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In this study, the genus Sorolopha Lower, 1901 in Japan was reviewed with five species including three that are described herein as new: S. sugiurai Sakagami, sp. nov., S. viridis Sakagami, sp. nov., and S. baribaris Sakagami, sp. nov. Adults of S. sugiurai and S. baribaris emerged from leaf shelters of Actinodaphne acuminata (Lauraceae). Larvae of S. viridis feed on young leaves of Illicium anisatum (Schisandraceae). S. viridis is known from Honshu and Yakushima Island, S. baribaris from Honshu and Kyushu, and S. sugiurai only from Honshu (Wakayama Prefecture). Diagnoses, biological information, and images of the adults, and male and female genitalia are provided.

Berries are agricultural products of great economic interest for Mexico, and their production has increased in recent years; however, crops are affected by tortricid leafrollers. From August 2019 to April 2021 in Michoacán and Guanajuato, Mexico, a study was conducted to determine the species of tortricids associated with blackberries (Rubus spp. L.), raspberries (Rubus idaeus L.) and strawberries (Fragaria × ananassa Duch.), as well as their altitudinal distribution. In 12 orchards located in these states, shoots, leaves and flowers infested by larvae were collected. The species were identified by male genitalia and were determined taxonomically as Amorbia cuneana (Walsingham, 1879), Argyrotaenia montezumae (Walsingham, 1914) and Platynota sp. Walker, 1859, found at elevations from 1290 to 2372 m. The most abundant species were A. cuneana and A. montezumae. Generally, these tortricids prefer to feed on tender vegetative parts of the plant, but the economic impact they have is not known. It is worth mentioning that the number of species found is lower than those reported in other countries, but it is necessary to broaden the study area to other berry-producing regions to determine whether their distribution is wider.

Five species of Acroclita Lederer, 1859 are reported from China, including Acroclita ovata sp. nov., which is described and illustrated from Hubei Province. The previously described species are listed along with references cited, host-plants, specimens examined, and distributions. A taxonomic key to Chinese species of Acroclita is presented based on characters of the male genitalia.

Three new species of Xenolepis Diakonoff, 1973 are described from Thailand: X. fissurarta Thonongtor & Pinkaew, sp. nov., X. apicula Thonongtor & Pinkaew, sp. nov., and X. cultrata Thonongtor & Pinkaew, sp. nov. The genus was previously known from Indonesia and India, so the new species represent the first records of the genus from Thailand. Illustrations of adults and male genitalia are provided. Females of the genus remain unknown. The new taxa increase the number of described Xenolepis species to five.

Pararrhaptica lysimachiana (Swezey) Presumed extinct This species is known only from the female holotype, reared from leaves of Lysimachia rotundifolia [= L. hillebrandii] (Primulaceae) by Swezey at Puʻu Hāpapa on Oʻahu in 1927. Collecting by KAA at the type locality did not result in any specimens, despite the host plant’s presence in the immediate vicinity, and many island-wide surveys., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 16, DOI: 10.5281/zenodo.8159762

Spheterista pernitida (Walsingham) Presumed extinct This species is known from a single female collected at Olaʻa on Hawaiʻi island in 1896 by Perkins (Walsingham 1907). However, in the absence of males, it is difficult to determine whether it belongs to Spheterista or another genus. If Spheterista, it likely fed on leaves or fruit of Cheirodendron or Polyscias (Araliaceae). The male mentioned by Walsingham (1907) as being “closely allied to pernitida ” is almost certainly not conspecific. The male genitalia closely resemble those of the Urticaceae-feeding “ Spheterista ” (“ S.” infaustana, “ S.” urerana)., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 17, DOI: 10.5281/zenodo.8159762, {"references":["Walsingham, L. 1907. Microlepidoptera. In Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge. pp. 469 - 759."]}

Spheterista argentinotata (Walsingham) Presumed extinct This species is known from two females collected by Perkins in Olaʻa and Hilo, Hawaiʻi island in 1895 and 1896, respectively (Walsingham 1907). In the absence of known males, it is difficult to ascertain for certain whether it belongs to Spheterista or another genus. If Spheterista, it likely fed on leaves or fruit of Cheirodendron or Polyscias (Araliaceae)., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 17, DOI: 10.5281/zenodo.8159762, {"references":["Walsingham, L. 1907. Microlepidoptera. In Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge. pp. 469 - 759."]}

Cydia crassicornis (Walsingham) Presumed extinct This species is known only from two males collected by R.C.L. Perkins in 1892 at 4000 ft near Kona, Hawaiʻi island (Walsingham 1907). Although possibly a color form of the koa-feeding C. walsinghamii (Butler), Oboyski (2011) considered it distinct based on subtle differences in the male genitalia. Perkins’ field notes for September 1892 indicates that he “collected … by sifting dead leaves at the foot of a big koa tree” (Evenhuis 2007). The forewings are very similar to C. conspicua (Walsingham), but the absence of a “sex pouch” (Zimmerman 1978) on the ventral surface of the male hindwing readily separates it from that species., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 14, DOI: 10.5281/zenodo.8159762, {"references":["Walsingham, L. 1907. Microlepidoptera. In Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge. pp. 469 - 759.","Oboyski, P. T. 2011. The Systematics, Evolution, and Ecology of Hawaiian Cydia (Lepidoptera: Tortricidae) [Unpublished doctoral dissertation]. University California, Berkeley.","Evenhuis, N. L., editor. 2007. \" Barefoot on Lava: The Journals and Correspondence of Naturalist R. C. L. Perkins in Hawaiʻi, 1892 - 1901. \" Bishop Museum Press, Honolulu. 412 pp.","Zimmerman, E. C. 1978. Microlepidoptera. Insects of Hawaii. 9. The University Press of Hawaii, Honolulu. 1,903 pp."]}

Cydia gypsograpta (Meyrick) Presumed extinct This species is known only from the male holotype supposedly collected by R.C.L. Perkins near Honolulu, Oʻahu in 1908 (Meyrick 1932; but see comments below). Oboyski (2011) found it difficult to distinguish from Canavalia -feeding Hawaiian Cydia., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 14, DOI: 10.5281/zenodo.8159762, {"references":["Meyrick, E. 1932. Exotic Microlepidoptera. Exotic Microlepidoptera. 4: 193 - 352.","Oboyski, P. T. 2011. The Systematics, Evolution, and Ecology of Hawaiian Cydia (Lepidoptera: Tortricidae) [Unpublished doctoral dissertation]. University California, Berkeley."]}

Austin, Kyhl A., Rubinoff, Daniel (2023): Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae). Proceedings of the Hawaiian Entomological Society 55: 11-27, DOI: http://doi.org/10.5281/zenodo.8159762, {"references":["Austin, K.A., and D. Rubinoff. 2023. New records of introduced Lepidoptera in the Hawaiian Islands for the year 2022. Bishop Museum Occasional Papers. 148: 175-184.","Banko, P., R. Peck, M. Munstermann, and K. Jaenecke. 2022. Host plant associations of Lepidoptera and implications for forest bird management at Hakalau Forest National Wildlife Refuge. Hawaii Cooperative Studies Unit Technical Report. 104: 1-39.","Davis, C.J., and N.L.H. Krauss. 1966. Recent introductions for biological control. Proceedings of the Hawaiian Entomological Society. 19: 201-207.","Davis, C.J. 1959. Recent introductions for biological control in Hawaii - IV. Proceedings of the Hawaii Entomological Society. 17: 62-66.","Davis, C.J. 1962. Cryptophlebia ombrodelta. Proceedings of the Hawaiian Entomological Society. 18: 2.","Diakonoff, A. 1977. Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen. 158: 1-55.","Evenhuis, N.L., editor. 2007. \"Barefoot on Lava: The Journals and Correspondence of Naturalist R.C.L. Perkins in Hawaiʻi, 1892- 1901.\" Bishop Museum Press, Honolulu. 412 pp.","Gagne, W.C. 1982. Working toward an assessment of the conservation status of Hawaii's endemic arthropods, with emphasis on the moths or Lepidoptera. Proceedings of the 4th Conference on Natural Sciences, Hawaii Volcanoes National Park Science Conference. University of Hawaii, Honolulu. pp. 63-72.","Gagne, W.C. and C.C. Christensen. 1985. Conservation status of native terrestrial invertebrates in Hawaii, pp. 105-126. In Stone, C.P. and Scott, J.M. (eds.), Hawai'i's terrestrial ecosystems: Preservation and Management. University of Hawaii Press, Honolulu.","Haines, W.P., J. Giffin, and D. Foote. 2004. Rediscovery of five species of Omiodes Guenee (Lepidoptera: Crambidae) on Hawai'i Island. Bishop Museum Occasional Papers. 79: 45-49.","Heddle, A.L. 2003. Systematics and Phylogenetics of the Endemic Genus Scotorythra (Lepidoptera:Geometridae) in the Hawaiian Islands [Unpublished doctoral dissertation]. University California, Berkeley.","Hembry, D.H., G. Bennett, E. Bess, I. Cooper, S. Jordan, J. Liebherr, K.N. Magnacca, D.M. Percy, D.A. Polhemus, D. Rubinoff, K.L. Shaw, and P.M. O'Grady. 2021. Insect Radiations on Islands: Biogeographic Pattern and Evolutionary Process in Hawaiian Insects. The Quarterly Review of Biology. 96: 247-296.","Kobayashi, S., C. A. Johns, and A.Y. Kawahara. 2021. Revision of the Hawaiian endemic leaf-mining moth genus Philodoria Walsingham (Lepidoptera: Gracillariidae): its conservation status, host plants and descriptions of thirteen new species. Zootaxa. 4944: 1-175.","Landry, J.-F. 2007. Taxonomic review of the leek moth genus Acrolepiopsis (Lepidoptera: Acrolepiidae) in North America. The Canadian Entomologist. 139: 319-353.","Magnacca, K.N. 2013. On the Locality of \"Kaumuohona.\" Proceedings of the Hawaiian Entomological Society. 45: 65-67.","Marr, K.L. and B.A. Bohm. 1997. A taxonomic revision of the endemic Hawaiian Lysimachia (Primulaceae) including three new species. Pacific Science. 51: 254-287.","Medeiros, M. J. 2009. A revision of the endemic Hawaiian genus Thyrocopa (Lepidoptera: Xyloryctidae: Xyloryctinae). Zootaxa. 2202: 1-47.","Meyrick, E. 1932. Exotic Microlepidoptera. Exotic Microlepidoptera. 4: 193-352.","Miller, S.E. and R.W. Hodges. 1995.Platynota stultana, the Omnivorous Leaf-Roller, Established in the Hawaiian Islands (Lepidoptera: Tortricidae). Bishop Museum Occasional Papers. 42: 36-39.","Nishida, G.M., editor.2002. Hawaiian Terrestrial Arthropod Checklist: Fourth Edition. Hawaii Biological Survey, Bishop Museum Technical Report No. 22. Bishop Museum Press, Honolulu. 313 pp.","Oboyski, P.T. 2011. The Systematics, Evolution, and Ecology of Hawaiian Cydia (Lepidoptera: Tortricidae) [Unpublished doctoral dissertation].University California, Berkeley.","Paxton, E.H., M. Laut, J.P. Vetter, and S.J. Kendall. 2018. Research and management priorities for Hawaiian forest birds. The Condor. 120: 557-565.","Pinzari, C.A., R.W. Peck, T. Zinn, D. Gross, K. Montoya-Aiona, K.W. Brinck, P.M. Gorresen, and F. J. Bonaccorso. 2019. Hawaiian hoary bat (Lasiurus cinereus semotus) activity, diet and prey availability at the Waihou Mitigation Area, Maui. Hawaii Cooperative. Studies Unit Technical Report. 90: 1-68.","Pogue, M.G. 1988. Revision of the genus Lorita Busck (Lepidoptera:Tortricidae: Cochylini), with a description of a new species. Proceedings of the Entomological Society of Washington. 90: 440-457.","Regnier, C., P. Bouchet, K.A. Hayes, N.W. Yeung, C.C. Christensen, D.J. Chung, B. Fontaine, and R.H. Cowie. 2015. Extinction in a hyperdiverse endemic Hawaiian land snail family and implications for the underestimation of invertebrate extinction. Conservation Biology. 29: 1715-1723.","Ronsted N., S.K. Walsh, M. Clark, M. Edmonds, T. Flynn, S. Heintzman, A. Loomis, D. Lorence, U. Nagendra, B. Nyberg, M. Opgenorth, L. Weisenberger, A. Williams, D. Wolkis, K.R. Wood, and M. Keir. 2022. Extinction risk of the endemic vascular flora of Kauaʻi, Hawaiʻi, based on IUCN assessments. Conservation Biology. 36: 1-11.","Swezey, O.H. 1915. New species of Hawaiian Moths. Proceedings of the Hawaiian Entomological Society. 3: 93-97.","Swezey, O.H. 1920. Some new Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 4: 376-386.","Swezey, O.H. 1933. New Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 8: 299-304.","Swezey, O.H. 1940. New species of Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 10: 461-465.","Swezey, O.H. 1946. New species of Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 12: 625-628.","USFWS. 1984. Endangered and Threatened Wildlife and Plants; Review of Invertebrate Wildlife for Listing as Endangered or Threatened Species. Federal Register. 49: 21664-21675.","USFWS. 1996. Endangered and Threatened","Wagner, W. L., D.R. Herbst, and S.H. Sohmer. 1999. Manual of the Flowering Plants of Hawaii. University of Hawaii Press, Honolulu. 1919 pp.","Walsingham, L. 1907. Microlepidoptera. In Sharp, D. (Ed.), Fauna Hawaiiensis. 1(5). Cambridge University Press, Cambridge. pp. 469-759.","Wood, K.R. 2013. Rediscovery of Lysimachia venosa (Wawra) H. St. John on Kaua'i, Hawaiian Islands. Bishop Museum Occasional Papers. 114: 37-38.","Zimmerman, E.C. 1948. Introduction. Insects of Hawaii. 1. The University Press of Hawaii, Honolulu. 206 pp.","Zimmerman, E.C. 1978. Microlepidoptera. Insects of Hawaii. 9. The University Press of Hawaii, Honolulu. 1,903 pp."]}

