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Abstract: During recent decades, Neanderthal diet has been a major research topic in palaeoanthropology. This has been accelerated by the maturation of different techniques, which have produced a plethora of new information. However, this proliferation of data has led to confusing and contradictory results. Furthermore, most of the ecological dietary studies have been carried out on specimens drawn from different time periods and regions, almost exclusively those characterized by cold, open environmental conditions. Subsistence models based on these fragmentary data have been applied to Neanderthals living in a variety of different regions and environments, even though their dietary strategies may have been as variable as regions they inhabited. In this paper we integrate different dietary approaches (studies of the zooarchaeology, stable isotopes and plant remains) from the central and southeastern Mediterranean coast of Iberia in order to develop a broader and more complex picture of Neanderthal diet in different Mediterranean environmental conditions. Our results suggest that there may have been some minor dietary variation due to climatic or environmental differences, but that Neanderthal diet focussed on large terrestrial game, supplemented by plant foods when these were available. [Copyright &y& Elsevier]

The resilience of Neanderthals towards changing climatic and environmental conditions, and especially towards severely cold climates in northern regions of central Europe, is still under debate. One way to address this is to investigate multi-layered occupation in different climatic intervals, using independently-compiled paleoenvironmental and chronological data. Unfortunately, most open-air sites on the northern European Plain lack a robust chronostratigraphy beyond the radiocarbon dating range, thereby often hampering direct links between human occupation and climate. Here we present the results of integrative research at the Middle Paleolithic open-air site of Lichtenberg, Northern Germany, comprising archaeology, luminescence dating, sedimentology, micromorphology, as well as pollen and phytolith analyses. Our findings clearly show Neanderthal presence in temperate, forested environments during the Mid-Eemian Interglacial, MIS 5e and the latest Brörup Interstadial, MIS 5c/GI 22 (Lichtenberg II). For the previously known occupation Lichtenberg I, we revise the chronology from the former early MIS 3 (57 ± 6 ka) to early MIS 4/GS 19 (71.3 ± 7.3 ka), with dominant cold steppe/tundra vegetation. The early MIS 4 occupation suggests that Neanderthals could adjust well to severely cold environments and implies recurring population in the region between MIS 5 and MIS 3. The artefact assemblages differ between the temperate and cold environment occupations regarding size, blank production, typology and tool use. We argue that this distinctness can partially be explained by different site functions and occupation duration, as well as the availability of large and high-quality flint raw material. Raw material availability is in turn governed by changing vegetation cover that hindered or fostered sediment redeposition as a provider of flint from the primary source of the glacial sediments nearby. • Integrative multidisciplinary approach of geo-biosciences and archaeology. • Precise chronology connects Neanderthal occupation to distinct climatic phases. • Neanderthals adapted to changing and also severely cold environments in northern Central Europe. • Variability of tool size, raw material and tool use in response to climate changes. • First numerical dates for early Weichselian interstadials in the type area. [ABSTRACT FROM AUTHOR]