repository of species at sites affected by forest harvesting; however, diaspore germination and establishment will be constrained by the local environment, including edaphic conditions and light intensity. [ABSTRACT FROM AUTHOR]

We compared understory vegetation composition and richness in aspen-dominated boreal mixedwood forest stands in Alberta, Canada, that had been burned by wildfire with those that burned and were subsequently salvage logged. Stands were examined at early and midsuccessional (2 and 34 years after disturbance(s), respectively) developmental stages. In comparison with wildfire stands, understory communities of early successional salvage-logged stands were characterized by greater species richness, weedy species presence, higher shrub abundance, and lower abundances of fire-specialist seed bank species. In constrained ordination, the understory community of early successional wildfire stands was related to greater canopy cover, sapling density, and moss depth, whereas that of salvage-logged stands was related to greater light, volume of downed deadwood, and litter and organic matter. Longer term effects of salvage logging on the understory community were minimal and, instead, reflected the influence of forest canopy redevelopment. In midsuccessional stands, understory composition was related to conifer density, litter cover, soil moisture, organic layer depth, tall shrub density, and bryophyte-covered microsite cover. Postfire salvage logging can have substantial short-term effects on the postfire understory plant community; in the longer term, effects will depend to a large extent on the influence of harvesting and subsequent management on canopy redevelopment. [ABSTRACT FROM AUTHOR]

We compared prefire and postfire organic-layer depths in boreal forest types (14 fires) across Canada, and examined tree recruitment as a function of depth. There was extensive within-stand variation in depth, much of it due to clustering of thinner organic layers around boles. There were no significant differences in postfire organic-layer depth among sites with different prefire forest species composition, but sites in the eastern boreal region had thicker postfire organic layers than those in the western boreal region. Mean organic-layer depth was much greater in intact stands than after fires; overall, fire reduced organic-layer depth by 60%, largely because of increases in the area of thin (<3 cm) organic layers (1% in intact stands vs. 40% in postfire stands). There was more variation in organic-layer depth within postfire than within prefire stands; notably, some areas in postfire stands were deeply combusted, while adjacent parts were only lightly combusted. We speculate that the diminished role of energy loss to latent heat around tree boles increased organic-layer consumption around tree boles. Seedlings were clustered around burned tree bases, where organic layers were thinner, and the dependence of a species on thin organic layers was an inverse function of seed size. [ABSTRACT FROM AUTHOR]

Abstract: We examined patterns of variation in richness, diversity, and composition of understory vascular plant communities in mixedwood boreal forests of varying composition (broadleaf, mixedwood, conifer) in Alberta, Canada, before and for 2 years following variable-retention harvesting (clearcut, 20 and 75% dispersed green tree retention, control). Broadleaf-dominated forests differed from mixedwood or conifer-dominated forests in that they had greater canopy cover, litter depth, soil nitrogen, warmer soils, as well as greater shrub cover, herb and shrub richness and diversity (plot scale). In contrast, conifer, and to a lesser extent mixedwood, forest had greater β diversity than broadleaf forest. Overall, mixedwood and conifer forests were similar to one another, both differed from broadleaf forest. Several species were found to be significant indicators of broadleaf forest but most of these also occurred in the other forest types. Understory composition was related to canopy composition and edaphic conditions. Variable-retention harvesting had little effect on understory cover, richness, or diversity but resulted in reduced richness and β diversity at a larger scale. The clearcut and 20% treatments affected composition in all forest types. Early successional species and those common in disturbed sites were indicators of harvesting while evergreen, shade-tolerant understory herbs were indicators of the control forest and 75% retention harvest. We conclude that it is important to maintain a range of variation in canopy composition of mixedwood forests in order to conserve the associated understory communities. The presence of conifers in these forests has a particularly important influence on understory communities. The threshold for a lifeboat effect of variable-retention harvesting is between 20 and 75% retention. Examination of richness and β diversity at a variety of scales can provide interesting information on effects of harvesting on spatial reorganization and homogenization of understory plant communities. [Copyright &y& Elsevier]

