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Clear cut harvest simplifies and eliminates old growth forest structure, negatively impacting biodiversity. Partial cut harvest has been hypothesized (1) to have less impact on biodiversity than clear cut harvest, and (2) to encourage old growth forest structures. Long-term studies are required to test this hypothesis as most studies are conducted soon after harvest. Using epixylic bryophytes as indicators, this study addresses this knowledge gap. Fourteen years after harvest, we examined changes in epixylic bryophyte community composition richness and traits, and their microhabitats (coarse woody debris characteristics and microclimate) along an unharvested, partial cuts and clear cuts harvest treatment in 30 permanent plots established in the boreal black spruce (Picea mariana) forests of northwestern Quebec, Canada. Our results were compared to those of an initial post-harvest study (year 5) and to a chronosequence of old growth forests to examine species changes over time and the similarity of bryophyte communities in partial cut and old growth forests. Coarse woody debris (CWD) volume by decay class varied among harvest treatments with partial cuts and clear cuts recording lower volumes of early decay CWD. The epixylic community was richer in partial cuts than in mature unharvested forests and clear cuts. In addition, species richness and overall abundance doubled in partial and clear cuts between years 5 and 14. Species composition also differed among treatments between years 5 and 14. Furthermore, conditions in partial cut stands supported small, drought sensitive, and old growth confined species that are threatened by conditions in clear cut stands. Lastly, over time, species composition in partial cuts became more similar to old growth forests. Partial cuts reduced harvest impacts by continuing to provide favorable microhabitat conditions that support epixylic bryophytes. Also, partial cut harvest has the potential to encourage old growth species assemblages, which has been a major concern for biodiversity conservation in managed forest landscapes. Our findings support the promotion of partial cut harvest as an effective strategy to achieve species and habitat conservation goals.

Understanding and measuring forest resistance and resilience have emerged as key priorities in ecology and management, particularly to maintain forest functioning. The analysis of the factors involved in a forest’s ability to cope with disturbances is key in identifying forest vulnerability to environmental change. In this study, we apply a procedure based on combining pathway analyses of forest composition and structure with quantitative indices of resistance and resilience to disturbances. We applied our approach to boreal forests affected by a major spruce budworm outbreak in the province of Quebec (Canada). We aimed to identify the main patterns of forest dynamics and the environmental factors affecting these responses. To achieve this goal, we developed quantitative metrics of resistance and resilience. We then compared forests with different pre-disturbance conditions and explored the factors influencing their recovery following disturbance. We found that post-outbreak forest dynamics are determined by distinct resistance and resilience patterns according to dominant species and stand composition and structure. Black spruce forests are highly resistant to spruce budworm outbreaks, but this resistance is conditioned by the length of the defoliation period, with long outbreaks having the potential to lead the system to collapse. In contrast, balsam fir forests easily change to a different composition after outbreaks but are highly resilient when mixed with hardwood species. Overall, the severity of the disturbance and the tree species affected are the main drivers contributing to boreal forest resistance and resilience. Our procedure is valuable to understand post-disturbance dynamics of a broad range of communities and to guide management strategies focused on enhancing the resistance and resilience of the system. This study was funded by MINECO via EST_RES Project (AGL2015-70425-R) and BIOCLIM (CGL2015-6741R) and the EC through the Marie Curie IRSES Project NEWFORESTS (PIRSES-GA- 2013-612645) and the ERA-NET Foresterra Project INFORMED (Grant Number: 29183). MINECO provided MSP with support through a predoctoral contract (BES-2013-063019) and AA through a Juan de la Cierva fellowship (FJCI-2014-20739). We thank Me´lanie Desrochers and Aure´lie Terrier for technical support and three anonymous reviewers for constructive comments on the manuscript.

