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Ophion klimenkoi Johansson sp. nov. urn:lsid:zoobank.org:act: 38AFA9D4-A900-4EAA-8C2E-E2B9417734D0 Figs 13A‒E Holotype: IRAN: ♀, Khorasan Province, SW part, Khezri, env., 33˚56’N, 58˚43’E, 1800 m. a. s., 23 May 2009, A. Klimenko leg. (MR). Paratypes: IRAN: 1♂, Khorasan Province, SW part, Khezri, env., 33˚56’N, 58˚43’E, 1800 m. a. s., 23 May 2009, A. Klimenko leg. (MR); 1♀, Khorasan Province, SE part, Q’aen area, Khunik pass, 33˚33’N, 59˚08’E, 2100 m. a. s., 25 May 2009, A. Klimenko leg. (MR). Type depositories: The holotype is housed in the ZSM, paratypes in MR. Etymology: The name refers to A. Klimenko who collected the only known specimens of the species. Diagnosis: Possibly a member of the O. mocsaryi species group. Distinguished from the other known members of the aggregate by the very densely and partly rugulose face in combination with the obtuse pleurosternal angles, which are situated distinctly anterior to the sternal angles. Description: Fore wing length 12.5‒13 mm. Body length 14‒15 mm. Antenna with 48‒54 flagellomeres. Temple narrowed behind compound eyes, in lateral view about 0.4 × as long as compound eye in female and male. Space between compound eye and lateral ocellus narrow, about 0.1‒0.2 × the diameter of ocellus in female and male. Occipital carina centrally evenly rounded. Face below antennal sockets shagreened with very dense punctures that partly form rugulose sculpture (Fig. 13E). Malar space narrow about 0.2 × as long as mandibular base in female and male (Fig. 13C). Mandibular gape acute, without internal angles, as in O. slaviceki Kriechbaumer. Antennae stout, first flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.2‒1.4 × as long as wide. Apical flagellomeres approximately 1.4 × as long as wide. Mesoscutum strongly polished, virtually without puctures. Mesopleuron weakly shagreened with punctures quite dense, space between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles strongly obtuse and indistinct, distinctly anterior to sternal angles (Fig. 13D). Scutellum without lateral carinae. Propodeum with the anterior transverse carina in some specimens forming an arcuate lobe centrally, with very weak rugulose sculpture, opaque. Posterior transverse carina indicated laterally (Fig. 13B). Propodeum polished, anterior to anterior transverse carina with distinct scarce puncture. Carination ranging from almost complete with only the posterior carina adjacent to area supeomedia absent to having the posterior carina and the lateral parts of the anterior transverse carina absent. Sclerotised part of first sternite ending posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 8.0 × as long as wide. Inner spur of hind tibia long, about 0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus absent or short, reaching 0.3 × the width of the discosubmarginal cell. Radius weakly sinuous, almost straight before junction with pterostigma. Nervellus broken below the middle by the discoidella. Colouration: Body testaceous. Inner and outer eye margins yellow. Mandibular teeth black. ovipositor sheath dark brown, contrasting in colour with posterior metasomal segments. DNA Barcode: The DNA barcode sequences of two specimens of Ophion klimenkoi sp. nov. are available at the BOLD systems database: sample id. NJP631/632, sequence id. LNU2467-21/2468-21. Remarks: A member of the O. mocsaryi -aggregate most closely related to Ophion rostralis, from which it shows a minimum sequence divergence of 1.79%.

Ophion rufoniger Kohl, 1905 Figs 16A‒B, F Ophion rufoniger Kohl, 1905: 223. Lectotype ♀ female from Azerbaijan in NHMW, examined. Material examined: IRAN: 1♀, Esfan dagheh-Djirouft, 20 Apr. 1971, Saf Zair leg. (HMIM). Remarks: Ophion rufoniger (Figs 16A‒B, F) was described from Goygol in Northwestern Azerbaijan (Kohl, 1905). It is a very rarely recorded species also known from Armenia and Georgia. The species is quite easily determined based on the buccate head and the dark reddish brown body with mainly black metasoma and legs. The depicted female specimen is from Georgia and housed in OÖLM., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 184, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Kohl, F. F. (1905) Hymenopteren. In: Penther, A. & Zederbauer, E. (Eds.), \" Ergebnisse einer naturwissenschaftlichen Reise zur Erdschias-Dagh (Kleinasien). \" Annalen des Naturhistorischen Museum in Wien, 20, pp. 220 - 246."]}

Ophion transcarinatus Johansson sp. nov. urn:lsid:zoobank.org:act: 37270A87-2573-4427-A74E-775EED4DB37D Figs 20A‒E Holotype: IRAN: 1♂, Golestan Province, Dasht, 37˚21’N, 56˚00’E, 1000 m. a. s., 11 May 2009, A. Klimenko leg. (MR). Paratypes: IRAN: 3♀♀, 1♂, Kohkiloyeh-va-Boyerahmad Province, Yasuj area, 10 km SE Sepidar, 5‒6 May 2007, A. Klimenko leg. (MR). Type depositories: The holotype is housed in the ZSM, paratypes in MR. Etymology: The name refers to the very distinct anterior transverse carina of propodeum. Diagnosis: A member of the Ophion mocsaryi aggregate with the contrasting black ovipositor sheaths in the female. Similar to Ophion crassicornis Brock, but with the face narrower and the lateral ocelli touching the inner margin of the compound eye. Similar also to O. mocsaryi and O. confusus but with the anterior transverse carina of propodeum more pronounced, centrally almost flange-like, and the pleurosternal angles less acute. Description: Fore wing length 16‒18 mm. Body length 17‒19 mm. Antenna with 57‒61 flagellomeres. Temple behind compound eyes, in lateral view about 0.6 × as long as compound eye in female and male. Space between compound eye and lateral ocellus narrow, about 0.1 × the diameter of ocellus in female and male. Occipital carina centrally evenly rounded. Face below antennal sockets shagreened with quite dense punctures, the interstices between punctures about equal to their diameter. Malar space narrow, about 0.1 × as long as mandibular base in female and male (Fig. 20C). Mandibular gape right angled, with internal angles. Antennae stout, first flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.2‒1.4 × as long as wide. Apical flagellomeres approximately 1.4 × as long as wide. Mesopleuron strongly shagreened, its surface partly rugulose as in O. crassicornis, with punctures quite dense, space between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles almost right-angled and indistinct, distinctly anterior to sternal angles (Fig. 20D). Scutellum without lateral carinae (Fig. 20B). Propodeum with the anterior transverse carina very strongly raised, in a couple of specimens almost flange-like. Posterior transverse carina only indicated laterally (Fig. 20B). Central longitudinal carinae quite weak. Propodeum weakly shagreened. Sclerotised part of first sternite ending almost level to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur quite slender, about 7.0 × as long as wide. Inner spur of hind tibia long, about 0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus distinct reaching about 0.3 × the width of the discosubmarginal cell. Radius weakly sinuous, almost straight before junction with pterostigma. Nervellus broken in the middle by the discoidella. Colouration: Body pale testaceous with paler areas as in the O. obscuratus aggregate, but less distinct, in a couple of specimens almost absent. Inner and outer eye margins widely yellow. Mandibular teeth black. ovipositor sheath dark brown, contrasting in colour with posterior metasomal segments. DNA Barcode: The DNA barcode sequences of two specimens of Ophion transcarinatus sp. nov. are available at the BOLD systems database: sample id. NJP636/669, sequence id. LNU2472-21/2505-21., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 191, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion crassicornis Brock 1980 Ophion crassicornis Brock, 1980: 84‒85; Figs 23, 24. Holotype ♂ from Britain in NHMUK, examined. Material Examined: IRAN: 1♂, 2♀♀, Mazandaran Province, Golestan forest, Tang-e-Gol, 700 m. a. s., 17 May 1993, Paz / Badii leg. (HMIM); 1♂, Mazandaran Province, Golestan forest, Dasht, 700 m. a. s., 16 May 1993, Paz / Badii leg. (HMIM); 1♀, 1♂, Golestan Province, P. M. Golestan, Tang-e-Gol, 5 May 1999, Mogh. / Bar. / Man. leg. (HMIM); 1♀, Golestan Province, 20 km S Gorgan city, 36˚43’N, 54˚35’E, 700 m. a. s., 10 May 2009, A. Klimenko leg. (MR); 1♀, Semnan Province, Shakhmirzad area, Chasm env. 35˚49’N, 53˚15’E 2200 m. a. s., 14 Jun. 2009, A. Klimenko leg. (MR); 1♂, Lorestan Province, Khorramabad area, Mahmudvand, 30 Apr 2007, A. Klimenko leg. (MR). Remarks: A widely distributed species that occurs all over Europe. It appears to be common also in southeastern Europe and the Middle East. New record