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Climate during the last glacial period was marked by abrupt instability on millennial timescales that included large swings of temperature in and around Greenland (Daansgard–Oeschger events) and smaller, more gradual changes in Antarctica (AIM events). Less is known about the existence and nature of similar variability during older glacial periods, especially during the early Pleistocene when glacial cycles were dominantly occurring at 41 kyr intervals compared to the much longer and deeper glaciations of the more recent period. Here, we report a continuous millennially resolved record of stable isotopes of planktic and benthic foraminifera at IODP Site U1385 (the "Shackleton Site") from the southwestern Iberian margin for the last 1.5 million years, which includes the Middle Pleistocene Transition (MPT). Our results demonstrate that millennial climate variability (MCV) was a persistent feature of glacial climate, both before and after the MPT. Prior to 1.2 Ma in the early Pleistocene, the amplitude of MCV was modulated by the 41 kyr obliquity cycle and increased when axial tilt dropped below 23.5 ∘ and benthic δ18O exceeded ∼3.8 ‰ (corrected to Uvigerina), indicating a threshold response to orbital forcing. Afterwards, MCV became focused mainly on the transitions into and out of glacial states (i.e. inceptions and terminations) and during times of intermediate ice volume. After 1.2 Ma, obliquity continued to play a role in modulating the amplitude of MCV, especially during times of glacial inceptions, which are always associated with declining obliquity. A non-linear role for obliquity is also indicated by the appearance of multiples (82, 123 kyr) and combination tones (28 kyr) of the 41 kyr cycle. Near the end of the MPT (∼0.65 Ma), obliquity modulation of MCV amplitude wanes as quasi-periodic 100 kyr and precession power increase, coinciding with the growth of oversized ice sheets on North America and the appearance of Heinrich layers in North Atlantic sediments. Whereas the planktic δ18O of Site U1385 shows a strong resemblance to Greenland temperature and atmospheric methane (i.e. Northern Hemisphere climate), millennial changes in benthic δ18O closely follow the temperature history of Antarctica for the past 800 kyr. The phasing of millennial planktic and benthic δ18O variation is similar to that observed for MIS 3 throughout much of the record, which has been suggested to mimic the signature of the bipolar seesaw – i.e. an interhemispheric asymmetry between the timing of cooling in Antarctica and warming in Greenland. The Iberian margin isotopic record suggests that bipolar asymmetry was a robust feature of interhemispheric glacial climate variations for at least the past 1.5 Ma despite changing glacial boundary conditions. A strong correlation exists between millennial increases in planktic δ18O (cooling) and decreases in benthic δ13C , indicating that millennial variations in North Atlantic surface temperature are mirrored by changes in deep-water circulation and remineralization of carbon in the abyssal ocean. We find strong evidence that climate variability on millennial and orbital scales is coupled across different timescales and interacts in both directions, which may be important for linking internal climate dynamics and external astronomical forcing. [ABSTRACT FROM AUTHOR]

Past glacials can be thought of as natural experiments in which variations in boundary conditions infl uenced the character of climate change. However, beyond the last glacial, an integrated view of orbital- and millennial-scale changes and their relation to the record of glaciation has been lacking. Here, we present a detailed record of variations in the land-ocean system from the Portuguese margin during the penultimate glacial and place it within the framework of ice-volume changes, with particular reference to European ice-sheet dynamics. The interaction of orbital- and millennial-scale variability divides the glacial into an early part with warmer and wetter overall conditions and prominent climate oscillations, a transitional mid-part, and a late part with more subdued changes as the system entered a maximum glacial state. The most extreme event occurred in the mid-part and was associated with melting of the extensive European ice sheet and maximum discharge from the Fleuve Manche river. This led to disruption of the meridional overturning circulation, but not a major activation of the bipolar seesaw. In addition to stadial duration, magnitude of freshwater forcing, and background climate, the evidence also points to the infl uence of the location of freshwater discharges on the extent of interhemispheric heat transport [ABSTRACT FROM AUTHOR]

The timing and rate of ecosystem response to abrupt climate change is a product of numerous complex interactions between biotic and abiotic drivers. Palaeoecological studies from long sedimentary records, particularly those that span periods of dynamic climate such as the last glacial cycle, can help to contextualise ecosystem responses to climate variability through time. Detailed studies that compare proxy data from multiple sites, with high chronological precision, have the potential to ascribe mutual climate drivers, and, therefore, track spatiotemporal variability in ecosystem responses. Here, we interrogate the vegetation impact of past climate change in the eastern Mediterranean, using three sub-centennially resolved pollen archives from Greece. The widespread Campanian Ignimbrite (CI/Y-5; ca. 39.85 ka BP) tephra marker is used as an isochron to directly correlate pollen records from Ioannina (NW Greece), Tenaghi Philippon (NE Greece), and Megali Limni (NE Aegean). Our results reveal spatiotemporal variability in the timing of vegetation response in the Mediterranean to climate forcing across Heinrich Stadial 4 (40.2–38.3 ka BP), a period of known abrupt climatic change. We identify a decline in tree pollen in all three sites, likely related to the onset of enhanced regional aridity, with vegetation at Tenaghi Philippon responding prior to the CI/Y-5, in contrast to at Megali Limni and Ioannina, where much of the vegetation change occurs following tephra deposition. • The Campanian Ignimbrite (CI/Y-5) tephra marker is used to interrogate the timing of vegetation response to HS4. • The timing of tree pollen minima across HS4 varies between three centennial-scale pollen records. • Temporal offsets may reflect site-specific characteristics and/or the fast and slow components of millennial-scale change. [ABSTRACT FROM AUTHOR]