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Elaphromyia yunnanensis Wang Elaphromyia yunnanensis Wang, 1990: 489. Type locality Lushui, Yunnan, China. Material examined: 1♂, INDIA: Arunachal Pradesh, Roing, Kebali, 10.ix.2014, Ramani, S. (UASB). Diagnosis (Figs 35, 36): Medium-sized (5.91 mm) fly; head as high as long, frons fulvous with three pairs of frontal setae (all black) and two pairs of orbital setae (posterior pair black), ocellar setae well developed, medial and lateral vertical setae, postocellar setae white, lanceolate; postocular setae black interspersed with white lanceolate setae. Face fulvous without any markings; scape, pedicel fulvous, first flagellomere shorter than face, arista short plumose. Scutum dark brown to black with golden yellow setulae except postpronotal lobe, notopleuron and beneath postsutural supra-alar seta. Anepisternum fuscous to dark brown with a single seta, anepimeron concolorous with anepisternum and with a single seta, katepisternum black with a fulvous anterior border and a single seta, anatergite and katatergite partly yellow and black. Chaetotaxy complete; 1 on postpronotal lobe, 1 presutural supra-alar, 1 anterior notopleural, 1 posterior notopleural, 1 postsutural supra-alar, 1 dorsocentral placed in line with postsutural supra-alar, 1 postalar, 1 intra-alar, 1 prescutellar acrostichal seta. Scutellum brown with two pairs of scutellar setae. All leg segments fulvous except hind femur with basal and medial black spots; forefemur with a single row of 6–7 ventral setae. Wing elongate (6.33 mm), parallel-sided and predominantly dark brown with numerous hyaline and subhyaline markings, costal and anal margins predominantly hyaline with faint brown markings, cells r 1 and r 2+3 with very small, closely packed hyaline spots, subhyaline markings appear to be transversely arranged, cell m with 4 prominent hyaline bands from the margin. Abdomen elongate-oval, all tergites dark brown to black with faint yellow markings laterally. Male genitalia (Figs 37–39): Epandrium broad, with no demarcation between lateral surstylus and epandrium, proctiger smaller than epandrium, apex of lateral surstylus pointed with numerous setae; epandrium and lateral surstyli round (posterior view) with apical extension of surstylus prominent; phallus elongate (4.54 mm) with a sclerotised rod like structure in glans of phallus, vesica as long as phallus. Distribution: China (Yunnan); India (Arunachal Pradesh: new record). Remarks: The specimen collected from India is comparatively larger than the type male and has transverse subhyaline markings on the wing and four elongate spots in cell dm, unlike Wang’s (1998) illustration, which depicts round, well separated spots and six prominent spots in cell m. Since only a single specimen is available and considering its overall appearance and dark abdomen, we provisionally regard the Indian specimen as a variant of Elaphromyia yunnanensis; however, further material might show it to be specifically distinct., Published as part of David, K. J., Hancock, D. L., Sachin, K., Ramya, R. S. & Ramani, S., 2021, Taxonomic notes on the genus Elaphromyia Bigot (Diptera: Tephritidae: Tephritinae: Pliomelaenini) in India, with description of a new species, pp. 251-262 in Zootaxa 5023 (2) on pages 259-261, DOI: 10.11646/zootaxa.5023.2.5, http://zenodo.org/record/5225884, {"references":["Wang, X. J. (1990) Six new species of Tephritini from Hengduan Mountains, China (Diptera: Tephritidae). Acta Zootaxonomica Sinica, 15 (4), 489 - 494.","Wang, X. J. (1998) The Fruit Flies (Diptera: Tephritidae) of the East Asian Region. Acta Zootaxonomica Sinica, Supplement 21, 1 - 338."]}

Elaphromyia Bigot Elaphromyia Bigot, 1859: 314. Type species Elaphromyia melas Bigot, 1859 (= Trypeta adatha Walker, 1849). Paralleloptera Bezzi, 1913: 154. Type species Paralleloptera pterocallaeformis Bezzi, 1913. Diagnosis: Small to medium-sized flies (2–5 mm), with elongate, parallel-sided, predominantly brown wing (Figs. 1, 6, 15–16) with a series of subhyaline spots/markings, a series of marginal spots and the dorsal setulae on vein R 1 without a gap posterior to the end of vein Sc. Epandrium bar-shaped in profile without demarcation between epandrium and lateral surstylus, proctiger smaller than epandrium, glans of phallus with acrophallus patterned and broadly bifid. Taeniae elongate, reaching more than three-quarters length of eversible membrane, aculeus with apex flattened in sagittal plane, two dark brown to black spermathecae with numerous projections., Published as part of David, K. J., Hancock, D. L., Sachin, K., Ramya, R. S. & Ramani, S., 2021, Taxonomic notes on the genus Elaphromyia Bigot (Diptera: Tephritidae: Tephritinae: Pliomelaenini) in India, with description of a new species, pp. 251-262 in Zootaxa 5023 (2) on page 252, DOI: 10.11646/zootaxa.5023.2.5, http://zenodo.org/record/5225884, {"references":["Bigot, J. M. F. (1859) Dipterorum aliquat nova genera. Revue et Magasin de Zoologie Pure et Appliquee, Paris, Series 2, 11, 305 - 315.","Bezzi, M. (1913) Indian trypaneids (fruit-flies) in the collection of the Indian Museum, Calcutta. Memoirs of the Indian Museum, 3, 53 - 175."]}

Elaphromyia juncta David, Hancock & Sachin, sp.n. urn:lsid:zoobank.org:act: B1C3991D-81E1-429C-AB2C-BFE4C1B12566 Description: Length of male (3.79 mm); female (4.27 mm). Head (Fig. 2): higher than long; frons fulvous with three pairs of frontal setae and two pairs of orbital setae; ocellar triangle dark fuscous with well-developed ocellar setae; medial vertical seta black; lateral vertical seta, paravertical seta and postocellar seta lanceolate and white; postocular setae black interspersed with white lanceolate setae. Face fulvous without any markings. Scape and pedicel fulvous, first flagellomere concolorous with frons, arista bare. Gena narrow with prominent genal seta, subvibrissal setae present. Thorax (Figs 3, 5): Scutum uniformly dark brown with three faint longitudinal lines. Chaetotaxy well developed: 1 postpronotal, 1 presutural supra-alar, 1 anterior notopleural, 1 posterior notopleural, 1 postsutural supra-alar, 1 dorsocentral placed in line with postsutural supra-alar, 1 postalar, 1 intra-alar, 1 prescutellar acrostichal seta. Scutellum brown with two pairs of scutellar setae. Anepisternum fulvous with two setae near phragma; anepimeron fulvous with two setae, katepisternum with a single seta; anatergite dark brown; katatergite and meron fulvous. Legs entirely fulvous without any black/dark markings; forefemur with a single row of 4–5 ventral setae and two rows of 6–7 stout dorsal setae. Wing (Fig. 6) elongate, parallel-sided (length, 3.79 mm in male; 4.02 mm in female), predominantly dark brown with a series of circular or elongate spots connected to each other or nearly connected and hyaline marginal spots. Cell sc with two marginal spots; cell r 1 with 4–5 hyaline spots along costal margin; cell r 2+3 with single hyaline spot near apex of vein R 2+3, cell m with three marginal hyaline spots; posterior subhyaline band extends from anal cell to vein Cu 1. Abdomen (Fig. 4): Fully fulvous without any dark markings on tergites in male but with a pair of faint black spots on tergite VI in female. Male genitalia: Epandrium broad with no demarcation between surstylus and epandrium (Fig. 7); apex of lateral surstylus not blunt (in lateral view), with well-developed dorsal lobe; proctiger short, much smaller than epandrium; medial surstylus shorter than lateral surstylus and with well-developed prensisetae (Fig. 8). Phallus elongate (1.57 mm); glans of phallus sclerotised with acrophallus patterned and broadly bifid apically (Fig. 9). Female genitalia: Oviscape orange-yellow, conical (1.18 mm) (Fig. 10); eversible membrane (0.87 mm) with elongate taeniae (three-quarters length of eversible membrane); spicules on distal end elongate-oval with blunt apex (Figs 11, 12); aculeus (0.78 mm) curved, flattened in sagittal plane in apical half, pointed with reduced preapical setae (Fig. 13); spermathecae (Fig. 14) dark brown to black, elongate-oval and covered with numerous projections. Type material: Holotype, 1♀, INDIA: Karnataka, Chikkamagaluru, Tarikere, Kemmangundi, K. R. hill station, 04.ii.2021, Ramya, R.S.; Paratype: 1♂, Karnataka, Chikkamagaluru, Tarikere, Kemmangundi, K. R. hill station, 04.ii.2021, Sachin, K. (NBAIR). Etymology: The specific epithet is an adjective derived from the Latin word junctus (=connected), denoting the connected hyaline spots in the wing. Distribution: Known only from western Karnataka in southern India. Remarks: This species was collected by sweep-netting over flowers of Wedelia sp. (Asteraceae) but that is unlikely to be its host plant. It is similar to E. pterocallaeformis by having the subhyaline spots arranged in distinct, subparallel transverse rows but can be differentiated by the presence of a fully fulvous abdomen without black spots in males and with only a single pair of spots (on tergite VI) in females, lateral surstylus without acutely pointed apex and an eversible membrane with elongate-oval spicules with blunt apices, whereas E. pterocallaeformis has two pairs of abdominal spots in both sexes, an acutely pointed lateral surstylus and an eversible membrane with apically pointed spicules. Elaphromyia juncta is similar to E. siva in having a fully yellow abdomen in males and only one pair of spots in females, but E. siva can be easily separated by the rounder, well separated subhyaline spots on the wing and semicircular-shaped spicules on the eversible membrane., Published as part of David, K. J., Hancock, D. L., Sachin, K., Ramya, R. S. & Ramani, S., 2021, Taxonomic notes on the genus Elaphromyia Bigot (Diptera: Tephritidae: Tephritinae: Pliomelaenini) in India, with description of a new species, pp. 251-262 in Zootaxa 5023 (2) on pages 253-256, DOI: 10.11646/zootaxa.5023.2.5, http://zenodo.org/record/5225884

