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Study of all flies (Diptera) collected for one year from a four-hectare (150 x 266 meter) patch of cloud forest at 1,600 meters above sea level at Zurquí de Moravia, San José Province, Costa Rica (hereafter referred to as Zurquí), revealed an astounding 4,332 species. This amounts to more than half the number of named species of flies for all of Central America. Specimens were collected with two Malaise traps running continuously and with a wide array of supplementary collecting methods for three days of each month. All morphospecies from all 73 families recorded were fully curated by technicians before submission to an international team of 59 taxonomic experts for identification. Overall, a Malaise trap on the forest edge captured 1,988 species or 51% of all collected dipteran taxa (other than of Phoridae, subsampled only from this and one other Malaise trap). A Malaise trap in the forest sampled 906 species. Of other sampling methods, the combination of four other Malaise traps and an intercept trap, aerial/hand collecting, 10 emergence traps, and four CDC light traps added the greatest number of species to our inventory. This complement of sampling methods was an effective combination for retrieving substantial numbers of species of Diptera. Comparison of select sampling methods (considering 3,487 species of non-phorid Diptera) provided further details regarding how many species were sampled by various methods. Comparison of species numbers from each of two permanent Malaise traps from Zurquí with those of single Malaise traps at each of Tapantí and Las Alturas, 40 and 180 km distant from Zurquí respectively, suggested significant species turnover. Comparison of the greater number of species collected in all traps from Zurquí did not markedly change the degree of similarity between the three sites, although the actual number of species shared did increase. Comparisons of the total number of named and unnamed species of Diptera from four hectares at Zurquí is equivalent to 51% of all flies named from Central America, greater than all the named fly fauna of Colombia, equivalent to 14% of named Neotropical species and equal to about 2.7% of all named Diptera worldwide. Clearly the number of species of Diptera in tropical regions has been severely underestimated and the actual number may surpass the number of species of Coleoptera. Various published extrapolations from limited data to estimate total numbers of species of larger taxonomic categories (e.g., Hexapoda, Arthropoda, Eukaryota, etc.) are highly questionable, and certainly will remain uncertain until we have more exhaustive surveys of all and diverse taxa (like Diptera) from multiple tropical sites. Morphological characterization of species in inventories provides identifications placed in the context of taxonomy, phylogeny, form, and ecology. DNA barcoding species is a valuable tool to estimate species numbers but used alone fails to provide a broader context for the species identified.

On the basis of re-evaluation of morphological characters of the Lycoriella group of genera and subgenera, generic rank is given to the two species groups belonging to Lycoriella (Hemineurina) Frey, 1942 and to Lycoriella (Coelostylina) Tuomikoski, 1960. The Lycoriella (Hemineurina) inflata group, including the type species of the subgenus, Sciara conspicua Winnertz, 1867, is treated as the genus Hemineurina stat. n. and the Lycoriella (Hemineurina) vitticollis group as the genus Trichocoelina gen. n. (type species Sciara vitticollis Holmgren, 1883). Coelostylina Tuomikoski, 1960 (type species Lycoriella (Coelostylina) freyi Tuomikoski, 1960) is a junior homonym of Coelostylina Kittl, 1894, and is renamed Stenacanthella nom. et stat. n. The genera are diagnosed and their phylogeny is discussed. Eight species are excluded from the Lycoriella group. They are transferred to the genera Bradysiopsis Tuomikoski, 1960, Camptochaeta Hippa & Vilkamaa, 1994, Merizomma Sasakawa, 2003 stat. n. and Scatopsciara Edwards, 1927 (five species) or are for the time being regarded as incertae sedis (two species) and as nomen nudum (one name). Numerous nomenclatural corrections are made also in the genera Hemineurina Frey, Stenacanthella Vilkamaa & Menzel and Trichocoelina Vilkamaa & Menzel. Altogether 42 new combinations, three changes in status and one new synonym are presented. A lectotype is designated for Hemineurina algida (Frey, 1948) and two Hemineurina species names are removed from synonymy and given full species status. The following species of Trichocoelina are newly described: Trichocoelina absidata sp. n. (Russia: Krasnodarsk region), T. aemula sp. n. (Finland, Russia: Krasnodarsk region), T. biplex sp. n. (Canada: Newfoundland and Labrador, Yukon), T. dicksoni sp. n. (Russia: Arkhangelsk oblast, Kemerovsk oblast, Krasnodarsk region), T. dispansa sp. n. (Russia: Krasnodarsk region), T. dividua sp. n. (Canada: Northwest Territories), T. hians sp. n. (Canada: Yukon), T. imitator sp. n. (Canada: Yukon), T. incrassata sp. n. (USA: Alaska), T. ithyspina sp. n. (Norway), T. jukkai sp. n. (Finland), T. magnifica sp. n. (Canada: Yukon), T. nefrens sp. n. (Russia: Krasnodarsk region), T. obesula sp. n. (Norway), T. oricillifera sp. n. (Finland, Norway), T. planilobata sp. n. (Finland), T. quintula sp. n. (Finland), T. semisphaera sp. n. (Finland, Norway), T. semusta sp. n. (Italy, USA: Alaska), and T. tecta sp. n. (Canada: Nunavut, Yukon, Russia: Krasnodarsk region, Yamalo-Nenets Autonomous Okrug, USA: Alaska). The Trichocoelina species are keyed, the 20 new species are described and illustrated, and the 9 previously known ones, transferred to the new genus, are briefly diagnosed and the taxonomically relevant literature regarding them is listed. Trichocoelina janetscheki (Lengersdorf, 1953) comb. n. and Trichocoelina brevicubitalis (Lengersdorf, 1926) comb. n. are redescribed. The genus Trichocoelina currently includes 29 species: 17 in the Palaearctic, 6 in the Nearctic and 6 in the Holarctic. All known species are northern or montane.

