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The precipitation orientation of α phases in a Ti-55511 alloy during the heat treatment and the subsequent hot deformation is investigated. It is observed that the β grains prominently influence the precipitation process of α phases, i.e., for the neighbouring β grains without similar {110} pole, the precipitation orientation of grain boundary (GB) α phases (α GB) keep {0001}//{110}β1 or {0001}//{110}β2 to maintain the burgers orientation relationship (BOR) with the neighbouring β grains. For the neighbouring β grains with similar {110} pole, the {0001} poles of α GB phases precipitate parallel to this similar {110} pole and maintain BOR with the neighbouring β grains. For the neighbouring β grains with multiple similar {110} poles, the {0001} poles of α GB phases precipitate along these similar poles and maintain BOR with the neighbouring β grains. Besides, the interface instability nucleation and sympathetic nucleation are the main nucleation modes of the GB widmanstatten α phase (α WGB), which primarily precipitate within β grains with the similar orientation of α GB phases. The intracrystalline widmanstatten phase (α WI) precipitate in a single β grain with 12 precipitation orientations. The BOR between α WI phases and β grains is destroyed during the subsequent hot compressive deformation. • Precipitation orientation of α phases in Ti-55511 alloy during heat treatment plus hot deformation is studied. • Precipitation orientation of GB α phases maintains the BOR with neighbouring β grains along different poles. • Interface instability and sympathetic nucleations are the main nucleation modes of GB widmanstatten α phases. • The intracrystalline widmanstatten phases (α WI) precipitate in a single β grain with 12 precipitation orientations. • The BOR between α WI phases and β grains is destroyed during the subsequent hot compressive deformation. [ABSTRACT FROM AUTHOR]