Nuritamburia phyllanthana (Swezey) Presumed extinct This species is known from the type series of seven specimens reared from Phyllanthus distichus (Phyllanthaceae) from Malamalama Ridge, Oʻahu in 1916 and 1925 (Swezey 1940). No other specimens are known, and targeted collecting near P. distichus on Oʻahu did not result in any specimens. The hostplant has not been encountered frequently by us, although Wagner et al. (1999) considered it “locally common in mesic forest, often on steep slopes or ridge tops, sometimes in dry shrubland” on Kauaʻi, Oʻahu, Molokaʻi, Lānaʻi, West Maui, and rarely on East Maui., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 15, DOI: 10.5281/zenodo.8159762, {"references":["Swezey, O. H. 1940. New species of Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 10: 461 - 465.","Wagner, W. L., D. R. Herbst, and S. H. Sohmer. 1999. Manual of the Flowering Plants of Hawaii. University of Hawaii Press, Honolulu. 1919 pp."]}

Pararrhaptica subsenescens (Walsingham) Presumed extinct This species is known from a single female collected by Perkins in 1893 on Molokaʻi (Walsingham 1907). No further locality data are given on the label, but the date (5 viii 1893) corresponds to a journal entry by Perkins from “near Waikolu” (Evenhuis 2007). We have collected in this general area and along the Hanalilolilo Trail but have not found this species. Like other Pararrhaptica, it likely fed on leaves and shoots of Myrsine spp. (Primulaceae)., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 16, DOI: 10.5281/zenodo.8159762, {"references":["Walsingham, L. 1907. Microlepidoptera. In Sharp, D. (Ed.), Fauna Hawaiiensis. 1 (5). Cambridge University Press, Cambridge. pp. 469 - 759.","Evenhuis, N. L., editor. 2007. \" Barefoot on Lava: The Journals and Correspondence of Naturalist R. C. L. Perkins in Hawaiʻi, 1892 - 1901. \" Bishop Museum Press, Honolulu. 412 pp."]}

Eccoptocera osteomelesana (Swezey) Presumed extinct This species is the most recent native Hawaiian tortricid to be described (Swezey 1946). It has been reared from leaves of Osteomeles anthyllidifolia (Rosaceae) and is known from small series of specimens collected in Mānoa and Wailupe Valley on Oʻahu and was most recently collected on the campus of UH Mānoa in 1969. However, it may have been introduced, as it more closely resembles some Pacific and Australian Eccoptocera than native Hawaiian ones. It is also notable that no other entomologist collected this species prior to F.X. Williams in 1929, despite it having been reared and collected in such heavily urbanized places. No specimens have been collected by us despite extensive searching near large patches of Osteomeles in the southern KoʻolauMountainsandextensivetrapping in areas it might be expected., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 15, DOI: 10.5281/zenodo.8159762, {"references":["Swezey, O. H. 1946. New species of Hawaiian Lepidoptera. Proceedings of the Hawaiian Entomological Society. 12: 625 - 628."]}

Cydia obliqua (Walsingham) Presumed extinct This species is known from three female specimens collected at Hualālai (5000 ft) and Kona (4000 ft) on Hawaiʻi island by R.C.L. Perkins in 1892. Oboyski (2011) considered it possibly just an extreme color form of the widespread and polymorphic C. plicatum (Walsingham), but refrained from synonymizing it because of significant differences in the female genitalia. The collection date given for the two female types from Hualālai (5 viii 1892) corresponds to a journal entry in which Perkins wrote, “there were two Tortricids on Koa” (Evenhuis 2007)., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 14, DOI: 10.5281/zenodo.8159762, {"references":["Oboyski, P. T. 2011. The Systematics, Evolution, and Ecology of Hawaiian Cydia (Lepidoptera: Tortricidae) [Unpublished doctoral dissertation]. University California, Berkeley.","Evenhuis, N. L., editor. 2007. \" Barefoot on Lava: The Journals and Correspondence of Naturalist R. C. L. Perkins in Hawaiʻi, 1892 - 1901. \" Bishop Museum Press, Honolulu. 412 pp."]}

Pararrhaptica punctiferanus (Walsingham) Presumed extinct This species was described from two specimens: the holotype male collected by Perkins in 1894 at 5000 ft on Haleakalā, Maui and a questionably associated female paratype, collected in the same year at 4000 ft elevation on Molokaʻi. The female paratype likely represents an undescribed species, as most Pararrhaptica appear to be single island endemics. Like most other Pararrhaptica, it likely fed on Myrsine spp. (Primulaceae)., Published as part of Austin, Kyhl A. & Rubinoff, Daniel, 2023, Rediscoveries and Presumed Extinctions of Hawaiian Leaf-roller Moths (Lepidoptera: Tortricidae), pp. 11-27 in Proceedings of the Hawaiian Entomological Society 55 on page 16, DOI: 10.5281/zenodo.8159762

Genus Exapate Hübner, 1825 Exapate Hübner, 1825. Type species: Phalaena Tinea gelatella Linnaues, 1761 [= [Phalaena] congelatella Clerck, 1759]. Scinipher Frölich, 1828. Type species: Phalaena congelatella Clerck, 1759. Cheimatophila Stephens, 1829. Preoccupied. Cheimaphasia Curtis, 1833. Type species: Phalaena Tinea gelatella Linnaues, 1761 [= [Phalaena] congelatella Clerck, 1759]. Cheimonophila Duponchel, 1838. Type species: Phalaena Tinea gelatella Linnaeus, 1761 [= [Phalaena] congelatella Clerck, 1759]. Enyphantes Hübner, 1806. Rejected. Exapate congelatella (Clerck, 1759) Phalaena congelatella Clerck, 1759 Phalaena paradoxa Sulzer, 1776 Lithosia gelata Fabricius, 1798 Tortrix gelatana Hübner, [1819] ? Exapate ignotella de Rougemont, 1906 Exapate congelatella ab. kenneli Schille, 1924 Exapate bicuspidella Bruun & Krogerus, 1996 Exapate duratella Heyden, 1864 Exapate aidasi sp. nov. Key to species Based on males 1 Small species (wingspan c. 17–19 mm), hindwings dark greyish; phallus with medial thorn............. E. aidaisi sp. nov. - Larger species (wingspan c. 21–24 mm), hindwings pale grey; phallus with subapical thorn.......................... 2 2 Forewing with brownish-grey ground colour, veins weakly suffused with brown; uncus moderately slender, with small lateral shoulders.......................................................................... E. congelatella (Clerck) - Forewing with greyish ground colour, veins distinctly suffused with grey-brown; uncus broad, with larger lateral shoulders...................................................................................... E. duratella Heyden, Published as part of Huemer, Peter & Mayr, Toni, 2023, A surprising new species of Exapate Hübner, 1825 (Lepidoptera, Tortricidae) from Armenia, pp. 75-82 in Zootaxa 5296 (1) on pages 76-77, DOI: 10.11646/zootaxa.5296.1.7, http://zenodo.org/record/7970231, {"references":["Bruun, H. H. & Krogerus, H. (1996) Exapate bicuspidella sp. n., a new species of Tortricidae from northeastenr Finland (Lepidoptera). Entomologica Fennica, 7, 77 - 86. https: // doi. org / 10.33338 / ef. 83895"]}

A new species of cold-season moths, Exapate aidasi sp. nov., is described from male specimens collected in Armenia. This is also the first record of the genus Exapate in Armenia and south of the Caucasus. The new species differs from the two previously known congeneric taxa both in phenotypic characteristics and in the structure of the male genitalia. In addition, DNA barcodes (cytochrome c-oxidase subunit 1) are clearly divergent from those of other species. Adult and male genitalia of all three species of Exapate are shown for comparison.

