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Deadwood is a major component of aboveground biomass (AGB) in tropical forests and is important as habitat and for nutrient cycling and carbon storage. With deforestation and degradation taking place throughout the tropics, improved understanding of the magnitude and spatial variation in deadwood is vital for the development of regional and global carbon budgets. However, this potentially important carbon pool is poorly quantified in Afrotropical forests and the regional drivers of deadwood stocks are unknown. In the first large-scale study of deadwood in Central Africa, we quantified stocks in 47 forest sites across Gabon and evaluated the effects of disturbance (logging), forest structure variables (live AGB, wood density, abundance of large trees), and abiotic variables (temperature, precipitation, seasonality). Average deadwood stocks (measured as necromass, the biomass of deadwood) were 65 Mg ha -1 or 23% of live AGB. Deadwood stocks varied spatially with disturbance and forest structure, but not abiotic variables. Deadwood stocks increased significantly with logging (+38 Mg ha -1 ) and the abundance of large trees (+2.4 Mg ha -1 for every tree >60 cm dbh). Gabon holds 0.74 Pg C, or 21% of total aboveground carbon in deadwood, a threefold increase over previous estimates. Importantly, deadwood densities in Gabon are comparable to those in the Neotropics and respond similarly to logging, but represent a lower proportion of live AGB (median of 18% in Gabon compared to 26% in the Neotropics). In forest carbon accounting, necromass is often assumed to be a constant proportion (9%) of biomass, but in humid tropical forests this ratio varies from 2% in undisturbed forest to 300% in logged forest. Because logging significantly increases the deadwood carbon pool, estimates of tropical forest carbon should at a minimum use different ratios for logged (mean of 30%) and unlogged forests (mean of 18%)., (© 2016 John Wiley & Sons Ltd.)

Deadwood in forests plays a critical role in maintaining the ecological functions. Small mammals use deadwood, and thus deadwood can mitigate the negative impacts of plantation on small mammals. This study focused on fallen dead trees in planted forests, and aimed to verify whether fallen dead trees affect behavioral patterns of Japanese squirrels. To clarify the use of fallen dead trees by Japanese squirrels in a Japanese cedar plantation, we observed squirrel behavior by camera trap surveys at 61 survey sites. Our findings showed that fallen dead trees play a crucial role in the behavior of Japanese squirrels, serving as landmarks for movement, vigilance, resting, and hoarding sites. These functions are critical for the survival of Japanese squirrels, suggesting that fallen dead trees in planted forests have positive impacts on their microenvironment use. The increase of deadwood due to disturbances such as heavy rainfall and snowfall resulting from climate change may provide benefits to arboreal small mammals in poorly managed planted forests. Additionally, leaving some of the deadwood generated during the harvesting process in properly managed forests can improve the quality of habitat for arboreal small mammals.

In the managed beech forests of Central Italy (Molise), the diversity of saproxylic species is potentially under threat by intensive management. To evaluate the impact of forestry on the biodiversity of these ecosystems, we analyzed the relationship between abundance of saproxylic beetles and (i) forest stand exposure and elevation, (ii) deadwood availability (coarse woody debris - CWD - and stumps), (iii) abundance of microhabitats. Four sampling sectors with different altitudes and exposure were identified in a 400-ha study area in the Appenine mountains. Fifteen circular plots (13 m radius) were established in each sector where deadwood and microhabitats were surveyed and saproxylic beetles sampled. We fitted joint species distribution models to quantify the relationship between forest attributes and saproxylic species’ abundance, including the interactions with their family and trophic category. Overall, 2334 specimens belonging to 64 species of saproxylic beetles were collected. Both abundance and species richness were higher in the sectors with high elevation (respectively, 55% and 44%) and South exposure (respectively, 28% and 44%). Average deadwood volumes were low (stumps: 7.6 m3 ha-1; CWD: 0.3 m3 ha-1; snags: 0.4 m3 ha-1), and insect galleries were the most abundant microhabitat (380 records over a total of 434). The most important variables affecting abundance were stump characteristics (model deviance = 81.2), elevation (deviance = 64.7), and CWD characteristics (deviance = 58.0). Our results show that topographical variables and forest structure jointly affect the abundance patterns of saproxylic beetle communities in managed beech forests. These ecological interactions imply that management has different impacts on the saproxylic communities in different topographic conditions. To acknowledge this complexity we advocate for a landscape-level forest management supporting the local beetle diversity maintaining a mosaic of semi-natural forest characteristics in different topographic contexts. The ecological value of the forest landscape will be further enhanced by the application of closer-to-nature management interventions based on deadwood retention, microhabitat creation and tree retention, in line with the guidelines of the new EU Forest Strategy for 2030.