Abstract: We examined forest structure and understory vascular plant communities of aspen (Populus tremuloides)-dominated mixedwood boreal forest in Alberta, Canada that had been burned by wildfire and then subjected to one of three treatments (salvage harvested with single-tree retention, salvage harvested with patch-retention, unsalvaged control). Both salvage harvesting treatments resulted in greater cover of regenerating aspen saplings, as compared to unsalvaged controls, but there were no significant effects on vascular plant richness per plot or β diversity. The salvage harvesting treatments differed from the unsalvaged control, but not from one another, in terms of vascular plant community composition. The significant indicators of the unsalvaged treatment were total moss cover and Geranium bicknellii, a fire specialist species. Another fire specialist, Corydalis sempervirens, was an indicator of the patch-retention treatment. Several environmental factors found to be significantly related to vascular plant composition reflected variation in burn severity, rather than the salvage-harvesting treatment. Burned-unsalvaged mixedwood forest was compared to three burned forest types that were unsalvaged because they are not considered merchantable [jack pine (Pinus banksiana), jack pine–black spruce (Picea mariana), black spruce]. The four types of forest were quite different in terms of their structure and vascular plant communities. Burned, non-merchantable forest will not serve as reserves of the post-fire plant communities typical of commercially desirable mixedwood forest, but they may harbor small populations of fire specialist species. Burn severity and pre-fire forest composition have an important influence on the post-fire community, perhaps even more than the salvage-harvesting treatments we examined. Partial salvage harvesting may be an effective way to mitigate some of the potentially negative impacts of post-fire salvaging. [Copyright &y& Elsevier]

Reclaimed landscapes after oil sands mining have saline soils; yet, they are required to have similar biodiversity and productivity as the predisturbance nonsaline landscape. Given that many species in the boreal forest are not tolerant of salinity, we studied the effects of soil salinity on plant communities in natural saline landscapes to understand potential plant responses during the reclamation process. Vegetation–soil relationships were measured along transects from flooded wetlands to upland forest vegetation in strongly saline, slightly saline, nonsaline, and reclaimed boreal landscapes. In strongly saline landscapes, surface soil salinity was high (>10 dS/m) in flooded, wet-meadow, and dry-meadow vegetation zones as compared to slightly saline (<5 dS/m) and nonsaline (<2 dS/m) landscapes. Plant communities in these vegetation zones were quite different from nonsaline boreal landscapes and were dominated by halophytes common to saline habitats of the Great Plains. In the shrub and forest vegetation zones, surface soil salinity was similar between saline and nonsaline landscapes, resulting in similar plant communities. In strongly saline landscapes, soils remained saline at depth through the shrub and forest vegetation zones (>10 dS/m), suggesting that forest vegetation can establish over saline soils as long as the salts are below the rooting zone. The reclaimed landscape was intermediate between slightly saline and nonsaline landscapes in terms of soil salinity but more similar to nonsaline habitats with respect to species composition. Results from this study suggest it may be unrealistic to expect that plant communities similar to those found on the predisturbance landscape can be established on all reclaimed landscapes after oil sands mining. [ABSTRACT FROM AUTHOR]

Despite the importance of seedbeds in the life histories of many plant species, there has been little study of the seedbeds created by wildfire in fire-prone vegetation types such as the boreal forest. Both within the interior and at the edge of a very large (>100 000 ha) 2001 wildfire in the mixedwood boreal region of Alberta, we examined the postfire duff depth and the percent coverage of seedbed types. Minimizing the effect of site and forest composition, we looked only at Picea glauca (Moench) Voss – Populus tremuloides Michx. sites burned during a single day of high fire intensity. Good seedbeds (thin humus and exposed mineral soil, with or without ash) averaged 35% coverage within the interior of the fire but varied enormously among stands. There was a weak but significant positive correlation between prefire percent white spruce basal area and percent mineral soil exposure; that is, there is some tendency for conifer stands to create the seedbeds best suited for their own germinants. Fire severity played a clear role in mineral soil exposure, which was greatest in areas with 100% canopy mortality. Mineral soil exposure was far less at the edges of the fire, averaging only 5% even in areas where all trees had been killed; the burn edge was characterized by superficial flaming combustion with no evidence of substantial duff removal via smoldering combustion. In short, the areas where white spruce seed will be most common after the fire, the edges, are where the worst seedbeds in the burn will be found. Regeneration microsites at fire edges appear to be better suited to regeneration of broadleaf species, via suckering; the persistence of white spruce in fire-prone landscapes continues to be difficult to explain. [ABSTRACT FROM AUTHOR]