Aim In managed forest landscapes, the tolerance of species to contemporary alteration of forest cover is often assumed to reflect their resilience to natural disturbances. We tested this central tenet of ecosystem-based management by comparing the structure of forest bird assemblages among four regions with contrasting historical natural disturbance regimes. Location Canada's boreal and northern hardwood forests. Methods Using point count data from four study regions across Canada, we first determined the relative sensitivity of individual bird species to the contemporary reduction of old forest cover at stand and ‘landscape-context’ (1-km radius) scales with log-linear models. The richness of species most sensitive to loss of old forest (hereafter ‘sensitive species’) was then modelled as a function of landscape-scale changes in old forest cover. Differences in the rate of decline in the richness of sensitive species with contemporary cover of old forest were compared among regions using ANCOVA. We then compared broken-stick regression models with linear models to detect thresholds, if present, in this relationship in each region. Results Bird assemblages from regions with relatively infrequent natural disturbances hosted more species sensitive to contemporary reduction in old forest cover. Those species were also more abundant than in regions with frequent natural disturbances, and the rate of decline in their richness with the loss of old forest was steeper in regions with infrequent natural disturbances than in those where they were frequent. However, we did not detect thresholds in this rate of decline in any study region. Main conclusions Our findings are consistent with the contention that historical natural disturbance regimes shape the response of biota to contemporary landscape alterations through evolutionary adaptation. We argue that forest management conducted within the natural range of variability in stand and landscape structure specific to a region is likely to be ecologically sustainable.

Habitat loss and degradation induced by human development are among the major threats to biodiversity worldwide. In this study, we tested our ability to predict the response of bird communities (128 species) to land-use changes in southern Quebec (~483,100 km2 ) over the last 30 yr (between 1984-1989 and 2010-2014) by using species distribution models (299,302 occurrences in 30,408 locations) from a hindcasting perspective. Results were grouped by functional guilds to infer potential impacts on ecosystem services, and to relate model transferability (i.e., ability of our models to be generalized to other times and scales) to specific functional and life-history traits. Overall, our models were able to accurately predict, both in space and time, habitat suitability for 69% of species, especially for granivorous, nonmigrant, tree-nesting species, and species that are tied to agricultural areas under intensive use. These findings indicate that model transferability depends upon specific functional and life-history traits, providing further evidence that species' ecologies affect the ability of models to accurately predict bird distributions. Declining bird species were mostly short-distance migrants that were associated with open habitats (agricultural and nonproductive forest) with aerial insectivorous or granivorous diets, which may be related to agricultural intensification and land abandonment. Land-use changes were positive for some forest bird species that were mainly associated with mixed and deciduous forests, generalist diets and tree-nesting strategies. Yet cavity-nesting birds have suffered substantial reductions in their distributions, suggesting that cumulative effects of intensive logging and wildfires on mature forests pose a threat for forest-specialist species. Habitat suitability changes predicted by our coarse-scale species distribution models partially agreed with the long-term trends reported by the North American Breeding Bird Survey. Our findings confirm land-use change as a key driving force for shaping bird communities in southern Quebec, together with the need to explicitly incorporate it into global change scenarios that better inform decision-makers on conservation and management.

Natural disturbances, such as fire and insect outbreaks, play important roles in natural forest dynamics, which are characterized over long time scales by changes in stand composition and structure. Individual-based forest simulators could help explain and predict the response of forest ecosystems to different disturbances, silvicultural treatments, or environmental stressors. This study evaluated the ability of the SORTIE-ND simulator to reproduce post-disturbance dynamics of the boreal mixedwoods of eastern Canada. In 1991 and 2009, we sampled all trees (including seedlings and saplings) in 431 (256 m2) plots located in the Lake Duparquet Research and Teaching Forest (western Quebec). These plots were distributed in stands originating from seven wildfires that occurred between 1760 and 1944, and which represented a chronosequence of post-disturbance stand development. We used the 1991 inventory data to parameterize the model, and simulated short- to long-term natural dynamics of post-fire stands in both the absence and presence of a spruce budworm outbreak. We compared short-term simulated stand composition and structure with those observed in 2009 using a chronosequence approach. The model successfully generated the composition and structure of empirical observations. In long-term simulations, species dominance of old-growth forests was not accurately estimated, due to possible differences in stand compositions following wildfires and to differences in stand disturbance histories. Mid- to long-term simulations showed that the secondary disturbance incurred by spruce budworm did not cause substantial changes in early successional stages while setting back the successional dynamics of middle-aged stands and accelerating the dominance of white cedar in late-successional post-fire stands. We conclude that constructing a model with appropriate information regarding stand composition and disturbance history considerably increases the strength and accuracy of the model to reproduce the natural dynamics of post-disturbance boreal mixedwoods.