for Iran., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 174, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion schmidti Johansson sp. nov. urn:lsid:zoobank.org:act: B004AAF8-3C20-469A-8D97-E701DBAC7178 Fig. 17A‒E. Holotype: IRAN: ♀, Ardabil Province, Bodalalou, 38˚01’30.87"N, 48˚30’23.7"E, 18 Sep. 2009, 2014 m. a. s., M. Manzari leg. (HMIM) Paratypes: IRAN: 2♀♀, 1♂, Ardabil Province, Bodalalou, 38˚01’30.87"N, 48˚30’23.7"E, 18 Sep. 2009, 2014 m. a. s., M. Manzari leg. (HMIM). Type depositories: Holotype and paratypes are stored in HMIM, except one female paratype, which is stored in NHRS. Etymology: The species is named in honour of the hymenopterist Stefan Schmidt in Munich. Diagnosis: Distinguished from Ophion minutus Kriechbaumer and Ophion golestanicus Johansson sp. nov. by the narrower face. Most similar to Ophion sistanicus Johansson sp. nov. and O. persicus Johansson sp. nov., but distinguished by the less punctate face and the acute angled mandibular gape. The two known females lack ramellus while the females of both sibling species have distinct ramellus. Description: Fore wing length 6‒8 mm. Body length 7‒9 mm. Antenna with 44‒47 flagellomeres. Head buccate, temple in lateral view 0.4 × as long as compound eye in female (Fig. 17E), 0.5 × in male. Ocelli large, touching compound eye. Face narrow (Fig. 17D). Occipital carina centrally evenly rounded. Face below antennal sockets polished with weak scarce punctures in female. Malar space absent. Mandibular gape strongly acute, without internal angles (Fig. 17C). First flagellomere 3.5 × as long as wide. Central flagellomeres slender, about 1.7‒1.8 × as long as wide. Apical flagellomeres approximately 1.8 × as long as wide. Mesoscutum polished with punctures very shallow and scarce. Mesopleuron in female polished with distinct punctures, interstices between punctures about 3.0 × their diameter, in male more shagreened, with denser, more distinct punctures. Epicnemial carina, in antero-ventral view, with pleurosternal angles obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum opaque, with weak rugulose sculpture. Propodeum with carinae entirely absent (Fig. 17B). Sclerotised part of first sternite ending distinctly posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 7.0‒8.0 × as long as wide. Inner spur of hind tibia long, about 0.4‒0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus absent (in females) or very short reaching 0.1 of the width of the discosubmarginal cell (males). Radius very weakly sinuous, conspicuously bent and widened before junction with pterostigma. Nervellus broken slightly below the middle by the discoidella. Colouration: Body yellow reddish, with the following paler yellowish: head apart, from a small spot in front of anterior ocellus, a small triangular area below the antennal scrobes and anterior margin of clypeus; four longitudinal stripes on mesoscutum; subalar prominence; spots on mesopleuron; a small spot in the lower posterior part of mesoscutum and one larger spot corresponding on metapleuron. Coxae with paler marks on the outer side. Metasoma with irregular pale markings on the posterior segments. Mandibular teeth black. Ovipositor sheath testaceous, concolourous with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 184-187, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion parvulus Kriechbaumer, 1879 Ophion parvulus Kriechbaumer, 1879: 104‒105. Lectotype ♂ in ZSM from Germany, examined. Material examined: IRAN: 1♂, Milan Province, Tutkabon near Roodbar, 36˚83’N, 49˚66’E 960 m. a. s., 16 Jun. 2010, Mi. Halada leg., (OÖLM); 1♂, Bakhtiyari, 20 km SE. Lordegan, 31˚21’N, 51˚09’E 1900 m. a. s., 31 May 2014, J. Halada leg., (OÖLM). Remarks: The aggregate nature of O. parvulus s. lat. was partly resolved by Johansson in Johansson & Cederberg (2019), but probably the aggregate still holds potentially cryptic species (see also remarks under O. fossulatus Hedwig)., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 180, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Kriechbaumer, J. (1879) Ophion minutus n. sp. Entomologische Nachrichten, 5, 105 - 106."]}

Ophion obscuratus (Fabricius, 1793) Ophion obscuratus Fabricius, 1798: 237. Lectotype ♂ in ZMUC, examined. Ophion obscurus Fabricius, 1804: 132. Ophion flavolineatus Brullé, 1846: 147‒148. Lectotype from Papua New Guinea, unsexed in MNHN, examined. Material examined: IRAN: 1♂, Mazandaran Province, Golestan forest, Tang-e-Gol, 700 m. a. s., 17 May 1993, Paz / Badii leg. (HMIM); 1♂, Mazandaran Province, Kelardasht, Rudbarak, Vandarbon, 2455 m. a. s., 23 May 2003, Gil. / Nem. leg. (HMIM). Remarks: The here presented specimens, though slightly smaller in size, corresponds very well morphologically with Ophion obscuratus, an European species occurring in late autumn in central and northern Europe. However, the species treated as Ophion obscuratus by previous authors has shown to be an aggregate consisting of an unknown number of species with slightly different characteristics. The period of flight for the recorded specimens indicates that it potentially could represent a previously undescribed species differentiated from the true O. obscuratus. The species was only recently properly defined (Johansson & Cederberg 2019) and any references in literature preceeding that date of publication may refer to any closely related and similar species., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 180, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion transcarinatus Johansson sp. nov. urn:lsid:zoobank.org:act: 37270A87-2573-4427-A74E-775EED4DB37D Figs 20A‒E Holotype: IRAN: 1♂, Golestan Province, Dasht, 37˚21’N, 56˚00’E, 1000 m. a. s., 11 May 2009, A. Klimenko leg. (MR). Paratypes: IRAN: 3♀♀, 1♂, Kohkiloyeh-va-Boyerahmad Province, Yasuj area, 10 km SE Sepidar, 5‒6 May 2007, A. Klimenko leg. (MR). Type depositories: The holotype is housed in the ZSM, paratypes in MR. Etymology: The name refers to the very distinct anterior transverse carina of propodeum. Diagnosis: A member of the Ophion mocsaryi aggregate with the contrasting black ovipositor sheaths in the female. Similar to Ophion crassicornis Brock, but with the face narrower and the lateral ocelli touching the inner margin of the compound eye. Similar also to O. mocsaryi and O. confusus but with the anterior transverse carina of propodeum more pronounced, centrally almost flange-like, and the pleurosternal angles less acute. Description: Fore wing length 16‒18 mm. Body length 17‒19 mm. Antenna with 57‒61 flagellomeres. Temple behind compound eyes, in lateral view about 0.6 × as long as compound eye in female and male. Space between compound eye and lateral ocellus narrow, about 0.1 × the diameter of ocellus in female and male. Occipital carina centrally evenly rounded. Face below antennal sockets shagreened with quite dense punctures, the interstices between punctures about equal to their diameter. Malar space narrow, about 0.1 × as long as mandibular base in female and male (Fig. 20C). Mandibular gape right angled, with internal angles. Antennae stout, first flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.2‒1.4 × as long as wide. Apical flagellomeres approximately 1.4 × as long as wide. Mesopleuron strongly shagreened, its surface partly rugulose as in O. crassicornis, with punctures quite dense, space between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles almost right-angled and indistinct, distinctly anterior to sternal angles (Fig. 20D). Scutellum without lateral carinae (Fig. 20B). Propodeum with the anterior transverse carina very strongly raised, in a couple of specimens almost flange-like. Posterior transverse carina only indicated laterally (Fig. 20B). Central longitudinal carinae quite weak. Propodeum weakly shagreened. Sclerotised part of first sternite ending almost level to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur quite slender, about 7.0 × as long as wide. Inner spur of hind tibia long, about 0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus distinct reaching about 0.3 × the width of the discosubmarginal cell. Radius weakly sinuous, almost straight before junction with pterostigma. Nervellus broken in the middle by the discoidella. Colouration: Body pale testaceous with paler areas as in the O. obscuratus aggregate, but less distinct, in a couple of specimens almost absent. Inner and outer eye margins widely yellow. Mandibular teeth black. ovipositor sheath dark brown, contrasting in colour with posterior metasomal segments. DNA Barcode: The DNA barcode sequences of two specimens of Ophion transcarinatus sp. nov. are available at the BOLD systems database: sample id. NJP636/669, sequence id. LNU2472-21/2505-21.

