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The giant panda, currently restricted to a small region of central China, was once widely dispersed throughout southern China and even some parts of continental Southeast Asia during the Pleistocene epoch. However, the evolutionary process and intraspecific variation remain poorly understood in light of limited fossil records, which are usually isolated teeth and a few fragmentary skulls and mandibles. Here we report three skulls, two of which are nearly complete, of Ailuropoda melanoleuca baconi from Yanjinggou in China, which represent the best-preserved skull material ever recorded for this paleosubspecies. A. m. baconi, characterized by a large and robust body, extremely developed zygomatic arches, a moderately developed sagittal crest, and a medium-rough external surface of the supraoccipital, exoccipital, and ascending ramus compared with Ailuropoda microta and other paleosubspecies and living subspecies of A. melanoleuca, can be regarded as a valid paleosubspecies. Ailuropoda fovealis or Ailuropoda melanoleuca fovealis turns out to be a synonym of A. m. baconi. The intraspecific variation of A. m. baconi is far more complicated than generally accepted. Based on comparisons with A. microta, A. m. wulingshanensis, and modern Ailuropoda melanoleuca, we propose that the evolutionary tendency of the skull appears to be much clearer than that of the teeth and that the increase in body size and the enhancement of the masticatory system can be deemed as the main evolutionary changes in this part of the Ailuropoda lineage. Meanwhile, our study for the first time provides a numerical age (ca. 0.35–0.26 Ma) for the Yanjinggou fauna, which is generally considered as a mixture of the middle Early Pleistocene fauna and the Mid-Late Pleistocene fauna. [ABSTRACT FROM AUTHOR]

Bianfudong (pinyin for Bat Cave) is a paleolithic site that was recently discovered in Heqing, Yunnan in southwestern China. Several hominin fossils, stone artifacts and more than 69,000 mammal fossils were unearthed in 2019. The hominin fossils and stone artifacts were excavated from the upper part (layers 1–12) of the section with an age range of 180–70 ka BP. Some large mammal fossils and micromammalian remains were collected from the lower part of the section (layers 22–30), which has an age >180 ka BP. This paper describes the micromammalian fossils collected from layers 25–30 and analyzes the fauna composition and paleohabitats. A total of 638 micromammal specimens represent 12 species, including 11 extant species i.e. Euroscaptor longirostris , Blarinella wardi , Anourosorex squamipes , Crocidura vorax , Eothenomys (Anteliomys) chinensis , E. (A.) proditor , Apodemus chevrieri , Ap. draco , Ap. latronum , Micromys minutus , and Niviventer confucianus , and one extinct species Anourosorex qianensis. The two arvicolid Eothenomys species dominate, both in the number of identified specimens and the minimum number of individuals. The composition of the Bianfudong micromammalian fauna can be compared with the faunas of Sanjiacun in Chenggong of Yunnan Province, Mawokoudong in Bijie and Xinglongdong in Fengjie, Guizhou Province. Using the random forest model with modern meteorological data, we calculated the mean annual precipitation (MAP) and the mean annual temperature (MAT) for all 11 living species in the Bianfudong. The MAP was inferred as 1008.04 mm, close to modern data in the Heqing area, while the MAT was slightly lower at 12.84 °C. The dominance of Eothenomys also supports a slightly cooler climate during the late Middle Pleistocene in the Heqing area. [ABSTRACT FROM AUTHOR]

Abstract The fossil evidence of hominins in China is crucial to understand the human evolution and dispersal in Eurasia. However, the dating of Chinese Early and Middle Pleistocene hominin record remains a serious problem. In this study, we applied coupled ESR/U-series dating method on Bailongdong hominin cave site in Hubei Province, central China, which is a key area to study early human migration between South and North China. The U-series analyses show that the fossil teeth samples from three horizons are close or beyond equilibrium, which indicate very early uptake or potential loss of uranium. Single saturation exponential fitting of 10 dose points irradiated up to 4 kGy was used for D E determination except one by double saturation exponential fitting with 16 dose points up to 63 kGy. Calculated by two uranium migration models, we obtain a weighted mean age of 509 ± 16 ka for five fossil teeth from layer 2. Combined with cosmogenic nuclides 26Al/10Be burial dating carried out on the cave deposits beneath the fossil layers in previous study, the multi-methods dating study places Bailongdong site at the early stage of Middle Pleistocene, and pinpoints the deposition of the fossil remains to the time of marine isotope stages 13. This is in agreement with the fauna composition and sedimentary characteristics which show a subtropical-tropical warm forest-grass environment. The multi-methods dating of Bailongdong site by both cosmogenic and coupled ESR/U-series methods is a good example for the future chronological studies of early human sites in China. Highlights • A Middle Pleistocene hominin site was dated by coupled ESR/U-series method. • Multi-methods dating study by ESR fossil dating and 26Al/10Be burial dating. • D E estimation by two fitting functions with different maximum doses was compared. [ABSTRACT FROM AUTHOR]