The genus Pseudoaerumnosa Rudzinski, 2006 is redefined. The genus includes the following species which are described, illustrated and keyed: Pseudoaerumnosa acinacea sp. n., P. ampliata sp. n., P. annae sp. n., P. awanensis sp. n., P. banari sp. n., P. clavidactyla sp. n., P. clivicola sp. n., P. collicola sp. n., P. consuota sp. n., P. cryptoloba sp. n., P. curvifalx sp. n., P. eminula sp. n., P. exacuta sp. n., P. filispicata sp. n., P. formosa sp. n., P. fragilis sp. n., P. impensa sp. n., P. inviolata Rudzinski, 2006, P. junciseta sp. n., P. obovata sp. n., P. pilicaudata sp. n., P. quadriquetra sp. n., P. saginata sp. n., P. tenuidens sp. n. and P. tkoci sp. n. Morphological characters and the phylogenetic position of Pseudoaerumnosa are discussed. For some species, the barcode (COI) sequence data were obtained. The genus is currently known from the Oriental (22 species), Malagasy (3 species) and Australasian regions (1 species).

Four Oriental species of Aerumnosa Mohrig, 1999 (Diptera: Sciaridae), a genus previously known only from Papua New Guinea, are newly described and illustrated: Aerumnosa bituberculata sp. n. (India), A. gemmifera sp. n. (Malaysia: Sabah), A. horrifica sp. n. (Brunei, Thailand) and A. impar sp. n. (Malaysia: Sabah). On the basis of the new material, the genus is redefined. A key to the known species of Aerumnosa is presented, including four new species. An updated molecular phylogenetic analysis based on four gene markers (18S, 28S, 16S and COI) shows Aerumnosa to be a member of the subfamily Cratyninae. The monophyly of Cratyninae is well supported, which clade also includes the genera Hyperlasion Schmitz, 1919, Pnyxiopalpus Vilkamaa & Hippa, 1999 and Pseudoaerumnosa Rudzinski, 2006. According to the present phylogenetic hypothesis, the monophyly of Cratyna Winnertz, 1967 s. l. needs to be revisited. The clade including Cratyna (s. str.) ambigua (Lengersdorf, 1934) appears as the sister group of Aerumnosa.

Six species of the genus Camptochaeta Hippa & Vilkamaa, 1994 and four of the genus Corynoptera Winnertz, 1867 from the Holarctic region are newly described and illustrated: Camptochaeta chilkatensis sp. n. (USA: Alaska), C. curtipilosa sp. n. (USA: Colorado), C. exquisita sp. n. (USA: Oregon), C. fitzgeraldi sp. n. (USA: Oregon), C. fulvistylata sp. n. (Canada: British Columbia), C. pertenuis sp. n. (Russia: Krasnodarsk region), Corynoptera bocheri sp. n. (Greenland; Canada: Quebec), C. hyperborea sp. n. (Russia: Krasnodarsk region), C. hystricina sp. n. (RUSSIA: Krasnodarsk region) and C. tenuichaeta sp. n. (Russia: Krasnodarsk region).