Exapate aidasi sp. nov. (Figs 2–4) Type material. Holotype ♁, Armenia, Vedi, Goravan Sands reserve, 39°33.4858′N; 44°43.2149′E, 930 m, 29.10.2021, Duda & Saldaitis leg.; DNA barcode ID TLMF _ Lep _32188 (TLMF). Paratypes: 7 ♁, Armenia, Areni, Novarank road, 39°41.8978′N; 45°12.6143′E, 1240 m, 5.10.2021, Duda & Saldaitis leg.; genitalia slide TOR 486 ♁ (RCTM, TLMF). 1 ♁, Armenia, near Garni, Azat riv. valley, 40°06.5280′N; 44°43.4364′E, 1200 m, 6.10.2021, Duda & Saldaitis leg. (RCTM). Diagnosis. The new species differs from both congeners externally by its smaller size and the darker grey hindwings, but to some extent also by the wing pattern (Figs 2–3). In E. congelatella the forewings are rather light brown to brownish grey, usually with well defined markings, whereas in E. duratella the darker coloured veins of the terminal part of the forewing are characteristic. Both species are distinctly larger on average (wingspan c. 21–23 mm in E. congelatella, c. 24–25 mm in E. duratella, c. 17–19 mm in E. aidasi sp. nov.) and the hindwings are light grey to grey-brown. Further, E. aidasi sp. nov. differs from the related species by subtle but diagnostic characters of the male genitalia, particularly the narrower uncus and valva and the absence of a subapical thorn of the pallus and the presence of a medial thorn (Figs 4–6). Finally, the DNA barcode sequence is clearly divergent at a distance of c. 2.5%. E. congelatella and E. duratella are extremely similar in genitalia and hardly discernible (Bruun & Krogerus 1996, Razowski 2002). Male genitalia differ slightly by the shape of the uncus. Description. Adult ( Figs 2–3). Head, thorax, tegulae and abdomen grey, frons and neck covered with brushlike hairy scales, labial palpi well developed, turned upwards; antenna dark grey, scape scaled, pedicellus and flagellomeres ciliated, with cilia of about half the length of the width of the flagellomeres; wingspan 17.3–19.2 mm; forewing grey with irregular dark grey mottling, particularly in the discal and discocellular area, on veins Sc and R 1, and in the terminal area; base of fringes with dark grey dots at end of veins, basal cilia line light grey-brown, distal part darker grey; hindwings dark greyish, cilia light grey brown and darker grey with darker basal line. Variation. The extent of the darker markings on the forewing varies considerably and is occasionally largely reduced. Male genitalia (Fig. 4). Uncus short and with a broad base about two-fifths the width of tegumen, medially abruptly widened with gradually constricted distal half; socii small, haired; gnathos medially short, ovate, lateral arms long, with short thorns; transtilla well developed, band-like, densely covered with medium-sized stiff setae in middle; juxta subtriangular, with long digitate appendices posteriorly; valva long and moderately slender, sacculus more strongly sclortised, with subbasal emargination and short finger-like appendix in middle; phallus long, slender, apically evenly pointed, with small postmedial thorn. Female genitalia. Unknown. Biology. Adults have been collected in early and late October at light. Host-plant and early stages are unknown, however, the congeneric species mainly feed on woody plants and the new species probably also feeds on such. One of the localities is a semi-desert with only a few woody plants which may be potential host-plants, e.g. Prunus incana or Rhamnus pallassii. Larix, the main host-plant for E. duratella according to Deutsch (2008), is not present in Armenia (Schmidt 1995). Distribution. Armenia. E. congelatella is widespread in temperate Europe, extending its eastern range to Ural mts. (Sinev et al. 2019) and allegedly Kazakhstan (Razowski 2002). E. duratella is restricted to European mountain systems: Alps, Pyrenees, Carpathians (Karle-Fendt and Wolf 2015). Etymology. The new species is named in honour of Aidas Saldaitis, co-collector of the type series, in recognition of his contribution in exploring Lepidoptera in many remote areas., Published as part of Huemer, Peter & Mayr, Toni, 2023, A surprising new species of Exapate Hübner, 1825 (Lepidoptera, Tortricidae) from Armenia, pp. 75-82 in Zootaxa 5296 (1) on pages 78-81, DOI: 10.11646/zootaxa.5296.1.7, http://zenodo.org/record/7970231, {"references":["Bruun, H. H. & Krogerus, H. (1996) Exapate bicuspidella sp. n., a new species of Tortricidae from northeastenr Finland (Lepidoptera). Entomologica Fennica, 7, 77 - 86. https: // doi. org / 10.33338 / ef. 83895","Razowski, J. (2002) Tortricidae (Lepidoptera) of Europe. Vol. 1. Tortricinae and Chlidanotinae. F. Slamka, Bratislava, 247 pp.","Deutsch, H. (2008) Exapate duratella Heyden 1864: Beitrag zur Biologie und Zuchtbericht (Lepidoptera, Tortricidae). Carinthia II, 198 / 118, 351 - 356.","Schmidt, W. C. (1995) Around the world with Larix: an introduction. In: Schmidt, W. C. & McDonald, K. J. (Eds.), Ecology and management of Larix forests: a look ahead. U. S. A. Gen. Tech. Rep. INT-GTR- 319. Intermountain Research Station, Ogden, Utah, pp. 6 - 18.","Sinev, S. Yu., Nedoshivina, S. V. & Dubatolov, V. V. (2019) Tortricidae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 120 - 156. [in Russian]","Karle-Fendt, A. & Wolf, W. (2015) Ein Vorkommen von Exapate duratella Heyden, 1864 in den Allgauer Hochalpen (Insecta: Lepidoptera: Tortricidae). Beitrage zur bayerischen Entomofaunistik, 15, 39 - 42."]}

Anthozela Meyrick, 1913 Anthozela Meyrick, 1913: 280. Type species: Anthozela chrysoxantha Meyrick, 1913. Gephyroneura Obraztsov, 1968: 181. Type species: Laspeyresia hemidoxa Meyrick, 1907. Preoccupied. Mehteria Koçak, 1981: 117. Replacement name for Gephyroneura., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Meyrick, E. (1913) Descriptions of South African Micro-lepidoptera. Annals of the Transvaal Museum, 3, 267 - 336. https: // doi. org / 10.5962 / bhl. title. 9702","Obraztsov, N. S. (1968) Descriptions and records of south Asiatic Laspeyresiini (Lepidoptera: Tortricidae). Journal of the New York Entomological Society, 76, 176 - 192.","Meyrick, E. (1907) Descriptions of Indian Micro-lepidoptera Lepidoptera I - XVIII. Journal of the Bombay Natural History Society, 18, 137 - 160.","Kocak, A. O. (1981) On the nomenclature of some genera of Lepidoptera. Priamus, 1 (3), 112 - 122."]}

Semnostola mystica Diakonoff, 1959 Semnostola mystica Diakonoff, 1959: 175. TL: India, Burma [Myanmar]. TD: Holotype (male), NHRS. Material examined. 1 ♁, Hualien Hsien, Losao, 1110 m, 2-IV-1984, A. Kawabe (Sun-137 ♁, USNM122510, A. Kawabe); 1 ♁, NanTow Co., Mei-feng, 30 km S Tayuling, 2200 m, 1-8-VI-1980, D. R. Davis. [USNM]. 1 ♁, Hsinchu /Miaoli Kuangwu, 2000 m, 24-25-VI-1985, J. Heppner & H. Wang; 5 ♁, Hsinchu Co. Kuangwu For. Sta., 2000 m, 18-25-VIII-1988, J. Heppner & H. Wang (Gen. Sl. Bae UFL-014 ♁); 1 ♁, Taichung Co. Chingshan, 1100 m, 27-III/ 1-IV-1990, J. Heppner & H. Wang; 3 ♁, Taichung Co. Chingshan, 1100 m, 8-11-V-1989, J. Heppner & H. Wang (TORT12045 ♁, FL226) [FSCA]. Distribution. China (Taiwan), India., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 373, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Diakonoff, A. (1959) Entomological results from the Swedish Expedition 1934 to Burma and British India, Lepidoptera collected by Rene Malaise. Microlepidoptera II. Arkiv for Zoologi, 12, 165 - 182."]}

Taiwancylis Razowski, 2000 Taiwancylis Razowski, 2000: 326. Type species: Taiwancylis cladodium Razowski, 2000., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 375, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Razowski, J. (2000) Tortricidae (Lepidoptera) collected in Taiwan, with description of one new genus and eight new species, and a comparison with some regional faunas. Zoological Studies, 39 (4), 319 - 327."]}

Pseudacroclita hapalaspis (Meyrick, 1931) Acroclita hapalaspis Meyrick, 1931: 144. TL: Japan (Honshu, Wakayama Prefecture, Hasimoto). TD: Holotype (male), BMNH. Pseudacroclita hapalaspis (Meyrick): Kawabe, 1992: 108. Material examined. 1 ♀, Suiryuto, 11-X-1935, S. Issiki; 1 ♁, Sozan, 10-VI-1934, S. Issiki (Sun-135 ♁, USNM144421, YH Sun); 1 ♀, Baibara, 25-III-43, S. Issiki (Sun- 136 ♀); 1 ♀, Kirai, 2-VI-1943, S. Issiki; 1 ♁, Kansirei, 18-X-1934, S. Issiki; 1 ♁, Taiwan, III-1929, S. Issiki; 1 ♀, Nantou Hsien, Yushih, 1700 m, 3-4-VIII-1987, A. Kawabe. [USNM]. 1 ♀, Taitung Co. Bishan Hot Spgs., 720 m, 26-29-X-1984, J. Heppner & H. Wang (TORT12020 ♀, FL175) [FSCA]. Distribution. China (Fujian, Hubei, Hunan, Guangdong, Guizhou, Taiwan), Japan., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 373, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109."]}

