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Pseudosubulina nouraguensis n. sp. (Fig. 5) TYPE MATERIAL. — Holotype: French Guiana, Réserve naturelle des Nouragues, Nouragues field station (c. 4°05’N, 52°41’W; K10 square), 85 m a.s.l., T. E. J. Ripken & O. Gargominy, 20.XI.1997, leaf litter on lateritic soil in a natural forest gap (INVMAR21054), MNHN 25073. Paratypes: same data as holotype, 2 specimens, MNHN 25074. TYPE LOCALITY. — Nouragues field station (c. 4°05’N, 52°41’W), Réserve naturelle des Nouragues, French Guiana. ETYMOLOGY. — The name of the species relates to its type locality. DIAGNOSIS. — Shell not umbilicate, slender, turrited with straight outline and obtuse apex; consists of up to 10 tightly coiled, slightly shouldered whorls with a deep straight crenulated suture. Protoconch glossy with faint radial and spiral sculpture. First whorl of teleoconch with well-defined radial ribs about same width as intercostal spaces.Surface of later teleoconch whorls with well-defined broad radial ribs much wider than intercostal spaces and with extremely faint, uneven and wavy spiral sculpture. Apertural dentition consists of columellar and basal lamellae and palatal tubercule, repeated half whorl from the apertural set within penultimate whorl. DESCRIPTION (HOLOTYPE) Shell of small size for the genus (height 6.8 mm, major diameter 1.9 mm), not umbilicate, dextral, slender, turrited with straight outline, rather thin, translucent, pale corneous. Whorls 10, slightly shouldered with a deep straight crenulated suture all the way to the aperture. Protoconch 2+ whorls, glossy with faint radial and spiral sculpture. Protoconch/teleoconch transition distinct because of change in macrosculpture. First whorl of teleoconch with well-defined radial ribs about same width as intercostal spaces. Surface of later teleoconch whorls with well-defined broad radial ribs much wider than intercostal spaces and with extremely faint, uneven and wavy spiral sculpture. Growth lines not visible. Aperture elongated, orthocline, with approximately 90° columellar-basal angle, cut by penultimate whorl, with faint whitish parietal callus enhanced on columellar edge. Apertural dentition of deeply sited, strong, short palatal tubercule, perpendicular short basal lamella about same distance from aperture edge as palatal one and large spirally curving downward columellar lamella. Penultimate whorl furnished with exactly the same dentition positioned half whorl from the apertural set. Peristome simple, acute, not reflected. HABITAT. — Primary forest, under leaf litter on lateritic soil. REMARKS With respect to shell morphology, Ischnocion triptyx (Pilsbry,1907) (forming the monotypical genus in the Subulinidae Fischer & Crosse, 1877) described from Colombia,is somewhat similar.However, P.nouraguensis n. sp. is almost definitely a spiraxid and Ischnocion has a parietal lamella that has never been observed in any species of Spiraxidae.Yet,the taxonomic position of P. nouraguensis n. sp. is problematic even within the Spiraxidae. The entire family is in great need of revision since the anatomy is not known for most of the described species (see Baker 1926). What can be observed from the limited material available is that the Recent Spiraxidae are probably remnants of an ancient clade represented today by few well-defined generic level taxa,many of which are monotypic in the current classification. We hypothesise that similarity of the shape of embryonic whorls is much stronger evidence of phylogenetic relationships than the presence or the lack of apertural lamellae in this family. We also suggest that the presence of apertural lamellae in the Spiraxidae is a symplesiomorphic character that has been lost independently in many taxa. All six species of Jamaican Spiraxis Adams, 1850 (including all its subgenera), as well as Venezuelan Spiraxis (Ravenia) blandi (Crosse, 1873) have pointed protoconch and therefore we cannot describe P. nouraguensis n. sp. within this genus despite some similarities with Spiraxis (Euspiraxis) Pfeiffer, 1854 in the apertural armature. Thus, we describe P. nouraguensis n. sp. as Pseudosubulina based on the turrited shape of the shell with an obtuse apex, despite the presence of the apertural lamellae,which have never been recorded in this genus; refraining from the description of another monotypic generic level taxon, pending the revision of the family Spiraxidae. This extends the diagnosis of the genus Pseudosubulina, which now includes species with palatal teeth and columellar lamellae., Published as part of Gargominy, Olivier & Muratov, Igor V., 2012, New taxa of land snails from French Guiana, pp. 783-792 in Zoosystema 34 (4) on pages 789-790, DOI: 10.5252/z2012n4a7, http://zenodo.org/record/5166717, {"references":["BAKER H. B. 1926. - The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela. Part IV. Occasional Papers of the Museum of Zoology University of Michigan 167: 1 - 49, pls 12 - 19."]}

Genus Pseudosubulina Strebel & Pfeffer, 1882 Pseudosubulina Strebel & Pfeffer, 1882: 117. TYPE SPECIES. — Pseudosubulina berendti Pfeiffer, 1862, by original designation. DISTRIBUTION. — Central America and Cuba.Record from Jamaica by Schileyko (2000:840, 841) is based on mistaken classification of Sigmataxis macrospira (Adams, 1851). EMENDATIONS TO THE DIAGNOSIS. — In addition to diagnostic characters indicated by Schileyko (2000: 840, 841), genus now includes species with proximal penial stimulator, as well as with apertural dentition (see descriptions of two new species below). REMARKS Thompson (2010: 197) apparently made a mistake stating the presence of an epiphallus in Pseudosubulina. There is no epiphallus in the genital apparatus of the type species (P. berendti) nor in the new species described below. The vas deferens enters the penis directly through the base of the penial retractor and there are no apparent structural differences between the proximal and distal parts of the penis that could suggest the possibility of a spermatophore creation in the former., Published as part of Gargominy, Olivier & Muratov, Igor V., 2012, New taxa of land snails from French Guiana, pp. 783-792 in Zoosystema 34 (4) on page 786, DOI: 10.5252/z2012n4a7, http://zenodo.org/record/5166717, {"references":["STREBEL H. & PFEFFER G. 1882. - Beitrag zur Kenntnis der Fauna mexikanischer Land- und Susswasser-Conchylien. Theil V. G. J. Herbst, Hamburg: 1 - 144, pls 1 - 19.","SCHILEYKO A. A. 2000. - Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydophoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica supplement 2: 729 - 880.","THOMPSON F. G. 2010. - Four species of land snails from Costa Rica and Panama (Pulmonata: Spiraxidae). Revista de Biologia Tropical 58 (1): 195 - 202."]}

