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Three types of ichnites from a new site discovered in Imilchil were studied and described. The structures of pes and manus prints and tail traces of tetrapods provide data to separate: real ichnites (theropod footprints and crocodile tail drag marks); and sauropod’ subtraces. Most of the theropod footprints are deformed by collapse of their walls, the sauropod subtraces show the strong deformation of about ten well-stratified sedimentary levels. The surface on which we can see theropod and crocodile ichnites is the tracking surface, i. e. the original surface on which they were printed, Se estudian tres tipos de icnitas de un nuevo yacimiento descubierto en Imilchil. Las estructuras de las huellas de pies, manos y cola de tetrápodos proporcionan datos para separar: icnitas reales (pisadas terópodas y marcas de arrastre de cola de cocodrilos) y calcos saurópodos? La mayor parte de las huellas terópodas están deformadas por colapso de sus paredes, y las subhuellas saurópodas muestran la fuerte deformación de unos diez niveles sedimentarios bien estratificados. La superficie en la que se encuentran las huellas terópodas y las icnitas de cocodrilo son la superficie de marcha, es decir la superficie original sobre las que se imprimieron

A new small site, not always outcropping, with large and giant theropod tracks in the Upper Jurassic of the Central High Atlas of Morocco is described. The ichnites and the typology of the rocks of the site are similar to those of the Iouaridène sites with which there is no cartographic continuity. The particular characters of the tracks and trackways are analyzed and highlighted and a hypothesis of distribution of this type of dinosaurs is pointed out Se describe un nuevo yacimiento pequeño, no siempre aflorante, con huellas terópodas grandes y gigantes en el Jurásico Superior del Alto Atlas Central de Marruecos. Las icnitas y la tipología de las rocas en las que yacen son similares a las de los yacimientos de Iouaridène con los que no hay continuidad cartográfica. Se analizan y destacan los caracteres particularesde las huellas y rastrilladas y se apunta una hipótesis de distribución de este tipo de dinosaurios

A new small site, not always outcropping, with large and giant theropod tracks in the Upper Jurassic of the Central High Atlas of Morocco is described. The ichnites and the typology of the rocks of the site are similar to those of the Iouaridène sites with which there is no cartographic continuity. The particular characters of the tracks and trackways are analyzed and highlighted and a hypothesis of distribution of this type of dinosaurs is pointed out, Se describe un nuevo yacimiento pequeño, no siempre aflorante, con huellas terópodas grandes y gigantes en el Jurásico Superior del Alto Atlas Central de Marruecos. Las icnitas y la tipología de las rocas en las que yacen son similares a las de los yacimientos de Iouaridène con los que no hay continuidad cartográfica. Se analizan y destacan los caracteres particularesde las huellas y rastrilladas y se apunta una hipótesis de distribución de este tipo de dinosaurios

Se describe un nuevo yacimiento con icnitas saurópodas del Cretácico Superior encontrado en Chott Tigri, en la zona de la Meseta Meridional Marroquí, entre el Atlas SaharianoyelAtlas Medio, separada del resto de yacimientos marroquíes. En conjunto hay tres rastrilladas titanosauriformes de ancho de via variable y otra de un dinosaurio desconocido que atribuimos a un ornitisquio cuadrúpedo no ornitópodo. En este trabajo se han aplicado cálculos basados en el análisis de las rastrilladas para deducir la distancia glenoacetabular y el tipo de marcha; en las tres rastrilladas saurópodas son congmentes con marcha amblar, In this paper a new Upper Cretaceous site with sauropod footprints is described . The outcrop, found in Chott Tigri, in the meridional are a of the High Plateau of Morocco, between the Saharan Atlas and the Middle Atlas is separate from other known sites with dinosaur footprints. We have found: three trackways of titanosauriform featu res and variable gauge, and o n e tra ckw ay o f an unknown dinosaur that we attribute to an ornithischian quadruped, but not an ornithopod. In this paper we have implemented calculations based on trackway analysis to deduce the glenoacetabular distance distance and the type of gait. In the three sauropod trackways the data are consistent with amble gait