Over the last 25 years, greater understanding of natural dynamics in the boreal forest has led to the integration of forest ecosystem management principles into forest policy of several Canadian provinces and, in turn, to greater interest in developing silvicultural treatments that are grounded in natural stand-level dynamics – often referred to as natural disturbance-based silviculture. As a result, alternative silvicultural practices including variants of partial cutting are increasingly being applied in the boreal forest as an approach to balancing economic and ecological management objectives. While the numerous benefits of partial cutting reported in the literature are acknowledged, the objective of this paper is to provide an overview of factors or constraints that potentially limit the application of these practices in boreal Canada in the context of forest ecosystem management and natural disturbance-based silviculture. Among constraining factors, numerous studies have reported elevated mortality rates of residual stems following partial cutting, initial growth stagnation of residual trees, problems related to recruitment of desirable species and, on certain flat or lowland sites, risks of long-term decline in site and stand productivity. A number of operational challenges to partial cutting in the boreal forest are also presented and several avenues of research are proposed.

In the eastern Canadian boreal forest, the term hardwood expansion or encroachment (enfeuillement in French) generally refers to the phenomenon of increasing cover of intolerant hardwoods, notably aspen (Populus tremuloides), following clearcutting. Such expansion is perceived as a local (stand-level) process that is reflected at a larger, regional scale. The objective of this study was to characterize the increase in aspen cover following clearcutting in the clay belt of Abitibi, in northwestern Quebec. Regional data from 3 province-wide forest inventories (1970, 1980, and 1990) conducted by the Quebec Ministry of Natural Resources and Wildlife were analyzed for 2 ecological regions, the Abitibi Plain (western balsam fir-white birch bioclimatic subdomain) and the Matagami Lake Plain (western black spruce-feathermoss bioclimatic subdomain). An assessment was done of forest cover transformations following clearcutting over 3 decades based on time since harvesting, surface deposit/drainage, and ecological region. Results show that mesic clay sites are particularly susceptible to aspen encroachment and that the overall rate of encroachment from the 1970s to the 1990s in the black spruce and balsam fir regions has been about 30%. Moreover, despite the greater abundance of poorly drained and organic sites, less favourable to hardwood establishment, in the more northerly black spruce region, the 2 regions appear to present the same sensitivity (or propensity) to intolerant hardwood expansion. On sensitive sites, management of black spruce forests should include adaptive practices to limit increases of the hardwood component.

Aim Climate is often regarded as the primary control determining the location of an ecotone between two vegetation zones. However, other ecological factors may also be important, especially when the northern limit of the dominant species of a vegetation zone extends further than the limit of the zone itself. This study aimed to identify the ecological variables explaining the transition between two zones within the boreal biome in Quebec (eastern Canada): the southern mixedwood forests dominated by balsam fir (Abies balsamea) and white birch (Betula papyrifera), and the northern coniferous forests dominated by black spruce (Picea mariana). Location Quebec (eastern Canada). Methods Data from 5023 sampling plots from the ecological inventory of the Quebec Ministry of Natural Resources distributed throughout the two bioclimatic zones were used in logistic regressions to determine the relationships between the presence or absence of balsam fir stands and different abiotic and biotic variables, at both stand and landscape scales. Results The presence of balsam fir stands was negatively related to the thick organic horizons, coarse xeric deposits and low positions on the slope, whereas stands were favoured by high elevations, steep slopes and moderate drainage. These results defined the suitable conditions for the development of balsam fir stands. In the coniferous zone these suitable conditions were less abundant. Furthermore, the saturation level of suitable sites was lower, as well as the incidence of balsam fir stands in unsuitable sites (overflow). Balsam fir stands were mostly located near lakes and rivers. All significant variables at both the stand and landscape scales explained between 34 and 42% of the location of the potential northern distribution limit of the mixedwood zone. Main conclusions Our results suggest the important role of historical factors related to post-glacial vegetation and past disturbances in determining the relative abundance of balsam fir in both zones of the boreal biome.

