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The Southern Hemisphere Westerlies (SHW) play a major role in the global climate system. The winds drive ocean circulation and affect the Southern Oceans’ ability to take up atmospheric CO2. Recently, the SHW core belt has strengthened and shifted south, but there is an insufficient understanding of its long-term behaviour. Palaeoclimatic records are key for capturing long-term variability through the SHW’s effect on surface temperature and moisture availability. However, terrestrial records are sparse in the Southern Hemisphere. We use a palynological record from Lake Diamond on sub-Antarctic South Georgia to provide reconstructions of vegetation and climate for the last ~10,000 years. Influx of long-distance transported pollen is used as a measure of surface wind strength. Changes in relative pollen abundance of native taxa occupying either upland (cold) or lowland (warm) environments indicate local climatic variability. On South Georgia, we find long-distance transported pollen from South American taxa, mainly Nothofagus and Ephedra. They show a general increase in abundance throughout the Holocene, with peak influxes between 5700–5400, 2800–1500 and 1000–500 cal yr BP. These intervals coincide with colder periods inferred from the palynological record, suggesting that SHW variation and temperature on South Georgia are highly connected. Agreement with palaeoecological records from eastern Patagonia show that climatic changes have been regionally consistent. The record from Lake Diamond further illustrates the importance of remote islands in contributing to a deeper understanding of atmospheric circulation and climatic variability in the sub-Antarctic.

The Southern Hemisphere Westerly Winds play a major role in the global climate system. By driving circulation in the Southern Ocean and its subsequent effect on the upwelling of carbon-rich deep water, the Westerlies affect the oceans ability to take up atmospheric CO2. Furthermore, by impacting temperature conditions and moisture availability, the Westerlies act as a first-order control on local environmental conditions. Uncovering long term natural climatic variability in the sub-Antarctic is therefore crucial to understand how the global system might react under future climate changes. Due to the lack of land mass on the Southern Hemisphere, sub-Antarctic islands are essential for studying climate variability in this region; terrestrial records provide valuable insights into both local and regional surface climate conditions. We use a pollen record from Lake Diamond to provide detailed reconstructions of vegetation and climate on sub-Antarctic South Georgia for the last ~9900 years. Westerly Wind strength and position is inferred from long-distance transport of pollen from South America, Africa, and New Zealand. Additionally, changes in relative pollen abundance of native taxa occupying either upland (cold) or lowland (warm) environments are used to infer local climatic variation, supported by additional sedimentological proxies. On South Georgia we find long-distance transported pollen from several South American taxa, mainly Nothofagus, Ephedra and Asteraceae. They show a general increase in abundance throughout the Holocene, with peak influx between 2800 and 1500 cal yr BP, most likely caused by changes in the strength of the Southern Hemisphere Westerly Winds. In both our record and others, this interval is seen as the end of the Neoglacial period.

A detailed, well-dated record of pollen and sedimentary ancient DNA (sedaDNA) for the period 15 000–9500 cal a bp describes changes at Lake Bolshoye Shchuchye in the Polar Ural Mountains, located far east of the classical Lateglacial sites in western Europe. Arctic tundra rapidly changed to lusher vegetation, possibly including both dwarf (Betula nana) and tree birch (B. pubescens), dated in our record to take place 14 565 cal a bp, coincident with the onset of the Bølling in western Europe; this was paralleled by increased summer temperatures. A striking feature is an early decline in Betula pollen and sedaDNA reads 300 years before the onset of the Younger Dryas (YD) in western Europe. Given the solid site chronology, this could indicate that the YD cooling started in Siberia and propagated westwards, or that the vegetation reacted to the inter-Allerød cooling at 13 100 cal a bp and did not recover during the late Allerød. During the YD, increases in steppe taxa such as Artemisia and Chenopodiaceae suggest drier conditions. At the onset of the Holocene, the vegetation around the lake reacted fast to the warmer conditions, as seen in the increase of arboreal taxa, especially Betula, and a decrease in herbs such as Artemisia and Cyperaceae. publishedVersion

Early agriculture can be detected in palaeovegetation records, but quantification of the relative importance of climate and land use in influencing regional vegetation composition since the onset of agriculture is a topic that is rarely addressed. We present a novel approach that combines pollen-based REVEALS estimates of plant cover with climate, anthropogenic land-cover and dynamic vegetation modelling results. This is used to quantify the relative impacts of land use and climate on Holocene vegetation at a sub-continental scale, i.e. northern and western Europe north of the Alps. We use redundancy analysis and variation partitioning to quantify the percentage of variation in vegetation composition explained by the climate and land-use variables, and Monte Carlo permutation tests to assess the statistical significance of each variable. We further use a similarity index to combine pollen based REVEALS estimates with climate-driven dynamic vegetation modelling results. The overall results indicate that climate is the major driver of vegetation when the Holocene is considered as a whole and at the sub-continental scale, although land use is important regionally. Four critical phases of land-use effects on vegetation are identified. The first phase (from 7000 to 6500 BP) corresponds to the early impacts on vegetation of farming and Neolithic forest clearance and to the dominance of climate as a driver of vegetation change. During the second phase (from 4500 to 4000 BP), land use becomes a major control of vegetation. Climate is still the principal driver, although its influence decreases gradually. The third phase (from 2000 to 1500 BP) is characterised by the continued role of climate on vegetation as a consequence of late-Holocene climate shifts and specific climate events that influence vegetation as well as land use. The last phase (from 500 to 350 BP) shows an acceleration of vegetation changes, in particular during the last century, caused by new farming practices and forestry in response to population growth and industrialization. This is a unique signature of anthropogenic impact within the Holocene but European vegetation remains climatically sensitive and thus may continue to respond to ongoing climate change. (C) 2017 Elsevier Ltd. All rights reserved.