Deadwood in forests plays a critical role in maintaining the ecological functions. Small mammals use deadwood, and thus deadwood can mitigate the negative impacts of plantation on small mammals. This study focused on fallen dead trees in planted forests, and aimed to verify whether fallen dead trees affect behavioral patterns of Japanese squirrels. To clarify the use of fallen dead trees by Japanese squirrels in a Japanese cedar plantation, we observed squirrel behavior by camera trap surveys at 61 survey sites. Our findings showed that fallen dead trees play a crucial role in the behavior of Japanese squirrels, serving as landmarks for movement, vigilance, resting, and hoarding sites. These functions are critical for the survival of Japanese squirrels, suggesting that fallen dead trees in planted forests have positive impacts on their microenvironment use. The increase of deadwood due to disturbances such as heavy rainfall and snowfall resulting from climate change may provide benefits to arboreal small mammals in poorly managed planted forests. Additionally, leaving some of the deadwood generated during the harvesting process in properly managed forests can improve the quality of habitat for arboreal small mammals. [ABSTRACT FROM AUTHOR]

The study aimed to quantify annual mortality and recruitment rates on permanent research plots in the Badínsky prales old-growth forest. The data measured in four stands originate from six censuses, which together cover a 48-year period. The mean annual mortality rate reached 4.2% (DBH > 2 cm) or 2.3% (DBH > 8 cm). The mortality peaked in the first part of the observation period, probably indicating an intermediate disturbance activity, and the subsequent declining trend led to minimum annual mortality during the last 20 years. We found substantially higher fir mortality in comparison with beech, primarily during the first decades of the measurement period. Among three size categories, the stems with DBH 2-8 cm showed the highest mortality rate. We detected the minimum fir recruitment rate throughout the observation period; on the other hand, the beech recruitment rate was relatively high responding to the open canopy. Standing and lying deadwood volumes indicated similar trends like mortality did, and high values of fir standing deadwood observed during the first decades can signify that a relatively high proportion of mortality was related to the fir decline caused by air pollution.

While healthy forest ecosystems deliver various services that can reduce flood risk, they can also contribute to flooding by providing wood that potentially contributes to the clogging of waterways and associated backwater effects. In this regard, deadwood, as a key aspect of healthy forests, is often in focus of post-flood disaster discourses. This research reflects on this ambiguity in the different forest management goals when it comes to managing forests for flood risk reduction versus forest health. A working definition of forest health will be presented and an overview of the different aspects of how a forest potentially can affect the flood hazard will be provided. This will refer to the ways forests influence (1) the discharge of water from the landscape into channels and (2) the characteristics of the channel and its riparian area and their respective influence on the transport of water, sediment, and debris. Often these two determining factors for the development of the flood peak are discussed separately and by different academic fields. This paper aims to connect the existing knowledge spheres and discusses the synergies and trade-offs. The review shows that the two objectives of forest health and flood risk reduction are largely synergetic. However, in direct proximity to watercourses trade-offs might occur. This is especially due to the ambivalent relation of living vegetation and deadwood to flood hazard. In places without susceptible infrastructures to clogging, deadwood and diverse vegetation structures should be supported due to their beneficial effects on water retention and channel characteristics. In places where susceptible infrastructures exist, trade-offs between the two objectives arise. Here the potential of freshly uprooted vegetation to cause damages should be reduced while maintaining the vegetation’s supportive characteristics, for example, concerning bank and slope stability. Where the risk of clogging is assessed as too high, also the selective removal or shortening of dead in-channel Large Wood can be considered. However, based on the literature review the risk deriving from dead Large Wood is evaluated as comparably low. This is related to its generally lower proportions and its smaller and less stable characteristics compared to freshly uprooted vegetation.

Riparian buffers are commonly used to mitigate the negative effects of forestry operations near water, particularly sediment transport to streams. In Sweden, current practices typically involve 5-7 m wide riparian buffers along small streams. Historical forest management, which favored conifers up to the channel edge, has resulted in these narrow buffers having a simplified tree species composition and structure, making them prone to windthrow. While windthrow can contribute large wood (LW) to streams, windthrow also risks increasing sediment inputs if rootwads are exposed near stream edges. This disturbance affects sediment connectivity, or the movement of particles through the fluvial system, but the interaction between LW dynamics and sediment connectivity in small boreal streams is not well understood. We investigated sediment connectivity at the Trollberget Experimental Area in northern Sweden, where six 100 m stream reaches had either 5 m or 15 m wide riparian buffers. Pre-harvest and one-year post-harvest data on windthrow, hydrology, and sediment yields were collected. Forest harvesting increased sediment connectivity in the streams regardless of buffer width, indicating that buffers wider than 15 m are necessary to reduce sediment input impacts in small headwater streams. Windthrow affecting stream channels was more common in the 5 m buffers, leading to significantly higher deposition of very fine sediments (<250 μm) compared to the 15 m buffers. Coarse (>1 mm) and fine sediments (250 μm – 1 mm) were also higher in the 5 m buffers. We found that sediment connectivity in streams was closely linked to LW dynamics, negatively before harvest but positively after harvest. Before harvest, LW trapped sediment and prevented downstream transport, but after harvest, the increased sediment input overwhelmed this function. Our results highlight a trade-off between the recruitment of LW and minimizing sediment connectivity, two key objectives in riparian buffer management. • Riparian buffers are important to protect stream ecosystems when the adjacent forest is cut. • We applied 5 m and 15 m wide buffers and followed the impact on sediment connectivity. • Instream wood trapped sediments until harvest; windthrow disrupted this function. • All size fractions of sediment increased after harvest, regardless of buffer width. • Current buffer practices prevent providing both instream wood and sediment trapping. [ABSTRACT FROM AUTHOR]

