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13 pages; International audience; Kisima Ngeda (KN), a spring on the northern margin of saline Lake Eyasi, Tanzania, sustains an Acacia-Hyphaene palm woodland and Typha swamps, while the surrounding vegetation is semi-desert. To study the vegetation changes associated with this spring, which represents a plausible modern analog for the fossil springs documented in the nearby paleoanthropological and archaeological sites of Olduvai Gorge, we analyzed the pollen content of a 43 cm-long sediment core that documents vegetation changes since the last ~1200 years (from cal yrs. C.E. 841 to 2011). Our results show that (1) Hyphaene palms, which require meso-halophytic soil conditions were most abundant in the area of the coring site until cal yrs. C.E. ~1150 when the groundwater supplying the KN spring was likely lower than at present, allowing intrusions of saline lake waters. (2) From cal yrs. C.E. ~1200, a peat began to develop, the palm woodland was replaced by a Mimosaceae woodland, and the increased presence of Typha pollen indicates the presence of more wetlands. (3) From cal yrs. C.E. 1600, the groundwater level of the KN spring increased and reached its highest level in the last 1200 years. (4) Peaks of wetland expansion, which reflect increased groundwater flow and level in response to amplified rainfall in the recharge area (Mt Oldeani, Ngorongoro Highlands), occurred at cal yrs. C.E. ~1200–1400 and ~1650–2011. These outflows of groundwater at Kisima Ngeda were linked to the intensity and frequency of positive Indian Ocean Dipole (IOD) events, which trigger heavy rains in eastern Africa. We conclude that the Kisima Ngeda hydrological system, which has been active for more than 1200 years, responds rapidly to regional climate change driven by changes in the sea surface temperatures (SSTs) of the Indian Ocean. Yet, it is also capable of remaining active during dry intervals as inferred from the Kisima Ngeda record prior to cal yrs. C.E. 1200. Our results support the hypothesis that this type of system helped to maintain Plio-Pleistocene hominin populations and activities in the arid lowlands of the rift on a multi-decennial scale.

International audience; Pliocene and Pleistocene sediments from West Turkana (Kenya, East African Rift System) form emblematic syn-rift successions for understanding the evolution of extensional basin and continental rifting. They also constitute world-renowned fossil-bearing strata from which >500 hominin fossils were discovered over the past decades, with >100 inventoried archeological sites. However, associated sedimentary dynamics and architectures as well as paleoenvironments are only partially reconstructed and the relative contribution of climate and tectonism to paleoenvironmental change over time remains unclear. Here, through the interpretation of sedimentary facies, the delineation of sequences and the analysis of δ13C in soil carbonates, we provide the first exhaustive reconstruction from ~4 to ~0.75 Ma of (i) fluctuations of the paleolake that occupied the Turkana Depression, (ii) the successive sedimentary dynamics and related paleolandscapes that characterized the West Turkana area and (iii) respective roles of climate and tectonism in the sedimentation. We show evidence for seven major transgression-regression (T-R) cycles between ~4 and ~ 0.75 Ma superimposed locally by lower amplitude T-R cycles., Comparing the sedimentological interpretations and the δ13C values in soil carbonates (literature data), we reveal that fluctuations of rainfall over the Ethiopian Dome, which hosts the drainage basin of the Omo River — the main tributary of Lake Turkana — controlled high-amplitude lake level fluctuations during the Plio-Pleistocene period. We also demonstrate that vegetation and tree cover evolved differently in the Omo Valley and West Turkana. Furthermore we outline that two different sedimentary systems reflecting two distinct modes of sedimentation alternated through times in the West Turkana area as a response to the variations in sediment supply coming from the western rift shoulder (i.e. Lapurr Range) that alternatively generated wave- or river-dominated sedimentary systems. In conclusion, we reveal that climate regulated water input, paleolake water-level fluctuations and vegetation. Tectonism determined sediment supply to the basin controlled in West Turkana by pulses of increased activity of the main border fault (i.e., the Murua Rith Lapurr Fault).