Single-tree selection cutting is sometimes believed to be similar to the natural gap disturbance regime of hardwood forests, but few studies have specifically compared the compositional and structural characteristics of old-growth hardwood stands, undergoing natural gap dynamics and hardwood stands previously subjected to partial cuts. This study characterized and compared the composition (saplings and trees) and structure (gaps, foliage distribution, tree diameter and density, snags and coarse woody debris) of old-growth stands (OG), 12-year-old selection cuts (SC), and 28-33-year-old diameter-limit cuts (DLC) in sugar maple (Acer saccharum)-dominated northern hardwood stands. Results showed marked structural differences between OG and harvested stands, with stronger differences between DLC and OG than between SC and OG. The synchronized formation of numerous canopy openings in harvested stands induced a massive post-harvest recruitment of advance regeneration in both SC and DLC that created a dense foliage layer in the understory. Large living trees (dbh > 39.1 cm) and defective trees were less numerous in SC than OG, which can have a detrimental impact on species dependent on these structural elements, and on the future availability and characteristics of coarse woody debris. Relatively few compositional differences were noticed among stand types, although a greater proportion of mid-tolerant species was found in the post-harvest recruitment cohorts of harvested stands compared to OG, and a lower proportion of beech (Fagus grandifolia Ehrh.) saplings was observed in DLC compared to OG and SC. We argue that even if selection cutting is closer to the natural disturbance regime of hardwood forests than diameter-limit cutting, and therefore representing progress toward the development and implementation of a natural-disturbance-based management, a recurring application of selection cutting might lead to a homogenization of forest structure and composition, a reduction of key structural features and a reduction in biological diversity at both the stand and landscape scales. Some management recommendations are proposed.

We studied community development and mortality among tree species at multiple spatial scales in a 236- year mixedwood chronosequence at Lake Duparquet, Quebec. Spatial relationships, species associations, and understory-overstory replacement patterns were studied using spatial statistics, patch indices, G tests, and transition matrices. Results of these analyses showed that shade-intolerant and shade-tolerant tree communities formed increas- ingly fine-grained patches in midsuccession. These fine-grained associations appear to be partly due to suppressed stems being released into a slowly thinning canopy and partly due to new recruitment of shade-tolerant conifers. In 1- ha plots, tree species richness and evenness peaked in 25-, 100-, and 400-m 2 subplots during midsuccession. Forest cover type diversity also peaked in midsuccessional landscapes assessed using aerial photographs. The oldest 1-ha plot was dominated by monospecific patches of eastern white cedar (Thuja occidentalis L.), which was the most likely re- placement for live and dead trees and the most abundant species in the numerous subplots that were occupied by trees smaller than 8 cm DBH. In spite of this dominance, long-lived paper birch (Betula papyrifera Marsh.) and residual fir (Abies balsamea (L.) Mill.) seedlings may maintain the mixedwood character of this stand into the future. The implica- tions of our findings for stand management and conservation are discussed. Resume : Les auteurs ont etudie le developpement et la mortalite de communautes d'arbres en peuplements mixtes a plusieurs echelles spatiales dans une chronosequence de 236 ans au lac Duparquet, Quebec. Les relations spatiales, les associations d'especes et les patrons de remplacement entre les especes du couvert dominant et celles du sous-bois ont ete etudies en utilisant des statistiques spatiales, des indices d'agglomeration, des tests de G et des matrices de transi- tion. Les resultats de ces analyses montrent que les communautes d'arbres intolerants et tolerants a l'ombre forment un nombre croissant d'agglomerations de petites tailles a l'etape du milieu de la succession. La formation de ces associa- tions de petites tailles semble partiellement attribuable, d'une part, aux tiges opprimees qui sont liberees a l'interieur d'un couvert qui s'eclaircit lentement et, d'autre part, au recrutement de coniferes tolerants a l'ombre. Dans des parcel- les de 1 ha, la richesse en especes et l'equitabilite ont culmine dans des sous-parcelles de 25, 100 et 400 m 2 de peu- plements parvenus a l'etape du milieu de la succession. La diversite du type de couvert forestier estimee a partir de photographies aeriennes a aussi culmine sur des territoires parvenus a l'etape de milieu de succession. La plus vieille placette de 1 ha etait dominee par des agglomerations monospecifiques de thuya occidental (Thuja occidentalis L.), l'espece la plus susceptible de remplacer les arbres morts et vivants et l'espece la plus abondante dans les nombreuses sous-parcelles occupees par des arbres dont le DHP est inferieu ra8c m. Malgre cette dominance, les especes a forte longevite que sont le bouleau a papier (Betula papyrifera Marsh.) et les semis residuels de sapin baumier (Abies balsa- mea (L.) Mill.) peuvent maintenir la mixite d'especes de ce type de peuplement dans le futur. Les implications de nos resultats sont discutees en regard de l'amenagement et de la conservation des forets.