In recent years, the loss of biodiversity occurred worldwide has called for the creation of various indicators aimed to assess the health status of the environment. Due to the increase of forest areas throughout Europe, different ecological indicators for the assessment of naturalness and health of forest ecosystems are being theorized and modelled, with relevant implications both for planning current and future management of forest ecosystems and to control their evolution over time. In this study, a new indicator of the ecological naturalness of forest habitats is proposed with the aim of providing an intuitive and straightforward tool for the harmonization and optimization of sampling efforts in similar projects (such as those proposed by Forest Europe). To this end, a limited number of variables concerning deadwood, climate, fauna, forest typology, and biotic and abiotic stress of trees have been combined into an indicator which reflects the overall health status and naturalness of forest ecosystems. Future studies and the collaboration between different research groups are desirable in order to validate the theorethical indicator proposed here and test its effectiveness and ease-of-use in projects focused on assessing the naturalness of forest ecosystems.

Relevance. The article discusses various options for assigning forest biogeocenoses to a certain dynamic position in the successional field of their development. The proposed method for dynamic assessment of the position of forest creates the possibility of assessing the total movement of tree volumes in the age generations of the age range of the forest biogeocenosis the possibility of analyzing the past changes in the age structure of the biogeocenosis and predicting its future development for a very distant future. Material and methods. The objects of research are native virgin spruce biogeocenoses of different ages in the northern taiga of the Kandalaksha forestry enterprise of the Nyamozersky forestry of the Murmansk region, the Severodvinsky forestry enterprise of the Arkhangelsk region; middle taiga spruce forests of the Vepssky forest reserve, Leningrad region. and the tract “Atleka” of the Andomsky forestry of the Vologda region; southern taiga spruce forests of the Kologrivsky forest reserve Kostroma region and the Central Forest Biosphere Reserve in the Tver region. The author studied the features of the structures of the age series of spruce forests, the most typical and productive types of growing conditions for spruce. Results and conclusions. All represented forest communities have different ratios of the number and volume of trees in the age generations of the age series and, as a consequence, different positions in the successional dynamics of development. Wood-destroying fungi of the biotrophic complex, as an endogenous structure of the forest community, actively participates in the dynamic processes of its development at all stages of its successional movement. The infection of trees by fungi of this complex increases from the last generations to the first to the maximum values in the oldest trees of the first generation. The dynamic position of the indigenous virgin forest community of different ages can be described with a wider range of assessments of the dynamics of its development in retrospect and perspective in terms of the ratio of quantitative and volumetric indicators of age generations of age series than in terms of the current structure of the age series. When assessing the successional stage of the forest community, it is necessary to take into account the quantity (volumes) and structure of the deadwood.

Earthworms are a significant group of soil macrofauna that provides a number of ecosystem functions in most terrestrial communities. Studies of the earthworm species composition and density of their populations were conducted in many regions of Russia, but the lumbricofauna of Novosibirsk region remained unexplored for a long time. The purpose of this investigation is to conduct a comparative analysis of the earthworm population of coniferous and small foliage forests in forest-steppe Ob region of the Novosibirsk area and identify the correlation of several species with the main physical and chemical properties of soil. The study was conducted in pine forests and birch-aspen forests. The main method of sampling was layer-by-layer excavation with manual destruction of soil samples and disassembling of forest deadwood. Some soil characteristics were also measured. There are data on the species composition and population density of earthworms for each habitat in results. Basing the ratio of earthworm living forms we created the classification of investigated habitats. We revealed that the diversity of epigeic and epi-endogeic earthworms in forests is largely provided by deadwood, and the main abiotic factor for worms is soil moisture. For the Asian subspecies Eisenia nordenskioldi nordenskioldi we performed the morphometric analysis in order to confirm characteristic polymorphism for it.

Deadwood plays a key role in the functioning and productivity of forest ecosystem and it is an important feature for the conservation of the biodiversity. In the past land-use and forest management have dramatically reduced the amount of deadwood in the European forests and only in the last decades the important role of the deadwood has been studied and recognised. At the political/administrative level the Ministerial Conference on the Protection of Forests in Europe (MCPFE) has recognized the importance of this forest component and the last year Italian Government has requested the regional forest laws and regulations to take into account deadwood and habitat/retention trees in the forest management. It is a current research priority to provide managers of European forests with a baseline of deadwood threshold retention value for decisions for their specific forest type and management target.

Earthworms are a significant group of soil macrofauna that provides a number of ecosystem functions in most terrestrial communities. Studies of the earthworm species composition and density of their populations were conducted in many regions of Russia, but the lumbricofauna of Novosibirsk region remained unexplored for a long time. The purpose of this investigation is to conduct a comparative analysis of the earthworm population of coniferous and small foliage forests in forest-steppe Ob region of the Novosibirsk area and identify the correlation of several species with the main physical and chemical properties of soil. The study was conducted in pine forests and birch-aspen forests. The main method of sampling was layer-by-layer excavation with manual destruction of soil samples and disassembling of forest deadwood. Some soil characteristics were also measured. There are data on the species composition and population density of earthworms for each habitat in results. Basing the ratio of earthworm living forms we created the classification of investigated habitats. We revealed that the diversity of epigeic and epi-endogeic earthworms in forests is largely provided by deadwood, and the main abiotic factor for worms is soil moisture. For the Asian subspecies Eisenia nordenskioldi nordenskioldi we performed the morphometric analysis in order to confirm characteristic polymorphism for it.