Genus Cyclopedus n. gen. TYPE SPECIES. — Cyclopedus anselini n. sp. by monotypy. ETYMOLOGY.— From the Greek, “cyclo-” for circle, which refers to the perfect spire of the shell and is related to the names of other genera in this super-family, “-pedus” for child, which refers to the small size of the shell. Cyclopède is also a dedication to the French humorist Pierre Desproges. DIAGNOSIS. — Genus characterised by the small shell size and unique teleoconch sculpture of fine impressed dots arranged in spiral lines. Operculum similar to that of Ptychocochlis Simpson, 1895 in having elevated translucent lamellae expanded at the summit and connected to the succeeding whorl, thus forming a completely closed channel in cross section. However, in Ptychocochlis these lamellae form visible steps on the outer surface of the operculum, while in Cyclopedus n. gen. this surface is flat. DESCRIPTION Shell small (height from 3.0to 3.5mm,major diameter from 3.6 to 4.2 mm), umbilicate, dextral, broadly conical with straight outline, elevated, rather thin, light corneous, semi-translucent. Whorls 3.8-4.0, regularly rounded with a deep straight suture all the way to the aperture. Protoconch two whorls, with eight spiral flat ribs.Protoconch/teleoconch transition distinct because of change in microsculpture and discontinuity of growth. Surface of teleoconch with growth lines and occasional varices (discontinuity in growth); sculpture of fine impressed dots arranged in spiral lines irregularly spaced from 0.1 to 0.75 mm and inconspicuous ribs (about 40 on the body whorl) separated by 5-10 growth lines.Aperture circular, not cut by penultimate whorl and clearly delineated but not detached from it in fully grown specimens.Peristome complete,simple,acute,not reflected.Umbilicus open, 5.5 times smaller than major diameter, V-shaped. Operculum almost circular (about 1.8 mm in diameter), corneous, externally concave and internally convex,without any visible steps on both sides,spirally coiled, with elevated translucent lamellae expanded at the summit and connected to the succeeding whorl, thus forming completely closed channel in cross section., Published as part of Gargominy, Olivier & Muratov, Igor V., 2012, New taxa of land snails from French Guiana, pp. 783-792 in Zoosystema 34 (4) on pages 784-785, DOI: 10.5252/z2012n4a7, http://zenodo.org/record/5166717

Bukobia uhehensis Verdcourt, 1965 Fig. 63 Bukobia uhehensis: Verdcourt 1965: 282, fig. 8 (Iringa, Tanzania; length 49 mm, sole width 6 mm). A rare eastern African species, previously known by a single specimen from the type locality in central Tanzania, about 600 km NW of the study area. Found at one station inland. Differs from all other species of the genus Bukobia, as well from two following species of the family, by the presence of a single large soft stimulator without any calcareous elements inside the shortly elongated diverticulum of the atrium., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 282, DOI: 10.5733/afin.051.0203

Tayloria (Colpanostoma) leroyi (Bourguignat, 1889) Fig. 31 Colpanostoma leroyi: Bourguignat 1889: 43, 48, pl. 1, figs 1–3 (Nguru Mountains, eastern Tanzania; 20× 25 mm). A common species found at 12 stations inland, previously known only from eastern Tanzania. The sculpture below the periphery is much less pronounced. Some of the shells are practically smooth basally. The shell illustrated here has a 0.69 h/w ratio. However, some other shells collected in the region are proportionally higher. An unfigured shell from station 17, for example, has similar proportions (17.0× 21.5 mm, h/ w 0.79) to the type (h/ w 0.80). Another two similar Tanzanian species: T. usambarica Craven, 1880 and T. helicoides (Boettger, 1913) do not have a sinuous lip., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 271, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["BOURGUIGNAT, J. R. 1889. Mollusques de l'Afrique equatoriale: de Moguedouchou a Bagamoyo et de Bagamoyo au Tanganika. Paris: Dumoulin et Cie.","CRAVEN, A. E. 1880. On a collection of land and freshwater shells made during a short expedition to the Usambara country in Eastern Africa, with descriptions of seven new species. Proceedings of the Scientific Meetings of the Zoological Society of London 48 (2): 216 - 219.","BOETTGER, C. R. 1913. Description of new species of land shells from Africa. Proceedings of the Malacological Society of London 10: 348 - 354."]}

Afroguppya rumrutiensis (Preston, 1911) Fig. 47 Thapsia rumrutiensis: Preston 1911: 466, pl. 11, fig. 9 (NW of Mount Kenya; 1.25× 2.50 mm). A common East African species with a general distribution from Kenya to the Eastern Cape in South Africa. A few damaged shells were found at one station on Cabo Delgado. Differs from the preceding species by the slightly larger shell with an angulated periphery and an always clearly visible microscopic spiral sculpture., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 278, DOI: 10.5733/afin.051.0203

Achatina immaculata Lamarck, 1823 Figs 19, 59 Achatina immaculata: Lamarck 1822: 128 (height c. 152 mm). A common, widespread eastern African species. Found at five stations inland and at one station on Cabo Delgado. Differs from all other species in the area by its large size (shell up to 200× 120 mm). Fresh shells differ from the similar most common A. fulica Bowdich, 1822 (not found in north-eastern Mozambique so far) by the pink columellar margin. Juvenile shells are even more common and can easily be separated from all other shells in the region by the large smooth protoconch., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 267, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["LAMARCK, J. - B. P. A. DE MONET, LE CHEVALIER DE. 1822. Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une Introduction offrant la determination des caracteres essentiels de l'Animal, sa distinction du vegetal et des autres corps naturels; enfin, l'exposition des principes fondamentaux de la Zoologie. T. 6 (2). Paris: A. Belin."]}