Se describe un nuevo yacimiento con icnitas saurópodas del Cretácico Superior encontrado en Chott Tigri, en la zona de la Meseta Meridional Marroquí, entre el Atlas SaharianoyelAtlas Medio, separada del resto de yacimientos marroquíes. En conjunto hay tres rastrilladas titanosauriformes de ancho de via variable y otra de un dinosaurio desconocido que atribuimos a un ornitisquio cuadrúpedo no ornitópodo. En este trabajo se han aplicado cálculos basados en el análisis de las rastrilladas para deducir la distancia glenoacetabular y el tipo de marcha; en las tres rastrilladas saurópodas son congmentes con marcha amblar In this paper a new Upper Cretaceous site with sauropod footprints is described . The outcrop, found in Chott Tigri, in the meridional are a of the High Plateau of Morocco, between the Saharan Atlas and the Middle Atlas is separate from other known sites with dinosaur footprints. We have found: three trackways of titanosauriform featu res and variable gauge, and o n e tra ckw ay o f an unknown dinosaur that we attribute to an ornithischian quadruped, but not an ornithopod. In this paper we have implemented calculations based on trackway analysis to deduce the glenoacetabular distance distance and the type of gait. In the three sauropod trackways the data are consistent with amble gait

Se describen dos huellas crocodyliformes de “les couches rouges” del Jurásico Medio del Alto Atlas de las que se destaca su importancia por su escasez en el registro mundial. Las huellas no son comparables con los icnogéneros descritos hasta ahora. La huella del pie es incompleta. Suponemos que es un crocodiliforme continental porque el ambiente sedimentario en el que están las icnitas es fluvial. Finalmente se indica, aunque con unos márgenes de inseguridad amplios, el orden de tamaño que suponemos para el autor de las pisadas Two Crocodyliform footprints from "les couches rouges" of the Middle Jurassic of the High Atlas are described.We highlight it for its scarcity in the global record. Footprints cannot be compared with other known ichnogenus. The pes print is incomplete.We assume that it is a continental crocodyliform because the footprints are in fluvial deposits. Finally there follows, although with wide margins of uncertainty, the size that we assume for the trackmake

Se describen dos huellas crocodyliformes de “les couches rouges” del Jurásico Medio del Alto Atlas de las que se destaca su importancia por su escasez en el registro mundial. Las huellas no son comparables con los icnogéneros descritos hasta ahora. La huella del pie es incompleta. Suponemos que es un crocodiliforme continental porque el ambiente sedimentario en el que están las icnitas es fluvial. Finalmente se indica, aunque con unos márgenes de inseguridad amplios, el orden de tamaño que suponemos para el autor de las pisadas, Two Crocodyliform footprints from "les couches rouges" of the Middle Jurassic of the High Atlas are described.We highlight it for its scarcity in the global record. Footprints cannot be compared with other known ichnogenus. The pes print is incomplete.We assume that it is a continental crocodyliform because the footprints are in fluvial deposits. Finally there follows, although with wide margins of uncertainty, the size that we assume for the trackmake

Se citan cuatro yacimientos en Ansous con icnitas terópodas, saurópodas y tireóforas de los que se describen una parte de las huellas terópodas y las tireóforas.Aunque se han publicado otras tres referencias de huellas tireóforas en Marruecos, esta es la primera asignación fundamentada de huellas cuadrúpedas de este tipo y de edad Jurásico Inferior (Pliensbachiense) en el país. Se proporcionan los datos biomórficos y morfométricos de las improntas terópodas y de las tireóforas. La atribución se hace por los caracteres biomórficos de las icnitas y por la edad de las rocas en las que están los yacimientos. Es recomendable continuar con la prospección en esta zona debido al excepcional potencial paleoicnológico, In this paper thyreophoran and theropod footprints contained in two of four new sites found in Ansous are described. In one of the sites there are also preserved sauropod tracks. Although there have been cited three other references to thyreophoran footprints in Morocco, this is the first of such quadrupedal footprints from the Lower Jurassic age (Pliensbachian) in the country.We provide biomorphic and morphometric data of booth footprint types (theropod and thyreophoran). The assignation is made by the biomorphic characters of the ichnites and the age of the rocks on which the sites are placed.We recommend continuing to explore in this area due to its exceptional potential of dinosaur palaeoichnology

Se citan cuatro yacimientos en Ansous con icnitas terópodas, saurópodas y tireóforas de los que se describen una parte de las huellas terópodas y las tireóforas.Aunque se han publicado otras tres referencias de huellas tireóforas en Marruecos, esta es la primera asignación fundamentada de huellas cuadrúpedas de este tipo y de edad Jurásico Inferior (Pliensbachiense) en el país. Se proporcionan los datos biomórficos y morfométricos de las improntas terópodas y de las tireóforas. La atribución se hace por los caracteres biomórficos de las icnitas y por la edad de las rocas en las que están los yacimientos. Es recomendable continuar con la prospección en esta zona debido al excepcional potencial paleoicnológico In this paper thyreophoran and theropod footprints contained in two of four new sites found in Ansous are described. In one of the sites there are also preserved sauropod tracks. Although there have been cited three other references to thyreophoran footprints in Morocco, this is the first of such quadrupedal footprints from the Lower Jurassic age (Pliensbachian) in the country.We provide biomorphic and morphometric data of booth footprint types (theropod and thyreophoran). The assignation is made by the biomorphic characters of the ichnites and the age of the rocks on which the sites are placed.We recommend continuing to explore in this area due to its exceptional potential of dinosaur palaeoichnology