Abstract Retention approaches in forest management are today common in several North European countries, integrated into the clearcutting practice as a way to promote biodiversity and maintain ecosystem functions. Individual green trees and retention patches (tree groups) are retained at final harvest, and deadwood is left at site or created. Here, we review research on retention in Sweden, Finland, Norway, the Baltic States, and NW Russia, with special focus on biodiversity. Following the first publication in 1994, about 180 peer-reviewed articles have been published. We present results from a systematic search of the retention literature, separated into the following topics: buffer zones, retention patches, high stumps, other types of deadwood, European aspen Populus tremula, and cost-efficiency. Russian literature is synthesized separately since studies from this region have so far almost exclusively been published in the Russian language. Furthermore, we describe six ongoing large-scale, replicated experiments with varying retention levels, five in Finland and one in Sweden, and summarize their main results. Among main conclusions for practice from the literature and experiments are that retention patches as large as 0.5 ha and 10-m-wide buffers to watercourses are not enough to maintain pre-harvest species composition but survival of forest species is still larger than on conventional clearcuts. Deadwood on clearcuts may present important habitats to saproxylic species, including rare and red-listed ones and a prioritization of tree species per stand is recommended. We identify several important future research directions including switch of focus towards the landscape as well as the species population level. Surveys in parts of European Russia where retention has been unintentionally implemented already for a century would indicate possible future trajectories of biodiversity and their drivers in other regions of Northern Europe. A stronger link to ecological theory would help in study designs and in the formulation of predicted outcomes.

The extinction of species is a non‐random process, and understanding why some species are more likely to go extinct than others is critical for conservation efforts. Functional trait‐based approaches offer a promising tool to achieve this goal. In forests, deadwood‐dependent (saproxylic) beetles comprise a major part of threatened species, but analyses of their extinction risk have been hindered by the availability of suitable morphological traits.To better understand the mechanisms underlying extinction in insects, we investigated the relationships between morphological features and the extinction risk of saproxylic beetles. Specifically, we hypothesised that species darker in colour, with a larger and rounder body, a lower mobility, lower sensory perception and more robust mandibles are at higher risk.We first developed a protocol for morphological trait measurements and present a database of 37 traits for 1,157 European saproxylic beetle species. Based on 13 selected, independent traits characterising aspects of colour, body shape, locomotion, sensory perception and foraging, we used a proportional‐odds multiple linear mixed‐effects model to model the German Red List categories of 744 species as an ordinal index of extinction risk.Six out of 13 traits correlated significantly with extinction risk. Larger species as well as species with a broad and round body had a higher extinction risk than small, slim and flattened species. Species with short wings had a higher extinction risk than those with long wings. On the contrary, extinction risk increased with decreasing wing load and with higher mandibular aspect ratio (shorter and more robust mandibles).Our study provides new insights into how morphological traits, beyond the widely used body size, determine the extinction risk of saproxylic beetles. Moreover, our approach shows that the morphological characteristics of beetles can be comprehensively represented by a selection of 13 traits. We recommend them as a starting point for functional analyses in the rapidly growing field of ecological and conservation studies of deadwood. [ABSTRACT FROM AUTHOR]

In recent decades, the conservation of biodiversity has become one of the main areas under consideration in managing forests in an ecologically sustainable way. Forest management practices are primary drivers of diversity and may enhance or decrease forest biodiversity, according to the measures applied (thinning options). We have focused on three beech (Fagus sylvatica L.) forests across a latitudinal gradient in Italy, characterised by different structures resulting from dissimilar management. We tested the short-term effects of differently-based silvicultural intervention vs. stands where no silvicultural practices were applied on biodiversity indicators and related proxies: deadwood amounts, microhabitat density, floristic richness and life form abundance. In each study area, the occurrence of the above indicators and proxies was evaluated before and after the implementation of crop tree thinning (CTT) and thinning from below (LT) methods, comparing them with control areas where no interventions were performed. After two years, the management options resulted in different responses of the investigated parameters. The CTT increased deadwood amounts in comparison with the LT ones, while stumps increased significantly after the LT thinning. Microhabitats increased significantly where intervention was not undertaken. On the contrary, they remained unaltered after the LT treatments. CTT thinning created favourable conditions for the development of microhabitats and their proliferation in the long term. Two years after the application of the CTT thinning treatment, all forest stands demonstrated a significant increase in their floristic richness and herb layer cover. Significant differences were also found in both the frequency and cover of life forms in relation to silvicultural treatment. These findings provide a better understanding of short-term effects of silvicultural treatment useful for maintaining biodiversity in mountain beech forests.