Ancylis Hübner, [1825] 1816 Ancylis Ḩbner, [1825] 1816: 376. Type species: Tortrix harpana Hübner, [1796−1799]. Epicharis Ḩbner, [1825] 1816: 376. Type species: Tortrix derasana Hübner, [1811−1813]. Preoccupied. Anchylopera Stephens, 1829: 47. Type species: Pyralis lundana Fabricius, 1776. Phoxopteris Treitschke, 1829: 232. Type species: Tortrix siculana Hübner, [1796−1799]. Anticlea Stephens, 1834: 113. Type species: Tortrix harpana Hübner [1796−1799]. Preoccupied. Philalcea Stephens, 1835: 396. Type species: Tortrix harpana Hübner [1796−1799]. Replacement name for Anticlea. Phoxopteryx Sodoffsky, 1837: 93, unjustified emendation Phoxopteris. Sideria Guenée, 1845: 156. Type species: Tortrix achatana [Denis & Schifferm̧ller, 1775]. Ancylopera Agassiz, 1848: 61. Unjustified emendation of Anchylopera. Siderea Stainton, 1859: 196. Misspelling of Sideria. Palaeobia Meyrick, 1881: 660. Type species: Palaeobia hibbertiana Meyrick, 1881. Lamyrodes Meyrick, 1910: 182. Type species: Lamyrodes phileris Meyrick, 1910. Phoxopteria Walcott, 1923: 201. Misspelling of Phoxopteris. Ancyloides Diakonoff, 1982: 63. Type species: Ancylis stenampyx Diakonoff, 1982. Preoccupied, subgenus of Ancylis. Remarks. No species had been recorded in Taiwan prior to this study., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 370, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Treitschke, F. (1829) Schaben-Geistchen. In: Ochsenheimer, F. (Ed.), Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer'schen Werks). Vol. 7. Gerhard Fleischer, Leipzig. 1 - 252.","Stephens, J. F. (1834) s. n. In: Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Vol. 4. Baldwin and Cradock, London, pp. 1 - 433.","Stephens, J. F. (1835) s. n. In: Illustrations of British Entomology; or a synopsis of indigenous insects, Mandibulata. Vol. 5. Baldwin and Cradock, London, pp. 369 - 447, pls. 24 - 27.","Sodoffsky, W. (1837) Etymologische Untersuchungen ueber die Gattungsnamen der Schmetterlinge. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 10 (6), 76 - 97. https: // doi. org / 10.5962 / bhl. title. 9388","Guenee, A. M. (1845) Essai sur une nouvelle classification des Microlepidopteres et catalogue des especes europeennes connues jusqu a jour. Annales de la Societe Entomologique de France, 3 (2), 105 - 192, 207 - 344.","Agassiz, L. (1848) Nomenclatoris zoologici index universalis. Continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum. Sumtibus et typis Jent et Gassmann, Soloduri, X + 1135 pp. https: // doi. org / 10.5962 / bhl. title. 1819","Stainton, H. T. (1859) A manual of British butterflies and moths, London, 2, 1 - 475.","Meyrick, E. (1881) Descriptions Australian Micro-Lepidoptera VI. Tortricina contimued. Proceedings of the Linnean Society of New South Wales, 6, 629 - 706. https: // doi. org / 10.5962 / bhl. part. 11888","Meyrick, E. (1910) Revision of Australian Tortricina. Proceedings of the Linnean Society of New South Wales, 35, 139 - 294. https: // doi. org / 10.5962 / bhl. part. 25545","Diakonoff, A. (1982) On a collection of some families of Microlepidoptera from Sri Lanka (Ceylon). Zoologische Verhandelingen, 193, 1 - 124."]}

Neoanathamna Kawabe, 1978 Neoanathamna Kawabe, 1978: 182. Type species: Neoanathama cerinus Kawabe, 1978. Remarks. No species had been recorded in Taiwan prior to this study., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 372, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kawabe, A. (1978) Descriptions of three new genera and fourteen new species of the subfamily Olethreutinae from Japan (Lepidoptera, Tortricidae). Tinea, 10 (19), 173 - 191."]}

Enarmonodes Danilevsky et Kuznetzov, 1968 Enarmonodes Danilevsky et Kuznetzov, 1968: 584. Type species: Grapholitha recreantana Kennel, 1900., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Danilevsky, A. S. & Kuznetzov, V. I. (1968) Tortricidae, Tribe Laspeyresiini. Fauna SSSR. Zoologicheskii Institut Akademii Nauk SSSR, Leningrad, 5 (1), 1 - 635.","Kennel, J. (1900) Neue palaarktische Tortriciden, nebst Bemerkungen uber einige bereits beschriebene Arten. Deutsche entomologische Zeitschrift Iris, 13, 124 - 160."]}

Anthozela hemidoxa (Meyrick, 1907) Laspeyresia hemidoxa Meyrick, 1907: 145. TL: India, Assam, Khasi Hills. TD: Lectotype (male), BMNH. Gephyroneura hemidoxa (Meyrick): Kawabe, 1992: 108. Mehteria hemidoxa (Meyrick): Brown, 2005: 416. Anthozela hemidoxa (Meyrick): Gilligan, 2018, in T@RTS. Material examined. 1 ♀, Tipon, 4-VII-1927, S. Issiki (Sun- 130 ♀); 1 ♀, Taihoku, 10-VII-1933, S. Issiki; 1 ♀, Taiwan: Taihoku, 6-V-1935, S. Issiki. [USNM]. 1 ♀, Ilan Co. Chilan Nsy., Tuchan [Chilan], 480 m, 15-17-X-1984, J. Heppner & H. Wang (TORT12043 ♀, FL207) [FSCA]. Distribution. China (Taiwan), India., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Meyrick, E. (1907) Descriptions of Indian Micro-lepidoptera Lepidoptera I - XVIII. Journal of the Bombay Natural History Society, 18, 137 - 160.","Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109.","Brown, J. W. (2005) Tortricidae (Lepidoptera). World Catalogue of Insects. Vol. 5. Apollo Books, Stenstrup, 741 pp.","Gilligan, T. M., Baixeras, J. & Brown, J. W. (2018) T @ RTS: Online World Catalogue of the Tortricidae. Version 4.0. Available from: http: // www. tortricidae. com / catalogue. asp (accessed 31 Mar 2023)"]}

Kennelia albifacies (Walsingham, 1900) Lipsotelus albifacies Walsingham, 1900: 570. TL: India. TD: Holotype (unknown), OUM. Argyroploce corthyntis Meyrick, 1909: 596. TL: Sri Lanka. Ceylon [Sri Lanka] (Patipola). TD: Lectotype (male), BMNH. Kennelia albifacies (Walsingham): Kawabe, 1992: 108. Material examined. 7 ♁, 2 ♀, NanTow Co. Mei-feng, 30 km S Tayuling, 2200 m, 1-8-VI-1980, D. R. Davis (Sun-164 ♁, USNM144436, YH Sun; Sun- 165 ♀, USNM144437, YH Sun); 1 ♁, NanTow Co. Tayuling, 2500 m, 9-18-VI-1980, D. R. Davis; 1 ♁, 4 ♀, Chiai Hsien, Fenchihu, 1400 m, 11-12-VIII-1985, A. Kawabe; 1 ♁, 3 ♀, Nantou Hsien, Lushan spa, 1200 m, 3-5-VIII-1985, A. Kawabe; 1 ♁, Wulai, Taipei Hsien, Formosa, 1-VIII-1974, H. Nakajima; 1 ♀, Wulai, Taipei Hsien, Formosa, 6-VIII-1974, H. Nakajima; 1 ♁, (Near Lushan) Nantou Hsien, 1200 m, 28-VII-1977, T. Shiraki; 2 ♁, Taiwan, S. Issiki; 2 ♁, Higasinoko, 3-VI-1943, S. Issiki; 1 ♁, Musya, 6-VI-1943, S. Issiki; 1 ♁, Raisya, 22-X-1934, S. Issiki. [USNM]. 1 ♁, 2 ♀, Taipei Co. Wulai, 200 m, 17-19-VI-1985, J. Heppner & H. Wang; 3 ♁, 1 ♀, Taichung Co. Chingshan, 1100 m, 31-VIII / 4-IX-1988, J. Heppner & H. Wang (Gen. Sl. Bae UFL-116 ♁); 2 ♁, Taoyuan Co. Upper Palin, 1500 m, 7-9-VII-1985, J. Heppner & H. Wang; 2 ♁, 1 ♀, Hsinchu /Miaoli Kuangwu, 2000 m, 24-25-VI-1985, J. Heppner & H. Wang; 1 ♁, Nantou Co. Tungpu, 1150 m, 1-VII-1985, J. Heppner & H. Wang [FSCA]. Distribution. China (Taiwan), India, Sri Lanka., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on pages 371-372, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Meyrick, E. (1909) Descriptions of Indian Micro-lepidoptera XI. Journal of the Bombay Natural History Society, 19, 582 - 607.","Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109."]}

Ancylis unculana (Haworth, 1811) Tortrix unculana Haworth, 1811: 453. TL: United Kingdom, Great Britain. TD: Syntype (s), BMNH. [Tortrix] dersana Ḩbner, [1811-1813]: pl. 32, fig. 206. TL: Europe. TD: Syntype (s), unknown. Ancylis unculana (Haworth): Kennel, 1916: 437. Material examined. 2 ♁, Nantou Hsien, Lushan spa, 1200 m, Aug. 9-10, 1987, A. Kawabe; 1 ♁, Hualien Hsien, Tayuling, 2560 m, Aug. 2-3, 1987, A. Kawabe; 1 ♁, Nantou Hsien, Lushan spa, 1200 m, July 27-29, 1983, A. Kawabe. [USNM]. 1 ♁, Kaohsiung Co. Shanpin For. Sta., [nr.] Liukuei, 750 m, 16-23-III-1990, J. Heppner & H. Wang (TORT12029 ♁, FL196); 1 ♁, 1 ♀, Taipei Co. Wulai, 200 m, 17-19-VI-1985, J. Heppner & H. Wang (Gen. Sl. Bae UFL-082 ♁; TORT12030, no abdomen) [FSCA]. Host plants. Ericaceae: Vaccinium vitis-idaea; Rosaceae: Prunus spinosa, Rubus sp.; Salicaceae: Populus sp.; Leguminosae: Pisum sp., Frangula alnus; Rhamnaceae: Rhamnua frangula, R. cathartica; Betulaceae: Corylus sp. (Bradley et al. 1979, Liu et al. 2002). Distribution. China (Anhui, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Hunan, Jilin, Shaanxi, Sichuan, Tianjin, Taiwan), Korea, Japan, Iran, Russia, Kazakhstan, Turkey, Europe. (Razowski, 2003) Remarks. This species is recorded for the first time in Taiwan., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 370, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Haworth, A. H. (1811) s. n. In: Lepidoptera Britannica, sistens digestionem novam Lepidopterorum quae in Magna Britannia reperiuntur. Part 3. Richard Taylor and Co., London, pp. 377 - 512.","Kennel, J. (1916) Die Palaearktischen Tortriciden. Zoologica, 21 (54), 398 - 545, pls 17 - 20.","Bradley, J. D., Trenewan, W. G. & Smith, A. (1979) British Tortricid moths. Tortricidae: Olethreutinae. The Ray Society, London, 36 pp., pls. 22 - 43.","Liu, Y. Q. & Li, G. W. (2002) Fauna Sinica, Insect. Vol. 27. Lepidoptera, Tortricidae. Science Press, Beijing, 463 pp.","Razowski, J. (2003) Tortricidae of Europe. Vol. 2. Olethreutinae. Frantisek Slamka, Krakow-Bratislava, 301 pp."]}

