Search

Bryophila schwingenschussi (Boursin, 1954) (Figs 3b and 7b) Material examined: Holotype: 1 male, “ Bône (Algerien), Juni 1931 (J. Staettermayer leg.) (Coll. Schwingenschuss)“ (Figs 3b and 7b). In NHMW. Taxonomic notes: It was described on a single male specimen, being compared with B. barbaria and B. blepharista, in a text where B. muralis and B. domestica were also mentioned as similar taxa. Boursin (1954) stated that this species and B. blepharista are very similar externally but B. schwingenschussi has shorter pectination of the male antennae, slenderer uncus, somewhat longer valvae and the vesica is armed with a strong cornutus which is missing from both B. blepharista and B. barbaria. As far as is known, it is restricted to Morocco. External and genital diagnosis: see under taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila barbaria Schawerda, 1934 (Figs 2a to 2d, 6c and 8c) Material examined: A large number of specimens of both sexes from the high altitude parts of the High Atlas (mainly from Oukaimeden). Apart from the lectotype and paralectotype (Figs 2a and 2b), the data of the dissected specimens are as follows: RL12467m—Morocco, Haut Atlas Mts, Oukaimeden, 2700 m, 10.VII.1986, leg. F. Fernandez-Rubio (coll. GR) and RL 12477m and RL12479f—Morocco, High Atlas Mts, Oukaimeden, 2400-2700 m, 5-21.VII.1971, leg. G. Friedel (coll. HNHM). Lectotype designation: Male, “Mauretania central, Atlas major, Sidi Chamarouche, Reraia-Tal, 2300 m, 27.6.1932, H. Dürck leg.”, “collectio Schawerda”, “ Bryophila muralis var barbaria Schaw., Type. ♂. Abgebildet ” (pink label), “PREPARATION | W.M. 409 | CH. BOURSIN” (coll. NHMW) (illustrated in Fig 2a). Taxonomic notes: The date of description is cited in most references as 1935, the date printed onto the volume is 1st December, 1934. It is a well-known species of the Atlas Mts in the North African Mediterranean. Schawerda (1934) wrote: „Im Grossen Atlas (Mauretanien = Barbaria) von Dr. Dürck (München) erbeutet. Sidi Chamarouche, Reraia-Tal, 2300 m, 27.6.1932. Types in my collection“. Despite the statement of Schawerda, arguing that one was male and the other female, the syntypes are both males. External diagnosis: This species has longer and broader wings than B. perloides and lighter forewing colouration, and the dark costal markings are always clearly visible (Figs 2a to 2d); the two species are well-separable by their external features. Genital diagnosis: In the male genitalia (Fig 6c), this species has, within the species complex, the longest uncus and strongest clasper, this longest too and with sharp apex. Also the valva is slenderest at the middle, with broadly rounded cucullus, compared to B. perloides. B. barbaria shows no cornuti in the vesica. In the female genitalia (Fig 8c), the ductus bursae is dorsally swollen at his posterior half (Fig 5c)., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on pages 60-61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Schawerda, K. (1934) Neue Heteroceren aus Algerien. Internationale Entomologische Zeitschrift, 28, 285 - 287."]}

Bryophila blepharista (Boursin, 1954) (Figs 3c and 7c) Material examined: Holotype (Figs 3c and 7c): 1 male, Taanzoult [Tazoult], Mittel-Atlas, French Marokko, 16. August 1950 (Ch. Rungs leg.). Paratypes: 1 male, from the same locality and date (Ch. Rungs leg.), 4 males, Aguelmane de Sidi Ali, Mittel-Atlas 2080 m, 5/ 25 July 1943 (Ch. Rungs leg.) (Coll. Rungs, Boursin and Schwingenschuss), 1 male, Bahlil, French Marokko, 20. August 1952 (Abdallah leg.) (Coll. Ch. Rungs). In MNHN. Taxonomic notes: The species was described from Morocco based on a few male specimens, showing differences in their antennae and certain features of the male genitalia (stout uncus and valval apical section, vesica without cornuti) compared with B. schwingenschussi and B. barbaria (Boursin 1954). External diagnosis: B. blepharista differs from B. perloides externally in the shape of the omega-shaped design, in which the inner marginal line is usually at a higher position in B. blepharista but close to the hind margin in B. perloides. Additionally, B. blepharista shows more or less pronounced antemarginal line, absent or almost absent in B. perloides. Genital diagnosis: The male genitalia of B. blepharista (Fig. 7c) are seemingly stouter than those of B. perloides (Figs 6a and 6b), according to the short series of examined specimens; the central clasper is comparatively shorter and gracile in B. perloides, although in both species shows spatulate apex. On the other hand, the uncus of B. blepharista has a narrow stem and incrassate subapical part, exactly like in B. perloides. The sacculus is short and flimsy, like in B. perloides too. Examination of larger series may clarify if these differences are actually interspecific or rather intraspecific. Epiphallus is cylindrical in B. blepharista and conical in B. perloides; but these differences may be due to different pressure of the cover glass., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 66, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila squamosa Schwingenschuss, 1936 (Figs 3a and 7a) Material examined: Two syntypes, labelled „ H. Atlas, Maroc, Goundafa, 1200 m, 15.-30.VI.1933, Schwingenschuss”, one of them designated here as lectotype (Fig 3a). In NHMW. Lectotype designation: male, “ H. Atlas, Maroc, Goundafa, 1200 m, 15.-30.VI.1933, Schwingenschuss”, “ Bryophila squamosa Schwingenschuss Type ♂ ”, slide No.: RL 12464m (coll. NHMW). Taxonomic notes: The taxon was described by Schwingenschuss in Zerny (1936). Most probably, the descriptions of B. barbaria and B. squamosa were made in parallel, and both may refer to the same taxon. Thus, the statement of Boursin (1954) about their synonymy may be correct. Nevertheless, we have found some differences between their male genitalia: the uncus of B. squamosa (Fig 7a) is more slender in its distal half than that of B. barbaria (Fig 6c), the apical section of valva is also narrower and less dilated at its tip, and the vesica is remarkably broader, more discoidal. We refrain from establishing the synonymy formally until we are able to study a longer series of specimens, including females. It has to be noted that Schwingenschuss itself remarked that “ Bryophila squamosa mihi is after Βοursin also just a barbaria. Since the design matches that of barbaria, but the wing-cut is different -especially in the female-, as well as the general impression, I consider it a subspecies, especially since the animals were found at 1200 m high in a hot basin in mid-June, while barbaria was found at 2200-2700 m in July.” External diagnosis: see under B. barbaria. The lectotype shows a much more blurred condition than most of the examined specimens of B. barbaria; it is thus probably an older specimen. Genital diagnosis (Fig. 7a): see under B. barbaria and here, under Taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 61, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Zerny, H. (1936) Die Lepidopterenfauna des Grossen Atlas in Marokko und seiner Randgebiete. Memoires de la Societe des Sciences Naturelles du Maroc, 42, 1 - 163.","Boursin, C. (1954) Zwei neue Cryphia Hb. (Bryophila) - Arten aus dem vorderasiatisch-mediterranean Faunenkreis. (Beitrage zur Kenntnis der \" Agrotidae-Trifinae \" 66). Zeitschrift der Wiener Entomologischen Gesellschaft, 39, 85 - 89."]}