Retention approaches in forest management are today common in several North European countries, integrated into the clearcutting practice as a way to promote biodiversity and maintain ecosystem functions. Individual green trees and retention patches (tree groups) are retained at final harvest, and deadwood is left at site or created. Here, we review research on retention in Sweden, Finland, Norway, the Baltic States, and NW Russia, with special focus on biodiversity. Following the first publication in 1994, about 180 peer-reviewed articles have been published. We present results from a systematic search of the retention literature, separated into the following topics: buffer zones, retention patches, high stumps, other types of deadwood, European aspen Populus tremula, and cost-efficiency. Russian literature is synthesized separately since studies from this region have so far almost exclusively been published in the Russian language. Furthermore, we describe six ongoing large-scale, replicated experiments with varying retention levels, five in Finland and one in Sweden, and summarize their main results. Among main conclusions for practice from the literature and experiments are that retention patches as large as 0.5 ha and 10-m-wide buffers to watercourses are not enough to maintain pre-harvest species composition but survival of forest species is still larger than on conventional clearcuts. Deadwood on clearcuts may present important habitats to saproxylic species, including rare and red-listed ones and a prioritization of tree species per stand is recommended. We identify several important future research directions including switch of focus towards the landscape as well as the species population level. Surveys in parts of European Russia where retention has been unintentionally implemented already for a century would indicate possible future trajectories of biodiversity and their drivers in other regions of Northern Europe. A stronger link to ecological theory would help in study designs and in the formulation of predicted outcomes. [ABSTRACT FROM AUTHOR]

One of the objectives of forest conservation is the set aside of unharvested areas. However, the fragmentation and lack of connectivity of protected areas make the integration of conservation measures in productive forests essential. Strategies to integrate conservation of saproxylic biodiversity in forest management have been developed, but often considering only specific aspects or remaining preliminary otherwise. As the impact of climate change and anthropogenic stresses increases, the development and the synthesis of this approach is crucial. We reviewed the key literature on forest management for biodiversity conservation, integrating forest science perspective to provide a practical management framework. Our goal is to present a management framework that could contribute to the effective preservation of forest insect biodiversity at the landscape scale, without high economic efforts, and addressing the conflicts that still jeopardize sustainable forest management. The results of our review support the creation of micro-reserves inside productive forests, to support large reserves in landscape conservation strategies. Micro-reserves increase the resilience of forest ecosystems to anthropogenic disturbances, through the development of a heterogeneous structure, maximizing microhabitat availability. Modeling forest management and harvest on local natural disturbance would extend the benefits of spatio-temporal heterogeneity in productive forests. Variable retention harvest systems, applied at the landscape scale, are a feasible and adaptable strategy to preserve and increase biodiversity, safeguarding structural legacies such as senescent trees and deadwood inside the productive matrix. The operational shift, from the stand to the forest landscape, is fundamental to extend the benefits of conservation measures. The Forest Biodiversity Artery, composed by several micro-reserves or îlots de senescence, connected by corridors of habitat trees and deadwood, constitutes a network that would deliver old-growth forests attributes to the productive matrix. This planning instrument would support forest connectivity, and socioeconomic constraints.

To describe the decay stage of coarse woody debris (CWD) a five decay-class system has been introduced and it is currently the most commonly applied. This system is based on visual, geometric and tactile features of the wood in the field; however, a detailed chemical characterization is often missing. Furthermore, the driving mechanisms (particularly substrate quality vs. environmental conditions) of deadwood decay are controversially discussed. Consequently, we investigated how typical major and minor chemical parameters of wood were correlated with the decay stage. The decomposition patterns of Norway spruce (Picea abies (L.) Karst) and European larch (Larix decidua Mill.) CWD of an Alpine setting were analyzed, and how the chemical and physical parameters were affected by the substrate and environmental conditions was checked. Two altitudinal sequences, having a different exposure (north- vs. south-facing sites), were sampled. We measured main biochemical compounds (lignin and cellulose), physical properties (density and water content), element concentrations (C, N, P, K, Ca, Mg, Fe, Mn), and the carbon isotopic signature (δ13C) of living trees and CWD at five decomposition stages (decay classes). Most investigated wood physico-chemical parameters such as wood density, water content, lignin and cellulose and even minor constituents (N, Ca, Mg, P, Fe, Mn) correlated well to the five decay-class system. Some important components, such as the carbon concentration and δ13C, did not vary with increasing decomposition. Our hypothesis that the different substrate should be traceable during CWD decay had to be rejected, although some statistically significant chemical differences between larch and spruce were measured in the living trees. The chosen tree species were probably not different enough to be chemically traceable in the CWD. Already in decay class 1, these differences were zeroed. The site conditions (expressed by the different altitudes and exposure) influenced only some of the investigated parameters, namely lignin, the δ13C isotopic ratio and nutrients such as P, Ca and K.

Deadwood is commonly known as an essential structural and functional element in forest communities. Thus it is important to understand its role and natural dynamics to improve the forest management strategies in both managed and unmanaged forests. This study analyzed both qualitative and quantitative characteristics of deadwoods asindicatorsof stand developmental stages in the unmanaged forests of Kheiroud in Mazandaran province, northern Iran. Three sample plots were established in the initial, optimal and decay stages, in which attributes such as diameter at breast height, height of all standing and dead trees as well as the decomposition stage of dead trees were recorded. Results showed differencesin the amount of deadwood per ha as well as in volume, type andsizeofthe coarse woody debris (CWD) in all the three stages. The frequency of snags was higher than logs in initial and decay stages, whereasthe proportion of logs was greater than snags in the optimal stage. Moreover, maximum and minimum volume ofthe CWD was observed in the initial and optimal stages. Amongst the species, beech returned the highest number and volume of deadwood in all three stages. In addition, the entire deadwood showed diameter