Ophion angularis Johansson & Cederberg, 2019 Ophion angularis Johansson & Cederberg, 2019: 37; Figs 9F, 22. Type ♀ in NHRS, examined. Material examined: IRAN: 1♀, 3♂♂, Golestan Province, 20 km S Gorgan city, 36˚43’N, 54˚35’E, 700 m. a. s., 10 May 2009, A. Klimenko leg. (MR). Remarks: Recently described from Northern Europe and the species has since also been recorded from Central and Southern Europe. It seem to prefer semi open xerothermic areas, but the host is still unknown.

Ophion angustigena Johansson sp. nov. urn:lsid:zoobank.org:act: 556A9109-DD41-4B4C-961F-776F3702A512 Figs 10A‒C. Holotype: IRAN: ♀, Hormozgan province, Minab, Raveng, 27˚4’37.36"N, 57˚10’22.09"E, 34 m. a. s., 4 May 2016, Afsari / Mozh. leg. (HMIM). Type depository: The holotype is housed in HMIM. Etymology: The name refers to the narrow temples of this species. Diagnosis: Ophion angustigena Johansson sp. nov. (Fig. 10A‒C) is easily distinguished by the very narrow temples, the narrow face, the large ocelli and the small mandibles. Potentially the species could be confused with Ophion pteridis Kriechbaumer, 1897, which have carinated scutellum, the face wider and the sclerotised part of first sternite ending distincly posterior to spiracle. Description: Male unknown. Fore wing length 15 mm. Body length 17 mm. Antenna with 56 flagellomeres. Temples strongly narrowed, in lateral view 0.1‒0.2 × as long as compound eye in female (Fig. 10C). Space between compound eye and lateral ocellus absent. Face very narrow and clypeus small (Fig. 10B). Occipital carina centrally evenly rounded. Face below antennal sockets polished laterally with distinct but quite sarce punctures, centrally shagreened with punctures almost dissolving. Malar space absent. Mandibular gape almost right angled, with weak indications of internal angles. Mandibles slightly worn in the holotype. First flagellomere 3.5 × as long as wide. Central flagellomeres stout, about 1.4‒1.5 × as long as wide. Apical flagellomeres approximately 1.6 × as long as wide. Mesoscutum polished with very weak indistinct punctures. Mesopleuron with indistinct very small punctures, space between punctures about 3.0 × their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles slightly obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum very weak rugulose sculpture, polished. Anterior transverse carina complete, angled backwards. Posterior transverse carina present laterally. Longitudinal carinae almost totally absent, only weakly indicated centrally. Sclerotised part of first sternite ending slighty posterior to spiracle. Hind trochantellus almost as long wide in dorsal view. Hind femur slender, about 8.0 × as long as wide. Inner spur of hind tibia long, about 0.4 × as long as hind metatarsus. Wing membrane clear. Ramellus distinct, reaching 0.3 × the width of the discosubmarginal cell. Radius sinuous, its inner part straight before junction with pterostigma. Nervellus broken slightly below the middle by the discoidella. Colouration: Body brownish yellow with extensive yellow markings, in distribution reminiscent of the members of the O. obscuratus aggregate, but much more indistinct. The following areas have pale markings: head apart from central part, four stripes on mesoscutum, scutellum laterally, subalar prominence, large crescent on mesopleuron, lower posterior part of metapleuron and posterior part of propodeum, legs except coxae, petioles and irregular spots on tergites 2‒3. Mandibular teeth black. Ovipositor sheath orange, concolorous with posterior metasomal segments. Remarks: A couple of males from Iran in HMIM have similar characteristics as the here described female, but differ in some respects, e.g. the carination of propodeum and the shape of the mandibles. It is possible that they represent the male sex of O. angustigena, but due to these differences, they are not included in the type series., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 170-172, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion rostralis Meyer, 1935 Ophion rostralis Meyer, 1935: 47. Type from Turkmenistan, lost. Material examined: TURKMENISTAN: 1 ♂ in ZIN determined by Meyer and referred to by Townes et al. (1965). IRAN: 9♀♀, Semnan Province, Shakhmirzad area, Chasm env. 35˚49’N, 53˚15’E 2200 m. a. s., 14 Jun. 2009, A. Klimenko leg. (MR); 1♀, Golestan Province, 20 km S Gorgan city, 36˚43’N, 54˚35’E, 700 m. a. s., 10 May 2009, A. Klimenko leg. (MR); 1♀, Khorasan Province, Western part, Bojnurd area, Gerivan N, 37˚17’N, 57˚13’E, 12 May 2009, A. Klimenko leg. (MR); 2♂♂, 1♀, Semnan Province, Damghan area 15 km SW Agareh v., 36˚06’N, 53˚43’E, 2000 m. a. s., 12 Jun. 2009, A. Klimenko leg. (MR). DNA Barcode: The DNA barcode sequences of two specimens of Ophion rostralis are available at the BOLD systems database: sample id. NJP740/741, sequence id. LNU2576-21/2577-21. Remarks: The type of Ophion rostralis, collected in Baýramaly in Turkmenistan is lost, as is one none type female determined by Meyer (Townes et al. 1965, Andrey Khalaim ZIN pers. comm.). The original description is quite vague and could be attributed to several species of Ophion occurring in the area. However, in the collection in St. Petersburg there is one male determined by Meyer, which is the closest one can get to a name bearing specimen for the time being. The male differs from the original description in having the mouthparts of normal constitution, the scutellum without lateral carinae and the body more yellow than brownish red, which may lead to the assumption that the superficially similar Ophion bipictor Aubert, 1980 was also included in the original description. The male in ZIN is similar to Ophion slaviceki Kriechbaumer, 1892, but has the temples more strongly narrowed and the apical flagellomeres stouter. The body is strongly polished with very weak punctures. The mandibles are lacking internal angles, thereby similar to the mandibles of Ophion slaviceki., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 184, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Meyer, N. F. (1935) [Parasitica of the family Ichneumonidae of the USSR and adjacent countries. Part 4. Ophioninae]. Akademia Nauk SSSR 16 (4). Akademia Nauk SSSR Press, Leningrad, 535 pp.","Townes, H. K., Momoi, S. & Townes, M. (1965) A catalogue and reclassification of the eastern Palearctic Ichneumonidae. Memoirs of the American Entomological Institute, No. 5, 1 - 661."]}

Ophion golestanicus Johansson sp. nov. urn:lsid:zoobank.org:act: AEB51640-1574-481B-BD1B-6361497A3E81 Figs 12A‒B, E Holotype: IRAN: ♀, Golestan Province, Dasht, 37˚21’N,56˚00’E, 1000 m.a.s., 11May 2009, A. Klimenko leg.(MR). Paratypes: IRAN: 12♀♀, 3♂♂, same data as holotype, (MR). Type depositories: The holotype is stored in ZSM, paratypes in MR according to original depository. Etymology: The name refers to the fact that the type series was collected in Golestan. Diagnosis: Similar to Ophion minutus Kriechbaumer, 1879, but usually with less pale markings on the mesosoma and on average slightly larger. The temples and the face is narrower and the flagellomeres are longer. In all known specimens, the posterior transverse carina is complete, m-shaped, while it usually is partly or completely missing in Ophion minutus. Also similar to the other three species of the O. minutus aggregate described in this study, but with more yellow pale markings, whitish on the other species, and with the face considerably wider. Description: Fore wing length 9‒10 mm. Body length 10‒12 mm. Antenna with 43‒46 flagellomeres. Temple buccate behind compound eyes, in lateral view about 0.7 × as long as compound eye in female and male (Fig. 12B). Space between compound eye and lateral ocellus narrow, about 0.2 × the diameter of ocellus in female and male. Face relatively narrow (Fig. 12A). Occipital carina centrally widely flattened. Face below antennal sockets with dense distinct punctures, the interstices