The forest fire regime was characterized for the boreal forest of western Quebec using the provincial government’s digital databases (1945-1998). Lightning- and human-caused fires account for 71% a...

In Canada, there are still extensive tracts of boreal forest consisting of stands that have resulted from natural disturbances. The country's forests are a mosaic made up to a large extent of old-growth forest that is beyond commercial harvesting age, especially in the boreal forest of eastern Canada. As areas of boreal forest under management steadily expand, as demand for forest products continues to grow and as rotation periods become shorter in response to silvicultural practices, the forest cover will inevitably become younger, causing changes to the structure and composition of the mosaic of forest stands that will affect the aspect of entire landscapes. These changes may have an adverse impact on biological diversity. Forest birds are one group of living organisms that may respond quickly to the advent of younger forest landscapes, thereby acting as a biological indicator. In this paper, we discuss some of the problems that birds face as a result of the truncation of the age-class distribution of managed forest landscapes in eastern Canada's coniferous boreal forest, using data obtained from our research in the Clay Belt region of Quebec and Ontario. More specifically, we look at how birds respond to changes in forest structure and composition in terms of time since natural disturbances, and to variation in dead trees availability. We then consider the impact of the prospective rejuvenation of the forest cover in managed forest landscapes, and possible solutions aimed at mitigating that impact through new management strategies based on the maintenance of forest ecosystem diversity. The ability of these new management strategies to maintain the ecological integrity of bird communities provides an indication of their potential as tools for contributing to the maintenance of biological diversity in a broader sense. Key words: bird communities, old black spruce forests, natural landscape age structure, stand structure, dead wood, multicohort management

Forest ecosystem management, based partly on a greater understanding of natural disturbance regimes, has many variations but is generally considered the most promising approach to accommodating biodiversity concerns in managed forested regions. Using the Lake Duparquet Forest in the southeastern Canadian boreal forest as an example, we demonstrate an approach that attempts to integrate forest and stand-level scales in biodiversity maintenance. The concept of cohorts is used to integrate stand age, composition and structure into broad successional or stand development phases. Mean forest age (MFA), because it partly incorporates historic variability of the regional fire cycle, is used as a target fire cycle. At the landscape level, forest composition and cohort objectives are derived from regional natural disturbance history, ecosystem classification, stand dynamics and a negative exponential age distribution based on a 140 year fire cycle. The resulting multi-cohort structure provides a framework for maintaining the landscape in a semi-natural age structure and composition. At the stand level, the approach relies on diversifying interventions, using both even-aged and uneven-aged silviculture to reflect natural stand dynamics, control the passage (‘‘fluxes’’) between forest types of different cohorts and maintain forest-level objectives. Partial and selective harvesting is intended to create the structural and compositional characteristics of mid- to late-successional forest types and, as such, offers an alternative to increasing rotation lengths to maintain ecosystem diversity associated with overmature and old-growth forests. The approach does not however supplant the necessity for complementary strategies for maintaining biodiversity such as the creation of reserves to protect rare, old or simply natural ecosystems. The emphasis on maintaining the cohort structure and forest type diversity contrasts significantly with current even-aged management in the Canadian boreal forest and has implications for stand-level interventions, notably in necessitating a greater diversification of silvicultural practices including more uneven-aged harvesting regimes. The approach also presents a number of operational challenges and potentially higher risks associated with multiply stand entries, partial cutting and longer intervals between final harvests. There is a need for translating the conceptual model into a more quantitative silvicultural framework. Silvicultural Forest Ecology and Management 155 (2002) 369‐385

Although the concept of forest ecosystem management based on natural disturbance has generated a great deal of interest, few concrete examples exist of FEM principles being put into application. Silvicultural practices that emulate natural disturbances are proposed with examples from the principal vegetation zones of Quebec. With the exception of the large-scale use of careful logging to protect advanced regeneration in ecosystems generally controlled by fire, stand-level silvicultural practices currently used are reasonably similar to natural disturbances, although important differences exist. In contrast, at the forest-level, even-aged management, as is currently practised, rarely permits adequate reproduction of the variety of age classes, stand types, and structural components normally found in the boreal forest. A model that allows an even-aged management approach inspired by natural dynamics is proposed.