Deadwood and burnt wood could be a great reserve in the construction and related industries. Although the properties and performance of healthy pine wood are known, there is practically no information on deadwood and burnt wood. Therefore, to further establish sustainable ways of using such wood, the biological durability and strength depending on operating conditions should be studied. The authors of the study determined some strength parameters on samples of pine deadwood from the Kiev region of different indicative age of desiccation (from 1 to 3 years) and on samples pine burnt wood of Zhytomyr region after grassroots fires. As a result of studies we founded hyphae of fungi on microsections of central and transitional zones of sap wood. Most hyphae were accumulated in cells of medullary rays and sap tracheids. The scientists have also shown the experimental results of some physical and mechanical properties of pine deadwood compared to healthy wood strength parameters and normative values for building constructions. These results suggest changes in micro- and macrostructure of wood, and therefore evidence of negative effects of fire and high temperatures on the strength parameters of harvested timber. Thus, we have concluded that pine deadwood and burnt wood can be used as a building material. However, in the case of manufacturing structural elements that have to bear the load on static bending, it is better to use deadwood indicative age of desiccation of one year and burnt wood from butt part of the trunk. The results of experimental studies allows some recommendations on using of pine deadwood and burnt wood and receive regulatory resistance values for the design and calculation of wooden structures.

The cavities that develop in veteran trees represent a key microhabitat for forest biodiversity and especially for secondary cavity nesters that rely on this resource for shelter. Since the availability of deadwood, veteran and hollow trees is threatened by forest management, we explored the possibility of increasing the presence of these scarce resources. To increase the abundance of dead wood-microhabitats, 113 trees of the hybrid planes (Platanus x acerifolia) were converted into new living structures, the Habitat Trees (HT). To investigate the potential of this resource on the native avian fauna, six types of cavities were designed according to the size requirements of the target bird species. The temporal evolution of the cavities and their use by birds were then studied for eight years. The artificial cavities generally did not compromise growth and stability of the trees, and the majority remained alive. These hollows offered better thermal insulation compared to traditional nest-boxes and natural cavities. Their use increased during the first three years, reaching the 80%. In the following years, the use declined (probably because of the increase of rot and displacement of the lids).This highlights the need of management to maintain their suitability through time. Our results suggest a possible cost-effective alternative use of alien tree species, which should be included in management actions to compensate the shortage of hollow trees in managed forests.

Due to their significance for carbon sequestration and ecosystem management, information on decay dynamic of deadwood and its total decay time are of great importance to ecologists and forest managers. This study was conducted in Kheirud forest of Nowshahr to study the decay process of beech and hornbeam deadwood. Therefore, 72 beech and hornbeam dead trees were randomly selected. For each sample tree, information on species, diameter, decay classes and slope gradient were recorded. The cubic cut samples with 5 cm length, width and height in two perpendicular aspects were taken from each decaying stump. 30 and 16 years of decaying time were shown to be needed for 95% dry mass of dead beech and hornbeam logs, respectively. In addition, the required periods for passing through all stages of decay for beech and hornbeam dead trees were 27 and 21 years, respectively. A strong relation (R2=0.8) between density and decay stage of dead trees was indicated. Therefore, the density of dead wood was concluded to significantly decline with increased decay stage in both species.

The paper presents the results of a naturalness assessments, and also of the first inventory in two newly established forest reserves (Voievodeasa and Loben) from North of Romanian Eastern Carpathians. The evaluation of naturalness used the historical data concerning the wood extraction, as were recorded in the forest management plans. In selected stands of higher local naturalness was further conducted an inventory, following the methodology recommended by COST E4 and based on a systematic grid with circular plots of 500 m2 . Expected, the results showed that extraction impacted the structural characteristics of stands, esspecially the standing/downing trees volume. The comparison of structural data resulted from inventory (density and volume of both alive and dead standing trees and coarse woody debris) with local references - “virgin“ forests located about 40 km apart of the forest reserves under study - suggested a degree of correspondence between 20-40%. The same data, compared with published information available from mixed forest of beech-fir-spruce accross temperate zone of Europe confirmed the departure of selected stands from the references of naturalness and their distinct grouping, compared with other forest reserves.

Intensive management implies harvesting large, old trees, which reduces the old-growth attributes. This negatively affects biodiversity, especially saproxylic beetles. In managed temperate oak forests, rotation extension induced by increasing the diameter threshold of final harvest by about 10 cm compared to conventional practices (i.e. DBH around 70 cm) might mitigate this negative effect. Here, we used a gradient of the proportion of overmature trees (DBH ≥ 80 cm) among mature trees (DBH ≥ 70 cm) across plots of high oak French forests to test the potential effect of increasing diameter threshold on (1) structural features and (2) species composition and diversity of saproxylic beetles communities. We assessed deadwood and microhabitats availability (i.e. volume/density and diversity) and canopy openness in 81 1-ha plots across eleven French forests. Results highlighted that a larger proportion of overmature trees, for a given density of mature trees, had limited effects on structural features: only cavities density showed a significant positive response, with no cascading effect on cavicolous beetles. Moreover, the proportion of overmature trees had no significant effect on the composition of saproxylic beetles communities (and ecological variables altogether explained only 17% of the composition inertia). By contrast, mature tree density enhanced microhabitat density and indirectly increased the abundance of rare species. Thus, shifting DBH from 70 to 80 cm in high oak forest could have no or limited effect on saproxylic beetles conservation. Improved strategies might rather stem from combining (1) longer rotation extension and (2) less intensive management practices in extended rotation stands. [ABSTRACT FROM AUTHOR]