International audience; Understanding Plio-Pleistocene paleoenvironments in the East African Rift and how they have changed over time is fundamental across geosciences, the most emblematic being perhaps their role in hominin evolution. In the northern Turkana Depression, among the three main Formations of the Omo Group, the Nachukui Formation is exposed on the western shore of present-day Lake Turkana. It consists of a > 700 m succession of fluvial-deltaic-paralic-lacustrine sediments deposited between ca. 4.00 and ca. 0.75 Ma, which have been extensively investigated using depositional environments and sequence analyses during the RiLakS project. Additionally, we use δ13C isotopic ratios in soil carbonates as a proxy of paleovegetation.Facies and sequence analyses reveal that proto-lake Turkana experienced 7 high-amplitude Transgression-Regression (T-R) cycles between ca. 4.00 and 1.25 Ma, which are locally superimposed by lower amplitude, secondary T-R cycles. Paralic or offshore depositional environments are observed quasi-continuously, at least in basinward sections, which suggest a more prolonged persistence of lacustrine conditions than hitherto accepted. Remarkable match between reconstructed evolution of palaeolake level and evolution of woody cover in the Lower Omo Valley suggests that rainfall in the drainage basin of the Omo river, i.e. rainfall over the Ethiopian dome, controlled high-amplitude paleolake fluctuations during the Plio-Pleistocene whereas rainfall in West Turkana had only limited influence. Additionally, we reveal that variation in sediment supply from the rift shoulder caused two different sedimentary dynamics, i.e. accommodation-dominated and supply-dominated systems, to alternate during the deposition of the Nachukui Formation, likely due to successive pulses of activity of the border fault.Finally, a reconstruction of paleolake fluctuation in the Turkana Depression is proposed and successive sedimentary paleolandscapes are presented. Respective influences of climate (local to regional) and tectonic (i.e. border fault activity) on paleoenvironments are deconvoluted. Our work highlights that the continuous reconstruction of depositional environments at various localities in a well-constrained chronological context is essential to reliably reconstruct paleoenvironments. However, until now, such powerful approach remains rarely associated with traditional site by site investigations.

The late Pliocene corresponds to a large cooling over Northern Hemisphere associated with sporadic occurrences of glaciations. The most important event occurred during the marine isotope stage M2 (MIS M2, 3.312–3.264 Ma) when a large glaciation took place with a sea level drop from 20 to 60 m, but its duration is short and the summer insolation forcing change at 65°N is weak. De Schepper et al (2013) invoked to explain the onset and termination of this glaciation with the opening and closing of the Central American Seaway (shallow CAS). Based on their hypothesis, we have intensively studied the onset mechanism of MIS M2 through a series of sensitivity experiments using the IPSL AOGCM and the asynchronous coupling with an Ice sheet model (GRISLI). Our results demonstrate that the shallow CAS helps to precondition the low-latitude oceanic circulation and affects the related northward energy transport, but cannot alone explain the onset of the M2 glaciation, the most important contribution on MIS M2 are from the large change of pCO2 as well as the internal feedbacks of vegetation and ice sheet. Moreover, we have also investigated the period from the late Pliocene to the early Pleistocene (3-2.5 Ma) through a transient-like simulation using the same AOGCM and ISM. This enables to simulate the Greenland Ice Sheet (GRIS) onset and development using the pCO2 reconstructions from different proxies. All these simulations were analyzed with emphasis on cryosphere and focused on the Northern Hemisphere (mid-to-high latitudes). Here we used the same modeling simulations but with a focus over the tropical Africa. We first depict the large changes of temperatures and hydrological cycle produced over this area during these two periods and compare our data to reconstructions. Moreover, by prescribing our climate results as inputs for the vegetation model (Biome4), we compare more directly the simulated plant functional types (PFTs) with that constructed by the pollen data. In addition, we further quantify the respective impact of various driving factors on these PFTs variations.

The Turkana Depression consists of several Oligocene to Pliocene North-South oriented half-grabens that connect the Ethiopian and Kenyan rift valleys within the eastern branch of the Cenozoic East African Rift System. In the northern portion of the Turkana Depression, exposed on the west side of modern Lake Turkana, is the Nachukui Formation that consists of a > 700 m pile of fluvial-deltaic-lacustrine sediments deposited between 4.2 and 0.7 Ma. The Nachukui Fm is a world-class fossil-bearing succession into which more than 500 hominin fossils were discovered, including major discoveries for the understanding of Human evolution and more than 100 archaeological sites. Most significant discoveries include Australopithecus anamensis, Kenyanthropus platyops, Paranthropus aethiopicus, Paranthropus boisei and specimens of Homo (i.e., H. rudolfensis and H. erectus) and early members of H. sapiens, as well as the earliest evidence of Acheulean stone tool technology and, more recently, the most primitive Lomekwian stone tool technology. Palaeoenvironmental changes may have had a strong influence on evolution, including that of the human lineage. However, in the Turkana Depression, palaeoenvironments are still very partially reconstructed and the respective role of climate and tectonism is still debated. Here, we used the interpretation of depositional environments, the delineation of depositional sequences and a record of d13C in pedogenic carbonates (i.e. paleovegetation proxy) to reconstruct 1) palaeolake Turkana fluctuations between ca. 4 and ca. 1.2 Ma and 2) the successive sedimentary palaeoenvironments and resulting landscapes that characterized the West Turkana area during the same time interval. Facies and sequence analyses reveal that palaeolake Turkana experienced eight low-frequency transgression-regression (T-R) cycles between ca. 4 and ca. 1.2 Ma; superimposed lower amplitude and shorter duration T-R cycles are also locally identified revealing subordinate-order fluctuations. In the same time, two different palaeolandscapes (labelled type-1 and type-2) alternated through times revealing variations in sediment supply coming from the western rift shoulder. A statistical treatment of the d13C record using a modified k-mean clustering allows us to confront a paleovegetation proxy and the sedimentological record. This combined approach reveals that the evolution of rainfall over the Ethiopian dome (i.e., drainage basin of the Omo river) controlled long-term palaeolake Turkana fluctuations during the Plio-Quaternary period while tectonism likely controlled the occurrence of different palaeolandscapes in West Turkana forced by changes in the rate of sediment supply. Finally, our study shows that traditional methods of basin geology (i.e., facies and sequence analysis) are key tools to provide large-scale paleolandscape reconstructions and palaeolake fluctuations needed for investigating the interactions between hominins and palaeoenvironments. Such a powerful procedure, however, is rare for hominins sites and has yet to be integrated in the workflow utilized by the paleontology and archeology communities.This is a contribution of the Rift Lake Sedimentology project (RiLakS).

