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SUMMARYFertilizer nitrogen (N) accounts for the majority of the greenhouse gas (GHG) emissions associated with intensive wheat production, and the form of fertilizer N affects these emissions. Differences in manufacturing emissions (as represented in the current carbon accounting methodologies) tend to favour urea, even when using the best available manufacturing technologies (BAT), whereas differences in fertilizer N efficiency and emissions of ammonia tend to favour ammonium nitrate (AN). To resolve these differences, data from 47 experiments in two large UK studies conducted from 1982 to 1987 and from 2003 to 2005 were reanalysed, showing that on average urea efficiency was 0·9 of AN (although mean ammonia emissions in 10 subsidiary experiments indicated an efficiency difference of 0·2); treating urea with a urease inhibitor (TU; AGROTAIN®, active ingredient N-(n-butyl) thiophosphoric triamide (n-BTPT)) brought its efficiency almost in line with AN; however, a significantly greater mean optimum N amount for TU (+0·1 of AN) was not fully explained. A standard response function relating wheat yield to applied AN was modified for degrees of relative inefficiency, thus enabling yields and GHG intensities (kg CO2e/tonne (t) grain) to be calculated using a PAS2050 compatible model for GHG emissions for any N amount of any N form. With AN manufactured by average European technology (AET), the estimated GHG intensity of wheat producing 8 t/ha was 451 kg/t; whereas with urea or TU made by AET it was 0.87–0.99 or 0.84–0.86 of this respectively. Using BAT for fertilizer manufacture, the grain's GHG intensity with AN and TU was 368 kg/t, but was 1·03–1·17 of this with untreated urea. The range of effects on GHG intensities arose mainly from remaining uncertainties in the inefficiencies of the N forms. Generally, economic margins and GHG intensities were not much affected by adjustments in N use for relative inefficiencies or different prices of urea-based fertilizers compared with AN. Overall, TU appeared to provide the best combination of economic performance and GHG intensity, unless the price for N as TU exceeded that for N as AN.

Knowledge of the changes in physiological traits associated with genetic gains in yield potential is essential to improve understanding of yield-limiting factors and to inform future breeding strategies. Recent advances in genetic yield potential and associated physiological changes in wheat (Triticum aestivum L.) are reviewed. Genetic gains in yield potential worldwide have been both positively correlated with harvest index (HI) and above-ground dry matter (AGDM), with more frequent reports of yield progress associated with biomass since about 1990. It is concluded that an important aim of future breeding will be the increase of biomass production while maintaining the present values of HI. In winter wheat recent biomass progress has been positively associated with pre-anthesis radiation-use efficiency (RUE) and water-soluble carbohydrate (WSC) content of stems at anthesis. Present results in two doubled-haploid (DH) populations show a positive linear relationship between stem WSC and grain yield in the UK environment. Results from various investigations worldwide in recent years have demonstrated that biomass increases have been associated with particular introductions of alien genes into wheat germplasm, e.g. the 1BL.1RS wheat-rye translocation and the 7DL.7Ag wheat-Agropyron elongatum translocation. Present results confirm a positive effect of 1BL.1RS on harvest biomass in two DH populations in the UK. The future prospects for identifying physiological traits to raise yield potential are considered with particular reference to winter wheat grown in northwestern Europe. It is proposed that optimized rooting traits, an extended stem-elongation phase, greater RUE, greater stem WSC storage and optimized ear morphology will be important for breeding progress in yield potential in future years.

In seeking better predictions of grain yield under light-limited conditions, shading was applied to field-grown winter wheat cv. Slejpner during each of five consecutive phases (canopy expansion, ear expansion, pre-flowering, grain expansion and grain filling). Absolute measures were taken of solar radiation and its effects on growth in three seasons, at a site where water and nutrient supplies were not limiting. Replicate mobile shades automatically occluded 0·80 of incident light when mean total solar radiation exceeded 250 J/m2 per s. Mean effects over seasons of shading on incident total solar radiation were −296, −139, −78, −157 and −357 MJ/m2 for the five phases respectively, and corresponding effects on shoot dry weight were −236, −184, −58, −122 and −105 g/m2. Estimated efficiency of radiation use after flowering was 1·2 g/MJ unshaded, tending to increase with shading. Shading in all phases reduced grain dry matter yield: mean effects over seasons were −106, −64, −61, −93 and −281 g/m2 for the five consecutive shading periods. Shading from GS31–39 increased mean maximum area of the two top leaves from 2700 to 3100 mm2 per leaf but, with fewer stems, canopy size remained unaffected. This and the next shading, from GS39–55, reduced specific leaf weight from 42 g/m2 by 4 and 3 g/m2 respectively, but effects on shoot dry weight were largely due to stem and ear. By flowering, stem weights, and especially their reserves of water-soluble carbohydrates, had partially recovered. Effects on yield of shading from GS31–39 were explained by a reduction in grains/m2 of 3070 from 26109. Shading from GS39–55 reduced grains/m2 by 4211 due to fewer grains per ear, whilst mean weight per grain increased in compensation. Shading from GS55–61 decreased grains/ear by 2·5. Shading from GS61–71 decreased ear growth and reduced stem weight, and at harvest resulted in 4·3 less grains/ear. Effects of the final shading from GS71–87 were fully explained by a reduction in mean dry weight/grain of 10·3 mg. Except for shading from GS71–87, source- and sink-based explanations of grain yield both proved feasible, within the precision of the measurements. Constraints to accurate comparison of source- and sink-based approaches are identified, and the implications for yield forecasting are discussed.