A checklist of the family Sciaridae (Diptera) recorded from Finland is provided. The genus Sciarosoma Chandler with a disputed family placement is also included in the list.

The phylogeny of the main groups of the Sciaroidea, including the fossil Antefungivoridae, Archizelmiridae, Mesosciophilidae, Pleciofungivoridae, Pleciomimidae, Protopleciidae and Bolitophilidae: Mangasinae, and an extant new taxon, was studied by parsimony analysis. Two cladistic analyses of seventy-eight morphological characters from adults were made. One analysis, with forty-one extant taxa in the ingroup and the other, with the addition of twelve fossil taxa, both produced two most parsimonious cladograms. The phylogenetic hypotheses obtained differed from each other, and in part also to a great extent from previous ones although most of the traditionally recognized groups appeared monophyletic, including the speciose Cecidomyiidae and Sciaridae. The Cecidomyiidae (fossil analysis) or the Keroplatidae-Ditomyiidae (extant analysis) appeared as the sister-group of the rest of the Sciaroidea. Following on from these analyses, we propose emending the current Sciaridae to include the following subfamilies: Archizelmirinae stat. n., Rangomaraminae stat. n., Sciarinae, Sciarosominae subfam. n. and Sciarotrichinae subfam. n. A new taxon from Namibia, Sciarotricha biloba gen. n., sp. n. is described, and, according to the phylogenetic analysis, is placed in the Sciaridae (Sciarotrichinae). The sister-group of the Sciaridae as newly defined is the Mycetophilidae group, in the extant analysis including the Mycetophilidae, Manotidae, Lygistorrhinidae, Pterogymnus and Sciaropota, and in the fossil analysis even including the Mesosciophilidae and the Ohakunea group (Ohakunea + Colonomyia).

The genus Peyerimhoffia Kieffer (type-species Sciara vagabunda Winnertz), currently classified as a subgenus of Cratyna Winnertz (type-species Cratyna atra Winnertz) is redefined and its phylogeny studied by parsimony analysis. Cladistic analysis using sixty-four morphological characters from adult males of sixty-six ingroup and one outgroup species produced thirty most parsimonious cladograms. The strict consensus cladogram of these showed unambiguously that the type-species of Peyerimhoffia and the species currently referred to the Corynoptera crassistylata group form a monophyletic unit. The phylogenetic hypothesis obtained also suggested that Cratyna s.l. in its current concept is non-monophyletic and that Peyerimhoffia deserves generic status. The genus is Holarctic-Oriental in distribution and includes the following species, which are keyed and described or diagnosed: Peyerimhoffia alpina (Mohrig) (Austrian Alps, Italy, Spain, Canada: Quebec), P. calva sp.n. (Nepal), P. collina sp.n. (Russia: Altay region), P. crassistylata (Frey) (central and northern Europe, Altay region, Kurile Islands), P. curvata (Mohrig & Mamaev) (Russia: Tuva region), P. infera sp.n. (northern Europe), P. ioculatoria (Mohrig) (Nepal), P. menzeli sp.n. (northern Europe), P. obtusicauda (Strobl) (southern Europe), P. semicurvata (Mohrig) (Nepal, Burma), P. subcurvata (Mohrig) (Nepal), P. thula sp.n. (northern Europe), P. ultima sp.n. (Nepal), and P. vagabunda (Winnertz) (Europe, eastern Palaearctic). Plastosciara perniciosa Edwards, Hyperlasion curtipennis Edwards, and Plastosciara hybrida Mohrig & Mamaev are excluded, the former appearing more closely related to Epidapus Haliday, the latter two to Spathobdella Frey.

The Northern Hemispheric sciarid species placed in the nigrohalteralis group of Corynoptera Winnertz are a monophyletic unit, which is described as a new genus Dichopygina gen. n. (type-species Bradysia (Chaetosciara) triseriata var nigrohalteralis Frey). The genus is Holarctic in distribution and includes the following species which are keyed and described: Dichopygina aculeata sp. n. (Holarctic), D. bernhardi sp. n. (Palaearctic), D. duplicis sp. n. (Nearctic), D. intermedia (Mohrig & Krivosheina) (Palaearctic), D. nigrohalteralis (Frey) (Holarctic), D. ramosa sp. n. (Palaearctic) and D. stricta sp. n. (Nearctic). Our previously published cladistic analysis suggests that the species now included in Dichopygina gen. n. are sister taxa to a large clade including the current Lycoriella Frey and elements of Corynoptera and Bradysia Winnertz. Corynoptera acanthostyla Tuomikoski and the New Zealand species referred to the Corynoptera nigrohalteralis group are excluded.