Trochonanina bloyeti Bourguignat, 1889 Fig. 50 Trochonanina bloyeti: Bourguignat 1889: 21 (Near Kondoa, Usambara, north-eastern Tanzania; 7× 15 mm). A rare eastern African species, with a general distribution from northern Tanzania to central Mozambique. A single shell found inland. Quite similar to the preceding species, differing from it by the wider spaced, stronger axial ribs on the upper side of the more depressed, uniformly coloured shell. The shell illustrated here has lighter spots in places of the damaged periostracum., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 280, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["BOURGUIGNAT, J. R. 1889. Mollusques de l'Afrique equatoriale: de Moguedouchou a Bagamoyo et de Bagamoyo au Tanganika. Paris: Dumoulin et Cie."]}

Nesopupa (Insulipupa) corrugata (Preston, 1912) Fig. 13 Jaminia corrugata: Preston 1912: 71, fig. 4 (Victoria Falls; 2.25×1.00 mm). A single, heavily worn shell found on Cabo Delgado.Previously recorded from the area near the type locality. There is also a single shell in the Natal Museum (L6653) collected on 19 February 1963 by A.C. and W.H. van Bruggen in the forest east of Gôndola in Central Mozambique. Characteristic strong corrugation (of the periostracum) can be observed only on fresh shells, and “a small, rather indistinct basal denticle situated well within and rather on the right-hand side of the shell” (Preston 1912: 71) can be missing (Pilsbry & Cooke 1920: 362, pl. 34, fig. 15). The shell from Cabo Delgado has just barely noticeable thickening in place of the lower palatal tooth, with a corresponding slight indentation on the outer surface of the last whorl. N. corrugata has a stronger corrugated periostracum and is proportionally higher than the preceding species., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on pages 264-265, DOI: 10.5733/afin.051.0203

Tropidophora insularis (Pfeiffer, 1852) Fig. 7 Cyclostoma insulare: Pfeiffer 1852: 64 (“ Isle de France ” – error; 13½ × 17 mm). A common species, recorded from seven inland stations, with a general distribution from South Africa to Kenya. “Described from a set in the Cuming collection labelled “ Mauritius,” a locality which Pfeiffer amended to “Natal” in his own working copy of his Monograph” (Connolly 1939: 547). Similar to, and probably a sister species of, T. ligata, with different ecological preferences since they do not co-occur at least in north-eastern Mozambique. Cases of co-occurrence in South Africa should be re-evaluated. Can easily be distinguished from T. ligata by the fairly strong spiral sculpture on the periphery as well as on the rest of the shell, the sculpture being weaker and not as dense and regular as in T. nyasana. It is much larger but otherwise very similar to the following species., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 263, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493

Tropidophora ligata (Müller, 1774) Figs 6, 56 Nerita ligata: Müller 1774: 181 (12.7–23.3× 12.7–21.2 mm). A common species, recorded from seven stations on Cabo Delgado and inland, with a general distribution from South Africa to Tanzania as well as on Madagascar and Mauritius. Differs from the other three species of Tropidophora in the region by the very weak spiral sculpture on the periphery of the last whorl, with most of the shells having noticeable spiral sculpture only in the umbilical area., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 263, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["MULLER, O. F. 1774. Vermivm terrestrium et fluviatilium, seu animalium infusoriorum, helminthicorum et testaceorum, non marinorum, succincta historia. 2. Havniae [Copenhagen] & Lipsiae [Leipzig]: Heineck et Faber."]}

Edentulina affinis Boettger, 1913 Fig. 30 Edentulina affinis: Boettger 1913: 349, pl. 15, fig. 8 (Kipatimu, eastern Tanzania; 31.5×15.0 mm). A common East African species, with a general distribution from Kenya to Mozambique. Found at ten stations inland and two stations on Cabo Delgado. The convexity of the whorls significantly varies amongst the shells from Mozambique and most of the shells do not have whorls “flatter” than in Comorian E. ovoidea (Bruguière, 1789), as stated in the original description of affinis., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 271, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["BOETTGER, C. R. 1913. Description of new species of land shells from Africa. Proceedings of the Malacological Society of London 10: 348 - 354.","BRUGUIERE, J. G. 1789 - 1792. L'Encyclopedie methodique ou par ordre de matieres par une societe de gens de lettres, de savants et d'artistes; precedee d'un Vocabulaire universel, servant de Table pour tout l'Ouvrage, ornee des Portraits de MM. Diderot et d'Alembert, premiers Editeurs de l'Encyclopedie. Histoire naturelle des vers. Tome Premier. Paris: Panckoucke."]}

Gittenedouardia metuloides (Smith, 1899) Fig. 15 Buliminus (Conulinus) metuloides: Smith 1899: 587, pl. 33, fig. 43 (Zomba Plateau; 10× 6 mm). A widespread but not very common species with a general distribution in Zambia, southern Malawi (type locality), Mozambique, Zimbabwe and the northern part of South Africa. A single worn shell (juvenile), found about 30 km inland, is noticeably wider than Bulimus badiolus Morelet, 1881 described from the Comoros and just slightly more slender than four specimens of G. metuloides in the Natal Museum (No. 2786, ex. coll. Connolly 1920) from the Victoria Falls. Differs from the three following species of Cerastidae by the slender shell with a deep suture., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 265, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493