The plant characteristics required to make wheat lodging-proof for the least investment of biomass are calculated using a validated model of the lodging process and information about the dry matter costs of altering lodging-associated characters. The plant characteristics required to give a crop yielding 8 t ha−1 with 500 shoots m−2 and 200 plants m−2, a lodging return period of 25 years include a height of 0.7 m, a root plate spread of 57 mm, and for the bottom internode a wall width of 0.65 mm with a diameter/material strength combination ranging from a diameter of 5.86 mm with a material strength of 20 MPa to a diameter of 4.00 mm with a material strength of 50 MPa. It is estimated that this ideotype would require 7.9 t ha−1 of stem biomass and would have a harvest index of 0.42. Observations of a wide range of varieties grown using crop management to maximise lodging resistance without reducing yield potential showed that the root plate of the best variety was 7 mm less than the ideotype target, the stem character targets were achieved but not all in one variety, and the height target was achievable with the use of plant growth regulators. Plant breeders must therefore focus on selecting for a wider root plate and combining the appropriate stem strength characteristics.

Using a pair of near-isogenic lines(NILs) of winter wheat (Triticumaestivum L.) contrasting for the Ppd-D1 and ppd-D1 alleles, in eachof Mercia and Cappelle-Desprez, experimentsin two seasons (1997/8 and 1998/9) on aloamy medium sand examined differences inflowering date, resource capture, biomassproduction and grain yield responses toirrigation. Drought did not occur for anysustained period in unirrigated conditionsin 1998 due to high seasonal rainfall. In1999, drought developed post-floweringunder unirrigated conditions. Ppd-D1on average advanced flowering by 12 days inMercia and 9 days in Cappelle-Desprez.Earlier flowering with Ppd-D1 was dueto a shorter thermal duration from cropemergence to GS31, with no effect on thethermal duration from GS31 to GS61. In bothgenetic backgrounds, Ppd-D1 decreasedabove-ground dry matter (AGDM) at harvestin irrigated conditions by 0.3–0.9 tha-1 (p< 0.05), but thiswas compensated for by increases inharvest index (HI), so that grain yield wasconserved. Although Ppd-D1 decreasedmaximum green area index (GAI) by 0.8–1.9this was countered by greater maintenanceof green area after flowering, so thatradiation interception during grain fillingwas conserved. The Ppd-D1 alleledecreased season-long crop water uptake inthe Mercia NILs in irrigated conditions by39 mm. Effects of drought in 1999,averaging across NILs, were todecrease machine-harvested grain yield by 0.6 t ha-1 in Mercia and by 1.8 tha-1 in Cappelle-Desprez (p

The timing of flag leaf senescence (FLS) is an important determinant of yield under stress and optimal environments. A doubled haploid population derived from crossing the photo period-sensitive variety Beaver,with the photo period-insensitive variety Soissons, varied significantly for this trait, measured as the percent green flag leaf area remaining at 14 days and 35 days after anthesis. This trait also showed a significantly positive correlation with yield under variable environmental regimes. QTL analysis based on a genetic map derived from 48 doubled haploid lines using amplified fragment length polymorphism (AFLP) and simple sequence repeat (SSR) markers, revealed the genetic control of this trait. The coincidence of QTL for senescence on chromosomes 2B and 2D under drought-stressed and optimal environments, respectively, indicate a complex genetic mechanism of this trait involving the re-mobilisation of resources from the source to the sink during senescence.

