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The avian fossil record of southern Brazil is scarce, consisting of few isolated remains. Here is described a ciconiid fossil from the assemblage of mammalian remains of the Santa Vitoria Formation (SVF). The specimen is a cervical vertebra morphologically similar to Ciconia maguari (maguari stork), common in southern Brazil today. Nevertheless, its larger dimensions suggest that it could be either a morphotype of that species or another, extinct species. The taphonomic modifications of the vertebra result of a combination of autogenic and allogenic factors including the aquatic habit, shape and transportation by flowing water and burial in a shallow stream. The fossil-bearing sediment dated by luminescence was deposited during the late Pleistocene, at 37.9 ka b2k, and exhibits physical features indicating deposition under variable discharge and seasonal oscillations of the water table. The presence of an aquatic bird in these sediments indicates a period of increased precipitation, chronocorrelated to one of the millenial-scale warming pulses recorded in ice cores from Antarctica (Antarctic Isotope Maxima) that characterized the interstadial MIS 3. On the other hand, the dry and cold climate of the following glacial MIS 2 may have forced ciconiids and aquatic mammals to retreat to suitable areas (refugia), as indicated by their absence in the loess deposits (Cordao Formation) overlying the SVF, until returning during the Holocene when climate became wet and warm again. The results presented here increase the avian fossil record of southern Brazil and help understand the role of climate change on the distribution of ciconiid birds and other taxa during the Quaternary in southern South America.

Here are described post mortem modifications (pseudopaleopathologies) in 762 specimens of extinct mammals from three middle to late Pleistocene assemblages (Campanha, Chui Creek and continental shelf) of the state of Rio Grande do Sul, southern Brazil. These assemblages include mostly large-sized herbivores preserved in fluvial facies, although the assemblage from the shelf consist of remains that were reworked several times by sea-level oscillations. Although some patterns of modification are similar between the assemblages (e.g. post-fossilization plain fractures), others such as the grade of abrasion or weathering varies according to the depositional setting. The scarcity of modifications that could be attributed to predators or scavengers indicate that carnivory was not a major process driving the formation of the assemblages. Most of the fossils from the Campanha are complete, but a relatively high proportion of them exhibits advanced stage of weathering, whereas those from Chui Creek and the continental shelf are mostly unweathered. Most of the remains from the Campanha and Chui Creek are unabraded, whereas all fossils from the shelf were subject to abrasion. Few fossils exhibit signs of modification by organisms such as plants and insects. Fossils from the continental shelf have crusts of beach rock, and some specimens from the Campanha and Chui Creek have crusts of calcium carbonate or iron oxide. Besides the abrasion, the main difference between fossils from the continental and marine assemblages is the dark color and higher mechanical resistance of the latter. This difference is related to post depositional geochemical processes in the marine environment. The chemical composition of fossils from Chui Creek and continental shelf was qualitatively analyzed using energy dispersive X-ray spectrometry (EDX) and compared with recent bones. The results show the presence in the fossils of several elements not found in recent bones, incorporated through diagenetic processes controlled by specific conditions of each depositional environment.

Sacisaurus agudoensis is the only silesaurid known from the Triassic beds of the Santa Maria Supersequence and the correlation of its type locality to the other Triassic deposits of south Brazil has always been controversial. In an attempt to improve this, a handful of dinosaur and cynodont remains found associated to S. agudoensis are here described and compared. The anatomy of the sauropodomorph is more similar to that of Norian forms such as Pantydraco caducus and Unaysaurus tolentinoi than to that of Carnian taxa such as Saturnalia tupiniquim and Pampadromaeus barberenai. The cynodonts recovered based on isolated teeth include a brasilodontid and a Riograndia-like form. This assemblage is consistent with a Norian age, as is also suggested by local stratigraphic correlation, which positions the site in the Caturrita Formation. O Sacisaurus agudoensis é o único silesaurídeo conhecido dos leitos Triássicos da Supersequência Santa Maria e da a correlação de sua localidade-tipo com os demais depósitos triássicos do sul do Brasil sempre foi controversa. Numa tentativa para melhorar isto, são aqui descritos alguns restos de dinossauros e cinodontes encontrados associados a S. agudoensis e comparado. A anatomia do sauropodomorfo é mais semelhante à das formas Norian, como Pantydraco caducus e Unaysaurus tolentinoi do que aos táxons Carnianos como Saturnalia tupiniquim e Pampadromaeus barberenai. Os cinodontes recuperados com base em dentes isolados incluem um brasilodontídeo e uma forma do tipo Riograndia. Esta montagem é consistente com uma idade Norian, como também é sugerido pela correlação estratigráfica local, que posiciona o sítio no Formação Caturrita.