En este artículo se estudia un nuevo yacimiento icnológico (34IGR) del sinclinal de Iouaridène, citado previamente por otros autores, pero no descrito. Las icnitas se caracterizan por su irregularidad, heterogeneidad y variación de tamaño. Los caracteres biomórficos y morfométricos tan variables son la respuesta, mostrada en el conjunto del sinclinal, a la diversidad biológica, morfológica y de estructuras inducidas por las propiedades físicas del barro. Este afloramiento es también la expresión de la variación de la cuenca, porque en este sector solo hay una capa con icnitas, frente a los yacimientos del oeste en los que hay superpuestas un número aún no determinado de capas fosilíferas, In this article is studied a new ichnological site (34IGR) of the Synclinal of Iouaridène already mentioned by the authors though still not described. The footprints are characterized by the ichnological heterogeneity and irregularity of his forms and dimensions. Both biomorphic and morphometricv characters answer to the variability showed in the ensemble of sinclinal sites relative to the biological diversity and to the morphology and structural response induced on the footprints by the physical characteristics of the mud. This site is also an expression of the evolution of the basin since he is in a sector that alone has one ichnological bed, against the deposits of the western part in which has been detected the overlapping of a number, still not assessed, of beds and levels with footprints

Se describe por primera vez un yacimiento con huellas de dinosaurio (Oumzawrou o 36IGR), que era conocido desde hace cuarenta años.Es el segundo en cuanto al número de huellas del conjunto de 43 yacimientos del sinclinal de Iouaridène (IGR) . En 36IGR concurren varias particularidades en relación con los demás de IGR que lo hacen destacable: a) las icnitas están en un solo nivel al contrario de lo que ocurre en el resto de yacimientos estudiados del sinclinal; b) el número de icnitas y rastrilladas semiplantígradas es relativamente muy grande; c) no hay rastrilladas con dirección E-O. La concentración y la cantidad de de huellas semiplantígradas es un hecho excepcional tanto en IGR como entre los yacimientos similares conocidos del resto del mundo. Dadas la condiciones estratigráficas, icnológicas y paleoambientales del yacimiento se postula que una relación debe condicionar la coexistencia de icnotipos y la conducta de los dinosaurios, We describe for the first time a dinosaur tracksite (Oumzawrou or 36IGR) which was known for forty years. It is the second in the number of prints of all 43 sites discovered in the Iouaridène syncline (IGR) In 36IGR concur several particularities that make it remarkable, in relation to others of IGR: a) the footprints are only on a single level, as opposite to the other tracksites analysed in the syncline; b) the number of semiplantigrade footprints and trackways is relatively very large; c) there are no trackways with E-W direction. The concentration and number of semiplantigrade footprins is an exceptional fact both in IGR and in similar outcrops in the world. According to the stratigraphic, ichnologic and palaeoenvironmental conditions of the tracksite is postulated that a relation must restrict the coexistence of ichnotypes and the behaviour of dinosaurs

Se describe por primera vez un yacimiento con huellas de dinosaurio (Oumzawrou o 36IGR), que era conocido desde hace cuarenta años.Es el segundo en cuanto al número de huellas del conjunto de 43 yacimientos del sinclinal de Iouaridène (IGR) . En 36IGR concurren varias particularidades en relación con los demás de IGR que lo hacen destacable: a) las icnitas están en un solo nivel al contrario de lo que ocurre en el resto de yacimientos estudiados del sinclinal; b) el número de icnitas y rastrilladas semiplantígradas es relativamente muy grande; c) no hay rastrilladas con dirección E-O. La concentración y la cantidad de de huellas semiplantígradas es un hecho excepcional tanto en IGR como entre los yacimientos similares conocidos del resto del mundo. Dadas la condiciones estratigráficas, icnológicas y paleoambientales del yacimiento se postula que una relación debe condicionar la coexistencia de icnotipos y la conducta de los dinosaurios We describe for the first time a dinosaur tracksite (Oumzawrou or 36IGR) which was known for forty years. It is the second in the number of prints of all 43 sites discovered in the Iouaridène syncline (IGR) In 36IGR concur several particularities that make it remarkable, in relation to others of IGR: a) the footprints are only on a single level, as opposite to the other tracksites analysed in the syncline; b) the number of semiplantigrade footprints and trackways is relatively very large; c) there are no trackways with E-W direction. The concentration and number of semiplantigrade footprins is an exceptional fact both in IGR and in similar outcrops in the world. According to the stratigraphic, ichnologic and palaeoenvironmental conditions of the tracksite is postulated that a relation must restrict the coexistence of ichnotypes and the behaviour of dinosaurs

