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A total of 58 (eight known and 50 new) species of the subgenus Stegana (Steganina) from China were surveyed and (re)described: S. (S.) bacilla Chen & Aotsuka, 2004, S. (S.) belokobylskiji Sidorenko, 1997, S. (S.) hirticeps Wang, Gao, & Chen, 2013, S. (S.) izu Sidorenko, 1997, S. (S.) kanmiyai Okada & Sidorenko, 1992, S. (S.) masanoritodai Okada & Sidorenko, 1992, S. (S.) maymyo Sidorenko, 1997, stat. rev., S. (S.) nigripes Zhang & Chen, 2015, S. (S.) alafoliacea Zhang & Chen, sp. nov., S. (S.) baoxing Li & Chen, sp. nov., S. (S.) bibarbata Li & Chen, sp. nov., S. (S.) bimai Cui & Chen, sp. nov., S. (S.) cinereipecta Zhang & Chen, sp. nov., S. (S.) cardua Cui & Chen, sp. nov., S. (S.) cordhirsuta Wang & Chen, sp. nov., S. (S.) cornuta Li & Chen, sp. nov., S. (S.) cucullata Li & Chen, sp. nov., S. (S.) cultella Cui & Chen, sp. nov., S. (S.) curvitabulata Cui & Chen, sp. nov., S. (S.) daiya Cui & Chen, sp. nov., S. (S.) dendrophila Zhang & Chen, sp. nov., S. (S.) flabella Li & Chen, sp. nov., S. (S.) flavipes Li & Chen, sp. nov., S. (S.) formosa Zhang & Chen, sp. nov., S. (S.) fusca Li & Chen, sp. nov., S. (S.) fuscipes Li & Chen, sp. nov., S. (S.) glaucopalpula Cui & Chen, sp. nov., S. (S.) haba Zhang & Chen, sp. nov., S. (S.) hirticlavata Cui & Chen, sp. nov., S. (S.) iaspidea Zhang & Chen, sp. nov., S. (S.) idiasta Cui & Chen, sp. nov., S. (S.) kanda Cui & Chen, sp. nov., S. (S.) labao Li & Chen, sp. nov., S. (S.) lancang Li & Chen, sp. nov., S. (S.) latifoliacea Wang & Chen, sp. nov., S. (S.) liusanjieae Li & Chen, sp. nov., S. (S.) magniflava Cui & Chen, sp. nov., S. (S.) mailangang Li & Chen, sp. nov., S. (S.) marenubila Cui & Chen, sp. nov., S. (S.) menghai Zhang & Chen, sp. nov., S. (S.) menglian Li & Chen, sp. nov., S. (S.) minutiflava Li & Chen, sp. nov., S. (S.) multiprocera Li & Chen, sp. nov., S. (S.) nayun Li & Chen, sp. nov., S. (S.) nigridentata Wang & Chen, sp. nov., S. (S.) nigripalpula Cui & Chen, sp. nov., S. (S.) otphylla Cui & Chen, sp. nov., S. (S.) radiciflava Zhang & Chen, sp. nov., S. (S.) rava Cui & Chen, sp. nov., S. (S.) sciophila Li & Chen, sp. nov., S. (S.) septencolorata Li & Chen, sp. nov., S. (S.) serrata Zhang & Chen, sp. nov., S. (S.) silvestrella Zhang & Chen, sp. nov., S. (S.) simola Cui & Chen, sp. nov., S. (S.) yani Li & Chen, sp. nov., S. (S.) yixiang Zhang & Chen, sp. nov., S. (S.) zaduo Cui & Chen, sp. nov., and S. (S.) zhuoma Cui & Chen, sp. nov. We also provided a complete list of Chinese Steganina species together with their geographical distributions. In addition, the majority of currently available DNA barcode (partial sequence of the mitochondrial cytochrome c oxidase subunit I (COI) gene) sequences of this subgenus (435 sequences of 102 spp.) were employed in a molecular analysis for species delimitation. Taken together, morphology- and molecular-based species delimitation results reached a consensus for an overwhelming majority of these Steganina species (98 of 102 spp.).

Stegana (Steganina) nigripes Zhang & Chen, 2015 (Fig. 20A–C) Stegana (Steganina) nigripes Zhang & Chen, 2015: 135. Remarks. Since the terminalia of the only male specimen was partly damaged (probably eaten by a cockroach) and undescribed, the specimens collected from Yingjiang, Yunnan showed no other significant morphological differences with the type material of S. (S.) nigripes from Menghai, Yunnan. They were conservatively identified as conspecific. Description. Female terminalia (Fig. 20A–C): Epiproct and cercus with numerous setae, lacking pubescence (Fig. 20A). Eighth sternite about 2 times as wide as long, with equally distributed numerous prensisetae (Fig. 20B). Hypoproct with numerous setae and much pubescence (Fig. 20C). Specimens examined. CHINA: 1♂ (SCAU, No. 125409, DNA #1985), 1♀ (SCAU, No. 125408, DNA #1981), Tongbiguan, Yingjiang, Yunnan, 24°37'04''N, 97°40'03''E, 19.v.2016, alt. 1310m, 13.v.2016, ex tussock, J. Huang; 1♀ (SCAU, No. 125410, DNA #1981), Botanic Garden, Ruili, Yunnan, 24°04'19''N, 97°49'08''E, alt. 1170m, 22.v.2016, ex tussock, Y.Q. Liu. Distribution. China (Yunnan).

Stegana (Steganina) serratoprocessata Chen & Chen, 2009 Stegana (Steganina) serratoprocessata Chen & Chen, 2009a: 499. Specimens examined. CHINA: 3♂, 3♀ (SCAU, Nos. 125530–35, DNA #1045), Maoershan, Xing’an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 22–24.ix.2006, ex tree trunks, J.J. Jiang, F. Zhao; 3♂, 3♀ (SCAU, Nos. 125536–41, DNA #1949, 1950), Jiuwanshan, Huanjiang, Guangxi, 25°18'48''N, 108°38'02''E, alt. 1500m, 14.ix.2015, ex tree trunks, Y.Q. Liu; 1♂ (SCAU, No. 125542, DNA #2612), Shengtangshan, Jinxiu, Guangxi, 23°58'23''N, 110°06'45''E, alt. 1300m, 31.vii.2017, ex tree trunk, H.W. Chen; 1♂ (SCAU, Nos. 125543, DNA #599), Wuliangshan, Jingdong, Yunnan, 24°20'40''N, 100°48'01''E, alt. 2200m, 27.vii.2006, ex tree trunk, J.J. Gao; 2♂, 1♀ (SCAU, Nos. 125544–46, DNA #2699, 2731), Botanic Garden, Ruili, Yunnan, 24°04'19''N, 97°49'08''E, alt. 1170m, 22.iv.2018, ex tree trunks, H.W. Chen, L. Gong, Y. Lin; 1♂ (SCAU, No. 125547, DNA #2352), Menglai, Cangyuan, Yunnan, 23°13'20''N, 99°15'04''E, alt. 1240m, 30.x.2017, ex tree trunk, B.X. Li; 13♂, 9♀, Yixiang, Pu’er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 11♂, 5♀ (SCAU, Nos. 125548–63, DNA #1415, 1445, 1442, 2191) on 13.v.2012, ex tree trunks, H.W. Chen, J.J. Gao, 2♂, 4♀ (SCAU, Nos. 125564–69, DNA #1902, 2560, 2561) on 28– 30.iii.2013, ex tussocks, J.J. Gao; 1♂ (Nos. 125570, DNA #997), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt. 1000m, 10.x.2010, ex tree trunk, L. Wu. Distribution. China (Guangdong, Hainan, Guangxi *, Yunnan *, Xizang *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 54-55, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Chen, X. P. & Chen, H. W. (2009 a) Four new species of the subgenus Steganina (Diptera: Drosophilidae) from southern China. Annales Zoologici, 59, 495 - 501. https: // doi. org / 10.3161 / 000345409 X 484874"]}

Stegana (Steganina) silvestrella Zhang & Chen, sp. nov. (Figs. 13G, 13H, 61) Diagnosis. This species is similar to S. (S.) flavipedes sp. nov. in the shape of male terminalia (Figs. 38, 61), but can be distinguished from the latter by having the pregonite blurry or absent (Fig. 61C, D); the aedeagus with numerous minute, serrated processes distally (Fig. 61C, D). Description. Male and female: First flagellomere yellow. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally (Fig. 13G). Katepisternum yellowish (Fig. 13H). Scutellum dark brown in male, brownish yellow in female, black near distal margin (Fig. 13G). Halter mostly yellowish (Fig. 13H). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites brown medially, yellow laterally (Fig. 13G, H); sternites yellow, about 1.8 times as wide as long. Male terminalia (Fig. 61): Epandrium with ca. 9 setae on dorsal to posterior portions per side (Fig. 61A). Surstylus with some setulae near distal margin, lacking serrated processes and pubescence (Fig. 61A, B). Cercus with much pubescence (Fig. 61A). Hypandrium roundly expanded anteromedially (Fig. 61C, D). Aedeagus tubular, distally expanded and rounded, lacking pubescence (Fig. 61C, D). Aedeagal sheath lacking serrated process (Fig. 61C, D). Subepandrial sclerite slightly half-rounded (Fig. 61G). Measurements and indices. BL = 2.63 mm in holotype (2.80–3.40 in 2♀ paratypes), ThL = 1.33 mm (1.33 in ♀), WL = 2.13 mm (2.00– 2.47 in ♀), WW = 1.00 mm (1.00– 1.13 in ♀), arb = 4/3 (4–5/3–4), avd = 1.00 (0.80–1.14), adf = 1.25 (1.00–1.40), flw = 2.42 (2.01–2.50), FW/HW = 0.45 (0.39–0.53), ch/o = 0.09 (0.08–0.11), prorb = 1.38 (0.83–1.33), rcorb = 0.75 (0.50–0.78), vb = 0.67 (0.60–0.80), dcl = 0.46 (0.46–0.48), presctl = 0.72 (0.70–0.82), sctl = 2.34 (2.11–2.50), sterno = 0.80 (0.60–1.11), orbito = 2.67 (2.12–2.67), dcp = 0.28 (0.25–0.28), sctlp = 1.86 (1.75–2.50), C = 1.67 (1.56–2.00), 4c = 1.29 (1.13–1.50), 4v = 1.91 (1.43–2.01), 5x = 2.43 (2.01–2.60), ac = 10.02 (9.00–12.01), M = 0.76 (0.73–0.84), C3F = 0.59 (0.64–0.87). Type materials. Holotype ♂ (SCAU, No. 125657, DNA #2752), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 700m, 25.iv.2007, ex tree trunk, H.W. Chen. Paratypes: 2♀ (SCAU, Nos. 125658, 59, DNA #2562), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt. 1320m, 10.v.2016, ex tree trunks, J. Huang, Y.L. Wang. Distribution. China (Yunnan). Etymology. From the Latin word " silvestris " (= arboreal), referring to the arboreal Steganina flies., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 96-98, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) cucullata Li & Chen, sp. nov. (Figs. 7C, 7D, 33) Diagnosis. This species closely resembles S. (S.) eurystoma Wang, Gao, & Chen, 2013 in the shape of male terminalia (as fig. 3 in Wang et al. 2013, Fig. 33), but can be distinguished from the latter by having the pregonite blurry or absent (Fig. 33C, D, J, K); the aedeagus lacking serrated process near distal margin (Fig. 33C, D, J, K); the aedeagal sheath distally slightly strong and dorsad curved (Fig. 33E, F, L, M). Remarks. The type specimens collected from Guangxi and Sichuan slightly differ from those from Chongqing in the surstylus lacking serrated process, with 1 peg-like tooth near distal margin (Fig. 33B, I), the cercus lacking pubescence (Fig. 33A, H), and the aedeagus with several minute, serrated processes distally and pubescence distoventrally (Fig. 33C, D, J, K). They were conservatively identified as conspecific. Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum dark brown, with brownish yellow longitudinal stripes submedially and laterally (Fig. 7C). Katepisternum yellowish (Fig. 7D). Scutellum dark brown (Fig. 7C). Halter mostly grey (Fig. 7D). Fore femur with a row of 4 or 5 sturdy setae on distal part of posteroventral surface. Abdominal tergites dark brown (Fig. 7C, D); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 33): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 33A, H). Surstylus with numerous setulae near distal margin (Fig. 33A, B, H, I). Hypandrium roundly expanded anteromedially (Fig. 33C, D, J, K). Pregonite with 1 or 2 sensilla (Fig. 33C, J, K). Aedeagus spoon-shaped, distally broadened (Fig. 33C, D, J, K). Subepandrial sclerite slightly vaulted, broadened distally (Fig. 33G, N). Measurement and indices. BL = 3.64 mm in holotype (2.35–3.15 in 5♂, 2.40–4.00 in 5♀ paratypes), ThL = 1.75 FW/HW = 0.37 (0.30–0.48), ch/o = 0.27 (0.18–0.36), prorb = 0.71 (0.76–1.38), rcorb = 0.53 (0.60–0.90), vb = 0.38 (0.50–0.78), dcl = damaged (0.36–0.75), presctl = 0.52 (0.31–0.78), sctl = 1.52 (1.56–2.13), sterno = 0.56 (0.55–1.17), orbito = 2.50 (1.20–2.40), dcp = 0.43 (0.15–0.34), sctlp = 1.64 (1.54–2.00), C = 1.71 (1.68–2.05), 4c = 1.24 (0.95–2.40), 4v = 1.88 (1.48–2.33), 5x = 1.57 (0.72–1.91), ac = 10.50 (7.33–12.67), M = 0.65 (0.53–0.82), C3F = 0.53 (0.54–0.72). Type materials. Holotype ♂ (SCAU, No. 121109, DNA #546), CHINA: Fengtongzhai, Baoxing, Sichuan, 30°32'42''N, 102°54'24''E, alt. 1700m, 19.ix.2005, ex tree trunk, H.W. Chen. Paratypes: 1♀ (SCAU, No. 121110), other data same as holotype; 5♂, 3♀ (SCAU, Nos. 125595–603, DNA #2609), 18.vii.2017, L. Gong, Y. Lin, Y.L. Wang, other data same as holotype; 1♂ (SCAU, No. 125604, DNA #983), Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 24.ix.2006, ex tree trunk, J.J. Jiang; 1♂ (SCAU, No. 125605, DNA #1782), Simianshan, Jiangjing, Chongqing, 28°37'19''N, 106°23'50''E, alt. 1108m, 7.vii.2014, ex tree trunk, Y.Q. Liu. Distribution. China (Guangxi, Chongqing, Sichuan). Etymology. From the Latin word " cucullatus " (= cucullate), referring to the spoon-shaped aedeagus.