In connection with the future Seine-North Europe Canal (Seine-Scheldt), a large-scale rescue archaeological survey was conducted at Havrincourt (northern France) between 2008 and 2011. The discovery of several levels of Palaeolithic flint artefacts embedded in a relatively thick loess sequence (ca 6–7 m) preserved on a gentle slope facing North-East, resulted in a 6000 m 2 excavation. This opened the opportunity for a detailed pedosedimentary and interdisciplinary geochronological survey ( 14 C, optically stimulated luminescence, thermoluminescence, ESR/U-series) that has allowed us date the sequence reliably. On the basis of these results we propose this sequence as a new pedostratigraphic and archaeological reference sequence for northern France. We present here the optical dating of the sequence performed on fine (4–11 μm) quartz grains extracted from 17 samples. The luminescence characteristics of these extracts indicate that the single-aliquot regenerative dose optically stimulated luminescence (SAR-OSL) procedure that was applied is well suited. A consistent set of optical ages was obtained for the loess deposited up to around 70 ka ago. Independent age control (pedostratigraphy, 14 C, ESR/U-series dates) allowed us to apply a Bayesian approach to build a chronometric model. This in turn enabled a regional chronostratigraphic framework to be built, to constrain the correlations with neighboring regions (northern France and Belgium) and to calculate a precise age for the four Palaeolithic levels discovered, including a unique occupation related to the early Upper Palaeolithic which was previously unknown in the area. [ABSTRACT FROM AUTHOR]

Gigantopithecus blacki was once a common primate during Early Pleistocene, distributed throughout southern China, but then it gradually declined with time and finally became extinct probably during Middle Pleistocene. Black Cave in Guangxi (the Zhuang Nationality Autonomous Region) was the first site yielding G. blacki teeth in-situ in 1956. It also produced numerous mammalian fossils belonging to the typical Middle Pleistocene Stegodon–Ailuropoda fauna from sediments overlying the G. blacki level. The present study carried out coupled ESR/U-series dating study of five Bovidae teeth from this palaeontological layer and obtained a weighted mean age of 383 ± 20 ka for this Stegodon–Ailuropoda fauna. This age estimate on fossil teeth is constrained by two 230 Th/U ages of 404 ± 24 ka and 382 ± 9 ka obtained on speleothems formed below and above the level of the Stegodon–Ailuropoda fauna. The Black Cave Stegodon–Ailuropoda fauna can be therefore correlated with MIS 11, and the remains of G. blacki should be older than 400 ka. [ABSTRACT FROM AUTHOR]

This paper reports the discovery of a new mammalian assemblage at Mocun cave in 2000 by the Natural History Museum of Guangxi. The Mocun site is located in a residual karst peak in southwest Guangxi, south China. Abundant mammalian fossil teeth and bones were extracted from relatively thick deposits, well constrained stratigraphically. The results show that 1) the Mocun fauna is composed of typical species of the " Ailuropoda - Stegodon association", including Stegodon, Ailuropoda, Rhinoceros, Pongo, Tapiru s, Megatapirus, and Elephas, common in the Middle to the Late Pleistocene, with a larger portion of modern species; 2) By employing U-series dating (MC-ICP-MS) method on both the soda straw stalactites and mammalian teeth collected from deposits, the age of the Mocun fauna has been constrained between 66 and 101 ka, a transitional period from the interglacial stage MIS5 to the beginning of the last glacial stage (MIS4), representing one of the securely dated Late Pleistocene sites in southern China so far; 3) The composition in large mammals suggests a forested environment which harboured a great biodiversity in large species; 4) The taphonomic analysis indicates that the fossil accumulation is mainly due to activities of porcupines and to water flows; 5) Interestingly, our taxonomic study reveals some changes in the occurrence of Tapirus sinensis and Megatapirus augustus between ∼66 and ∼86 Ka, along with a change in the abundance of artiodactyls versus primates. It remains to demonstrate with more comprehensive records if these changes may be related to the mid-Late Pleistocene climatic transition. • a new mammalian fauna cave site (Mocun cave) with accurate date to 66 and 101 ka in southwest Guangxi. • The composition in large mammals suggests a compatible habitat for a great biodiversity of species. • Some changes between layers with Tapirus sinensis replaced by Megatapirus augustus between ∼66 and ∼86 Ka, along with a change in the abundance of artiodactyls versus primates. [ABSTRACT FROM AUTHOR]