International audience; A fossil hominin cranium was discovered in mid-Pliocene deltaic strata in the Godaya Valley of the northwestern Woranso-Mille study area in Ethiopia. Here we show that analyses of chemically correlated volcanic layers and the palaeomagnetic stratigraphy, combined with Bayesian modelling of dated tuffs, yield an age range of 3.804 ± 0.013 to 3.777 ± 0.014 million years old (mean ± 1σ) for the deltaic strata and the fossils that they contain. We also document deposits of a perennial lake beneath the deltaic sequence. Mammalian fossils associated with the cranium represent taxa that were widespread at the time and data from botanical remains indicate that the vegetation in the lake and delta catchment was predominantly dry shrubland with varying proportions of grassland, wetland and riparian forest. In addition, we report high rates of sediment accumulation and depositional features that are typical of a steep topographic relief and differ from younger Woranso-Mille fossil localities, reflecting the influence of active rift processes on the palaeolandscape. Q1 Until recently, field work in the Woranso-Mille area (Afar, Ethiopia) has concentrated on exposures along the Mille River that include more than 120 m of fossiliferous sedimentary and volcanic strata that are between 3.8 and 3.2 million years (Myr) old. East of Korsi Dora (Fig. 1a), sedimentary sequences that are younger than the extra-regional approximately 3.57-Myr-old Kilaytoli tuff 1 have yielded specimens of at least two hominin species, Australopithecus afarensis and Australopithecus deyiremeda 2-5

International audience; At present, Lake Chad (∼ 13 • N, ∼ 14 • E) is a shallow freshwater lake located in the Sahel/Sahara region of central northern Africa. The lake is primarily fed by the Chari-Logone river system draining a ∼ 600 000 km 2 watershed in tropical Africa. Discharge is strongly controlled by the annual passage of the in-tertropical convergence zone (ITCZ) and monsoon circulation leading to a peak in rainfall during boreal summer. During recent decades, a large number of studies have been carried out in the Lake Chad Basin (LCB). They have mostly focused on a patchwork of exposed lake sediments and outcrops once inhabited by early hominids. A dataset generated from a 673 m long geotechnical borehole drilled in 1973, along with outcrop and seismic reflection studies, reveal several hundred metres of Miocene-Pleistocene lacustrine deposits. CHADRILL aims to recover a sedimentary core spanning the Miocene-Pleistocene sediment succession of Lake Chad through deep drilling. This record will provide significant insights into the modulation of orbitally forced changes in northern African hydroclimate under different climate boundary conditions such as high CO 2 and absence of Northern Hemisphere ice sheets. These investigations will also help unravel both the age and the origin of the lake and its current desert surrounding. The LCB is very rich in early hominid fossils (Australop-ithecus bahrelghazali; Sahelanthropus tchadensis) of Late Miocene age. Thus, retrieving a sediment core from this basin will provide the most continuous climatic and environmental record with which to compare hominid migrations across northern Africa and has major implications for understanding human evolution. Furthermore, due to its dramatic and episodically changing water levels and associated depositional modes, Lake Chad's sediments resemble maybe an analogue for lake systems that were once present on Mars. Consequently, the study of the subsurface biosphere contained in these sediments has the potential to shed light on microbial biodiversity present in this type of depositional environment. We propose to drill a total of ∼ 1800 m of poorly to semi-consolidated lacustrine, fluvial, and eolian sediments down to bedrock at a single onshore site close to the shoreline of present-day Lake Chad. We propose to locate Published by Copernicus Publications on behalf of the IODP and the ICDP. 72 F. Sylvestre et al.: The Lake CHAd Deep DRILLing Project (CHADRILL) our drilling operations onshore close to the site where the geotechnical Bol borehole (13 • 28 N, 14 • 44 E) was drilled in 1973. This is for two main reasons: (1) nowhere else in the Chad Basin do we have such detailed information about the lithologies to be drilled; and (2) the Bol site is close to the depocentre of the Chad Basin and therefore likely to provide the stratigraphically most continuous sequence.