VI. Stegana (Steganina) shirozui species group (26 spp.) alianya Zhang & Chen, 2021; China (Yunnan). ctenaria Nishiharu, 1979; Russia (Far East), China (Jilin, Liaoning, Taiwan), Japan (Hokkaido, Honshu, Kyushu), Korea. danbaensis Chen & Chen, 2012; China (Sichuan). dianensis Chen & Chen, 2012; China (Yunnan). diodonta Zhang & Chen, 2021; China (Yunnan). glabra Chen & Chen, 2012; China (Shaanxi *, Hubei *, Sichuan *, Guizhou, Yunnan). jiajinshanensis Chen, Gao, & Chen, 2009; China (Sichuan, Yunnan). jianfenglingensis Chen, Gao, & Chen, 2009; China (Hainan, Guangxi *). maoershanensis Chen, Gao, & Chen, 2009; China (Guangxi). masanoritodai Okada & Sidorenko, 1990; China (Jilin, Liaoning *), Russia (Far East), Japan (Hokkaido, Honshu, Kyushu). montana Chen & Chen, 2012; China (Yunnan). multidentata Chen, Gao, & Chen, 2009; China (Hubei, Sichuan *). oligochaeta Chen & Chen, 2012; China (Yunnan). parvispina Chen & Chen, 2012; China (Shaanxi). pianmaensis Chen & Chen, 2012; China (Yunnan). protuberans Chen & Chen, 2012; China (Yunnan). qinlingensis Chen, Gao, & Chen, 2009; China (Shaanxi, Sichuan). shennongi Chen, Gao, & Chen, 2009; China (Hubei). shirozui Okada, 1971; China (Taiwan). tentaculifera Chen & Chen, 2012; China (Hubei). unidentata Takada, 1968; Russia (Far East), China (Jilin, Liaoning, Taiwan), Korea, Japan (Hokkaido, Honshu, Kyushu). wuyishanensis Chen, Gao, & Chen, 2009; China (Jiangxi, Fujian, Yunnan). xanthosticta Chen, Gao, & Chen, 2009; China (Hubei). xishuangbanna Chen & Chen, 2012; China (Yunnan). zebromyia Zhang & Chen, 2021; China (Yunnan). zopheria Zhang & Chen, 2021; China (Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 105-106, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Nishiharu, S. (1979) Three new species of Drosophilidae from Japan (Diptera). Kontyu, Tokyo, 47, 38 - 43.","Chen, X. P. & Chen, H. W. (2012) Ten new species of the Stegana (Steganina) shirozui species group (Diptera, Drosophilidae) from China. Zootaxa, 3333 (1), 24 - 37. http: // dx. doi. org / 10.11646 / zootaxa. 3333.1.2","Okada, T. (1971) A revision and taxometric analysis of the genus Stegana Meigen of Japan and adjacent countries (Diptera, Drosophilidae). Mushi, 45, 81 - 99.","Takada, H. (1968) Drosophila survey of Hokkaido, XXVI. Descriptions of three new species of Drosophilidae from Japan. Journal of the Faculty of General Education, Sapporo University, 1, 119 - 127."]}

Stegana (Steganina) flabella Li & Chen, sp. nov. (Figs. 8C, 8D, 37) Diagnosis. This species closely resembles S. (S.) euryphylla Chen & Chen, 2009 in the shape of male terminalia (as figs. 14–20 in Chen & Chen 2009, Fig. 37), but can be distinguished from the latter by having the aedeagus distomedially and subdistolaterally lacking much pubescence (Fig. 37). Description. Male and female: First flagellomere black. Clypeus yellow. Mesonotum brown to dark brown medially, yellow to brownish laterally (Fig. 8C). Katepisternum yellowish (Fig. 8D). Scutellum dark brown (Fig. 8C). Halter mostly brownish (Fig. 8D). Fore femur with 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 8C, D); sternites brown, slightly wider than long. Male terminalia (Fig. 37): Epandrium with ca. 11 setae on dorsal to posterior portions per side (Fig. 37A). Surstylus with minute, serrated processes and numerous setulae on outer surface, 1 peg-like tooth near distal margin (Fig. 37A, B). Cercus lacking pubescence (Fig. 37A). Hypandrium expanded anteromedially (Fig. 37C, D). Pregonite small, with 3 sensilla (Fig. 37C, D). Aedeagus distally broadened and with numerous minute, serrated processes, with a row of serrated processes laterally and some pubescence distoventrally (Fig. 37C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 37E, F). Subepandrial sclerite rounded distally (Fig. 37G). Measurements and indices. BL = 2.37 mm in holotype (2.19–2.39 in 3♂, 2.34–2.41 in 3♀ paratypes), ThL = 1.09 mm (1.09–1.12 in ♂, 0.95–1.21 in ♀), WL = 2.12 mm (2.03–2.13 in ♂, 1.91–2.32 in ♀), WW = 0.95 mm (0.87–0.97 in ♂, 0.84–1.01 in ♀), arb = 6/4 (6–7/4–5), avd = 0.71 (0.71–0.86), adf = 1.46 (1.05–1.34), flw = 1.85 (1.67–2.05), FW/HW = 0.44 (0.38–0.42), ch/o = 0.32 (0.25–0.38), prorb = 0.83 (0.68–1.32), rcorb = 0.67 (0.59– 0.92), vb = 0.51 (0.50–0.71), dcl = 0.76 (0.53–0.81), presctl = 0.65 (0.55–0.78), sctl = 1.96 (1.61–2.13), sterno = 0.71 (0.55–1.39), orbito = 2.00 (1.50–2.08), dcp = 0.29 (0.27–0.42), sctlp = 1.39 (1.39–1.82), C = 1.99 (1.71–2.13), 4c = 1.25 (1.00–1.59), 4v = 2.25 (1.57–2.37), 5x = 1.34 (1.32–1.75), ac = 8.44 (8.87–12.29), M = 0.71 (0.59–0.96), C3F = 0.60 (0.47–0.61). Type materials. Holotype ♂ (SCAU, No. 125710, DNA #2659), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 670m, 25.iv.2017, ex tree trunk, H.W. Chen. Paratypes: 3♂, 3♀ (SCAU, Nos. 125711–16, DNA #2691), other data same as holotype. Distribution. China (Yunnan). Etymology. From the Latin word " flabellum " (= knife), referring to the distally broadened aedeagus.

Stegana (Steganina) izu Sidorenko, 1997 (Fig. 24) Stegana (Steganina) izu Sidorenko, 1997b: 167. Remarks. The male specimens from Ryukyu Islands, Japan slightly differ from the type materials from Izu Islands, Japan in the distal shape of aedeagus and aedeagal sheath (as figs. 4–7 in Sidorenko 1997b, Fig. 24C–F). They were conservatively identified as conspecific. Description. Female terminalia (Fig. 24H–J): Epiproct with several setae, lacking pubescence (Fig. 24H). Cercus with numerous setae and much pubescence (Fig. 24H). Eighth sternite about 1.5 times as wide as long, glabrous on basal 2/3, with some setae near posterior margin (Fig. 24I). Hypoproct with numerous setae and much pubescence (Fig. 24J). Specimens examined. CHINA: 9♀ (SCAU, Nos. 125427–35, DNA #1893, 1977–79), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 660m, 21–25.iv.2007, ex tree trunks, H.W. Chen, J.J. Gao; JAPAN: 2♂ (SCAU, Nos. 125806, 07, DNA #1872), Iriomote Island, Ryukyu Islands, 24°20'31''N, 123°49'20''E, alt. 330m, 14,18. vi.1999, ex tree trunks, H.W. Chen. Distribution. China (Taiwan, Yunnan*), Japan (Honshu, Ryukyu Islands*)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 51-52, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Sidorenko, V. S. (1997 b) New Asian species and new records of the genus Stegana Meigen (Diptera, Drosophilidae). II. Subgenera Pseudostegana Okada and Steganina Wheeler. Annales de la Societe entomologique de France, New Series, 33, 165 - 172."]}

I. Stegana (Steganina) biprotrusa species group (5 spp.) acantha Wu, Gao, & Chen, 2010; China (Hainan). ancistrophylla Wu, Gao, & Chen, 2010; China (Guangdong, Guangxi, Guizhou *, Yunnan *). langufoliacea Wu, Gao, & Chen, 2010; China (Taiwan, Guangxi, Yunnan*). otocondyloda Wu, Gao, & Chen, 2010; China (Yunnan). iaspidea Zhang & Chen, sp. nov.; China (Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 104, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen, 2009 Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen in Chen et al. 2009: 1914. Specimens examined. CHINA: 1♂ (SCAU, Nos. 125426, DNA #2656), Nonggang, Longzhou, Guangxi, 22°32'26''N, 106°43'18''E, alt. 230m, 23.viii.2004, ex tree trunk, H.W. Chen. Distribution. China (Hainan, Guangxi *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 49, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) zhuoma Cui & Chen, sp. nov. (Figs. 14G, 14H, 65D–F) Diagnosis. This species is similar to S. (S.) zaduo sp. nov. in the shape of female terminalia (Fig. 65), but can be distinguished from the latter by having the yellowish katepisternum brown on anterior corner (Fig. 14H); the 8th sternite nearly squared, with several setae and much pubescence on distolateral 1/3 (Fig. 65E). Description. Female: First flagellomere brown. Clypeus black. Mesonotum black medially, brown laterally (Fig. 14G). Scutellum black, yellow at tip (Fig. 14G). Halter mostly white (Fig. 14H). Fore femur with a row of 3 long, strong setae on distal part of posteroventral surface. All abdominal tergites dark brown to black (Fig. 14G, H); sternites brown, about 2 times as wide as long. Female terminalia (Fig. 65D–F): Epiproct with several setae, lacking pubescence (Fig. 65D). Cercus and hypoproct with numerous setae and much pubescence (Fig. 65D, F). Eighth sternite glabrous on basal 2/3 (Fig. 65E). Measurements and indices. BL = 2.70 mm in holotype, ThL = 1.45 mm, WL = 2.92 mm, WW = 1.26 mm, arb = 7/5, avd = 0.85, adf = 1.17, flw = 2.22, FW/HW = 0.39, ch/o = 0.25, prorb = 1.02, rcorb = 0.76, vb = 0.74, dcl = 0.48, presctl = 0.57, sctl = damaged, sterno = 0.77, orbito = 1.90, dcp = 0.32, sctlp = 1.45, C = 1.66, 4c = 1.35, 4v = 1.95, 5x = 1.45, ac = 12.89, M = 0.71, C3F = 0.63. Type material. Holotype ♀ (SCAU, No. 125789, DNA #2757), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt. 2000m, 11.x.2010, ex tree trunk, L. Wang. Distribution. China (Xizang). Etymology. From the word " zhuoma " in Zang language from the Zang nationality living in Xizang, China, referring to women., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 102, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) bimai Cui & Chen, sp. nov. (Figs. 6E, 6F, 29) Diagnosis. This species closely resembles S. (S.) clavispinula Chen & Chen, 2009 in the shape of epandrium, surstylus, cercus (as figs. 1, 2 in Chen & Chen 2009, Fig. 29A, B), and aedeagal sheath in ventral view (as fig. 6 in Chen & Chen 2009, Fig. 29E), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth near distal margin (Fig. 29B); the aedeagus expanded distally in ventral view (Fig. 29D), lacking serrated process in median process (Fig. 29C, D); the subepandrial sclerite slightly X-shaped (Fig. 29G). Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum brown medially, yellow laterally (Fig. 6E). Scutellum brown, yellow along margin (Fig. 6E). Katepisternum yellowish (Fig. 6F). Halter mostly grey (Fig. 6F). Fore femur with 3 thin, long setae on distal part of posteroventral surface. Abdominal tergites brown (Fig. 6E, F); sternites brownish, about 2 times as wide as long in male, about 3 times as wide as long in female. Male terminalia (Fig. 29): Epandrium with ca. 15 setae on dorsal to posterior portions per side (Fig. 29A). Surstylus slightly squared, with minute, serrated processes distally and several setulae near distal margin (Fig. 29A, B). Cercus lacking pubescence (Fig. 29A). Hypandrium expanded anteromedially (Fig. 29C, D). Pregonite small, with 3 sensilla (Fig. 29C, D). Aedeagus with some pubescence distoventrally (Fig. 29C). Aedeagal sheath with several minute, serrated processes distally (Fig. 29E, F). Measurements and indices. BL = 2.63 mm in holotype (2.52–2.61 in 2♂, 2.48–2.65 in 2♀ paratypes), ThL = 1.31 mm (1.31–1.34 in ♂, 1.26–1.28 in ♀), WL = 2.32 mm (2.28–2.51 in ♂, 2.31–2.61 in ♀), WW = 1.00 mm (0.97–1.01 in ♂, 0.95–1.08 in ♀), arb = 6/4 (7/4–5), avd = 0.65 (0.84–0.93), adf = 1.12 (1.08–1.20), flw = 2.12 (1.87–2.39), FW/HW = 0.40 (0.36–0.44), ch/o = 0.26 (0.24–0.27), prorb = 0.68 (0.87–1.07), rcorb = 0.71 (0.60– 0.71), vb = 0.60 (0.61–0.89), dcl = damaged (0.55–0.72), presctl = damaged (0.46–0.66), sctl = 2.19 (2.17–2.89), sterno = 0.39 (0.47–0.93), orbito = 1.80 (1.80–1.98), dcp = 0.30 (0.28–0.36), sctlp = 1.78 (1.50–1.73), C = 1.67 (1.72–1.98), 4c = 1.17 (1.22–1.28), 4v = 1.83 (1.88–2.19), 5x = 1.81 (1.44–1.86), ac = 15.17 (9.00–13.14), M = 0.74 (0.61–0.71), C3F = 0.55 (0.60–0.64). Type materials. Holotype ♂ (SCAU, No. 125744, DNA #2706), CHINA: Muyiji Park, Ximeng, Yunnan, 22°37'23''N, 99°36'11''E, alt. 1100m, 16.iv.2018, ex tree trunk, L. Zhu. Paratypes: 2♀ (SCAU, Nos. 125745, 46), other data same as holotype; 2♂ (SCAU, Nos. 125747, 48, DNA #2727), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 860m, 24.xii.2003, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. From the word " bimai " in Dai language from the Dai nationality living in Yunnan, China, referring to the Water-Sprinkling Festival (new year)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 57-59, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) hirticlavata Cui & Chen, sp. nov. (Figs. 9G, 9H, 44) Diagnosis. This species is similar to S. (S.) jianqinae Zhang, Tsaur, & Chen, 2014 in the shape of male terminalia (as fig. 8 in Zhang et al. 2014, Fig. 44), but can be distinguished from the latter by having the first flagellomere yellow on basal 2/3, black on distal 2/3 (Fig. 9H); the pregonite fused basally, with much long pubescence distally (Fig. 44C, D). Description. Male and female: Face mostly yellow, black near lower margin; facial carina absent. Clypeus yellow. Mesonotum yellow, with blurry longitudinal stripes (Fig. 9G). Katepisternum yellowish (Fig. 9H). Scutellum yellow, brown along margin in male (Fig. 9G). Halter mostly yellow (Fig. 9H). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites brownish (Fig. 9G, H); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 44): Epandrium with 4 setae near dorsal portion per side (Fig. 44A). Surstylus with numerous setulae on outer surface and some pubescence medially (Fig. 44A, B). Cercus with much pubescence (Fig. 44A). Hypandrium expanded anteromedially (Fig. 44C, D). Aedeagus expanded distally, with a pair of rod-like processes (Fig. 44C, D). Aedeagal sheath slightly saddle-shaped (Fig. 44E, F). Subepandrial sclerite slightly quadrate (Fig. 44G). Measurements and indices. BL = 2.64 mm in holotype (2.54 in 1♂, 2.52–2.88 in 5♀ paratypes), ThL = 1.33 mm (1.53 in ♂, 1.31–1.57 in ♀), WL = 2.65 mm (2.56 in ♂, 2.32–2.85 in ♀), WW = 1.28 mm (1.40 in ♂, 1.21–1.34 in ♀), arb = 7/5 (7–8/5–6), avd = 0.82 (0.74–0.80), adf = 1.31 (1.13–1.56), flw = 2.33 (2.17–2.72), FW/HW = 0.34 (0.34–0.39), ch/o = 0.17 (0.12–0.22), prorb = 0.88 (0.82–1.01), rcorb = 0.74 (0.63–0.77), vb = 0.82 (0.56–0.86), dcl = 0.44 (0.35–0.68), presctl = 0.45 (0.45–0.73), sctl = 1.84 (1.74–2.00), sterno = 0.74 (0.70–0.93), orbito = 2.00 (1.74–2.04), dcp = 0.14 (0.14–0.29), sctlp = 1.75 (1.31–1.98), C = 1.74 (1.83–1.95), 4c = 1.06 (0.97–1.10), 4v = 1.39 (1.28–1.55), 5x = 1.31 (1.17–1.27), ac = 13.00 (8.17–15.43), M = 0.52 (0.43–0.53), C3F = 0.70 (0.59–0.69). Type materials. Holotype ♂ (SCAU, No. 125571, DNA #1970), CHINA: Baihualing, Baoshan, Yunnan, 25°17'50''N, 98°48'12''E, alt. 1700m, 21.vi.2013, ex tussock, H.W. Chen. Paratypes: 4♀ (SCAU, Nos. 125572–75, DNA #2177), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt. 1320m, 6.v.2016, ex tree trunks, J. Huang, Y.Q. Liu, Y.L. Wang, L. Zhu; 1♀ (SCAU, No. 125577, DNA #1890), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 28.v.2015, ex tree trunk, H.W. Chen; 1♂ (SCAU, No. 125576, DNA #2181), Qimaba, Lüchun, Yunnan, 22°49'08''N, 102°17'25''E, alt. 1280m, 3.viii.2016, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. A combination of the Latin words '' hirtus '' (= hirsute) and '' clavatus '' (= rod-like), referring to the pregonite distally with long pubescence., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 75-76, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Zhang, Y., Tsaur, S. C. & Chen, H. W. (2014) Survey of the genus Stegana Meigen (Diptera, Drosophilidae) from Taiwan, with"]}