between punctures about 0.2‒0.5 × the diameter of punctures. Malar space about 0.1 × as long as mandibular base in female and male. Mandibular gape slightly acute, without internal angles, but often worn. First flagellomere 4.0 × as long as wide. Central flagellomeres slender, about 2.0 × as long as wide. Apical flagellomeres approximately 2.0 × as long as wide (Fig. 12E). Mesoscutum polished with scarce punctures, space between punctures about 5.0 × their diameter. Mesopleuron weakly shagreened with distinct punctures, interstices between punctures about equal to their diameter Epicnemial carina, in antero-ventral view, with pleurosternal angles obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum polished with rugulose sculpture. Posterior transverse carina often complete, but in small specimens partly weakened. Sclerotised part of first sternite ending distinctly posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 7.0‒8.0 × as long as wide. Inner spur of hind tibia long, about 0.4‒0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus reaching 0.1‒0.2 of the width of the discosubmarginal cell. Radius very weakly sinuous, conspicuously bent and widened before junction with pterostigma. Nervellus broken in the middle or slightly below the middle by the discoidella. Colouration: Body yellow reddish, with the following paler yellow: mandibles, clypeus, inner and outer orbits, subalar prominence, tegulae, scutellum and postskutellum diffusely, a larger diffuse spot in front of the epicnemial carina on mesopleuron, sometimes a small diffuse spot in the lower posterior part of mesoscutum and one small spot corresponding on metapleuron. Mandibular teeth black. Ovipositor sheath testaceous-brown, concolorous with posterior metasomal segments. DNA Barcode: The DNA barcode sequence of one specimen of Ophion golestanicus sp. nov. is available at the BOLD systems database: sample id. NJP627, sequence id. LNU2463-21. Remarks: The barcoded specimen of Ophion golestanicus Johansson sp. nov. shows a minimum sequence divergence of 3.35% from its closest neighbor, Ophion minutus in BOLD.

Ophion al Johansson sp. nov. urn:lsid:zoobank.org:act: CCB675F3-A628-49AF-B859-78AE80463769 Figs 9A‒B, D‒E Holotype: IRAN: ♀, Kohkiluyeh & Boyerahmad, Kakan, Hoseynkhani, 26 May 1995, 2100 m. a. s., Hash. / Badii / Safs. leg. (HMIM). Paratype: IRAN: 1♀, Chaharmahal & Bakhtiari Province, Kuhrang, Hiregan, 32˚20’31"N, 50˚26’56"E, 6 Jun. 2005, 2305 m. a. s., Ebra / Hai. leg. (HMIM). Type depositories: The holotype is stored in HMIM and the paratype in NHRS Etymology: The scientific name al refers to a nocturnal creature feeding on babies in ancient Persian mythology, hence expressing a similar ecology as Ophion al Johansson sp. nov. Diagnosis: Most similar to Ophion bipictor Aubert, 1980 (Figs 9C, F) but with the pleurosternal angles more obtuse, less numerous flagellomeres, wider face (Fig. 9B) and temples (Fig. 9E). Ophion al Johansson sp. nov. also lacks the typical colouration of O. bipictor, which has two spots in front of the scutellum and the subalar prominence whitish (Fig. 9F). Also similar to the Western Palaearctic species Ophion slaviceki Kriechbaumer, 1892 and O. kevoensis Jussila, 1965 but distinguished by the stouter flagellomeres, the more distinct carination of propodeum and the black ovipositor sheath. Description: Male unknown. Fore wing length 13‒14 mm. Body length 15‒16 mm. Antenna with 56 flagellomeres. Head buccate, temple in lateral view 0.8‒0.9 × as long as compound eye (Fig. 9E). Space between compound eye and lateral ocellus wide, about 0.2 × the diameter of ocellus. Face relatively wide (Fig. 9B). Occipital carina centrally very weakly angulate. Face below antennal sockets polished with distinct, but rather scarce punctures. Malar space about 0.1 × as long as mandibular base. Mandibular gape acute, but one female with indication of internal angles. Mandibles heavily worn in the two known specimens. First flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.2‒1.3 × as long as wide. Apical flagellomeres approximately 1.5 × as long as wide. Mesoscutum and mesopleuron strongly polished with distinct punctures, space between punctures about three × their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles right angled and distinct, anterior to sternal angles (Fig. 9D). Scutellum without lateral carinae. Propodeum with carination complete.Anterior transverse carina in one specimen becoming almost absent laterally. Sclerotised part of first sternite ending level to spiracle. Hind trochantellus as long as wide in dorsal view. Hind femur about 7.0 × as long as wide. Inner spur of hind tibia long, about 0.4 × as long as hind metatarsus. Wing membrane clear. Ramellus quite short, reaching 0.3 × the width of the discosubmarginal cell. Radius weakly sinuous, its inner part before junction with pterostigma straight. Nervellus broken slightly below the middle by the discoidella. Colouration: Body brownish reddish, quite dark compared to other Ophion species. Inner and outer eye margins whitish, the outer widely, the inner narrow. Mandibular teeth black. Ovipositor sheath black, contrasting in colour with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 168-170, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion breviflagellator Johansson sp. nov. urn:lsid:zoobank.org:act: 2F04FCE7-8DAD-4192-9C44-8B0C429A50D2 Figs 11A‒E Holotype: IRAN: ♀, Khorasan Province, Golulsara ni, 2800 m. a. s., 16‒17 Jul. 1997, Mirz./Ziai, (HMIM). Paratypes: IRAN: 1♀, Hormozgan province, Minab, Raveng, 27˚4’37.36"N, 57˚10’22.09"E, 34 m. a. s., 4 May 2016, Afsari / Mozh. leg. (HMIM); 1♀, Ardebil Province, Azlandooz, 39˚26’15.1"N, 47˚24’05.9"E, 182 m. a. s., Mogh. / Nas. leg. (HMIM); 1♂, Golestan Province, P. M. Golestan, 17‒18 Jul. 1996, 1650 m. a. s., Ebra. / V. Naz. leg. (HMIM) Type depositories: The holotype and paratypes is stored in HMIM, one female paratype is housed in NHRS. Etymology: The name refers to the stout basal flagellomeres. Diagnosis: Distinguished from other species of Ophion by the quite polished mesosoma with very scarce punctures, the carination of propodeum with complete carination, the anterior transverse carina conspicuously bent in front of the area superomedia and the stout flagellomeres. Description: Fore wing length 13‒15 mm. Body length 16‒18 mm. Antenna with 49‒53 flagellomeres. Temple narrowed behind compound eyes, in lateral view about 0.3 × as long as compound eye in female (Fig. 11D), 0.5 × in male. Space between compound eye and lateral ocellus narrow, about 0.1 × the diameter of ocellus in female and male. Face narrow (Fig. 11C). Occipital carina centrally angulate. Face below antennal sockets polished, centrally with distinct punctures, which becomes weaker laterally. Malar space about 0.2 × as long as mandibular base in female and 0.3 × in males. Mandibular gape almost right angled, with internal angles. First flagellomere 3.0 × as long as wide (Fig. 11E). Central flagellomeres stout, about 1.2‒1.3 × as long as wide. Apical flagellomeres approximately 1.4 × as long as wide. Mesoscutum and mesopleuron polished with punctures quite distinct, space between punctures about 2.0 × their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles strongly obtuse and indistinct, distinctly anterior to sternal angles. Scutellum with lateral carinae indicated in basal 0.2. Propodeum polished with very weak, rugulose sculpture. Carination complete. Anterior tranverse carina distinctly arcuate at upper part of area superomedia (Fig. 11B). Sclerotised part of first sternite ending posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 8.0 × as long as wide. Inner spur of hind tibia quite short, about 0.3 × as long as hind metatarsus. Wing membrane clear. Ramellus reaching 0.3 of the width of the discosubmarginal cell. Radius weakly sinuous, straight before junction with pterostigma. Nervellus broken