Although forest edges have been studied extensively as an important consequence of fragmentation, a unifying theory of edge influence has yet to be developed. Our objective was to take steps toward the development of such a theory by (1) synthesizing the current knowledge of patterns of forest structure and composition at anthropogenically created forest edges, (2) developing hypotheses about the magnitude and distance of edge influence that consider the ecological processes influencing these patterns, and (3) identifying needs for future research. We compiled data from 44 published studies on edge influence on forest structure and composition in boreal, temperate, and tropical forests. Abiotic and biotic gradients near created forest edges generate a set of primary responses to edge creation. Indirect effects from these primary responses and the original edge gradient perpetuate edge influence, leading to secondary responses. Further changes in vegetation affect the edge environment, resulting in ongoing edge dynamics. We suggest that the magnitude and distance of edge influence are a direct function of the contrast in structure and composition between adjacent communities on either side of the edge. Local factors such as climate, edge characteristics, stand attributes, and biotic factors affect patch contrast. Regional factors define the context within which to assess the ecological significance of edge influence (the degree to which the edge habitat differs from interior forest habitat). Our hypotheses will help predict edge influence on structure and composition in forested ecosystems, an important consideration for conservation. For future research on forest edges in fragmented landscapes, we encourage the testing of our hypotheses, the use of standardized methodology, complete descriptions of study sites, studies on other types of edges, synthesis of edge influence on different components of the ecosystem, and investigations of edges in a landscape context. [ABSTRACT FROM AUTHOR]

Patterns of moss and liverwort species diversity -- species richness and species turnover (b diversity) -- in three conifer-dominated boreal forest stands of northern Alberta, Canada are described. We examined the relationship between bryophyte species diversity and micro-environment at two sample grains, the microsite -- substrate types for moss colonization: logs, stumps, tree bases, undisturbed patches of forest floor (dominated by feather moss species), and disturbed patches of forest floor -- and the mesosite (25 m x 25 m plots). Microsite type and properties (e.g. decay class, hardwood vs softwood, pH) were the principal predictors of bryophyte species diversity and not micro-environment variation among mesosites. Microsite type was the strongest predictor of microsite species richness and b diversity was higher among microsites and types and within microsites than among mesosites or stands. Microsite properties were significant predictors of species richness for all microsite types. Log and stump decay classes,influenced also by hardwood vs soft-wood predicted species richness of woody microsite types and soil pH and moisture predicted species richness of forest floor microsites. β-diversity was highest for tree bases and disturbed patches of forest floor and lowest for logs. Mesosite b diversity was lower than that among microsites, and mesosite species richness was not well explained by measured environmental parameters. Results suggest that in conifer-dominated boreal stands, species richness of microsites is only negligibly influenced by within-stand variation at the mesosite grain and that substrate characteristics are the most important predictors of bryophyte species diversity in this ecosystem. [ABSTRACT FROM AUTHOR]

Abstract. We investigated the spatial and temporal pattern of edge influence on primary (forest structure) and secondary (understory structure and composition) responses at forest edges next to regenerating clear-cuts in Populus-dominated boreal forest in Alberta, Canada. We used before/after-harvest comparisons to determine whether there were significant effects of edge creation in the first and second year after harvest. We also investigated 5- and 16-yr-old edges to determine whether their structure and composition were significantly different from interior forest. Distance of edge influence (20 - 60 m) was not extensive compared to findings from studies conducted in other forest types. In the first two years after harvest, there were significant primary responses to edge creation including increased tree mortality, snag breakage and amount of downed coarse woody debris; as well as secondary responses including prolific vegetative regeneration of Populus spp. and a decrease in total shrub cover. Significant edge effects were still detectable 5 and 16 yr after edge creation, despite the rapid regeneration of Populus -dominated forest in the adjacent clear-cuts. These older edges showed evidence of past primary and secondary responses, including a more heterogeneous canopy, along with additional changes in understory composition. At both younger and older edges the distance of edge influence was greater for secondary responses (up to 60 m) than for primary responses (10 - 20 m). We hypothesize that secondary responses to edge creation start later than primary responses and that the distance of edge influence is greater for secondary (vs. primary) responses. [ABSTRACT FROM AUTHOR]