Kennelia Rebel, 1901 Anomalopteryx Kennel, 1900: 157. Type species: Anomalopteryx xylinana Kennel, 1900. Preoccupied. Kennelia Rebel, 1901: 263. Type species: Anomalopteryx xylinana Kennel, 1901., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Rebel, H. (1901) Catalog der Lepidopteren des palaearctischen Faunengebietes. Vol. 2. R. Friedlander & Sohn, Berlin, 1 - 368.","Kennel, J. (1900) Neue palaarktische Tortriciden, nebst Bemerkungen uber einige bereits beschriebene Arten. Deutsche entomologische Zeitschrift Iris, 13, 124 - 160."]}

Pseudacroclita Oku, 1979 Pseudacroclita Oku, 1979: 591. Type species: Acroclita hapalaspis Meyrick, 1931., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 373, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Oku, T. (1979) Acrolita and some allied genera (Lepidoptera: Tortricidae) from Japan, with descriptions of new taxa. Kontyu, 47 (4), 586 - 592."]}

Eleven genera and thirteen species of the tortricid tribe Enarmoniini documented from Taiwan are listed. One species, Semnostola tristrigma sp. nov., is described as new to science. Two species, Ancylis unculana (Haworth, [1811]) and Neoanathamna pallens Kawabe, 1980 along with their genera, are newly recorded for Taiwan. Images of the adults and genitalia are provided for the new species.

Enarmoniodes furcula Kuznetzov, 1973 Enarmoniodes furcula Kuznetzov, 1973: 694. TL: China (Taiwan). TD: MGAB. Distribution. China (Taiwan)., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kuznetzov, V. I. (1973) Descriptions of east-Asian Tortricids of the subfam. Olethreutinae (Lepidoptera, Tortricidae). Entomologicheskoe Obozrenie, 52 (3), 682 - 699."]}

Enarmonodes aeologlypta (Meyrick, 1936) Laspeyresia aeologlypta Meyrick, 1936: 615. TL: Japan, Honshu, Yumoto. TD: Holotype (male), BMNH. Enarmonodes aeologlypta (Meyrick): Kawabe, 1992: 108. Material examined. 1 ♀, Hualien Co. Kuyuan, Taroko Gorge, 750 m, 6-7-VII-2005, J. Heppner & H. Wang (TORT5634 ♀); 1 ♀, Hsinchu /Miaoli Kuangwu, 2000 m, 24-25-VI-1985, J. Heppner & H. Wang (TORT11871 ♀, FL101) [FSCA]. Distribution. China (Taiwan), Japan., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109."]}

Loboschiza Diakonoff, 1968 Loboschiza Diakonoff, 1968: 93. Type species: Argyroploce clytocarpa Meyrick, 1920. Rhadinoscolops Obraztsov, 1968: 187. Type species: Pyralis koenigiana Fabricius, 1775., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 372, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Diakonoff, A. (1968) Microlepidoptera of the Philippine Islands. Bulletin of the United States National Museum, 257, 1 - 484, 846 figs. https: // doi. org / 10.5479 / si. 03629236.257.1","Obraztsov, N. S. (1968) Descriptions and records of south Asiatic Laspeyresiini (Lepidoptera: Tortricidae). Journal of the New York Entomological Society, 76, 176 - 192.","Fabricius, J. C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species adiectis synonymis, locis, descriptionibus, observationibus. Korte, Flensburgi and Lipsiae, xxx + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510"]}

Taiwancylis cladodium Razowski, 2000 Taiwancylis cladodium Razowski, 2000: 326. TL: Taiwan (Kaohsiung, Shanpin). TD: Holotype (male), CMNH. Distribution. China (Taiwan)., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 375, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Razowski, J. (2000) Tortricidae (Lepidoptera) collected in Taiwan, with description of one new genus and eight new species, and a comparison with some regional faunas. Zoological Studies, 39 (4), 319 - 327."]}

Enarmoniodes Ghesquièe, 1940 Enarmoniodes Ghesquièe, 1940. Type species: Enarmoniodes mirabilis Ghesquièe, 1940., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 371, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346

Loboschiza koenigiana (Fabricius, 1775) Pyralis koenigiana Fabricius, 1775: 653. TL: India. TD: Syntype (s) (unknown), unkonwn. Pyralis koenigana Fabricius, 1787: 237. no type. Hemerosia aurantiana Pryer, 1877: 235. TL: China [Chekiang, Shanghai]. TD: Holotype (female), BMNH. Eucelis vulnerata Walsingham, 1900: 571. TL: India. Burma [Myanmar] (Tenasserim). TD: Holotype (unknown), OUM. Grapholitha delectana Snellen, 1902: 72. TL: Indonesia (Java, Tegal). TD: Syntypes (unknown), NCB. Loboschiza koenigiana (Fabricius): Kuznetzov, 1988: 74. Rhadinoscolops koenigiana: Liu et Li, 2002: 367. Material examined. 1 ♁, Nantou Hsien, Lushan spa, 1200 m, 9-10-VIII-1987, A. Kawabe; 1 ♁, 1 ♀, Hualien Hsien, Hungyeh spa, 200 m, 29-30-III-1984, Y. Shibata; 1 ♀, Taiwan, 19-VIII-1933, S. Issiki; 1 ♀, Taihoku, 17-XII-1934, S. Issiki (Sun- 134 ♀); 1 ♁, Ranrum, 8-VII-1925, S. Issiki. [USNM]. 2 ♁, Pingtung Co. Kenting Park, 50 m, 1-5-IX-1983, J. Heppner; 2 ♁, Pingtung Co. Kenting Park, 50 m, 29-31-VIII-1983, J. Heppner; 2 ♁, Ilan Co. Chilan Nsy., Tuchan [Chilan], 480 m, 15-17-X-1984, J. Heppner & H. Wang; 1 ♁, Nantou Co. 5 Km N. Chitou, 500 m, 29- VI-1985, J. Heppner & H. Wang; 1 ♁, Pingtung Co. Kenting Park, 50 m, 31-X/ 1-XI-1984, J. Heppner & H. Wang; 2 ♁, Pingtung Co. Kenting Park, 255 m, 23-28-IV-1989, J. Heppner & H. Wang; 1 ♁, Hualien Co. Tienhsiang, 435 m, 25-VIII-1983, J. Heppner (Gen. Sl. Bae UFL-065 ♁); 1 ♀, Pingtung Co. Kenting Park, 255 m, 23-28-IV-1989, J. Heppner & H. Wang [FSCA]. Host plants. Meliaceae: Melia azedarach. (Liu et al. 2002). Distribution. China (Anhui, Heilongjiang, Henan, Hubei, Hunan, Fujian, Guangdong, Guangxi, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang, Taiwan), India, Indonesia, Korea, Japan, Sri Lanka, Pakistan, New Guinea, Australia., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 372, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Fabricius, J. C. (1775) Systema entomologiae, sistens insectorum classes, ordines, genera, species adiectis synonymis, locis, descriptionibus, observationibus. Korte, Flensburgi and Lipsiae, xxx + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510","Fabricius, J. C. (1787) Mantissa Insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus. Vol. 2. Christ. Gottl. Proft, Hafniae, 382 pp.","Pryer, W. B. (1877) Descriptions of new species of Lepidoptera from North China. Cistula Entomologica, 2 (18), 231 - 236.","Snellen, P. C. T. (1902) Beschrijvingen van nieuwe exotische Tortricinen, Tineinen en Pterophorinen benevens aanteekeningen over reeds bekend gemaakte soorten, II. Tijdschrift voor Entomologie, 44, 67 - 89.","Kuznetzov, V. I. (1988) New species of Tortricid moths of subfamily Olethreutinae (Lepidoptera, Tortricidae) of the fauna of North Vietnam. Entomologicheskoe Obozrenie, 67, 615 - 631.","Liu, Y. Q. & Li, G. W. (2002) Fauna Sinica, Insect. Vol. 27. Lepidoptera, Tortricidae. Science Press, Beijing, 463 pp."]}

Semnostola Diakonoff, 1959 Semnostola Diakonoff, 1959: 174. Type species: Semnostola mystica Diakonoff, 1959. Ancyloides Kuznetzov, 1964: 882. Type species: Eucosmomorpha (Ancyloides) magnifica Kuznetzov, 1964. Subgenus of Eucosmomorpha., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 373, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Diakonoff, A. (1959) Entomological results from the Swedish Expedition 1934 to Burma and British India, Lepidoptera collected by Rene Malaise. Microlepidoptera II. Arkiv for Zoologi, 12, 165 - 182.","Kuznetzov, V. I. (1964) New genera and species of leaf-rollers (Lepidoptera: Tortricidae) from the Far-East. Entomologicheskoe obozrenie, 43, 873 - 889. [in Russian]"]}

Tetramoera schistaceana (Snellen, 1890) Grapholitha schistaceana Snellen, 1890: 95. TL: Indonesia (Java, Tegal). TD: Syntypes (unknown), RMNH. Tetramoera schistaceana (Snellen): Kawabe, 1992: 108. Material examined. 1 ♀, Heito, 24-VII-1934, Yamaho (Sun- 138 ♀, USNM144422, YH Sun) [USNM]. Distribution. China (Guangdong, Taiwan), Vietnam, Indonesia, Philippine, Madagascar, Micronesia, Hawaii., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 375, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Snellen, P. C. T. (1890) Aanteekeningen over Lepidoptera schadelijk voor het suikerriet. Mededelingen van het Profstation suikerbieteelt W-Java, Kagok-Tegal, 1, 94 - 97.","Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109."]}