Bryophila katiba (Rungs, 1972) (Figs 4a and 4b; 8d) Material examined (but not yet dissected): Holotype male (Fig 4a) and two male paratypes: Maroc, Anti Atlas, Tafraout, 18- VI11-1946, in MNHN. After Rungs (1972): “Wingspan: 25 mm. A female captured with the male, shows the bottom of the wings shrimpy pink and no deep white: leander n.f. «. 1 female (dissected) (Figs 4b and 8d), Morocco, Anti-Atlas Mts, Tizi’n Tarakatine pass, 1600 m, 29°45’11”N, 8°50’02”W, 29-V-2005, leg. Gy. M. László & G. Ronkay (in coll. GR). Taxonomic notes: The taxonomic status of this taxon remains somehow uncertain, as no type material has yet been dissected, but it differs externally from the other members of the species-group by its smaller size, paler colouration and dark markings (Figs 4a and 4b). The types resemble more B. barbaria than any other taxa, though they are remarkably smaller and paler coloured than specimens of B. barbaria from the higher altitude places of the Moroccan Atlas. Besides, the genitalia of a female collected in the Anti Atlas mountains (Fig 8d) clearly differ from those of B. barbaria (Fig 8c) in the shape of the appendix bursae and in the width of the ductus bursae, much narrower in B. katiba and without the characteristic swelling of B. barbaria. External and genital diagnosis: see under taxonomic notes., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 66, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Rungs, C. E. E. (1972) Lepidopteres nouveaux du Maroc et de la Mauritanie. Bulletin du Museum National d'Histoire Naturelle, Series 3, 60, Zoologie 46, 669 - 692."]}

Bryophila perloides Guenée, 1852, stat. rev. (Figs 1a to 1d, 6a and 6b) Material examined: 2 male syntypes in coll. NHMUK, “ Andalousie ”, one of them here designated as lectotype (Figs 1a and 6b) (specimen labelled as 010918082). 1 male, 7-9-2010, Finca La Molina, Casares, Málaga, Spain (30 STF93), Penny Hale leg., coll. MF in coll. ZMC (Figs 1b and 6a). 2 males, 24.8.2018 and 21.9.2018, Campo de Gibraltar, Algeciras, Cádiz, Spain (30 STE79), José Manuel Gaona Ríos phot. (Figs 1c and 1d). Lectotype designation: male, “ Andalousie ”, “ Perloides ”, “ Syntype ” (blue ring label), “Ex MUSAEO Dris BOISDUVAL”, “Fig par J. CULOT, Noct. et Géom. d’Europe, Pl. 24 Fig 12”, “Vu par Stgr pour Catal. 1900”, “Ex Oberthür Coll. Brit. Mus. 1927-3.” (specimen labelled as 010918082, coll. NHMUK) (illustrated in Fig 1a). Taxonomic notes: the species is listed in most recent works as subordinate to Bryopsis muralis (e.g. Poole 1989, Fibiger et al. 2009; but see Fibiger et al. 2011: 33). This statement is erroneous; this species in not a Bryopsis but a true Bryophila, according to the genital configuration (see above). All five known specimens are from Andalucía, S. Spain; therefore perloides is the only European member of the group. External diagnosis: Ground colour bright olive green (Figs 1a and 1b) to dull greyish (Figs 1c and 1d). Forewings relatively small and tapered to the apex. Unlike B. domestica, subbasal and antemedian lines fused basally by an inner marginal line, forming the omega-shaped design typical of the species-group, which is completely closed in perloides; this feature is shared with the species of Bryopsis (e.g. B. muralis), where it is open to almost open in its basal part (Figs 1e and 1f). Dark spots at the costa very subtle. Unlike B. muralis, but in common with the rest of the species of the group, black X-mark on the forewings proximal to reniform stigma present. Antemarginal line absent or very subtle, in contrast to B. muralis, where it is quite apparent. Genital diagnosis: male genitalia simple, as in the rest of the species of the group; all species have similar genitalia, mostly differing in subtle details (Figs 6a to 7c). Those of B. perloides (Figs 6a and 6b) are