Experiments in three dry years, 1993/94, 1994/95 and 1995/96, on a medium sand at ADAS Gleadthorpe, England, tested responses of six winter wheat cultivars to irrigation of dry-matter growth, partitioning of dry matter to leaf, stem and ear throughout the season, and to grain at final harvest. Cultivars (Haven, Maris Huntsman, Mercia, Rialto, Riband and Soissons) were selected for contrasts in flowering date and stem soluble carbohydrate. Maximum soil moisture deficit (SMD) exceeded 140 mm in all years, with large deficits (>75 mm) from early June in 1994 and from May in 1995 and 1996. The main effects of drought on partitioning of biomass were for a decrease in the proportion of the crop as lamina in the pre-flowering period, and then earlier retranslocation of stem reserves to grains during the first half of grain filling. Restricted water availability decreased grain yield by 1·83 t/ha in 1994 (PPPPPP2 in the unirrigated crop; greater accumulation appeared to be associated with better maintenance of HI under drought. It is concluded that high stem-soluble carbohydrate reserves could be used to improve drought resistance in the UK's temperate climate, but that early flowering seems less likely to be useful.

Six sites of variable soil type and environments in England and Scotland were sown with winter wheat (Triticum aestivum L. cv. Mercia) in the autumns of 1992 and 1993 with optimum inputs for growth. Crop monitoring between February and grain maturity provided data to investigate environmental and physiological factors important in controlling biomass accumulation.With increasing use of crop modelling as a tool for interpreting experiments and as crop management decision support systems, it is important that all influences on crop productivity are understood. The ‘radiation use efficiency’ or radiation conversion coefficient provides a convenient basis to study these influences.Significant differences in seasonal radiation conversion coefficients were observed between sites (P < 0·001), ranging from 2·82 to 3·87 g total dry matter/MJ absorbed photosynthetically active radiation. A series of simple dry matter models were developed to help explain biomass accumulation in relation to a number of environmental variables (using the measured green area index as an input) with correlation coefficients [ges ] 0·98 obtained across all sites. Apart from sunlight, differences in the canopy's ability to accumulate nitrogen and maintenance respiration costs were the most significant factors (P < 0·001). The nitrogen effect suggests changes in the conversion of assimilates with nitrogen availability, despite relatively high nutrition levels ([ges ] 180 kg N/ha).Over all sites the canopy extinction coefficient could be described as a linear function of the site spring time plant population (P < 0·01). A more detailed model considered canopy light attenuation to vary between sites according to sowing date and autumn/winter plant establishment and environment.

An experiment was conducted over seven sites in eastern England sown to winter cereals, whereby the soil mineral nitrogen (N) and N recovery in the crop were measured frequently during the active growth period. Plots received 100 kg/ha fertilizer N on one of three dates in spring (28 March, 11 April and 25 April) on each site in each of the three years (1994 to 1996), and these were compared with controls with no applied N. At one site, uncropped plots were included, and measurements of gaseous N emission, soil respiration, soil microbial biomass and root tissue N concentration were also made. The fertilizer applications boosted yields by at least 3 t/ha at all sites, but apparent recovery of fertilizer N varied throughout the previously known range of 45–85%. Soil type and timing of application had no effect on N recovery, or on final yields. N was mineralized from soil sources throughout the growth period, but mineralization was outweighed by ‘‘immobilization’’ of large amounts of N (around 30 kg/ha) in the soil, chiefly during May when crop growth was most rapid. This occurred on all sites with a crop, but not where the crop was absent. Measured losses of N2 and N2O were very small (