Elaphromyia siva Frey Elaphromyia siva Frey, 1917: 19. Type locality: Anuradhapura, North-Central Sri Lanka. Elaphromyia pterocallaeformis: David & Ramani (2011, fig. 104) [misidentification]. Material examined: 1♂, INDIA: Karnataka: Bangalore, Attur, 07. i.2013, David, K.J.; 1♀, Bangalore, Hebbal, 07.ii.2013, David, K.J.; 1♀, INDIA: Karnataka, Bangalore, Attur, 13.i.2014, David, K.J.; 1♀, Bangalore, Attur, 15.xi.2017, Prabhu, G.; 1♀, Bangalore, Attur, 31.x.2020, Sachin, K. (NBAIR). 1♂, INDIA: Karnataka, West Chikkaballapur, 03.viii.1976, C.A., Viraktamath. (UASB). Diagnosis (Figs 15, 16): Small species (male- 2.45 mm; female- 3.02 mm). Head higher than long; frons with three pairs of frontal setae (posterior frontal seta white) and two pairs of orbital setae (posterior orbital seta white); ocellar triangle dark fuscous with well-developed ocellar setae; medial vertical seta black; lateral vertical seta, paravertical seta and postocellar seta stubby white; postocular setae black interspersed with white stubby setae. Face fulvous with deep concavity. Scape and pedicel fulvous, first flagellomere concolorous with frons, arista short pilose. Gena narrow, with genal seta. Scutum uniformly dark brown with three faint longitudinal lines, chaetotaxy well developed: 1 on postpronotal lobe, 1 presutural supra-alar, 1 anterior notopleural, 1 posterior notopleural, 1 postsutural supra-alar, 1 dorsocentral placed in line with postsutural supra-alar, 1 postalar, 1 intra-alar, 1 prescutellar acrostichal seta. Scutellum brown with two pairs of scutellar setae. Anepisternum fulvous with two or three setae near phragma; anepimeron fulvous with two setae, katepisternum with a single seta; anatergite dark brown; katatergite and meron fulvous. All segments of leg fulvous without any black/dark markings; forefemur with a single row of 4–5 ventral setae. Wing (2.74 mm in male; 3.04 mm in female) shorter than in E. juncta, parallel-sided, predominantly dark brown with a series of rounded/circular subhyaline markings not arranged in distinct subparallel transverse rows and hyaline marginal spots. Cell sc with a single marginal spot; cell r 1 with 3–4 hyaline spots along the costal margin; cell r 2+3 with a single hyaline spot near the apex of vein R 2+3, cell m with three marginal hyaline spots; posterior subhyaline band extends from anal cell to vein Cu 1. Abdomen fully fulvous without any markings on tergites in male; but with a pair of faint black spots on tergite VI in female. Male genitalia: Epandrium broad with no demarcation between surstylus and epandrium (Fig. 17); apex of lateral surstylus blunt (in profile view); proctiger short, much smaller than epandrium; medial surstylus shorter than lateral surstylus and with well-developed prensisetae (Fig. 18). Phallus (1.00 mm) relatively shorter than in E. juncta; glans of phallus sclerotised with acrophallus patterned and broadly bifid apically (Fig. 19). Female genitalia: Oviscape orange-yellow (Fig. 20), conical (0.79 mm); eversible membrane (0.66 mm) with elongate taeniae (three-quarters length of eversible membrane); spicules on distal end semicircular (Figs 21, 22); aculeus (0.56 mm) curved, flattened in sagittal plane, pointed with reduced preapical setae (Fig. 23); spermathecae dark brown to black, elongate-oval with a prominent neck and covered with numerous projections (Fig. 24). Distribution: Sri Lanka (Type locality) and southern India (new record). Remarks: This species is much smaller than the other two species recorded from India (E. juncta and E. pterocalleformis) and is the smallest species known. Apart from the difference in size, it can be differentiated from the other two species by the presence of rounder subhyaline spots that are not aligned into subparallel transverse rows and the presence of a single spot in the subcostal cell. It can be differentiated further from E. pterocalleformis in lacking acutely pointed lateral surstylus and by the elongate-oval spicules on the distal end of the eversible membrane and abdominal markings. Described originally from Sri Lanka, this species is recorded for the first time from India, having been previously misidentified as E. pterocallaeformis by David & Ramani (2011: fig. 104). The identity of the Indian specimens has been confirmed by comparison with photographs of the Holotype female from Sri Lanka., Published as part of David, K. J., Hancock, D. L., Sachin, K., Ramya, R. S. & Ramani, S., 2021, Taxonomic notes on the genus Elaphromyia Bigot (Diptera: Tephritidae: Tephritinae: Pliomelaenini) in India, with description of a new species, pp. 251-262 in Zootaxa 5023 (2) on pages 256-258, DOI: 10.11646/zootaxa.5023.2.5, http://zenodo.org/record/5225884, {"references":["Frey, R. (1917) Ein Beitrag zur Kenntnis der Dipteren fauna Ceylon. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar (A), 59 (20), 1 - 36.","David, K. J. & Ramani, S. (2011) An illustrated key to fruit flies (Diptera: Tephritidae) from Peninsular India and the Andaman and Nicobar Islands. Zootaxa, 3021 (1), 1 - 31. https: // doi. org / 10.11646 / zootaxa. 3231.1.4"]}

Bactrocera (Bactrocera) apiconigroscutella Drew Bactrocera (Bactrocera) apiconigroscutella Drew (in Drew & Raghu, 2002: 333; Drew & Romig, 2013). Material examined: 8♂♂, INDIA, Maharashtra, Konkan Region, H. R. Sawai. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5B). Lateral surstylus shorter than epandrium, posterior lobe of surstylus as long as anterior lobe; ventral rim of lateral surstylus 3��4 times longer than dorsal rim (Fig. 3B). Proctiger quadrate and smaller than epandrium (Fig. 3B). Medial surstylus with a pair of thick prensisetae. Phallus 2.0�� 2.2 mm long, excluding glans (0.4 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8B)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 109, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Dacus (Mellesis) insulosus Drew & Hancock Dacus (Callantra) insulosus Drew & Hancock, in Drew et al., 1998: 599; David & Ramani, 2011:4. Dacus (Mellesis) insulosus (in Drew & Romig, 2013: 387). Material examined: 1♀, INDIA, Andaman and Nicobar Islands, Mount Harriet, 25. viii. 1993, Ranganath, H. R. (UASB). Diagnosis: Female with dorsoventrally flattened, conical oviscape (1.87 mm); eversible membrane 2.11 mm long, spicules on distal end of eversible membrane with 8̄10 sharp projections, the medial one 5̄6 times higher than basal ones (Fig. 12L); aculeus (1.72 mm) broad with three pairs of preapical setae (Fig. 14L).

Tribe Dacini Diagnosis. Scutum black/brown/reddish-brown with or without yellow medial and lateral postsutural vittae; reduced head and thoracic chaetotaxy; face with a medial transverse ridge (protuberance in profile view); wing with cell bm deeper/broader than bcu; extension of cell bcu longer than bcu; males usually with pecten; ceromata present; proctiger hyaline, smaller than epandrium; glans of phallus with well developed acrophallus made of single semitubular lobe; basal pouch like structure prominent; subapical lobe hammer shaped and basal lobe present; aculeus dorsoventrally flattened; well developed taeniae; base of eversible membrane with conical spicules; two convoluted and mulberry shaped spermathecae., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 104, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635

Dacus (Mellesis) discophorus (Hering) Callantra discophora Hering, 1956: 64. Dacus (Callantra) discophorus (in Drew et al., 1998: 586; Norrbom et al., 1999: 130; David & Ramani, 2011: 4). Dacus (Mellesis) discophorus (Hering) (in Drew & Romig, 2013: 381). Material examined: 3♂, INDIA, Karnataka, Chettalli, 26.viii. 2012, Ranganath, H. R., 2♂, INDIA, Karnataka, Bangalore, 14.x.2013, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Attur Farm, 16.ix.2013, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Attur Farm, 02.ix.2013, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Attur Farm, 16.vi.2014, David, K. J. Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7F). Lateral surstylus shorter than epandrium, apex of posterior lobe of surstylus lanceolate, 2��3 times longer than anterior lobe (Fig. 4R). Proctiger triangular and smaller than epandrium. Medial surstylus longer than lateral surstylus, tapering towards apex with a pair of prensisetae (Fig. 7F). Phallus 2.4 mm long excluding glans (0.38 mm); 0.75 of glans sclerotised with (hexagonal) patterned praeputium; subapical lobe and basal lobe present (Fig. 10P)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 125-126, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","David, K. J. & Ramani, S. (2011) An illustrated key to fruit flies (Diptera: Tephritidae) from Peninsular India and the Andaman and Nicobar Islands. Zootaxa, 3021, 1 - 31.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Bactrocera) limbifera (Bezzi) Chaetodacus ferrugineus var. limbiferus Bezzi, 1919: 424. Strumeta pedestris var.limbiferus Perkins, 1938a: 126. Dacus (Strumeta) limbiferus (in Hardy & Adachi, 1954: 172). Dacus (Strumeta) limbifer (in Hardy, 1974: 35). Bactrocera (Bactrocera) limbifera (in Norrbom et al., 1999: 92). Material examined: 3♂♂, INDIA, Andaman and Nicobar Islands, Middle Andamans, Kadamtala, N 11 0 50.295’E 92 0 46.415’, 10.iii.2012, David, K. J., 3♂♂, INDIA, Andaman and Nicobar Islands, Sippighat, 08.iii.2012, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5K). Lateral surstylus shorter than epandrium; posterior lobe of surstylus broad, curved back, not longer than anterior lobe. Proctiger quadrate and smaller than epandrium (Fig. 8K). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae (Fig. 3K). Phallus 2.5 mm long, excluding glans (0.48 mm); 0.70 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8K). ‘ Bactrocera (Bactrocera) melastomatos Drew & Hancock Bactrocera (Bactrocera) melastomatos Drew & Hancock, 1994: 38. Material examined: 1♂, INDIA, Andaman Inslands, H. R. Ranganath (UASB) Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5L). Lateral surstylus shorter than epandrium; posterior lobe of surstylus blunt, not longer than anterior lobe. Proctiger quadrate and smaller than epandrium (Fig. 3L). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 3.2 mm long excluding glans (0.24 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8L)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 120-121, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, 2 (Supplement), iii + 68 pp."]}