in the middle or slightly above the middle by the discoidella. Colouration: Body yellow reddish, with the following paler whitish: inner and outer orbits, stemmaticum, four longitudinal stripes on mesoscutum (the central ones narrower), subalar prominence, speculum, a small spot in the lower posterior part of mesoscutum and one small spot corresponding on metapleuron, lateral margins of scutellum. Mandibular teeth black. Ovipositor sheath brown, slightly contrasting with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 172-174, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion sistanicus Johansson sp. nov. urn:lsid:zoobank.org:act: DD5A6DEB-B4C3-45E1-AD92-54E2EE618960 Figs 19A‒E Holotype: IRAN: ♀, Sistan-Baluchstan Province, Khash, Kousheh, 28˚37’38.7"N, 61˚00’56.3"E, 1715 m. a. s., Agu. 16‒30, 2015, 2583 m. 22 Apr. 2012, Afs. / Mogh. / Moj leg. (HMIM). Paratype: 1♂, same data as holotype, (HMIM). Type depositories: The holotype is stored in HMIM and the paratype in NHRS Etymology: The name refers to the Sistan-Baluchstan province in Iran where the types were collected. Diagnosis: A member of the Ophion minutus aggregate. Easily separable from Ophion minutus and O. golestanicus Johansson sp. nov. by the narrower face and the more whitish markings. Distinguished from the very similar Ophion schmidti Johansson sp. nov. by the more strongly polished mesosoma in the male and female, the more elongate flagellomeres, the narrower temples and the right-angled mandibular gape with the upper mandibular tooth slightly larger than the lower tooth. The polished mesosoma also distinguishes this species from Ophion persicus Johansson sp. nov. Description: Fore wing length 7 mm. Body length 8‒9 mm. Antenna with 44 flagellomeres. Head narrowed behind compound eyes, temple in lateral view 0.3 × as long as compound eye in female and male (Fig. 19B). Ocelli large, touching inner margin of compound eye. Face narrow (Fig. 19D). Occipital carina centrally slightly flattened. Face below antennal sockets shagreened with rather dense punctures in female. The interstices between punctures about equal to their diameter. Malar space absent. Mandibular gape right angled, without internal angles (Fig. 19C). First flagellomere 4.5 × as long as wide. Central flagellomeres slender, about 2.0 × as long as wide. Apical flagellomeres approximately 1.8 × as long as wide. Mesoscutum polished with virtually no punctuation. Mesopleuron in both sexes polished with distinct, but very scarce punctures, interstices between punctures about 3.0 × in female, 5.0 × in male. Epicnemial carina, in antero-ventral view, with pleurosternal angles obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum opaque, with weak rugulose sculpture. Propodeum with carinae entirely absent. Sclerotised part of first sternite ending distinctly posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 7.0‒8.0 × as long as wide. Inner spur of hind tibia long, about 0.4 × as long as hind metatarsus. Wing membrane clear. Ramellus distinct, short reaching 0.1 of the width of the discosubmarginal cell in the female, 0.3 × in the male. Radius very weakly sinuous, conspicuously bent and widened before junction with pterostigma. Nervellus almost straight broken distinctly below the middle by the discoidella. Colouration: Body red brownish, with the following paler whitish: head apart, from a small spot in front of anterior ocellus, a small triangular area below the antennal scrobes and anterior margin of clypeus; four longitudinal stripes on mesoscutum; subalar prominence; spots on mesopleuron; a spot in the lower posterior part of mesoscutum and one larger spot corresponding on metapleuron (Fig. 19E). Propodeum basally. Coxae with paler marks on the outer side. Metasoma with irregular pale markings on the posterior segments. Mandibular teeth black. Ovipositor sheath testaceous, concolourous with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 189, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion semipullatus Kokujev, 1909 Figs 16C‒E Ophion semipullatus Kokujev, 1909: 246‒247. Type ♀ collected in Uzbekistan, lost. Material examined: IRAN: 1♀, Hormozgan Province, Geno Mt. 1550 m. a. s., 18 Apr. 1994, Parch / Ebra. leg. (HMIM); 1♀, Bakhtiari, Zarbkuh, 2400 m. a. s., 16 Aug. 1976, Bruomand / Pazuki leg. (HMIM); 1♀, Bakhtiari / Ardol, Gandaman, kuhe-kallar, 2750 m. a. s., 13‒14 Jul. 1982, Bruomand / Pazuki leg. (HMIM). Remarks: A very distinct large and dark brownish species (Figs 16C, E) described from Buchara in present day Uzbekistan. The type is lost and the interpretation presented here is based on the original description and a series of females housed in the Meyer collection in ZIN. The dates on the labels indicate that the species occurs in two generations in Iran. The antennae has about 60 flagellomeres, but are noticeably short due to the very stout flagellomeres. New record for Iran., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 187, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Kokujev, N. R. (1909) Hymenoptera asiatica nova IX., Revue Russe d'Entomologie, 9, 246 - 248."]}

Ophion golestanicus Johansson sp. nov. urn:lsid:zoobank.org:act: AEB51640-1574-481B-BD1B-6361497A3E81 Figs 12A‒B, E Holotype: IRAN: ♀, Golestan Province, Dasht, 37˚21’N,56˚00’E, 1000 m.a.s., 11May 2009, A. Klimenko leg.(MR). Paratypes: IRAN: 12♀♀, 3♂♂, same data as holotype, (MR). Type depositories: The holotype is stored in ZSM, paratypes in MR according to original depository. Etymology: The name refers to the fact that the type series was collected in Golestan. Diagnosis: Similar to Ophion minutus Kriechbaumer, 1879, but usually with less pale markings on the mesosoma and on average slightly larger. The temples and the face is narrower and the flagellomeres are longer. In all known specimens, the posterior transverse carina is complete, m-shaped, while it usually is partly or completely missing in Ophion minutus. Also similar to the other three species of the O. minutus aggregate described in this study, but with more yellow pale markings, whitish on the other species, and with the face considerably wider. Description: Fore wing length 9‒10 mm. Body length 10‒12 mm. Antenna with 43‒46 flagellomeres. Temple buccate behind compound eyes, in lateral view about 0.7 × as long as compound eye in female and male (Fig. 12B). Space between compound eye and lateral ocellus narrow, about 0.2 × the diameter of ocellus in female and male. Face relatively narrow (Fig. 12A). Occipital carina centrally widely flattened. Face below antennal sockets with dense distinct punctures, the interstices between punctures about 0.2‒0.5 × the diameter of punctures. Malar space about 0.1 × as long as mandibular base in female and male. Mandibular gape slightly acute, without internal angles, but often worn. First flagellomere 4.0 × as long as wide. Central flagellomeres slender, about 2.0 × as long as wide. Apical flagellomeres approximately 2.0 × as long as wide (Fig. 12E). Mesoscutum polished with scarce punctures, space between punctures about 5.0 × their diameter. Mesopleuron weakly shagreened with distinct punctures, interstices between punctures about equal to their diameter Epicnemial carina, in antero-ventral view, with pleurosternal angles obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum polished with rugulose sculpture. Posterior transverse carina often complete, but in small specimens partly weakened. Sclerotised part of first sternite ending distinctly posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 7.0‒8.0 × as long as wide. Inner spur of hind tibia long, about 0.4‒0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus reaching 0.1‒0.2 of the width of the discosubmarginal cell. Radius very weakly sinuous, conspicuously bent and widened before junction with pterostigma. Nervellus broken in the middle or slightly below the middle by the discoidella. Colouration: Body yellow reddish, with the following paler yellow: mandibles, clypeus, inner and outer orbits, subalar prominence, tegulae, scutellum and postskutellum diffusely, a larger diffuse spot in front of the epicnemial carina on mesopleuron, sometimes a small diffuse spot in the lower posterior part of mesoscutum and one small spot corresponding on metapleuron. Mandibular teeth black. Ovipositor sheath testaceous-brown, concolorous with posterior metasomal segments. DNA Barcode: The DNA barcode sequence of one specimen of Ophion golestanicus sp. nov. is available at the BOLD systems database: sample id. NJP627, sequence id. LNU2463-21. Remarks: The barcoded specimen of Ophion golestanicus Johansson sp. nov. shows a minimum sequence divergence of 3.35% from its closest neighbor, Ophion minutus in BOLD., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 175-177, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Kriechbaumer, J. (1879) Ophion minutus n. sp. Entomologische Nachrichten, 5, 105 - 106."]}