Kennelia protocyma (Meyrick, 1936) Argyroploce protocyma Meyrick, 1936: 613. TL: Taiwan, Formosa [Taiwan] (Pinansyusan, Hinode). TD: Holotype (female), BMNH. Kennelia protocyma (Meyrick): Kawabe, 1992: 108. Material examined. 3 ♀, Taiwan, S. Issiki (Sun- 166 ♀, USNM144438, YH Sun); 1 ♀, NanTowCo. Tayuling, 2500 m, 9-18-VI-1980, D. R. Davis [USNM]. Distribution. China (Taiwan)., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 372, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kawabe, A., Komai, F. & Razowski, J. (1992) Tortricoidea. In: Heppner, J. B. & Inoue, H. (Eds.), Lepidoptera of Taiwan. Vol. 1. Part 2. Checklist. Scientific Publishers, Gainesville, Florida, pp. 103 - 109."]}

Semnostola tristrigma sp. nov. (Figure 1) Type material. CHINA, Yunnan: Holotype ♁, Botanical Garden of Xishuangbanna South Medicine, Jinghong City, 585 m, 18-IV-1995, leg. Hongjiang Wang, genitalia slide no. YHL0244 (NKU). Paratypes: 5 ♁, 1 ♀, Kaohsiung Co. Dona, 16 Km NE. Maolin, 500 m, 3-XI-1984, J. Heppner & H. Wang; 3 ♁, Pingtung Co. Kenting Park, 255 m, 23-28-IV-1989, J. Heppner & H. Wang, genitalia slides nos. FL220 (♁), FL122 (♀), FL145 (♁), FL146 (♁), FL176 (♁) [FSCA]. Diagnosis. The new species is similar to S. grandaedeaga Zhang et Wang, 2006 (Zhang & Wang 2006) in male genitalia. It can be distinguished from the latter in the forewing by the distinct three dark brown fasciae, tegumen with terminal subrectangular portion in the male genitalia, and the corpus bursae with two signa in the female genitalia. In S. grandaedeaga, the forewing has one distinct dark brown fascia, tegumen has terminal triangular portion. Description. Adult (Fig. 1a). Wing expanse 10.5–11.5 mm. Vertex and forehead dark brown. Antenna brown. Labial palpus yellowish brown mixed with black scales, second segment 1.5 times length of diameter of compound eyes, third segment about 1/5 length of second segment. Thorax and tegula brown. Forewing ground color brownish black on upperside, mixed with yellowish brown scales; subbasal, median and postmedian fasciae dark brown, extending obliquely outward from costal margin; costa with nine white strigulae from base to vein R 4, strigulae 1 and 2 indistinct, strigulae 3 and 4 between subbasal and median fasciae, strigulae 5 and 6 appearing as two marks, pairs of strigulae 7–9 distributed between veins R 1 -R 2, R 2 -R 3 and R 3 -R 4, respectively, becoming confluent and extending obliquely to R 5 and M 1 at termen; cilia brownish black. Hindwing and cilia brownish black. Fore- and midlegs yellowish brown, tarsi with black rings; hindleg yellowish brown. Abdomen with upperside dark brown and underside brown. Male genitalia (Fig. 1b): Tegumen high, with terminal subrectangular portion. Uncus absent. Socius with long hairs, terminal end round. Valva with basal half broad, narrow to distal half; costa nearly straight and with long hairs distally; basal cavity about half length of valva; sacculus moderately setose; cucullus subrectangular with a dorsl pollex bearing a stout spine. Aedeagus tapering from base to apex, strongly sclerotized, length 0.8 times of valva; cornuti absent. Female genitalia (Fig. 1c): Papilla analis hairy, with anterior half narrower than posterior half. Apophyses posteriores slightly shorter than apophyses anteriores. Sterigma broadly U-shaped. Antrum distinct, about 1/5 length of ductus bursae. Ductus bursae as long as corpus bursae, widened toward corpus bursae anteriorly; ductus seminalis from anterior 2/5 of ductus bursae. Corpus bursae broadly pear-shaped, with two round signa placed near opening of corpus bursae. Etymology. The specific epithet is derived from the Latin tri - (= three) and - strigma (= stripe), indicating three dark brown stripes distributed at the subbasal fasciae, median fascia and postmedian fascia respectively. Distribution. China (Taiwan, Yunnan)., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on pages 373-375, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Zhang, X. & Wang, S. X. (2006) Systematic Study of the genus Kennelia Rebel from China (Lepidoptera, Tortricidae, Olethreutinae). Acta Zootaxonomica Sinica, 31 (3), 613 - 616."]}

Neoanathamna pallens Kawabe, 1980 Neoanathamna pallens Kawabe, 1980: 22. TL: Japan (Shikoku, Kagawa Prefecture, Shionoe, Hitotsu-uchu). TD: Holotype (male), USNM. Material examined. 1 ♁, Taiwan: Raisya, Nov. 23, 1934, S. Issiki; 1 ♁, Taiwan: Taihoku, June 16, 1946, S. Issiki; 1 ♁, Taiwan: Taihoku, June 18, 1946, S. Issiki. [USNM]. 1 ♁, 2 ♀, Hualien Co. Fuyan, 400 m, 7-III-1990, J. Heppner & H. Wang (Gen. Sl. Bae UFL-064 ♁); 2 ♁, 2 ♀, Taipei Co. Wulai, 200 m, 17-19-VI-1985, J. Heppner & H. Wang (TORT11872 ♁, FL071; TORT11873 ♀, FL072); 1 ♁, Taichung Co. Chingshan, 1100 m, 31-VIII/ 4-IX-1988, J. Heppner & H. Wang; 1 ♀, Kaohsiung Co. Shanpin For. Sta., [nr.] Liukuei, 750 m, 16-23-III-1990, J. Heppner & H. Wang; 1 ♀, Ilan Co. Fushan For. Sta., 650 m, 4-11-IV-1990, J. Heppner; 1 ♀, Ilan Co. 5 Km N. Nanao, 100 m, 11- 13-X-1984, J. Heppner & H. Wang [FSCA]. Distribution. China (Hunan, Taiwan), Japan. Remarks. This species is recorded for the first time in Taiwan., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on pages 372-373, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Kawabe, A. (1980) Descriptions of fourteen new species of the micro moths from Japan (Lepidoptera: Tortricidae, Cochylidae, Carposinidae). Tinea, 11 (3), 17 - 31."]}

Tetramoera Diakonoff, 1968 Tetramoera Diakonoff, 1968: 68. Type species: Grapholitha schistaceana Snellen, 1890., Published as part of Sun, Yinghui & Li, Houhun, 2023, Checklist of the tribe Enarmoniini Diakonoff, 1953 (Lepidoptera: Tortricidae) of Taiwan, with description of one new species, pp. 369-377 in Zootaxa 5278 (2) on page 375, DOI: 10.11646/zootaxa.5278.2.9, http://zenodo.org/record/7906346, {"references":["Diakonoff, A. (1968) Microlepidoptera of the Philippine Islands. Bulletin of the United States National Museum, 257, 1 - 484, 846 figs. https: // doi. org / 10.5479 / si. 03629236.257.1","Snellen, P. C. T. (1890) Aanteekeningen over Lepidoptera schadelijk voor het suikerriet. Mededelingen van het Profstation suikerbieteelt W-Java, Kagok-Tegal, 1, 94 - 97."]}