Summary Pnyxiopalpus gen.n. (type-species P. raptor sp.n.), Oriental in distribution, includes the following species: P. acanthipes sp.n. (Malay Peninsula), P. aculeatus sp.n. (Borneo), P. adebratti sp.n. (Borneo), P. aphrodite sp.n. (Sumatra), P. dentaneus sp.n. (Sulawesi), P. fossor sp.n. (Borneo), P. fuscinellus sp.n. (Borneo), P. hamatus sp.n. (Borneo), P. latifalx sp.n. (Borneo), P. macrocellus sp.n. (Malay Peninsula), P. microdon sp.n. (Sumatra), P. nepenthophilus sp.n. (Malay Peninsula), P. noona sp.n. (Palawan), P. raptor sp.n. (Borneo), P. reticulatus sp.n. (Malay Peninsula) and P. simplex sp.n. (Borneo). Males and females are keyed and described. Pnyxiopalpus has a number of characters previously unknown in Sciaridae. In contrast to other Sciaridae, interspecific morphological variation is greater for the females compared to the males. Based on a parsimony analysis of sixty-four morphological characters, Pnyxiopalpus is monophyletic, and its sister group appears to be Spathobdella Frey + {[Peyerimhoffia Kieffer + (Faratsiho Paulian + Pnyxia Johannsen)] + [Hyperlasion Schmitz + (Hermapterosciara Mohrig & Mamaev + Parapnyxia Mohrig & Mamaev)]}. According to the most parsimonious solution, the monophyly of Hyperlasion, Lycoriella Frey and Plastosciara Berg in their current sense is questioned. Aptery and one-segmented maxillary palp, usually regarded as important in sciarid classification, show a lot of homoplasy.

Chvála, M. 2005: The Empidoidea (Diptera) of Fennoscandia and Denmark. IV. Genus Hilara. Fauna Entomologica Scandinavica, Vol. 40. Brill Academic Publishers, Leiden-Boston. ISBN 9004147993, 233 pp. Price 99 EUR. Lyneborg, Leif&Barkemeyer,Werner 2005: The Genus Syritta, A World Revision of the Genus Syritta Le Peletier & Serville, 1828 (Diptera: Syrhidae). Entomonograph Volume 15, A series facing global biodiversity in insects. Apollo books, Kirkeby Sand 19, DK-5771 Stenstrup, Denmark. ISBN 87-88757-53-6, Hardback, 224 pp. Price 420.00 DKK.

Lobosciara Steffan, characterized by unusual modifications in the male genitalia, is the sister group of Dolichosciara Tuomikoski. The concept of the genus is redefined on the basis of the new material. Lobosciara includes the following species which are keyed and described: L. spinipennis (Sasakawa) (Thailand, Micronesia), L. adebratti sp. n. (Borneo), L. bilobata sp. n. (Malay Peninsula, Borneo, Sulawesi), L. latiloba sp. n. (Malay peninsula, Borneo) and L. trilobata sp. n. (Sulawesi). The phylogenetic interrelationship of the species is as follows: [spinipennis + trilobata] + [latiloba + (adebratti + bilobata)].

Five new species of Keilbachia Mohrig are described: Keilbachia adstricta, K. constricta, K. cracens, K. fratercula and K. truncata. The species are the first sciarids recognized from New Caledonia and are similar in structure to the New Guinean K. indigena Mohrig and the Indomalayan K. adunca Hippa & Vilkamaa. A key to the Australasian species of Keilbachia is provided.

Camptochaeta prolixa sp. n. (Diptera, Sciaridae) is described from caves in Nevada, and three other congeneric species are recorded from caves in Nevada and Arkansas, United States. The new species shows some indication to a subterranean mode of life, including long antenna and legs, and in some specimens, reduction of the eye bridge.