Experiments at three sites in 1993, six sites in 1994 and eight sites in 1995, mostly after oilseed rape, tested effects of previous fertilizer N (differing by 200 kg/ha for 1993 and 1994 and 300 kg/ha for 1995) and date of sowing (differing by about 2 months) on soil mineral N and N uptake by winter wheat cv. Mercia which received no fertilizer N. Soil mineral N to 90 cm plus crop N (‘soil N supply’; SNS) in February was 103 and 76 kg/ha after large and small amounts of previous fertilizer N respectively but was not affected by date of sowing. Previous fertilizer N seldom affected crop N in spring because sowing was too late for N capture during autumn, but it did affect soil mineral N, particularly in the 60–90 cm soil horizon, presumably due to over-winter leaching. Tillering generally occurred in spring, and was delayed but not diminished by later sowing. Previous fertilizer N increased shoot survival more than it increased shoot production. Final shoot number was affected by previous fertilizer N, but not by date of sowing. Overall, there were 29 surviving tillers/g SNS.N uptakes at fortnightly intervals from spring to harvest at two core sites were described well by linear rates. The difference between sowings in the fitted date with 10 kg/ha crop N was 1 month; these dates were not significantly affected by previous fertilizer. N uptake rates were increased by both previous fertilizer N and late sowing. Rates of N uptake related closely to soil mineral N in February such that ‘equivalent recovery’ was achieved in late May or early June. At one site there was evidence that most of the residue from previous fertilizer N had moved below 90 cm by February, but N uptake was nevertheless increased. Two further ‘satellite’ sites behaved similarly. Thus at 14 out of 17 sites, N uptake until harvest related directly and with approximate parity to soil mineral N in February (R2 = 0·79), a significant intercept being in keeping with an atmospheric contribution of 20–40 kg/ha N at all sites.It is concluded that, on retentive soils in the UK, SNS in early spring was a good indicator of N availability throughout growth of unfertilized wheat, because the N residues arising from previous fertilizer mineralized before analysis, yet remained largely within root range. The steady rates of soil mineral N recovery were taken as being dependent on progressively deeper root development. Thus, even if soil mineral N equated with a crop's N requirement, fresh fertilizer applications might be needed before ‘equivalent recovery’ of soil N, to encourage the earlier processes of tiller production and canopy expansion. The later process of grain filling was sustained by continued N uptake (mean 41 kg/ha) coming apparently from N leached to the subsoil (relating to previous fertilizer use) as well as from sources not related to previous fertilizer use; significant net mineralization was apparent in some subsoils.

The effects of eight rates of nitrogen (N) application (0–245 kg/ha) on the performance of winter wheat over five seasons (1991–1995) on a long-term field experiment (established 1978) at Ropsley (UK) are described. In each of the five seasons, N was withdrawn from replicate plots in order to study the residual effect of fertilizer. N applications in excess of 140 kg/ha left significant residues as soil mineral N (SMN) in the autumn which, despite some loss over-winter, had a significant effect on the yield and N offtake of the subsequent crop. The amount of N carried over was equivalent to 8–20% of the fertilizer N and was observed at N applications up to 40 kg/ha lower than the optimum rate (c. 200 kg/ha). Part of the unrecovered N was also considered to contribute to the long-term build-up of fertility at the site. The results suggest that restrictions on N use to below the optimum will reduce leachable N, but may have an impact on soil fertility and future crop productivity. In addition, the rate of N applied to preceding crops should be taken into account when formulating fertilizer advice on retentive soils.

A simple model of canopy expansion and senescence with accumulated thermal time from sowing was used to describe differences in canopy development and potential size across a range of sites in the UK between 1992 and 1995. The principal model inputs were nitrogen, temperature and sowing date. The model was calibrated across six sites within the major wheat growing areas using canopy data collected during the 1992/93 and 1993/94 growing seasons and was validated using data collected in the 1994/95 season. The model was able to predict the time course of green area index (GAI) over a season with an r2[ges ]0·87 for the five English sites, and an r2=0·68 for the single Scottish site. This model may prove to be a useful approach to forecasting the potential canopy size, based upon an estimate or measure of the total nitrogen available to the crop (both from applied fertilizer and the soil).

Amounts of spring nitrogen (N) fertilizer (0–240 kg/ha), combined with three timing treatments (single, divided early or divided late), were tested at 14 sites in England and Wales between 1984 and 1988 to determine the optimum fertilizer N requirement for winter oats. The trials were superimposed on commercial crops of the cultivars Pennal (9 sites) or Peniarth (5 sites). Optimum amounts of N ranged from nil to 202 kg/ha (mean 119) and optimum yields varied between 5·8 and 9·9 t/ha (mean 7·3). Much (c. 60%) of the inter-site variation in N optimum was explained by differences in soil N supply, as indicated by N offtake in the grain at nil applied N. Mean yield differences between single and early (+0·08 t/ha) or late (−0·04 t/ha) divided dressings were slight, although significant (PLodging occurred at 11 of the 12 sites where lodging scores were recorded and always increased significantly (PPGrain N concentrations increased significantly (P