More than 40 years of scientific investigations of the hominin bearing Plio-Pleistocene sediments of northern Tanzania have provided a number of paleobotanical data, which, taken as a whole, provide today a way to investigate vegetation changes between 4 and 1 Ma, at a time when our early ancestors emerged. Here, I have integrated the data from all vegetation proxies obtained for the paleontological sites of Laetoli, Olduvai, and Peninj (i.e. macroscopic plant remains, pollen and phytolith assemblages, carbon and oxygen isotopic ratios measured on carbonates, and organic biomarkers). This important, yet discontinuous botanical record suggests some similarities between past and present-day vegetation at the regional scale: Afromontane forests with Olea , Podocarpus , Juniperus , Hagenia abyssinica in the highlands, and wooded grasslands with grasses and drought-adapted Acacia , Commiphora , Capparidaceae, and Chenopodiaceae and/or Amaranthaceae in the lowlands were present in the southern Serengeti–Crater Highlands region since 4 Ma. Grasses of the C 4 photosynthetic type made their first appearance in the record at ∼3.7 Ma, i.e. during the mid-Pliocene, ∼700 ky before major p CO 2 and temperature decline. C 4 grasses became dominant in the vegetation soon after (∼3.66 Ma), probably in response to reduced precipitation. At ∼2.6 Ma, phytolith and isotope indicate grassland with abundant C 4 xerophytic grasses that document strong aridity during the Intensification of Northern Hemisphere Glaciation period (∼2.7–2.5 Ma). After 2 Ma, the detailed and diverse record at Olduvai indicates complex vegetation patterns linked to oscillating precipitation, varying lake levels, and the presence of geological faults. Hence, despite low (basin-averaged) reconstructed paleo-precipitation amounts of ∼250–700 mm/y, C 4 -grasslands, closed woodlands, wetlands, and palm-groves co-occurred on short spatial scales near saline Lake Olduvai. Freshwater wetlands and palm-rich woodlands occurred in highly localized areas on the lake margins, where aquifer barriers and/or outcrops caused by geologic faults allowed groundwater discharge. Botanical evidence of paleo-springs at Laetoli, Olduvai, and Peninj includes the marked presence of Typha (cattail) and Hyphaene (palm tree) in association with Acacia pollen grains, and (at Olduvai) abundant forest indicator phytoliths and organic and isotopic biomarkers. At Olduvai, freshwater wetlands were most developed when lake level and fluvial competence where low, i.e. during dry periods. When wet–dry climate variability was extreme in East Africa (∼1.9–1.7 Ma), freshwater springs may therefore have offered a sustainable habitat (i.e. refuge) for several species, including hominins, and favored hominin and artifact concentration at these specific places, particularly during environmentally stressful times.

We have analyzed the phytolith content of 37 samples from the paleosol capped by Tuff IC dated 1.84 Ma within Middle Bed I at Olduvai Gorge, Tanzania. Samples were obtained from different trenches sampling an area of about 25 ha to document the paleovegetation associated with the abundant faunal remains and hominin artifacts in FLK, AMK, PTK and DS sites. The analysis reveals the abundance of globular granulate and forest indicator phytoliths (from 3% to 86% relative to the total number of phytoliths in the assemblages) attributed to woody plants, and of short cell grass phytoliths (up to 22%). The presence of palm phytoliths (up to 57%) attests for meso-haline soils in the area in agreement with the presence of the saline/alkaline Olduvai paleolake and one (or more) groundwater discharge areas (freshwater springs and/or wetland). Phytoliths provide botanical evidence for a mixed paleovegetation dominated by forest and woodland with patches of open-spaces occupied by grasses, prior to the deposition of Tuff IC. The presence of fern phytoliths in some assemblages suggest shaded and humid habitat, which supports the presence of freshwater (watercourses or springs) that might have attracted hominins and other animals. Yet, we have found that our phytolith extractions (residues) also include micrometric zeolites (chabazite, clinoptilolite, phillipsite), which attest that the phytolith signal in the Zinj paleosols is certainly affected by dissolution. This study completes the spatial analysis of the area and reveals that the wooded area extends north, south, and west of FLK Zinj, with palm trees preferentially located in the periphery, like the Zinj, AMK, and DS sites. The phytolith-inferred distribution of plants does not support the hypothesis that topography was influential in the location of these sites, particularly given that trees and/or shrubs were dominant all over. Rather, it is the presence of freshwater ponds that seem to have been the attractive factor for hominins at this particular place and time.