Collection materials for specimens of the genus Keilbachia Mohrig, 1987 (Sciaridae) were studied. The following new species are described and figured: Keilbachia acumina sp. n. (Japan), K. adjuncta sp. n. (Taiwan), K. bifida sp. n. (Nepal), K. brevicoxa sp. n. (India), K. camptospina sp. n. (Vietnam), K. cornuta sp. n. (Japan), K. criniloba sp. n. (India), K. inscissa sp. n. (Nepal), K. macripes sp. n. (Nepal), K. megacantha sp. n. (Japan), and K. trispinoides sp. n. (India). Additionally, Sciara neglecta Johannsen, 1912, is combined in the genus Keilbachia Mohrig. New faunistic records are presented for the species Keilbachia ferrata (Japan, Finland, Sweden), K. neglecta (USA), K. praedicata (Taiwan) and K. sasakawai (Japan). All 52 species, currently included in Keilbachia, are listed, and a complete literature synopsis on the genus is provided.

The concept of Phytosciara Frey with the subgenera P. (Phytosciara), P. (Dolichosciara) and P. (Prosciara) is polyphyletic and the subgenera are treated as genera. Prosciara includes 1 Holarctic, 8 Palearctic and 28 Oriental species. The species are keyed and described, including the following new species: P. angustiloba sp. n. (Thailand), P. biceps sp. n. (Burma), P. bifida sp. n. (Burma), P. biloba sp. n. (Burma), P. collina sp. n. (Burma), P. crassidens sp. n. (Burma), P. decamera sp. n. (Burma), P. dolichochaeta sp. n. (Burma), P. duplex sp. n. (Burma), P. ensfera sp. n. (Burma), P. filichaeta sp. n. (Burma, Nepal), P. furcifera sp. n. (Burma), P. gemellata sp. n. (Burma), P. gibbosa sp. n. (Burma), P. glomerata sp. n. (Burma), P. latifurca sp. n. (Burma), P. latilingula sp. n. (Burma), P. megacera sp. n. (Burma), P. megachaeta sp. n. (Burma), P. mima sp. n. (Burma), P. pectinifera sp. n. (Japan), P. pentacanta sp. n. (Nepal), P. pentadactylasp. n. (Burma), P. plusiochaeta sp. n. (Finland), P. pollex sp. n. (Burma), P. processifera sp. n. (Burma), P. quantula sp. n. (Burma), P. tetrix sp. n. (Burma) and P. triloba sp. n. (Burma). Xenopygina Frey is not a synonym of Prosciara but an independent genus. It includes two species: X. hastata (Johannsen) n. comb. (USA) and X. paradoxa (Frey) (USSR).

Hippa, Heikki, Vilkamaa, Pekka (2006): Recharacterization of Rhynchoheterotricha Freeman (Diptera, Sciaroidea), with description of R. chandleri sp. n. from South Africa. Zootaxa 1167: 61-68, DOI: 10.5281/zenodo.2645499, {"references":["Chandler, P. (2002) Heterotricha Loew and allied genera (Diptera: Sciaroidea): offshoots of the stem group of Mycetophilidae and/or Sciaridae? Annales de la Societe Entomologique de France (n.s.), 38 (1-2), 101-144.","Freeman, P. (1951) Diptera of Patagonia and South Chile. Part III-Mycetophilidae. The British Museum (Natural History), London, 138 pp., plates I-XLIX.","Freeman, P. (1960) A new genus and species of Sciaridae (Diptera, Nematocera) from South Africa. Annals of the Natal Museum, 25, 75-77","Hippa, H. & Vilkamaa P. (2005) The genus Sciarotricha gen. n. (Sciaridae) and the phylogeny of recent and fossil Sciaroidea (Diptera). Insect Systematics & Evolution, 36, 121-144."]}

A cladistic analysis of the extant Sciaroidea was made, using re-evaluated morphological characters, with some additional taxa and with or without the Mesozoic Mesosciophilidae in the ingroup. When the new data were incorporated, the conflict between our earlier phylogenies, with extant taxa only and with both extant and fossil taxa in the ingroup, was resolved. The present result is similar to the one we reached earlier with extant + fossil taxa, which showed the Cecidomyiidae as the sister group of all other Sciaroidea, instead of the Cecidomyiidae appearing as the sister group of the Mycetophilidae + Sciaridae. In the new hypothesis, the main clades of the Sciaroidea outside the Cecidomyiidae were as follows: Ditomyiidae + (Diadocidiidae + ((Keroplatidae + (((Bolitophilidae + ((((Mesosciophila + (((((Starkomyia + rest of Sciaroidea))))). The placement of the recently described, enigmatic New Zealand genus Starkomyia Jaschhof was similar regardless of whether the fossil Mesosciophilidae were included in the analysis or not. The two stillundescribed Neotropical taxa that we included fell in the Rangomaraminae clade within the Sciaridae. Unlike our earlier result with extant taxa only, and similar to our extant + fossil result, the Ohakunea + Colonomyia clade now appears as the sister group of the MycetophilidaeLygistorrhinidae-Sciaridae lineage.

