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Scirtothrips quercus sp.n. (Figs 1–11) Female macroptera. Body yellow including legs with brown markings on pronotum, anterior part of mesonotum and unguitractor plates; antennal segment I pale; segments II–VIII brown; fore wings brown with apex paler (Fig. 1). Head broader than long with transverse striations; ocellar setae III located between fore and hind ocelli, close to fore ocelli; with four pairs of postocular setae, poI and poII longer, poIII minute (Fig. 3). Antennae 8-segmented; III and IV each with forked sense cone (Fig. 10); II–VI with rows of microtrichia. Pronotum broader than long with 12 discal setae, pmII two times longer than pmIII, but pmI subequal to pmIII (Fig. 3); Mesonotum with transverse striations, anteromedian campaniform sensilla absent, median pair of setae far away from posterior margin; metanotum reticulate (Fig. 3), median pair of setae located on anterior margin, campaniform sensilla absent; fore wing first vein with 9 setae, second vein with 3 setae, clavus with four veinal setae and one discal seta with apical seta longest, posterior fringes straight but wavy at tips (Fig. 9). Abdominal tergites II–VII with setae S1 small, closer together than their length, lateral microtrichial field of abdominal tergites with 3 discal setae (Fig. 7), tergite VIII with few microtrichia anteromedially and posteromarginal comb complete (Fig. 5); tergite IX without microtrichia; antecostal ridge on tergites and sternites III–VIII dark; sternites with microtrichia restricted to lateral areas only and absent in middle (Fig. 8). Measurements. (Holotype female in microns). Body length 1007. Head length (width across cheeks) 91 (111); compound eye dorsal length (width) 59 (35); pronotum median length (width) 81 (145). Fore wing length 569. Ovipositor length 163. Antenna segments I–VIII length (width): 23 (22), 34 (24), 39 (18), 40 (17), 38 (16), 38 (15), 7 (6), 11 (4). Male macroptera. Similar to female except pale yellow body (Figs 2, 4). Antennal segments I–II pale; III–VIII brown (Fig. 11). Abdominal tergite IX with drepanae (Fig. 6). Abdominal sternites without pore plate. Measurements (Male paratype in microns): Body length 681. Head length (width across cheeks) 64 (99); compound eye dorsal length (width) 44 (36); pronotum median length (width) 68 (111). Fore wing length 443. Antennal segments I–VIII length (width): 18 (19), 29 (20), 31 (15), 33 (14), 29 (14), 33 (13), 5 (6), 9 (3) Material studied. Holotype female, INDIA, Uttarakhand, Almora, (N29.57, E79.48, 1601 m) on Quercus sp. (Fagaceae), 11.iii.2020, (Reg. No. 12700/H17), Tyagi and Patidar (in NZC, Kolkata, India). Paratypes: 27 females, 2 males, all from same locality as holotype (Reg. No. 11311/H17 to 11336/H17,12701/H17 to 12703/H17); INDIA, Himachal Pradesh, Mandi (N31.97, E76.84, 1425 m), two females from general vegetation, 15.x.2013, Kumar and Tyagi (Reg. No.6517/H17, 6518/H17); Mandi (N31.43, E77.07, 2022 m), 4 females on general vegetation, 05.vi.2014, Kumar and Tyagi (Reg. No.6519/H17 to 6522/H17); Mandi (N31.42, E77.07, 1925 m), one female on fern, 04.vi.2014, Kumar and Tyagi (Reg. No.6523/H17) (in NZC, Kolkata, India); INDIA, Jammu & Kashmir, Chassana, Reasi (N33.21, E74.44, 1220 m), one female on Coriandrum sativum (Apiaceae), 19.iv.2022, Pal (Reg. No. 12368/H17); Paikul, Doda, (N33.06, E75.51, 1720 m), 4 females on Cucumis sativus (Cucurbitaceae), 9.viii.2022, Pal (Reg. No. 12704/H17 to 12707/H17) (in NZC, Kolkata, India). Etymology: The species is named after the Quercus sp. Comments. According to the key to Scirtothrips species from India, this new species is very close to S. kenyensis Mound. They can be distinguished by the following character states: ocellar III between the fore and hind ocelli (close together on tangent between anterior margins of hind ocelli); head and ocellar triangle without brown shading (head anterior margin and ocellar triangle with shading in kenyensis); fore wing posteromarginal cilia straight but tips wavy (completely straight in kenyensis); tergite II–IX without shading medially (with shading medially in kenyensis); pmII two times longer than pmIII (pmII three times longer than pmIII in kenyensis); fore wing first vein with 9 setae with 3+3+1+1+1 arrangement (8 setae with 3+2+1+1+ 1 in kenyensis). Further, this species is close to S. dobroskyi Moulton but it can be differentiated by ocellar III located between fore and hind ocelli (between hind ocelli in dobroskyi); pmII two times longer than pmI (3 times in dobroskyi), fore wing posteromarginal cilia straight, wavy at tips (straight in dobroskyi), tergite VIII with few rows of microtrichia medially near to anterior margin (absent in dobroskyi), tergite IX without microtrichia (weak in dobroskyi). According to available keys to species of genus Scirtothrips from Australia and Africa, the new species is similar to Scirtothrips longipennis (Bagnall), and shares the position of ocellar setae III, colour of fore wings, and tergal antecostal ridges. The new species differs from longipennis by the following characters: antennal segment II brown (II yellow in longipennis); pronotum with very close striations (wide apart in longipennis); clavus with 4 veinal setae (3 in longipennis); fore wing posteromarginal cilia straight, wavy at tips (wavy in longipennis); tergite VIII with few rows of microtrichia medially (absent in longipennis); tergite IX without microtrichia medially (with microtrichia in longipennis).