We reconstruct mean July temperature (Tjul) from three sites on the island of Andoya, northern Norway for the period 15 000–9000 cal yr BP using three biotic proxies. A probabilistic indicator species approach (pdf method) was used for plant macrofossil data from all sites (M-Tjul). Reconstructions based on pollen (P-Tjul) (2 sites) and chironomids C-Tjul) (1 site) were done using weighted averaging and weighted-averaging partial least-squares, respectively. A sediment proxy for catchment glacier development was measured at the main site, Lusvatnet. The July temperature reconstructions from all the sites show a similar development through the deglaciation. An initial July temperature of 4 °C increased to a maximum of 7–8 °C during the Allerod at c.13 200 cal yr BP. There was no marked cooling at the start of the Younger Dryas (YD) chronozone, at Lusvatnet, C-Tjul decreased after 12 700 cal yr BP to a low of 6 °C at about 12 550 cal yr BP before increasing markedly around 12 400 cal yr BP. Conversely, P-Tjul rose slowly through the early YD. Both proxies exceeded the Allerod temperatures for around 300 years before decreasing towards a minimum of 6 °C at c. 12 000 cal yr BP. M-Tjul maintained a mean of c. 8 °C throughout the YD. Different sensitivities of proxies are discussed. Their different responses during the YD at the three sites, along with the glacial evidence, highlight the role of aridity. At c. 11 500 cal yr BP, July temperatures increased rapidly over Andoya to about 10 °C, and then rose gradually to maximum values of 12 °C at c. 9700 cal yr BP. The temperature reconstructions are consistent with the flow of Atlantic water into the Nordic Seas except during the YD, when variability in sea ice may have been more important. During the early Holocene there are some differences between sites, resulting from local site factors such as altitude and exposure, soil development, and catchment instability.

Sedimentary pollen offers excellent opportunities to reconstruct vegetation changes over past millennia. Number of different pollen taxa or pollen richness is used to characterise past plant richness. To improve the interpretation of sedimentary pollen richness, it is essential to understand the relationship between pollen and plant richness in contemporary landscapes. This study presents a regional-scale comparison of pollen and plant richness from northern Europe and evaluates the importance of environmental variables on pollen and plant richness. We use a pollen dataset of 511 lake-surface pollen samples ranging through temperate, boreal and tundra biomes. To characterise plant diversity, we use a dataset formulated from the two largest plant atlases available in Europe. We compare pollen and plant richness estimates in different groups of taxa (wind-pollinated vs. non-wind-pollinated, trees and shrubs vs. herbs and grasses) and test their relationships with climate and landscape variables. Pollen richness is significantly positively correlated with plant richness (r = 0.53). The pollen plant richness correlation improves (r = 0.63) when high pollen producers are downweighted prior to estimating richness minimising the influence of pollen production on the pollen richness estimate. This suggests that methods accommodating pollen-production differences in richness estimates deserve further attention and should become more widely used in Quaternary pollen diversity studies. The highest correlations are found between pollen and plant richness of trees and shrubs (r = 0.83) and of wind-pollinated taxa (r = 0.75) suggesting that these are the best measures of broad-scale plant richness over several thousands of square kilometres. Mean annual temperature is the strongest predictor of both pollen and plant richness. Landscape openness is positively associated with pollen richness but not with plant richness. Pollen richness values from extremely open and/or cold areas where pollen production is low should be interpreted with caution because low local pollen production increases the proportion of extra-regional pollen. Synthesis. Our results confirm that pollen data can provide insights into past plant richness changes in northern Europe, and with careful consideration of pollen-production differences and spatial scale represented, pollen data make it possible to investigate vegetation diversity trends over long time-scales and under changing climatic and habitat conditions.

A new palaeoclimatic reconstruction of mid-summer (July) temperatures for the last 7.5 ka in northern Fennoscandia is presented. It is based on two botanical proxies: spectra of fossil pollen and tree rings of Scots pine logs recovered from lacustrine sediments in the Arctic tree-line region. A newly developed method of proxy fusion is used to integrate the proxy-specific reconstructions of past summer temperature variability based on the pollen-stratigraphic and dendroclimatic data. The rationale behind the method is that the two proxies are likely to be connected to climate variability in a timescale-dependent fashion and, accordingly, the new reconstruction makes use of the low- and high-frequencies from pollen-stratigraphic and tree-ring data, respectively. The most prominent features of the new reconstruction are: (1) the long-term decline of temperatures by 2.0 °C over the past 7.5 ka, (2) the mid-Holocene warmth culminating between 5 and 4 ka as a deviation from the cooling trend, (3) the Little Ice Age cool phase between 0.7 and 0.1 ka, and (4) the subsequent warming during the past century. These periods are superimposed on year-to-year variations in climate as dated to calendar-year accuracy by dendrochronology. Within the modern period, the years 1934 and 1937 are among the warmest, and the years 1903 and 1910 are among the coldest summers in the context of the past 7.5 ka. On average, the reconstructed Holocene climate was approximately 0.85 °C warmer than the twentieth century.