Achatina randabeli Bourguignat, 1889 Fig. 20 Achatina randabeli: Bourguignat 1889: 84, pl. 5, fig. 6 (Tabora, Ounyanyembé; 55× 28 mm). A rather uncommon species, previously known only from the type locality in central Tanzania. Found at five stations inland and two station on Cabo Delgado. One of the smaller species of Achatina. The largest shell from north-eastern Mozambique is 51× 26 mm. Achatina arctespirata (57× 32 mm) described by Bourguignat (1889: 83, pl. 5, fig. 7) from the southern region of Tanganyika is probably a synonym based on a freak specimen. Both species were considered subspecies of the Tanzanian A. craveni Smith, 1881 (type: 81× 37 mm) by Bequaert (1950: 50). Differs from all other species of the genus by the narrow-conical apex with straight outlines. Can easily be separated from similar sized species of Limicolaria by the strongly truncated columella., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 267, DOI: 10.5733/afin.051.0203, {"references":["BOURGUIGNAT, J. R. 1889. Mollusques de l'Afrique equatoriale: de Moguedouchou a Bagamoyo et de Bagamoyo au Tanganika. Paris: Dumoulin et Cie.","SMITH, E. A. 1881. On a collection of shells from lakes Tanganyika and Nyassa and other localities in East Africa. Proceedings of the scientific meetings of the Zoological Society of London 49 (2): 276 - 300.","BEQUAERT, J. 1950. Studies in the Achatinidae, a group of African land snails. Bulletin of the Museum of Comparative Zoology at Harvard College, in Cambridge 105 (1): 3 - 216."]}

Kempioconcha kirki (Dohrn, 1865) Fig. 22 Buliminus kirki: Dohrn 1865: 232 (Cabaçeira, Mozambique; 23.8× 11.1 mm). A common species, with a general distribution in Mozambique and Zimbabwe. Found at three stations inland and three on Cabo Delgado. The type (7 whorls) is slightly smaller than most of the shells of this species in the study area. However, the h/w ratio is almost exactly the same in the type (2.14) and in the shell illustrated here (2.17). It is smaller than the preceding species and much larger than any of the following species of the family. The type locality of Buliminus liederi Martens, 1895 in south-eastern Tanzania (Kitohaui on the plateau between Ukuledi and Umbekuru, District Mgao) is only about 150 km northwest of the study area. It has a more elongated shell (h/ w 2.55): 28× 11 mm with 7½ whorls in the original description (von Martens 1895: 180). Van Bruggen (2008: 360–368, fig. 10) reported Pseudoglessula (Kempioconcha) liederi (8 whorls subadult: 27.6× 12.1 mm) from Malawi. However, he followed the Verdcourt (1967) key where liederi is separated from the similarly sized K. kirki (Dohrn, 1865) only by “apical whorls with costae only slightly more widely spaced than on the lower whorls”. This character was mentioned by von Martens in comparison with “ Limicolaria borellii ” (=? Glandina boivini Morelet, 1860, see Connolly 1925: 171) and Pseudoglessula conradi in his second description of liederi (von Martens 1897: 61, pl. 3, fig. 32) and he separated Buliminus from Pseudoglessula based on this character, in addition to the truncated columella in the latter (von Martens 1897: 116). However, in the same work von Martens (1897: 62) classified kirki Dohrn, 1865 as Buliminus, thus acknowledging that the axial ribs are not stronger and wider spaced on the apical whorls in both liederi and kirki. Unfortunately, he did not compare these two species, concentrating his attention on the penultimate whorl sculpture of his apparently subadult specimen of kirki. I prefer to use the earlier name here, since variability of the shells from station 6 allows the application of both names, pending a proper revision of the genus., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on pages 268-269, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["DOHRN, H. 1865. List of the land and freshwater shells of the Zambesi and Lake Nyassa, Eastern Tropical Africa, collected by John Kirk, M. D., F. L. S., & c. Proceedings of the Scientific Meetings of the Zoological Society of London 33 (1): 231 - 234."]}

Tropidophora zanguebarica (Petit, 1850) Figs 8, 57 Cyclostoma zanguebarica: Petit 1850: 53, pl. 3, fig. 5 (Ile de Zanzibar; 10–12 × 10–12 mm). Recorded from all three stations on Cabo Delgado, with a general coastal distribution from Kenya to Mozambique. Its relations with T. letourneuxi (Ancey in Bourguignat, 1887) should be re-evaluated (see remarks in Rowson 2007: 432). I prefer to use the earlier name here since variability of the shells from Cabo Delgado allows the application of both names. Resembles a small T. insularis but, unlike that species, co-occurs with T. ligata. Possibly just a coastal variation of T. insularis with size variability from 10.5×10.5 to 13.0×12.0 mm at a single locality., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 263, DOI: 10.5733/afin.051.0203, {"references":["PETIT, M. 1850. Notice sur le genre Cyclostoma, et catalogue des especes appartenant a ce genre. Journal de conchyliologie, comprenant l'Etude des Animaux, des Coquilles vivantes et des Coquilles fossiles 1: 36 - 55.","ROWSON, B. 2007. Land molluscs of Zanzibar island (Unguja), Tanzania with the description of a new species of Gulella (Pulmonata: Streptaxidae). Journal of Conchology 39 (4): 425 - 466."]}

Gulella perissodonta (Sturany, 1898) Fig. 36 Ennea perissodonta: Sturany 1898: 1898: 26, pl. 1, fig. 18 (Delagoa Bay, Mozambique; 4.2× 2.1 mm). Found at one station on Cabo Delgado. Two shells (2.7× 1.4 mm) were collected by van Hoepen (van Bruggen 2006: 124) on Ilha Vumba only 49 km south of station 7 on Cabo Delgado. Prior to that, perissodonta was known only from south-eastern Mozambique, Swaziland and north-eastern KwaZulu-Natal where it is quite common. The type, described from the area near Maputo, is larger than illustrated here, but the shells from northern KwaZulu-Natal vary from 3.2×1.7 to 6.2× 2.5 mm (van Bruggen 1969: 47)., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 274, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["STURANY, R. 1898. Catalog der bisher bekannt gewordenen sudafrikanischen Land- und Susswasser-Mollusken mit besonderer Berucksichtigung des von Dr. Penther gesammelten Materiales. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe 67: 537 - 642.","BRUGGEN, A. C. VAN. 1969. Studies on the land molluscs of Zululand with notes on the distribution of land molluscs in southern Africa. Zoologische verhandelingen uitgegeven door het Rijksmuseum van Natuurlijke Historie te Leiden (Ministerie van Cultuur, Recreatie en Maatschappelijk Werk) 103: 1 - 116, 4 pls."]}