Pal, Shash, Patidar, Abhishek, Panjaliya, Rakesh Kumar, Kumar, Vikas, Tyagi, Kaomud (2023): Species of the genus Scirtothrips from India (Thysanoptera, Thripidae). Zootaxa 5306 (3): 392-396, DOI: 10.11646/zootaxa.5306.3.7, URL: http://dx.doi.org/10.11646/zootaxa.5306.3.7

Key to ten species of Scirtothrips from India (asterisk indicates species included from literature) 1. Ocellar I setae absent [tergites III–V with S1 setae wider apart than their length, male tergite IX with drepanae]... malayensis -. Ocellar I setae present................................................................................. 2 2. Microtrichia on abdominal sternites IV–VI extend completely across median area at least on posterior half.............. 3 -. Microtrichia on abdominal sternites IV–VI restricted to lateral areas; male tergite IX with drepanae................... 7 3. Antecostal ridge on tergites dark but interrupted medially [antennae and fore wings dark, male tergite IX without drepanae]................................................................................................. hitam -. Antecostal ridge on tergites not interrupted medially......................................................... 4 4. Abdominal tergites with pale antecostal ridge and median area [lateral microtrichial field of tergites with 4–5 discal setae, male tergite IX without drepanae].................................................................... oligochaetus -. Abdominal tergites with dark antecostal ridge............................................................... 5 5. Abdominal tergites without dark median area, lateral microtrichial field of tergites with 4–5 discal setae; male tergite IX with drepanae....................................................................................... fulleri * -. Abdominal tergites with dark median area, lateral microtrichial field of tergites with 3 discal setae; male tergite IX without drepanae............................................................................................ 6 6. Sternites with dark transverse antecostal ridge......................................................... dorsalis -. Sternites without dark transverse antecostal ridge................................................... pteridicola * 7. Fore wing posteromarginal cilia either completely wavy or wavy at tips.......................................... 8 -. Fore wing posteromarginal cilia straight................................................................... 9 8. Ocellar setae III placed between fore and hind ocelli, close to fore ocelli; antecostal ridge on tergites dark; lateral microtrichial field of abdominal tergites with 3 setae; microtrichia absent on tergite IX; fore wing dark with pale apex, posteromarginal cilia wavy at tips................................................................................ quercus sp.n. -. Ocellar setae III placed anterior to tangent between anterior margins of hind ocelli; antecostal ridge on tergites pale; lateral microtrichial field of abdominal tergites with 4 setae; microtrichia present on tergite IX; fore wing pale, posteromarginal cilia completely wavy.............................................................................. mangiferae 9. Ocellar setae III placed between the hind ocelli toward the posterior margin of hind ocelli; pronotal pmII four times longer than pmI; metanotum with reticulate sculpture, reticles with internal markings in middle, median setae far behind anterior margin; fore wing second vein with one seta; tergite IX with microtrichia medially................................ bispinosus -. Ocellar III placed anterior to hind ocelli, just at the line of anterior margin of hind ocelli; pronotal pmII......... three times longer than pmI; metanotum with reticulate sculpture, reticles without internal markings; median setae at anterior margin; fore wing second vein with 3 setae; tergite IX without microtrichia medially................................... kenyensis, Published as part of Pal, Shash, Patidar, Abhishek, Panjaliya, Rakesh Kumar, Kumar, Vikas & Tyagi, Kaomud, 2023, Species of the genus Scirtothrips from India (Thysanoptera, Thripidae), pp. 392-396 in Zootaxa 5306 (3) on pages 392-393, DOI: 10.11646/zootaxa.5306.3.7, http://zenodo.org/record/8063100