Stegana (Steganina) nigrolimbata Duda, 1924 Stegana (Steganina) nigrolimbata Duda, 1924: 181; Okada, 1971: 83; Cao & Chen, 2008: 29. Specimens examined. CHINA: 7♂, 4♀ (SCAU, Nos. 125307–17, DNA #1961, 1964), Maolan, Libo, Guizhou, 25°14'45''N, 107°57'31''E, alt. 750m, 19.ix.2015, ex tree trunks, J. Huang, Y.Q. Liu, Y.L. Wang, Y. Zhang. Distribution. China (Taiwan, Guangdong, Hainan, Guangxi, Guizhou*, Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 44, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Duda, O. (1924) Beitrag zur Systematik der Drosophiliden unter besonderer Berucksichtigung der palaarktischen u. orientalischen Arten (Dipteren). Archiv f ¸ r Naturgeschichte (A), 90, 172 - 234.","Okada, T. (1971) A revision and taxometric analysis of the genus Stegana Meigen of Japan and adjacent countries (Diptera, Drosophilidae). Mushi, 45, 81 - 99.","Chen, X. P. & Chen, H. W. (2008) The Stegana coleoptrata species group (Diptera: Drosophilidae) from mainland China. Zootaxa, 1891 (1), 55 - 65. https: // doi. org / 10.11646 / zootaxa. 1891.1.5"]}

Stegana (Steganina) bacilla Chen & Aotsuka, 2004 (Fig. 16) Stegana (Steganina) bacilla Chen & Aotsuka, 2004: 2785. Remarks. The specimens collected from Yunnan, China slightly differ from the type material from Ryukyu Islands, Japan (as fig. 9 in Chen & Aotsuka 2004) in the shape of distal portion of aedeagus and aedeagal sheath in ventral view (Fig. 16C, E). They were conservatively identified as conspecific. Specimens examined. CHINA: 1♂, 5♀ (SCAU, Nos. 125202–07, DNA #2730), Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 23–25.x.2017, ex tree trunks, H.W. Chen, L. Gong, B.X. Li; 9♂, 7♀ (SCAU, Nos. 125208–23, DNA #2636, 2661, 2627), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 580m, 5♂, 7♀ on 16,17. iv.2007, 4♂ on 19.xii.2017, H.W. Chen, J.J. Gao, L. Gong; 8♂ (SCAU, Nos. 125224–31, DNA #2639), Guanlei, Mengla, Yunnan, 21°38'39''N, 101°09'52''E, alt. 560m, 24.iv.2016, ex tree trunks, J. Huang, Y.Q. Liu; 3♂, 1♀ (SCAU, Nos. 125232–35, DNA #1344, 1886), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 680m, 2♂ on 27.ix.2011, 1♂, 1♀ on 30.ix.2011, ex tree trunks, H.W. Chen, J.J. Gao. Distribution. China (Taiwan, Yunnan*), Japan (Ryukyu Islands).

Stegana (Steganina) flabella Li & Chen, sp. nov. (Figs. 8C, 8D, 37) Diagnosis. This species closely resembles S. (S.) euryphylla Chen & Chen, 2009 in the shape of male terminalia (as figs. 14–20 in Chen & Chen 2009, Fig. 37), but can be distinguished from the latter by having the aedeagus distomedially and subdistolaterally lacking much pubescence (Fig. 37). Description. Male and female: First flagellomere black. Clypeus yellow. Mesonotum brown to dark brown medially, yellow to brownish laterally (Fig. 8C). Katepisternum yellowish (Fig. 8D). Scutellum dark brown (Fig. 8C). Halter mostly brownish (Fig. 8D). Fore femur with 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 8C, D); sternites brown, slightly wider than long. Male terminalia (Fig. 37): Epandrium with ca. 11 setae on dorsal to posterior portions per side (Fig. 37A). Surstylus with minute, serrated processes and numerous setulae on outer surface, 1 peg-like tooth near distal margin (Fig. 37A, B). Cercus lacking pubescence (Fig. 37A). Hypandrium expanded anteromedially (Fig. 37C, D). Pregonite small, with 3 sensilla (Fig. 37C, D). Aedeagus distally broadened and with numerous minute, serrated processes, with a row of serrated processes laterally and some pubescence distoventrally (Fig. 37C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 37E, F). Subepandrial sclerite rounded distally (Fig. 37G). Measurements and indices. BL = 2.37 mm in holotype (2.19–2.39 in 3♂, 2.34–2.41 in 3♀ paratypes), ThL = 1.09 mm (1.09–1.12 in ♂, 0.95–1.21 in ♀), WL = 2.12 mm (2.03–2.13 in ♂, 1.91–2.32 in ♀), WW = 0.95 mm (0.87–0.97 in ♂, 0.84–1.01 in ♀), arb = 6/4 (6–7/4–5), avd = 0.71 (0.71–0.86), adf = 1.46 (1.05–1.34), flw = 1.85 (1.67–2.05), FW/HW = 0.44 (0.38–0.42), ch/o = 0.32 (0.25–0.38), prorb = 0.83 (0.68–1.32), rcorb = 0.67 (0.59– 0.92), vb = 0.51 (0.50–0.71), dcl = 0.76 (0.53–0.81), presctl = 0.65 (0.55–0.78), sctl = 1.96 (1.61–2.13), sterno = 0.71 (0.55–1.39), orbito = 2.00 (1.50–2.08), dcp = 0.29 (0.27–0.42), sctlp = 1.39 (1.39–1.82), C = 1.99 (1.71–2.13), 4c = 1.25 (1.00–1.59), 4v = 2.25 (1.57–2.37), 5x = 1.34 (1.32–1.75), ac = 8.44 (8.87–12.29), M = 0.71 (0.59–0.96), C3F = 0.60 (0.47–0.61). Type materials. Holotype ♂ (SCAU, No. 125710, DNA #2659), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 670m, 25.iv.2017, ex tree trunk, H.W. Chen. Paratypes: 3♂, 3♀ (SCAU, Nos. 125711–16, DNA #2691), other data same as holotype. Distribution. China (Yunnan). Etymology. From the Latin word " flabellum " (= knife), referring to the distally broadened aedeagus., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 68-69, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) tongi Wang, Gao, & Chen, 2011 Stegana (Steganina) tongi Wang, Gao, & Chen in Wang et al. 2011: 510. Specimens examined. CHINA: 1♂, 2♀, Yixiang, Pu’er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m: 1♂ (SCAU, No. 125237, DNA #2601) on 2.x.2011, ex tree trunk, H.W. Chen, 2♀ (SCAU, Nos. 125238, 39, DNA #2192) on 28–30.iii.2013, ex tussocks, J.J. Gao; 1♂, 1♀ (SCAU, Nos. 125240, 41, DNA #2633), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 560m, 19.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Taiwan, Guangdong, Hainan, Guangxi, Yunnan*)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 41, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, J. Q., Gao, J. J. & Chen, H. W. (2011) Stegana castanea species group (Diptera, Drosophilidae) from the Oriental region. Journal of Natural History, 45, 505 - 519. https: // doi. org / 10.1080 / 00222933.2010.534239"]}

Stegana (Steganina) lancang Li & Chen, sp. nov. (Figs. 10E, 10F, 46) Diagnosis. This species is similar to S. (S.) kanmiyai Okada & Sidorenko, 1992 in the shape of male terminalia (as fig. 3E–G in Okada & Sidorenko 1992, Fig. 46A–G), but can be distinguished from the latter by having the brownish yellow body color (Fig. 10E, F); the surstylus with 1 peg-like tooth near distal margin (Fig. 46A, B); the aedeagus lacking pubescence distolaterally (Fig. 46C, D). Description. Male: Clypeus and palpus yellow. Mesonotum brownish yellow (Fig. 10E). Katepisternum yellowish (Fig. 10F). Scutellum brownish yellow, brownish near distal margin (Fig. 10E). Halter mostly grey (Fig. 10F). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites brownish yellow (Fig. 10E, F); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 46A–G): Epandrium with 13 setae on dorsal to posterior portions per side (Fig. 46A). Surstylus with numerous setulae distally and some minute, serrated processes distoventrally (Fig. 46C, D). Aedeagal sheath acute apically, with 1 pair of acute processes subapically (Fig. 46E, F). Subepandrial sclerite vaulted (Fig. 46G). Female terminalia (Fig. 46H–J): Epiproct with several setae on distal 1/3 (Fig. 46H). Cercus and hypoproct with numerous setae and much pubescence (Fig. 46H, J). Eight sternite about 2 times as long as wide, mostly glabrous except with some setae distally (Fig. 46I). Measurements and indices. BL = 2.74 mm in holotype (2.33–2.67 in 5♂, 2.39–2.73 in 4♀ paratypes), ThL = 1.32 mm (1.21–1.31 in ♂, 1.21–1.37 in ♀), WL = 2.48 mm (2.16–2.43 in ♂, 2.27–2.63 in ♀), WW = 0.99 mm (0.95–1.02 in ♂, 0.95–1.05 in ♀), arb = 6/4 (6–8/4–5), avd = 0.88 (0.74–0.89), adf = 1.05 (1.00–1.74), flw = 1.81 (1.56–2.15), FW/HW = 0.47 (0.34–0.43), ch/o = 0.27 (0.22–0.30), prorb = 0.93 (0.77–1.53), rcorb = 0.60 (0.62–0.91), vb = 0.52 (0.46–0.83), dcl = 0.87 (0.59–0.87), presctl = 0.52 (0.60–0.84), sctl = damaged (1.41–1.94), sterno = 0.52 (0.63–1.08), orbito = 1.80 (1.59–1.96), dcp = 0.30 (0.24–0.35), sctlp = 1.67 (1.27–2.19), C = 1.76 (1.65–2.11), 4c = 1.26 (0.96–1.30), 4v = 1.86 (1.44–1.84), 5x = 1.59 (1.26–1.76), ac = 10.67 (8.17–11.00), M = 0.67 (0.55–0.72), C3F = 0.68 (0.57–0.66). Type materials. Holotype ♂ (SCAU, No. 125697, DNA #2637), CHINA: Guanlei, Mengla, Yunnan, 21°38'39''N, 101°09'52''E, alt. 560m, 24.iv.2016, ex tree trunk, J. Huang. Paratypes: 4♀ (SCAU, Nos. 125698–701, DNA #2637), other data same as holotype; 8♂ (SCAU, Nos. 125702–09, DNA #2638), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 560m, 17.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Yunnan). Etymology. From the word " lancang " in Dai language from the Dai nationality living in Yunnan, China, referring to the Lancang River., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 79-80, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Okada, T. & Sidorenko, V. S. (1992) New or unrecorded species of Stegana (Steganina) (Diptera, Drosophilidae) from the Old World, especially Japan and Russia. Japanese Journal of Entomology, 60, 415 - 426."]}