Bactrocera (Tetradacus) minax (Enderlein) Polistomimetes minax Enderlein, 1920: 358. Lectotype male in BMNH. Mellesis citri Chen, 1940: 133; White & Wang, 1992: 276 (synonym). Dacus (Polistomimetes) minax (in Drew, 1979: 76). Bactrocera (Polistomimetes) minax (in Drew, 1989: 13). Bactrocera (Tetradacus) minax (in White & Wang, 1992: 276, Lectotype designation: Norrbom et al., 1998: 101). Material examined: 5♂♂, 5♀♀, INDIA, West Bengal, Kalimpong, 8.xi.2016, Rakseh Pashi Diagnosis: Male with epandrium and surstyli inverted U-shaped (posterior view) (Fig. 7A). Lateral surstylus much reduced than epandrium; apex of lateral surstylus not differentiated into anterior or posterior lobes. Proctiger quadrate and smaller than epandrium (Fig. 4M). Phallus elongate (6.17 mm long) excluding glans (0.5 mm); threefourth of glans sclerotised with fully patterned praeputium; subapical lobe and basal lobe present (Fig. 10K). Female with brown, bottle shaped oviscape (1.20 mm); elongate eversible membrane (3.35 mm) with spiclues on distal end conical (Fig. 12H); aculeus (3.28 mm) with acute apex (Fig. 14H). Two black, convoluted mulberry shaped spermathecae., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 123, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["White, I. M. & Elson-Harris, M. M. (1992) Fruit flies of economic significance: their identification and bionomics. CAB International, Wallingford, 601 pp.","Drew, R. A. I. (1989) The tropical fruit flies (Diptera: Tephrititidae: Dacinae) of the Australasian and Oceanic regions. Memoirs of the Queensland Museum, 26, 1 ‾ 521."]}

Bactrocera (Bactrocera) digressa Radhakrishnan Bactrocera (Bactrocera) digressa Radhakrishnan, 1999: 1. Bactrocera (Daculus) yercaudiae Drew (in Drew & Raghu, 2002) 2002: 346. (synonymised by David & Ramani, 2011). Bactrocera (Daculus) digressa Radhakrishnan (in Drew & Romig, 2013: 199). Bactrocera (Bactrocera) digressa Radhakrishnan (in Hancock, 2015: 42). Material examined: 5♀♀, INDIA, Karnataka, Gowribidanur, Kolar, 23.vii. 2007, Naveen Kumar, V., 1♂, INDIA, Karnataka, Bangalore, Attur farm, 18.ii.2013, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Hebbal, 13.ii.2013, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Hebbal, 30.xi.2012, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5H), Lateral surstylus shorter than epandrium; posterior lobe of surstylus blunt (in profile view), curved in, as long as anterior lobe (Fig. 3H). Proctiger membranous, quadrate and smaller than epandrium (Fig. 3H). Medial surstylus, longer than lateral surstylus with a pair of thick prensisetae (Fig. 5H). Phallus 3.5 mm long, excluding glans (0.4 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8H). Female with reddishbrown, dorsoventrally flattened oviscape (1.05 mm); eversible membrane (1.06 mm) with spicules on distal end having 1��3 sharp projections (Fig. 11F); aculeus short (0.9 mm) with bifid apex and four pairs of preapical setae (Fig. 13F); two black convoluted berry shaped spermathecae (Fig. 15E)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 118, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["David, K. J. & Ramani, S. (2011) An illustrated key to fruit flies (Diptera: Tephritidae) from Peninsular India and the Andaman and Nicobar Islands. Zootaxa, 3021, 1 - 31.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Parazeugodacus) conica David and Ramani, sp. nov. Description: Medium sized (5.9 mm), black species. Frons fulvous with two frontal setae and single orbital seta, ocellar triangle black, ocellus fuscous, outer vertical seta and medial vertical seta present; occiput fuscous with a row of thin, black postocular setae; face fulvous with elongate, black spot in each antennal furrow, gena fulvous with subocular spot and seta. Scape, pedicel fuscous; first flagellomere dark fuscous; combined length of pedicel and first flagellomere as long as the vertical length of face (Fig. 1A). Scutum (2.07 mm long; 1.67 mm wide) predominantly black without lateral and medial postsutural vitta. Yellow markings as follows; postpronotal lobe, notopleuron, narrow anepisternal stripe (as broad as notopleuron, not continued as a small spot on katepisternum), 0.6 of katatergite and 0.9 of anatergite. Scutellum yellow with a broad, medial black patch and two pairs of scutellar setae; mediotergite and subscutellum black. Chaetotaxy; 2 scapular setae, 1 anterior notopleural seta, 1 notopleural seta, 1 anepisternal seta, 1 postsutural supra-alar seta, 1 post-alar seta, 1 intra-alar seta, 1 prescutellar acrostichal seta and 1 scutellar seta. Legs; coxa, trochanter black, all femora with apical ring like fuscous markings, tibia fuscous, tarsal segments fulvous (Fig. 1B,C). Wing (4.20 mm) predominantly hyaline without costal band, pterostigma dark brown, base of cell br hyaline without microtrichia, extension of cell bcu slightly longer than bcu, anal streak lacking (Fig. 1E). Abdomen (2.07 mm long; 2.09 mm wide) oval, wholly black (dark brown in Fig. 1D after processing in KOH) except two hyaline spots on tergite II (Fig. 1D). Female genitalia: Oviscape black (0.7 mm); spicules on distal end of eversible membrane (0.96 mm) conical (Fig. 1G), Acuelus (Fig. 1F) needle shaped (0.9 mm), length to width ratio, 5.3, aculeus tip acute without distinct preapical setae (Fig. 1G); black, convoluted, mulberry shaped spermathecae (Fig. 1I). Material examined: Holotype 1♀, INDIA, Karnataka, Chikmagalur, Mudigere, 03.vii.2012, David, K. J. (deposited at NBAIR) Etymology: The species is named for its conical-shaped spicules on distal end of eversible membrane. Remarks: Bactrocera conica is similar to B. bipustulata (Bezzi) in possessing hyaline wing without costal band, scutellum with a black medial patch; but it can be differentiated by the presence of separate black spots on the face (Fig. 1A), scutum without lateral postsutural vitta (Fig. 1B), katepisternum without yellow spot (Fig. 1C), femur not predominantly fuscous, abdomen fully black and conical shaped spicules on distal end of eversible membrane (Fig 1H). In contrast, B. bipustulata possess entirely black face (Fig. 1J), scutum with lateral postsutural vitta extending upto postsutural supra-alar seta (Fig. 1K), femur predominantly fuscous, katepisternum with yellow spot (Fig. 1L), abdomen not wholly black (Fig. 1M) and broad spicules with 3-5 projections (Fig. 1P) on distal end of eversible membrane. Since males were not available for the study, it is retained in the genus Bactrocera., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 105, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635

Bactrocera (Bactrocera) prabhui David, sp. nov. Description: Medium sized (6.6̄ 6.9 mm), black species. Frons fulvous with two frontal setae and single orbital seta, ocellar triangle black, ocellus fuscous, outer vertical seta and medial vertical seta present; occiput fuscous with a row of thin, black postocular setae; face fulvous with elongate, black spot in each antennal furrow and a medial longitudinal black band, gena fulvous with subocular spot and seta. Scape, pedicel fuscous; first flagellomere dark fuscous; combined length of pedicel and first flagellomere as long as the vertical length of face (Fig. 2A). Scutum (2.5̄ 2.9 mm long; 1.9̄ 2.2 mm wide) black with narrow, yellow lateral postsutural vitta tapering to end before intra-alar seta. Yellow markings as follows; postpronotal lobe, notopleuron, anepisternal stripe (as broad as notopleuron, continued as a small spot on katepisternum), 0.6 of katatergite and 0.9 of anatergite. Scutellum yellow with narrow, black basal band and a pair of scutellar setae, mediotergite and subscutellum black. Chaetotaxy; 2 scapular setae, 1 anterior notopleural seta, 1 notopleural seta; 1 anepisternal seta, 1 post-alar seta, 1 intra-alar seta, 1 prescutellar acrostichal seta, 1 scutellar seta. Legs; coxa, trochanter black, all femora with fuscous markings (0.9 of fore femur 0.25 of midfemur and hindfemur), tibia fuscous, tarsal segments fulvous (Fig. 2B,C). Wing (5.2̄ 5.9 mm) predominantly hyaline with narrow costal band from cell sc to wing apex slightly overlapping vein R 2+3, expanded slightly beyond vein R 2+3, base of cell br fully covered with microtrichia, extension of cell bcu longer than bcu and well developed anal streak (Fig. 2E). Abdomen (2.6̄ 2.8 mm long; 2.9 mm wide), oval, predominantly black except fulvous band on tergites I ⅈ broad, medial longitudinal, black band from tergites IIĪV present; lateral regions of tergites IIĪV predominantly black (Fig. 2D). Female genitalia: Oviscape black (1.39 mm); spicules on distal end of eversible membrane (2 mm) conical with three to four pointed projections (Fig. 2J). Aculeus (Fig. 2F) elongate (1.24 mm) with length to width ratio, 6.2; aculeus tip mucronate with two pairs of weakly developed preapical setae (Fig. 2G, H); two black, convoluted, mulberry shaped spermathecae (Fig. 2I). Material examined: Holotype ♀, INDIA, Karnataka, Kodagu, Somwarpet, 24-26.vi.2017, Prabhu, G. (NBAIR). Paratype 1♀, same data as of Holotype (NBAIR), Paratype 1♀, same data as of Holotype (UASB). Etymology: The species name is after its collector, Prabhu, G. Remarks: Bactrocera prabhui is similar to B. caryeae and B. furcata in having a predominantly black scutum with narrow lateral postsutural vitta, predominantly black abdomen, elongate-oval black facial spots and markings on all femora; but can be clearly differentiated from the former by the femoral markings (Fig. 2C) (extensive markings covering entire femur), costal band overlapping vein R 2+3 (Fig. 2E), mucronate aculeus tip (acute in B. caryeae), two pairs of weakly developed preapial setae in aculeus (Fig. 2H) (four pairs in B.caryeae), conical spicules of eversible membrane (Fig. 2J) (broad spicule with several projections of equal size in B.caryeae) and dense aggregation of microtrichia in the base of cell br and along vein Cu 2. B. prabhui can be distinguished from B. furcata by mucronate aculeus tip (bifid in B. furcata), conical spiclues of eversible membrane with 3-4 projections (single cone like projections in B. furcata) and costal band overlapping vein R 2+3. It is similar to B. fastigata by lacking postsutural supra-alar seta, but differs with respect to narrow anepisternal stripe (broad stripe in B. fastigata reaching anterior notopleural seta), parallel sided narrow postsutural vitta of uniform width (lateral vitta tapering posteriorly in B. prabhui), costal band overlapping vein R 2+3, broad anal streak, densely microtrichose base of cell br, extensive fuscous markings on femora and extensive black markings on all tergites.