Kruger, L. C., Paus, A., Svendsen, J. I. & Bjune, A. E. 2011: Lateglacial vegetation and palaeoenvironment in W Norway, with new pollen data from the Sunnmore region. Boreas, 10.1111/j.1502-3885.2011.00213.x. ISSN 0300-9483. Two sediment sequences from Sunnmore, northern W Norway, were pollen-analytically studied to reconstruct the Lateglacial vegetation history and climate. The coastal Dimnamyra was deglaciated around 15.3 ka BP, whereas Lokjingsmyra, further inland, became ice-free around 14 ka BP. The pioneer vegetation dominated by snow-bed communities was gradually replaced by grassland and sparse heath vegetation. A pronounced peak in Poaceae around 12.9 ka BP may reflect warmer and/or drier conditions. The Younger Dryas (YD) cooling phase shows increasing snow-bed vegetation and the local establishment of Artemisia norvegica. A subsequent vegetation closure from grassland to heath signals the Holocene warming. Birch forests were established 500–600 years after the YD–Holocene transition. This development follows the pattern of the Sunnmore region, which is clearly different from the Empetrum dominance in the Lateglacial interstadial further south in W Norway. The Lateglacial oscillations GI-1d (Older Dryas) and GI-1b (Gerzensee) are hardly traceable in the north, in contrast to southern W Norway. The southern vegetation was probably closer to an ecotone and more susceptible to climate changes.

In the DOORMAT (Direct Observation of Recent Macrofossils Across Treeline) project, the modern representation of local vegetation by pollen and plant remains (plant macrofossils) across a west Norwegian tree line, composed of Betula pubescens and Pinus sylvestris, has been studied over 2 years. The aim was to discover if the modern tree line could be detected and therefore how precisely past tree-line movements could be reconstructed and related to Holocene climate changes by using one proxy or a combination of both. Traps were placed in the vegetation from 663 to 1,120 m a.s.l., spanning the pine altitudinal species limit, the birch tree-line ecotone, and the vegetation zones up to the mid-alpine zone. Three traps were also set in the small lake Trettetjorn close to the modern tree line at 800 m a.s.l. Traps were emptied twice a year to sample both summer and winter seasons. Macrofossils represent their local vegetation well. However, tree Betula remains were trapped above the tree line and Pinus and Picea remains were recorded 1.0–1.5 km away from their sources, demonstrating considerable dispersal capacity. This shows that rare macrofossil remains do not necessarily represent the local presence of these trees. Aerial tree pollen deposition in traps at the upper limit of pine woodland and in the subalpine birch woodland was unexpectedly low, whereas pollen accumulation rates (PAR) were orders of magnitude higher in the lake traps. We hypothesise that the lake receives regional pollen rain washed in from its catchment by snow meltwater and that high values in traps are due to continuous suspension of pollen in the lake water during summer. The interpretation of tree-line changes from existing Holocene pollen and plant macrofossil data from Trettetjorn was supported and refined by the DOORMAT macrofossil data, but the modern pollen data were anomalous.

Bjune, A. E., Birks, H. J. B., Peglar, S. M. & Odland, A. 2010: Developing a modern pollen–climate calibration data set for Norway. Boreas, Vol. 39, pp. 674–688. 10.1111/j.1502-3885.2010.00158.x. ISSN 0300-9483. Modern pollen–climate data sets consisting of modern pollen assemblages and modern climate data (mean July temperature and mean annual precipitation) have been developed for Norway based on 191 lakes and 321 lakes. The original 191-lake data set was designed to optimize the distribution of the lakes sampled along the mean July temperature gradient, thereby fulfilling one of the most critical assumptions of weighted-averaging regression and calibration and its relative, weighted-averaging partial least-squares regression. A further 130 surface samples of comparable taphonomy, taxonomic detail and analyst became available as a result of other projects. These 130 samples, all from new lakes, were added to the 191-lake data set to create the 321-lake data set. The collection and construction of these data sets are outlined. Numerical analyses involving generalized linear modelling, constrained ordination techniques, weighted-averaging partial least-squares regression, and two different cross-validation procedures are used to asses the effects of increasing the size of the calibration data set from 191 to 321 lakes. The two data sets are used to reconstruct mean July temperature and mean annual precipitation for a Holocene site in northwest Norway and a Lateglacial site in west-central Norway. Overall, little is to be gained by increasing the modern data set beyond about 200 lakes in terms of modern model performance statistics, but the down-core reconstructions show less between-sample variability and are thus potentially more plausible and realistic when based on the 321-lake data set.

The Holocene lake history, vegetation history and climate history of Brurskardstjorni, an alpine lake in the Jotunheimen Mountains of south-central Norway, are reconstructed. The reconstructions are based on fossil pollen, plant macrofossils, diatoms, chironomids and sediment characteristics. Subsequent to deglaciation, the lake was formed at about 11,000 cal years BP. A diverse chironomid assemblage quickly colonised the lake, whereas the first diatoms were found about 400 years later. At that time, the lake water was turbid with a high pH. The surrounding soils were immature and unstable and dominated by open pioneer vegetation. Compared to the present, summer temperatures were warmer and there was less winter precipitation. From about 10,000 cal years BP, local organic production increased rapidly and from about 9,500 cal years BP a few macrofossils and a high pollen influx of birch suggest that the tree-line was close to the lake. Pine most likely reached its highest tree-line altitude around 9,000 cal years BP and has receded since that time. From about 5,000 cal years BP, the total amount of trees and shrubs decreased and the landscape became more open, probably due to decreasing temperatures and increasing effective moisture lowering the birch tree-line. Coinciding with a cooling during the last 3,000 years, lake-water pH decreased. There is large incongruence between the Holocene July temperatures inferred from pollen and chironomids. The biological proxies responded to a combined effect of environmental change and biotic interactions. This response is interpreted with reference to taxon-environment relationships in the modern calibration data sets and with reference to the latent structure and ecological demands of the fossil assemblages.