Jammuthrips paikulensis sp.n. (Figs 1 –12) Female macroptera. Body brown, fore wing banded with three white and four dark bands, one small grey patch just above the clavus anterior margin, second grey patch before middle and third grey patch in middle and fourth in posterior half, extreme distal end white; legs yellow, fore tibia with small grey patch in middle, unguitractor plates dark; antennal segments I–II white and III–VIII dark (Fig. 1). With the character states in the generic diagnosis: Head with ocellar setae II smaller than III, maxillary palp 2-segmented, but in one female, maxillary palp 2- or 3-segmented (Fig. 2). Antennal segment VI with a long sensorium on inner margin, reaching middle of VIII, VII with outer sense cone reaching to apical margin of VIII (Fig. 9). Mesothoracic furca with spinula (Fig. 10). Fore wing with 34 costal seta, first vein with 13 setae, second vein with one seta; clavus with 4 setae (Fig. 7). Abdominal tergites II–VII with reticulation near median setae and laterally, reticles with wrinkles; length of median setal pair on tergites I–VIII gradually increasingly (Figs 4, 12); median pair of setae on tergite IX slightly longer than lateral pair, with median campaniform sensilla; tergite IX with numerous microtrichia on posterior half medially. Sternite II with two pairs of posteromarginal setae, III–VII with three pairs. Measurements (holotype female, in microns). Body length 870. Head, length 40, width across genae (across eyes) 98 (96); compound eye dorsal length 37, ocellar III length 23; maxillary palp length 47. Antenna length 180. Pronotum median length 86 width at anterior end (width at posterior end) 115(132). Fore wing length 605. Hind tibia length 169, fore tarsus length 32, mid tarsus length 32, hind tarsus length 45. Ovipositor length 144. Antennal segments I–VIII length (width): 14 (16), 28 (23), 22 (14), 30 (16), 31 (16), 30 (11), 7 (4) and 12 (3). Male Unknown. Material studied. Holotype female: JAMMU and KASHMIR: Doda: Paikul, on leaves of Sorghum vulgare, 30.viii.2021, (Reg. No. 12113/H17) by Shash Pal, deposited at the NZC, Kolkata, India. Paratypes: 6 females same data as holotype (Reg. No. 12114/H17 to 12119/H17). Etymology: The species named after the type locality.

Jammuthrips paikulensis sp.n. (Figs 1 –12) Female macroptera. Body brown, fore wing banded with three white and four dark bands, one small grey patch just above the clavus anterior margin, second grey patch before middle and third grey patch in middle and fourth in posterior half, extreme distal end white; legs yellow, fore tibia with small grey patch in middle, unguitractor plates dark; antennal segments I–II white and III–VIII dark (Fig. 1). With the character states in the generic diagnosis: Head with ocellar setae II smaller than III, maxillary palp 2-segmented, but in one female, maxillary palp 2- or 3-segmented (Fig. 2). Antennal segment VI with a long sensorium on inner margin, reaching middle of VIII, VII with outer sense cone reaching to apical margin of VIII (Fig. 9). Mesothoracic furca with spinula (Fig. 10). Fore wing with 34 costal seta, first vein with 13 setae, second vein with one seta; clavus with 4 setae (Fig. 7). Abdominal tergites II–VII with reticulation near median setae and laterally, reticles with wrinkles; length of median setal pair on tergites I–VIII gradually increasingly (Figs 4, 12); median pair of setae on tergite IX slightly longer than lateral pair, with median campaniform sensilla; tergite IX with numerous microtrichia on posterior half medially. Sternite II with two pairs of posteromarginal setae, III–VII with three pairs. Measurements (holotype female, in microns). Body length 870. Head, length 40, width across genae (across eyes) 98 (96); compound eye dorsal length 37, ocellar III length 23; maxillary palp length 47. Antenna length 180. Pronotum median length 86 width at anterior end (width at posterior end) 115(132). Fore wing length 605. Hind tibia length 169, fore tarsus length 32, mid tarsus length 32, hind tarsus length 45. Ovipositor length 144. Antennal segments I–VIII length (width): 14 (16), 28 (23), 22 (14), 30 (16), 31 (16), 30 (11), 7 (4) and 12 (3). Male Unknown. Material studied. Holotype female: JAMMU and KASHMIR: Doda: Paikul, on leaves of Sorghum vulgare, 30.viii.2021, (Reg. No. 12113/H17) by Shash Pal, deposited at the NZC, Kolkata, India. Paratypes: 6 females same data as holotype (Reg. No. 12114/H17 to 12119/H17). Etymology: The species named after the type locality., Published as part of Pal, Shash, Kumar, Vikas, Panjaliya, Rakesh Kumar & Tyagi, Kaomud, 2022, A new genus and species of subfamily Dendrothripinae (Thysanoptera: Thripidae) from India, pp. 383-388 in Zootaxa 5175 (3) on pages 384-387, DOI: 10.11646/zootaxa.5175.3.5, http://zenodo.org/record/7003696