Bactrocera (Bactrocera) carambolae Drew & Hancock Bactrocera (Bactrocera) carambolae Drew & Hancock, 1994: 11. Material examined: 1♂, INDIA, Andaman and Nicobar Islands, Andaman Islands, Chouldari, Bloomsdale, 07.iii.2012, David, K. J., 1♂, same data except date 08.iii.2012. (NBAIR); 1♂, INDIA, Andaman and Nicobar Islands, Mt. Harriet, S. Andamans, 22. xi. 1993, H. R. Ranganath, reared from Areca triandra Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5D). Lateral surstylus shorter than epandrium; posterior lobe of lateral surstylus pointed, not longer than anterior lobe (Fig. 3D). Medial surstylus with a pair of thick prensisetae. Phallus 2.3 mm long excluding glans (0. 34 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8D)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 112-113, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, 2 (Supplement), iii + 68 pp."]}

Zeugodacus (Zeugodacus) cucurbitae (Coquillett) Dacus cucurbitae Coquillett, 1899: 129. Bactrocera cucurbitae Bezzi, 1913: 96. Chaetodacus cucurbitae Bezzi, 1916: 109. Strumeta cucurbitae Perkins, 1938: 127. Dacus (Strumeta) cucurbitae Hardy & Adachi, 1954: 164. Dacus (Zeugodacus) cucurbitae Drew, 1973: 23. Bactrocera (Zeugodacus) cucurbitae (in Drew, 1989: 212; Drew & Romig, 2013: 279). Zeugodacus (Zeugodacus) cucurbitae (in Doorenweerd et al., 2018 -45). Material examined: 4♂♂, INDIA, Odisha, Bhubaneshwar, OUAT Campus, 05.ii.2016, Sunil Joshi. 1♂, INDIA, Karnataka, Bangalore, Attur farm, 13.xi.2012, David, K. J.; 1♀, INDIA, Karnataka, Bangalore, Attur farm, 10.xi.2012, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 4E). Lateral surstylus longer than epandrium; posterior lobe of surstylus pointed, 10��12 times longer than anterior lobe. Proctiger triangular and smaller than epandrium (Fig. 6I). Medial surstylus, longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.10 mm long excluding glans (0.41 mm); 0.75 of glans sclerotised with patterned praeputium; subapical lobe and basal lobe present (Fig. 10C). Female with dorsoventrally flattened, brown oviscape (1.0 mm); eversible membrane (1.5 mm) with spicules on distal end possessing 5��6 stout, pointed projections (medial one higher than lateral projections) (Fig. 12B). Aculeus 1.3 mm long, tip pointed with 4 pairs of preapical setae (Fig. 14B), apical ones 10��12 times longer than basal ones, two black, tightly coiled mulberry shaped spermathecae (Fig. 15R)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 129-130, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. (1989) The tropical fruit flies (Diptera: Tephrititidae: Dacinae) of the Australasian and Oceanic regions. Memoirs of the Queensland Museum, 26, 1 ‾ 521.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Zeugodacus) havelockiae (Drew & Romig) Bactrocera (Zeugodacus) havelockiae Drew & Romig, 2013: 297. Zeugodacus (Zeugodacus) havelockiae (Drew & Romig) (in Doorenweerd et al., 2018: 46). Material examined: 20♂♂, INDIA, Andaman and Nicobar Islands, Kadamtala, 09.iii.2012, David, K. J. (NBAIR), 1♀, INDIA, Andaman and Nicobar Islands, Mt. Harriet, 11.vii. 1993, H.R. Ranganath, 1♀, INDIA, Andaman and Nicobar Islands, 14.xi.1996, H. R. Ranganath (UASB). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6L). Lateral surstylus longer than epandrium; posterior lobe of surstylus 12̄14 times longer than anterior lobe. Proctiger triangular and smaller than epandrium. Medial surstylus as long as lateral surstylus (Fig. 4H). Phallus elongate (2.73 mm), excluding glans (0.46 mm); glans sclerotised with patterned praeputium, trumpet shaped subapical lobe and short basal lobe (Fig. 10F). Female with red-brown, dorsoventrally flattened oviscape (1.5 mm), eversible membrane (2.0 mm long), with spicules on distal end possessing 5̄6 broad lanceolate projections (Fig. 12D). Aculeus (1.73 mm) with acute apex and 4 pairs of preapical setae (Fig. 14D); two black, highly convoluted, mulberry shaped spermathecae (Fig. 15T)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 130, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Genus Bactrocera Macquart Bactrocera Macquart, 1835; 452. Type species Bactrocera longicornis Macquart, 1835, by monotypy. Diagnosis. Abdominal tergites free; abdomen generally oval, rarely petiolate; scutellum triangular. Epandrium round to oval (posterior view), posterior lobe of surstylus as long as or shorter than anterior lobe; upper rim of lateral surstylus 0.5 of lower rim except B. rubigina and B. paraosbeckiae. Glans of phallus with well developed acrophallus and unpatterned praeputium (apomorphy) except subgenus Tetradacus, dorsal sclerite of glans with hexagonal scultpuring (visible clearly under 100x objective). Aculeus usually with four pairs of preapical setae, rarely two/not traceable in few species. Spicules on distal end of eversible membrane usually broad with several, small spine like projections, rarely with single or few conical projections., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 104, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635

Zeugodacus (Parasinodacus) binoyi Drew Bactrocera (Sinodacus) binoyi Drew (in Drew & Raghu, 2002: 347). Bactrocera (Parasinodacus) binoyi Drew (in Drew & Romig, 2013: 226). Zeugodacus (Parasinodacus) binoyi Drew (in Doorenweerd et al., 2018: 44). Material examined: 1♂, INDIA, Kerala, Nelliyampathy, 1110 m, 10 0 32���N 77 0 42���E, 21.v.2010, Sudha, M. Cue lure (UASB). Diagnosis: Male with epandrium and surstyli bulb-shaped (posterior view) (Fig. 6D), with hyaline, setose, triangular proctiger. Posterior lobe of lateral surstylus broad, 7��8 times longer than anterior lobe, both lobes curved in (Fig. 3X). Medial susrtylus as long as lateral surstylus with well developed prensisetae. Phallus elongate, glans of phallus sclerotised with well developed, patterned (elongate tubular/conical pattern) praeputium; basal lobe and subapical lobe present (Fig. 9L)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 128, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Dacus (Mellesis) ramanii Drew & Hancock Dacus (Callantra) ramanii Drew & Hancock (in Drew et al., 1998: 623; David & Ramani, 2011: 4). Material examined: 2♂, INDIA, Karnataka, Bangalore, 0 4. ii. 2014, David, K. J.; 2♂, INDIA, Karnataka, Bangalore, 11. xi. 2013, David, K. J.; 2♂, INDIA, Karnataka, Bangalore, 16. xii. 2013, David, K. J.; 2♂, INDIA, Karnataka, Bangalore, 0 2. xii. 2013, David, K. J.; 2♂, INDIA, Karnataka, Bangalore, 15. iv. 2014, David, K. J.; 2♂, INDIA, Karnataka, Bangalore, 19. v. 2014, Prabhu, G.; 1♂, INDIA, Karnataka, Bangalore, 15. i. 2013, David, K. J.; 1♂, INDIA, Karnataka, Bangalore, 0 4. xi. 2013, David, K. J.; 1♂, INDIA, Karnataka, Bangalore, 30. vi. 2014, David, K. J.; 1♂, INDIA, Karnataka, Bangalore, 10. ii. 2014, David, K. J; 1♂, INDIA, Karnataka, Chikkaballapur, 27. iii. 2013, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7G). Lateral surstylus broad; posterior lobe of surstylus lanceolate, 3.5 times longer than anterior lobe, anterior lobe pointed (Fig. 4S). Medial surstylus as long as lateral surstylus with a pair of well developed prensisetae. Proctiger triangular not larger than epandrium. Phallus 2.12 mm long excluding glans (0.4 mm), 0.75 of glans sclerotised with well developed fully patterned (polygonal pattern) praeputium, subapical and basal lobe present (Fig. 10Q)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 126, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["David, K. J. & Ramani, S. (2011) An illustrated key to fruit flies (Diptera: Tephritidae) from Peninsular India and the Andaman and Nicobar Islands. Zootaxa, 3021, 1 - 31."]}