Ophion sinuosus Johansson sp. nov. urn:lsid:zoobank.org:act: 569E4C1C-8FBF-42F4-863B-187E75B42691 Figs 18A‒D Holotype: IRAN: ♀, Khorasan Province, Shomali-Ghuchan-Chakane, dga/ Pahne, 1780 m. a. s., 11‒12 Jun. 2009, Mof. leg. (HMIM). Paratype: IRAN: 1♀, same data as holotype (HMIM). Type depositories: The holotype female stored in HMIM and the paratype female stored in NHRS Diagnosis: Similar to Ophion rostralis (as here defined and delimited), but with fewer flagellomeres, wider face in anterior view and different carination of the propodeum with the anterior transverse carina distinctly sinuate, the posterior transverse carina only present laterally and all trace of central and lateral longitudinal carinae missing. Etymology: The name refers to the sinuate anterior transverse carina of the propodeum. Description: Fore wing length 14‒15 mm. Body length 15‒16 mm.Antenna with 49‒50 flagellomeres. Temple in lateral view narrow, about 0.3‒0.4 × as long as compound eye in female (Fig. 18C). Lateral ocelli touching inner margin of compound eye. Face wide (Fig. 18D). Occipital carina centrally evenly rounded. Face below antennal sockets strongly polished with very weak and scattered punctures. Malar space about 0.2 × as long as mandibular base (fig. 18D). Mandibular gape acute, without internal angles. First flagellomere 3.5 × as long as wide. Central flagellomeres stout, about 1.3 × as long as wide. Apical flagellomeres approximately 1.3 × as long as wide. Mesoscutum strongly polished without any obvious puncture. Mesopleuron polished with punctures quite dense, but very weak. Epicnemial carina, in antero-ventral view, with pleurosternal angles weakly obtuse, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum strongly polished. Anterior transverse carina distinct and sinuate (Fig. 18B). Posterior transverse carina strong laterally, centrally widely interrupted. Longitudinal carinae absent. Sclerotised part of first sternite level to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur quite stout, about 6.0‒7.0 × as long as wide. Inner spur of hind tibia short, about 0.3 × as long as hind metatarsus. Wing membrane clear. Ramellus absent or short, reaching 0.2‒0.3 the width of the discosubmarginal cell. Radius weakly sinuous in outer part, its inner part straight before junction with pterostigma. Nervellus broken slightly below the middle by the discoidella. Male unknown. Colouration: Body pale brownish, light compared to other Ophion species. Head and mesosoma with pale pattern reminiscent of the members of the Ophion parvulus aggregate. Mandibular teeth black. ovipositor sheath concolorous with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 187-189, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion pravinervis Kokujev, 1906 Figs 15A‒E Ophion pravinervis Kokujev, 1906: 166. Type from Georgia, lost. Material examined: IRAN: 1♀, Kohkiloyeh-va-Boyerahmad Province, Yasuj area, 10 km SE Sepidar, 5‒6 May 2007, A. Klimenko leg. (MR). Redescription female: Fore wing length 11 mm. Body length 13.5 mm.Antenna with 49 flagellomeres. Temple in lateral view narrow 0.5 × as long as compound eye in female (15E). Space between compound eye and lateral ocellus narrow, about 0.1 × the diameter of ocellus. Face narrow (Fig. 15D). Occipital carina centrally flattened. Face below antennal sockets polished with very distinct and dense punctures. Punctures often merging. Clypeus with larger, slightly scarcer punctures. Malar space very narrow about 0.05 × as long as mandibular base. Mandibular gape acute, mandibles in the examined specimen without internal angles. First flagellomere 3.5 × as long as wide. Central flagellomeres quite stout, about 1.5‒1.6 × as long as wide. Apical flagellomeres approximately 1.7 × as long as wide. Mesoscutum polished with distinct punctures, space between punctures about three × their diameter. Mesopleuron shagreened with quite small dense punctures that often merge (Fig. 15B). Epicnemial carina, in anteroventral view, with pleurosternal angles right angled and distinct, anterior to sternal angles. Scutellum without lateral carinae. Propodeum with quite coarse rugulose sculpture, centrally with two almost parallel carinae which are joined anteriorly by a very short transverse carina that is completely absent laterally (Fig. 15C). The parallel carinae does not extend onto the petiolar area. Posterior transverse carina distinct laterally, widely interrupted centrally. Hind femur about 6.0 × as long as wide. Inner spur of hind tibia long, about 0.3 × as long as hind metatarsus, about two × as long as outer spur. Wing membrane clear. Ramellus quite short, reaching 0.2 × the width of the discosubmarginal cell. Radius weakly sinuous, its inner part before junction with pterostigma conspicuously bent (Fig. 15A). Nervellus broken distinctly below the middle by the discoidella. Male unknown. Colouration: Body brownish reddish, quite dark compared to other Ophion species. Inner and outer eye margins whitish, the outer widely, the inner narrow. Mandibular teeth black. Subalar prominence with a whitish spot. Scutellum and postscutellum whitish. Scutellum with a brownish spot centrally at the anterior margin. Posterior metasomal segments infuscate ventrally. Ovipositor sheath concolorous with posterior metasomal segments. DNA Barcode: The DNA barcode sequences of one specimen of Ophion pravinervis is available at the BOLD systems database: sample id. NJP745, sequence id. LNU2581-21. Remarks: Ophion pravinervis is previously only known from the now lost type material collected in Georgia (Kokujev 1906). Proposed as a senior synonym of Ophion cortesi Ceballos, 1938 in Rey del Castillo and Izquierdo Moya, based on communication with Pier-Luigi Scaramozzino (1990). This is clearly erroneous as O. cortesi is a valid species of different constitution., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 182-184, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion angularis Johansson & Cederberg, 2019 Ophion angularis Johansson & Cederberg, 2019: 37; Figs 9F, 22. Type ♀ in NHRS, examined. Material examined: IRAN: 1♀, 3♂♂, Golestan Province, 20 km S Gorgan city, 36˚43’N, 54˚35’E, 700 m. a. s., 10 May 2009, A. Klimenko leg. (MR). Remarks: Recently described from Northern Europe and the species has since also been recorded from Central and Southern Europe. It seem to prefer semi open xerothermic areas, but the host is still unknown., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 170, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion vasi Johansson sp. nov. urn:lsid:zoobank.org:act: 6EBBB168-DABB-4B10-B2F0-5A9EF523A696 Figs 21A‒E Holotype: IRAN: ♀, Lorestan Province, Khorramabad area, Mahmudvand, 30 Apr. 2007, A. Klimenko leg. (MR). Paratype: IRAN: 1♀, Hormozgan Province, Minab, Raveng, 27˚4’37.36"N, 57˚10’22.09"E, 34 m. a. s., 4 May 2016, Afsari / Mozh leg. (HMIM). Type depositories: Holotype stored in HMIM, paratype in NHRS Etymology: Named in honour of the Hungarian hymenopterist Zoltán Vas. Diagnosis: With the pale markings of the mesosoma reminiscent of members of the Ophion obscuratus aggregate, however distinguished by the combination of the narrow face, the narrow temples, the very weakly developed pleurosternal angles and the uncarinated or almost uncarinated and polished propodeum. Description: Male unknown. Fore wing length 14 mm. Body length 15 mm.Antenna with 56‒57 flagellomeres. Temples in lateral view narrow, about 0.4 × as long as compound eye. Lateral ocelli touching compound inner margin of compound eye (Fig. 21C). Face narrow (Fig. 21D). Occipital carina centrally evenly rounded. Face below antennal sockets shagreened with dense, irregular punctures that partially merge. Malar space about 0.1 × as long as mandibular base. Mandibular gape right angled, with internal angles. First flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.4‒1.5 × as long as wide. Apical flagellomeres approximately 1.6 × as long as wide. Mesoscutum polished with quite dense, but very weak and irregular punctuation. Mesopleuron polished with punctures quite dense, space between punctures about equal to two × their diameter. Epicnemial carina, in anteroventral view, with pleurosternal angles strongly obtuse and indistinct, distinctly anterior to sternal angles (Fig. 21E). Scutellum without lateral carinae. Propodeum polished with only the anterior transverse carina weakly indicated centrally in one of the types (Fig. 21B). Sclerotised part of first sternite ending level to spiracle. Hind trochantellus as long as wide in dorsal view. Hind femur slender, about 8.0 × as long as wide. Inner spur of hind tibia short, about 0.3 × as long as hind metatarsus. Wing membrane clear. Ramellus reaching 0.3 the width of the discosubmarginal cell. Radius weakly sinuous, straight before junction with pterostigma. Nervellus broken in the middle or slightly above the middle by the discoidella. Colouration: Body yellow reddish, with the following paler whitish: inner and outer orbits, four longitudinal stripes on mesoscutum (the central ones narrower), subalar prominence, scutellum laterally and postscutellum, a diffuse spot on the speculum or below the subalar prominence, a small spot in the lower posterior part of mesoscutum and one larger spot corresponding on metapleuron. Mandibular teeth black. Ovipositor sheath brown, slightly contrasting with posterior metasomal segments., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 193, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion xrafstra Johansson sp. nov. urn:lsid:zoobank.org:act: FCCABB31-30E6-4C3C-8DB2-9622323C7935 Figs 22A‒E Holotype: IRAN: ♀, Khorasan Province, Shomali-Ghuchan-Chakane, dga/ Pahne, 1780 m. a. s., 11‒12 Jun. 2009, Mof. leg. (HMIM) Paratype: 1♀, same data as the holotype. Type depositories: The holotype is stored in HMIM and the paratype is stored in NHRS. Etymology: The name refer to xrafstra, which are evil, usually nocturnal spirits in Persian mythology that entered the human world, sometimes disguised as wasps. Diagnosis: Similar to Ophion turcomanicus, but with narrower face and fewer teeth on the hind tarsal claws. Description: Male unknown. Fore wing length 15 mm. Body length 18 mm.Antenna with 44‒45 flagellomeres. Temples in lateral view buccate, about 0.8 × as long as compound eye. Gap between lateral ocelli and inner margin of compound eye wide, about 0.2‒0.3 × the diameter of ocellus (Fig. 22C). Face relatively narrow (Fig. 22E). Occipital carina centrally evenly rounded. Face below antennal sockets strongly polished with weak very scarce punctures. Malar space about 0.1 × as long as mandibular base. Mandibular gape strongly acute and mandibular teeth very long (Fig. 22E). First flagellomere 4.0 × as long as wide. Central flagellomeres stout, about 1.4‒1.5 × as long as wide. Apical flagellomeres approximately 1.6 × as long as wide. Mesoscutum strongly polished with quite dense, but very weak and irregular punctuation. Mesopleuron polished with very weak punctures, space between punctures about two × their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles strongly obtuse and indistinct, distinctly anterior to sternal angles. Scutellum without lateral carinae. Propodeum quite polished with only the anterior transverse carina distinct throughout its length. Posterior transverse carina widely interrupted centrally (Fig. 22B). Two short parallel carinae present centrally at the indicated junction with the posterior transverse carina. Sclerotised part of first sternite ending level to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur about 6.0 × as long as wide. Inner spur of hind tibia about 0.3 × as long as hind metatarsus, outer spur 0.6 × as long as outer spur. Hind tarsal claws with about six spines with quite large insterstices (Fig. 22D). Wing membrane clear. Ramellus reaching 0.2 × the width of the discosubmarginal cell. Radius weakly sinuous in outer part, straight before junction with pterostigma. Nervellus broken in the middle or slightly below the middle by the discoidella. Colouration: Body pale yellowish with typical coloration of desert species (Fig. 22A). Mandibular teeth black., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 196, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion klimenkoi Johansson sp. nov. urn:lsid:zoobank.org:act: 38AFA9D4-A900-4EAA-8C2E-E2B9417734D0 Figs 13A‒E Holotype: IRAN: ♀, Khorasan Province, SW part, Khezri, env., 33˚56’N, 58˚43’E, 1800 m. a. s., 23 May 2009, A. Klimenko leg. (MR). Paratypes: IRAN: 1♂, Khorasan Province, SW part, Khezri, env., 33˚56’N, 58˚43’E, 1800 m. a. s., 23 May 2009, A. Klimenko leg. (MR); 1♀, Khorasan Province, SE part, Q’aen area, Khunik pass, 33˚33’N, 59˚08’E, 2100 m. a. s., 25 May 2009, A. Klimenko leg. (MR). Type depositories: The holotype is housed in the ZSM, paratypes in MR. Etymology: The name refers to A. Klimenko who collected the only known specimens of the species. Diagnosis: Possibly a member of the O. mocsaryi species group. Distinguished from the other known members of the aggregate by the very densely and partly rugulose face in combination with the obtuse pleurosternal angles, which are situated distinctly anterior to the sternal angles. Description: Fore wing length 12.5‒13 mm. Body length 14‒15 mm. Antenna with 48‒54 flagellomeres. Temple narrowed behind compound eyes, in lateral view about 0.4 × as long as compound eye in female and male. Space between compound eye and lateral ocellus narrow, about 0.1‒0.2 × the diameter of ocellus in female and male. Occipital carina centrally evenly rounded. Face below antennal sockets shagreened with very dense punctures that partly form rugulose sculpture (Fig. 13E). Malar space narrow about 0.2 × as long as mandibular base in female and male (Fig. 13C). Mandibular gape acute, without internal angles, as in O. slaviceki Kriechbaumer. Antennae stout, first flagellomere 3.0 × as long as wide. Central flagellomeres stout, about 1.2‒1.4 × as long as wide. Apical flagellomeres approximately 1.4 × as long as wide. Mesoscutum strongly polished, virtually without puctures. Mesopleuron weakly shagreened with punctures quite dense, space between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles strongly obtuse and indistinct, distinctly anterior to sternal angles (Fig. 13D). Scutellum without lateral carinae. Propodeum with the anterior transverse carina in some specimens forming an arcuate lobe centrally, with very weak rugulose sculpture, opaque. Posterior transverse carina indicated laterally (Fig. 13B). Propodeum polished, anterior to anterior transverse carina with distinct scarce puncture. Carination ranging from almost complete with only the posterior carina adjacent to area supeomedia absent to having the posterior carina and the lateral parts of the anterior transverse