Cyathopoma diegoense Fischer-Piette, Blanc & Vukadinovic, 1974 Fig. 3 Cyathopoma diegoense: Fischer-Piette, Blanc & Vukadinovic 1974: 467, figs 2–4 (Cap Diego, northern Madagascar; c. 2× 3 mm). A single shell without an operculum, collected on Cabo Delgado, is very similar to the voucher specimen described and illustrated by Emberton (2003: 30–31, fig. 40) from the type locality. The photograph of the shell from Cabo Delgado matches the photograph of the shell from Cap Diego almost exactly when superimposed at 113% magnification. The characteristic sculpture of the spiral cords with the gap on the whorl shoulder (“missing” cord below the suture) and the minute dense axial striation between the spiral cords also can be observed on both shells. Cyathopoma azaniense Verdcourt, 1978 from Kenya is smaller (1.4×2.0 mm) and does not have the characteristic gap between the suture and the first spiral cord on the following whorl. The original description of C. diegoense was based on a single adult specimen “environ” (circa) 2 mm high and 3 mm wide. The same size was repeated in comparison with C. artatum Sykes, 1897 on the same page just below the description. However, the figure of the holotype on the same page has a 0.88 h/w ratio, which clearly does not match the proportions stated in the description. Emberton (2003) did not give any explanation for his decision to include the 2.1 mm high and 2.0 mm wide C. duboisi Fischer-Piette, Blanc, Blanc & Salvat, 1993, as well as his 1.9 mm high and 2.9 wide “voucher specimen” in the range of variation of C. diegoense. Thus, the specimen from Cabo Delgado is very likely C. diegoense sensu Emberton, 2003 but the extent of the name diegoense is doubtful., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 261, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["FISCHER-PIETTE, E., BLANC, F. & VUKADINOVIC, D. 1974. Additions aux Mollusques terrestres de Madagascar. Bulletin du Museum National d'Histoire Naturelle, Paris, 3 e serie, N ° 218, mai - juin 1974, Zoologie 146: 465 - 526.","EMBERTON, K. 2003. Madagascan Cyathopoma sensu lato (Gastropoda: Cyclophoridae). Archiv fur Molluskenkunde 132: 9 - 91.","FISCHER-PIETTE, E., BLANC, Ch. P., BLANC, F. & SALVAT, F. 1993. Gasteropodes terrestres prosobranches. Faune de Madagascar 80: 1 - 281."]}

Streptostele (Raffraya) herma Connolly, 1912 Fig. 38 Streptostele (Raffraya) herma: Connolly 1912: 89, pl. 2, fig. 3 (Victoria Falls; 4.5× 1.3 mm). A rather uncommon East African species, with a general distribution from south-eastern Kenya to north-eastern parts of South Africa. All shells found at all three stations on Cabo Delgado are slightly smaller than the type of this species. Differs from all other snails in the area by the small, elongated shell with a weak parietal tooth and slight palatal thickening., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 277, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["CONNOLLY, M. 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11 (3): 59 - 306."]}

Pupoides coenopictus (Hutton, 1834) Fig. 10 Pupa coenopicta: Hutton 1834: 85, 93 (Beana, India; height c. 5.29 mm). A common, widespread species recorded in large numbers from all three stations on Cabo Delgado, with a general distribution in tropical Asia and Africa., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 264, DOI: 10.5733/afin.051.0203, {"references":["HUTTON, T. 1834. On the land shells of India. The Journal of the Asiatic Society of Bengal 3: 81 - 93."]}

Euonyma tugelensis (Melvill & Ponsonby, 1897) Fig. 27 Subulina tugelensis: Melvill & Ponsonby 1897: 637, pl. 17, fig. 9 (Lower Tugela River, Natal; 14× 4 mm). Five damaged shells found at two stations inland, although similar in size to Opeas cressyi Connolly, 1922 (type: 10.8× 2.8 mm) from central Mozambique, are not as elongated as that species. The largest (damaged) shell from the study area measures 8.2× 3.1 mm and is similar in size and proportions to Opeas durbanense (8.7× 3.3 mm) described by Sturany (1898: 61, pl. 2, figs 42–44) from the coast of KwaZulu-Natal. Herbert and Kilburn (2004: 133) treated durbanense as a synonym of tugelensis. Differs from the following species, as well as from common and similarly sized Allopeas clavulinum (Potiez & Michaud, 1838), by the extremely narrow umbilical slit., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 270, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["MELVILL, J. C. & PONSONBY, J. H. 1897. Descriptions of eleven new species of land and freshwater Mollusca from South Africa. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, 6 th series 19 (114): 633 - 639.","STURANY, R. 1898. Catalog der bisher bekannt gewordenen sudafrikanischen Land- und Susswasser-Mollusken mit besonderer Berucksichtigung des von Dr. Penther gesammelten Materiales. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe 67: 537 - 642.","HERBERT, D. & KILBURN, R. 2004. Field guide to the land snails and slugs of eastern South Africa. Pietermaritzburg: Natal Museum.","POTIEZ, V. L. V. & MICHAUD, A. L. G. 1838. Galerie des Mollusques, ou Catalogue methodique, descriptif et raisonne, des Mollusques et Coquilles du Museum de Douai. 1. Paris: J. B. Bailliere."]}

Elisolimax flavescens (Keferstein, 1866) Fig. 64 Parmarion flavescens: Keferstein 1866: 70, pl. 2, figs 2–8 (Inhambane, Mozambique; length 35 mm – contracted). A common eastern African species, with a general distribution from southern Tanzania to the Eastern Cape in South Africa. Found at two stations on Cabo Delgado. Differs from other continental species of the genus Elisolimax, as well as from the preceding and the following species of the family, by the long, narrow diverticulum of the atrium with two large longitudinal folds inside., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 283, DOI: 10.5733/afin.051.0203, {"references":["KEFERSTEIN, W. 1866. Ueber Parmarion flavescens sp. n. aus Mossambique. Malakozoologische Blatter 13: 70 - 76."]}