Balsam fir (Abiesbalsamea (L.) Mill.) mortality caused by the last spruce budworm (Choristoneurafumiferana (Clem.)) outbreak (1970–1987) was studied in 624 sites belonging to a complex natural forest mosaic originating from different fires in northwestern Quebec. Multiple regression analyses were used to assess the respective effects of stand structure, species composition, site characteristics, and the forest composition surrounding the stand on observed stand mortality. Mortality was observed to increase in relation to diameter of the trees, basal area of balsam fir, and the number of stands dominated by conifers in the forest mosaic. All of these factors showed significant independent effects, but 60% of the variance remained unexplained. Site characteristics, however, did not show a significant relationship to stand mortality. The results suggest that forest composition at both the stand and the forest mosaic levels may be responsible for differing degrees of defoliation that result in differences in stand mortality. Forest management strategies that favor the presence of mixed compositions both at the stand level and at the mosaic level may contribute to decreased stand vulnerability.

A study of forest succession was conducted in an 11 ha old-growth beech-maple stand located at the northern limit of the temperate deciduous forest of North America. Eventual changes in dominance were simulated from three transition matrix models based on understory composition or observed replacement in forest gaps. All the models suggested that the forest composition is not currently at equilibrium. If the transition tendency persists, American beech (Fagus grandifolia Ehrh.) will sharply increase in abundance at the expense of sugar maple (Acer saccharum Marsh.). Possible mechanisms responsible for this replacement are discussed.

The aim of this study was to evaluate if and how the previous land use (here: agriculture versus forest) has influenced the floristic composition now observed in the understory of jack pine plantations of Abitibi-Temiscamingue (North-Eastern Canada). Floristic inventories were conducted in jack pine plantations which had been established on forest sites after logging and on sites previously utilized for agriculture. These were compared to one another and to inventories conducted in natural forests and on agricultural sites which had not (not yet) been planted with trees (old fields). The vegetation community composition of the sites is analysed using several multivariate techniques. Results show that the flora of plantations on old agricultural fields is markedly different than that of plantations established on sites which were previously forested and that the two types of plantation are unlikely to become floristically the same before the anticipated harvest time. Comparing old and young plantations and old fields in light of natural forests sites, we show that the previous land-use is a key determinant of future compositional differences in plantations. This must be considered when we evaluate the impact of establishing plantations on sites with different land-use histories.

In natural boreal forests, disturbances such as fire and variation in surficial deposits create a mosaic of forest stands with different species composition and age. At the landscape level, this variety of stands can be considered as the natural mosaic diversity. In this paper, we describe a model that can be used to estimate the natural diversity level of landscapes. We sampled 624 stands for tree species composition and surficial deposits in eight stand-age classes corresponding to eight fire episodes in the region of Lake Duparquet, Abitibi, Quebec at the southern fringe of the Boreal Forest. For six surficial deposit types, stand composition data were used to define equations for vegetation changes with time for a chronosequence of 230 years for four forest types. Using Van Wagner’s (1978) model of age class distribution of stands, the proportion of each forest type for several lengths of fire cycle were defined. Finally, for real landscapes (ecological districts) of the ecological region of the “Basses-Terres d’Amos”, the proportion of forest types were weighted by the proportion of each surficial deposit type using ecological map information. Examples of the possible uses of the model for management purposes, such as biodiversity conservation and comparisons of different landscapes in terms of diversity and sensitivity to fire regime changes, are discussed.