Zeugodacus (Zeugodacus) tau (Walker) Dasyneura tau Walker, 1849: 1074. Dacus hageni de Meijere, 1911: 375. Dacus caudatus var. nubilus Hendel, 1912 (synonymised by White & Wang, 1992: 278) Dacus nubilus ssp. femoralis Hendel, 1934:11. Dacus (Zeugodacus) tau Hardy, 1977: 60. Bactrocera (Zeugodacus) tau (in Wang, 1996: 72). Dacus (Zeugodacus) elegantis Tseng, Chen & Chu, 1992: 61 (synonymised by Drew & Romig, 2013: 353) Dacus (Zeugodacus) flavus Tseng, Chen & Chu, 1992:63. (synoymised by Drew & Romig, 2013: 353) Bactrocera (Zeugodacus) tau (Walker) (in Drew & Romig, 2013: 353). Zeugodacus (Zeugodacus) tau (Walker) (in Doorenweerd et al., 2018: 48). Material examined: 9♂♂, INDIA, Tripura, Agartala, 10.iii.2016, Sunil Joshi, 1♀, INDIA, Sikkim, Tadong, 26 Oct ̄ 3 Nov 2014, Sunil Joshi (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6N). Lateral surstylus longer than epandrium; posterior lobe of surstylus pointed, 9̄10 times longer than anterior lobe (Fig. 4J). Proctiger quadrate (when inflated) and smaller than epandrium. Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 3.2 mm excluding glans (0.45 mm); 0.75 of glans sclerotised with entirely patterned praeputium (conical), subapical lobe and basal lobe present (Fig. 10H). Female with brown, dorsoventrally flattened oviscape (2.05 mm); spicules on distal end of eversible membrane (3.2 mm) with 10̄12 pointed projections (medial ones 3̄4 times higher than lateral ones) (Fig. 12E). Aculeus (2.4 mm long), tip pointed with 4 pairs of preapical setae, apical setae 10-12 times longer than basal ones (Fig.14E); two black, tightly coiled mulberry shaped spemathecae (Fig. 15U)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 131, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["White, I. M. & Elson-Harris, M. M. (1992) Fruit flies of economic significance: their identification and bionomics. CAB International, Wallingford, 601 pp.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bactrocera) fastigata Tsuruta & White Bactrocera (Afrodacus) fastigata Tsuruta & White, 2001: 72. Bactrocera (Bactrocera) fastigata (Copeland et al., 2004: 151). Bactrocera (Afrodacus) fastigata (in Drew & Romig, 2013: 29). Bactrocera (Bactrocera) fastigata (in Hancock, 2015: 42). Material examined: 2♂♂, INDIA, Karnataka, Uduppi, Brahmavara, 31.i.2008, David, K. J. (UASB). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5I). Lateral surstylus shorter than epandrium; posterior lobe of surstylus blunt, not longer than anterior lobe (Fig. 3I). Proctiger quadrate and smaller than epandrium (Fig. 3I). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.40 mm long excluding glans (0.34 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8I)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 118-119, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Tsuruta, K. & White, I. M., (2001) Eleven new species of the genus Bactrocera Macquart (Diptera: Tephritidae) from Sri Lanka. Entomolgical Science, 4, 69 ‾ 87.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Bactrocera) nigrofemoralis White & Tsuruta Bactrocera (Bactrocera) nigrofemoralis White & Tsuruta (in Tsuruta & White, 2001) 2001: 79. Material examined: 1♂, INDIA, Karnataka, Mudigere, ZAHRS, 02.i.2015. Prasanna R.Y. 1♀, INDIA, Karnataka, Mudigere, ZAHRS, 02.i.2015. Prasanna R. Y. Attur farm, 29.vii.2013, David, K.J. (NBAIR). Diagnosis: Male with epandrium and surstyli spherical in outline (Fig. 5M) (posterior view). Lateral surstylus shorter than epandrium in profile; posterior lobe of surstylus blunt, as long as anterior lobe. Proctiger quadrate and smaller than epandrium (Fig. 3M). Phallus 1.9�� 2.2 mm long, excluding glans (0.4 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 9A). Female with black oviscape (1.16 mm); eversible membrane (1.4 mm) with spicules on distal end having four-five broad projections (Fig. 11H). Aculeus (1.2 mm long) with acute apex and four pairs of preapical setae (Fig. 13H), two black mulberry shaped spermathecae (Fig. 15H)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 121, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Tsuruta, K. & White, I. M., (2001) Eleven new species of the genus Bactrocera Macquart (Diptera: Tephritidae) from Sri Lanka. Entomolgical Science, 4, 69 ‾ 87."]}

Zeugodacus (Javadacus) scutellarius (Bezzi) Chaetodacus scutellarius Bezzi, 1916:110. Dacus (Bactrocera) scutellarius Hardy, 1977: 52. Bactrocera (Bactrocera) scutellaria (in Norrbom et al., 1999:95). Bactrocera (Javadacus) scutellaria (in Drew & Raghu, 2002: 328. Drew & Romig, 2013: 211). Zeugodacus (Javadacus) scutellarius (in Doorenweerd et al., 2018: 48). Material examined: 2♂♂, INDIA, Kerala, Kaniyambetta, 23.x.2005, David K. J. (UASB). Diagnosis: Male genitalia with epandrium and lateral surstyli bulb-shaped (posterior view). Lateral surstylus longer than epandrium; posterior lobe of surstylus narrow, curved in, 8̄10 times longer than anterior lobe. Proctiger triangular and smaller than epandrium (Fig. 3U). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae (Fig. 6A). Phallus 1.8̄ 2.1 mm long excluding glans (0.35 mm); 0.75 of glans sclerotised with patterned praeputium; subapical lobe and basal lobe present (Fig. 9I)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 127, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Zeugodacus) scutellaris (Bezzi) Bactrocera scutellaris Bezzi, 1913: 98. Zeugodacus malaisei Hering, 1938: 4. (synonymised by Norrbom et al., 1999: 103) Dacus pusaensis Kapoor & Katiyar, 1970:252. (synonymised by Norrbom et al., 1999: 103) Dacus (Zeugodacus) scutellaris, Hardy, 1973: 68. Bactrocera (Zeugodacus) fraserensis Chua, 2010: 483. (synonymised by Drew & Romig, 2013: 340) Bactrocera (Zeugodacus) scutellaris (Bezzi) (in Drew & Romig, 2013: 340). Zeugodacus (Zeugodacus) scutellaris (Bezzi) (in Doorenweerd et al., 2018: 48). Material examined: 32♂♂, INDIA, Himachal Pradesh, Shimla, Pawabo, September 2015, Sunil Joshi (UASB). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6M). Lateral surstylus longer than epandrium; posterior lobe of surstylus broad, curved in, 10 times longer than anterior lobe (Fig. 4I). Proctiger triangular and smaller than epandrium (Fig. 4I). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.1 mm excluding glans (0.42 mm); 0.75 of glans sclerotised with patterned praeputium and apex of acrophallus with a ring like structure; subapical lobe and basal lobe present (Fig. 10G)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 131, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Genus Dacus Fabricius Dacus Fabricius, 1805: 272. (in Drew, 1989: 16). Type species: Dacus armatus Fabricius, 1805, by subsequent designation of Speiser, 1924: 140 and confirmed by Hendel (1927: 25). Diagnosis. Abdomen elongate-oval or petiolate, abdominal tergites fused. Scutum with or without lateral and medial postsutural vittae, chaetotaxy reduced compared to Bactrocera and Zeugodacus Hendel; scutellum short and broad. Posterior margin of sternite V of male with shallow to moderate concavity (emargination). Male genitalia with epandrium and surstyli distinctly bulb-shaped (posterior view); posterior lobe of surstylus 2��3 times longer than anterior lobe except in D. longicornis. Glans of phallus with single tubular acrophallus, praeputium fully patterned, predominantly hexagonal pattern except elongate pattern in D. persicus. Aculeus tip with three pairs of preapical setae. It is similar to Bactrocera and Zeugodacus in general appearance but can be easily separated by the presence of fused abdominal tergites. Subgenus Sinodacus Zia of genus Zeugodacus and subgenera Semicallantra Drew and Tetradacus Miyake of Bactrocera possess petiolate abdomen as in Dacus but can be differentiated by free abdominal tergites., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 123, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. (1989) The tropical fruit flies (Diptera: Tephrititidae: Dacinae) of the Australasian and Oceanic regions. Memoirs of the Queensland Museum, 26, 1 ‾ 521."]}

Zeugodacus (Hemigymnodacus) diversus (Coquillett) Dacus diversus Coquillett, 1904: 139. Asiadacus diversa Perkins, 1937: 57; 1938a: 134 (as diversus) Dacus (Gymnodacus) diversus Hardy, 1954: 18. Dacus (Hemigymnodacus) diversus Hardy, 1973: 19. Dacus quadrifidus Hendel, 1928: 343 (synonymised by Norrbom et al., 1999: 99). Dacus citronellae Kapoor & Katiyar, 1969: 123 (synonymised by Norrbom et al., 1999: 99). Dacus latifae Anwar Cheema, 1964: 302 (synonymised by Drew & Romig, 2013). Bactrocera (Paratridacus) diversa (in Norrbom et al., 1999: 99). Bactrocera (Hemigymnodacus) diversa (in Drew & Romig, 2013). Zeugodacus (Hemigymnodacus) diversus (in Doorenweerd et al., 2018: 45). Material examined: 1♂, INDIA, Karnataka, Bangalore, Attur farm, 11.xi.2013, David, K. J., 3♀♀, INDIA, Maharashtra, Konkan Region, September 2012, H. R. Sawai. (NBAIR). Diagnosis: Male with epandrium and surstyli bulb-shaped, i.e., with prominent constriction between epandrium and surstyli (posterior view) (Fig. 5T). Lateral surstylus longer than epandrium; posterior lobe of surstylus 10 times longer than anterior lobe (Fig. 3T). Proctiger triangular shaped and smaller than epandrium. Medial surstylus as long as lateral surstylus (Fig. 5T). Phallus 2.35 mm long excluding glans (0.39 mm); glans sclerotised with patterned praeputium; trumpet-shaped subapical lobe and short basal lobe (Fig. 9H). Female with dark-brown to black oviscape (1.72 mm); eversible membrane (2.02 mm) with spicules on distal end possessing 8̄11 small, blunt, projections (Fig. 11K); aculeus tip acute, needle shaped, with four pairs of preapical setae (Fig. 13K); two tightly coiled, mulberry shaped black spermathecae (Fig. 15P)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 127, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Zeugodacus) fuscoalatus (Drew & Romig) Bactrocera (Zeugodacus) fuscoalata Drew & Romig, 2013: 295. Zeugodacus (Zeugodacus) fuscoalatus (Drew & Romig) (in Doorenweerd et al., 2018: 46). Material examined: 1♂, 1♀, INDIA, Andaman and Nicobar Islands, Andaman Islands, South Andamans, Mt. Harriet, 25. iv. 1995, H. R. Ranganath. (UASB). Diagnosis: Male genitalia with epandrium and surstyli bulb-shaped (posterior view) (Fig. 6J). Lateral surstylus longer than epandrium; posterior lobe of surstylus 10̄12 times longer than anterior lobe. Medial surstylus as long as lateral surstylus with thick prensisetae (Fig. 4F). Phallus elongate (4.3 mm), excluding glans (0.47 mm); glans sclerotised with well developed patterned praeputium (triangular), trumpet-shaped subapical lobe and short basal lobe (Fig. 10D). Female with dorsoventrally flattened, brown oviscape (2.44 mm); eversible membrane (5.5 mm) with spicules on distal end having 6̄7 stout, lanceolate projections (Fig. 12C). Aculeus 2.7 mm long, tip trilobed with 4 pairs of preapical setae (Fig. 14C); two black, tightly coiled mulberry shaped spemathecae (Fig. 15S)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 130, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Zeugodacus) zahadi (Mahmood) Bactrocera (Zeugodacus) zahadi Mahmood, 1999: 232 Zeugodacus (Zeugodacus) zahadi Mahmood (in Doorenweerd et al. 2018: 49). Material examined: 1♂, INDIA, Karnataka, Vittal, CPCRI, 18.ii.2015, Prabhu, G., 2♂♂, INDIA, Karnataka, Mudigere, September 2015, Sunil Joshi, Agartala, 10.iii.2016, Sunil Joshi, 1♂, INDIA, Tamil Nadu, Ooty, Wellington, 18.ii.2016, Prabhu, G., 1♂, INDIA, Karnataka, Bangalore, Attur farm, 09.vii.2013, David, K. J.; 1♀, INDIA, Karnataka, Madikeri, CHES, Chettalli, 05-09.xi.2012., David, K.J. (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6P). Lateral surstylus longer than epandrium; posterior lobe of surstylus pointed, 9̄10 times longer than anterior lobe (Fig. 4L). Proctiger membranous, triangular and smaller than epandrium. Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.7 mm long excluding glans (0.48 mm); 0.75 of glans sclerotised with patterned praeputium (conical projections on basal half), subapical lobe and basal lobe present (Fig. 10J). Female with dorsoventrally flattened brown oviscape (1.5 mm); eversible membrane (2.2 mm) with spicules on distal end having 5-6 stout, pointed projections (Fig. 12G); aculeus (1.7 mm) tip pointed with 4 pairs of preapical setae (Fig. 14G), apical ones 10-12 times longer than basal ones, two black, tightly coiled mulberry shaped spemathecae (Fig 15W)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 132, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bactrocera) vishnu Drew & Hancock Bactrocera (Bactrocera) vishnu Drew & Hancock, 1994: 65. Material examined: 1♂, INDIA, Karnataka, Madikeri, CHES, Chettalli, 05-09.xi.2012, David, K. J., 1♂, INDIA, Karnataka, Bangalore, Attur, 30. vi. 2014, Prabhu, G. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (Fig. 5R) (posterior view). Lateral surstylus shorter than epandrium; posterior lobe of surstylus broad, not longer than anterior lobe (Fig. 3R). Ventral rim of lateral surstylus 3��4 times longer than dorsal rim. Proctiger membranous, quadrate smaller than epandrium (Fig. 3R). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.0�� 2.3 mm long excluding glans (0.38 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 9F)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 122, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, 2 (Supplement), iii + 68 pp."]}