AMK (Amin Mturi Korongo) is a newly discovered site situated under Tuff IC (Bed I, 1.84 Ma). It contains several fossiliferous levels and the top one is situated on the same paleosurface as FLK-Zinj. For the first time this allows sampling the “Zinj” paleoenvironment well into the Secondary Gorge and expands the known area of this paleolandscape. Fossils found at this site show exceptional preservation. Several articulated units have been discovered, indicating minimal postdepositional disturbance and rapid sedimentation. This assemblage allows a general estimation of time span (the most elusive variable in archaeological analyses) for the formation of AMK. Phytolith analyses have discovered a dense palm forest at the site, expanding the forested area known on the slightly elevated platform that contains the FLK-Zinj – FLK-NN – PTK sites. Although a few artifacts have been discovered in the vicinity of AMK, the site was mostly naturally (i.e., non-anthropogenically) formed. This is of major relevance to determine that factors other than forested habitats must have influenced the formation of anthropogenic sites on the same platform as AMK in the Olduvai lacustrine basin.

Paleovegetation studies are essential to ecologically frame hominin evolution. The analysis of the phytolith content of 22 paleosol samples collected along the vertical sequence of the channel containing the BK (Bell's Korongo) archaeological levels, provided evidence of the paleovegetation that accompanied the evolution of the riverine system that formed this site 1.353 million years ago, when climate was particularly arid. The analysis reveals the abundance of forest indicator phytoliths (46% to 92% of the assemblages), rare grass silica short cells and sedges phytoliths (up to 15% of assemblages), and palms (up to 10% of the phytolith assemblages). The vegetation of the BK fluvial system was therefore characterized by abundant woody plants. It did not vary significantly diachronically in the period of time under scrutiny. The alluvial paleovegetation corresponds well to the fluvial dynamic that formed BK, including the presence of sterile samples recovered in areas where the water traction changed the soils frequently. This spot of dense paleovegetation together with the fluvial watercourse (in a paleolandscape likely dominated by sparse/open vegetation) could have been attractive, seasonally, for animals and hominins.

We analyzed phytolith and diatom remains preserved at 45 Miocene and Pliocene localities dated between 8 and 1 Ma in northern Chad (16-17 degrees N). Some of these localities yielded cranial remains, lower jaws, and teeth of the hominin species Australopithecus bahrelghazali (similar to 3.6 Ma) and Sahelanthropus tchadensis (similar to 7 Ma). Of the 111 sediment samples analyzed, 41 yielded phytoliths, 20 yielded diatoms, and seven yielded both phytoliths and diatoms. Freshwater planktonic and tychoplanktonic diatom species, indicative of lacustrine conditions, are dominant (>91%) in the samples. The phytolith assemblages indicate an opening of the vegetation and a general trend toward an expansion of grass-dominated environments during the time spanning the two hominin occurrences in Chad. The phytoliths suggest the presence of a mosaic environment, including closed forest patches, palm groves, and mixed/grassland formations, between 7.5 and 7 Ma, the replacement by palm grove-like vegetation at approximately 6.5-5 Ma, and the presence of exclusive grass-dominated formations after 4.5 Ma. The type-locality of S. tchadensis (TM266) was likely similar to modern palm grove formations with an arboreal cover percentage >= 40%. The type locality of A. bahrelghazali (KT12) was a grass-dominated ecosystem (likely savanna) with an unrated percentage of arboreal cover. Furthermore, the grass phytolith data support the existence of a (recurrent) Sahelian-like dry climate in northern Chad since at least 8 Ma. Therefore the local closed vegetation formations in the Djurab region at 7.5-7 Ma were sustained by aquatic systems (such as lakes or related rivers, marshes) rather than by extensive annual precipitation.

This study presents mineralogical and geochemical data from a borehole drilled near the locality of Bol (13 degrees 27'N, 14 degrees 44'E), in the eastern archipelago of the modern Lake Chad (Chad). Samples were taken from a similar to 200 m long core section forming a unique sub-continuous record for Central Africa. Among these samples, 25 are dated between 6.4 and 2.4 Ma. Dominant minerals are clays (66% average) mixed with varying amounts of silt and diatomite. The clay fraction consists of Fe-beidellite (87% average), kaolinite, and traces of illite. Clay minerals originate from the erosion of the vertisols that surrounded the paleolake Chad. Sedimentological data indicate that a permanent lake (or recurrent lakes) existed from 6.7 until 2.4 Ma in the vicinity of Bol. By comparison with modern latitudinal distribution of vertisols in Africa the climate was Sudanian-like. Changes in the sedimentation rate suggest a succession of wetter and dryer periods during at least six million years in the region during the critical time period covering the Miocene-Pliocene transition.