Stegana (Steganina) kanmiyai Okada & Sidorenko, 1992 (Fig. 25) Stegana (Steganina) kanmiyai Okada & Sidorenko, 1992: 419. Diagnosis. This species is similar to S. (S.) lancang sp. nov. in the shape of male terminalia (Figs. 25, 46A–G), but can be distinguished from the latter by having the cercus lacking pubescence (Fig. 25A); the aedeagus with numerous minute, serrated processes along anterior margin and pubescence anteromedially (Fig. 25C, D). Specimens examined. CHINA: 2♂ (SCAU, Nos. 125436, 37, DNA #785), Mangshan, Yizhang, Hunan, 24°56'53''N, 112°54'01''E, alt. 1260m, 3.x.2004, ex tree trunk, M.F. Xu; 1♂ (SCAU, No. 125438, DNA #2602), Chebaling, Shixing, Guangdong, 24°43'22''N, 114°31'49''E, alt. 780m, 11.ix.2017, ex tussock, L. Gong; 5♂, 11♀ (SCAU, Nos. 125439–54, DNA #786, 1046, 1184, 2759, 2760), Conghua, Guangdong, 23°43'30''N, 113°48'20''E, alt. 500m, 3♂, 6♀ on 3.xi.2003, 2♂, 5♀ on 10.x.2005, ex tree trunks, H.W. Chen; 1♂, 4♀ (SCAU, Nos. 125455– 59, DNA #457, 2709), Dinghushan, Zhaoqing, Guangdong, 23°10'17''N, 112°32'31''E, alt. 580–670m, ex tree trunks, 5,6. vii.2005, J.J. Jiang, MF Xu; 1♀ (SCAU, No. 125460, DNA #2185), Tianluhu, Guangzhou, Guangdong, 23°13'43''N, 113°25'34''E, alt. 360m, 6.ix.2015, ex tree trunk, J. Huang; 4♂, 7♀ (SCAU, Nos. 125461–71, DNA #1193), Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 23.x.2004, ex tree trunks and tussocks, H.L. Cao, H.W. Chen, MF Xu; 1♀ (SCAU, No. 125472, DNA #1962), Mulun, Huanjiang, Guangxi, 25°07'13''N, 108°02'34''E, alt. 500m, 16.ix.2015, ex tree trunk, J. Huang; 3♂, 2♀ (SCAU, Nos. 125473–77, DNA #2621), Guposhan, Hezhou, Guangxi, 24°36'10''N, 111°35'18''E, alt. 670m, 29.vii.2017, ex tree trunks, H.W. Chen; 1♂, 1♀ (SCAU, Nos. 125478, 79, DNA #2610), Shengtangshan, Jinxiu, Guangxi, 23°58'23''N, 110°06'45''E, alt. 1300m, 31.vii.2017, ex tree trunks, H.W. Chen; 1♀ (SCAU, No. 125480, DNA #1466), Bapen, Fusui, Guangxi, 22°32'36'' N, 107°53'18''E, alt. 210m, 18.viii.2004, ex tree trunk, H.W. Chen; 1♂, 5♀ (SCAU, Nos. 125481–86, DNA #796), Emeishan, Leshan, Sichuan, 29°36'14''N, 103°28'56''E, alt. 700m, 5.vii.2007, ex tussocks, H.Z. Cao, J.J. Gao, J.J. Jiang; 1♀ (SCAU, No. 125487, DNA #1896), Baihualing, Baoshan, Yunnan, 25°17'50''N, 98°48'12''E, alt. 1420m, 25.viii.2013, ex tussock, Q.S. Gao; 1♂, 1♀ (SCAU, Nos. 125488, 89, DNA #2351), Menglai, Cangyuan, Yunnan, 23°13'20''N, 99°15'04''E, alt. 1240m, 30.x.2017, ex tree trunks, H.W. Chen, L. Gong; 1♂, 2♀ (SCAU, Nos. 125490–92, DNA #1444), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 13.v.2012, ex tree trunks, H.W. Chen; 1♂, 1♀ (SCAU, Nos. 125493, 94, DNA #2664), Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 13,14. iv.2018, ex tree trunks, H.W. Chen. Distribution. China (Hunan *, Taiwan, Guangdong*, Guangxi*, Sichuan*, Yunnan*), Japan (Honshu)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 52-54, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Okada, T. & Sidorenko, V. S. (1992) New or unrecorded species of Stegana (Steganina) (Diptera, Drosophilidae) from the Old World, especially Japan and Russia. Japanese Journal of Entomology, 60, 415 - 426."]}

Stegana (Steganina) cordhirsuta Wang & Chen, sp. nov. (Figs. 6I, 6J, 31) Diagnosis. This species resembles S. (S.) sphaerica Wang, Gao, & Chen, 2013 in the shape of aedeagal sheath (as fig. 10E, F in Wang et al. 2013, Fig. 31C), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth near distal margin (Fig. 31A); the subepandrial sclerite slightly vaulted, deeply split medially and broadened laterally (Fig. 31D). Description. Male and female: First flagellomere yellow on basal 1/3, black on distal 2/3. Clypeus brown medially, yellow laterally. Mesonotum mostly brownish, with 3 indistinct, longitudinal yellow stripes medially and sublaterally (Fig. 6I). Katepisternum mostly yellowish, brown on posterior corner (Fig. 6J). Scutellum mostly brownish, brown anteriorly and black along posterior margin (Fig. 6I). Halter mostly grey (Fig. 6J). Fore femur with a row of 5 or 6 long setae on distal part of posteroventral surface. Abdominal tergites mostly yellow, dark brown near distal margin, lacking pubescence (Fig. 31A). Cercus with much pubescence (Fig. 31A). Hypandrium roundly expanded anteromedially (Fig. 31B). Pregonite large, lacking sensillum (Fig. 31B, C). Aedeagus rounded distally, with much pubescence distoventrally (Fig. 31B, C). Aedeagal sheath distally dorsad curved and with some minute, serrated processes (Fig. 31C). Measurements and indices. BL = 3.40 mm in holotype (2.97–3.27 in 5♂, 3.29 in 1♀ paratypes), ThL = 1.50 mm (1.37–1.70 in ♂, 1.80 in ♀), WL = 2.50 mm (1.90–2.50 in ♂, 3.07 in ♀), WW = 1.00 mm (1.03–1.20 in ♂, 1.10 in ♀), arb = 11/5 (11–13/4–6), avd = 0.90 (0.67–1.11), adf = 2.00 (1.50–3.00), flw = 2.40 (2.00–2.50), FW/HW = 0.32 (0.34–0.45), ch/o = 0.22 (0.14–0.29), prorb = 0.87 (1.46–1.88), rcorb = 0.40 (0.54–0.88), vb = 0.50 (0.33–0.67), dcl = 0.26 (0.32–0.50), presctl = 0.58 (0.53–0.67), sctl = 2.14 (1.73–2.50), sterno = 0.60 (0.56–0.71), orbito = 1.43 (1.67–2.50), dcp = 0.21 (0.18–0.23), sctlp = 2.00 (1.43–2.22), C = 2.03 (1.73–2.41), 4c = 1.25 (1.04–1.38), 4v = 2.06 (1.69–2.20), 5x = 1.57 (1.29–2.40), ac = 20.00 (14.50–22.00), M = 0.69 (0.51–0.75), C3F = 0.70 (0.68–0.80). Type materials. Holotype ♂ (SCAU, No. 121099, DNA #1042), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 660m, 21.iv.2007, ex tree trunk, H.W. Chen. Paratypes: 15♂, 1♀ (SCAU, 121100– 08, 125579–85, DNA #1345, 1411), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1420m, 14♂, 1♀ on 30.ix.2011, 1♂ on 9.v.2012, ex tree trunks, H.W. Chen, J.J. Gao. Distribution. China (Yunnan). Etymology. A combination of the Latin words " cordatus " (= cord) and '' hirsutus '' (= hirsute), referring to the aedeagus medially with pubescence.

Stegana (Steganina) baoxing Li & Chen, sp. nov. (Figs. 6C, 6D, 28) Diagnosis. This species is similar to S. (S.) alafoliacea sp. nov. in the dark body color (Fig. 6A–D), but can be distinguished from the latter by having the surstylus lacking serrated process (Fig. 28A); the aedeagus rounded distally in ventral view (Fig. 28B); the aedeagal sheath lacking serrated process (Fig. 28D, E). Description. Male: First flagellomere brown. Clypeus dark brown medially, yellow laterally. Mesonotum and scutellum dark brown to black (Fig. 6C, D). Katepisternum brownish (Fig. 6D). Halter mostly grey (Fig. 6C). Fore femur with 3 thin, long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 6C, D); sternites brown, about 2 times as wide as long. Male terminalia (Fig. 28): Epandrium with ca. 8 setae on dorsal to posterior portions per side (Fig. 28A). Surstylus slightly curved subdistally, with 1 peg-like tooth and several setulae near distal margin (Fig. 28A). Cercus with much pubescence (Fig. 28A). Hypandrium expanded anteromedially (Fig. 28B, C). Pregonite small, lacking sensillum (Fig. 28B, C). Aedeagus slightly expanded distally, with numerous minute, serrated processes distolaterally and pubescence medially (Fig. 28B, C). Subepandrial sclerite slightly split distally (Fig. 28F). Measurements and indices. BL = 3.16 mm in holotype, ThL = 1.45 mm, WL = 2.99 mm, WW = 1.04 mm, arb = 5/4, avd = 0.80, adf = 1.14, flw = 1.81, FW/HW = 0.34, ch/o = 0.25, prorb = 1.21, rcorb = 0.77, vb = 0.78, dcl = 0.42, presctl = 0.50, sctl = 3.16, sterno = 0.79, orbito = 1.80, dcp = 0.37, sctlp = 1.56, C = 2.07, 4c = 1.43, 4v = 2.65, 5x = 2.22, ac = 12.11, M = 0.93, C3F = 0.59. Type material. Holotype ♂ (SCAU, No. 125693, DNA #2607), CHINA: Fengtongzhai, Baoxing, Sichuan, 30°32'42''N, 102°54'24''E, alt. 1500m, 18.vii.2017, ex tree trunk, L. Gong. Distribution. China (Sichuan). Etymology. From the name of the type locality "Baoxing".

Stegana castanea species group Stegana (Steganina) castanea species group Wang, Gao, & Chen in Wang et al. 2011: 506. Diagnosis. Pregonite rod-shaped distinctly, basally fused to or connected with hypandrium (e.g., Figs. 16C, 16D, 17C, 17D), Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 40, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, J. Q., Gao, J. J. & Chen, H. W. (2011) Stegana castanea species group (Diptera, Drosophilidae) from the Oriental region. Journal of Natural History, 45, 505 - 519. https: // doi. org / 10.1080 / 00222933.2010.534239"]}

Stegana (Steganina) angulistrata Wu & Chen, 2010 Stegana (Steganina) angulistrata Wu & Chen, 2010: 69. Specimens examined. CHINA: 1♂ (No. 125500, DNA #1274), Huanggangshan, Qianshan, Jiangxi, 28°00'43''N, 117°44'59''E, alt. 1700m, 19.vi.2004, ex tree trunk, H.W. Chen; 1♂ (SCAU, No. 125501, DNA #812), Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 23.x.2004, ex tree trunk, H.W. Chen; 7♂, 10♀ (SCAU, Nos. 125502–18, DNA #2611, 2615), Shengtangshan, Jinxiu, Guangxi, 23°58'23''N, 110°06'45''E, alt. 1300–1500m, 2♂, 6♀ on 30.vii.2017, 5♂, 4♀ on 31.vii.2017, ex tree trunks, H.W. Chen. Distribution. China (Jiangxi *, Guangxi, Yunnan).