Comparison among methods for the assessment of deadwood volume in a former holm oak coppice. The paper aims to compare three methods for the assessment of deadwood volume, i.e., LIS (Line Intersect System), FAS (Fixed Area Sampling), and WM (Weighings Method). The control data are represented by the outputs of Xylometric measurement. The study was carried out in a former holm oak (Quercus ilex L.) coppice located in the nature park of Montioni in southern Tuscany. LIS and FAS overestimated significantly the quantity of deadwood (+12% and +50%, respectively). The error become higher as the minimum threshold sampling increases. The WM, besides the operational complexity of its application, led to more promising and precise results.

Chestnut (Castanea sativa MILL.) is one of the most important forest tree species in Europe, and it is considered a symbol of the natural vegetation in southern Europe. In Calabria (southern Italy) chestnut forest covers an area of approximately 87000 hectares, most of which (about 80%) managed as coppice. In this study a growth and yield table has been elaborated. Thurthermore, assortment type and quantity of deadwood have been evaluated according to age of coppice and forest fire prevention, respectively. The study site is located in the “Presila of Catanzaro” and the research was carried out in 15 plots; the age of the examined stands ranged from 2 to 50 years old. More than 30000 shoots per hectare were recorded in the first two years after coppicing. As opposed, about 2300 and 1000 shoots per hectare were observed 15 and 50 years after coppicing, respectively. The culmination of the mean annual increment of the forest standing volume (16 m3 ha-1 year-1) was highlighted 25 years after coppicing, while the current annual increment culmination (21 m3 ha-1 year-1) was observed at 15 years. Fifteen years after coppicing, most of the wood production was constituted by small dimension assortments. Twenty five years after coppicing small and large poles were the prevailing assortments while telegraph poles and timber beams increased after 50 years. The amount of deadwood in forest ranged between 11.9 and 68.7 m3 ha-1. The largest component was represented by standing dead shoots. The results show that coppice management can be adopted even if the main purpose of the chestnut stand is the production of large size assortments. In chestnut coppice, highly vulnerable to fire, the reduction of stand density with silvicultural practices (thinning and displacement) is the main way to promote the efficiency of forest and a higher strength and resiliency against forest fire.

Current National Forest Inventory highlight the further increase over the last two decades of coppice area under the position of mature standing crop or in the post-cultivation phase, both being developed throughout the original cultivation area. This pattern, mainly due to the unprofitable fuel wood harvesting, also involved holm oak and beech forests, some of the most diffused forest covers in Sardinia, along the Apennines and pre-Alps. The alternative management option to ageing (the pro-active way of coppice conversion into high forest) has been also practiced in the public domain, but on much smaller areas as compared with those undergoing post-cultivation phase. Aged coppices located into medium-good site classes showed a positive growth pattern resulting in a high, age-related, wood matter storage. At the meantime, regular mortality occurring since former rotation into the fully-stocked shoot populations, stocked up high deadwood amounts, this becoming an outstanding attribute of these types. Carbon storage is becoming one of the major tasks attributable to these systems within the post-cultivation phase. Purposes are here to: (i) estimate living woody and standing + lying deadwood mass densities; (ii) determine deadwood/living mass ratio; (iii) verify lying deadwood decay class; (iv) analyse diversity between two sites aged likewise but different as for geographical location and tree species. A holm oak coppice aged 55 in Sardinia and a beech coppice aged 57 in Tuscany were selected at the purpose. Both stands have been developing the post-cultivation phase since two-three times the traditional rotation and represent the maximum ages in this position. Living and standing dead woody dry mass density were determined in each site by specific allometric functions. Lying deadwood amount was assessed by a sampling design covering systematically the full test area. Three decay classes were determined according to Hunter (modified). The tree species, both shade-tolerant, showed a similar pattern as for time of wood formation and its transfer to deadwood, as well. The main difference consists only in their evergreen and deciduous habit. Auto-ecology seems to be the main driver of stand development as for carrying capacity, mortality rate and survival time into the overstocked stand structures. Leaf standing mass and leaf /thin branches ratio are two and three times higher in holm oak, consistently with its biology. Living woody mass density is made basically by stem and thick branches (88%) and varies from 322 Mg ha-1 (beech) to 225 Mg ha-1 (holm oak). Deadwood amount ranges from 28 to 25 Mg ha-1 i.e., from 9 to 11% of living woody mass. Standing deadwood is 70 to 73% of total amount. As for lying material, the intermediate decay class is prevailing: 74%, (beech) and 66% (holm oak). Both dynamics and age-related data suggest the active role these types may play in soil conservation, improvement of forest quality and carbon storage. The need of a consistent monitoring of the further post-cultivation progress is finally stressed to recognize driving forces acting, the onset of possible limiting factors and feedbacks.