Bactrocera (Bactrocera) ranganathi Drew & Romig Bactrocera (Bactrocera) ranganathi Drew & Romig, 2013: 168. Material examined: 2♂♂, INDIA, Andaman Islands, South Nicobar, Campbell Bay, 16. vi. 1997, H. R. Ranganath. (UASB). Diagnosis: Male with epandrium and surstyli oval shaped (posterior view) (Fig. 5O). Lateral surstylus shorter epandrium; posterior lobe of surstylus pointed, as long as anterior lobe, forked (lateral view) (Fig. 3O). Proctiger quadrate and smaller than epandrium. Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 1.7 mm long excluding glans (0. 41 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 9C)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 121, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Parazeugodacus) bipustulata (Bezzi) Bactrocera bipustulata Bezzi, 1914: 153. Bactrocera (Zeugodacus) bipustulata (in Norrbom et al., 1999: 102). Bactrocera (Parazeugodacus) bipustulata (in Drew & Romig, 2013: 245; Hancock & Drew, 2015: 93). Material examined: 1♀, INDIA, Tamil Nadu, Gudalur, 8. iii. 2017, Saranya Babu; 2 ♂♂, INDIA, Karnataka, Bangalore, Hebbal, 16.viii.2011, David, K. J., 1♂, INDIA, Karnataka, 0 6. xi. 2012, David, K. J. (NBAIR) Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 6F). Lateral surstylus shorter than epandrium in profile; posterior lobe of surstylus blunt, as long as anterior lobe (Fig. 4B). Proctiger quadrate and smaller than epandrium (Fig. 4B). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 2.2 mm long excluding glans (0.42 mm). Three-fourth of glans sclerotised with patterned praeputium (Fig. 9N); acrophallus with dorsal notch; subapical lobe present. Female with black (1.13 mm) oviscape; eversible membrane (1.05 mm) with spicules on distal end conical with 2-3 projections (Fig. 12 A); aculeus (1.06 mm) with acute apex (Fig. 14A); two black, convoluted mulberry shaped spermathecae (Fig. 15L)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 123, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Zeugodacus (Zeugodacus) gavisus (Munro) Dacus gavisus Munro, 1935: 15. Bactrocera (Zeugodacus) gavisa (in Norrbom et al., 1999: 102; Drew & Romig 2013: 296). Zeugodacus (Zeugodacus) gavisus (Munro) (in Doorenweerd et al., 2018: 46). Material examined: 2♂♂, INDIA, Karnataka, Bangalore, Attur farm, 01.x.2012, David, K.J., 1♂ same data except date 26.v.2014 and collector, Prabhu, G., 1♂ same data of previous except date 30.vi.2014 (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6K). Lateral surstylus longer than epandrium; posterior lobe of surstylus 10 times longer than anterior lobe (Fig. 4G). Medial surstylus as long as lateral surstylus. Proctiger triangular and smaller than epandrium. Phallus 2.3 mm long excluding glans (0.35 mm); glans sclerotised with well developed patterned praeputium (triangular and circular pattern), trumpet-shaped subapical lobe and short basal lobe; vesica sclerotised slightly to form hook-like structure (Fig. 10E)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 130, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Javadacus) trilineatus (Hardy) Dacus (Afrodacus) trilineatus Hardy, 1955: 12. Bactrocera (Javadacus) trilineata (Hardy) (in Norrbom et al., 1999: 98; Drew & Romig, 2013 -213). Zeugodacus (Javadacus) trilineatus (Hardy) (in Doorenweerd et al., 2018 -49). Material examined: 3♂♂, INDIA, Tamil Nadu, Hosur, Rayakottai, 29.ii.2016, Prabhu, G, 1♀, INDIA, Karnataka, Mudigere, KVK, 13.iii.2014, Rajesh, S. (NBAIR). Diagnosis: Male with epandrium and surstyli bulb-shaped (posterior view) (Fig. 6B). Lateral surstylus longer than epandrium; posterior lobe of surstylus broad, curved in, 10 times longer than anterior lobe. Proctiger triangular and smaller than epandrium (Fig. 3V). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 1.79̄2.0 mm long excluding glans (0.35 mm); 0.75 of glans sclerotised with patterned praeputium; subapical lobe, basal lobe present (Fig. 9J). Female with black oviscape (1.2 mm); eversible membrane 1.7 mm long, spicules on distal end broad with four-five teeth like projections (Fig. 11L); aculeus (1.2 mm) with apex triangular and four pairs of preapical setae (Fig. 13L); two black, spermatheca (highly convoluted, mulberry shaped) (Fig. 15 Q)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 128, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bactrocera) paraosbeckiae Drew Bactrocera (Bactrocera) paraosbeckiae Drew (in Drew & Raghu, 2002) 2002: 340. Material examined: 2♂♂, INDIA, Kerala, Calicut, Maroor, 13. xi. 2012, Devaraj (NBAIR). Diagnosis: Male with epandrium and surstyli inverted U-shaped in outline (Fig. 5N) (posterior view). Lateral surstylus (0.1 mm wide) longer than epandrium; posterior lobe of surstylus pointed, curved back, not longer than anterior lobe. Ventral rim of lateral surstylus as long as the dorsal rim (0.2 mm). Proctiger dome shaped and smaller than epandrium (Fig. 3O). Medial surstylus tapering towards apex with a pair of thick prensisetae. Phallus 1.8��2.0 mm long excluding glans (0.40 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 9B)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 121, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635