Gulella (Molarella) subhyalina (Smith, 1890) Fig. 35 Ennea subhyalina: Smith 1890: 165, pl. 6, fig. 13 (Mamboya, north-eastern Tanzania; 6.00× 2.25 mm). A not very common eastern African species, with a general distribution from northern Tanzania to northern Mozambique. Found at one station inland. Differs from other Streptaxidae in the area by the smooth cylindrical shell with a single large parietal lamella, two upper palatal, one deeply positioned lower palatal, one basal lamella and two columellar lamellae. The type of Ennea subhyalina does not have the lower palatal tooth. There are several other species and forms with a smooth cylindrical shell and similar dentition. Ennea ingens described by Sturany (1898: 23, pl. 1, fig. 9) from the coast of KwaZulu-Natal (Durban) is much larger (9.0× 3.2 mm) and also has only two palatal teeth. Ennea gwendolineae described by Preston (1910: 527, fig. 3) from Shimba Hills south-west of Mombasa in south-eastern Kenya, also has only two palatal teeth and is smaller (5.0× 1.5 mm) but otherwise very similar to subhyalina. Surprisingly, Preston did not mention subhyalina in the original description of gwendolineae, instead comparing his new species with the much more different E. subflavescens, which was described by Smith (1890: 165, pl. 6, fig. 14) on the same page with subhyalina and illustrated next to it. Connolly (1922 b: 501, pl. 14, fig. 27) described G. (M.) gwendolineae scissidens from Tanzania (Dar es Salaam) with an additional “more deep-set” lower palatal tooth. With the exception of its much smaller size (4.1× 1.6 mm), scissidens is otherwise almost identical in shell shape, dentition and lack of sculpture to subhyalina from north-eastern Mozambique. Considering the very close similarities between gwendolineae + scissidens, and subhyalina + shells with an additional lower palatal tooth from north-eastern Mozambique, it is quite possible that all of them may well be just variations of one species with a distribution from south-eastern Kenya to north-eastern Mozambique. Therefore, it seems unnecessary to give a separate name to the seventoothed variety from north-eastern Mozambique., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 273, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["STURANY, R. 1898. Catalog der bisher bekannt gewordenen sudafrikanischen Land- und Susswasser-Mollusken mit besonderer Berucksichtigung des von Dr. Penther gesammelten Materiales. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftlichen Classe 67: 537 - 642.","PRESTON, H. B. 1910. Additions to the non-marine molluscan fauna of British and German East Africa and Lake Albert Edward. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, 8 th series 6 (35): 526 - 536."]}

Trochonanina albopicta (von Martens, 1869) Fig. 49 Nanina mossambicensis var. albopicta: von Martens 1869 a: 56, pl. 1, fig. 2 (East Africa; 11× 18 mm). Ledoulxia albopicta (von Martens, 1869): Connolly 1925: 135. A common East African species, with a general distribution from southern Kenya to central Mozambique. Found at four stations inland. Possibly just a colour variety of Trochonanina mossambicensis (Pfeiffer, 1855), as it was originally described by von Martens. Can easily be recognized by the characteristic pattern of lighter, nontranslucent specks oriented in spiral lines on a darker, semi-translucent, corneous background, as well as alternating zones of lighter and darker close-set axial riblets on the upper side of the shell with only microscopic spiral striation on the base. The shell illustrated here is much faded., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 280, DOI: 10.5733/afin.051.0203

Gonaxis denticulatus (Dohrn, 1878) Fig. 33 Streptaxis denticulatus: Dohrn 1878: 152 (Mombasa, south-eastern Kenya; c. 7.0× 4.5 mm). A widespread, common East African species with a general distribution from Kenya to Mozambique. Found at eight stations inland. This species is smaller and has a less distorted axis than the preceding species. It differs from other similarly sized Gonaxis by the smooth shell with a strongly serrated suture. Dohrn had a quite unusual way of measuring distorted shells. He gave “Diam. maj. 7, min. 4½, alt. 3½ mill.” in the original description of denticulatus. Apparently, he let the shell lie down freely on the flat surface and measured the height of the shell above the surface, calling that measurement “alt.”, which corresponds to the minor diameter in modern terminology. Then his “diam. maj.” roughly corresponds to the height (being slightly larger) and “diam. min.” roughly corresponds to the major diameter (being slightly smaller). Thus, the size from the original description in modern terminology would be: height ~ 7 mm, diameter major ~ 4.5 mm and diameter minor – 3.5 mm. (This became apparent in the original description of the similarly sized but much stronger sculptured Streptaxis kirkii Dohrn, 1865, where he added the diameter of the aperture: “Diam. maj. 5/16, min. 3/16, alt. 3/16, apert. diam. 1/8 poll.” (Dohrn 1865: 232), which corresponds to: height – 7.94 mm, diameter major and minor – 4.76 mm, aperture diameter – 3.18 mm in modern terminology.) Surprisingly, von Martens (1897: 30, 32) copied the dimensions given by Dohrn into his comparison table, which is probably the reason why he listed separately the very similar Streptaxis ordinarius Smith, 1890. The latter species measures 6.5×4.0 mm in the original description (Smith 1890: 160, pl. 6, figs 2 and 2a) and was synonymised with Streptaxis denticulatus by Thiele (1911: 186), who apparently compared the types of both species. The similarly sized Gonaxis gibbonsi Taylor, 1877 (7.0× 4.4 mm) has a proportionally more elongated and slightly more distorted shell. G. cressyi Connolly, 1922 is smaller (6.2× 3.8 mm) and has slightly less distorted shell. G. mozambicensis (Smith, 1881) is slightly larger: 7.5× 4.5–4.8 mm (Smith 1881: 280, pl. 32, figs 6, 6a) but may prove to be a synonym of denticulatus., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 272, DOI: 10.5733/afin.051.0203, {"references":["DOHRN, H. 1865. List of the land and freshwater shells of the Zambesi and Lake Nyassa, Eastern Tropical Africa, collected by John Kirk, M. D., F. L. S., & c. Proceedings of the Scientific Meetings of the Zoological Society of London 33 (1): 231 - 234.","THIELE, J. 1911 [1912]. Mollusken der Deutschen Zentralafrika-Expedition. In: Schubotz, H., ed., Deutsche Zentral-Afrika-Expedition, 1907 - 1908. 3, Zoologie I (6) Mollusca. Leipzig: Klinkhardt & Bergmann, pp. 175 - 214, 6 pls.","TAYLOR, J. W. 1877. Description of new species of land shells from the East coast of Africa. The Quarterly Journal of Conchology 1 (12): 251 - 255.","SMITH, E. A. 1881. On a collection of shells from lakes Tanganyika and Nyassa and other localities in East Africa. Proceedings of the scientific meetings of the Zoological Society of London 49 (2): 276 - 300."]}