The Sanga do Cabral Formation of southern Brazil has a rich fossil tetrapod assemblage and is suggested to have an Early Triassic age mainly based on the presence of the parareptile Procolophon trigoniceps . However, a Permo-Triassic age can be also suggested for this unit taking into account the presence of putative Permian taxa and some previous stratigraphic assessments. We describe here several large vertebrae from the Sanga do Cabral Formation that display a distinctive morphology that includes the presence of a transverse distance across postzygapophyses more than twice the transverse width of the centrum, and accessory articulation structures in the neural arch that remind the hyposphene and hypantrum present in some basal parareptiles and diadectomorphs. Vertebrae with a similar large size and morphology had been previously reported from the same locality as belonging to the genus Procolophon based on their parareptile appearance (mainly the presence of a swollen neural arch) and the fact that the vertebrae were collected at the same locality where a large fragmentary skull assigned to this taxon was found. However, these vertebrae lack a comparable consistent morphology with those of Procolophon and basic statistical analyses demonstrate that these vertebrae are significantly larger than those expected in the largest known Procolophon skulls of South Africa. The morphology of these vertebra is consistent with that present in seymouriamorphs, pareiasaurs and diadectomorphs, but the absence of exclusive diagnostic characters precludes an assure assignation to either of these taxa. According to their current stratigraphic range, seymouriamorphs are the most plausible postulation, as their younger representatives are known from Late Permian deposits of Russia, but the other candidates cannot be excluded. The presence of any representative of those groups in the Sanga do Cabral Formation would be important because: (1) they would represent the first and only known record of seymouriamorphs or diadectomorphs in Gondwana, regarding the characters that the described vertebrae share with these groups; (2) they would suggest a survivorship for pareisaurs up to the Latest Permian or through the Permian-Triassic boundary, according to geochronological data currently available for this unit; (3) they might also suggest a Late Permian age for at least part of the Sanga do Cabral Formation if the intraformational conglomerates that yielded the vertebrae resulted from the rejuvenation of older levels of the same unit, and they do not include reworking of stratigraphically older strata.

Unlike most rauisuchians, which are known based on partially preserved specimens, fossils attributed to Decuriasuchus quartacolonia include a monotaxonomic assemblage com- posed of nine associated individuals (MCN-PV10.105a-i), three with almost complete skulls (MCN-PV10.105a,c,d), and a partial disarticulated skull (MCN-PV10.004) collected in the Mid- dle Triassic (Ladinian, Dinodontosaurus Biozone) beds of the Santa Maria Formation, in south Brazil. Because of its completeness and possible phylogenetic position, as one of the most basal loricatans, D. quartacolonia is a key taxon for anatomic, evolutionary and biomechanical stud- ies of rauisuchians. The comparative description of its osteology reveals that the skull and mandi- ble of D. quartacolonia are very similar to those of cf. Prestosuchus chiniquensis and Saurosuchus galilei, sharing a drop-shaped subnarial fenestra, a subtriangular antorbital fenestra with an elon- gated and narrow anterior point, a 'roman nosed' nasal, and a posteroventrally oriented ridge on the lateral surface of the ventral ramus of the squamosal. Among the differences are the autapomor- phies of D. quartacolonia: numerous maxillary teeth (17), lateral expansion of the nasal/lacrimal covering the antorbital fenestra dorsally, and squamosal and quadratojugal forming a subtriangular projection that invades the lower temporal fenestra. The study of rauisuchians began in the middle of the 20th century, when the German palaeontolo- gist Friedrich von Huene discovered archosaur fossils in Brazil. Huene (1938b, 1942) described four new taxa currently considered as rauisuchians: Prestosuchus chiniquensis, Prestosuchus loricatus, Procerosuchus celer and Rauisuchus tiradentes. The generic name of the latter taxon, in honour of Guilherme Rau, a Brazilian-German dentist who collaborated with Huene, achieved family and class status in later works (e.g. Huene 1956; Krebs 1976; Bonaparte 1984; Parrish 1993; Brusatte et al. 2010; Nesbitt 2011). Since then, several fur- ther rauisuchian remains have been discovered in Brazil, including rather complete specimens such as that of cf. Prestosuchus chiniquensis (UFRGS-PV0156T; Barberena 1978). However, more recently, about 70 years after Huene's discov- eries, a new taxon was proposed (Franca et al. 2011a). Named Decuriasuchus quartacolonia, the taxon was based on a monotaxonomic assemblage