Jammuthrips gen. n. (Figs 1 –12) Type species: Jammuthrips paikulensis sp.n. Female macroptera. Head broader than long, genae short, slightly concave, with two pairs of ocellar setae, pair I absent, pair III arising outside the ocellar triangle, without postocular setae and without prominent sculpture (Figs 3, 8); eyes slightly enlarged ventrally, without pigmented facets; mouth cone long and cone shaped, extending to mesosternum; maxillary palps long, 2-segmented (Figs 2, 10). Antennae 8-segmented, III with simple and IV with forked sense cone (Fig. 9). Pronotum wider than long, with irregular reticulations, reticles with internal wrinkles, with one pair of long anteromarginal setae, and two pairs of posteroangular setae, discal setae long, subequal to or shorter than posteroangular setae (Figs 5, 8); ferna divided (Fig. 10). Mesonotum without anteromedian campaniform sensilla; median pair of setae far ahead of posterior margin, mesothoracic sternoplural sutures present; metanotum with reticulate sculpture, reticles with wrinkles, median pair of setae located far behind the anterior margin, submedian pair located near anterior margin, without campaniform sensilla (Fig. 11). Fore wing posterior margin straight, surface uniformly covered with microtrichia, costal setae small, first vein setae slightly longer than costal setae, apex bluntly rounded, with one small apical seta (Figs 6, 7); anteromarginal cilia arise far behind the anterior margin; posterior fringes straight. All tarsi 1-segmented; hind tarsi elongate, longer than fore and mid tarsi. Median setae on tergites I–VIII long and close together, distance between them less than their length, tergite I with small median setae; lateral thirds of tergites II–VIII with reticulation; reticles with wrinkles, VIII without posteromarginal comb of microtrichia (Fig. 12). Sternites III–VII each with three pairs of small marginal setae. Comments. This new genus is similar to the genus Dendrothrips Uzel in the following character states: fore wing anteromarginal cilia arising far behind anterior margin, prosternal ferna divided, maxillary palps 2-segmented and fore wing with relatively straight posterior margin. In contrast to Dendrothrips, the new genus has the fore wing costal setae extending to anterior margin, maxillary palps long, pronotal anteromaginal setae prominent, mouthcone reaching to middle of mesosternum.

Jammuthrips gen. n. (Figs 1 –12) Type species: Jammuthrips paikulensis sp.n. Female macroptera. Head broader than long, genae short, slightly concave, with two pairs of ocellar setae, pair I absent, pair III arising outside the ocellar triangle, without postocular setae and without prominent sculpture (Figs 3, 8); eyes slightly enlarged ventrally, without pigmented facets; mouth cone long and cone shaped, extending to mesosternum; maxillary palps long, 2-segmented (Figs 2, 10). Antennae 8-segmented, III with simple and IV with forked sense cone (Fig. 9). Pronotum wider than long, with irregular reticulations, reticles with internal wrinkles, with one pair of long anteromarginal setae, and two pairs of posteroangular setae, discal setae long, subequal to or shorter than posteroangular setae (Figs 5, 8); ferna divided (Fig. 10). Mesonotum without anteromedian campaniform sensilla; median pair of setae far ahead of posterior margin, mesothoracic sternoplural sutures present; metanotum with reticulate sculpture, reticles with wrinkles, median pair of setae located far behind the anterior margin, submedian pair located near anterior margin, without campaniform sensilla (Fig. 11). Fore wing posterior margin straight, surface uniformly covered with microtrichia, costal setae small, first vein setae slightly longer than costal setae, apex bluntly rounded, with one small apical seta (Figs 6, 7); anteromarginal cilia arise far behind the anterior margin; posterior fringes straight. All tarsi 1-segmented; hind tarsi elongate, longer than fore and mid tarsi. Median setae on tergites I–VIII long and close together, distance between them less than their length, tergite I with small median setae; lateral thirds of tergites II–VIII with reticulation; reticles with wrinkles, VIII without posteromarginal comb of microtrichia (Fig. 12). Sternites III–VII each with three pairs of small marginal setae. Comments. This new genus is similar to the genus Dendrothrips Uzel in the following character states: fore wing anteromarginal cilia arising far behind anterior margin, prosternal ferna divided, maxillary palps 2-segmented and fore wing with relatively straight posterior margin. In contrast to Dendrothrips, the new genus has the fore wing costal setae extending to anterior margin, maxillary palps long, pronotal anteromaginal setae prominent, mouthcone reaching to middle of mesosternum., Published as part of Pal, Shash, Kumar, Vikas, Panjaliya, Rakesh Kumar & Tyagi, Kaomud, 2022, A new genus and species of subfamily Dendrothripinae (Thysanoptera: Thripidae) from India, pp. 383-388 in Zootaxa 5175 (3) on page 384, DOI: 10.11646/zootaxa.5175.3.5, http://zenodo.org/record/7003696