Stegana (Steganina) dendrophila Zhang & Chen, sp. nov. (Figs. 8A, 8B, 36) Diagnosis. This species resembles S. (S.) cyclophylla Chen & Chen, 2009 in the shape of hypandrium, pregonite, and aedeagus (as figs. 10, 11 in Chen & Chen 2009, Fig. 36B, C), but can be distinguished from the latter by having the surstylus lacking peg-like tooth (Fig. 36A); the aedeagus with numerous minute, serrated processes distally, lacking pubescence (Fig. 36B, C); the subepandrial sclerite slightly squared, concaved anteromedially (Fig. 36D). Description. Male: First flagellomere yellow on basal 1/3, black on distal 2/3. Clypeus brown medially, yellow laterally. Mesonotum and scutellum black (Fig. 8A). Katepisternum yellowish, brown on anterior corner (Fig. 8B). Halter mostly brownish (Fig. 8B). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites dark brown (Fig. 8A, B); sternites brown, slightly wider than long. Male terminalia (Fig. 36): Epandrium with ca. 20 setae on dorsal to posterior portions per side (Fig. 36A). Surstylus with several setulae and some pubescence distally (Fig. 36A). Cercus with much pubescence (Fig. 36A). Hypandrium expanded anteromedially (Fig. 36B, C). Pregonite fused with hypandrium and small, with 4 sensilla (Fig. 36B, C). Aedeagal sheath with some numerous minute, serrated processes distally (Fig. 36B, C). Measurements and indices. BL = 3.20 mm in holotype, ThL = 1.30 mm, WL = 1.93 mm, WW = 1.07 mm, arb = 9/6, avd = 0.83, adf = 1.71, flw = 1.29, FW/HW = 0.46, ch/o = 0.14, prorb = 1.45, rcorb = 1.00, vb = 0.33, dcl = 0.66, presctl = 0.70, sctl = 2.77, sterno = 1.00, orbito = 2.50, dcp = 0.19, sctlp = 2.50, C = 1.84, 4c = 1.23, 4v = 1.80, 5x = 1.33, ac = 9.25, M = 0.53, C3F = 0.62. Type material. Holotype ♂ (SCAU, No. 125633, DNA #1338, 2770), CHINA: Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 680m, 28.ix.2011, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. From the Greek word '' dendrophilus '' (= dendrophilous), referring to the arboreal Steganina flies., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 66-67, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) nigridentata Li & Chen, sp. nov. (Figs. 12E, 12F, 55) Diagnosis. This species is similar to S. (S.) menghai sp. nov. in the shape of male terminalia (Figs. 50, 55), but can be distinguished from the latter by having the pregonite blurry or absent (Fig. 50B, C); the aedeagus laterally deeply dehiscent and with 4 small black teeth per side, distally lacking serrated process (Fig. 55B, C). Description. Male and female: First flagellomere dark brown. Clypeus dark brown medially, yellow laterally. Mesonotum brown to black medially, brownish yellow to brown laterally (Fig. 12E). Katepisternum yellow above, brownish below (Fig. 12F). Scutellum dark brown to black (Fig. 12E). Halter mostly grey (Fig. 12F). Fore femur with a row of 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites mostly black (Fig. 12E, margin, lacking pubescence (Fig. 55A). Cercus with much pubescence (Fig. 55A). Hypandrium expanded anteromedially (Fig. 55B, C). Aedeagal sheath distally dorsad curved and with numerous minute, serrated processes (Fig. 55D, E). Subepandrial sclerite vaulted (Fig. 55F). Measurements and indices. BL = 3.28 mm in holotype (2.88–3.52 in 5♂, 3.24–4.20 in 5♀ paratypes), ThL = 1.44 mm (1.43–1.80 in ♂, 1.76–2.05 in ♀), WL = 2.44 mm (2.36–3.00 in ♂, 2.72–3.24 in ♀), WW = 1.20 mm (1.10–1.40 in ♂, 1.208–1.60 in ♀), arb = 14/5 (8–14/4–6), avd = 0.60 (0.44–0.91), adf = 1.43 (1.57–2.25), flw = 1.57 (1.50–2.60), FW/HW = 0.38 (0.27–0.38), ch/o = 0.22 (0.13–0.27), prorb = 1.25 (0.75–1.14), rcorb = 0.92 (0.58–1.17), vb = 0.75 (0.29–0.67), dcl = 0.75 (0.27–0.75), presctl = 0.63 (0.50–0.93), sctl = 1.50 (1.24–1.93), sterno = 0.92 (0.59–0.82), orbito = 2.60 (1.67–2.50), dcp = 0.21 (0.18–0.26), sctlp = 1.73 (1.50–2.27), C = 2.62 (1.73–2.25), 4c = 0.81 (1.09–1.26), 4v = 1.63 (1.63–2.19), 5x = 1.60 (1.36–2.00), ac = 13.00 (10.00–21.25), M = 0.50 (0.50–0.81), C3F = 0.57 (0.69–0.82). Type materials. Holotype ♂ (SCAU, No. 121132), CHINA: Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 23.x.2004, ex tree trunk, H.W. Chen. Paratypes: 6♂, 18♀ (SCAU, Nos. 121133–56, DNA #1461, 1462), 19.x.2006, other data same as holotype; 2♀ (SCAU, Nos. 125606, 07, DNA #2190), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 13.v.2012, ex tree trunks, J.J. Gao. Distribution. China (Guangxi, Yunnan). Etymology. A combination of the Latin words " niger " (= black) and " dentatus " (= dentate), referring to the aedeagus laterally with 4 small black teeth., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 90-91, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) mailangang Li & Chen, sp. nov. (Figs. 11C, 11D, 49) Diagnosis. This species slightly resembles S. (S.) cyclophylla Chen & Chen, 2009 in the shape of hypandrium, pregonite, and subepandrial sclerite (as figs. 9–11 in Chen & Chen 2009, Fig. 49C, D, G), but can be distinguished from the latter by having the aedeagus distally with some minute, serrated processes (Fig. 49C, D); the aedeagal sheath lacking serrated process (Fig. 49E, F). Description. Male: First flagellomere black. Clypeus yellow. Mesonotum black medially, dark brown laterally (Fig. 11C). Katepisternum yellowish (Fig. 11D). Scutellum black (Fig. 11C). Halter mostly black (Fig. 11D). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites black (Fig. 11C, D); sternites brown, about 2 times as wide as long. Male terminalia (Fig. 49): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 49A). Surstylus with 1 peg-like tooth near distal margin and some setulae distally, lacking pubescence (Fig. 49A, B). Cercus with much pubescence (Fig. 49A). Hypandrium roundly expanded anteromedially (Fig. 49C, D). Pregonite fused with hypandrium, with 3 sensilla (Fig. 49C, D). Aedeagus roundly expanded distally, with some pubescence distoventrally (Fig. 49C, D). Subepandrial sclerite vaulted, expanded dorsally (Fig. 49G). Measurements and indices. BL = 2.77 mm in holotype, ThL = 1.33 mm, WL = 2.75 mm, WW = 1.32 mm, arb = 7/5, avd = 0.68, adf = 1.35, flw = 1.78, FW/HW = 0.34, ch/o = 0.23, prorb = 1.21, rcorb = 0.97, vb = 1.45, dcl = 0.59, presctl = 0.49, sctl = 1.55, sterno = 0.76, orbito = 1.78, dcp = 0.30, sctlp = 1.69, C = 2.18, 4c = 1.14, 4v = 1.95, 5x = 1.65, ac = 8.09, M = 0.72, C3F = 0.69. Type material. Holotype ♂ (SCAU, No. 125694, DNA #2623), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 660m, 17.xii.2017, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. From the word " mailangang " in Dai language from the Dai nationality living in Yunnan, China, referring to the meaning of the type locality (Wangtianshu: sky tree)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 83-84, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) masanoritodai Okada & Sidorenko, 1992 (Figs. 5G, 5H, 22) Stegana (Steganina) masanoritodai Okada & Sidorenko, 1992: 415. Remarks. The specimens collected from Liaoning, China slightly differ from those from Kumamoto, Japan (as figs. 6, 7 in Chen et al. 2009) in the shape of aedeagal sheath (Fig. 22D, E); the aedeagal sheath lacking serrated, minute process (Fig. 22D, E); the subepandrial sclerite narrowed distally (Fig. 22F). They were conservatively identified as conspecific. Specimens examined. CHINA: 2♂ (SCAU, Nos. 125416, 17, DNA #2489), Guojiabaozi, Kuandian, Liaoning, 40°48'50''N, 124°45'34''E, alt. 340m, 26.vii.2018, ex tree trunks, L. Gong; 1♂ (SCAU, No. 125418, DNA #2499), Laobiangou, Benxi, Liaoning, 41°05'32''N, 124°30'07''E, alt. 530m, 21.vii.2018, ex tussock, B.X. Li. Distribution. China (Jilin, Liaoning *), Russia (Far East), Japan (Hokkaido, Honshu, Kyushu).

Stegana (Steganina) tongi Wang, Gao, & Chen, 2011 Stegana (Steganina) tongi Wang, Gao, & Chen in Wang et al. 2011: 510. Specimens examined. CHINA: 1♂, 2♀, Yixiang, Pu’er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m: 1♂ (SCAU, No. 125237, DNA #2601) on 2.x.2011, ex tree trunk, H.W. Chen, 2♀ (SCAU, Nos. 125238, 39, DNA #2192) on 28–30.iii.2013, ex tussocks, J.J. Gao; 1♂, 1♀ (SCAU, Nos. 125240, 41, DNA #2633), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 560m, 19.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Taiwan, Guangdong, Hainan, Guangxi, Yunnan*).

VII. Stegana (Steganina) undulata species group (8 spp.) biflava Li, Zhang, & Chen, 2019; China (Guangxi, Guizhou, Yunnan). flaviclypeata Chen & Chen, 2011; China (Sichuan, Yunnan). flavipalpata Chen & Chen, 2011; China (Yunnan). flavivittata Li, Zhang, & Chen, 2019; China (Yunnan). leucothorax Chen & Chen, 2011; China (Yunnan). melanocheilota Chen & Chen, 2011; China (Yunnan). melanothorax Chen & Chen, 2011; China (Yunnan). undulata de Meijere, 1911; China (Hainan, Guangxi, Yunnan), Myanmar (Yangon), Indonesia (Java, Sumatra, Kalimantan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 106, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["de Meijere, J. C. H. (1911) Studien uber sudostasiatische Dipteren. VI. Tijdschrift voor Entomologie, 54, 258 - 432."]}

Stegana (Steganina) nigripalpula Cui & Chen, sp. nov. (Figs. 5C, 5D, 20G–I) Diagnosis. This species differs from the other species of the ornatipes species group in having the large and black palpus (Fig. 5D); the first flagellomere entirely black. Description. Female: Frons entirely black. Face black, with 1 yellow band medially; facial carina absent. Clypeus black. Mesonotum and scutellum dark brown (Fig. 5C, D). Katepisternum dark brown above, yellowish to yellow below (Fig. 5D). Halter mostly black. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites black medially, dark brown laterally (Fig. 5C, D); sternites brown, about 3 times as wide as long. Female terminalia (Fig. 20G–I): Epiproct and cercus lacking pubescence (Fig. 20G). Eighth sternite about 2 times as wide as long, glabrous on basal 2/3 portion, with numerous prensisetae near posterior margin (Fig. 20H). Hypoproct with numerous setae and much pubescence (Fig. 20I). Measurements and indices. BL = 2.65 mm in holotype, ThL = 1.33 mm, WL = 2.61 mm, WW = 1.09 mm, arb = 8/6, avd = 0.73, adf = 1.56, flw = 2.83, FW/HW = 0.35, ch/o = 0.11, prorb = 0.95, rcorb = 0.74, vb = 0.57, dcl = 0.59, presctl = 0.67, sctl = damaged, sterno = 0.62, orbito = 1.62, dcp = 0.27, sctlp = 1.46, C = 1.79, 4c = 1.14, 4v = 1.45, 5x = 1.11, ac = 8.00, M = 0.61, C3F = 0.62. Type material. Holotype ♀ (SCAU, No. 125416, DNA #1876, 2787), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 570m, 29.ix.2011, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. A combination of the Latin words " niger " (= black) and " palpulus '' (= palpus), referring to the black palpus., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 48, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) bibarbata Li & Chen, sp. nov. (Figs. 4G, 4H, 19) Diagnosis. This species differs from the other species of the nigrolimbata species group in having the yellow body color (Fig. 4G, H); the surstylus slightly triangular, lacking scale-like process (Fig. 19A, B); the aedeagal sheath not expanded distolaterally, with some minute, serrated processes (Fig. 19E, F). Description. Male: Frons yellowish brown, lacking black band. First flagellomere yellow. Face black, with 1 yellow band medially. Clypeus black medially, yellow laterally. Palpus yellow basally, dark brown distally. Gena narrow and yellow. Mesonotum and scutellum mostly yellow (Fig. 4G). Katepisternum yellowish above, brownish below (Fig. 4H). Halter brownish basally, grey distally (Fig. 4H). Fore femur with 4 thin, long setae on distal part of posteroventral surface. Abdominal tergites yellow (Fig. 4G, H); sternites brownish, slightly wider than long. Male terminalia (Fig. 19): Epandrium with ca. 28 setae on dorsal to posterolateral portions per side (Fig. 19A). Surstylus with numerous setae near margin, lacking pubescence (Fig. 19A, B). Cercus lacking pubescence (Fig. 19A). Hypandrium expanded anteromedially (Fig. 19C, D). Pregonite blurry or absent (Fig. 19C, D). Aedeagus with much pubescence basally (Fig. 19C, D). Phallapodeme (aedeagal apodeme in McAlpine 1981) podgy (Fig. 19C, D). Measurements and indices. BL = 2.93 mm in holotype, ThL = 1.46 mm, WL = 2.64 mm, WW = 1.28 mm, arb = 6/3, avd = 0.67, adf = 1.31, flw = 1.95, FW/HW = 0.36, ch/o = 0.10, prorb = 0.32, rcorb = 0.75, vb = 0.49, dcl = 0.61, presctl = 0.50, sctl = 1.17, sterno = 0.71, orbito = 1.58, dcp = 0.29, sctlp = 1.50, C = 1.78, 4c = 1.15, 4v = 1.55, 5x = 0.95, ac = 10.20, M = 0.44, C3F = 0.65. Type material. Holotype ♂ (SCAU, No. 125407, DNA #2460), CHINA: Husa, Longchuan, Yunnan, 24°28'21''N, 97°54'06''E, alt. 1340m, ex tree trunk, 7.xi.2017, H.W. Chen. Distribution. China (Yunnan). Etymology. A combination of the Latin words " bi -" (= two) and " barbatus " (= beard), referring to the pair of long, filar processes of aedeagus.

Stegana (Steganina) latigena Wang, Gao, & Chen, 2013 Stegana (Steganina) latigena Wang, Gao, & Chen in Wang et al. 2013: 2003. Specimens examined. CHINA: 1♂ (SCAU, No. 125529, DNA #1947), Maolan, Libo, Guizhou, 25°14'45''N, 107°57'31''E, alt. 750m, 19.ix.2015, ex tussock, Y.Q. Liu. Distribution. China (Guangdong, Guangxi, Guizhou *, Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 54, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, L., Gao, J. J. & Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055"]}