Nesopupa (Afripupa) bisulcata (Jickeli, 1873) Fig. 14 Pupa bisulcata: Jickeli 1873: 107 (Abyssinia; 1.9×1.0 mm). A widespread but not very common species with a general distribution from Eritrea (type localities) to Mozambique. A single shell found on Cabo Delgado. Differs from both preceding species of Insulipupa by its smaller size and stronger angular tooth, which is almost as large as the parietal lamella., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 265, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["JICKELI, C. F. 1873 [1872]. Diagnosen neuer Mollusken meiner Reiseausbeute. Malakozoologische Blatter 20: 99 - 108."]}

Tropidophora nyasana (Smith, 1899) Fig. 5 Pomatias nyasanus: Smith 1899: 591, pl. 35, fig. 5 (Mount Chiradzulu, Nyika Range, Zomba Plateau; 20× 21 mm). Recorded from two stations on Cabo Delgado; has a general distribution in Malawi (type localities), Mozambique and Zimbabwe. Differs from the other three species of Tropidophora in the region by the presence of numerous regularly spaced spiral cords over the entire shell surface., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 263, DOI: 10.5733/afin.051.0203

Gastrocopta klunzingeri (Jickeli, 1873) Fig. 11 Pupa klunzingeri: Jickeli 1873: 106 (Abyssinia; 2.25× 1.25 mm). Recorded from two stations on Cabo Delgado, with a general distribution from Eritrea (type localities) to Mozambique. Differs from similar sized species of Vertiginidae by the fused angular and parietal lamellae, the narrow apex and the strongly convex whorls separated by a deep suture., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 264, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["JICKELI, C. F. 1873 [1872]. Diagnosen neuer Mollusken meiner Reiseausbeute. Malakozoologische Blatter 20: 99 - 108."]}

Microcystina minima (Adams, 1867) Fig. 46 Macrochlamys minima: Adams 1867: 303, pl. 19, fig. 2 (near Port Louis, Mauritius; c. 1.3× 0.5 mm). A single shell was found on Cabo Delgado.Previously known only from the Mascarene Islands where it is quite common and recently recorded from Zanzibar (Rowson et al. in press). The smallest species in the study area. Can be easily distinguished from the following species by the lack of spiral sculpture. The shell was originally described “... depressa, discoidea... Diam. 1⅓, alt. ½ mill.” and the apex is not visible on the original illustration. The measurements in the original description were clearly approximate. Griffiths and Florens (2006: 130, pl. 27, fig. E) illustrated M. minima with a protruding apex and gave “Diameter: 1.3 mm; height 0.5 mm ” for the size of the shell. However, if calculated from the proportions of that illustration, assuming that diameter is 1.3 mm, the height of the shell would be 0.82 mm. Thus, in view of the fact that there are no other shells with a narrow umbilicus, without any spiral sculpture and 1.3 mm in diameter found so far on Mauritius, it would be safe to assume that M. minima has variably protruding apical whorls., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on pages 277-278, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["ADAMS, H. 1867. Descriptions of new species of shells collected by Geoffrey Nevill, Esq.; at Mauritius. Proceedings of the Scientific Meetings of the Zoological Society of London 35 (2): 303 - 307.","GRIFFITHS, O. L. & FLORENS, V. F. B. 2006. A field guide to the non-marine molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion and the Northern Dependencies of Mauritius). Mauritius: Bioculture Press."]}

Kempioconcha boivini (Morelet, 1860) Figs 23, 60 Glandina boivini: Morelet 1860: 72, pl. 5, fig. 5 (mainland Kenya coast near Mombasa; 18× 8 mm). A common East and Central African species. Found at ten stations inland. Differs from the following two species of the genus by the proportionally wider shell., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 269, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493

Quickia concisa (Morelet, 1848) Fig. 52 Succinia concisa: Morelet 1848: 351 (Gabon; 7× 4 mm). A common African species, with a general distribution in tropical areas from Sierra Leone to Kenya and Tanzania, as well as on the Comoros, Seychelles and Mascarene Islands. Found at all three stations on Cabo Delgado., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 283, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["MORELET, A. 1848. Testacea quaedam Africae occidentalis terrestria et fluviatilia. Revue Zoologique par La Societe Cuvierienne; Association universelle pour l'Avancement de la Zoologie, de l'Anatomie comparee et de la Palaeontologie 11: 351 - 355."]}

Rachis cunctatoris Van Bruggen, 1975 Fig. 17 Rachis cunctatoris: Bruggen 1975: 217, figs 7–9 (Lower Shire Valley, southern Malawi; 18.7× 8.1 mm). A rather uncommon species previously known only from the dry lowlands of Malawi. Found at four stations inland. Differs from the previous species by the more slender shell, with not more than two dark bands (on the periphery and just above the suture) and by the complete absence of spotted coloration., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 267, DOI: 10.5733/afin.051.0203