particularly similar to those of B. barbaria (Fig 6c), but the uncus is more incrassate subapically, with shorter tip, whereas that of B. barbaria is more gradually tapered towards its apex. Additionally, the aedeagus of B. perloides shows a narrow coecum penis and broadens to its distal end, whereas in B. barbaria and other closely related species it is more or less cylindrical. Costal and hind margins of the valvae of B. perloides are almost parallel; the sacculus is weaker and smaller, the clasper smaller too, fully embedded onto the central area of valva, with slightly spatulate apex. Vesica without cornuti. The female genitalia has not yet been studied, no females have been collected so far., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on pages 56-60, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Guenee, A. (1852 - 1854) Noctuelites (II). In: Boisduval, J. A. & Guenee, A., Histoire naturelle des lnsectes. Species general des Lepidopteres. Vol. 6. Librairie Encyclopedique de Roret, Paris, pp. 29. [nº 30]","Poole, R. W. (1989) Noctuidae. In: Heppner, J. B. (Ed.), Lepidopterorum Catalogus. New Series. Vol. 118. Part. 1. E. J. Brill / Flora & Fauna Publications, Leiden, pp. 1 - 500.","Fibiger, M., Ronkay, L., Steiner, A. & Zilli, A. (2009) Noctuidae Europaeae. Vol. 11. Pantheinae, Dilobinae, Acronictinae, Eustrotiinae, Nolinae, Bagisarinae, Acontiinae, Metoponiinae, Heliothinae and Bryophilinae. Entomological Press, Soro, 504 pp.","Fibiger, M., Yela, J. L., Zilli, A., Ronkay, G. & Ronkay, L. (2011) Check list of the Quadrifid Noctuoidea of Europe. In: Witt, T. J. & Ronkay, L. (Eds.), Noctuidae Europaeae. Vol. 13. Entomological Press, Soro, pp. 23 - 44."]}

Rediscovery of Bryophila perloides On the night of 7 September 2010, a male specimen resembling B. perloides -according to the descriptions and illustrations in Guenée (1852), Rambur (1858 -[1866]) and Culot (1909-1913)- was photographed by Penny Hale at Finca La Molina, Casares, Málaga, Spain (30STF93) (Fig 1b). Following the suggestion of MF, the specimen was retained and sent to him for dissection. Examining the picture of the moth, JLY suggested B. perloides, an assumption shared by the rest of the team and confirmed by genitalia examination (Fig 6a). The genitalia showed crucial differences from those of Bryophila domestica (see e.g. Fibiger et al. 2009 and Fig 5a in this paper) and of Bryopsis muralis (Fig 5b; for taxonomic nomenclature, see Discussion), although they are much closer to those of B. domestica, from which it differs by the much shorter central claspers, wider and cygnated uncus and epiphallus broader and without distal spicules. The genitalia of one of the syntypes of B. perloides (Fig 6b) matched those of the specimen collected in Casares. It is to be noted that differences in width of the uncus between figs 6a and 6b are due to different pressure of the cover glass on the slides; same argument is applicable to the apparently different insertion position of the median claspers. The vesica is only partially everted in the syntype (Fig 6b) due to collapsing, probably related to the age of the specimen. Inspection of pictures taken by entomologists in S. Spain and uploaded to the Internet allowed us to find two additional specimens, both recorded at the Campo de Gibraltar, Algeciras, Cádiz, Spain, 30STE79, phot. José Manuel Gaona Ríos, one of them on 24.8.2018 (https://www.facebook.com/photo.php?fbid=1753590898021857&set=pcb.1753597151354565) (Fig 1c) and the other one on 21.9.2018 (https://www.facebook.com/photo.php?fbid=17850 87104872236&set=pcb.1785087411538872) (Fig 1d), showing on the forewings both key elements of most of the species of the perloides group, the basal omega-like design and