The responses to fertilizer N of winter wheat and winter barley grown on sandy soils were measured in 74 experiments in England from 1991 to 1994. A series of curves was fitted to the data from each of the experiments to determine which best described the yield response to fertilizer N. The linear plus exponential model, adopted for data from a range of other soil types, with the rate parameter (r) determined for each experiment, proved to be the most satisfactory for these data. The median value of r at 0·987 was less than has usually been determined for other soil types, and was reduced by increasing moisture stress in June, suggesting that the reduced efficiency with which recovered N produces assimilate is partly a result of moisture stress. The model produced a mean Nopt of 143 kg/ha following cereals.Soil N supply in spring (SNSs) was small at c. 45 kg/ha and commensurate with a large requirement for fertilizer N. Apparent fertilizer recovery (AFR) at 0·70 was greater than that previously measured on clay or chalk soils, suggesting that AFR was not restricted by lack of soil moisture. The larger AFR is considered to be a consequence of more rapid remobilization of fertilizer N in these sandy soils. The greater recovery of fertilizer N reduced the optimum requirement for fertilizer N. Nitrogen harvest index (NHI) was decreased from 0·76 to 0·70 as fertilizer N increased from 0 to 300 kg/ha, whereas dry matter harvest index (HI) decreased from 0·53 to 0·50. The economic optimum requirement for fertilizer N, at 143 kg/ha, was less than reported for other soil types.The increase in grain yield from applied fertilizer N was less in these experiments than in those reported on other soil types in the UK. However, this overall observation masks two counterbalancing differences. Fertilizer N was recovered more efficiently on these soils; the recovered fertilizer N was used with similar efficiency to produce grain dry matter. The explanation proposed is that on these soils a large proportion of soil water is readily available to crops, and this was taken up rapidly in May accompanied by fertilizer N. However, shortage of water in June restricted the ability of the crop to produce assimilate from fertilizer N applications >c. 150 kg/ha.

The response of cultivars to applied nitrogen was examined in 11 seasons, 1982–92, in two experiments per year, normally testing seven cultivars at seven rates of fertilizer nitrogen. In all, 27 cultivars were tested in 22 experiments throughout Nottinghamshire, Lincolnshire, Northamptonshire and Suffolk. Cultivars ranged in their date of introduction from Maris Huntsman (1969) to Hereward (1988). For each cultivar in each experiment, the economic optimum yield (Yopt), the amount of fertilizer N needed to produce it (Nopt), the grain %N at Nopt, the offtake of N in the grain at nil N (Noff(N0)) and Nopt (Noff(opt)) and the estimated recovery of fertilizer in the grain at Nopt (AFRopt) were estimated by fitting linear plus exponential curves to data for grain yield and two-straight-line models to data for grain N offtake. From cross-site analysis, normalized cultivar means were calculated for each variate. Over the 20-year period relating to the cultivars in the trial, the contribution of new genotypes to grain yield improvement was 1·92 t/ha, Yopt increasing by 96 kg/ha per year. There was no change in grain %N at Nopt. The effect of changes through breeding from 1969 to 1988 was to increase Noff(opt) by 42 kg/ha (2·1 kg/ha per year), that was associated with a decrease in Noff(N0) (equivalent of soil N offtake) of 15 kg/ha (0·77 kg/ha per year). Part of the increased requirement for fertilizer N was fulfilled by an increase in AFRopt of 18% over the 20-year period. The net effect was for Nopt itself to increase by 56 kg/ha (2·8 kg/ha per year). Since survey evidence indicates no general increase in N use on wheat by farmers since the mid-1980s, it appears that current fertilizer use by farmers may be underestimating the requirement for N now. Alternatively in previous years N requirements may have been overestimated. The change in N available for loss to the environment, from the balance of grain Noff(opt) and Nopt, was from 11 kg N/ha in 1969 compared to 25 kg N/ha in 1988. It seems possible that the potential increase in nitrate levels in groundwater associated with plant type may not have been realised because farmers have conserved the amount of N they use.

Functions which predict rate of leaf emergence and final number of leaves, used in a model to predict the date at which growth stages occur, were tested in an experiment on winter wheat cv. Mercia grown with standardized husbandry at six sites in 1992/93, 1993/94 and 1994/95. A study of the number of detectable nodes on the culm and leaf length was also made.The predicted rate of leaf emergence was mostly within 5% of the observed value. The difference between observed and predicted final number of leaves was mostly less than half a leaf but suspected errors in leaf counts resulted in some differences of more than two leaves.Variable extension of the basal internode impaired confidence in the detection of nodes. The mean number of detectable nodes differed significantly among sites and between seasons from 3·7 to 4·8 but could not be related to sowing date or final number of leaves. Further information on factors affecting extension of the basal internode is desirable to standardize node detection and improve prediction of culm leaf appearance.Culm leaves showed successively longer laminae up to the penultimate leaf. There was a significant relationship between length of the flag leaf and the final number of leaves, but it was positive in 1993/94 and negative in 1994/95. This may have been due to greater water stress in 1994/95.