Stegana (Steganina) menglian Li & Chen, sp. nov. (Figs. 11G, 11H, 51) Diagnosis. This species is similar to S. (S.) lancang sp. nov. in the shape of male terminalia (Figs. 46A–G, 51), but can be distinguished from the latter by having the surstylus lacking serrated process (Fig. 51A, B); the aedeagus lacking a row of serrated processes distoventrally (Fig. 51C, D). Description. Male and female: First flagellomere yellowish brown. Clypeus yellow. Mesonotum dark brown medially, brown laterally (Fig. 11G). Katepisternum mostly yellowish, brownish on anterior corner (Fig. 11H). Scutellum dark brown, yellow at tip (Fig. 11G). Halter mostly white (Fig. 11H). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites black (Fig. 11G, H); sternites brown, about 2 times as wide as long. Male terminalia (Fig. 51): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 51A). Surstylus with 1 peg-like tooth and some setulae near distal margin, lacking pubescence (Fig. 51A, B). Cercus with much pubescence (Fig. 51A). Hypandrium roundly expanded anteromedially (Fig. 51C, D). Pregonite with 2 sensilla (Fig. 51C, D). Aedeagus roundly expanded distally, with several minute, serrated processes distolaterally and some pubescence distoventrally (Fig. 51C, D). Aedeagal sheath distally strongly dorsad curved and with several minute, serrated processes (Fig. 51E, F). Subepandrial sclerite nearly vaulted (Fig. 51G). Measurements and indices. BL = 2.21 mm in holotype (2.43 in 1♂, 2.73–2.79 in 2♀ paratypes), ThL = 1.08 mm (1.21 in ♂, 1.32–1.36 in ♀), WL = 2.09 mm (2.12 in ♂, 2.46–2.52 in ♀), WW = 0.88 mm (0.91 in ♂, 0.99–1.04 in ♀), arb = 7/4 (5–6/3–4), avd = 0.84 (0.77–0.88), adf = 0.97 (1.11–1.26), flw = 1.77 (1.73–1.93), FW/HW = 0.40 (0.41–0.45), ch/o = 0.30 (0.25–0.27), prorb = 1.18 (1.04–1.20), rcorb = 0.78 (0.57–0.71), vb = 0.71 (0.71–0.72), dcl = 0.74 (0.53–0.80), presctl = 0.67 (0.47–0.88), sctl = 1.44 (1.57–1.92), sterno = 0.79 (0.60–0.85), orbito = 1.46 (1.19–1.40), dcp = 0.30 (0.32–0.37), sctlp = 1.92 (1.41–1.79), C = 1.68 (1.62–1.82), 4c = 1.10 (1.30–1.36), 4v = 1.42 (1.86–2.07), 5x = 1.55 (1.34–1.50), ac = 10.86 (9.18–10.22), M = 0.70 (0.62–0.65), C3F = 0.61 (0.55–0.62). Type materials. Holotype ♂ (SCAU, No. 125725, DNA #2673), CHINA: Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 25.x.2017, ex tree trunk, H.W. Chen. Paratypes: 1♀ (SCAU, No. 125726, DNA #2692), other data same as holotype; 1♂, 1♀ (SCAU, Nos. 125727, 28, DNA #2720), Longtan Park, Ximeng, Yunnan, 22°38'55''N, 99°36'02''E, alt. 1150m, 17.iv.2018, ex tree trunks, L. Gong, Y. Lin. Distribution. China (Yunnan). Etymology. From the name of the type locality "Menglian"., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 85-86, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) nayun Li & Chen, sp. nov. (Figs. 12C, 12D, 54) Diagnosis. This species is similar to S. (S.) cylindrica Wang, Gao, & Chen, 2013 in the shape of male terminalia (as fig. 2 in Wang et al. 2013, Fig. 54), but can be distinguished from the latter by having the tubular aedeagus distoventrally with 3 rugae per side, lacking pubescence (Fig. 54C, D); the subepandrial sclerite lacking pubescence (Fig. 54G). Description. Male and female: First flagellomere brownish. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally (Fig. 12C). Katepisternum yellow (Fig. 12D). Scutellum brown medially, dark brown laterally, yellow at tip (Fig. 12C). Halter mostly grey (Fig. 12D). Fore femur with a row of 4 long setae on distal part of posteroventral surface. Abdominal tergites brown to dark brown; sternites brown, about 2 times as wide as long. Male terminalia (Fig. 54A–G): Epandrium with ca. 11 setae on dorsal to posterior portions per side (Fig. 54A). Surstylus with 1 peg-like tooth and some setulae near distal margin, lacking pubescence (Fig. 54A, B). Cercus lacking pubescence (Fig. 54A). Hypandrium strongly expanded anteromedially (Fig. 54C, D). Pregonite small, with 2 sensilla (Fig. 54C, D). Aedeagus lacking serrated process (Fig. 54C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 54E, F). Subepandrial sclerite slightly rectangle distally (Fig. 54G). Female terminalia (Fig. 54H–J): Epiproct with several setae on distal 1/2 (Fig. 54H). Cercus and hypoproct with numerous setae and much pubescence (Fig. 54H, J). Eight sternite about 1.5 times as long as wide, glabrous on basal 4/5, with several setae and some pubescence on distal 1/5 (Fig. 54I). HW = 0.38 (0.42), ch/o = 0.29 (0.38), prorb = 1.05 (1.50), rcorb = 0.79 (0.88), vb = 0.64 (0.93), dcl = 0.43 (0.35), presctl = 0.70 (0.70), sctl = 3.15 (2.39), sterno = 1.19 (0.93), orbito = 1.89 (1.89), dcp = 0.21 (0.21), sctlp = 1.78 (2.22), C = 1.98 (1.95), 4c = 1.05 (1.39), 4v = 1.20 (2.21), 5x = 1.54 (1.91), ac = 10.25 (13.00), M = 0.55 (0.76), C3F = 0.61 (0.70). Type materials. Holotype ♂ (SCAU, No. 125730, DNA #2674), CHINA: Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 25.x.2017, ex tree trunk, H.W. Chen. Paratypes: 1♀ (SCAU, No. 125731, DNA #2693), other data same as holotype; 1♀ (SCAU, No. 125787, DNA #2608), Fengtongzhai, Baoxing, Sichuan, 30°32'42''N, 102°54'24''E, alt. 1500m, 18.vii.2017, ex tree trunk, Y.L. Wang. Distribution. China (Sichuan, Yunnan). Etymology. From the name of the type locality "Jinshan" belonging to "Nayun, Menglian, Yunnan "., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 88-90, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, L., Gao, J. J. & Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055"]}

Stegana (Steganina) multidentata Chen, Gao, & Chen, 2009 (Fig. 5I, J) Stegana (Steganina) multidentata Chen, Gao, & Chen in Chen et al. 2009: 1917. Specimens examined. CHINA: 2♂, 5♀ (SCAU, Nos. 125419–25, DNA #2472), CHINA: Gexigou, Yajiang, Sichuan, 30°02'58''N, 100°57'36''E, alt. 2800m, 4.vi.2018, ex tussocks, Y. Lin, Y.L. Wang. Distribution. China (Hubei, Sichuan *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 50, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

VIII. Ungrouped Stegana (Steganina) species (67 spp.) angulistrata Wu & Chen, 2010; China (Jiangxi *, Guangxi, Yunnan). clavispinula Chen & Chen, 2009; China (Hainan). convena Wang, Gao, & Chen, 2013; China (Yunnan). cyclophylla Chen & Chen, 2009; China (Guangxi *, Yunnan). cyclostoma Wu & Chen, 2010; China (Yunnan). cylindrica Wang, Gao, & Chen, 2013; China (Hainan, Guangxi *, Yunnan *). euryphylla Chen & Chen, 2009; China (Taiwan, Yunnan). eurystoma Wang, Gao, & Chen, 2013; China (Yunnan). hamata Wang, Gao, & Chen, 2013; China (Yunnan). hirticeps Wang, Gao, & Chen, 2013; China (Guangdong *, Guangxi, Yunnan *). izu Sidorenko, 1997; China (Taiwan, Yunnan*), Japan (Honshu, Ryukyu Islands*). jianqinae Zhang, Tsaur, & Chen, 2014; China (Taiwan, Hainan, Guangxi). kanmiyai Okada & Sidorenko, 1992; China (Hunan *, Taiwan, Guangdong*, Guangxi*, Sichuan*, Yunnan*), Japan (Honshu). latigena Wang, Gao, & Chen, 2013; China (Guangdong, Guangxi, Guizhou *, Yunnan). maymyo Sidorenko, 1997; China * (Yunnan), Myanmar (Pyin Oo Lwin). nigrifrons de Meijere, 1911: China (Taiwan), Japan (Honshu, Kyushu), Nepal, Indonesia (Java, Sumatra), Africa. planiceps Wang, Gao, & Chen, 2013; China (Hainan, Yunnan). polytricapillum Wu & Chen, 2010; China (Hainan, Yunnan). rhomboica Wang, Gao, & Chen, 2013; China (Guangxi, Yunnan). serratoprocessata Chen & Chen, 2009; China (Guangdong, Hainan, Guangxi *, Yunnan *, Xizang *). setivena Wang, Gao, & Chen, 2013; China (Yunnan). sphaerica Wang, Gao, & Chen, 2013; China (Hainan, Yunnan). wulai Zhang, Tsaur, & Chen, 2014; China (Taiwan). alafoliacea Zhang & Chen, sp. nov.; China (Guangxi). baoxing Li & Chen, sp. nov.; China (Sichuan). bimai Cui & Chen, sp. nov.; China (Yunnan). cinereipecta Zhang & Chen, sp. nov.; China (Yunnan). cordhirsuta Wang & Chen, sp. nov.; China (Yunnan). cornuta Li & Chen, sp. nov.; China (Yunnan). cucullata Li & Chen, sp. nov.; China (Guangxi, Chongqing, Sichuan). cultella Cui & Chen, sp. nov.; China (Yunnan). curvitabulata Cui & Chen, sp. nov.; China (Hainan). daiya Cui & Chen, sp. nov.; China (Sichuan). dendrophila Zhang & Chen, sp. nov.; China (Yunnan). flabella Li & Chen, sp. nov.; China (Yunnan). flavipes Li & Chen, sp. nov.; China (Yunnan). formosa Zhang & Chen, sp. nov.; China (Taiwan). fusca Li & Chen, sp. nov.; China (Yunnan). fuscipes Li & Chen, sp. nov.; China (Yunnan). glaucopalpula Cui & Chen, sp. nov.; China (Taiwan). haba Zhang & Chen, sp. nov.; China (Yunnan). hirticlavata Cui & Chen, sp. nov.; China (Yunnan). idiasta Cui & Chen, sp. nov.; China (Yunnan). kanda Cui & Chen, sp. nov.; China (Yunnan). labao Cui & Chen, sp. nov.; China (Yunnan). lancang Li & Chen, sp. nov.; China (Yunnan). latifoliacea Wang & Chen, sp. nov.; China (Yunnan). liusanjieae Li & Chen, sp. nov.; China (Guangxi). magniflava Cui & Chen, sp. nov.; China (Yunna). mailangang Li & Chen, sp. nov.; China (Yunnan). menghai Zhang & Chen, sp. nov.; China (Yunnan). menglian Li & Chen, sp. nov.; China (Yunnan). minutiflava Li & Chen, sp. nov.; China (Guangdong, Hainan, Yunnan). multiprocera Li & Chen, sp. nov.; China (Hainan). nayun Li & Chen, sp. nov.; China (Sichuan, Yunnan). nigridentata Wang & Chen, sp. nov.; China (Guangxi, Yunnan). otphylla Cui & Chen, sp. nov.; China (Sichuan, Yunnan). radiciflava Zhang & Chen, sp. nov.; China (Yunnan). rava Cui & Chen, sp. nov.; China (Yunnan). sciophila Li & Chen, sp. nov.; China (Yunnan). septencolorata Li & Chen, sp. nov.; China (Yunnan). silvestrella Zhang & Chen, sp. nov.; China (Yunnan). simola Cui & Chen, sp. nov.; China (Yunnan). yani Li & Chen, sp. nov.; China (Yunnan). yixiang Zhang & Chen, sp. nov.; China (Yunnan). zaduo Cui & Chen, sp. nov.; China (Yunnan). zhuoma Cui & Chen, sp. nov.; China (Xizang)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 106-107, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wu, L. & Chen, H. W. (2010) Three new species of Stegana (Steganina) (Diptera: Drosophilidae) from southern China. Oriental Insects, 44, 69 - 75. https: // doi. org / 10.1080 / 00305316.2010.10417607","Okada, T. & Sidorenko, V. S. (1992) New or unrecorded species of Stegana (Steganina) (Diptera, Drosophilidae) from the Old World, especially Japan and Russia. Japanese Journal of Entomology, 60, 415 - 426.","de Meijere, J. C. H. (1911) Studien uber sudostasiatische Dipteren. VI. Tijdschrift voor Entomologie, 54, 258 - 432."]}

Stegana (Steganina) lancang Li & Chen, sp. nov. (Figs. 10E, 10F, 46) Diagnosis. This species is similar to S. (S.) kanmiyai Okada & Sidorenko, 1992 in the shape of male terminalia (as fig. 3E–G in Okada & Sidorenko 1992, Fig. 46A–G), but can be distinguished from the latter by having the brownish yellow body color (Fig. 10E, F); the surstylus with 1 peg-like tooth near distal margin (Fig. 46A, B); the aedeagus lacking pubescence distolaterally (Fig. 46C, D). Description. Male: Clypeus and palpus yellow. Mesonotum brownish yellow (Fig. 10E). Katepisternum yellowish (Fig. 10F). Scutellum brownish yellow, brownish near distal margin (Fig. 10E). Halter mostly grey (Fig. 10F). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites brownish yellow (Fig. 10E, F); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 46A–G): Epandrium with 13 setae on dorsal to posterior portions per side (Fig. 46A). Surstylus with numerous setulae distally and some minute, serrated processes distoventrally (Fig. 46C, D). Aedeagal sheath acute apically, with 1 pair of acute processes subapically (Fig. 46E, F). Subepandrial sclerite vaulted (Fig. 46G). Female terminalia (Fig. 46H–J): Epiproct with several setae on distal 1/3 (Fig. 46H). Cercus and hypoproct with numerous setae and much pubescence (Fig. 46H, J). Eight sternite about 2 times as long as wide, mostly glabrous except with some setae distally (Fig. 46I). Measurements and indices. BL = 2.74 mm in holotype (2.33–2.67 in 5♂, 2.39–2.73 in 4♀ paratypes), ThL = 1.32 mm (1.21–1.31 in ♂, 1.21–1.37 in ♀), WL = 2.48 mm (2.16–2.43 in ♂, 2.27–2.63 in ♀), WW = 0.99 mm (0.95–1.02 in ♂, 0.95–1.05 in ♀), arb = 6/4 (6–8/4–5), avd = 0.88 (0.74–0.89), adf = 1.05 (1.00–1.74), flw = 1.81 (1.56–2.15), FW/HW = 0.47 (0.34–0.43), ch/o = 0.27 (0.22–0.30), prorb = 0.93 (0.77–1.53), rcorb = 0.60 (0.62–0.91), vb = 0.52 (0.46–0.83), dcl = 0.87 (0.59–0.87), presctl = 0.52 (0.60–0.84), sctl = damaged (1.41–1.94), sterno = 0.52 (0.63–1.08), orbito = 1.80 (1.59–1.96), dcp = 0.30 (0.24–0.35), sctlp = 1.67 (1.27–2.19), C = 1.76 (1.65–2.11), 4c = 1.26 (0.96–1.30), 4v = 1.86 (1.44–1.84), 5x = 1.59 (1.26–1.76), ac = 10.67 (8.17–11.00), M = 0.67 (0.55–0.72), C3F = 0.68 (0.57–0.66). Type materials. Holotype ♂ (SCAU, No. 125697, DNA #2637), CHINA: Guanlei, Mengla, Yunnan, 21°38'39''N, 101°09'52''E, alt. 560m, 24.iv.2016, ex tree trunk, J. Huang. Paratypes: 4♀ (SCAU, Nos. 125698–701, DNA #2637), other data same as holotype; 8♂ (SCAU, Nos. 125702–09, DNA #2638), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 560m, 17.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Yunnan). Etymology. From the word " lancang " in Dai language from the Dai nationality living in Yunnan, China, referring to the Lancang River.