Dacus (Callantra) longicornis Wiedemann Dacus longicornis Wiedemann, 1830: 524. Callantra smieroides Walker, 1860: 154. (synonymised by Drew et al., 1998) Dacus vespoides Bezzi, 1909: 262. Mellesis destillatoria Bezzi, 1916: 118. (synonymised by Drew et al., 1998) Mellesis eumenoides Bezzi, 1916: 437. (synonymised by Drew et al., 1998) Mellesis bioculata Bezzi, 1919: 437 (in Drew et al., 1998: 604) (synonymised by Drew & Romig, 2013) Callantra vespoides Enderlein, 1920: 358. (synoymised by Hardy, 1977) Callantra smieroides Malloch, 1939: 411. Callantra destillatoria Hardy, 1973:9. (synonymised by Drew et al., 1998) Callantra eumenoides Hardy, 1973:11.(synonymised by Drew et al., 1998) Callantra bioculata Hardy, 1974: 6. (synonymised by Drew et al., 1998) Callantra vareigata Wang, 1990: 73.(synonymised by Drew et al., 1998) Dacus (Callantra) vareigata (in Liang et al., 1993: 139). Dacus (Callantra) eumenoides (in Kapoor, 1993: 83). Callantra longicornis (in Hardy, 1977: 45). Callantra unifasciatus (in Hardy, 1982: 184).(synonymised by Drew et al., 1998) Dacus (Callantra) longicornis (in Drew & Romig, 2013: 373). Material examined: 1♂, INDIA, Manipur, Imphal, Lamphelpat, 779 m, 24.83 o N, 93.93 o E, 26.ix. 2013, Yeshwanth, H.M., 1♂, INDIA, Karnataka, Bangalore, 09.xi. 2009. (UASB), 1♀, Host. Jackfruit, Locn. Gakulpur, Date 1.1.0 7, Coll. J. Das., Agarwala Collection (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view). (Fig. 7B). Lateral surstylus shorter than epandrium; apex of posterior lobe of surstylus pointed, as long as anterior lobe. Proctiger quadrate and smaller than epandrium (Fig. 4N). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae (Fig. 7B). Phallus 2.3 to 2.5 mm long excluding glans (0.425 mm); 0.75 of glans sclerotised with patterned (hexagonal) praeputium; subapical lobe and basal lobe present (Fig. 10L). Female with brown oviscape (1.7 mm); elongate eversible membrane (4.03 mm), spiclues on distal end of eversible membrane with 5̄6 projections, tapering laterally (Fig. 12I); aculeus 2.4 mm long, with acute apex and three pairs of preapical setae (Fig. 14 I). Two brown, convoluted mulberry shaped spermathecae (Fig. 15 M)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 124, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Dacus (Didacus) ciliatus Loew Dacus ciliatus Loew, 1862:7. Dacus brevistylus Bezzi, 1908:149. Dacus (Bactrocera) cocciniae Premlata & Singh, 1987: 401. Dacus (Afrodacus) cocciniae Agarwal et al., 1992: 20. Dacus (Didacus) ciliatus (in Drew et al., 1998: 646; Drew & Romig, 2013: 374). Material examined: 6♂, INDIA, New Delhi, IARI, 12. vii. 2012, Shakti, K. Singh; 1♀, INDIA, Tamil Nadu, Periyakulam, 30. iii. 2012, David, K. J.; 8♂, INDIA, Meghalaya, Barapani, February 2014, Boopathi (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7C). Lateral surstylus surstylus broad, shorter than epandrium; posterior lobe of surstylus 3 times longer than anterior lobe, hook shaped (Fig. 4O); medial surstylus as long as lateral surstylus, with a pair well developed prensisetae at its apex. Proctiger triangular and not larger than epandrium (Fig. 4O). Phallus (2 mm long) excluding glans (0.3 mm); glans sclerotised with well developed fully patterned praeputium, subapical lobe and basal lobe present (Fig. 10M). Female with fuscous oviscape (1.21 mm), eversible membrane (2.1 mm) with spicules on distal end having 4��5 projections of equal height (Fig. 12J); aculeus (1.43 mm) tip pointed with three pairs of preapical setae (Fig. 14J). Two black, convoluted mulberry shaped spermathecae (Fig. 15N)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 124, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Bactrocera) dorsalis (Hendel) Musca ferruginea Fabricius, 1794: 342. Preoccupied by Musca ferruginea Scopoli, 1763 Dacus ferrugineus Fabricius, 1805: 274. Dacus dorsalis Hendel, 1912: 18 Bactrocera ferruginea Bezzi, 1913: 95. Chaetodacus ferrugineus var. dorsalis Hendel, 1915: 426. Chaetodacus ferrugineus Bezzi, 1916: 104. Chaetodacus ferrugineus dorsalis Bezzi, 1916: 104. Chaetodacus ferrugineus var. okinawanus Shiraki, 1933: 62 (synonymised by Hardy, 1969: 402.) Dacus (Strumeta) dorsalis (in Hardy & Adachi, 1956). Strumeta dorsalis (in Hering, 1956: 63). Strumeta ferruginea Hering, 1956: 63 Strumeta dorsalis okinawana Shiraki, 1968: 23. Dacus (Bactrocera) dorsalis Hardy, 1977: 49. Dacus (Bactrocera) semifemoralis Tseng, Chen & Chu, 1992: 46. (synonymised by Drew & Romig, 2013: 76) Dacus (Bactrocera) yilanensis Tseng, Chen & Chu, 1992, 52. (synonymised by Drew & Romig, 2013: 76) Bactrocera (Bactrocera) dorsalis Drew & Hancock, 1994: 17. Bactrocera (Bactrocera) variabilis Lin & Wang, in Lin et al., 2011b: 896. (synonymised by Drew & Romig, 2013: 76). Bactrocera (Bactrocera) invadens Drew, Tsuruta & White, 2005: 149.(synonymised by Schutze et al., 2014) Bactrocera (Bactrocera) papayae Drew & Hancock, 1994: 48 (synonymised by Schutze et al., 2014). Bactrocera (Bactrocera) philippinensis Drew & Hancock, 1994: 52 (synonymised by Schutze et al., 2014) Material examined: 1♂, INDIA, Andhra Pradesh, Thirupati, 22.v.2015, Pampapathi, 9♂♂, INDIA: Karnataka, Bangalore, Ramagiri, August 2014, Raghavendra, A; 2♂♂, INDIA, Karnataka, Bangalore, Attur, 10.xi. 2014, Prabhu, G; 1♂, same data except Hebbal, 22.ii.2013, Prabhu, G. 1♀, INDIA, Karnataka, Bangalore, Attur, 0 2. iv. 2014, Prabhu, G; 1♀, INDIA, Karnataka, Bangalore, IIHR, Hessaraghatta, 12.v.2010, Rajeswari. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5G). Lateral surstylus shorter than epandrium; posterior lobe of surstylus not longer than anterior lobe (Fig. 3G). Proctiger quadrate and smaller than epandrium (Fig. 3G). Medial surstylus longer than lateral surstylus, tapering towards apex with a pair of thick prensisetae. Phallus 1.80 mm long, excluding glans (0.42 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8G). Female with reddish brown, dorsoventrally flattened, oviscape (1.4 mm); eversible membrane (1.70 mm) with spicules on distal end, broader than high, with11-12 small pointed projections (Fig. 11E); aculeus (1.5 mm) with acute apex and four pairs of preapical setae (Fig. 13E); two black, convoluted, mulberry shaped spermathecae (Fig. 15F)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 116-117, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Drew, R. A. I. & Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, 2 (Supplement), iii + 68 pp."]}

Bactrocera (Bactrocera) zonata (Saunders) Dasyneura zonatus Saunders, 1842: 61. Bactrocera maculigera Doleschall, 1858: 122 (synoymized by Norrbom et al., 1999) Rivellia persicae Bigot, 1890: 192. Dacus ferrugineus var. mangifera e Cotes, 1893: 17. Dacus (Strumeta) zonatus Saunders (in Hardy, 1973: 54). Bactrocera (Bactrocera) zonata (in Norrbom et al., 1999: 96). Material examined: 5♂♂, INDIA, Karnataka, Hassan, Arthur, 23.v.2015, Abraham Verghese, 14♀♀, INDIA: New Delhi, IARI, 30.iv.2012, Shakti, K. Singh (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (Fig. 5S) (posterior view). Epandrium as long as lateral surstylus in profile; posterior lobe of surstylus blunt, as long as anterior lobe (Fig. 3S). Proctiger membranous, quadrate smaller than epandrium (Fig. 3S). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 1.9̄ 2.12 mm long excluding glans (0.36 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 9G). Female with reddishbrown, dorsoventrally flattened oviscape (1.2 mm); eversible membrane (1.42 mm) with broad spicules on distal end having 4-5 blunt projections (Fig. 11 I), aculeus short (1.01 mm) with acute apex and four pairs of preapical setae (Fig. 13 I); two black convoluted mulberry shaped spermathecae (Fig. 15J)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 122, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251."]}

Zeugodacus (Zeugodacus) assamensis White Bactrocera (Zeugodacus) assamensis White (in White & Evenhuis, 1999: 522). Bactrocera (Zeugodacus) gansuica Chen, Han & Zhou, in Chen et al., 2012: 352. (synonymised by Drew & Romig, 2013) Bactrocera (Zeugodacus) assamensis White (in Drew & Romig, 2013: 268). Zeugodacus (Zeugodacus) assamensis White (in Doorenweerd et al., 2018: 44). Material examined: 2♂♂, INDIA, Himachal Pradesh, Shimla, June 2011, Sushma, ex. Cherry, (UASB). Diagnosis: Male genitalia with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6G). Lateral surstylus longer than epandrium; posterior lobe of surstylus pointed, punctated, 8 times longer than anterior lobe (Fig. 4C). Proctiger triangular and smaller than epandrium. Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 3 mm long excluding glans (0.35 mm); 0.75 of glans sclerotised with patterned/granulated praeputium; subapical lobe and basal lobe present (Fig. 10A)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 129, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bulladacus) cinnabaria Drew & Romig Bactrocera (Bulladacus) cinnabaria Drew & Romig, 2013:197. Material examined: 1♀, INDIA, Andaman and Nicobar Islands, Car Nicobar Island, 0 4. vii. 1995, H. R. Ranganath (UASB) Diagnosis: Female with reddish-brown, dorsoventrally flattened oviscape (0.7 mm); eversible membrane (0.6 mm) with broad spicules on distal end having 10��12 projections (Fig. 11J); aculeus short (0.8 mm) with acute apex and four pairs of preapical setae (Fig. 13J); two black, convoluted mulberry shaped spermathecae (Fig. 15K)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 123, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Dacus (Leptoxyda) persicus Hendel Dacus (Leptoxyda) persicus Hendel, 1927a: 29. Dacus ferrugineus -Becker, 1913:641[Misidentification] Leptoxyda sp. nr. longistyla Bezzi, 1913: 92. Dacus longistylus Wiedemann, 1830 (Asian records) [misidentifcations by various authors] Dacus (Leptoxyda) persicus Hendel (in Drew & Romig, 2013: 375). Material examined: INDIA: 1♀, Karnataka, Mysore, 9.v. 2007, Rajeswari, E. K., 1♂, INDIA, Karnataka, Bangalore, Hebbal, 28.vii. 2014, Prabhu, G., 1♀, INDIA, Karnataka, Bangalore, Hebbal, 13.x.2014, Prabhu, G., 1♀, INDIA, Karnataka, Bangalore, Attur, 19.vii.2014, Verghese, A., 1♀, INDIA, Karnataka, Bangalore, Hebbal, 30.vii.2014, Prabhu, G., 1♀, INDIA, Karnataka, Bangalore, Hebbal, 18.vi.2014, Prabhu, G., 1♀, INDIA, Karnataka, Bangalore, Hebbal, 13.v.2014, Prabhu, G., INDIA, Karnataka, Bangalore, Hebbal, 0 2. viii.2012, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7D). Lateral surstylus 3��4 times longer than anterior lobe, apex of posterior lobe of surstylus hook shaped, curved in (Fig. 4P). Proctiger quadrate, almost as high as epandrium. Medial surstylus flattened, longer than lateral surstylus with a pair of short prensisetae (Fig. 7D). Phallus 4.8��5.0 mm long excluding glans (0.30�� 0.32 mm); 0.75 of glans sclerotised with elongate patterned praeputium, well developed vesica and basal lobe present (Fig. 10N). Female with bottle shaped oviscape (bulbous base with a narrow neck) 2.3 �� 2.7 mm long; eversible membrane (5 mm) with elongate spicules on distal end having 18��20 sharp projections (medial ones higher than lateral ones) (Fig. 12K). Aculeus 3.6 to 3.8 mm long with three pairs of preapical setae and rounded apex (Fig. 14K). Two brown, mulberry shaped, spermathecae, (Fig. 15O)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 125, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Bactrocera (Bactrocera) affinis (Hardy) Dacus (Neodacus) affinis Hardy, 1954:7. Dacus (Polistomimetes) oscinae Agarwal & Kapoor, 1983: 171. (synonymized by Norrbom et al., 1999:87) Bactrocera (Bactrocera) affinis (in Norrbom et al., 1999:87). Material examined: 2♂♂, INDIA, Karnataka, Bangalore, Attur farm, 29.vii.2013, David, K.J. (NBAIR) Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5A). Lateral surstylus shorter than epandrium, posterior lobe of surstylus pointed, as long as anterior lobe (Fig. 3A). Proctiger quadrate and smaller than epandrium. Medial surstylus with a pair of thick prensisetae (Fig. 5A). Phallus 2.8̄3.0 mm long excluding glans (0.40 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8A)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 109, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251."]}