Sorolopha baribaris Sakagami, sp. nov. (Figs 9, 10, 15, 20) urn:lsid:zoobank.org:act: 15FEB3A8-8E02-4652-BFE0-C441A4634A5E Diagnosis. Sorolopha baribaris has a mean forewing length of 6.8 mm (5.5–7.9 mm, n = 37), with little sexual dimorphism except for scale tuffs on the abdomen. Males have scale tuffs on the abdomen, but females don’t. The ground color of the forewing ranges from deep green to brown. The forewing, which is relatively narrow, has a brown subcircular marking in apical area. The male genitalia of S. baribaris are characterized by a valva with a neck, a wide ventral process, and a short cucullus. The male genitalia of this species are similar to those of S. ferruginosa, described from Thailand (cf., Kawabe 1989: 30, fig. 106), but they can be distinguished by the gently curved sacculus, the shorter ventral process of the valva, the longer cucullus, and the gently curved phallus. This species is also superficially similar to S. sphaerocopa, also recorded from Japan (cf., Le et al. 2013: 523, fig. 6; Nasu 2013: 202, pl. 24, figs. 5–6), but it can be distinguished by the round markings and the position of the secondary scale tuff in the male, which is on the hindwing in S. sphaerocopa, but on the abdomen in this species. Description. Head (Fig. 10): Upper frons greenish brown or dark brown, lower frons yellowish brown, vertex roughly scaled, greenish brown or dark brown with blue illumination, labial palpus yellowish brown; antenna brown. Thorax: Dark green or dark brown, tegulae dark green to dark brown. Legs, except tarsus, light brown dorsally; tarsus dark brown, light brown at apex of each segment; legs dark brown ventrally. Forewing (Fig. 9) mean length 6.5 mm in male (5.5–7.3 mm, n = 23), 7.2 mm in female (6.0–7.9 mm, n=14), broadened distally, costa gently curved in distal half, with scattered dark brown spots; nine pairs of strigulae from base to R 4; first to fourth strigulae forming grayish green or light brown fasciae, fascia extending from first strigula merged to that from second strigula at middle, reaching dorsum, fascia extending from third strigula merged with that from fourth strigula at middle, reaching dorsum and curved outward, fascia dark brown between fourth and fifth strigulae, reaching near tornus, oblique grayish green stria extending from strigula fifth, merged with that from sixth strigula on R 1, reaching tornus, seventh, eighth, and nineth strigulae gray or light brown; preterminal spot large, circular, dark brown; apex dark brown; cilia dark brown. Hindwing dark brown, paler towards base. Cilia dark brown. Abdomen: Dark brown dorsally; beige ventrally, and with black scales from A1 to A3. Scale tuffs on posterior end of A 2 in male. Male genitalia (Fig. 15) with tegumen high, narrowed towards apex, erect subtriangular. Uncus small, semicircular. Socius about 2/5 length of tegumen, slightly curved, haired at clavate apex. Valva narrow at base third; sacculus well defined with small prominence, densely haired especially on ventral margin; cucullus short, with ventral process wide, circular, short, with spines along ventral margin, rounded at apex. Phallus lightly sclerotized, moderate in length, curved. Female genitalia (Fig. 20) with sterigma strongly sclerotized, funnel-shaped, with wide frontal cleft. Ductus bursae 1.2 times as long as corpus bursae, sclerotized in posterior 1/3, ductus seminalis originating from middle with spherical bulla seminalis. Corpus bursae mushroom-shaped, narrowed in anterior 2/3, with two signa close to ductus bursae, one short horn shaped, another small, round scobination. Material examined. Holotype. JAPAN: Wakayama Pref.: Hirai, (33°38′14″N, 135°41′23″E, alt. 142 m), ♂, emerged on 27.V.2018, reared from leaves of Actinodaphne acuminata collected on 8.V.2018, K. Sakagami leg. genitalia slide KS226♂. Paratypes. JAPAN: HONSHU: Wakayama Pref.: Shitaki (33°51′25″N, 135°50′25″E, alt. 137m), 10♂, 8♀, emerged on 27–28. V.2019, reared from leaves of A. acuminata collected on 11. V.2019, K. Sakagami leg., genitalia slide 197♂, 198♀, 200♂, 202♂, 208♂, 210♂, 216♀; Unehata (33°45′23″N, 135°47′52″E, alt. 147m), 5♂, 3♀, emerged on 21. V.–4. VI.2019, reared from leaves of A. acuminata collected on 11. V.2019, K. Sakagami leg., genitalia slide KS 201♀; type locality, 6♂, 3♀, emerged on 23–28. V.2018, reared from leaves of A. acuminata collected on 8. V.2018, K. Sakagami leg., genitalia slide KS199♂, 215♀. KYUSHU: Miyazaki Pref.: Gônohara (31°41′52″N, 131°22′00″E, alt. 60 m), Nichinan-shi, 1♂, collected on 23.VII.2022, K. Sakagami leg., genitalia slide KS222♂. Distribution. Japan (Honshu, Kyushu). Host plant. Actinodaphne acuminata (Lauraceae) Biology. Adults emerge from leaf shelters of the host between May and June in Honshu. An adult was collected by a light trap in Kyushu in late July. Larvae tie together young leaves of A. acuminata and feed on the leaves within the leaf shelters between April and May in Honshu (Fig. 21). This species emerges approximately 10 days before S. plinthograpta and S. sugiurai under laboratory conditions. Etymology. This species is named from “baribarinoki,″ which is the Japanese name for the host plant Actinodaphne acuminata. Remarks. This species shows similarities to members of the cyclotoma group in various morphological features, such as the male genitalia. However, the signa of this species differs from that of the cyclotoma group. Further investigation is required to accurately determine the taxonomic position of this species., Published as part of Sakagami, Kota, 2023, Review of the species of Sorolopha Lower (Lepidoptera: Tortricidae) in Japan with descriptions of three new species, pp. 131-142 in Zootaxa 5278 (1) on pages 138-141, DOI: 10.11646/zootaxa.5278.1.6, http://zenodo.org/record/7895134, {"references":["Kawabe, A. (1989) Records and descriptions of the subfamily Olethreutinae (Lepidoptera: Tortricidae) from Thailand. Microlepidoptera of Thailand, 2, 23 - 82.","Le, X., Qi, M. J., Park, K. T. & Bae, Y. S. (2013) A survey of the genus Sorolopha Lower, 1901 in the northern part of Vietnam (Lepidoptera: Tortricidae, Olethreutinae). Zootaxa, 3652 (5), 519 - 532. https: // doi. org / 10.11646 / zootaxa. 3652.5.2","Nasu, Y. (2013) Olethreutinae (except Grapholitini). In: Nasu, Y., Hirowatari, T. & Kishida, Y. (Eds.), The Standard of Moths in Japan IV. Gakken Education Publishing, Tokyo, pp. 198 - 272. [in Japanese]","Meyrick, E. (1931) Eucosmidae. In: Meyrick, E. (Ed.), Exotic Microlepidoptera 4. Taylor and Francis, London, pp. 126 - 153.","Meyrick, E. (1930) Description of new Microlepidoptera from Tonkin. In: de Joannis, J. (Ed.) Lepidopteres heteroceresdu Tonkin. 3 re Partie. Annales de la Societe Entomologique de France, 98 (Suppl.), 559 - 835."]}

Sorolopha sugiurai Sakagami, sp. nov. (Figs 3, 4, 12, 17) urn:lsid:zoobank.org:act: E5ECA10A-C46E-419E-9F0D-5622891B0FFF Diagnosis. Sorolopha sugiurai has a mean forewing length of 8.4 mm (6.9–9.3 mm, n = 32), with little or no sexual dimorphism. The forewing is expanded posteriorly, and is greenish-brown with scattered white scales, dark-brown shades, and inverted V-shaped white markings in the apical area. The apex of the forewing is relatively distinct, and the termen is slightly curved. S. sugiurai is characterized by gently curved long socii and long slender cucullus. This species is similar to S. herbifera Meyrick, 1909 in external appearance and in genitalia, especially those of the female (cf., Diakonoff 1973: 72, figs. 112, 113; Le et al. 2013: 530, figs. 9, 17). It can be distinguished from the latter by the darker coloration of head, thorax, and forewing, and by the gently curved basal area of the socii and the shorter ventral process of cucullus in the male genitalia. Description. Head (Fig. 4): Frons and vertex dark green; labial palpus ferruginous; antenna dark brown. Thorax: Dark brown; tegula dark green. Foreleg dark brown dorsally, beige with scattered black scales ventrally; midleg femur beige tinged with blackish scales dorsally, beige ventrally, tibia and tarsus dark brown; hindleg femur scattered with blackish scales dorsally, femur and tibia beige ventrally, dorsal side of tibia and tarsus dark brown. Forewing (Fig. 3) mean length 8.3 mm in male (6.9–9.1 mm, n = 23), 8.8 mm in female (8.1–9.3 mm, n = 9); costa gently curved, apex distinct, termen near apex slightly depressed, partly greenish-brown with white scales and dark brown areas. Costa with nine pairs of whitish strigulae from base to apex, each pair with a whitish stria extending rather obliquely; first and second strigulae forming greenish-brown patch extending to base of dorsum; third and fourth strigulae forming wide greenish-brown band reaching 1/2 length of dorsum and curved outwardly; fifth and sixth strigulae forming greenish-brown fascia running obliquely to tornus; seventh to nineth strigulae white, merging on horizontal line near costa. Inverted V-shaped white marking just below strigulae fifth‒eighth and inside dark brown spot near apex. Cilia dark brown. Hindwing with basal half light brown, distal half dark brown. Cilia dark brown. Abdomen: Dark brown dorsally, beige with scattered dark brown scales ventrally; female with hind margin of S7 bearing wide, semi-elliptical excavation. Male genitalia (Fig. 12) with tegumen high and narrowed toward apex. Uncus obtuse with round apex. Socius slender, long, reaching mid length of tegumen, gently curved at base, haired at apical clavate. Valva long, slender, curved dorsally at apex; sacculus with nearly rectangular prominence, very densely haired on posterior edge of sacculus; cucullus slender with ventral triangular process ending in single slightly curved spine, slightly curved about 2/5 length of terminal part of cucullus. Phallus short without cornutus, slightly curved, obliquely truncate. Female genitalia (Fig. 17) with papilla analis narrow. Antrum cone-shaped, sclerotized. Ductus bursae membranous, slender, same length as corpus bursae; ductus seminalis originating from close to corpus bursae, with small oval bulla seminalis. Corpus bursae distorted ovate, signa two roundish sclerites of unequal size. Material examined. Holotype. JAPAN: HONSHU: Wakayama Pref.: Unehata (33°45′N, 135°47′E, alt. 147 m), ♂, emerged on 10.VI.2019, reared from leaves of Actinodaphne acuminata collected on 11.V.2019, K. Sakagami leg., genitalia slide KS232♂. Paratypes. JAPAN: HONSHU: Wakayama Pref.: Wadagawa (Unehata) (33°46′N, 135°48′E, alt. 109 m), 5♂, 3♀, emerged on 31. V.–12. VI.2017, 2♂, emerged date unknown, reared from leaves of A. acuminata collected on 14. V.2017, K. Sakagami leg., genitalia slide KS 12♀, 16♂, 83♂, 84♀, wing slide KS 15♀; Unehata, Wakayama (33°45′N, 135°47′E, alt. 147 m), 9♂, 3♀, emerged on 8–11. VI.2018, reared from leaves of A. acuminata collected on 9. V.2018, K. Sakagami leg., genitalia slide KS91♂, 92♀, 188♂; same locality, 7♂, 5♀, emerged on 6–11. VI.2019, reared from leaves of A. acuminata collected on 11. V.2019, K. Sakagami leg., genitalia slide KS 190♀, 207♂; same locality, 2♂, 2♀, emerged on 6–12. VI.2022, reared from leaves of A. acuminata collected on 30.IV.2022, K. Sakagami leg. Distribution: Japan (Wakayama Pref.). Host plant: Actinodaphne acuminata (Lauraceae). Biology. Adults emerge from leaf shelters of the host plant in June. Larvae use silk to tie together young leaves of A. acuminata, and they feed on the leaves within the leaf shelters. The leaf shelters frequently include larvae of other Sorolopha species (Fig. 21). Larvae cut and fold back the edge of a single leaf and pupate inside it. Despite extensive sampling in the southern part of Kii Peninsula, larvae of this species were collected only from A. acuminata at a very small area. Etymology. This species is named as a patronym for Dr. Shinji Sugiura, who first discovered larvae of this species at the type locality, Wakayama Pref. Remarks. This species belongs to the herbifera group (Diakonoff 1973)., Published as part of Sakagami, Kota, 2023, Review of the species of Sorolopha Lower (Lepidoptera: Tortricidae) in Japan with descriptions of three new species, pp. 131-142 in Zootaxa 5278 (1) on page 134, DOI: 10.11646/zootaxa.5278.1.6, http://zenodo.org/record/7895134, {"references":["Diakonoff, A. (1973) The south Asiatic Olethreutini (Lepidoptera, Tortricidae). Zo ˆ logische Monographieen van het Rijksmuseum van Natuurlijke Historie No. 1, 700 pp.","Le, X., Qi, M. J., Park, K. T. & Bae, Y. S. (2013) A survey of the genus Sorolopha Lower, 1901 in the northern part of Vietnam (Lepidoptera: Tortricidae, Olethreutinae). Zootaxa, 3652 (5), 519 - 532. https: // doi. org / 10.11646 / zootaxa. 3652.5.2"]}