In 1977, a four-course rotation was set up at Ropsley (UK) to study crop response to eight rates of nitrogen application (35–265 kg ha−1). This rotation continued until 1990 when continuous winter wheat was introduced. Results from 1978 to 1990 provide an opportunity to study the initial phase of cumulative effects from different rates of N fertilizer application on the recovery of N by cereals and the retention of N in the soil.From 1978 to 1990, considerable variation in the recovery of nitrogen by winter wheat was observed. Neither rainfall nor drainage, as indicators of possible denitrification or leaching losses, provided a useful explanation for this, possibly because of the relatively dry conditions prevailing after spring fertilizer application. There was no evidence of increased soil N fertility, beyond single year residues, as a result of large N applications over the 13-year period.In order to achieve the economic optimum grain yield, it was necessary to use N applications which produced inefficient recovery of N. Thus, greater return of N in crop residues and immobilization at relatively large N applications (>150 kg ha−1) contributed to an observed build-up in soil organic N over the period of study. Plots receiving, on average, 265 kg ha−1 appeared to gain c. 250 kg ha−1 N over control plots (35 kg ha−1) after 13 years of N application. Reducing the N application rate from the economic optimum to a more biologically efficient N rate (156 kg ha−1) was calculated to result in an average yield loss of 0·305 t ha−1 and cause an estimated £17 ha−1 loss in profit.

In 1992 and 1993, eight rates of 15N-labelled fertilizer (0–245 kg ha−1) were applied to winter wheat growing on the Ropsley long term field trial where eight different N amounts had been applied annually since 1978. The fate of the labelled N in the crop and topsoil (0–23 cm) was determined at harvest in the year of 15N application and in the first and second residual years.By harvest in the second residual year, 60–77% of the original labelled application had been recovered in the crop and topsoil with 23–40% lost. These losses virtually all occurred within the first two growing seasons; there was no significant loss during the second residual year. Significant changes in the 15N balance were observed at N applications in excess of the range 140–175 kg ha−1 which suggested a marked decrease in the efficiency of N use and an increase in residual labile N in the soil at harvest. At low N applications (−1), a positive added nitrogen interaction (ANI) was observed: 40–50% of this was a residual ANI due to the short or long term effect of applying N fertilizer, and the remainder was probably an apparent ANI due to pool substitution in the immobilization process. At large N applications (>175 kg ha−1) a negative ANI was observed: large N applications resulted in a net suppression of soil N uptake due to substitution by fertilizer for a limited plant N demand.

In 74 experiments carried out in England from 1990 to 1994, cereal test crops were grown on sandy soils prone to nitrate leaching, following cereals, sugarbeet and potatoes. Measurements were made of the effects of the previous crops on soil mineral N, and on N recovery by the cereal test crop.Soil N supply in autumn (SNSa) was greater following potatoes (c. 100 kg/ha N) than following sugarbeet or cereals (c. 60 kg/ha N). However when potato crops to which organic manures had been applied were excluded, mean SNSa after potatoes was only c. 60 kg/ha. Soil N supply in spring (SNSs) following sugarbeet and potatoes was similar and at c. 56 kg/ha, c. 10 kg/ha greater than following cereals. Seasonal differences in SNSs were related to excess winter rainfall and soil water-holding capacity. Modelled leaching losses gave good agreement with overwinter changes in SNS (r=0·87), although SNSs was usually greater than predicted by the model. This discrepancy was considered to be due to overwinter mineralization, which was estimated from the intercept of the regression lines to be c. 40 kg/ha following cereals and potatoes, and c. 50 kg/ha following sugarbeet.Apparent net mineralization (AM) of N during the test crop growing season was c. 37, 53 and 63 kg/ha following cereals, sugarbeet and potatoes respectively. However, AM was c. 24 kg/ha N greater if the preceding potato crop had been given organic manures, but there was no difference in AM following manured and unmanured sugarbeet crops.Nitrogen offtake by the cereal test crop without fertilizer N (NoffN0) was c. 15·20 kg/ha greater following potatoes and sugarbeet than following cereals. The greater N offtake following potatoes is considered to be due to mineralization of organic manures applied to the potato crop, while the effect following sugarbeet appears to be due to mineralization of sugarbeet residues. At harvest, SMN was c. 38 kg/ha and similar for all three previous crops. It is concluded that mineralization of sugarbeet residues has taken place more quickly on these sandy soils than previously reported on soils of greater clay content. Only potatoes grown with organic manures leave greater N residues than cereals.