Little is known about the relationships between fossil oribatid mite assemblages in lake and mire sediments and the composition, abundance, and richness of their living communities. Because oribatid mites are a relatively new area of palaeolimnological study, there is a great lack of knowledge about the taphonomic processes that might affect fossil mite assemblages. The DOORMAT (Direct Observation Of Recent Macrofossils Across Tree-line) project was designed to study the transport and deposition of oribatid mites and plant remains in the tree-line area of western Norway. The present study also compares modern oribatid assemblages with fossil oribatid assemblages in a Holocene lake-sediment sequence from the nearby Trettetjorn, and considers the optimal location for studying fossil oribatid mites within a lake basin. Seven novel terrestrial traps (50 × 80 cm Astroturf doormats) were placed at major vegetational transitions along an altitudinal transect crossing the tree-line ecotone from 633 to 1,120 m a.s.l. at Upsete, west Norway. Three sediment traps were placed in Trettetjorn (810 m a.sl.) at the inlet, the middle, and near the outlet. In each terrestrial trap, the oribatid assemblage was found to be characteristic of the surrounding habitat. The lake-trap analyses showed that aquatic or moist-habitat species had the highest chance of being incorporated into the lake sediments; the number of terrestrial species decreased considerably from both outlet and inlet traps to the central trap in the deepest water. The area adjacent to the inlet of Trettetjorn would therefore be the optimal location for a sediment core for oribatid analysis. However, this conclusion is not supported when the modern trap results are compared with the Trettetjorn sequence from the lake centre.

With global warming, the distribution of warmth through the year is likely to change in the future and comparable changes may have occurred over the course of the Holocene. Its effect on vegetation composition and species distribution can be compared to that of a continental versus an oceanic climate. The distribution of five major tree species along a continentality gradient was studied in Fennoscandia based on distribution maps and on their proportions of pollen in surface-sediment samples. Both analyses indicate that the five arboreal species show similar patterns of response to a continentality index in the order Ulmus glabra, Corylus avellana, Quercus robur, Tilia cordata, and Picea abies from the most oceanic to most continental. Continentality, growing degree days above 5 degrees C, and January temperature were reconstructed quantitatively from four pollen diagrams using transfer functions based on a combined Fennoscandian pollen surface-sample data-set. Quantitative reconstructions indicate that the climate in Fennoscandia has become increasingly more continental over the last 7000 years, and this is largely an effect of winter cooling. Early Holocene vegetation composition has poor analogues to the present vegetation in Fennoscandia, which hampers quantitative reconstructions. Qualitative reconstructions suggest that the early Holocene in Fennoscandia was the most oceanic period, but probably with a high variability in temperature. (C) 2008 Elsevier Ltd. All rights reserved.

Pollen and plant macrofossil analyses from Svanavatnet in northern Norway provide records of past vegetation and climate in this region from c. 8700 cal.yr BP until the present. Pollen accumulation rates and the presence of plant macrofossils indicate that Betula pubescens (birch) was present from c. 8600 cal.yr BP and Pinus sylvestris (pine) from c. 8200 cal.yr BP. Quantitative climate is reconstructed using modern pollen-climate transfer functions based on weighted-averaging partial least squares regression. A rapid increase in mean July temperature (T-jul) and mean annual precipitation (P-ann) is inferred for the early Holocene. At times when tree abundance is at its highest and most diverse, inferred Tjul indicates maximum temperatures during the mid-Holocene of about 2 degrees C warmer than at present. During the same time period, inferred Pann is 200 - 300mm above present-day conditions until c. 3000 cal.yr BP. Mean January temperatures (T-jan) are reconstructed to be about 2 degrees C warmer than today from 8000 to 3500 cal.yr BP. After 3500 cal.yr BP until today, a gradual decrease is seen in all the reconstructed climate parameters, together with a reduction in tree abundance and the development of a mosaic of open vegetation with grasses, dwarf shrubs and wet areas, and of woodland containing B. pubescens, P. sylvestris and Picea abies (spruce).

Two records of July temperature and two records of reconstructed winter precipitation along the western coast of Norway have been combined to examine regional patterns in glacier activity and winter precipitation during the Holocene. The maritime glaciers in western Norway are mainly controlled by winter precipitation. Hence, fluctuations in magnitude and equilibrium-line altitude (ELA) of these glaciers reflect variability in wintertime atmospheric circulation patterns in the northeast Atlantic region. By combining an independent proxy for summer temperature with reconstructed ELAs during the Holocene, it is possible to reconstruct former winter precipitation. We track the dominant position of the westerlies during the Holocene by comparing records of winter precipitation along a 2000 km south–north coastal transect in Norway. Analyses of modern data on the evolution of the polar vortex indicate that there is a strong relationship between polar vortex and storm-track variability over the North Atlantic. Periods with increased winter precipitation along the coast of Norway are associated with a stronger effect of the westerlies, where differences in the distribution of precipitation are assumed to reflect changes in the position of the westerlies. The largest precipitation anomalies caused by a dominant southerly position of the westerlies are found around 2.8, 1.2 and 0.4 ka yrs BP. However, a general humid phase prevailed between 2.3 and 0.9 ka yrs BP.