Sorolopha plinthograpta (Meyrick, 1931) (Figs 1, 2, 11, 16) Argyroploce plinthograpta: Meyrick, 1931: 135 (♂). TL: Taiwan. Olethreutes plinthograpta: Clarke, 1958: 539, pl. 268, fig. 1 (left wing). Eudemis plinthograpta: Issiki, 1957: 69, pl. 11, fig. 332. Sorolopha plinthograpta: Diakonoff, 1973: 84, figs. 131–133 (♂, ♀ genit.); Kawabe, 1982: 93, pl. 22, figs. 15–16; Park & Ahn, 1987: 186, fig. 12; Kuznetsov, 2001: 287, pl. 171, fig. 1 (♂ genit.); Nasu, 2011: 680, pl. 68, figs. 271a–e; Yu & Li, 2009: 10; Nasu, 2013: 202, pl. 24, fig. 4. Diagnosis. Sorolopha plinthograpta has a mean forewing length of 7.9 mm (6.3–9.0 mm, n = 30), little or no sexual dimorphism. The forewing is sub-rectangular, expanding posteriorly, and brick-red or blackish-brown with dark-brown shades on the center. The male genitalia of S. plinthograpta are similar to those of S. semiculta (Meyrick, 1909) (cf., Diakonoff 1973: 76, fig. 121), but those of S. plinthograpta have slenderer socii, lacking modified hairs at the apex, and an obliquely truncate phallus. The male genitalia of S. plinthograpta are also similar to those of S. autoberylla (Meyrick, 1932) and S. plumboviridis Diakonoff, 1973 (cf., Diakonoff 1973: 73, fig. 115; 83, fig. 135), but S. plinthograpta has fewer spines on the apical edge of the sacculus, and the terminal part of the cucullus is shorter. Material examined. JAPAN: HONSHU: Wakayama Pref.: Tanago (33°47′03″N, 135°53′24″E, alt. 59 m), 1♂, emerged on 10. VI.2019, reared from leaves of Actinodaphne acuminata collected on 11. V.2019, K. Sakagami leg., genitalia slide KS184♂; Wadagawa (33°46′02″N, 135°48′41″E, alt. 109 m), 1♂, 2♀, emerged on 8–11. VI.2017, reared from leaves of A. acuminata collected on 14. V.2017, K. Sakagami leg., genitalia slide KS 41♀, 81♂, 82♀; Unehata (33°45′23″N, 135°47′52″E, alt. 147 m), 1♂, emerged on 10. VI.2019, reared from leaves of A. acuminata collected on 11. V.2017, K. Sakagami leg.; Hirai (33°38′14″N, 135°41′23″E, alt. 142 m), 7♂, 4♀, emerged on 3–14. VI.2018, reared from leaves of A. acuminata collected on 8. V.2018, K. Sakagami leg., genitalia slide KS115♂, 116♂, 117♀, 118♂, 127♂, 183♀, 185♂, 186♂, 187♀, 189♀, 205♂; Mt. Nachisan, 1♀, collected on 27.VIII.1959, S. Moriuti leg., (OMU). KYUSHU: Fukuoka Pref.: Inunaki-tôge, collected on 24.IV.1960, K. Yano leg., (ELKU). Ôita Pref.: Beppu, collected on 1937, T. Torinaga leg., (ELKU). Kagoshima Pref.: Ôsumi, Mt. Kirishima, 1♂, collected on 13.X.1953, T. Yasuda leg., (OMU); Satamisaki, 1♂, collected on 14.X.1955, T. Kodama leg., (OMU). NANSEI ISLANDS: Nakanoshima Is.: Mt. Nanatsuyama (29°50′N, 129°53′E, alt. 134 m), 1♂, collected on 13.IX.2018 by light traps, K. Takahashi leg. Takarajima Is.: Ôgomori (29°09′20″N, 129°13′00″E, alt. 5 m), 1♂, emerged on 3.X.2018, reared from leaves of Cinnamomum daphnoides collected on 9.IX.2018, K. Sakagami leg.; Ôgomori (29°09′22″N, 129°12′54″E, alt. 6 m), 1♀, emerged on 1. V.2019, reared from leaves of C. yabunikkei collected on 7.IV.2019, 1♀, emerged on 1–2. V.2019, reared from leaves of Machilus thunbergii collected on 7.IV.2019, K. Sakagami leg.; Mt. Megamiyama (29°09′09″N, 129°11′58″E, alt. 55 m), 1♀, emerged on 3.IX.2018, reared from leaves of M. thunbergii collected on 11.IX.2018, K. Sakagami leg.; Mt. Imakiradake (29°08′49″N, 129°12′07″E, alt. 176 m), collected on 10.IX.2018, K. Sakagami. Amami Ôshima Is.: Chinase (28°21′04″N, 129°26′13″E, alt. 40 m), 1♂, 1♀, emerged on 6. V.2019, reared from leaves of A. acuminata collected on 12.IV.2019, K. Sakagami leg., genitalia slide KS 193♀, 195♂. OKINAWA IS.: Yona (26°45′N, 128°12′E), Kunigami, 1♂, 1♀, emerged on 27.IV.2015, reared from leaves of A. acuminata collected on 29.III.2015, F. Komai leg., genitalia slide KS 17♀, 62♂. Ishigakijima Is.: Mt. Omotodake (24°25′N, 125°11′E), 1♀, emerged on 28.III.2015, reared from leaves of Machilus thunbergii collected on 4.II.2015, F. Komai leg., genitalia slide KS 61♀; Takada-rindou (24°24′N, 124°11′E), 1♂, emerged on 25.III.2012, reared from leaves of M. thunbergii collected on 5.II.2012, F. Komai leg., genitalia slide KS18♂, wing slide 19♂; Ban′na-kôen (24°22′N, 124°10′E), 1♂, emerged on 24.III.2015, reared from leaves of M. thunbergii collected on 25.II.2015, F. Komai leg., genitalia slide KS20♂. Distribution. China, Taiwan, Japan (Honshu, Shikoku, Kyushu, Nansei Islands), Korea, Russia, Thailand, Vietnam, Indonesia (Java). Host plant. Actinodaphne acuminata, Cinnamomum daphnoides, C. yabunikkei, C. camphora, Litsea coreana, Machilus thunbergii (all Lauraceae), Elaeocarpus japonicus (Elaeocarpaceae) (Kawabe 1982, Nasu 2011, 2013). Biology. Adults have been taken from June to October in Honshu, Japan (Nasu 2013), and are commonly collected using light traps. Larvae tie together multiple young leaves of host plants and feed on the leaves in the shelters. Larvae of this species are often found with larvae of other Sorolopha species within the leaf shelters of host plants (Fig. 21). Larvae cut and fold back the edge of a single leaf and pupate inside it. Remarks. Diakonoff (1973) assigned this species to the herbifera group. Adults collected from the Nansei Islands (Okinawa Is. and Ishigakijima Is.) tend to be darker than those from other regions (e.g., Honshu and Kyushu) in Japan. Papers citing Kawabe (1982) indicating Pueraria montana (Fabaceae) is the host plant of S. plinthograpta (e.g., Le et al. 2013) are incorrect. Kawabe (1982) actually identified the host plant as C. camphora (Lauraceae) (Kawabe 1982)., Published as part of Sakagami, Kota, 2023, Review of the species of Sorolopha Lower (Lepidoptera: Tortricidae) in Japan with descriptions of three new species, pp. 131-142 in Zootaxa 5278 (1) on pages 132-133, DOI: 10.11646/zootaxa.5278.1.6, http://zenodo.org/record/7895134, {"references":["Meyrick, E. (1931) Eucosmidae. In: Meyrick, E. (Ed.), Exotic Microlepidoptera 4. Taylor and Francis, London, pp. 126 - 153.","Clarke, J. F. G. (1958) Catalogue of Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 3. British Museum (Natural History), London, 600 pp.","Issiki, S. (1957) Tortricidae. In: Esaki, T., Issiki, S., Mutuura, A., Inoue, H., Ogata, M., Okagaki, H. & Kuroko, H. (Eds.), Moths of Japan in Color Vol. I, Hoikusha Publishing Co., Ltd., Higashiosaka, pp. 372 - 483. [in Japanese]","Diakonoff, A. (1973) The south Asiatic Olethreutini (Lepidoptera, Tortricidae). Zo ˆ logische Monographieen van het Rijksmuseum van Natuurlijke Historie No. 1, 700 pp.","Kawabe, A. (1982) Tortricidae. In: Inoue, H., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. & Owada, M. (Eds.), Moths of Japan. Vol. 2. Kodansha, Tokyo, pp. 158 - 181. [in Japanese]","Park, K. & Ahn, S. (1987) Newly recorded species of Tortricidae (Lepidoptera) from Korea (I) Thirteen species of the subfamily Olethreutinae. Korean Journal of Applied Entomology, 26 (4), 181 - 187. [in Korean]","Kuznetsov, V. I. (2001) Tortricidae. In: Ler, P. A. (Ed.), Key to the insects of Russian Far East. Vol. V. Trichoptera and Lepidoptera. Pt 3. Dal'nauka, Vladivostok, pp. 11 - 472. [in Russian]","Nasu, Y. (2011) Sorolopha plinthograpta (Meyrick) In: Komai, F., Yoshiyasu, Y., Nasu, Y. & Saito, T. (Eds.), A Guide to the Lepidoptera of Japan. Tokai University Press, Kanagawa, pp. 680. [in Japanese]","Yu, H. L. & Li, H. (2009) Review of the genus Sorolopha Lower (Lepidoptera: Tortricidae, Olethreutinae) from Mainland China, with descriptions of two new species. Zootaxa, 2062 (1), 1 - 14. https: // doi. org / 10.11646 / zootaxa. 2062.1.1","Nasu, Y. (2013) Olethreutinae (except Grapholitini). In: Nasu, Y., Hirowatari, T. & Kishida, Y. (Eds.), The Standard of Moths in Japan IV. Gakken Education Publishing, Tokyo, pp. 198 - 272. [in Japanese]","Le, X., Qi, M. J., Park, K. T. & Bae, Y. S. (2013) A survey of the genus Sorolopha Lower, 1901 in the northern part of Vietnam (Lepidoptera: Tortricidae, Olethreutinae). Zootaxa, 3652 (5), 519 - 532. https: // doi. org / 10.11646 / zootaxa. 3652.5.2"]}