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Competing Interests: The authors declare no conflicts of interest.

The Pliocene–Pleistocene Transition (PPT), from around 3.2 to 2.5 million years ago (Ma), represented a major shift in the climate system and was characterized by a gradual cooling trend and the appearance of large continental ice sheets over northern Eurasia and North America. Paleo evidence indicates that the PPT was accompanied and possibly caused by a decrease in atmospheric CO 2 , but the temporal resolution of CO 2 reconstructions is low for this period of time and uncertainties remain large. Therefore, instead of applying existent CO 2 reconstructions we solved an ‘inverse’ problem by finding a schematic CO 2 concentration scenario that allows us to simulate the temporal evolution of key climate characteristics in agreement with paleoclimate records. To this end, we performed an ensemble of transient simulations with an Earth system model of intermediate complexity from which we derived a best guess transient CO 2 scenario for the interval from 3.2 to 2.4 Ma that gives the best fit between the simulated and reconstructed benthic δ 18 O and global sea surface temperature evolution. Our data-constrained CO 2 scenarios are consistent with recent CO 2 reconstructions and suggest a gradual CO 2 decline from 375–425 to 275–300 ppm, between 3.2 and 2.4 Ma. In addition to a gradual decline, the best fit to paleoclimate data requires the existence of pronounced CO 2 variability coherent with the 41-kyr (1 kyr = 1000 years) obliquity cycle. In our simulations the long-term CO 2 decline is accompanied by a relatively abrupt intensification of Northern Hemisphere glaciation at around 2.7 Ma. This is the result of a threshold behaviour of the ice sheets response to gradual CO 2 decrease and orbital forcing. The simulated Northern Hemisphere ice sheets during the early Pleistocene glacial cycles reach a maximum volume equivalent to a sea level drop of about 40 m. Both ice volume and benthic δ 18 O are dominated by 41-kyr cyclicity. Our simulations suggest that before 2.7 Ma Greenland was ice free during summer insolation maxima and only partly ice covered during periods of minimum summer insolation. A fully glaciated Greenland comparable to its present-day ice volume is modelled only during glacial maxima after 2.7 Ma and more continuously after 2.5 Ma. [ABSTRACT FROM AUTHOR]

Changes in ocean circulation have been proposed as a trigger mechanism for the large coupled climate and carbon cycle perturbations at the Paleocene-Eocene Thermal Maximum (PETM, ca. 55 Ma). An abrupt warming of oceanic intermediate waters could have initiated the thermal destabilization of sediment-hosted methane gas hydrates and potentially triggered sediment slumps and slides. In an ensemble of fully coupled atmosphere-ocean general circulation model (AOGCM) simulations of the late Paleocene and early Eocene, we identify such a circulation-driven enhanced intermediate-water warming. Critically, we find an approximate twofold amplification of Atlantic intermediate-water warming when CO2 levels are doubled from 2× to 4× preindustrial CO2 compared to when they are doubled from 1× to 2×. This warming is largely focused on the equatorial and South Atlantic and is driven by a significant reduction in deep-water formation from the Southern Ocean. This scenario is consistent with altered PETM circulation patterns inferred from benthic carbon isotope data and the intensity of deep-sea carbonate dissolution in the South Atlantic. The linkage between intermediate-water warming and gas hydrate destabilization could provide an important feedback in the establishment of peak PETM warmth. [ABSTRACT FROM AUTHOR]