Few proxies exist to identify aridity in the depositional record, although drylands cover ca 30% of the modern continental surface. New exposures in a siliciclastic and carbonate sequence in an arid to hyperarid basin at Olduvai Gorge, Tanzania provide a unique multi‐proxy record of a 1·85 Ma landscape that was exploited by early humans. The 2 m thick sequence of clastics and carbonates that are exposed along a 450 m outcrop records climate change over a single precession (dry‐wet‐dry) cycle. Siliciclastic data (sedimentary structures, grain size, mineralogy) and biological data are combined with data for a 10 to 35 cm thick limestone (stable isotopes, elemental geochemistry, petrography) to generate a depositional facies model for a site DK (Douglass Korongo) on this dry rift basin landscape. This site was situated on a low gradient, distal portion of a volcaniclastic alluvial fan. The clastics are intercalated distal alluvial fan sandy silts and lake clays that accumulated in a low energy environment. Groundwater discharge and the alkaline springs and seeps during wet‐to‐dry change in climate made a freshwater carbonate‐rich environment. Bedded lithofacies (a lime mudstone with fossils) were deposited in shallow standing (spring‐fed) pools, while nodular lithofacies with calcite spherulites indicate permanently saturated ground (seeps). Both environments experienced similar diagenesis, that is, the precipitation of authigenic barite from supersaturated groundwater, desiccation and pedogenesis, and late‐stage calcite precipitation. Compositional and isotopic data suggest that a fresh groundwater‐fed system was available to early humans even during dry intervals of the precession cycle. ISSN:2055-4877

A geomorphological, sedimentological, stratigraphic, and geometric study of 30 trenches excavated around FLK Zinj (Bed I, Olduvai Gorge) has enabled the partial reconstruction of the paleolandscape surrounding this site for a radius of similar to 1000 m. This is the largest sample of geological and archaeological information yet available to reconstruct the topography, ecology, and geomorphology of the Zinj paleosurface and the hominin activities preserved within it. Contrary to previous interpretations, which place FLK Zinj on an isolated and narrow peninsula, it appears that the site was located on the edge of an elevated platform traceable for hundreds of meters. Hominins created FLK Zinj (and other sites, such as the recently discovered PTK and AMK) within the wooded habitats of this platform rather than the more open and grassy environments situated on lower portions of the lacustrine floodplain. Input areas, probably in the form of alluvial fans, existed to the south, following a North-South direction. These input areas are partially responsible for changes in the type sequence. Restricted erosion documented on the wooded platform was mostly caused by runoff processes. An archaeological study of the excavated trenches reveals a sharp contrast in fossil and stone tool density between FLK Zinj and the surrounding landscape, further supporting the contention that the site may have acted as a "central place" where repeated carcass transport, butchery, and consumption took place. Taphonomic studies indicate that at this stage of human evolution, hominins had primary access to carcasses and were not dependent on other carnivores for obtaining meat.

International audience; The multi component FLK North archaeological site was discovered over 50 years ago, and its interpretation has been highly controversial since. Explanations of the dense bone and stone tool accumulation range from a site on a featureless lake margin that is dominantly anthropogenic in origin to a site near a freshwater wetland that is dominated by carnivore activity (e.g. felids and hyenas). FLK North occurs stratigraphically between the Ng'eju Tuff (1.818 AE 0.006 Ma) and Tuff IF (1.803 AE 0.002 Ma), and is composed of 9 distinct levels. Analysis of newly recovered fossil bones and artifacts has shown that the bones of large animals are largely the product of felid hunting and feeding behavior, followed by hyena gnawing and breakage of some bones. The expanded sample of felid prey remains is significant for understanding the contrasts between the mortality profiles of fossil assemblages produced by carnivores and those produced by hominins. Geologic mapping in the environs of the site has revealed rich sedi-mentological and paleoecological records and a thin, but persistent tuff (here named Kidogo Tuff) that is w1.5 m below Tuff IF. Electron microprobe analyses of the tuff mineralogy revealed a unique geochemical fingerprint that permits its use for correlation of widely separated outcrops and facilitates the high resolution reconstruction of the landscape at the time of site formation. The 9 archaeological levels comprise a relatively continuous record through a Milankovitch precession cycle (dry-wet-dry). As the lake receded into the central basin during the dry part of the cycle, surface water supplies dwindled and groundwater-fed springs and wetlands became the dominant freshwater supply. The FLK North archaeological record essentially ended when level 1 was covered with 0.4 m of Tuff IF in a violent volcanic eruption of nearby Mt. Olmoti. However, the overlying Bed II sediments contain scattered archaeological material and a freshwater carbonate deposit that is similar to those found associated with other Bed II archaeological sites, e.g. VEK, HWK and HWKE. The recognition of the ecological association of springs, wetlands and archaeological remains is a powerful predictive tool for locating new archaeological sites in this region that is known for hominin remains.