The South American Teiidae fossil record is restricted to the Cenozoic, and the most conspicuous remains were found in Early to Late Miocene of Argentina and Middle Miocene of Colombia and Peru, all represented by Tupinambinae lizards. Here, we describe a right fragmentary dentary and one dorsal vertebra collected in the Solimoes Formation at the Talisma locality, situated on the Purus River, in the southwestern Brazilian Amazonia (Late Miocene). The material is tentatively conferred to the extinct genus Paradracaena. It represents the first record of lizards for the Neogene southwestern Brazilian Amazonia.

The Triassic of Southern Brazil is well-known for its rich tetrapod body fossils. However, trace fossils such as tetrapod trackways and trails were discovered only recently from the Predebon outcrop (upper part of the Alemoa Member, Santa Maria Formation), providing new information on small-sized animals. The trackways can be identified as Rhynchosauroides retroversipes isp. nov., Rhynchosauroides isp., Rhynchosauroides? isp. and drag marks produced by swimming animals of lacertoid affinity. The preservation of the tracks and trackways was influenced by the water level whilst they were being made and subsequent subaerial exposure. The trackmaker of R. retroversipes isp. nov. corresponds to a lacertoid quadruped tetrapod with primitive autopodia, a sprawling gait and possibly a long tail, whilst the pes was directed postero-laterally. The trackmaker may have been a climbing animal, with adaptations that include curved claws and reversion of the pes. On the ground this animal would have had a slow, lumbering locomotion, although it could attain higher speeds over short distances using a bipedal gait. These trackways can be attributed to lacertoid reptiles, possibly sphenodontids, whose skeletons are encountered in the Caturrita Formation, which overlies the Santa Maria Formation.

The South American Late Triassic offers the most comprehensive window to the early radiation of dinosaurs. This is enhanced by the discovery of Sacisaurus agudoensis, a new dinosauriform from the Caturrita Formation of Brazil. Various morphological features suggest its close phylogenetic affinity to Silesaurus, and both may be basal ornithischian dinosaurs. Sacisaurus has a pair of elements forming the tip of its lower jaw, hypothesized to be equivalent to the ornithischian predentary. This suggests that during an initial stage of their evolution, those dinosaurs had a paired predentary, which later fused into a single structure. As an originally paired bone, the predentary is comparable to elements that more often form the vertebrate mandible, such as the mentomeckelian bone. Although synapomorphic for ornithischians, the predentary does not seem neomorphic for the group, but primarily homologous to parts of the symphyseal region of the lower jaw of other vertebrates.

In their monograph Review of the Pelycosauria, Romer and Price (1940), proposed that the earliest synapsids (“pelycosaurs”) were cosmopolitan, despite the observation that amniotes appeared to be restricted to the paleotropics during the Late Carboniferous and Early Permian (290–282 Ma). Romer and Price (1940) accounted for the scarcity of terrestrial tetrapods, including “pelycosaurs,” in Lower Permian beds elsewhere to the absence of coeval continental deposits beyond North America and Europe. Indeed, most workers recognized a geographical and temporal gap between Permo-Carboniferous “pelycosaurs” and therapsid synapsids. Recent research has confirmed that varanopid and caseid “pelycosaurs” were components of therapsid-dominated Late Permian faunas preserved in Russia and South-Africa (Tatarinov and Eremina, 1975; Reisz, 1986; Reisz et al., 1998; Reisz and Berman, 2001). In this note, we report the first record of basal synapsids for South America (Fig. 1) and discuss its paleogeographic and temporal implications. These remains come from outcrops assigned to the Buena Vista Formation (Goso et al., 2001; Fig. 2), which filled part of the Parana Basin during Late Permian and probably Early Triassic (Bossi and Navarro, 1991). The Parana Basin is one of the major sedimentary basins of Gondwana, covering an area of nearly 1.5 million km2, in Brazil, Argentina, Paraguay and Uruguay (Franca et al., 1995; Zalan et al., 1990). At the beginning of basin development the Devonian marine and brackish conditions prevailed, but subsequently, during the Late Permian, there was a gradual change to continental sedimentation and the deposition of fossil-bearing red beds. Buena Vista Formation is characterized by reddish fine sandstone interbedded with lenticular clay layers and intraformational conglomerates (Fig. 2), (Bossi and Navarro, 1991; Goso et al., 2001). The paleoenvironment is related to the final withdrawal of the sea, and the upper part of this …