Dacus (Neodacus) sphaeroidalis (Bezzi) Mellesis sphaeroidalis Bezzi, 1916:115. Callantra sphaeroidalis Hardy, 1973: 11; Hardy, 1977: 46. Dacus (Callantra) sphaeroidalis (in Liang et al., 1993: 139; Drew et al., 1998:629: Norrbom et al., 1998: 131). Sinodacus chonglui Chao & Lin, 1996: 131. (synonymised by Drew & Romig, 2013) Dacus (Neodacus) sphaeroidalis (in Drew & Romig, 2013: 402). Material examined: 3♂♂, INDIA, Chandigarh, Panchkula, 12.ix.2009, Shakti Singh (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7H). Lateral surstylus shorter than epandrium, apex of posterior lobe of surstylus blunt, 3 times longer than anterior lobe. Proctiger triangular and smaller than epandrium (Fig. 4T). Medial surstylus longer than lateral surstylus with well developed prensisetae. Phallus 3.7 mm long excluding glans (0.5 mm); 0.75 of glans sclerotized, preaputium patterned (Fig. 10R)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 126, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Dacus (Mellesis) crabroniformis (Bezzi) Monacrostichus crabroniformis Bezzi, 1914b: 153. Mellesis crabroniformis -Bezzi, 1916: 117. Callantra crabroniformis - Hardy, 1977: 45. Dacus (Callantra) crabroniformis (in Kapoor, 1993: 83, Drew et al., 1999: 584; Norrbom et al., 1999: 130; David & Ramani, 2011:4; Drew & Romig, 2013: 379). Material examined: 1♂, INDIA, Karnataka, Bangalore, Attur, 0 7. xii. 2014., Prabhu, G., 1♂, INDIA, Karnataka, Bangalore, Acharlahalli, 2. ii. 2011, Basanth, Y. S. (NBAIR). Diagnosis: Epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 7E). Lateral surstylus shorter than epandrium, apex of posterior lobe of surstylus blunt, 4 times longer than anterior lobe. Proctiger triangular and smaller than epandrium (Fig. 4Q). Medial surstylus longer than lateral surstylus with well developed prensisetae. Phallus 1.6 mm long, excluding glans (0. 36 mm); 0.75 of glans sclerotized, preaputium patterned (Fig. 10 O)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 125, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Hancock, D. L. (1999) Phylogeny of the Tribe Dacini (Dacinae) based on morphological, distributional, and biological data. In: Aluja, M. & Norrbom, A. L. (Eds.), Fruit Flies (Tephritidae): Phylogeny and Evolution of Behavior. CRC Press, Boca Raton, pp. 491 - 504.","Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","David, K. J. & Ramani, S. (2011) An illustrated key to fruit flies (Diptera: Tephritidae) from Peninsular India and the Andaman and Nicobar Islands. Zootaxa, 3021, 1 - 31.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Zeugodacus (Parasinodacus) incisus (Walker) Dacus incisus Walker, 1861d: 323. Dacus (Strumeta) incisus (in Hardy, 1959: 175). Bactrocera (Bactrocera) incisa (in Norrbom et al., 1999: 91). Dacus poonensis Kapoor, 1971: 478 (synonymised by Norrbom et al., 1999: 91) Bactrocera (Zeugodacus) incisa (in Chen et al., 2011: 60). Bactrocera (Parasinodacus) incisa (in Drew & Romig, 2013: 230). Zeugodacus (Parasinodacus) incisus (in Doorenweerd et al., 2018: 46). Material examined: 2♂♂, INDIA, Andaman Islands, 16. vi. 1995, H. R. Ranganath, Cue lure (UASB). Diagnosis: Male with epandrium and surstyli bulb-shaped (posterior view) (Fig. 6E). Lateral surstylus longer than epandrium; posterior lobe of lateral surstylus broad, 7̄8 times longer than anterior lobe, both lobes curved in (Fig. 4A). Medial susrtylus as long as lateral surstylus with well developed prensisetae. Phallus 4.7 mm, glans of phallus sclerotised with well developed, patterned praeputium; basal lobe and subapical lobe present (Fig. 9M)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on pages 128-129, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Zeugodacus (Zeugodacus) caudatus (Fabricius) Dacus caudatus Fabricius, 1805; 276. Bactrocera maculipennis Doleschall, 1856:412. Dacus (Zeugodacus) caudatus (in Hardy & Adachi, 1954: 186). Bactrocera (Zeugodacus) caudata (in Norrbom et al., 1999: 102; Drew & Romig, 2013: 277). Zeugodacus (Zeugodacus) caudatus (Fabricius) (in Doorenweerd et al., 2018: 44). Material examined: 2♂♂, INDIA, Tripura, Agartala, 10.iii.2016, Sunil Joshi (NBAIR). Diagnosis: Male with epandrium and surstyli bulb-shaped (posterior view) (Fig. 6H). Lateral surstylus longer than epandrium, posterior lobe of surstylus 10 times longer than anterior lobe (Fig. 4D). Proctiger as high as long smaller than epandrium. Medial surstylus as long as lateral surstylus. Phallus 2.33 mm long, excluding glans (0.38 mm); glans sclerotised with well developed patterned praeputium (triangular pattern on basal half), praeputium, trumpet-shaped subapical lobe and short basal lobe (Fig. 10B)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 129, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bactrocera) blairiae Drew & Romig Bactrocera (Bactrocera) blairiae Drew & Romig, 2013: 57. Material examined: 2♂♂, INDIA, Andaman and Nicobar Islands, Andaman Islands, Mt. Harriet, 11.iii. 2012, David, K. J., 3♂♂, INDIA, Andaman and Nicobar Islands, Andaman Islands, Chouldari, Bloomsdale, 8.iii. 2012, David, K. J. (NBAIR); 1♀, INDIA, Andaman and Nicobar Islands, Andaman Islands, Mt. Harriet, 25.iv. 1995, H. R. Ranganath (UASB). Diagnosis: Male with epandrium and surstyli oval in outline (posterior view) (Fig. 5C). Lateral surstylus shorter than epandrium; posterior lobe of surstylus broad, curved in, as long as anterior lobe; ventral rim of lateral surstylus 3 times longer than dorsal rim (Fig. 3C). Proctiger quadrate and smaller than epandrium (Fig. 3C). Medial surstylus with a pair of thick prensisetae (Fig. 5C). Phallus 2.8��3.0 mm long excluding glans (0.42 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present (Fig. 8C). Female with dorsoventrally flattened, reddish-brown oviscape (1.89 mm); eversible membrane (2.70 mm) with conical spicules on distal end (Fig. 11B); aculeus short (1.9 mm), apex acute with smooth preapical steps (Fig. 13B), four pairs of preapical setae; two black convoluted mulberry shaped spermathecae (Fig. 15B)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 112, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp."]}

Zeugodacus (Zeugodacus) watersi (Hardy) Dacus (Neodacus) watersi Hardy, 1954: 12. Bactrocera (Paradacus) watersi (in Norrbom et al., 1999: 99). Bactrocera (Sinodacus) watersi (in Drew & Raghu, 2002: 328). Bactrocera (Zeugodacus) watersi (in Drew & Romig, 2013: 368). Zeugodacus (Zeugodacus) watersi (in Doorenweerd et al., 2018: 49). Material examined: 8♂♂, 10♀♀, INDIA, Karnataka, Madikeri, CHES, Chettalli, 05-09. xi.2012, David, K. J. (NBAIR). Diagnosis: Male with epandrium and surstyli distinctly bulb-shaped (posterior view) (Fig. 6O). Lateral surstylus longer than epandrium; posterior lobe of surstylus 10̄11 times longer than anterior lobe (Fig. 4K). Proctiger triangular shaped and smaller than epandrium. Medial surstylus as long as lateral surstylus with well developed prensisetae. Phallus elongate, 4.5 mm long, excluding glans (0.44 mm); glans sclerotised with well developed patterned praeputium, trumpet shaped subapical lobe and short basal lobe (Fig. 10 I). Female with redbrown, elongate oviscape (2.5 mm), eversible membrane long (4.12 mm) with spicules on distal end possessing 2̄3 broad lanceolate projections (Fig. 12F); aculeus (2.93 mm) tip trilobed with 4 pairs of preapical setae (Fig. 14F); two black, highly convoluted mulberry shaped spermathecae (Fig. 15V)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 132, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251.","Drew, R. A. I. & Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, 653 pp.","Doorenweerd, C., Leblanc, L., Norrbom, A. L., San Jose, M. & Rubinoff, D. (2018) A global checklist of the 932 fruit fly species in the tribe Dacini (Diptera: Tephritidae). Zookeys, 730, 17 ‾ 54. https: // doi. org / 10.3897 / zookeys. 730.21786"]}

Bactrocera (Bactrocera) versicolor (Bezzi) Chaetodacus ferrugineus var. versicolor Bezzi, 1916: 105. Bactrocera (Bactrocera) versicolor (in Norrbom et al., 1999: 96). Material examined: 42♂♂, INDIA, Karnataka, Coorg, Chettalli, 14.ix. 2012, Ranganath, H. R. (NBAIR). Diagnosis: Male with epandrium and surstyli oval in outline (Fig. 5Q) (posterior view). Lateral surstylus shorter than epandrium; posterior lobe of surstylus pointed at apex (in profile view), curved in, as long as anterior lobe. Proctiger quadrate and smaller than epandrium (Fig. 3Q). Medial surstylus longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus 3.7̄ 4.1 mm long excluding glans (0.4 mm); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe, and basal lobe present (Fig. 9E)., Published as part of David, K. J. & Ramani, S., 2019, New species, redescriptions and phylogenetic revision of tribe Dacini (Diptera: Tephritidae: Dacinae) from India based on morphological characters, pp. 101-146 in Zootaxa 4551 (2) on page 122, DOI: 10.11646/zootaxa.4551.2.1, http://zenodo.org/record/2622635, {"references":["Norrbom A. L., Carroll, L. E., Thompson, F. C., White I. M. & Freidberg, A. (1999). Systematic database of names. In: Thompson F. C. (Ed.), Myia. Uol. 9. Fruit fly expert identification system and systematic information database. Backhuys Publishers, Leiden, pp. 65 - 251."]}