International audience; Recent excavations in Level 4 at BK (Bed II, Olduvai Gorge, Tanzania) have yielded nine hominin teeth, a distal humerus fragment, a proximal radius with much of its shaft, a femur shaft, and a tibia shaft fragment (cataloged collectively as OH 80). Those elements identified more specifically than to simply Hominidae gen. et sp. indet are attributed to Paranthropus boisei. Before this study, incontrovertible P. boisei partial skeletons, for which postcranial remains occurred in association with taxonomically diagnostic craniodental remains, were unknown. Thus, OH 80 stands as the first unambiguous, dentally associated Paranthropus partial skeleton from East Africa. The morphology and size of its constituent parts suggest that the fossils derived from an extremely robust individual who, at 1.338±0.024 Ma (1 sigma), represents one of the most recent occurrences of Paranthropus before its extinction in East Africa.

International audience; Recent excavations at FLK North (Olduvai Gorge, Tanzania) have produced new information on the orientation of archaeological materials at various levels of the site. This information includes the uniform distribution of material azimuths, which contrasts with previous inferences of highly patterned orientations of materials in the Bed I archaeological sites. Those previous inferences of patterned material orientations are based on Mary Leakey's 50-year-old drawings of artifact and fossil bone distribution, but are not verified by our precise measurements of archaeological objects made in situ. Nor do those previous results agree with the general lack of geological, geomorphological, and/or taphonomic data that would indicate significant post-depositional movement of archaeological materials in the sites. We argue here that Leakey's drawings are incomplete (only portions of each assemblage were drawn) and inaccurate in their representation of the original locations, shapes and orientations of most archaeological specimens. This argument is supported by several important mismatches in object representations between a photograph taken of a small portion of the FLK 22 Zinjanthropus site floor before the removal of the archaeological items, and the sketch of the same area drawn by Leakey. Thus, we conclude that primary orientation data of excavations (i.e., direct measurements taken from items) generated prior to object removal are the only valid indicators of the relative isotropy or anisotropy of these important paleoanthropological assemblages.

FLK North is one of the densest concentrations of fossils found in Olduvai Gorge. A recent taphonomic re-evaluation of the collection excavated by Leakey at the site suggests that it was a palimpsest in which most of the animals were accumulated and modified by carnivores. The lithic tools therefore seem to have an independent depositional history from most of the fauna. The present study, based on new excavations, expands the evidence supporting this interpretation and demonstrates a thicker deposit than was reported by Leakey, including new archaeological levels. It also shows that in the few instances where hominins butchered carcasses, meat, not marrow, was their main target. This argues against passive scavenging hypotheses, which emphasize the dietary role of marrow, and instead underscores the importance of meat in the diet of early Pleistocene hominins.

Renewed excavations at FLK Zinj and its surrounding landscape have yielded valuable information regarding its paleoecological situation and the prehistoric behavioral function of the site. The density of materials at the main cluster of the site excavated by Leakey contrasts with the bone and lithic scatters surrounding the site. The location of FLK Zinj, situated a few hundred meters away from a freshwater spring, would have enabled hominins access to water, plants and game. The appeal of the spot for hominins (also explained by the presence of a wooded habitat) is confirmed by inferences of its redundant use prior and during the formation of the FLK Zinj paleosol, as witnessed by materials accumulated both under and on the waxy clay deposit that constitutes the FLK Zinj stratum. The single-cluster nature of the site indicates central-place behavior and evidence is provided that hominins occupied the site at a time of very low predation hazards in the area.

The records of early hominins are commonly localized both temporally and spatially even in archaeologically rich basins like Olduvai Gorge, Tanzania. The FLK North site was discovered in 1960, but the reason for the exact location of this dense concentration of fossils and stone tools on a lake-margin flat has not been explained. We present new geological and geochemical information from six excavations in upper Bed I, which revealed up to 1.4-m-thick carbonate deposits, attesting to the presence of freshwater springs in the area surrounding FLK North. The δ18O signatures of the tufa are typical for meteoric water that has evolved during evaporation. Tuff IF that caps the sequence was deposited on uneven topography with the highest area a low-relief ridge between two faults at the archaeological site and lowest areas being sites of groundwater discharge. The model proposed here is that rainfall on adjacent highlands was transported to the basin where faults acted as conduits for water. Similar hydrogeological settings at modern lakes Manyara and Eyasi, currently support lush groundwater forest and woodland despite arid climate. FLK North may have been an “oasis” offering freshwater and shelter for consuming meat by both carnivores and hominins.