Twenty-six experiments were carried out in England and Wales from 1983 to 1985 to compare urea and ammonium nitrate (AN) as N top-dressing for cereals over the range 100–300 kg/ha N. Most of the experiments were sited on chalky or other soils of pH>7·0 where the greatest differences in effectiveness were expected.The results showed that while relative N offtake in grain was 2·5% greater from AN, there was no difference in grain yield. Splitting the main N application increased N offtake from urea but not from AN. Overall, urea effectiveness from a single application at GS 31 increased with increasing rainfall on the day of application, but was not increased by subsequent rainfall. On chalk soils, urea effectiveness increased with increasing cumulative rainfall up to the fifth day after application. There was little effect of rainfall on urea effectiveness when the dressing was split. Although grain yield was unaffected by the type of fertilizer N applied, grain N concentration was usually less from urea.The fertilizer N requirement for optimum yield (Nopt) wassimilar for both fertilizers. Splitting the main N application had no effect on yield or Nopt. We concludethat urea is a satisfactory source of N for cereals.

SUMMARYSoil mineral nitrogen (Nmin) was measured to 90 cm at a total of 12 sites in the UK in the autumn after an oilseed rape experiment, which measured responses to fertilizer N. On average, Nmin, increased by 15 kg/ha per 100 kg/ha fertilizer nitrogen (N) applied to the rape, up to the economic optimum amount of N (Nmin). There were larger increases in Nmin where fertilizer applications exceeded Nopt, thus super-optimal fertilizer applications disproportionately increased the amount of nitrate likely to leach over-winter. The small effects of sub-optimal N on Nmin were associated with large increases in N offtake by the oilseed rape, whereas the larger effects of super-optimal N on Nmin were associated with only small increases in N offtake. Over 70% of the variation in autumn Nmin was explained by the previous rape's N fertilizer rate and the topsoil organic matter content.Nitrogen applied to the rape increased grain yields of the succeeding wheat crops when no further fertilizer N was applied to the wheat. It was concluded that N applied to oilseed rape significantly affected Nmin after harvest, and these effects were not completely nullified by leaching over-winter, so soil N supply to the succeeding wheat crop was significantly increased. Responses in grain yield indicated that each 100 kg/ha N applied to the rape provided N equivalent to c. 30 kg/ha for the following cereal. Each 1% of soil organic matter further contributed N to the wheat, equivalent to 25 kg/ha.It is important to ensure that oilseed rape receives no more than the optimum amount of fertilizer N if subsequent leaching is to be minimized. Reductions below optimum amounts will have only a small effect on leaching. Substantial changes in the economic optimum N for rape production should be accompanied by adjustment in fertilizer N application to following wheat crops. Fertilizer recommendation systems for wheat should take account of the fertilizer N applied to the preceding oilseed rape and the topsoil organic matter content.

SUMMARYAt four pairs of sites in the UK, two cultivars of wheat, bred either for autumn (cv. Mercia) or for spring (cv. Tonic) sowing were sown in November 1988 and 1989 and their yield response to springapplied fertilizer N assessed over the range 0–320 kg/ha N. Grain samples were taken at harvest and analysed for those qualities regarded as indicators of suitability for milling. In five experiments, the grain yield of Mercia was up to 1 t/ha greater than that of Tonic, but overall there was no consistent difference between the two cultivars in the optimum economic yield (Yopt)or in the amount of fertilizer N needed to produce it (Nopt). Both cultivars produced c. 17·5 kg of grain per kg of fertilizer N applied up to Nopl. In the five experiments in which Tonic yielded less, it produced grain of greater N concentration than Mercia when given the same amount of fertilizer N. Differences in grain N concentration, while small (< 0·3%), were significant when analysed across sites. In six experiments both cultivars produced grain of 11 % protein at Nopt. There were no consistent or worthwhile increases in specific weight, Hagberg falling number (HFN) or response to the sodium dodecyl sulphate (SDS) sedimentation test of either cultivar from applying more fertilizer N than Nopl. The average specific weight of Mercia grain was 2·5 kg/hi greater than that of Tonic, while Tonic gave larger SDS values of c. 8. There was little difference in HFN. It was concluded that the yield requirement for fertilizer N was similar for these two cultivars of wheat when sown in late autumn, and there was no need to apply differential fertilizer N applications to the cultivars to produce grain of the quality needed for breadmaking. In four experiments the application of plant growth regulator (PGR) increased the yields of both cultivars by c. 0·5 t/ha.