Stegana (Steganina) septencolorata Li & Chen, sp. nov. (Figs. 13E, 13F, 60) Diagnosis. This species is similar to S. (S.) nayun sp. nov. in the shape of male terminalia (Figs. 54, 60), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth and some minute, serrated processes near distal margin (Fig. 60A, B); the cercus with much pubescence (Fig. 60A); the aedeagus with some pubescence distoventrally (Fig. 60C, D); the subepandrial sclerite vaulted, roundly expanded distally (Fig. 60G). Description. Male: First flagellomere brownish. Clypeus brownish yellow. Mesonotum brownish medially, yellow laterally (Fig. 13E). Katepisternum yellowish, brownish on anterior corner (Fig. 13F). Scutellum brown, yellow at tip (Fig. 13E). Halter mostly white (Fig. 13F). Fore femur with a row of 3 long, suberect setae on ventral surface. Abdominal tergites brownish to brown (Fig. 13E, F); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 60): Epandrium with ca. 7 setae on dorsal to posterior portions per side (Fig. 60A). Surstylus with numerous setulae near distal margin, lacking pubescence (Fig. 60A, B). Hypandrium roundly expanded anteromedially (Fig. 60C, D). Pregonite small, with 5 sensilla (Fig. 60C, D). Aedeagus tubular, distally expanded, distoventrally with 5 rugae per side, lacking serrated process (Fig. 60C, D). Aedeagal sheath with some minute, serrated processes distally (Fig. 60E, F). Measurements and indices. BL = 2.23 mm in holotype, ThL = 0.96 mm, WL = 2.07 mm, WW = 0.93 mm, arb = 5/3, avd = 0.85, adf = 1.12, flw = 1.73, FW/HW = 0.34, ch/o = 0.17, prorb = 0.90, rcorb = 0.23, vb = 0.48, dcl = 0.50, presctl = 0.55, sctl = 2.00, sterno = damaged, orbito = 2.33, dcp = damaged, sctlp = 1.53, C = 1.89, 4c = 0.71, 4v = 1.11, 5x = 1.36, ac = 11.29, M = 0.38, C3F = 0.78. Type material. Holotype ♂ (SCAU, No. 125668, DNA #1879), CHINA: Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1420m, 13.v.2012, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. A combination of the Latin words " septen -" (= seven) and " coloratus " (= colored), referring to the type locality " Yunnan " as a colorful place in China., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 95-96, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) nigripes Zhang & Chen, 2015 (Fig. 20A–C) Stegana (Steganina) nigripes Zhang & Chen, 2015: 135. Remarks. Since the terminalia of the only male specimen was partly damaged (probably eaten by a cockroach) and undescribed, the specimens collected from Yingjiang, Yunnan showed no other significant morphological differences with the type material of S. (S.) nigripes from Menghai, Yunnan. They were conservatively identified as conspecific. Description. Female terminalia (Fig. 20A–C): Epiproct and cercus with numerous setae, lacking pubescence (Fig. 20A). Eighth sternite about 2 times as wide as long, with equally distributed numerous prensisetae (Fig. 20B). Hypoproct with numerous setae and much pubescence (Fig. 20C). Specimens examined. CHINA: 1♂ (SCAU, No. 125409, DNA #1985), 1♀ (SCAU, No. 125408, DNA #1981), Tongbiguan, Yingjiang, Yunnan, 24°37'04''N, 97°40'03''E, 19.v.2016, alt. 1310m, 13.v.2016, ex tussock, J. Huang; 1♀ (SCAU, No. 125410, DNA #1981), Botanic Garden, Ruili, Yunnan, 24°04'19''N, 97°49'08''E, alt. 1170m, 22.v.2016, ex tussock, Y.Q. Liu. Distribution. China (Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 45-46, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Zhang, Y. & Chen, H. W. (2015) Four new species of the Stegana ornatipes species group (Diptera: Drosophilidae) from Yunnan, China, with DNA barcoding information. Zootaxa, 3905 (1), 131 - 137. https: // doi. org / 10.11646 / zootaxa. 3905.1.8"]}

Stegana (Steganina) alafoliacea Zhang & Chen, sp. nov. (Figs. 6A, 6B, 27) Diagnosis. This species is similar to S. (S.) baoxing sp. nov. in the dark body color (Fig. 6A–D), but can be distinguished from the latter by having the surstylus with numerous minute, serrated processes on outer surface (Fig. 27A); the aedeagus strongly expanded apicolaterally in ventral view (Fig. 27B); the aedeagal sheath with numerous minute, serrated processes (Fig. 27D, E). Description. Male: First flagellomere black. Clypeus black. Mesonotum and scutellum dark brown (Fig. 6A, as wide as long. Male terminalia (Fig. 27): Epandrium with ca. 5 setae on dorsal to posterior portions per side (Fig. 27A). Surstylus curved submedially, with 1 peg-like tooth and several setulae distally (Fig. 27A). Cercus with much pubescence (Fig. 27A). Hypandrium expanded anteromedially (Fig. 27B, C). Pregonite blurry or absent (Fig. 27B, C). Aedeagal sheath erect, slightly dorsad curved (Fig. 27D, E). Subepandrial sclerite deeply split medially (Fig. 27F). Measurements and indices. BL = 2.35 mm in holotype, ThL = 1.27 mm, WL = 2.00 mm, WW = 0.93 mm, arb = 7/4, avd = 1.60, adf = 1.25, flw = 2.50, FW/HW = 0.45, ch/o = 0.10, prorb = 1.22, rcorb = 0.67, vb = 0.75, dcl = 0.73, presctl = 0.28, sctl = 1.80, sterno = 0.80, orbito = 2.00, dcp = 0.20, sctlp = 1.71, C = 1.88, 4c = 1.33, 4v = 1.83, 5x = 1.67, ac = 8.00, M = 0.83, C3F = 0.63. Type material. Holotype ♂ (SCAU, No. 125631, DNA #1459), CHINA: Maoershan, Xingan, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 17.x.2004, ex tree trunk, H.W. Chen. Distribution. China (Guangxi). Etymology. A combination of the Latin words " alaris " (= alary) and '' foliaceus " (= foliaceous), referring to the apicolaterally strongly extended aedeagus.

Stegana (Steganina) menglian Li & Chen, sp. nov. (Figs. 11G, 11H, 51) Diagnosis. This species is similar to S. (S.) lancang sp. nov. in the shape of male terminalia (Figs. 46A–G, 51), but can be distinguished from the latter by having the surstylus lacking serrated process (Fig. 51A, B); the aedeagus lacking a row of serrated processes distoventrally (Fig. 51C, D). Description. Male and female: First flagellomere yellowish brown. Clypeus yellow. Mesonotum dark brown medially, brown laterally (Fig. 11G). Katepisternum mostly yellowish, brownish on anterior corner (Fig. 11H). Scutellum dark brown, yellow at tip (Fig. 11G). Halter mostly white (Fig. 11H). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites black (Fig. 11G, H); sternites brown, about 2 times as wide as long. Male terminalia (Fig. 51): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 51A). Surstylus with 1 peg-like tooth and some setulae near distal margin, lacking pubescence (Fig. 51A, B). Cercus with much pubescence (Fig. 51A). Hypandrium roundly expanded anteromedially (Fig. 51C, D). Pregonite with 2 sensilla (Fig. 51C, D). Aedeagus roundly expanded distally, with several minute, serrated processes distolaterally and some pubescence distoventrally (Fig. 51C, D). Aedeagal sheath distally strongly dorsad curved and with several minute, serrated processes (Fig. 51E, F). Subepandrial sclerite nearly vaulted (Fig. 51G). Measurements and indices. BL = 2.21 mm in holotype (2.43 in 1♂, 2.73–2.79 in 2♀ paratypes), ThL = 1.08 mm (1.21 in ♂, 1.32–1.36 in ♀), WL = 2.09 mm (2.12 in ♂, 2.46–2.52 in ♀), WW = 0.88 mm (0.91 in ♂, 0.99–1.04 in ♀), arb = 7/4 (5–6/3–4), avd = 0.84 (0.77–0.88), adf = 0.97 (1.11–1.26), flw = 1.77 (1.73–1.93), FW/HW = 0.40 (0.41–0.45), ch/o = 0.30 (0.25–0.27), prorb = 1.18 (1.04–1.20), rcorb = 0.78 (0.57–0.71), vb = 0.71 (0.71–0.72), dcl = 0.74 (0.53–0.80), presctl = 0.67 (0.47–0.88), sctl = 1.44 (1.57–1.92), sterno = 0.79 (0.60–0.85), orbito = 1.46 (1.19–1.40), dcp = 0.30 (0.32–0.37), sctlp = 1.92 (1.41–1.79), C = 1.68 (1.62–1.82), 4c = 1.10 (1.30–1.36), 4v = 1.42 (1.86–2.07), 5x = 1.55 (1.34–1.50), ac = 10.86 (9.18–10.22), M = 0.70 (0.62–0.65), C3F = 0.61 (0.55–0.62). Type materials. Holotype ♂ (SCAU, No. 125725, DNA #2673), CHINA: Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 25.x.2017, ex tree trunk, H.W. Chen. Paratypes: 1♀ (SCAU, No. 125726, DNA #2692), other data same as holotype; 1♂, 1♀ (SCAU, Nos. 125727, 28, DNA #2720), Longtan Park, Ximeng, Yunnan, 22°38'55''N, 99°36'02''E, alt. 1150m, 17.iv.2018, ex tree trunks, L. Gong, Y. Lin. Distribution. China (Yunnan). Etymology. From the name of the type locality "Menglian".

Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen, 2009 Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen in Chen et al. 2009: 1914. Specimens examined. CHINA: 1♂ (SCAU, Nos. 125426, DNA #2656), Nonggang, Longzhou, Guangxi, 22°32'26''N, 106°43'18''E, alt. 230m, 23.viii.2004, ex tree trunk, H.W. Chen. Distribution. China (Hainan, Guangxi *).

Stegana (Steganina) zhuoma Cui & Chen, sp. nov. (Figs. 14G, 14H, 65D–F) Diagnosis. This species is similar to S. (S.) zaduo sp. nov. in the shape of female terminalia (Fig. 65), but can be distinguished from the latter by having the yellowish katepisternum brown on anterior corner (Fig. 14H); the 8th sternite nearly squared, with several setae and much pubescence on distolateral 1/3 (Fig. 65E). Description. Female: First flagellomere brown. Clypeus black. Mesonotum black medially, brown laterally (Fig. 14G). Scutellum black, yellow at tip (Fig. 14G). Halter mostly white (Fig. 14H). Fore femur with a row of 3 long, strong setae on distal part of posteroventral surface. All abdominal tergites dark brown to black (Fig. 14G, H); sternites brown, about 2 times as wide as long. Female terminalia (Fig. 65D–F): Epiproct with several setae, lacking pubescence (Fig. 65D). Cercus and hypoproct with numerous setae and much pubescence (Fig. 65D, F). Eighth sternite glabrous on basal 2/3 (Fig. 65E). Measurements and indices. BL = 2.70 mm in holotype, ThL = 1.45 mm, WL = 2.92 mm, WW = 1.26 mm, arb = 7/5, avd = 0.85, adf = 1.17, flw = 2.22, FW/HW = 0.39, ch/o = 0.25, prorb = 1.02, rcorb = 0.76, vb = 0.74, dcl = 0.48, presctl = 0.57, sctl = damaged, sterno = 0.77, orbito = 1.90, dcp = 0.32, sctlp = 1.45, C = 1.66, 4c = 1.35, 4v = 1.95, 5x = 1.45, ac = 12.89, M = 0.71, C3F = 0.63. Type material. Holotype ♀ (SCAU, No. 125789, DNA #2757), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt. 2000m, 11.x.2010, ex tree trunk, L. Wang. Distribution. China (Xizang). Etymology. From the word " zhuoma " in Zang language from the Zang nationality living in Xizang, China, referring to women.