The Middle Pleistocene archaeological site of Guado San Nicola was discovered in 2005 in a fossil fluvial terrace of the Volturno River, close to the village of Monteroduni, Molise, Italy. Palaeontological remains and lithic artefacts, including both handaxes and Levallois, discoid and opportunistic debitage, were recovered in fluvial and slope sediments rich in volcanoclastic materials. This site includes four distinct human occupation levels. In two of them both “shaping-façonnage” and “knapping-débitage” technologies are highlighted, placing this site at the Lower/Middle Palaeolithic transition. In the present study, geochronological analyses by 40 Ar/ 39 Ar on single-crystal and ESR/U-series on teeth were performed to precise the chronological framework of the occupations. The 40 Ar/ 39 Ar data obtained securely bracket the human occupation levels at the transition between the interglacial and glacial marine isotopic stages MIS 11 (i.e. 400 ± 9 ka) and MIS 10 (i.e. 345 ± 9ka). The weighted mean age obtained from ESR/U-series dating of six teeth (i.e. 364 ± 36 ka) is in very good agreement with the 40 Ar/ 39 Ar results. The radio-isotopic constraints we presented place the Guado San Nicola site as one of the earliest testimonies of Levallois debitage in Western Europe and confirm the potential and accuracy of paleo-dosimetric methods to date Middle Pleistocene sites. [ABSTRACT FROM AUTHOR]

Several caves of the Guangxi Zhuang Autonomous Region, southern China, have delivered Gigantopithecus blacki remains, an extinct Pleistocene giant ape, in association with abundant mammalian faunas. To determine their geological ages, fossil teeth from Mohui and Sanhe Caves were dated using the coupled ESR/U-series method. The teeth from Mohui Cave gave age estimates of 1.69 ± 0.22 Ma and 1.29 ± 0.11 Ma. The Sanhe Cave samples had age estimates ranging from 910 ± 200 ka to 600 ± 150 ka with error weighted mean ages of 890 ± 130 ka and 720 ± 90 ka for the layers 5 and 4, respectively. Our results and previous paleomagnestism data place the Gigantopithecus fauna at Mohui Cave between Olduvai and Jaramillo subchrons and suggest that it was coeval with Chuifeng, Longgupo and Liucheng assemblages. The Sanhe fauna is younger, of late early Pleistocene age, and can be dated to the period between Jaramillo subchron and B/M boundary. [ABSTRACT FROM AUTHOR]

Abstract: The combined U-series/electron spin resonance (ESR) dating method was applied to nine teeth from two Early Pleistocene archaeological sites located in the Orce area (Guadix-Baza Basin, Southern Spain): Fuente Nueva-3 (FN-3) and Barranco León (BL). The combination of biostratigraphy and magnetostratigraphy places both sites between the Olduvai and Jaramillo subchrons (1.78–1.07Ma). Our results highlight the difficulty of dating such old sites and point out the limits of the combined U-series/ESR dating method based on the US model. We identified several sources of uncertainties that may lead to inaccurate age estimates. Seven samples could not be dated because the dental tissues had (230Th/234U) activity ratios higher than equilibrium, indicating that uranium had probably leached from these tissues. It was however possible to calculate numerical estimates for two of the teeth, both from FN-3. One yielded a Middle Pleistocene age that seems to be strongly underestimated; the other provided an age of 1.19±0.21Ma, in agreement with data obtained from independent methods. The latter result gives encouragement that there are samples that can be used for routine dating of old sites. [Copyright &y& Elsevier]