Nesopupa (Insulipupa) peilei Madge, 1938 Fig.12 Nesopupa peilei: Madge 1938: 16, pl. 3, fig. 2 (Flic-en-Flac, Mauritius; 2.1× 1.1 mm). A common species on the Mascarene Islands, where it appears to be restricted to the coastal and drier areas inland (Griffiths & Florens 2006: 85). A single shell found on Cabo Delgado. Can be separated from the following species by the less elongated shell., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 264, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["MADGE, E. H. 1938. Notes on some Mollusca of Mauritius, with descriptions of four new species. Mauritius Institute Bulletin 1 (3): 15 - 29.","GRIFFITHS, O. L. & FLORENS, V. F. B. 2006. A field guide to the non-marine molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion and the Northern Dependencies of Mauritius). Mauritius: Bioculture Press."]}

Kempioconcha cf. obtusa auct. non (Boettger, 1913) Fig. 21 Pseudoglessula cf. obtusa: Verdcourt 1967: 50–52, fig. 23. A rare and possibly not previously recognised species. Only a few damaged shells were found at four stations in a 6× 20 km area west of Palma. The size of the shell varies significantly, the largest (8 whorls; subadult) being 40× 20 mm. The same-sized Krapfiella magnifica Preston, 1913 a has a spirally sculptured protoconch. Differs from all other large species of Kempioconcha by the narrowly conical apex. Apparently the largest species of the genus. Verdcourt (1967: 50–52, fig. 23) most probably illustrated the same species (7 whorls subadult) from the Rondo Plateau in south-western Tanzania and gave the following shell dimensions (26.0–29.5×12.0–15.0 mm) for this species, which he called “ cf. obtusa Boettger ” and suspected that it “will prove to be conspecific” with true obtusa Boettger, 1913 and obesa Germain, 1916. However, Kempioconcha obtusa (Boettger, 1913), the largest known species of the genus (32.0× 15.5 mm, with obesa apparently being a synonym), has a noticeably wider, obtuse apex, which is the defining character of that species according to Boettger (1913: 352, pl. 17, fig. 2)., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 268, DOI: 10.5733/afin.051.0203, {"references":["BOETTGER, C. R. 1913. Description of new species of land shells from Africa. Proceedings of the Malacological Society of London 10: 348 - 354."]}

Maizania wahlbergi (Benson, 1852) Fig. 4 Cyclostoma wahlbergi: Benson 1852: 271 (Natal; 7× 14 mm). A common species, recorded from nine stations on Cabo Delgado and inland, with a general coastal distribution from Kenya to the Eastern Cape, South Africa., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 262, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["BENSON, W. H. 1852. Notes on the genus Cyclostoma; and characters of some new species from India, Borneo, and Natal. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology. 2 nd ser. 10: 268 - 272."]}

Opeas hannense (Rang, 1831) Fig. 28 Helix (Cochlicelle) hannensis: Rang 1831:41, pl.3, fig.8 (Cape Verde peninsula,WestAfrica;4–5 × 1.5–2 mm). One of the most common and widespread (mostly through introduction) tropical species; found at one station inland. This is apparently the first record of this species from eastern Africa. The illustrated shell is almost the same in size and proportions as Opeas araeum Connolly, 1923: 652, pl. 19, fig. 35 (type: 5.3× 1.8 mm from Kenya), which is possibly a synonym of hannense. Can be easily separated from the two other small subulinids in the area by the narrow but decisively open umbilicus., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 270, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["RANG, S. 1831. Annales des sciences naturelles comprenant la physiologie animale et vegetale, l'anatomie comparee des deux regnes, la zoologie, la botanique, la mineralogie et la geologie. Paris: Crochard."]}

Limicena nyasana (Smith, 1899) Fig. 18 Buliminus (Conulinus) nyasanus: Smith 1899: 586, pl. 33, figs 41, 42 (Nyika Plateau, Mount Chiradzulu and Zomba Plateau; 21× 15 mm). A widespread but not very common species, with a general distribution from southern Tanzania to southern Zambia, Malawi (type localities) and Mozambique. Found at one station on Cabo Delgado. Differs from the three preceding species of Cerastidae by the wider bulimoid shell with a cylindrical umbilicus, and by the characteristic protoconch sculpture of seven strong spiral ridges well illustrated by Mordan (1998: 62, fig. 3)., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 267, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["MORDAN, P. 1998. The anatomy and phylogeny of the African land snail Limicena Connolly, 1925 (Pulmonata: Cerastidae). Zoologische Mededelingen 72: 59 - 72."]}

Urocyclus kirki Gray, 1864 Fig. 65 Urocyclus kirki: Gray 1864: 251 (Near the mouth of the River Zambezi, Mozambique). A common eastern African species, with a general distribution from southern Tanzania to Zululand in South Africa. Found at a single station inland. Differs from all other species of the family by the large, muscular diverticulum of the atrium with a single large calcareous dart inside., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 283, DOI: 10.5733/afin.051.0203, http://zenodo.org/record/7913493, {"references":["GRAY, J. 1864. On Urocyclus, a new genus of terrestrial gasteropodous Mollusca from Africa. Proceedings of the Scientific Meetings of the Zoological Society of London 32 (2): 250 - 251."]}

Kempioconcha cf. subolivacea (Smith, 1890) Fig. 24 A single shell found inland is about the same size but not as widely conical with straight outlines as in the original description of Buliminus olivaceus J.S. Gibbons in Taylor 1877 preoccupied: 253, pl. 2, fig. 5; (19.35× 7.92 mm) from “Bawri Island, Zanzibar, Channel”. Differs from the following species by the proportionally higher shell with much smaller apical whorls., Published as part of Muratov, Igor V., 2010, Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique, pp. 255 in African Invertebrates 51 (2) on page 269, DOI: 10.5733/afin.051.0203, {"references":["TAYLOR, J. W. 1877. Description of new species of land shells from the East coast of Africa. The Quarterly Journal of Conchology 1 (12): 251 - 255."]}