Holocene lake and catchment environmental history and regional climate are reconstructed from lake sediments at Trettetjørn, a small lake situated close to the present-day treeline in western Norway. Sediments began to accumulate in the lake ca. 8575 (± 115) cal yr BP. Pollen-inferred mean July temperatures (Tjul) fluctuated below 12°C with two cooler phases ca. 8400 and 8200 cal yr BP. The pollen-inferred annual precipitation (Pann) was lowest during the early Holocene and varied around 1600 mm yr−1. At the same time the highest diatom-inferred pH values (6.8) were reconstructed, probably due to input of base cations from the immature catchment soils. Betula pubescens became established around the lake ca. 8270 cal BP, soon followed by Pinus sylvestris. Maximum Tjul of ca. 12.5°C occurred from 7760 to 5200 cal yr BP. The oribatid mite assemblages confirm the vegetation development from semiopen grassland to forest. Inferred Tjul was variable after 5200 cal yr BP and declined markedly around 4175 cal yr BP. At the same time more oceanic conditions are inferred with changes in the vegetation and mite assemblages suggesting the expansion of mires. After ca. 5000 cal yr BP and towards the present, the diatom concentration in the core becomes low and variable, thickness of the valves within the same taxon varies, and sometimes diatom frustules are completely absent from the sediment column. The diatom valve dissolution has affected both the diatom assemblages and the diatom-inferred pH up to the present-day. All proxies suggest that human impact affected the catchment after ca. 2000 cal yr BP followed by more intensive impact after ca. 1555 cal yr BP. The enlargement of the settlement at Upsete during the construction of the Bergen–Oslo railway (1894–1909 AD) is reflected by a charcoal peak and the presence of spheroidal carbonaceous particles coincides with a change in the diatom assemblages. Published version

Analyses of organic content, magnetic susceptibility, grain size and pollen in sediments from the proglacial lake Vanndalsvatnet in western Norway provide a high-resolution terrestrial record and pollen-based quantitative estimates of mean July and January temperatures and annual precipitation across the ∼ 8200 cal. yr BP event. Glaciers in the catchment melted away at approximately 8600 cal. yr BP. Immediately following deglaciation, a series of thin minerogenic layers indicate several abrupt, short-lived glacial episodes peaking at ∼ 8550, 8450, 8350, 8250, 8200, 7900, 7300 and 7150 cal. yr BP. A single, mid-Holocene glacial episode occurred at 4900-4800 cal. yr BP. Between 2000 and 1400 cal. yr BP, six short-lived glacial episodes occurred ∼ 2000, 1900, 1800, 1700, 1600, and 1500 cal. yr BP. The part of Spørteggbreen that drains to Vanndalsvatnet has existed continuously since ∼ 1400 cal. yr BP. Just prior to a first loss-on-ignition minimum reflecting a glacial episode centred at 8200 cal. yr BP, pollen-inferred July temperatures were relatively high, January temperatures were low, and annual precipitation was relatively low. During the period 8200-7900 cal. yr BP, July temperatures showed a falling trend. Both January temperature and annual precipitation, however, were relatively high. After 7900 cal. yr BP, July temperatures increased, but both January temperatures and annual precipitation were lower than in the preceding period. The pollen analytical and sedimentary data suggest that the glacial advance during the Finse event seems not to have been a response to cooler summers, but to milder winters and increasing precipitation (similar to a positive North Atlantic Oscillation weather mode).

Long-term tree-line fluctuations have been studied using pollen and plant macrofossils preserved in lake sediments from three sites on an oceanic to continental transect in southern Norway. After deglaciation the early Holocene vegetation developed from an open pioneer herb-dominated vegetation into dwarf-shrub heath with shrubs, which was soon colonised by Betula and later also by Pinus sylvestris at the two lower sites. Maximum tree-line altitudes occurred in the early-to mid-Holocene. P. sylvestris reached 100–150 m higher than at present in continental areas, and 35–100 m higher on the west coast during the Holocene. Betula pubescens grew at altitudes that today reach 1300 m a.s.l. in Jotunheimen and 800 m a.s.l. in western Norway. The Pinus forest was at the two southwestern sites mixed with Betula and Alnus was closer to the sites between 8500 and 7500 cal years BP. A shift from mixed pine-birch woodland to birch woodland is seen from ca. 4300 cal years BP with the development of a sub-alpine birch belt followed by further recession of the birch forest. On the west coast, birch dominated only during the last 500–1000 years. Further decrease of woodland and opening of the landscape in the last 2000 years occurred due to climatic change and human impact such as sheep and cattle grazing.

Reconstructions of mean July temperature (Tjul) and winter precipitation (Pw) for the last 11/500 years on the Folgefonna peninsula are presented. Tjul was reconstructed using pollen-climate transfer functions and Pw was reconstructed based on the exponential relationship between mean solid winter precipitation and ablation-season temperature at the equilibrium-line altitude (ELA) with a reconstructed former ELA, using Tjul as the proxy for ablation-season temperature. The reconstructions from the Folgefonna peninsula suggest that the early Holocene was relatively cool and dry until c. 8000 cal. yr BP, followed by a warm and humid mid-Holocene until c. 4000 cal. yr BP with inferred Tjul above 12°C and Pw reaching as high as 225% of the present day. Subsequent to c. 4000 cal. yr BP a reduction is seen in both inferred Tjul and Pw with large fluctuations during the last 500 years. In addition, new calculations of Pw from two glaciers (Hardangerjøkulen and Jostedalsbreen) in southern Norway are presented. The results show that Pw varied in phase at all glaciers, probably as a response to the same climate forcing factor. During the early Holocene a major shift is suggested between winds from the west and the east.