The stratigraphic sequences of numerous Palaeolithic sites of Central and Southern Italy, very rich in both archaeological and palaeontological remains, have also recorded Pleistocene volcanic events through volcanic ash deposits (tephra). They allow the establishment of an accurate chronological framework by comparing results obtained by 40Ar/39Ar dating method on single volcanic K-feldspar crystals, with those derived from ESR and ESR/U-series analyses on fluvial bleached quartz and tooth enamel respectively. Since 2009, these three methods were hence applied on samples of several Middle Pleistocene sites of Central and Southern Italy including both volcanic and archaeological levels (from the west to the east): La Polledrara di Cecanibbio, Isoletta, (Latium), Guado San Nicola, Isernia La Pineta (Molise), Valle Giumentina (Abruzzo) and Venosa Notarchirico (Basilicata). The obtained results, covering a time range from 660 to 350 ka, prove that such a multi-method approach when possible is essential to constrain the chronology of each site and allow the recognition of the specific limitations due to the lack of quartz for ESR or to complex geochemical histories in teeth rendering difficult the ESR/U-series method. Despite these limitations, the ESR framework is globally in agreement with the 40Ar/39Ar chronology, while ESR/U-series dates can be underestimated for the oldest sites. In such cases, an isochron approach attests however of the quite good reliability of the palaeodosimetric reconstruction and the observed age underestimation could be related to other factors affecting the ESR age determination. [ABSTRACT FROM AUTHOR]

The Pongo fossil record of China extends from the Early Pleistocene to the Late Pleistocene, but to date, no late Middle Pleistocene samples of Pongo with precise absolute dating have been identified in southern China. Here, we report the recovery of 106 fossil teeth of Pongo from Ganxian Cave in the Bubing Basin, Guangxi, southern China. We dated the speleothems using Uranium-series and dated the two rhinoceros teeth using coupled electron spin resonance/Uranium-series dating methods to between 168.9 ± 2.4 ka and 362 ± 78 ka, respectively. These dates are consistent with the biostratigraphic and magnetostratigraphic age estimates. We further describe the fossil teeth from Ganxian Cave and compare them metrically to samples of fossil Pongo (i.e., Pongo weidenreichi , Pongo duboisi , Pongo palaeosumatrensis , Pongo javensis , and Pongo sp.) from the Early, Middle, and Late Pleistocene and to extant Pongo (i.e., Pongo pygmaeus and Pongo abelii) from Southeast Asia. Based on overall dental size, a high frequency of lingual cingulum remnants on the upper molars, and a low frequency of moderate to heavy wrinkling on the molars, we attribute the Ganxian fossils to P. weidenreichi. Compared with Pongo fossils from other mainland Southeast Asia sites, those from Ganxian confirm that dental size reduction of Pongo occurred principally during the Early and Middle Pleistocene. From the Middle to Late Pleistocene, all teeth except the P3 show little change in occlusal area, indicating that the size of these teeth remained relatively stable over time. The evolutionary trajectory of the Pongo dentition through time may be more complex than previously thought. More orangutan fossils with precise dating constraints are the keys to solving this issue. [ABSTRACT FROM AUTHOR]

Recently, rich fossiliferous deposits dated to the late Cenozoic have been discovered from Jinyuan Cave at Luotuo Hill, an extremely large-sized cave site situated within in the Dalian Puwan Economic Zone of Dalian Municipality, Liaoning Province in Northeast China. Abundant and diverse vertebrate fossils unearthed from different fossiliferous layers of Jinyuan Cave, a site of enormous size with a sedimentary thickness of over 40 m and its deposits can be roughly divided into three fossiliferous deposit units, altogether containing six fossiliferous layers belonging to different ages from the Late Pliocene to the Middle Pleistocene. Based on a combination of biostratigraphic and geochronological evidence, the fossil assemblages from Jinyuan Cave can be divided into three successive faunas that span the Late Cenozoic (ca. 3.60–0.35 Ma): the Wanghai fauna from upper faunal unit (Middle Pleistocene, 0.78–0.35 Ma), the Jinyuan fauna from middle-lower faunal unit (Early Pleistocene, 2.60–0.78 Ma) and the Luotuoshan fauna of the bottommost faunal unit (Late Pliocene, 3.6–2.6 Ma). The mammalian faunal evolution of Jinyuan Cave can be divided into four temporal stages during the late Cenozoic. The new discovered Jinyuan Cave provides a unique and nearly-continuous faunal and sedimentary sequence stretching from the Late Pliocene to the Middle Pleistocene, hugely significant for establishing the Quaternary Mammal Fauna Sequence of Northeast China for the first time based on a combination of biostratigraphy, geochronology, and for discussing the environmental background. [ABSTRACT FROM AUTHOR]