the x-mark next to the reniform spot (red arrows in the figures)., Published as part of Vargas-Rodríguez, Suraya M., Fibiger, Michael, Honey, Martin R., Ronkay, Gábor, Yela, José Luis, Zilli, Alberto & Ronkay, László, 2020, Bryophila perloides Guenée, 1852 (Lepidoptera: Noctuidae: Bryophilinae), an overlooked member of the European Noctuidae fauna, with upgrading of Bryopsis Boursin, 1970 to generic level (Taxonomic studies on the western Palaearctic Bryophilinae, No. 1), pp. 53-70 in Zootaxa 4845 (1) on page 55, DOI: 10.11646/zootaxa.4845.1.4, http://zenodo.org/record/4406308, {"references":["Guenee, A. (1852 - 1854) Noctuelites (II). In: Boisduval, J. A. & Guenee, A., Histoire naturelle des lnsectes. Species general des Lepidopteres. Vol. 6. Librairie Encyclopedique de Roret, Paris, pp. 29. [nº 30]","Rambur, P. (1858 - [1866]) Catalogue systematique des Lepidopteres d'Andalousie. Bailliere, Paris. 412 pp.","Fibiger, M., Ronkay, L., Steiner, A. & Zilli, A. (2009) Noctuidae Europaeae. Vol. 11. Pantheinae, Dilobinae, Acronictinae, Eustrotiinae, Nolinae, Bagisarinae, Acontiinae, Metoponiinae, Heliothinae and Bryophilinae. Entomological Press, Soro, 504 pp."]}

[EN A new species of genus Lygephila Billberg, 1820 (Lepidoptera, Noctuidae) is described. It is named Lygephila herrerai n. sp., and his locus typicus is the Sierra of Cazorla, in Jaen (Spain). General comments on genus Lygephila and on its Iberian species are added., [ES] Se describe una nueva especie del género Lygephila Billberg, 1820 (Lepidoptera, Noctuidae), que es denominada Lygephila herrerai n. sp. Su localidad típica es la Sierra de Cazorla, en Jaén (España). Se añaden comentarios generales sobre el género Lygephila y sus especies ibéricas.

[EN] The Catocalinae species (Lepidoptera, Noctuidae) that inhabit the Madrid province are studied, giving data on their morphology, distribution and ecology. The description of the genitalic armours of those not belonging to the genus Catocala is emphasized, since the Catocala species were already studied by AGENJ0 (1959). However, the genitalia of Catocala mariana Rambur, 1858, bona species, is examined, comparing it with that of Catocala eutychea (Treitschke, 1835). One species, Ophiusa tirhaca (Cramer, 1777), is mentioned for the first time of Madrid. Some data about Clytie illunaris (Hübner, [1813]) and Clytie sancta (Staudinger, 1897), both previously mentioned as existing in Madrid, are given, explaining the most important differences that could be observed in their genitalia. As it is commented, the latter and Clytie syriaca (Bougnion, 1837) must be eliminated from the lists of the Iberian fauna until their presence in our peninsula is shown by fiable data., [ES] Se estudian, aportando datos tanto de morfología como de distribución y ecología, las especies integrantes de la subfamilia Catocalinae (Lepidoptera, Noctuidae) encontradas en la provincia de Madrid, poniendo mayor énfasis en la descripción de las estructuras genitales de aquellas que no pertenecen al género Catocala, ya tratado por AGENJO (1959). Sí se trata, sin embargo, la genitalia de Catocala mariana Rambur, 1858, bona species, comparándola con la de Catocala eutychea (Treitschke, 1835). Se cita una especie como nueva para Madrid, Ophiusa tirhaca (Cramer, 1777). Se ofrecen algunos datos de interés sobre Clytie illunaris (Hübner, [1813 ]) y Clytie sancta (Staudinger, 1897) (ambas citadas de Madrid), indicándose las diferencias más importanes que pueden encontrarse entre sus estructuras genitales; como se comenta, tanto esta última especie como Clytie syriaca (Bugnion, 1837) han de ser eliminadas de las listas de la fauna ibérica mientras no se demuestre con datos fiables su presencia en nuestra Península.