The 1959 discovery of the hominin fossil Zinjanthropus boisei brought the world's attention to the rich records at Olduvai Gorge, Tanzania. Subsequent excavations of archaeological level 22 (FLK Zinj) Bed I uncovered remains of Homo habilis and a high-density collection of fossils and Oldowan stone tools. The occurrence of this unusual collection of bones and tools at this specific location has been controversial for decades. We present paleoecological data that provide new insights into the origin of FLK Zinj. Our recent excavations 200 m north of the site uncovered a 0.5-m-thick tufa mound draped by Tuff IC, in the same stratigraphic horizon as level 22. Stable isotope analyses indicate that the carbonates were deposited by a freshwater spring. Phytolith analysis of the waxy clay under Tuff IC revealed abundant woody dicotyledon and palm phytoliths, indicating that the site was wooded to densely wooded. The time equivalency and close physical proximity of the two environments indicate the two are related. This study has provided the first documented evidence of springs in Bed I and these data have important implications for the interpretation of hominin behavior in meat acquisition and the ongoing debate on scavenging versus hunting.

Sediments containingArdipithecus ramiduswere deposited 4.4 million years ago on an alluvial floodplain in Ethiopia’s western Afar rift. The Lower Aramis Member hominid-bearing unit, now exposed across a >9-kilometer structural arc, is sandwiched between two volcanic tuffs that have nearly identical40Ar/39Ar ages. Geological data presented here, along with floral, invertebrate, and vertebrate paleontological and taphonomic evidence associated with the hominids, suggest that they occupied a wooded biotope over the western three-fourths of the paleotransect. Phytoliths and oxygen and carbon stable isotopes of pedogenic carbonates provide evidence of humid cool woodlands with a grassy substrate.

International audience; We present a synthesis of modem phytolith studies from Africa, to infer the potential and limitations of phytolith assemblages to reconstruct vegetation and tree cover density. The modem dataset includes 149 phytolith assemblages of surface soil samples from 10 phytogeographical zones and sub-zones from East and West Africa, as well as 500 m-resolution satellite estimates of the percent tree cover at the sampling sites. To test the potential of phytolith assemblages to discriminate vegetation types we used principal component analysis. For each phytogeographical zone and sub-zone, we also provided the mean values, standard effors, and 95% confidence intervals for the means obtained on the modem African dataset for the abundance of the 13 most common phytolith types preserved in soil samples, and for four phytolith indices. Results from the modem African dataset show that 1) the relative abundances of 11 (out of 13) phytolith types allow discrimination of all vegetation zones but the Somalia-Masai steppe region, which at elevation < 600 in asl, exhibits a high proportion of rondel and trapeziform short cell phytoliths, like in the Afromontane region; 2) the co-occurrence of rondels and trapeziform polylobates characterises zones above 1900 m asl and/or current annual temperatures < 19 degrees C; 3) the relative abundance of globular phytoliths (granulate, smooth, and echinate) is better correlated to 500 m-resolution satellite estimates of the tree cover (R-2 = 0.60 for n = 149, and R-2 = 0.57 for n = 85, p < 0.005) than is the abundance of arboreal pollen (R-2 = 0.42, p < 0.005, only for n = 85). The tree cover, however, is largely under-estimated in the Afromontane zone, where globular phytoliths do not trace high-elevation forests. Limitations in our interpretation do exist, but could be overcome in the future through additional studies along an elevation/temperature gradient in the Somalia-Masai region of East Africa, and with more precise identifications of phytolith types and sub-types.

International audience; Phytolith assemblage analysis offers the potential to refine our knowledge of paleoecosystems where grasses and sedges predominate. In this work, Holocene and Pleistocene sediments from an arid tropical region in Ethiopia have been analyzed for their phytolith content, presented as detailed counts and diagrams according to the Twiss classification. The aim is to test the usefulness of phytolith assemblages to indicate paleoenvironments at Middle Awash, where fossil pollen grains are poorly preserved in sediments that yielded abundant archaeological remains. The vegetation in the Middle Awash subdesertic valley is currently a shrub steppe dominated by C4 grasses adapted to arid conditions, with a narrow riparian forest limited to the Awash River. Our results show that modern surface samples, Holocene and Pleistocene sediments contain well-preserved and different phytolith assemblages, and therefore that no translocation processes from modern soil to geological strata seem to occur. Fossil records and modern assemblages are interpreted using phytolith ratios to estimate the density of the tree cover, the aridity and the proportion of C3 versus C4 grasses, as applied to phytolith assemblages from North America and West Africa. The phytolith assemblages from modern soil samples correctly reflect the proportion of trees and shrubs versus grasses, different in the riparian vegetation and the shrub steppe. Modern phytolith assemblages appear to be a mixed signature of local and regional vegetation. Phytolith analysis of the Holocene sample suggests a grassland, where the grass community is constituted by the Chloridoideae subfamily, adapted to warm and dry conditions and where C3-Pooideae cover the highlands. Phytolith analysis of the Pleistocene sample evidences grassland formation with scattered woody elements, where C4-Panicoideae grasses, adapted to warm and humid conditions dominate the grass cover. However, these conclusions need to be confirmed by more complete study on phytolith assemblages from modern vegetation from Ethiopia.