Holocene variations of Bjornbreen, Smorstabbtinden massif, west-central Jotunheimen are reconstructed from the lithostratigraphy of two alpine stream-bank mires flooded episodically by meltwater. The approach uses multiple sedimentological indicators (weight loss-on-ignition, mean grain size, grain-size fractions, bulk density, moisture content and magnetic susceptibility), an a priori model of overbank deposition of suspended glaciofluvial sediments, a detailed chronology based on 56 radiocarbon dates, and a Little Ice Age sedimentological analogue. Rapid, late-Preboreal deglaciation was indicated by immigration of Betula pubescens by 9700 cal. BP. An interval of at least 3000 years in the early Holocene when glaciers were absent was interrupted by two abrupt episodes of glacier expansion around the time of the Finse Event, the first at ca 8270–7900 cal. BP (Bjornbreen I Event) and the second at ca 7770–7540 cal. BP (Bjornbreen II Event). Neoglaciation began shortly before ca 5730 cal. BP with gradual build-up to the maximum of the Bjornbreen III Event at ca 4420 cal. BP. Later maxima occurred at ca 2750 cal. BP (Bjornbreen IV Event) and at 1300, 1260, 1060 and 790 cal. BP (all within the Bjornbreen V Event). Glaciers were smaller than today and possibly melted away on several occasions in the late Holocene (ca 3950, 1410 and 750 cal. BP). Minor maxima also occurred at ca 660 and 540 cal. BP, within the late Mediaeval Warm Period and the early Little Ice Age, respectively. The Little Ice Age maximum was dated to 213±25 BP (ca 205 cal. BP). The relative magnitudes of the main glacier maxima were determined: Erdalen Event>Little Ice Age Event (Bjornbreen VI)>Bjornbreen I (Finse Event) ≈ Bjornbreen II>Bjornbreen V⩾Bjornbreen IV>Bjornbreen III. These episodic events of varying magnitude and abruptness were used in conjunction with an independent summer-temperature proxy to reconstruct variations in equilibrium-line altitude (ELA) and a Holocene record of winter precipitation. Since the Preboreal, ELA varied within a range of about 390 m, and winter precipitation ranged between 40 and 160% of modern values. Winter precipitation variations appear to have been the main cause of these century- to millennial-scale Holocene glacier variations.

Palaeoecological records provide a rich source of information to explore how plant distribution ranges respond to climate changes, but their use is complicated by the fact that, especially when based on pollen data, they are often spatially too inaccurate to reliably determine past range limits. To solve this problem, we focus on hazel ( Corylus avellana), a tree species with large and heavy fruits (nuts), which provide firm evidence of the local occurrence of species in the past. We combine the fossil nut records of hazel from Fennoscandia, map its maximum distribution range during the Holocene thermal maximum (HTM) and compare the fossil record with the Holocene hazel range shift as simulated by the LPJ-GUESS dynamic vegetation model. The results show that the current northern range limit of hazel in central and eastern Fennoscandia is constrained by too short growing seasons and too long and cold winters and demonstrate that the species responded to the HTM warming of about 2.5°C (relative to the present) by shifting its range limit up to 63–64°N, reached a rough equilibrium with the HTM climatic conditions and retreated from there to about 60°N during the last 4000 years in response to the late-Holocene cooling. Thus, the projected future warming of about 2.5°C would reverse the long-term southward retraction of species’ northern range limit in Europe and is likely to lead to hazel being a common, regeneratively reproductive species up to 63–64°N. In addition to the accuracy of the projected warming, the likelihood of this scenario will depend on inter-specific competition with other tree taxa and the potential of hazel to migrate and its population to grow in balance with the warming. In general, the range dynamics from the HTM to the present suggest a tight climatic control over hazel’s range limit in Fennoscandia.

A contribution to the PAGES Focus 4 Land Use and Cover Theme ABSTRACT: We present quantitative reconstructions of regional vegetation cover in north western Europe western Europe north of the Alps and eastern Europe for five time windows in the Holocene [around 6k 3k 0.5k 0.2k and 0.05k calendar years before present (bp)] at a 1° × 1° spatial scale with the objective of producing vegetation descriptions suitable for climate modelling. The REVEALS model was applied on 636 pollen records from lakes and bogs to reconstruct the past cover of 25 plant taxa grouped into 10 plant functional types and three land cover types [evergreen trees summer green (deciduous) trees and open land]. The model corrects for some of the biases in pollen percentages by using pollen productivity estimates and fall speeds of pollen and by applying simple but robust models of pollen dispersal and deposition. The emerging patterns of tree migration and deforestation between 6k bp and modern time in the REVEALS estimates agree with our general understanding of the vegetation history of Europe based on pollen percentages. However the degree of anthropogenic deforestation (i.e. cover of cultivated and grazing land) at 3k 0.5k and 0.2k bp is significantly higher than deduced from pollen percentages. This is also the case at 6k in some parts of Europe in particular Britain and Ireland. Furthermore the relationship between summer green and evergreen trees and between individual tree taxa differs significantly when expressed as pollen percentages or as REVEALS estimates of tree cover. For instance when Pinus is dominant over Picea as pollen percentages Picea is dominant over Pinus as REVEALS estimates. These differences play a major role in the reconstruction of European landscapes and for the study of land cover–climate interactions biodiversity and human resources.