During the 2009's summer courses (universities Rabat-La Rioja) 6 new paleoichnological sites are found in the Bou Guemez river. The previously know Ibaqalliwn-Talsnant sites are located in the close west. All new, and Ibaqualliwn-Talsnant fossil traces, are print over the upper levels of marine limestones from Arganane Formation (Domerian). The likewise Domerian lowermost part continental sediments of the superposed Azilal Formation is placed to 5 m above the ichnites. The Utological composition, invertebrate fossil content, and number ofichnological levels is similar in the AssifAit-n-Bou Cuemez-Assif-n-Rbat zone. In spite ofthe Quaternary aluvial sediments cover, the geological properties are enough to define exactly the same stratigraphic position of the sites. The Arganane Formation dissapear to east part and the new sites are the east end ofthe Bou Guemez dinosaur footprints outcrops. It is important show the sauropod supremacy in the Bou Guemez zone

The Iouaridène syncline is a is famous palaeontological site among the ichnologists for several reasons: i) the time of the site is known - the first footprint paper were published in 1937; ii) the discovery of the sauropod trackway Breviparopus, Dutuit and Ouazzou, 1980 used as narrow gauge morphotipe by Farlow (1992); iii) the succesive discussions about the age of the “red beds” where the ichnites were found, and in consequence, the variations in the footprint age attributions; and iv) the discussion about the swimmer dinosaurs in manus-only sauropod trackways. The exploration on the area and the bibliographical review have allowed to verify many of the former suppositions and to adjust the age of the mentioned sauropod trackway, fact of great importancy in the dinosaur palaeoichnology

The Iouaridène syncline is a is famous palaeontological site among the ichnologists for several reasons: i) the time of the site is known - the first footprint paper were published in 1937; ii) the discovery of the sauropod trackway Breviparopus, Dutuit and Ouazzou, 1980 used as narrow gauge morphotipe by Farlow (1992); iii) the succesive discussions about the age of the “red beds” where the ichnites were found, and in consequence, the variations in the footprint age attributions; and iv) the discussion about the swimmer dinosaurs in manus-only sauropod trackways. The exploration on the area and the bibliographical review have allowed to verify many of the former suppositions and to adjust the age of the mentioned sauropod trackway, fact of great importancy in the dinosaur palaeoichnology

The first found tracks of dinosaur in the North of the Goulmima-Tinerhir line (Inner Haut Atlas Central, Morocco) are described. They are located in the Arhbalou Formation and come from isolated and grouped prints mainly of theropod dinosaur. They have importance by the enlargement of the geographical area with dinosaur paleoichnological sites and by they constitute the first dated Carixian dinosaur footprints founds in Morocco.

On the nor th of the Goulmima-Tinerhir line three new dinosaur ichnological sites were discovered. The new ichnites and sites (21 and 3) are added to the 23 footprints and 4 outcrops known until now. All sites are in the Aghbalou Formation (Carixian) being those described in this work, older that previously known. Eubrontes, other not identified theropod and possibly sauropod ichnites are described in the new localities. The litology and the direct and indirect structures accompanying of the footsteps also suppose a contribution for the extension of variation both in the sedimentary materials and morphologic ichnological types in the region

The first found tracks of dinosaur in the North of the Goulmima-Tinerhir line (Inner Haut Atlas Central, Morocco) are described. They are located in the Arhbalou Formation and come from isolated and grouped prints mainly of theropod dinosaur. They have importance by the enlargement of the geographical area with dinosaur paleoichnological sites and by they constitute the first dated Carixian dinosaur footprints founds in Morocco.

On the nor th of the Goulmima-Tinerhir line three new dinosaur ichnological sites were discovered. The new ichnites and sites (21 and 3) are added to the 23 footprints and 4 outcrops known until now. All sites are in the Aghbalou Formation (Carixian) being those described in this work, older that previously known. Eubrontes, other not identified theropod and possibly sauropod ichnites are described in the new localities. The litology and the direct and indirect structures accompanying of the footsteps also suppose a contribution for the extension of variation both in the sedimentary materials and morphologic ichnological types in the region

There are not known too much dinosaur trackways with the metatarsus imprints. The existent dates mention abouth 30 trackways. This is the first time, one or this kind of trackway is described in Morocco. The shape of this footprints are conditioned on the mud behaviour and probably on the trackmaker speed