Stegana (Steganina) rava Cui & Chen, sp. nov. (Figs. 13A, 13B, 58) Diagnosis. This species is similar to S. (S.) flavipes sp. nov. in the shape of male terminalia (Figs. 38, 58), but can be distinguished from the latter by having the surstylus laterally with some minute, serrated processes (Fig. 58A, B); the aedeagus with some pubescence distoventrally (Fig. 58C, D); the subepandrial sclerite vaulted (Fig. 58G). Description. Male: First flagellomere brown. Clypeus yellow. Mesonotum yellow (Fig. 13A). Katepisternum yellowish (Fig. 13B). Scutellum yellow, brownish near distal margin (Fig. 13A). Halter mostly brownish (Fig. 13B). Fore femur with 4 thin, long setae on distal part of posteroventral surface. Abdominal tergites brownish (Fig. 13A, B); sternites yellow, about 1.6 times as wide as long. Male terminalia (Fig. 58): Epandrium with ca. 8 setae on dorsal to posterior portions per side (Fig. 58A). Surstylus slightly oval, with some setulae along distal margin and inner surface, lacking pubescence (Fig. 58A, B). Cercus with sparse pubescence (Fig. 58A). Hypandrium expanded anteromedially (Fig. 58C, D). Pregonite small, with 2 sensilla (Fig. 58C, D). Aedeagus tubular, distally slightly expanded, distoventrally with 3 rugae per side, lacking serrated process (Fig. 58C, D). Aedeagal sheath small, lacking serrated process (Fig. 58E, F). Measurements and indices. BL = 2.67 mm in holotype, ThL = 1.15 mm, WL = 2.32 mm, WW = 1.01 mm, arb = 6/5, avd = 0.76, adf = 0.93, flw = 1.76, FW/HW = 0.37, ch/o = 0.28, prorb = 0.87, rcorb = 0.68, vb = 0.60, dcl = 0.61, presctl = 0.80, sctl = 2.10, sterno = 0.76, orbito = 1.67, dcp = 0.21, sctlp = 1.51, C = 1.66, 4c = 1.07, 4v = 1.63, 5x = 1.23, ac = 9.27, M = 0.52, C3F = 0.62. Type material. Holotype ♂ (SCAU, No. 125656, DNA #2705), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 560m, 16,17. iv.2007, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. From the word " rava " in Dai language from the Dai nationality living in Yunnan, China, referring to the meaning "a happy place"., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 93-94, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) fusca Li & Chen, sp. nov. (Figs. 8I, 8J, 40) Diagnosis. This species differs from the other Steganina species in having the small body size (BL ranged from 2.33–2.85 mm in measured specimens) and the relatively light body color (Fig. 8I, J); the aedeagal opening nearly triangular in ventral view, with thick pubescence (Fig. 40B). Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum brownish yellow in male, brown medially, brownish laterally in female (Fig. 8I). Katepisternum yellowish (Fig. 8J). Scutellum brownish medially, brown laterally (Fig. 8I). Halter mostly brownish (Fig. 8J). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites yellow to brownish in male (Fig. 8I, J), brown in female; sternites brown, slightly wider than long. Male terminalia (Fig. 40): Epandrium with ca. 10 setae on dorsal to posterior portions per side (Fig. 40A). Surstylus with 1 peg-like tooth and several setulae near distal margin, lacking pubescence (Fig. 40A). Cercus lacking pubescence (Fig. 40A). Hypandrium roundly expanded anteromedially (Fig. 40B, C). Pregonite with 5 sensilla (Fig. 40B, C). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 40D, E). Subepandrial sclerite slightly vaulted (Fig. 40F). Measurements and indices. BL = 2.75 mm in holotype (2.33–2.73 in 2♂, 2.37–2.85 in 3♀ paratypes), ThL = 1.29 mm (1.15–1.34 in ♂, 1.21–1.34 in ♀), WL = 2.21 mm (2.25–2.68 in ♂, 2.41–2.65 in ♀), WW = 1.14 mm (1.01–1.24 in ♂, 0.95–1.16 in ♀), arb = 7/3 (6–8/4–5), avd = 0.63 (0.74–0.89), adf = 1.24 (1.03–1.50), flw = 1.85 (2.36–3.13), FW/HW = 0.41 (0.37–0.43), ch/o = 0.26 (0.16–0.39), prorb = 1.35 (0.83–1.06), rcorb = 1.10 (0.57– 0.72), vb = 0.89 (0.61–0.75), dcl = 0.66 (0.54–0.75), presctl = 0.64 (0.52–0.78), sctl = 1.90 (1.87–2.52), sterno = 0.89 (0.67–1.14), orbito = 1.62 (1.63–2.05), dcp = 0.30 (0.29–0.36), sctlp = 1.39 (1.33–1.93), C = 1.65 (1.49–1.82), 4c = 1.04 (1.24–1.56), 4v = 1.44 (1.68–2.14), 5x = 1.53 (1.45–1.85), ac = 11.33 (9.91–13.00), M = 0.50 (0.59–0.74), C3F = 0.69 (0.56–0.70). Type materials. Holotype ♂ (SCAU, No. 125686, DNA #2172), CHINA: Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt. 1320m, 10.v.2016, ex tree trunk, Y.L. Wang. Paratypes: 2♂, 3♀ (SCAU, Nos. 125687– 91, DNA #2729), Muyiji Park, Ximeng, Yunnan, 22°37'13''N, 99°35'45''E, alt. 1100m, 30.iv.2016, ex tree trunks, J. Huang, Y.Q. Liu, Y.L. Wang, L. Zhu. Distribution. China (Yunnan). Etymology. From the Latin word " fuscus " (= brown), referring to the brown body color., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 71, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) hirticeps Wang, Gao, & Chen, 2013 (Fig. 23) Stegana (Steganina) hirticeps Wang, Gao, & Chen in Wang et al. 2013: 2002. Remarks. The specimens collected from Guangdong and Yunnan slightly differ from the type materials from Guangxi in having the aedeagus with several minute, serrated processes laterally (Fig. 23B, C); the aedeagal sheath lacking serrated process (Fig. 23D, E). They were conservatively identified as conspecific. Specimens examined. 1♂ (SCAU, No. 125523, DNA #2604), CHINA: Nanling, Shaoguan, Guangdong, 24°56'41''N, 113°03'04''E, alt. 1300m, 29.v.2018, ex tree trunk, Y. Lin; 1♂ (SCAU, No. 125524, DNA #2680), Husa, Longchuan, Yunnan, 24°28'21''N, 97°54'06''E, alt. 1340m, 7.xi.2017, ex tree trunk, H.W. Chen; 1♂, 3♀ (SCAU, Nos. 125525–28, DNA #2679), Hesong, Menghai, Yunnan, 21°50'17''N, 100°05'48''E, alt. 1600m, 7.v.2012, ex tree trunks, H.W. Chen. Distribution. China (Guangdong *, Guangxi, Yunnan *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 51, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, L., Gao, J. J. & Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055"]}

Stegana (Steganina) latifoliacea Wang & Chen, sp. nov. (Figs. 10G, 10H, 47) Diagnosis. This species is similar to S. (S.) cucullata sp. nov. in the shape of male terminalia (Figs. 34, 47), but can be distinguished from the latter by having the mesonotum brownish yellow (Fig. 10G); the pregonite blurry or absent (Fig. 47C, D). Description. Male and female: First flagellomere yellow on basal 1/3, brownish on distal 2/3. Clypeus yellow. Katepisternum yellowish (Fig. 10H). Scutellum brownish to dark brown (Fig. 10G). Halter mostly grey (Fig. 10H). Fore femur with a row of 4 setae on distal part of posteroventral surface. Abdominal tergites dark brown medially, brownish laterally in male (Fig. 10G, H), mostly dark brown in female; sternites brownish, slightly wider than long. Male terminalia (Fig. 47): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 47A). Surstylus with 1 peg-like tooth and several setulae near distal margin, lacking pubescence (Fig. 47A, B). Cercus with much pubescence (Fig. 47A). Hypandrium roundly expanded anteromedially (Fig. 47C, D). Aedeagus dramatically expanded distally, with some pubescence distoventrally (Fig. 47C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 47E, F). Subepandrial sclerite slightly vaulted (Fig. 47G). Measurements and indices. BL = 2.67 mm in holotype (2.54–2.95 in 5♂, 2.79–3.05 in 5♀ paratypes), ThL = 1.30 mm (1.26–1.51 in ♂, 1.19–1.36 in ♀), WL = 1.51 mm (2.14–2.61 in ♂, 2.19–2.64 in ♀), WW = 0.80 mm (0.94–1.14 in ♂, 0.97–1.04 in ♀), arb = 10/6 (7–9/5–7), avd = 1.00 (0.73–0.94), adf = 1.00 (1.00–1.53), flw = 2.29 (1.65–2.29), FW/HW = 0.31 (0.34–0.39), ch/o = 0.14 (0.26–0.34), prorb = 1.25 (0.68–1.34), rcorb = 0.83 (0.71– 1.02), vb = 0.67 (0.63–0.90), dcl = 0.19 (0.38–0.71), presctl = 0.38 (0.43–0.78), sctl = 1.93 (1.65–2.09), sterno = 0.55 (0.62–1.23), orbito = 2.00 (1.56–1.96), dcp = 0.21 (0.27–0.36), sctlp = 1.90 (1.45–1.90), C = 1.83 (1.48–1.91), 4c = 1.21 (1.06–1.29), 4v = 2.00 (1.48–1.86), 5x = 1.13 (1.23–1.74), ac = 11.50 (9.00–14.57), M = 0.47 (0.54–0.72), C3F = 0.63 (0.53–0.68). Type materials. Holotype ♂ (SCAU, No. 125614, DNA #002), CHINA: Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 600m, 16.iv.2007, ex tree trunk, H.W. Chen. Paratypes: 9♂, 5♀ (SCAU, Nos. 125615–28, DNA #1439, 1936), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 660m, 22–25.iv.2007, ex tree trunks, H.W. Chen, J.J. Gao. Distribution. China (Yunnan). Etymology. A combination of the Latin words " latus " (= broad) and " foliaceus " (= foliaceous), referring to the distally expanded aedeagus., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 80, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) magniflava Cui & Chen, sp. nov. (Fig. 11A, 11B) Diagnosis. This species differs from the other Steganina species in having the large body size (BL = 3.32 mm in holotype) and the yellow body color (Fig. 11A, B). Remarks. The terminalia of the only male specimen was partly damaged (probably eaten by a cockroach) and 11A). Katepisternum yellowish (Fig. 11B). Halter mostly grey (Fig. 11B). Fore femur with a row of 4 or 5 strong setae on distal part of posteroventral surface.Abdominal tergites brownish yellow (Fig. 11A, B); sternites brownish, about 2 times as wide as long. Measurements and indices. BL = 3.32 mm in holotype, ThL = 1.49 mm, WL = 3.19 mm, WW = 1.33 mm, arb = 7/4, avd = 0.84, adf = 1.17, flw = 2.67, FW/HW = 0.35, ch/o = 0.31, prorb = 0.85, rcorb = 0.67, vb = 0.66, dcl = 0.53, presctl = 0.59, sctl = 1.96, sterno = 0.84, orbito = 1.67, dcp = 0.32, sctlp = 1.74, C = 1.77, 4c = 1.02, 4v = 1.48, 5x = 1.37, ac = 10.73, M = 0.48, C3F = 0.67. Type material. Holotype ♂ (SCAU, No. 125788, DNA #2078), Muyiji Park, Ximeng, Yunnan, 22°37'15''N, 99°35'42''E, alt. 1200m, 29.iv.2016, ex tree trunk, J. Huang. Distribution. China (Yunnan). Etymology. A combination of the Latin words " magnus " (= large) and " flavus " (= yellow), referring to the large body size and the yellow body color., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 82-83, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) sciophila Li & Chen, sp. nov. (Figs. 13C, 13D, 59) Diagnosis. This species is similar to S. (S.) hirticeps Wang, Gao, & Chen, 2013 in the shape of male terminalia (as fig. 5 in Wang et al. 2013, Fig. 59), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth near distal margin (Fig. 59A, B); the subepandrial sclerite vaulted, bifurcated basally per side, roundly expanded distally (Fig. 59G). Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum dark brown medially, brownish laterally (Fig. 13C). Katepisternum yellowish (Fig. 13D). Scutellum mostly yellow, brown anteromedially (Fig. 13C). Halter mostly brownish (Fig. 13D). Fore femur with a row of 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 13C, D); sternites brown in male, yellow in female, about 2.2 times as wide as long. Male terminalia (Fig. 59): Epandrium with ca. 8 setae on dorsal to posterior portions per side (Fig. 59A). Surstylus with some minute, serrated processes on outer surface and setulae near distal margin, lacking pubescence (Fig. 59A). Cercus lacking pubescence (Fig. 59A). Hypandrium roundly expanded anteromedially (Fig. 59C, D). Pregonite small, with 2 sensilla (Fig. 59C, D). Aedeagus with much pubescence distomedially (Fig. 59C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 59E, F). Measurements and indices. BL = 3.07 mm in holotype (2.54–2.88 in 4♂, 3.13–3.21 in 3♀ paratypes), ThL = 1.48 mm (1.21–1.36 in ♂, 1.36–1.49 in ♀), WL = 2.97 mm (2.33–2.78 in ♂, 2.64–2.91 in ♀), WW = 1.23 mm (0.92–1.04 in ♂, 1.01–1.31 in ♀), arb = 7/5 (5–7/4–5), avd = 0.76 (0.73–0.91), adf = 1.44 (0.96–1.30), flw = 2.05 (1.53–2.21), FW/HW = 0.34 (0.33–0.39), ch/o = 0.19 (0.26–0.34), prorb = 1.09 (1.07–1.46), rcorb = 0.78 (0.69– 1.04), vb = 0.87 (0.60–0.82), dcl = 0.43 (0.49–0.82), presctl = 0.49 (0.55–0.83), sctl = 2.33 (1.66–3.04), sterno = 0.81 (0.69–1.45), orbito = 1.43 (1.56–2.00), dcp = 0.31 (0.31–0.37), sctlp = 1.73 (1.14–1.92), C = 1.82 (1.60–1.95), 4c = 1.19 (1.02–1.22), 4v = 1.84 (1.51–1.69), 5x = 1.94 (1.33–1.69), ac = 9.33 (9.91–14.00), M = 0.66 (0.51–0.60), C3F = 0.62 (0.55–0.64). Type materials. Holotype ♂ (SCAU, No. 125717, DNA #2667), CHINA: Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 25.x.2017, ex tree trunk, H.W. Chen. Paratypes: 3♂, 1♀ (SCAU, Nos. 125718– 21), L. Gong, B.X. Li, other data same as holotype; 1♂, 2♀ (SCAU, Nos. 125722–24, DNA #2668), Muyiji Park, Ximeng, Yunnan, 22°37'23''N, 99°36'11''E, alt. 1100m, 16.iv.2018, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Yunnan). Etymology. From the Greek word " sciophilous " (= sciophilous), referring to the sciophilous Steganina flies.

Stegana (Steganina) simola Cui & Chen, sp. nov. (Figs. 13I, 13J, 62) Diagnosis. This species is similar to S. (S.) cinereipecta sp. nov. in the shape of male terminalia except aedeagus and subepandrial sclerite (Figs. 30, 62), but can be distinguished from the latter by having the surstylus with numerous minute, serrated processes on outer surface and lacking pubescence (Fig. 62A, B); the aedeagus distally roundly expanded, lacking pubescence (Fig. 62C, D); the subepandrial sclerite slightly vaulted (Fig. 62G). Description. Male and female: First flagellomere yellow. Clypeus yellow. Mesonotum brown medially, yellow laterally (Fig. 13I). Katepisternum yellowish (Fig. 13J). Scutellum dark brown (Fig. 13I). Halter mostly grey (Fig. 13J). Fore femur with a row of 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black medially, yellow laterally (Fig. 13I, J); sternites yellow, about 1.6 times as wide as long. Male terminalia (Fig. 62): Epandrium with ca. 11 setae on dorsal to posterior portions per side (Fig. 62A). Surstylus with 1 peg-like tooth and numerous setulae near distal margin (Fig. 62A, B). Cercus with much pubescence (Fig. 62A). Hypandrium roundly expanded anteromedially (Fig. 62C, D). Pregonite small, with 2 or 3 sensilla (Fig. 62C, D). Aedeagus with numerous minute, serrated processes distally (Fig. 62C, D). Aedeagal sheath distally with numerous minute, serrated processes (Fig. 62E, F). Measurements and indices. BL = 2.73 mm in holotype (2.73–2.80 in 2♂, 2.80–3.40 in 2♀ paratypes), ThL = 1.33 mm (1.27–1.33 in ♂, 1.33 in ♀), WL = 2.13 mm (2.13–2.20 in ♂, 2.00– 2.47 in ♀), WW = 1.00 mm (0.93–1.00 0.67 (0.60–0.80), dcl = 0.51 (0.48–0.51), presctl = 0.77 (0.69–0.77), sctl = 2.00 (2.00–2.50), sterno = 0.80 (0.60– 1.11), orbito = 2.67 (2.00–2.67), dcp = 0.22 (0.17–0.22), sctlp = 1.86 (1.75–2.50), C = 1.67 (1.56–2.00), 4c = 1.29 (1.13–1.50), 4v = 1.71 (1.43–2.33), 5x = 2.50 (1.67–2.50), ac = 9.00 (9.00), M = 0.71 (0.63–0.83), C3F = 0.56 (0.56–0.67). Type materials. Holotype ♂ (SCAU, No. 125650, DNA #1185), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 700m, 25.iv.2007, ex tree trunk, H.W. Chen. Paratypes: 2♂, 2♀ (SCAU, Nos. 125651–54, DNA #2662), other data same as holotype. Distribution. China (Yunnan). Etymology. From the word " simola " in Va language from the Va nationality living in Yunnan, China, referring to the meaning "a happy place"., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 98-99, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657