The coastal plain of the state of Rio Grande do Sul, in southern Brazil, includes four barrier-lagoon depositional systems formed by successive Quaternary sea-level highstands that were correlated to marine isotope stages (MIS) 11, 9, 5 and 1, despite the scarcity of absolute ages. This study describes a sea-level highstand older than MIS 5, based on the stratigraphy, ages and fossils of the shallow marine facies found in coastal barrier (Barrier II). This facies outcrops along the banks of Chuí Creek, it is composed of fine, well-sorted quartz sand and contains ichnofossils Ophiomorpha nodosa and Rosselia sp., and molluscan shells. The sedimentary record indicates coastal aggradation followed by sea-level fall and progradation of the coastline. Thermoluminescence (TL) and electron spin resonance (ESR) ages from sediments and fossil shells point to an age of ∼220 ka for the end of this marine transgression, thus correlating it to MIS 7 (substage 7e). Altimetric data point to a maximum amplitude of about 10 meters above present-day mean sea-level, but tectonic processes may be involved. Paleoceanographic conditions at the time of the highstand and correlations with other deposits in the Brazilian coasts are also discussed. A planície costeira do estado do Rio Grande do Sul, no sul do Brasil, inclui quatro sistemas deposicionais do tipo laguna-barreira formados por sucessivas transgressões marinhas do Quaternário, correlacionadas aos estágios isotópicos marinhos (MIS) 11, 9, 5 e 1, apesar da escassez de idades absolutas. Este estudo descreve um máximo transgressivo marinho mais antigo que o MIS 5, com base na estratigrafia, datações e fósseis encontrados na fácies marinho raso de uma barreira costeira (Barreira II). Esta fácies encontra-se exposta ao longo das barrancas do Arroio Chuí, é composto por areia quartzosa fina, bem-selecionada, com icnofósseis Ophiomorpha nodosa e Rosselia sp., e conchas de moluscos. O registro sedimentar indica agradação da costa seguida por regressão do nível do mar e progradação da linha de costa. Datações por termoluminescencia (TL) e ressonância do spin do elétron (ESR) em sedimentos e conchas fósseis indicam uma idade de ∼220 ka para o final desta transgressão, correlacionando-a, portanto, ao MIS 7 (sub-estágio 7e). Dados altimétricos indicam amplitude máxima de aproximadamente 10 metros acima do nível atual do mar, mas processos tectônicos podem estar envolvidos. Condições paleoceanográficas à época da transgressão e correlações com outros depósitos da costa brasileira também são discutidos.

A primitive ‘ictidosaur’ from lower Norian beds of southern Brazil, Riograndia guaibensis gen. et sp. nov., represented by a fragmentary skull and a lower jaw bearing a complete dentition, shows a more generalized morphology than Chaliminia from the Upper Triassic of Argentina and PachygenelusDiarthrognathus from the Lower Jurassic of South Africa, Canada and Greenland. The frontal bone borders the orbit, and ventrally contacts the dorsal process of the palatine. The secondary bony palate extends back to the last postcanine. I1 and i2 are reduced, whereas I2-3 and i1 are hypertrophied. Both PC 1–7 and pc 1–7 have blade-like crowns without cingula and with 5–9 small sharp cuspules. The upper postcanine crowns are semicircular in labial view with the cuspules around their margins. The lower postcanine crowns are asymmetrical with most of the cuspules dorsodistally distributed. The possible origin of this peculiar dentition is interpreted as the retention of the juvenile dentition of ancestors. The hypothesis that Riograndia guaibensis and the so-called ‘ictidosaurs’ might have been derived from gomphodont cynodonts is presented.

This paper describes a possible fossil egg proceeding from the Bauru Group, Late Cretaceous of Paraná Basin, Brazil. External morphology, dimensions, texture and shell ornamentation examined under electron microscopy show close resemblance to the Recent podocnemidid chelonian eggs. Association with bony material in the outcrop suggests that it is related to a species of Podocnemis. Computerized tomography reveals a high density outer region corresponding to the shell component layers, and successive layers with decreasing density towards the nucleus. An area of high density in the central region may represent remains of an embryo.