Studies were undertaken to investigate the composition, structure and diversity of four different types of North Mongolian montane forest near the southern border of the taiga. These forest types, classified as willow-poplar, larch-birch, spruce-fir and Siberian-pine forests, were found to be significantly different with respect to the elevational gradient. In the study area, two fire regimes could be distinguished at lower and drier elevations, frequent surface fires resulted in less diverse forests comprising coniferous species, which in their adult form were found to be fire-resistant, burn-colonizing and light-demanding deciduous tree species. In contrast, the forests colonizing the moist, higher elevations and shaded slopes had a higher diversity of tree species with greater basal area, higher proportion of deadwood, and more regenerating trees; here the coniferous shade trees experienced infrequent but destructive treetop fires. Overall, our results showed that the four forest types differed in composition according to the tree species, diameter and height classes.

Although coarse woody debris (CWD) represents one of the major carbon pools in natural forest ecosystems, little information is available about its CWD carbon stocks. This study demonstrates the importance of proper estimation of carbon stocks in CWD, which accounts for the decay process of CWD, on an example of a natural mountainous spruce forest located in Central Europe. The study accounts for aboveground coarse woody debris including standing dead trees, lying deadwood, and naturally formed stumps. Basic mensurational information (diameter, height, decay class) about dead wood was collected in the field during the inventory of the forests of the nature reserve Babia hora. The data were used for the calculation of CWD timber volume. In the next step, CWD timber volume was converted to carbon stock using the carbon proportion of 50.1% and density values of decay classes derived from the information published elsewhere. The analysis revealed that when CWD timber volume was converted to carbon stocks using the basic wood density of fresh wood, C stocks were overestimated by 40% or more depending on the developmental stage and elevation. The results also revealed that as the elevation increases, CWD carbon stocks decrease and the differences between the developmental stages diminish.

Deadwood plays a key role in the functioning and productivity of forest ecosystem and it is an important feature for assessing and monitoring biodiversity. The third Ministerial Conference on the Protection of Forests in Europe (MCPFE) has recognized the importance of this forest component at the European level. More recently the national forest inventories have tried to harmonize the procedures including the quantitative and qualitative measurements of deadwood. This paper reports on the methods and results of Italian National Inventory of Forests and Forest Carbon Sinks (INFC) and compares the deadwood data with the results of other main European inventories and some local studies. In Italy the volume of deadwood (logs, snags and stumps) is on average 8.8 m3 ha-1.

The Forest Biodiversity Test phase Assessments (ForestBIOTA) project was carried out on 97 intensive monitoring plots located in 12 European countries. It aimed at the development and test implementation of monitoring methods for forest biodiversity across Europe. More specifically it covered the fields of (i) stand structure, (ii) forest deadwood, (iii) ground vegetation and (iv) epiphytic lichens. A forest type classification was applied in addition. The project took into account policy requirements stated by a number of political processes and institutions including the Convention on Biological Diversity, the Ministerial Conference for the Protection of Forests in Europe, the Environment for Europe Ministerial Conference and the European Union. Whereas detailed results of the assessments are presented in additional publications (Stofer et al. 2007, in prep, Meyer et al. 2007, submitted), this paper draws a number of general conclusions and gives recommendations for future development of forest biodiversity monitoring in Europe.

Necromass has been included within the indicators for the biodiversity and naturality assessment of the forest ecosystems and for the sustainable forest management. Taking into account this fact, the purpose of this work has been the quantification and qualification of the necromass in ageing coppices of the Gargano. To pursue this aim, three experimental areas, with 2500 m2 of surface each one, where the necessary reliefs have been effected, have been realized. The estimated deadwood volume is 34 m3 ha-1 and the stored carbon contents is 12.62 Mg C ha-1. The stand analysis pointed out that the quantity of present necromass is intrinsically linked whether to the culture abandonment or to the meteoric disturbances. Moreover the quantity and quality of necromass can be an indicator whether of the forest stability or of the stand evolutive trend in relatively recent times. The study of the quali-quantitative characteristics of the necromass in wood helps to understand the evolutive phases where a forest stand is in, and it can be, with other parameters, a guide to the planning and management. And today, just in the planning and management of the forest resources, we must give a higher importance to the presence of necromass in wood, not only in quantitative terms but also in qualitative ones.

The mycoflora was investigated under the conditions of climax spruce stands in the Krkonoše Mountains in relation to wood decomposition. The areas under observation have been affected more or less by air pollution since the eighties. The average mass of deadwood found on the plots is 124 m3 per ha - the mass of fallen trunks is about 32 m3 per ha, mean value from total average. About 128 species of macrofungi were identified that besides others included 43 species of wood-decaying fungi. Also 54 mycorrhizal species were identified. Among the mycorrhizal fungi about 10 species were dominant, such as Laccaria laccata (Scop.: Fr.) Cooke, Lactarius helvus Fr., Lactarius mitissimus Fr., Lactarius rufus (Scop.) Fr., Russula emetica (Schaeff.: Fr.) Pers. and Russula ochroleuca Pers. etc. Concerning the volume of decomposed wood on monitored plots in climax spruce stands, the prevalent wood-decaying fungi are brown rot fungi. The proportion of brown rot fungi in wood decomposition is 60-95% of deadwood mass on the plots of climax spruce stands. A dominant species is Fomitopsis pinicola (Sw.: Fr.) P. Karst. causing the brown rot. Concerning the group of white rot fungi, the most important is Stereum sanguinolentum (Alb. & Schw.: Fr.) Fr., participating by 17% in wood decomposition on plots damaged by deer.