In this study, we characterised the composition and configuration of post-fire residual habitats belonging to two physiographic zones of the black spruce–moss domain in western Quebec. Thirty-three large fires (2000–52 000 ha) were selected and extracted on classified Landsat satellite imagery. The results show that a minimum of 2% and a maximum of 22% of burned areas escaped fire, with an overall average of 10.4%. The many forest patches that partially or entirely escaped fire formed residual habitats (RHs). It was found that although the area of RHs follows a linear relationship with fire size, their proportion appears relatively constant. Spatial analyses showed that the fires could be separated into two groups depending on the physiographic zones (East-Canadian Shield v. West-Clay Belt Lowlands). Fires in the west zone generate less RHs and appear to be associated with more extreme weather conditions. In most cases there was no association with water or wetlands; in some fires the presence of RHs is associated with the proximity of water bodies. The failure to find an association between RHs and wetlands suggests that this type of environment is part of the fuel. Coniferous woodland with moss appears particularly overrepresented within RHs. Our results suggest that the local and regional physiographic conditions strongly influence the creation of RHs; therefore, it is important to consider those differences when applying ecosystem-based management.

Newfoundland’s west coast has long been a center of attention for taxonomists and botanists interested in floristic and phytogeographical studies because of its peculiar flora. The importance of this area was first recognized by Fernald (1911, 1926, 1933) in the early part of this century. His pioneering explorations led to many important discoveries and resulted in two landmark papers concerning the phytogeography of the region of the Gulf of St Lawrence (Fernald 1924, 1925).

Bird community response to both landscape-scale and local (forest types) changes in forest cover was studied in three boreal mixed-wood forest landscapes modified by different types of disturbances: (1) a pre-industrial landscape where human settlement, agriculture, and logging activities date back to the early 1930s, (2) an industrial timber managed forest, and (3) a forest dominated by natural disturbances. Birds were sampled at 459 sampling stations distributed among the three landscapes. Local habitat and landscape characteristics of the context surrounding each sampling station (500-m and 1-km radius) were also computed. Bird communities were influenced by landscape-scale changes in forest cover. The higher proportion of early-successional habitats in both human-disturbed landscapes resulted in significantly higher abundance of early-successional bird species and generalists. The mean number of mature forest bird species was significantly lower in the industrial and pre-industrial landscapes than in the natural landscape. Landscape-scale conversion of mature forests from mixed-wood to deciduous cover in human-disturbed landscapes was the main cause of changes in mature forest bird communities. In these landscapes, the abundance of species associated with mixed and coniferous forest cover was lower, whereas species that preferred a deciduous cover were more abundant. Variation in bird community composition determined by the landscape context was as important as local habitat conditions, suggesting that predictions on the regional impact of forest management on songbirds with models solely based on local scale factors could be misleading. Patterns of bird species composition were related to several landscape composition variables (proportions of forest types), but not to configuration variables (e.g., interior habitat, amount of edge). Overall, our results indicated that the large-scale conversion of the southern portion of the boreal forest from a mixed to a deciduous cover may be one of the most important threats to the integrity of bird communities in these forest mosaics. Negative effects of changes in bird communities could be attenuated if current forestry practices are modified toward maintaining forest types (deciduous, mixed-wood, and coniferous) at levels similar to those observed under natural disturbances.

Over the past decades, there has been an increasing interest in the development of forest management approaches that are based on an understanding of historical natural disturbance dynamics. The rationale for such an approach is that management to favor landscape compositions and stand structures similar to those of natural ecosystems should also maintain biological diversity and essential ecological functions. In fire-dominated landscapes, this approach is possible only if current and future fire frequencies are sufficiently low, comparing to pre-industrial fire frequency, that we can substitute fire by forest management. We address this question by comparing current and future fire frequency to historical reconstruction of fire frequency from studies realized in the Canadian boreal forest. Current and simulated future fire frequencies using 2 and 3 x CO2 scenarios are lower than the historical fire frequency for many sites, suggesting that forest management could potentially be used to recreate the forest age structure of fire-controlled pre-industrial landscapes. There are however, important limitations to the current even-age management.

The distribution of phytogeographical groups and life-form categories of the total vascular flora (711 taxa) is analysed for thirty-five habitats, ranging from seashore tidal flats to tundra barrens. Classification and ordination techniques (canonical correlation analyses) are used to study the interrelationships. Most of the phytogeographical groups and the life-forms have high correlations with the distribution of the vegetation types of Gros Morne National Park. CCA provides not only a synthetic account of the rich flora of the study area, but it helps to establish interesting relationships between plant distributions, life-forms and vegetation types, even at a regional level.