carina absent. Sclerotised part of first sternite ending posterior to spiracle. Hind trochantellus shorter than wide in dorsal view. Hind femur slender, about 8.0 × as long as wide. Inner spur of hind tibia long, about 0.5 × as long as hind metatarsus. Wing membrane clear. Ramellus absent or short, reaching 0.3 × the width of the discosubmarginal cell. Radius weakly sinuous, almost straight before junction with pterostigma. Nervellus broken below the middle by the discoidella. Colouration: Body testaceous. Inner and outer eye margins yellow. Mandibular teeth black. ovipositor sheath dark brown, contrasting in colour with posterior metasomal segments. DNA Barcode: The DNA barcode sequences of two specimens of Ophion klimenkoi sp. nov. are available at the BOLD systems database: sample id. NJP631/632, sequence id. LNU2467-21/2468-21. Remarks: A member of the O. mocsaryi -aggregate most closely related to Ophion rostralis, from which it shows a minimum sequence divergence of 1.79%., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 177-179, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion mocsaryi Brauns, 1889 Ophion mocsaryi Brauns, 1889: 89. Lectotype ♀ from Hungary in HMHN, examined. Material examined: IRAN: 1♀, Semnan Province, Shakhmirzad area, Chasm env. 35˚49’N, 53˚15’E 2200 m. a. s., 14 Jun. 2009, A. Klimenko leg. (MR); 1♀, Elburz Geb. Südseite, 30 km N. of Teheran, 1800 m. a. s., 27 Aug. 1971, Vartian leg. (OÖLM). Remarks: This species belongs to a species group of closely related species commonly treated under the name Ophion mocsaryi Brauns. The here treated species fits with the interpretation presented by Johansson & Cederberg, (2019). However, one should be aware that the presumably aggregate nature might call for future adjustements in nomenclature. The species was only recently properly defined (Johansson & Cederberg 2019) and any references in literature preceeding that date of publication may refer to any closely related and similar species., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 180, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion fossulatus Hedwig, 1957 stat. rev. Ophion fossulatus Hedwig, 1957: 106. Holotype ♀ in SMNS from Iran, examined. Material examined: IRAN: Holotype ♀, Iran, Masanderan, Sulede, 27. April 1951. 100 m. a. s., Shäuffele leg. (SMNS); 5♀♀, 4♂♂, Teheran Province, Firooz kuh, 35˚45’48.21"N, 52˚45’24.40"E, 1981 m. a. s., 10 Jun. 2007, Lofalizadeh leg. (HMIM); 1♀, Kohkiloye-va-Boyerahmad Province, Yasuj area, 10 km SE Sepidar, 5‒6 May 2007, A. Klimenko leg. (MR); 5♀♀, 4♂♂, Golestan Province, Dasht, 37˚21’N, 56˚00’E, 1000 m. a. s., 11 May 2009, A. Klimenko leg. (MR); 1♀, Lorestan Province, Khorramabad area, Mahmudvand, 30 Apr. 2007, A. Klimenko leg. (MR); 1♀, Golestan Province, Elburs mts, Maghazy Valley, Lux, 36˚64’92.3"N, 54˚20’70.1"E, 11 Jun. 2007, 1630 m. a. s., N. Pöll leg. (OÖLM). DNA Barcode: The DNA barcode sequences of two specimens of Ophion fossulatus are available at the BOLD systems database: sample id. NJP742/744, sequence id. LNU2578-21/2580-21. Remarks: Previously synonymized with Ophion mocsaryi Brauns, 1889. Study of the type shows that this species belongs to the Ophion parvulus aggregate, which is characterized by the evenly curved radius and the weakly defined pleurosternal angles. Very similar to the Northern and Central European species Ophion parvulus Kriechbaumer, but with the carination of propodeum more strongly raised, usually both anterior and posterior transverse carinae and central and lateral longitudinal carinae complete, and the pleurosternal angles more prominent. The two barcoding sequences that were extracted, cluster in BOLD with O. parvulus and obviously the two species are very closely related. The minor differences in morphology motivates the treatment of O. fossulatus as a separate species, but undoubtedly the taxonomy of the aggregate would benefit from a more thorough study using integrative methods., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 175, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655, {"references":["Hedwig, K. (1957) Ichneumoniden und Braconiden aus den Iran 1954 (Hymenoptera)., Jahresheft des Vereins fur Vaterlaendische Naturkunde, 112 (1), 103 - 117."]}

Ophion turcomanicus Szépligeti, 1905 Figs 15F‒G Ophion turcomanicus Szépligeti, 1905: 521. Lectotype ♀ and paralectotype from Turkmenistan in HMHN, examined. Ophion asiaticus Kokujev, 1906: 167‒168. Lectotype ♀ from Iran in ZIN, examined. Ophion obscuratus var. heratensis Hedwig, 1961:291. Syn. nov. Holotype ♀ from Afghanistan in SMNK, examined. Material examined: IRAN: Lectotype ♀ of Ophion asiaticus, Western Iran (ZIN); 1♀, Teheran Province, Shahriar, Khoshnam, 19‒20 May 1991, Ebra./ Badii leg. (HMIM); 1♀, Sistan-Baluchstan Province, Khash, Kousheh, 28˚37’38.7"N, 61˚00’56.3"E, 1715 m. a. s., 16‒30 Aug. 2015, 2583 m. 22 Apr. 2012, Afs. / Mogh. / Moj leg. (HMIM); 1♀, Khorasan Province, Torbat-e-Haydarieh area, Zharf v., 14 May 2007, 1600 m. a. s., A. Klimenko leg. (MR); 1♀, Khorasan Province, SE part, Q’aen area, Khunik pass, 33˚33’N, 59˚08’E, 2100 m. a. s., 25 May 2009, A. Klimenko leg. (MR); 1♀, Chaharmahal & Bakhtiari, Kuhrang, Hiregan, 32˚20’N, 50˚26’E, 2305 m. a. s., 6 Jun. 2005, Haj / Ebra leg. (HMIM); 1♀, W. Azerbaijan Province, Rezaieh, Ghushchi, 9 Jun. 1975, Abai leg. (HMIM); 1♀ (“obscuratus heratensis” Hedwig 1961), Khorasan Province, Western part, Bojnurd area, Gerivan N, 37˚17’N, 57˚13’E, 12 May 2009, A. Klimenko leg. (MR) (this specimen has very large ocelli and less prominent pleurosternal angles). Remarks: Similar to Ophion eremita Kokujev but distinguished by the less numerous flagellomeres (44‒50) (44 and 46 in the studied types of O. turcomanicus, 50 in the type of O. asiaticus), the often complete, but rather weak carination of propodeum, which usually have at least the anterior and posterior transverse carinae present and the area superomedial discernible. The mandibles, which are often worn, are similar to Ophion slaviceki with an acute angled gap without internal angles and thereby differing from the mandibles of O. eremita, which usually have distinct internal angles in a right-angled gap. On average also smaller than O. eremita. The here studied specimens display a considerable amount of variation which may indicate cryptic species. O. turcomanicus and O. eremita belong to an aggregate of similar and closely related desert and steppe species also including Ophion geyri Habemehl, 1921 and a couple of undescribed species. Of particular interest is a single female in coll. MR with unusually large ocelli and weakly defined pleurosternal angles. This specimen corresponds with the holotype of Ophion obscuratus var. heratensis Hedwig, 1961 housed in the SMNK. It is here treated as a variety of Ophion turcomanicus, but if this form is found to be a valid species, it has to be assigned a new name given the fact that it was described as a variety in 1961., Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on page 192, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/5225655

Ophion obscuratus FABRICIUS,1798 M a t e r i a l e x a m i n e d: Markazi site 4: 1♀ 6.-10.v.2018, leg. M. Parachami-Araghi; Markazi site 1: 2♀♀ 1♂ 11.vi.-20.vii.2018, leg. M. Parachami-Araghi; Zanjan site 2: 1♂ 21.v.2016, leg. H. Nasserzadeh. D i s t r i b u t i o n: Palaearctic and Oriental regions, known from Iran (BARAHOEI et al. 2012)., Published as part of Riedel, Matthias, Ameri, Ali, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2019, Contribution to the Ɨchneumonidae (Hymenoptera) oI Ɨran, with descriptions oI seven new species, pp. 361-390 in Linzer biologische Beiträge 51 (1) on page 378, DOI: 10.5281/zenodo.3758345, {"references":["BARAHOEI H., RAKHSHANI E. & M. RIEDEL (2012): A checklist of Ichneumonidae (Hymenoptera: Ichneumonoidea) from Iran. 2 Iranian Journal of A nimal Biosystematics 8: 83 - 132."]}