The following species of the genus Epidapus Haliday, 1851 are recognized from New Caledonia: Epidapus (s. str.) aciculatus sp. n., E. (Pseudoaptanogyna) angulatus sp. n., E. (s. str.) formosus sp. n., E. (Zuhalia) primus Mohrig, 2004 and E. (s. str.) triquetrus sp. n. The new species are described, and a key to the local species is given.

The genus Baeosciara Tuomikoski, 1960, Holarctic in distribution, includes the following species: B. discolor (Lengersdorf) (Europe), B. scotica (Edwards) (Europe, northern North America), B. sinuata (Menzel & Mohrig) (Europe) and B. pectinata sp. n. (western Canada). The new species is described, the other species are redescribed or diagnosed. Based on my earlier cladistic analysis of the Trichosia group of genera, the original generic rank of Baeosciara is restored.

The following species of the genus Pseudolycoriella Menzel & Mohrig, 1998 are recognized from New Caledonia, allnewly described: Pseudolycoriella acicula sp. n., P. bisulca sp. n., P. capillosa sp. n., P. latiflagellata sp. n., P. pallidula sp. n., P. senticosa sp. n., P. simplex sp. n., P. spicata sp. n., P. tenuis sp. n., P. tribulosa sp. n., P. trispicata sp. n., P. tumidior sp. n. and P. villosa sp. n. A key to the species is given.

Phylogeny of the Oriental genus Pnyxiopalpus Vilkamaa & Hippa, 1999 was re-studied in the light of the recently described, putatively related genus Vulagisciara Evenhuis, 2007, from Fiji, and a new species of Pnyxiopalpus, P. persimplex sp. n. from Selangor, Malaysia. Characters of these taxa were added to a previously published data matrix . The parsimony analysis yields two most parsimonious cladograms. In the strict concensus cladogram, Vulagisciara appears as the sister group of a monophyletic Pnyxiopalpus. Pnyxiopalpus persimplex sp. n. is described. The new species shares the basic structure of Pnyxiopalpus, but lacks some of the unique morphological modifications present in congeneric species. P. persimplex appears as the sister species to P. simplex, these two species together appearing as the sister group to all other Pnyxiopalpus.

When we (Hippa, Mattsson and Vilkamaa 2005a) proposed the genus name Blagorrhina gen. n., with two new species, we missed to include the type species designation for the genus. The subsequent correction (Hippa, Mattsson and Vilkamaa 2005b), in which we did not indicate the name to be ‘new’, failed to make the generic name available according to ICZN Article 16.1, and it remained a nomen nudum (Evenhuis et al. 2008).

Eight new, strictly sympatric species of the genus Keilbachia Mohrig, 1987, from Burma (Myanmar) are described and illustrated: Keilbachia aculeata, K. apprima, K. arcuata, K. arrecta, K. demissa, K. mira, K. scutica, and K. tenuicula. Keilbachia flagrispina Mohrig is redescribed on the basis of Burmese specimens. A key to all currently known species of Keilbachia is presented.

When preparing the final copy of our paper on Lygistorrhinidae (Hippa et al. 2005), the type speciesdesignation for the genus Blagorrhina was lost. We here designate Blagorrhina blagoderovi Hippa,Mattsson & Vilkamaa as the type species of Blagorrhina Hippa, Mattsson & Vilkamaa.We thank Dr. Christian Thompson (Washington, D.C.) for pointing out the missing type species designation.

The phylogeny of 50 genera or subgenera of the Sciaridae was studied by parsimony analysis. A cladistic analysis using 138 morphological characters from the adults of fifty-one ingroup and one outgroup species produced one most parsimonious cladogram. Schwenckfeldina, Chaetosciara, and Scythropochroa appeared successively as the sister groups of all other genera included in the analysis. The solution obtained was, in part, different from earlier, traditional views of sciarid phylogeny. The new Oriental genus Xenosciara had the phylogenetic relationship [Metangela + (Pnyxiopalpus + Xenosciara)]. Xenosciara invisa gen. n., sp. n., is described and illustrated. The new monotypic genus is exceptional in its hypopygial structures.