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Abstract This letter uses simultaneous observations from Magnetosphere Multiscale (MMS) and Time History of Events and Macroscale Interactions during Substorms (THEMIS) to address the dynamics of the magnetopause and magnetosheath boundary layers during the main phase of a storm during which the interplanetary magnetic field (IMF) reverses from south to north. Near the dawn terminator, MMS observes two boundary layers comprising open and closed field lines and containing energetic electrons and ring current oxygen. Some closed field line regions exhibit sunward convection, presenting an avenue to replenish dayside magnetic flux lost during the storm. Meanwhile, THEMIS observes two boundary layers in the pre‐noon sector which strongly resemble those observed at the flank by MMS. Observations from the three THEMIS spacecraft indicate the boundary layers are still evolving several hours after the IMF has turned northward. These observations advance our knowledge of the dynamic magnetopause and magnetosheath boundary layers under the combined effects of an ongoing storm and changing IMF.