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Competing Interests: This work was supported by the US National Institute of Allergy and Infectious Diseases (grant numbers R01 AI132606 and R01 AI153521) and the US Centers for Disease Control and Prevention (contract number 200–2016–91779), paid to YHG. The findings, conclusions, and views expressed are those of the authors and do not necessarily represent the official position of the Centers for Disease Control and Prevention. KIOR declares no competing interests.