SUMMARYAt 14 sites in the UK, spring wheat (Triticum aestivum) cv. Tonic, was sown on three or four dates at each site between October and March in the 1988/89, 1989/90 and 1990/91 seasons. Responses to spring-applied fertilizer N over the range 0–320 kg/ha were determined. Earlier sowing did not increase uptake of soil N by the crop. Fertilizer N increased grain N offtake by between 25 and 140 kg/ha and yield by between 0·3 and 5·5 t/ha, although grain yield was less responsive to fertilizer N at later sowing dates. Apparent recovery of fertilizer N (AFR) also decreased as sowing was delayed but there was no effect of delayed sowing on the amount of grain produced from each kg of fertilizer N recovered. Because fertilizer N recovery decreased with later sowing, the amount of fertilizer N needed to produce the optimum economic grain yield was not reduced. Neither AFR nor optimum fertilizer (Nopt) was related to optimum yield. Regression of Nopt on the difference between optimum yield and yield without fertilizer N (△y) explained 77% of the variance in Nopt. There was an inverse relationship between △y and soil mineral N (SMN) in spring; regression of △y, on SMN in spring accounted for 29% of the variance in △y Current advisory systems which adjust economic fertilizer N recommendations according to anticipated yield are not justified by these results. Moreover the adjustments made, based on yield expectation, appear about three times as large as those needed to minimize residues of fertilizer N left unrecovered by the crop and to reduce the risk of nitrate leaching in the following winter.

SUMMARYNitrogen nutrition of two succeeding wheat crops was studied after ploughing of grassland in July 1987 on a clay soil at ADAS Drayton. The four plots of grassland had received 100, 250, 450 and 750 kg N/ha per year for 4 years from 1984 and were grazed by beef cattle at stocking densities which varied according to grass growth.Determinations of soil mineral N taken to 60 cm every 3 weeks from July to the following May were particularly variable. However, in the first 2 years after ploughing the means of the series of mineral N measurements were directly proportional to the amounts of fertilizer N applied to the grass.N offtake in winter wheat grain without fertilizer N was directly proportional to fertilizer N applied to grass but this had little effect on maximum grain yields. Large soil N supplies did not necessarily predispose the wheat crops to large grain N concentrations because fertilizer N caused grain N offtake to reach a similar maximum, irrespective of previous grass N.Optimum amounts of fertilizer N for the wheat were 188, 147, 87 and nil kg/ha in 1988 and 152,130, 89 and 25 kg/ha in 1989 after 100, 250, 450 and 750 kg N/ha per year applied to the grass. Soil N supply as indicated by both the amount of fertilizer applied to grass and means of mineral N measurements accounted for almost all of this variation. Mean soil mineral N over winter was no better as an indicator of soil N supply than the amount of N applied to the grass. However, before adopting N applied to grass as a more general index of N supply, it would need to be adjusted for variation in N removed and lost during grass growth; these were controlled in this experiment.

SUMMARYThree experiments carried out at Bridget's Experimental Husbandry Farm, Hampshire between 1984 and 1987 on clay loam and silty clay loam soils over chalk investigated the response of winter wheat grown after grass to spring-applied fertilizer nitrogen. The influence of method of establishment and the use of a nitrification inhibitor were also studied. Yields were similar whether the crops were established by direct drilling or following ploughing. Crops required between 88 and 209 kg N/ha nitrogen fertilizer for optimum yield. There was no consistent difference in nitrogen requirement nor were there consistent differences in nitrogen offtake (which is the amount of N in kg/ha removed in harvested grain) or apparent fertilizer recovery following ploughing or direct drilling. The use of a nitrification inhibitor increased grain yield when applied in the autumn without fertilizer, but not when applied with fertilizer in the spring.

SUMMARYThe fertilizer-nitrogen (N) requirement for wheat grown in the UK varies from field to field. Differences in the soil type, climate and cropping history result in differences in (i) the crops’ demands for N, (ii) the supply of N from the soil (SNS) and (iii) the recovery of the fertilizer by the crops. These three components generally form the basis of systems for N recommendation. Three field experiments were set out to investigate the variation of the N requirement for wheat within fields and to explore the importance of variation in the crops’ demands for N, SNS and fertilizer recovery in explaining the differences in the economic optima for N. The N optima were found to vary by >100 kg N/ha at two of the sites. At the other site, the yield response to N was small. Yields at the optimum rate of N varied spatially by c. 4 t/ha at each site. Soil N supply, which was estimated by the unfertilized crops’ harvested N, varied spatially by 120, 75 and 60 kg/ha in the three experiments. Fertilizer recovery varied spatially from 30% to >100% at each of the sites. There were clear relationships between the SNS and the N optima at all the three sites. The expected relationship between the crop's demand for N and N optima was evident at only one of the three sites. There was no consistent relationship between the N recovery and the N optima. A consistent relationship emerged, however, between the optimal yield and SNS; areas with a greater yield potential tending to also supply more N from the soil. This moderated the expected effect of the SNS and the crop's demand for N on the N optima.