• Site wetness influenced understory response to retention harvesting. • Within each forest type, responses varied along the depth-to-water gradient. • Conifer stands had the greatest change in understory-DTW relationships. • Depth-to-water index can be used to guide placement of retention. To inform biodiversity conservation efforts in managed forest landscapes, we explore if a topographic moisture index (depth-to-water, based on remotely-sensed (LIDAR) data) can provide insight into responses of understory vegetation to retention harvesting in the boreal mixedwood forests of northwestern Alberta, Canada. Sample plots were placed along the depth-to-water moisture gradient in three forest types: coniferous, mixedwood, and deciduous (broadleaf), and in four retention harvesting treatments: unharvested (control), 50% (dispersed green-tree) retention, 20% retention, and clearcut (2% retention). Understory diversity, abundance, and composition were assessed 15 years after harvest. Harvesting affected the relationships between understory variables and the depth-to-water index, with the effects differing between forest types. Coniferous stands showed the most dramatic responses to harvesting, in that most relationships between understory attributes and the depth-to-water index changed due to harvesting. For instance, harvested coniferous stands had higher diversity on wetter sites, rather than on drier sites as was seen in the unharvested stands. In mixedwood stands only the relationship between composition and depth-to-water was affected by harvesting. Broadleaf stands were intermediate; abundance and composition showed a significant depth-to-water by harvesting treatment interaction. Abundance and depth-to-water relationships were weaker in harvested, as compared to unharvested, broadleaf stands. Within each forest type, the effects of harvesting also varied along the depth-to-water gradient. In coniferous and mixedwood forest types, wetter sites were most sensitive to harvesting while in broadleaf stands drier sites were more sensitive. Our study shows that the depth-to-water index can be used to better understand and predict the response of understory vegetation to harvesting and can be useful for guiding the placement of retention. [ABSTRACT FROM AUTHOR]

Retention harvesting, or the approach of leaving live mature trees behind during forest harvest, is used in natural disturbance‐based management to mitigate the effects of logging on biodiversity. However, responses of many boreal vertebrates to variable retention harvesting are unknown. We investigated the influence of different retention levels in forest harvests on stand use by wildlife 15–18 yr post‐harvest using a combination of surveys of wildlife signs (scats, middens) and camera trapping. Site‐level measures of forest structure, including canopy cover, horizontal cover, tree height, tree diameter, basal area, cover of downed coarse woody material, and understory plant cover, were used to describe post‐harvest differences in habitats used by common wildlife species in northwest Alberta's boreal forest. Stand use of six species (black bear, coyote, fisher, red squirrel, wolverine, woodland caribou) increased with level of retention, while stand use of two species (grouse, snowshoe hare) declined with retention level. Retention level did not significantly affect stand use of five species (American marten, Canada lynx, deer, gray wolf, moose). Higher levels of retention characterized by greater canopy cover, basal area, and abundance of deadwood were associated with use of forest habitats by late‐seral species. Woodland caribou, a species of conservation concern, was detected only in harvested stands with at least 20% retention. Greater understory and horizontal cover characterized lower levels of retention being attractive for early‐seral species. These findings demonstrate the value of retention harvesting for conservation of wildlife species in boreal forest, while highlighting the challenge of managing forests for multiple species with different habitat preferences. [ABSTRACT FROM AUTHOR]

Provides information on the 4th International Workshop on Disturbance Dynamics in Boreal Forest held at the University of Northern British Columbia on August 9 to 12, 2002. Issues involving forest management discussed at the event; Keynote speakers who were present in the workshop; Discussion on the importance and role of disturbance in shaping boreal forest.