International audience; This study aims to evaluate the direct effects of anthropogenic deforestation on simulated climate at two contrasting periods in the Holocene, ∼ 6 and ∼ 0.2 k BP in Eu-rope. We apply the Rossby Centre regional climate model RCA3, a regional climate model with 50 km spatial resolution, for both time periods, considering three alternative descriptions of the past vegetation: (i) potential natural vegetation (V) simulated by the dynamic vegetation model LPJ-GUESS, (ii) potential vegetation with anthro-pogenic land use (deforestation) from the HYDE3.1 (History Database of the Global Environment) scenario (V + H3.1), and (iii) potential vegetation with anthropogenic land use Published by Copernicus Publications on behalf of the European Geosciences Union. 662 G. Strandberg et al.: Sensitivity to changes in anthropogenic deforestation from the KK10 scenario (V + KK10). The climate model results show that the simulated effects of deforestation depend on both local/regional climate and vegetation characteristics. At ∼ 6 k BP the extent of simulated deforestation in Europe is generally small, but there are areas where deforestation is large enough to produce significant differences in summer temperatures of 0.5–1 • C. At ∼ 0.2 k BP, extensive deforestation , particularly according to the KK10 model, leads to significant temperature differences in large parts of Europe in both winter and summer. In winter, deforestation leads to lower temperatures because of the differences in albedo between forested and unforested areas, particularly in the snow-covered regions. In summer, deforestation leads to higher temperatures in central and eastern Europe because evapotranspiration from unforested areas is lower than from forests. Summer evaporation is already limited in the south-ernmost parts of Europe under potential vegetation conditions and, therefore, cannot become much lower. Accordingly , the albedo effect dominates in southern Europe also in summer, which implies that deforestation causes a decrease in temperatures. Differences in summer temperature due to deforestation range from −1 • C in southwestern Europe to +1 • C in eastern Europe. The choice of anthropogenic land-cover scenario has a significant influence on the simulated climate, but uncertainties in palaeoclimate proxy data for the two time periods do not allow for a definitive discrimination among climate model results.

We present pollen-based reconstructions of the spatio-temporal dynamics of northern European regional vegetation abundance through the Holocene. We apply the Regional Estimates of VEgetation Abundance from Large Sites (REVEALS) model using fossil pollen records from eighteen sites within five modern biomes in the region. The eighteen sites are classified into four time-trajectory types on the basis of principal components analysis of both the REVEALS-based vegetation estimates (RVs) and the pollen percentage (PPs). The four trajectory types are more clearly separated for RVs than PPs. Further, the timing of major Holocene shifts, rates of compositional change, and diversity indices (turnover and evenness) differ between RVs and PPs. The differences are due to the reduction by REVEALS of biases in fossil pollen assemblages caused by different basin size, and inter-taxonomic differences in pollen productivity and dispersal properties. For example, in comparison to the PPs, the RVs show an earlier increase in Corylus and Ulmus in the early-Holocene and a more pronounced increase in grassland and deforested areas since the mid-Holocene. The results suggest that the influence of deforestation and agricultural activities on plant composition and abundance from Neolithic times was stronger than previously inferred from PPs. Relative to PPs, RVs show a more rapid compositional change, a largest decrease in turnover, and less variable evenness in most of northern Europe since 5200 cal yr BP. All these changes are primarily related to the strong impact of human activities on the vegetation. This study demonstrates that RV-based estimates of diversity indices, timing of shifts, and rates of change in reconstructed vegetation provide new insights into the timing and magnitude of major human distribution on Holocene regional, vegetation, feature that are critical in the assessment of human impact on vegetation, land-cover, biodiversity, and climate in the past. (C) Elsevier Ltd.All tights reserved.

The late-glacial vegetation development in northern Norway in response to climate changes during the Allerod, Younger Dryas (YD), and the transition to the Holocene is poorly known. Here we present a high-resolution record of floral and vegetation changes at lake Lusvatnet, south-west Andoya, between 13500 and 8000 cal b.p. Plant macrofossil and pollen analyses were done on the same sediment core and the proxy records follow each other very closely. The core has also been analyzed using an ITRAX XRF scanner in order to check the sediment sequence for disturbances or hiatuses. The core has a good radiocarbon-based chronology. The Saksunarvatn tephra fits very well chronostratigraphically. During both the Allerod and the Younger Dryas time-periods arctic vegetation prevailed, dominated by Salix polaris associated with many typically arctic herbs such as Saxifraga cespitosa, Saxifraga rivularis and Oxyria digyna. Both periods were cold and dry. Between 12450 and 12250 cal b.p. during the Younger Dryas chronozone, the assemblage changed, particularly in the increased abundance of Papaver sect. Scapiflora and other high-Arctic herbs, suggesting the development of polar desert vegetation mainly as a response to increased aridity. After 11520 cal b.p. a gradually warmer and more oceanic climate initiated a succession to dwarf-shrub vegetation and the establishment of Betula woodland after 1,000 years at c. 10520 cal b.p. The overall late-glacial aridity contrasts with oceanic conditions in southern Norway and is probably related to sea-ice extent.

Mid to high latitude forest ecosystems have undergone several major compositional changes during the Holocene. The temporal and spatial patterns of these vegetation changes hold potential information to their causes and triggers. Here we test the hypothesis that the timing of vegetation change was synchronous on a sub-continental scale, which implies a common trigger or a step-like change in climate parameters. Pollen diagrams from selected European regions were statistically divided into assemblage zones and the temporal pattern of the zone boundaries analysed. The results show that the temporal pattern of vegetation change was significantly different from random. Times of change cluster around 8.2, 4.8, 3.7, and 1.2 ka, while times of higher than average stability were found around 2.1 and 5.1 ka. Compositional changes linked to the expansion of Corylus avellana and Alnus glutinosa centre around 10.6 and 9.5 ka, respectively. A climatic trigger initiating these changes may have occurred 0.5 to 1 ka earlier, respectively. The synchronous expansion of C. avellana and A. glutinosa exemplify that dispersal is not necessarily followed by population expansion. The partly synchronous, partly random expansion of A. glutinosa in adjacent European regions exemplifies that sudden synchronous population expansions are not species specific traits but vary regionally.