Biochronology is the most widely used method for organizing the successions of continental vertebrate faunas in geological time and for allowing the correlation of continental deposits. However, due to the fragmentary nature of the paleontological record and the diachronicity of first and last occurrences of vertebrate taxa in different areas (i.e., bioevents), it is difficult to precisely define the temporal boundaries between biochronological units. That is why it is crucial to calibrate biochronological data with independent proxies wherever possible. Here, thanks to an interdisciplinary approach that combines sedimentology and stratigraphy, vertebrate paleontology, micropaleontology (ostracods), palynology, and geochronology, we provide a chronological framework and a paleoenvironmental reconstruction of the paleontological site of Pantalla (Central Italy). The combination between biochronological/biostratigraphic and geochronological (U-series/ESR dating and paleomagnetism) data allows us to refer the site to c. 2.2 Ma, i.e., in the transition phase between the Middle and Late Villafranchian in the European biochronological scheme. We reconstruct the environment as a fluvial area with a frequently flooded wet floodplain and marked (probably seasonal) variations of the water level of the channel river. The depositional system was surrounded by a conifer-dominated forest, which is suggestive of a glacial phase. • Pantalla is a paleontological site yielding well-preserved terrestrial mammals. • Geological and paleontological data are combined in an interdisciplinary approach. • A chronological and paleoenvironmental framework of Pantalla is provided. • Age of the site (c. 2.2 Ma) is close to the Middle-Late Villafranchian transition. • The environment was a fluvial area with a frequently flooded wet floodplain. [ABSTRACT FROM AUTHOR]

The archaeological site of Guado San Nicola (Monteroduni, Central Italy), under excavation since 2008, is located in the Upper Volturno Valley, near the top of a fluvial terrace attributed to the Middle Pleistocene. Several anthropic levels have been recognized within a stratigraphic sequence of more than two meters in thickness, characterized by alternating gravelly and sandy fluvial layers rich in pyroclastic materials, attributable to the end of MIS 11–beginning of MIS 10, according to the obtained radiometric dates ( 40 Ar/ 39 Ar and ESR/U-series). In this paper we discuss the chrono-stratigraphic record, the paleontological and zooarchaeological data and we provide a detailed techno-economic analysis of the lithic industry, characterized by the coexistence of handaxes and Levallois debitage. The site of Guado San Nicola represents the oldest evidence of the emergence of prepared-core technologies in the Italian Peninsula, adding new data to the debate on the Lower/Middle Palaeolithic transition in Western Europe. [ABSTRACT FROM AUTHOR]

ESR dating was applied to fossil teeth and optically bleached quartz grain samples from two units of the sequence at Vallparadís (Barcelona, Spain): weighted mean ESR age estimates of 858 ± 87 ka and 849 ± 48 ka were obtained for EVT-7, which includes the archaeological level 10, and EVT-8, respectively. These results are in good agreement with the existing magneto-biostratigraphic framework that constrain these deposits between 780 and 990 ka, and indicate that Vallparadís EVT-7 has a chronology very close to that of Atapuerca Gran Dolina TD-6 (Spain). [ABSTRACT FROM AUTHOR]

Paleolithic rock paintings are known from many parts of the world, including Africa, Australia, Europe and Island Southeast Asia. However, no Paleolithic rock paintings have yet been confirmed in the East Asia mainland, partly due to the lack of suitable dating materials. The rock paintings distributed at more than 70 sites in the Jinsha River valley, Yunnan Province, southwestern China, include the naturalistic outlines of large mammals, which superficially resemble the Magdalenian rock art in Europe. Wanrendong Cave in Tiger Leaping Gorge, has preserved typical Jinsha River rock paintings, associated with some precipitates of secondary carbonates (i.e., speleothem). High precision U-series dating of 13 small speleothems, with a total of 36 subsamples overlying or underlying the pigment layers, revealed that these red paintings were created during the Pleistocene – Holocene transition, representing the oldest absolutely dated rock paintings from China so far. Our results show that at least three painting phases in Wanrendong Cave can be precisely constrained: ∼13,000–13,580 yr B.P., ∼10,540–10,830 yr B.P., and ∼8370–8700 yr B.P. These intervals are substantially earlier than the timing of agriculture in Yunnan Province (∼4600 yr BP), indicating that these paintings were created by Paleolithic hunter-gatherers. Additionally, these intervals coincide with the Bølling-Allerød and early Holocene warming periods, suggesting a possible link between the rock painting activity and climate changes. • High-precision U-series dating was applied to the speleothems associated with the rock paintings in Wanrendong Cave. • Wanrendong rock paintings were dated to three phases: 1) ∼13,000–13,580 yr B.P., 2) ∼10,540–10,830 yr B.P., and 3) ∼8370–8700 yr B.P. • The ages of Wanrendong rock paintings indicate that these paintings were created by Paleolithic hunter-gatherers. [ABSTRACT FROM AUTHOR]