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The fossil fuel crisis is a major concern across the globe, and fossil fuels are being exhausted day by day. It is essential to promptly change from fossil fuels to renewable energy resources for transportation applications as they make a major contribution to fossil fuel consumption. Among the available energy resources, a fuel cell is the most affordable for transportation applications because of such advantages as moderate operating temperature, high energy density, and scalable size. It is a challenging task to optimize PEMFC operating parameters for the enhancement of performance. This paper provides a detailed study on the optimization of PEMFC operating parameters using a multilayer feed-forward neural network, a genetic algorithm, and the design of a multiport high-gain DC–DC converter for hybrid electric vehicle application, which is capable of handling both a 6 kW PEMFC and an 80 AH 12 V heavy-duty battery. To trace the maximum power from the PEMFC, the most recent SFO-based MPPT control technique is implemented in this research work. Initially, a multilayer feed-forward neural network is trained using a back-propagation algorithm with experimental data. Then, the optimization phase is separately carried out in a neural-power software environment using a genetic algorithm (GA). The simulation study was carried out using the MATLAB/R2022a platform to verify the converter performance along with the SFO-based MPPT controller. To validate the real-time test bench results, a 0.2 kW prototype model was constructed in the laboratory, and the results were verified. [ABSTRACT FROM AUTHOR]