UV-B radiation currently represents c. 1.5% of incoming solar radiation. However, significant changes are known to have occurred in the amount of incoming radiation both on recent and on geological timescales. Until now it has not been possible to reconstruct a detailed measure of UV-B radiation beyond c. 150yr ago. • Here, we studied the suitability of fossil Pinus spp. pollen to record variations in UV-B flux through time. In view of the large size of the grain and its long fossil history, we hypothesized that this grain could provide a good proxy for recording past variations in UV-B flux. • Two key objectives were addressed: to determine whether there was, similar to other studied species, a clear relationship between UV-B-absorbing compounds in the sporopollenin of extant pollen and the magnitude of UV-B radiation to which it had been exposed; and to determine whether these compounds could be extracted from a small enough sample size of fossil pollen to make reconstruction of a continuous record through time a realistic prospect. • Preliminary results indicate the excellent potential of this species for providing a quantitative record of UV-B through time. Using this technique, we present the first record of UV-B flux during the last 9500yr from a site near Bergen, Norway. © 2011 The Authors. New Phytologist © 2011 New Phytologist Trust.

The major objectives of this paper are: (1) to review the pros and cons of the scenarios of past anthropogenic land cover change (ALCC) developed during the last ten years, (2) to discuss issues related to pollen-based reconstruction of the past land-cover and introduce a new method, REVEALS (Regional Estimates of VEgetation Abundance from Large Sites), to infer long-term records of past land-cover from pollen data, (3) to present a new project (LANDCLIM: LAND cover – CLIMate interactions in NW Europe during the Holocene) currently underway, and show preliminary results of REVEALS reconstructions of the regional land-cover in the Czech Republic for five selected time windows of the Holocene, and (4) to discuss the implications and future directions in climate and vegetation/land-cover modeling, and in the assessment of the effects of human-induced changes in land-cover on the regional climate through altered feedbacks. The existing ALCC scenarios show large discrepancies between them, and few cover time periods older than AD 800. When these scenarios are used to assess the impact of human land-use on climate, contrasting results are obtained. It emphasizes the need for methods such as the REVEALS model-based land-cover reconstructions. They might help to fine-tune descriptions of past land-cover and lead to a better understanding of how long-term changes in ALCC might have influenced climate. The REVEALS model is demonstrated to provide better estimates of the regional vegetation/landcover changes than the traditional use of pollen percentages. This will achieve a robust assessment of land cover at regional- to continental-spatial scale throughout the Holocene. We present maps of REVEALS estimates for the percentage cover of 10 plant functional types (PFTs) at 200 BP and 6000 BP, and of the two open-land PFTs “grassland” and “agricultural land” at five time-windows from 6000 BP to recent time. The LANDCLIM results are expected to provide crucial data to reassess ALCC estimates for a better understanding of the land suface-atmosphere interactions. NordForsk LANDCLIM VR LANDCLIM MERGE

The Holocene epoch, which followed the last major pulse of glaciation (the Younger Dryas) at the end of the last glaciation, encompasses a period before there was any substantial anthropogenic forcing of climate. A synthesis of climatic development during the Holocene (ca. 11,500 cal. yr BP to the present) is based on pollen-based quantitative temperature reconstructions, tree-line variations, chironomids, tree-ring records, speleothem data, glacier variations, and marine records (stable isotopes, species abundance, lithological changes) from the Nordic Seas. The Holocene has been regarded as a period of relatively stable climate, but recent evidence suggests there have been several significant millennial-scale climate fluctuations (larger than the post mid-19th century warming trend) throughout the Holocene. A general climate warming in the first part of the Holocene was punctuated by a few, abrupt climate reversals, centred at 10,000, 9,700, and 8,200 cal. yr BP. The data suggest there was a period of relatively warm conditions in the first half of the Holocene, in many areas warmer than in the 20th century, after which temperatures generally declined. The temperature decline was punctuated by centennial-scale warmer and colder periods with the most recent cold episode (∼AD 1550-1925), including the Little Ice Age, being one of the coldest of the entire Holocene. The kind of data presented here can be used for detecting mechanisms and forcing factors behind the reconstructed climate variations and to study leads and lags in the Earth's climate system.

Changes in tree-line, mean July temperature (T-jul ) and mean annual precipitation (P-ann ) for the last 10 200 cal. yr BP are reconstructed on the basis of pollen and plant macrofossils preserved in lake sediments from two sites near the present-day tree-line in Troms, northern Norway. Quantitative climate reconstructions are performed using pollen-climate transfer functions based on WA-PLS regression. Early Holocene Betula pubescens forests were gradually replaced by Pinus sylvestris at Dalmutladdo (355 m a.s.l.) starting about 7000 cal. yr BP. The local presence of pine woodland at that time is supported by finds of stomata and plant macrofossils and by high pollen accumulation rates. Until about 4000 cal. yr BP the P. sylvestris tree-line was 250-300 m higher than today, suggesting T-jul about 2.0degreesC higher than at present. The later part of the Holocene has a cooler and moister climate and an increasing development of mires and fern-rich vegetation, as shown by increases of Sphagnum and fern spores and the re-establishment of B. pubescens woodland. The reconstructed T-jul from the two sites shows similar trends to previously published data, with T-jul 1-2degreesC warmer between 9500 cal. yr BP and 2000 cal. yr BP T-jul . Maximum T-jul values occur between 8500 and 4500 cal. yr BP, after which there is a gradual decrease in T-jul.